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<document ID-DOI="10.1590/S1679-62252003000200002" ID-ISSN="1982-0224" ID-Zenodo-Dep="4647103" LSID="urn:lsid:zoobank.org:act:4947C013-E9B9-47A6-87F4-429C67BE7B1D" approvalRequired="8" approvalRequired_for_textStreams="8" checkinTime="1617110689260" checkinUser="carolina" docAuthor="Britto, Marcelo R. &amp; Lima, Flávio C. T." docDate="2003" docId="03F6EE50E02F762710ACFA3EFA32C769" docLanguage="en" docName="NeotropIchthyol.1.2.83-91.pdf" docOrigin="Neotropical Ichthyology 1 (2)" docSource="http://www.scielo.br/scielo.php?script=sci_arttext&amp;pid=S1679-62252003000200002&amp;lng=en&amp;tlng=en" docStyle="DocumentStyle:85A40B78B715C4497F837D9D78407CE5.3:NeotropIchthyol.2003-2013.journal_article" docStyleId="85A40B78B715C4497F837D9D78407CE5" docStyleName="NeotropIchthyol.2003-2013.journal_article" docStyleVersion="3" docTitle="Corydoras tukano Britto &amp; Lima 2003, new species" docType="treatment" docUuid="4947C013-E9B9-47A6-87F4-429C67BE7B1D" docUuidSource="ZooBank" docVersion="2" lastPageId="5" lastPageNumber="88" masterDocId="FFCF9628E02E76221379FFFDFF98C150" masterDocTitle="Corydoras tukano, a new species of corydoradine catfish from the rio Tiquié, upper rio Negro basin, Brazil (Ostariophysi: Siluriformes: Callichthyidae)" masterLastPageNumber="91" masterPageNumber="83" pageId="1" pageNumber="84" updateTime="1617111093233" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Corydoras tukano, a new species of corydoradine catfish from the rio Tiquié, upper rio Negro basin, Brazil (Ostariophysi: Siluriformes: Callichthyidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Britto, Marcelo R.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lima, Flávio C. T.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Neotropical Ichthyology</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2003</mods:date>
<mods:detail type="pubDate">
<mods:number>2003-12-31</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>83</mods:start>
<mods:end>91</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://www.scielo.br/scielo.php?script=sci_arttext&amp;pid=S1679-62252003000200002&amp;lng=en&amp;tlng=en</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1590/S1679-62252003000200002</mods:identifier>
<mods:identifier type="ISSN">1982-0224</mods:identifier>
<mods:identifier type="Zenodo-Dep">4647103</mods:identifier>
<mods:identifier type="ZooBank">BBA084B6-1A25-4780-AD28-C074290DC6A1</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:zoobank.org:act:4947C013-E9B9-47A6-87F4-429C67BE7B1D" httpUri="http://treatment.plazi.org/id/03F6EE50E02F762710ACFA3EFA32C769" lastPageId="5" lastPageNumber="88" pageId="1" pageNumber="84">
<subSubSection box="[981,1325,1475,1499]" pageId="1" pageNumber="84" type="nomenclature">
<paragraph blockId="1.[981,1325,1475,1531]" box="[981,1325,1475,1499]" pageId="1" pageNumber="84">
<heading box="[981,1325,1475,1499]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="84" reason="2">
<emphasis bold="true" box="[981,1325,1475,1499]" pageId="1" pageNumber="84">
<taxonomicName authority="Britto &amp; Lima, 2003" authorityName="Britto &amp; Lima" authorityYear="2003" box="[981,1179,1475,1499]" class="Actinopterygii" family="Callichthyidae" genus="Corydoras" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="84" phylum="Chordata" rank="species" species="tukano" status="new species">
<emphasis bold="true" box="[981,1179,1475,1499]" italics="true" pageId="1" pageNumber="84">Corydoras tukano</emphasis>
</taxonomicName>
,
<taxonomicNameLabel box="[1193,1325,1475,1499]" pageId="1" pageNumber="84" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[1107,1198,1508,1531]" pageId="1" pageNumber="84" type="description">
<paragraph blockId="1.[981,1325,1475,1531]" box="[1107,1198,1508,1531]" pageId="1" pageNumber="84">
<figureCitation box="[1107,1198,1508,1531]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="1.[151,190,744,768]" captionStartId-1="2.[298,337,716,740]" captionStartId-2="2.[298,337,1411,1435]" captionStartId-3="3.[253,292,718,742]" captionTargetBox-0="[283,1375,203,690]" captionTargetBox-1="[299,1359,191,692]" captionTargetBox-2="[299,1359,803,1387]" captionTargetBox-3="[222,1417,191,691]" captionTargetId-0="figure-603@1.[243,1387,159,723]" captionTargetId-1="figure-365@2.[299,1359,191,692]" captionTargetId-2="figure-366@2.[299,1359,803,1387]" captionTargetId-3="figure-488@3.[222,1417,191,691]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="2" captionTargetPageId-3="3" captionText-0="Fig. 1. Corydoras tukano, holotype, MZUSP 82100, presumably a female, 40.9 mm SL; Brazil, Amazonas state, rio Tiquié, comunidade de Caruru. Caudal-fin margin reconstructed from MZUSP 81194, 37.4 mm SL." captionText-1="Fig. 2. Corydoras tukano, holotype, MZUSP 82100, presumably a female, 40.9 mm SL; Brazil, estado" captionText-2="Fig. 3. Corydoras tukano, paratype, MZUSP 81179, presumably a mature male, 36.1 mm SL; Brazil," captionText-3="Fig. 4. Dorsal (left) and ventral (right) views of head and pectoral fins of Corydoras tukano, holotype, MZUSP 82100, 40.9 mm SL; Brazil, estado do Amazonas, rio Tiquié, comunidade de Caruru." figureDoi-0="http://doi.org/10.5281/zenodo.4647105" figureDoi-1="http://doi.org/10.5281/zenodo.4647107" figureDoi-2="http://doi.org/10.5281/zenodo.4647109" figureDoi-3="http://doi.org/10.5281/zenodo.4647111" httpUri-0="https://zenodo.org/record/4647105/files/figure.png" httpUri-1="https://zenodo.org/record/4647107/files/figure.png" httpUri-2="https://zenodo.org/record/4647109/files/figure.png" httpUri-3="https://zenodo.org/record/4647111/files/figure.png" pageId="1" pageNumber="84">Figs. 1-4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection lastPageId="2" lastPageNumber="85" pageId="1" pageNumber="84" type="materials_examined">
<paragraph blockId="1.[832,1475,1571,1979]" pageId="1" pageNumber="84">
<materialsCitation collectingDate="2002-10-20" collectingDateMax="2002-10-21" collectingDateMin="2002-10-20" collectionCode="MZUSP, UTM" collectorName="de Caruru &amp; Flavio C. T. Lima" country="Brazil" latitude="0.27469447" location="rio Tiquie" longLatPrecision="1" longitude="-69.914894" pageId="1" pageNumber="84" specimenCode="MZUSP 82100, UTM 19" specimenCount="1" stateProvince="Amazonas" typeStatus="holotype">
<emphasis bold="true" box="[832,942,1571,1595]" pageId="1" pageNumber="84">
<typeStatus box="[832,938,1571,1595]" pageId="1" pageNumber="84">Holotype</typeStatus>
.
</emphasis>
<specimenCode box="[950,1114,1572,1595]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="84">MZUSP 82100</specimenCode>
(
<quantity box="[1130,1228,1572,1595]" metricMagnitude="-2" metricUnit="m" metricValue="4.09" pageId="1" pageNumber="84" unit="mm" value="40.9">40.9 mm</quantity>
SL),
<collectingCountry box="[1291,1359,1572,1595]" name="Brazil" pageId="1" pageNumber="84">Brazil</collectingCountry>
,
<collectingRegion country="Brazil" name="Amazonas" pageId="1" pageNumber="84">estado do Amazonas</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:03F6EE50E02F762710ACFA3EFA32C769:8E80099DE02F762310C3F9B9FBBBC70B" box="[954,1059,1604,1627]" country="Brazil" latitude="0.27469447" longLatPrecision="1" longitude="-69.914894" name="rio Tiquie" pageId="1" pageNumber="84" stateProvince="Amazonas">rio Tiquié</location>
, comunidade
<collectorName box="[1201,1307,1604,1627]" pageId="1" pageNumber="84">de Caruru</collectorName>
, beaches in pool below the fall,
<geoCoordinate box="[975,1106,1636,1659]" degrees="0" direction="north" minutes="16" orientation="latitude" pageId="1" pageNumber="84" precision="1" seconds="28.9" value="0.27469447">0°1628.9&quot;N</geoCoordinate>
,
<geoCoordinate box="[1114,1261,1636,1659]" degrees="69" direction="west" minutes="54" orientation="longitude" pageId="1" pageNumber="84" precision="1" seconds="53.6" value="-69.914894">69°5453.6&quot;W</geoCoordinate>
(
<specimenCode box="[1272,1357,1636,1659]" collectionCode="UTM" pageId="1" pageNumber="84">UTM 19</specimenCode>
N0398191/ 0030366);
<collectorName box="[942,1116,1668,1691]" pageId="1" pageNumber="84">Flávio C.T. Lima</collectorName>
<emphasis box="[1120,1163,1668,1691]" italics="true" pageId="1" pageNumber="84">et al</emphasis>
.,
<date box="[1181,1342,1668,1691]" pageId="1" pageNumber="84" value="2002-10-20" valueMax="2002-10-21" valueMin="2002-10-20">
<collectingDate box="[1181,1342,1668,1691]" pageId="1" pageNumber="84" value="2002-10-20" valueMax="2002-10-21" valueMin="2002-10-20">20-21 Oct 2002</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph blockId="1.[832,1475,1571,1979]" pageId="1" pageNumber="84">
<materialsCitation collectingDate="2000-10-23" collectingDateMax="2000-11-29" collectingDateMin="2000-10-23" collectionCode="MZUSP, CAS, ANSP, UTM" collectorName="de Boca do Sal &amp; de Cunuri &amp; M. L. Lopes" country="Brazil" county="All" latitude="0.20638889" location="Boca do Sal" longLatPrecision="21" longitude="-69.37444" municipality="rio Tiquie" pageId="1" pageNumber="84" specimenCode="MZUSP 81194, CAS 217960, MZUSP 64096, MZUSP 65689, ANSP 179201, UTM 19, MZUSP 81179" specimenCount="1" stateProvince="Amazonas" typeStatus="paratype">
<emphasis bold="true" box="[832,953,1699,1723]" pageId="1" pageNumber="84">
<typeStatus box="[832,948,1699,1723]" pageId="1" pageNumber="84">Paratypes</typeStatus>
.
</emphasis>
<collectingCounty box="[961,995,1700,1723]" pageId="1" pageNumber="84">All</collectingCounty>
from
<collectingCountry box="[1064,1132,1700,1723]" name="Brazil" pageId="1" pageNumber="84">Brazil</collectingCountry>
,
<collectingRegion box="[1145,1374,1700,1723]" country="Brazil" name="Amazonas" pageId="1" pageNumber="84">estado do Amazonas</collectingRegion>
:
<specimenCode collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="84">MZUSP 81194</specimenCode>
(5,
<quantity box="[929,1068,1732,1755]" metricMagnitude="-2" metricUnit="m" metricValue="3.7399999999999998" metricValueMax="4.029999999999999" metricValueMin="3.45" pageId="1" pageNumber="84" unit="mm" value="37.4" valueMax="40.3" valueMin="34.5">34.5-40.3 mm</quantity>
SL);
<specimenCode box="[1119,1249,1732,1755]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="1" pageNumber="84" type="Museum">CAS 217960</specimenCode>
(1,
<quantity box="[1282,1371,1732,1755]" metricMagnitude="-2" metricUnit="m" metricValue="3.46" pageId="1" pageNumber="84" unit="mm" value="34.6">34.6 mm</quantity>
SL); same data as holotype.
<specimenCode box="[1022,1186,1764,1787]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="84">MZUSP 64096</specimenCode>
(4, 28.2-38.0 mm SL),
<collectingMunicipality pageId="1" pageNumber="84">rio Tiquié</collectingMunicipality>
, comunidade
<collectorName box="[1058,1230,1796,1819]" pageId="1" pageNumber="84">
de
<location LSID="urn:lsid:plazi:treatment:03F6EE50E02F762710ACFA3EFA32C769:8E80099DE02F7623173DF8F9FB56C64B" box="[1092,1230,1796,1819]" country="Brazil" county="All" latitude="0.20638889" longLatPrecision="21" longitude="-69.37444" municipality="rio Tiquie" name="Boca do Sal" pageId="1" pageNumber="84" stateProvince="Amazonas">Boca do Sal</location>
</collectorName>
;
<location LSID="urn:lsid:plazi:treatment:03F6EE50E02F762710ACFA3EFA32C769:8E80099DE02F762317A5F8F9FABDC64B" box="[1244,1317,1796,1819]" country="Brazil" county="All" latitude="0.20638889" longLatPrecision="21" longitude="-69.37444" municipality="rio Tiquie" name="Rafael" pageId="1" pageNumber="84" stateProvince="Amazonas">Rafael</location>
,
<date box="[1330,1471,1796,1819]" pageId="1" pageNumber="84" value="2000-10-23">
<collectingDate box="[1330,1471,1796,1819]" pageId="1" pageNumber="84" value="2000-10-23">23 Oct 2000</collectingDate>
</date>
.
<specimenCode box="[832,983,1828,1851]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="84">MZUSP 65689</specimenCode>
(2,
<quantity box="[1017,1157,1828,1851]" metricMagnitude="-2" metricUnit="m" metricValue="3.585" metricValueMax="3.85" metricValueMin="3.32" pageId="1" pageNumber="84" unit="mm" value="35.85" valueMax="38.5" valueMin="33.2">33.2-38.5 mm</quantity>
SL);
<specimenCode box="[1208,1353,1828,1851]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="1" pageNumber="84">ANSP 179201</specimenCode>
(1,
<quantity box="[1386,1475,1828,1851]" metricMagnitude="-2" metricUnit="m" metricValue="3.5799999999999996" pageId="1" pageNumber="84" unit="mm" value="35.8">35.8 mm</quantity>
SL);
<location LSID="urn:lsid:plazi:treatment:03F6EE50E02F762710ACFA3EFA32C769:8E80099DE02F7623100BF8B9FC41C60B" box="[882,985,1860,1883]" country="Brazil" county="All" latitude="0.20638889" longLatPrecision="21" longitude="-69.37444" municipality="rio Tiquie" name="rio Tiquie" pageId="1" pageNumber="84" stateProvince="Amazonas">rio Tiquié</location>
, one hour by boat below comunidade
<collectorName box="[1366,1470,1860,1883]" pageId="1" pageNumber="84">de Cunuri</collectorName>
, below
<location LSID="urn:lsid:plazi:treatment:03F6EE50E02F762710ACFA3EFA32C769:8E80099DE02F762310FCF899FBF0C62B" box="[901,1128,1892,1915]" country="Brazil" county="All" latitude="0.20638889" longLatPrecision="21" longitude="-69.37444" municipality="rio Tiquie" name="Cachoeira do Tucano" pageId="1" pageNumber="84" stateProvince="Amazonas">Cachoeira do Tucano</location>
,
<geoCoordinate box="[1137,1255,1892,1915]" degrees="0" direction="north" minutes="12" orientation="latitude" pageId="1" pageNumber="84" precision="15" seconds="23" value="0.20638889">0°1223”N</geoCoordinate>
,
<geoCoordinate box="[1265,1401,1892,1915]" degrees="69" direction="west" minutes="22" orientation="longitude" pageId="1" pageNumber="84" precision="15" seconds="28" value="-69.37444">69°2228”W</geoCoordinate>
(
<specimenCode collectionCode="UTM" pageId="1" pageNumber="84">UTM 19</specimenCode>
N458666/22437; coordinates are from Cunuri);
<collectorName box="[1343,1470,1924,1947]" pageId="1" pageNumber="84">M. L. Lopes</collectorName>
,
<date box="[832,967,1956,1979]" pageId="1" pageNumber="84" value="2000-11-29">
<collectingDate box="[832,967,1956,1979]" pageId="1" pageNumber="84" value="2000-11-29">29 Nov 2000</collectingDate>
</date>
.
<specimenCode box="[974,1131,1956,1979]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="84">MZUSP 81179</specimenCode>
(2, 34.0-
<quantity box="[1222,1312,1956,1979]" metricMagnitude="-2" metricUnit="m" metricValue="3.61" pageId="1" pageNumber="84" unit="mm" value="36.1">36.1 mm</quantity>
SL),
<location LSID="urn:lsid:plazi:treatment:03F6EE50E02F762710ACFA3EFA32C769:8E80099DE02F7623162FF859FA25C6EB" box="[1366,1469,1956,1979]" country="Brazil" county="All" latitude="0.20638889" longLatPrecision="21" longitude="-69.37444" municipality="rio Tiquie" name="rio Tiquie" pageId="1" pageNumber="84" stateProvince="Amazonas">rio Tiquié</location>
</materialsCitation>
,
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4647107" ID-Zenodo-Dep="4647107" box="[298,1356,716,740]" httpUri="https://zenodo.org/record/4647107/files/figure.png" pageId="2" pageNumber="85" startId="2.[298,337,716,740]" targetBox="[299,1359,191,692]" targetPageId="2">
<paragraph blockId="2.[298,1356,716,772]" box="[298,1356,716,740]" pageId="2" pageNumber="85">
<emphasis bold="true" box="[298,366,716,740]" pageId="2" pageNumber="85">Fig. 2.</emphasis>
<taxonomicName authorityName="Britto &amp; Lima" authorityYear="2003" box="[372,561,717,740]" class="Actinopterygii" family="Callichthyidae" genus="Corydoras" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="85" phylum="Chordata" rank="species" species="tukano">
<emphasis box="[372,561,717,740]" italics="true" pageId="2" pageNumber="85">Corydoras tukano</emphasis>
</taxonomicName>
, holotype, MZUSP 82100, presumably a female, 40.9 mm SL; Brazil, estado
</paragraph>
</caption>
<paragraph blockId="2.[298,1356,716,772]" box="[298,823,749,772]" pageId="2" pageNumber="85">
<materialsCitation box="[298,819,749,772]" collectorName="de Caruru" location="rio Tiquie" pageId="2" pageNumber="85" specimenCount="1" stateProvince="Amazonas">
do
<collectingRegion box="[331,447,749,772]" country="Venezuela" name="Amazonas" pageId="2" pageNumber="85">Amazonas</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:03F6EE50E02F762710ACFA3EFA32C769:8E80099DE02C762012B1FD10FDADC254" box="[456,565,749,772]" name="rio Tiquie" pageId="2" pageNumber="85" stateProvince="Amazonas">rio Tiquié</location>
, comunidade
<collectorName box="[710,819,749,772]" pageId="2" pageNumber="85">de Caruru</collectorName>
</materialsCitation>
.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4647109" ID-Zenodo-Dep="4647109" box="[298,1359,1411,1435]" httpUri="https://zenodo.org/record/4647109/files/figure.png" pageId="2" pageNumber="85" startId="2.[298,337,1411,1435]" targetBox="[299,1359,803,1387]" targetPageId="2">
<paragraph blockId="2.[298,1359,1411,1467]" box="[298,1359,1411,1435]" pageId="2" pageNumber="85">
<emphasis bold="true" box="[298,367,1411,1435]" pageId="2" pageNumber="85">Fig. 3.</emphasis>
<taxonomicName authorityName="Britto &amp; Lima" authorityYear="2003" box="[373,565,1412,1435]" class="Actinopterygii" family="Callichthyidae" genus="Corydoras" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="85" phylum="Chordata" rank="species" species="tukano">
<emphasis box="[373,565,1412,1435]" italics="true" pageId="2" pageNumber="85">Corydoras tukano</emphasis>
</taxonomicName>
, paratype, MZUSP 81179, presumably a mature male, 36.1 mm SL; Brazil,
</paragraph>
</caption>
<paragraph blockId="2.[298,1359,1411,1467]" box="[298,957,1444,1467]" pageId="2" pageNumber="85">
<materialsCitation box="[298,952,1444,1467]" collectorName="de Boca do Sal" location="Boca do Sal" municipality="rio Tiquie" pageId="2" pageNumber="85" specimenCount="1" stateProvince="Amazonas">
<collectingRegion box="[298,523,1444,1467]" country="Brazil" name="Amazonas" pageId="2" pageNumber="85">estado do Amazonas</collectingRegion>
,
<collectingMunicipality box="[533,642,1444,1467]" pageId="2" pageNumber="85">rio Tiquié</collectingMunicipality>
, comunidade
<collectorName box="[789,952,1444,1467]" pageId="2" pageNumber="85">
de
<location LSID="urn:lsid:plazi:treatment:03F6EE50E02F762710ACFA3EFA32C769:8E80099DE02C7620104CFA59FC20C4EB" box="[821,952,1444,1467]" municipality="rio Tiquie" name="Boca do Sal" pageId="2" pageNumber="85" stateProvince="Amazonas">Boca do Sal</location>
</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph blockId="2.[159,802,1512,1983]" pageId="2" pageNumber="85">
comunidade de Boca do Sal,
<geoCoordinate box="[448,586,1512,1535]" degrees="0" direction="north" minutes="16" orientation="latitude" pageId="2" pageNumber="85" precision="1" seconds="22.5" value="0.27291667">0°1622.5” N</geoCoordinate>
,
<geoCoordinate box="[594,731,1512,1535]" degrees="69" direction="west" minutes="54" orientation="longitude" pageId="2" pageNumber="85" precision="15" seconds="03" value="-69.90083">69°5403” W</geoCoordinate>
(
<collectionCode box="[742,802,1512,1535]" pageId="2" pageNumber="85">UTM</collectionCode>
19N0399757/0030167); N. P. Marques,
<date box="[575,705,1544,1567]" pageId="2" pageNumber="85" value="2002-10-25">25 Oct 2002</date>
.
<materialsCitation collectionCode="MZUSP" pageId="2" pageNumber="85" specimenCode="MZUSP 81244" specimenCount="1">
<specimenCode collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="2" pageNumber="85">MZUSP 81244</specimenCode>
(16,20.3-41.4 mmSL)
</materialsCitation>
;
<materialsCitation box="[457,796,1576,1599]" collectionCode="INPA" pageId="2" pageNumber="85" specimenCode="INPA 21423" specimenCount="1">
<specimenCode box="[457,581,1576,1599]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="2" pageNumber="85">INPA 21423</specimenCode>
(2,25.6-36.0mmSL)
</materialsCitation>
;
<materialsCitation collectingDate="2002-10-29" collectionCode="MNRJ, UTM" collectorName="Tarcisio &amp; N. P. Marques" latitude="0.28311113" location="Coracao de Maria" longLatPrecision="1" longitude="-69.8345" municipality="Cabari" pageId="2" pageNumber="85" specimenCode="MNRJ 25355, UTM 19" specimenCount="1">
<specimenCode box="[159,295,1608,1631]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="2" pageNumber="85">MNRJ 25355</specimenCode>
(2, 24.4-32.0 mm SL); igarapé
<collectingMunicipality box="[600,670,1608,1631]" pageId="2" pageNumber="85">Cabari</collectingMunicipality>
, comunidade de
<location LSID="urn:lsid:plazi:treatment:03F6EE50E02F762710ACFA3EFA32C769:8E80099DE02C762013B9F995FE1DC72F" box="[192,389,1640,1663]" latitude="0.28311113" longLatPrecision="1" longitude="-69.8345" municipality="Cabari" name="Coracao de Maria" pageId="2" pageNumber="85">Coração de Maria</location>
,
<geoCoordinate box="[400,549,1640,1663]" degrees="0" direction="north" minutes="16" orientation="latitude" pageId="2" pageNumber="85" precision="1" seconds="59.2" value="0.28311113">0°1659.2” N</geoCoordinate>
,
<geoCoordinate box="[560,725,1640,1663]" degrees="69" direction="west" minutes="50" orientation="longitude" pageId="2" pageNumber="85" precision="1" seconds="04.2" value="-69.8345">69°5004.2” W</geoCoordinate>
(
<specimenCode collectionCode="UTM" pageId="2" pageNumber="85">UTM 19</specimenCode>
N0407140/ 0031302);
<collectorName box="[412,494,1672,1695]" pageId="2" pageNumber="85">Tarcísio</collectorName>
&amp;
<collectorName box="[522,665,1672,1695]" pageId="2" pageNumber="85">N. P. Marques</collectorName>
,
<date box="[673,799,1672,1695]" pageId="2" pageNumber="85" value="2002-10-29">
<collectingDate box="[673,799,1672,1695]" pageId="2" pageNumber="85" value="2002-10-29">29 Oct 2002</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation collectingDate="2002-11-07" collectionCode="MZUSP, UTM" latitude="0.27291667" location="Boca do Sal" longLatPrecision="15" longitude="-69.90083" municipality="rio Tiquie" pageId="2" pageNumber="85" specimenCode="MZUSP 81276, UTM 19" specimenCount="1" stateProvince="Lima">
<specimenCode box="[159,318,1704,1727]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="2" pageNumber="85">MZUSP 81276</specimenCode>
(5,
<quantity box="[356,502,1704,1727]" metricMagnitude="-2" metricUnit="m" metricValue="2.3049999999999997" metricValueMax="2.62" metricValueMin="1.9899999999999998" pageId="2" pageNumber="85" unit="mm" value="23.049999999999997" valueMax="26.2" valueMin="19.9">19.9-26.2 mm</quantity>
SL),
<collectingMunicipality box="[558,663,1704,1727]" pageId="2" pageNumber="85">rio Tiquié</collectingMunicipality>
, comunidade de
<location LSID="urn:lsid:plazi:treatment:03F6EE50E02F762710ACFA3EFA32C769:8E80099DE02C762013C2F935FEABC78F" box="[187,307,1736,1759]" latitude="0.27291667" longLatPrecision="15" longitude="-69.90083" municipality="rio Tiquie" name="Boca do Sal" pageId="2" pageNumber="85" stateProvince="Lima">Boca do Sal</location>
,
<geoCoordinate box="[315,451,1736,1759]" degrees="0" direction="north" minutes="16" orientation="latitude" pageId="2" pageNumber="85" precision="1" seconds="22.5" value="0.27291667">0°1622.5” N</geoCoordinate>
,
<geoCoordinate box="[458,589,1736,1759]" degrees="69" direction="west" minutes="54" orientation="longitude" pageId="2" pageNumber="85" precision="15" seconds="03" value="-69.90083">69°5403”W</geoCoordinate>
(
<specimenCode box="[599,684,1736,1759]" collectionCode="UTM" pageId="2" pageNumber="85">UTM 19</specimenCode>
N0399757/ 0030167); F. C. T.
<collectingRegion box="[352,408,1768,1791]" country="Peru" name="Lima" pageId="2" pageNumber="85">Lima</collectingRegion>
<emphasis box="[413,458,1769,1792]" italics="true" pageId="2" pageNumber="85">et al</emphasis>
.,
<date box="[476,600,1768,1791]" pageId="2" pageNumber="85" value="2002-11-07">
<collectingDate box="[476,600,1768,1791]" pageId="2" pageNumber="85" value="2002-11-07">7 Nov 2002</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation collectionCode="MZUSP" collectorName="de Caruru &amp; M. L. Lopes" latitude="0.26666668" location="Boca de Sal" longLatPrecision="1308" longitude="-69.9" municipality="rio Tiquie" pageId="2" pageNumber="85" specimenCode="MZUSP 81153" specimenCount="1" stateProvince="Lima">
<specimenCode box="[609,767,1768,1791]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="2" pageNumber="85">MZUSP 81153</specimenCode>
(9,
<quantity box="[159,304,1800,1823]" metricMagnitude="-2" metricUnit="m" metricValue="3.63" metricValueMax="3.72" metricValueMin="3.54" pageId="2" pageNumber="85" unit="mm" value="36.3" valueMax="37.2" valueMin="35.4">35.4-37.2 mm</quantity>
SL, 6 cs,
<quantity box="[401,547,1800,1823]" metricMagnitude="-2" metricUnit="m" metricValue="2.96" metricValueMax="3.8299999999999996" metricValueMin="2.09" pageId="2" pageNumber="85" unit="mm" value="29.599999999999998" valueMax="38.3" valueMin="20.9">20.9-38.3 mm</quantity>
SL),
<collectingMunicipality box="[602,706,1800,1823]" pageId="2" pageNumber="85">rio Tiquié</collectingMunicipality>
, between the comunidades
<collectorName box="[335,436,1832,1855]" pageId="2" pageNumber="85">de Caruru</collectorName>
and
<location LSID="urn:lsid:plazi:treatment:03F6EE50E02F762710ACFA3EFA32C769:8E80099DE02C7620129BF8D5FDC4C66F" box="[482,604,1832,1855]" latitude="0.26666668" longLatPrecision="1308" longitude="-69.9" municipality="rio Tiquie" name="Boca de Sal" pageId="2" pageNumber="85" stateProvince="Lima">Boca de Sal</location>
,
<geoCoordinate box="[612,690,1832,1855]" degrees="0" direction="north" minutes="16" orientation="latitude" pageId="2" pageNumber="85" precision="925" value="0.26666668">0°16N</geoCoordinate>
,
<geoCoordinate box="[699,797,1832,1855]" degrees="69" direction="west" minutes="54" orientation="longitude" pageId="2" pageNumber="85" precision="925" value="-69.9">69°54W</geoCoordinate>
;
<collectorName box="[159,282,1864,1887]" pageId="2" pageNumber="85">M. L. Lopes</collectorName>
<emphasis box="[286,329,1865,1888]" italics="true" pageId="2" pageNumber="85">et al</emphasis>
., 2001-2002
</materialsCitation>
.
<materialsCitation collectingDate="2002-11-07" collectionCode="MZUSP" collectorName="de Sao Domingos Savio &amp; Jabuti" location="rio Tiquie" pageId="2" pageNumber="85" specimenCode="MZUSP 81216" specimenCount="1" stateProvince="Lima">
<specimenCode box="[467,623,1864,1887]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="2" pageNumber="85">MZUSP 81216</specimenCode>
(2,
<quantity box="[658,801,1864,1887]" metricMagnitude="-2" metricUnit="m" metricValue="1.92" metricValueMax="2.13" metricValueMin="1.71" pageId="2" pageNumber="85" unit="mm" value="19.2" valueMax="21.3" valueMin="17.1">17.1-21.3 mm</quantity>
SL),
<location LSID="urn:lsid:plazi:treatment:03F6EE50E02F762710ACFA3EFA32C769:8E80099DE02C762013ADF895FED9C62F" box="[212,321,1896,1919]" name="rio Tiquie" pageId="2" pageNumber="85" stateProvince="Lima">rio Tiquié</location>
, between the comunidades
<collectorName pageId="2" pageNumber="85">de São Domingos Sávio</collectorName>
and
<collectorName box="[264,328,1928,1951]" pageId="2" pageNumber="85">Jabuti</collectorName>
, 0°15-16N, 69°51-54W; F.C.T.
<collectingRegion box="[667,722,1928,1951]" country="Peru" name="Lima" pageId="2" pageNumber="85">Lima</collectingRegion>
<emphasis box="[727,770,1929,1952]" italics="true" pageId="2" pageNumber="85">et al</emphasis>
.,
<date pageId="2" pageNumber="85" value="2002-11-07">
<collectingDate pageId="2" pageNumber="85" value="2002-11-07">7 Nov 2002</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="3" lastPageNumber="86" pageId="2" pageNumber="85" type="diagnosis">
<paragraph blockId="2.[839,1483,1512,1983]" lastBlockId="3.[151,1275,115,139]" lastPageId="3" lastPageNumber="86" pageId="2" pageNumber="85">
<emphasis bold="true" box="[839,950,1512,1536]" pageId="2" pageNumber="85">Diagnosis.</emphasis>
<taxonomicName authorityName="Britto &amp; Lima" authorityYear="2003" box="[957,1144,1513,1536]" class="Actinopterygii" family="Callichthyidae" genus="Corydoras" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="85" phylum="Chordata" rank="species" species="tukano">
<emphasis box="[957,1144,1513,1536]" italics="true" pageId="2" pageNumber="85">Corydoras tukano</emphasis>
</taxonomicName>
can be distinguished from all its congeners, except
<taxonomicName authority="Myers &amp; Weitzman" authorityName="Myers &amp; Weitzman" authorityYear="1960" box="[1042,1399,1544,1568]" class="Actinopterygii" family="Callichthyidae" genus="Corydoras" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="85" phylum="Chordata" rank="species" species="reynoldsi">
<emphasis box="[1042,1174,1545,1568]" italics="true" pageId="2" pageNumber="85">C. reynoldsi</emphasis>
Myers &amp; Weitzman
</taxonomicName>
and
<taxonomicName authority="Nijssen" authorityName="Nijssen" authorityYear="1971" class="Actinopterygii" family="Callichthyidae" genus="Corydoras" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="85" phylum="Chordata" rank="species" species="weitzmani">
<emphasis italics="true" pageId="2" pageNumber="85">C. weitzmani</emphasis>
Nijssen
</taxonomicName>
, by its color pattern consisting of a light ground coloration, with three dark, large, roughly rectangular blotches: the first one (“mask”) on the head, across the eye; the second one on the trunk at the level of dorsal fin; and the third one on the trunk at the level of the adipose fin. It can be distinguished from
<taxonomicName authorityName="Myers &amp; Weitzman" authorityYear="1960" box="[1047,1176,1737,1760]" class="Actinopterygii" family="Callichthyidae" genus="Corydoras" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="85" phylum="Chordata" rank="species" species="reynoldsi">
<emphasis box="[1047,1176,1737,1760]" italics="true" pageId="2" pageNumber="85">C. reynoldsi</emphasis>
</taxonomicName>
by the larger dark blotches, which in
<taxonomicName authorityName="Britto &amp; Lima" authorityYear="2003" box="[933,1031,1769,1792]" class="Actinopterygii" family="Callichthyidae" genus="Corydoras" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="85" phylum="Chordata" rank="species" species="tukano">
<emphasis box="[933,1031,1769,1792]" italics="true" pageId="2" pageNumber="85">C. tukano</emphasis>
</taxonomicName>
are broad, with the trunk blotches extending across 5-6 series of lateral plates. In
<taxonomicName authorityName="Myers &amp; Weitzman" authorityYear="1960" box="[1208,1332,1801,1824]" class="Actinopterygii" family="Callichthyidae" genus="Corydoras" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="85" phylum="Chordata" rank="species" species="reynoldsi">
<emphasis box="[1208,1332,1801,1824]" italics="true" pageId="2" pageNumber="85">C. reynoldsi</emphasis>
</taxonomicName>
, the first trunk blotch (
<emphasis box="[919,950,1833,1856]" italics="true" pageId="2" pageNumber="85">i.e.</emphasis>
, at the level of dorsal fin), is a discrete, verticallyelongated bar, and the second trunk blotch extends only across 2-3 series of lateral plates (see Myers &amp; Weitzman, 1960: fig. 2; Nijssen &amp; Isbrücker, 1983a: fig.10). See “Discussion” for additional evidence for the recognition of both as distinct species. 86
<taxonomicName authorityName="Britto &amp; Lima" authorityYear="2003" box="[358,553,116,139]" class="Actinopterygii" family="Callichthyidae" genus="Corydoras" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="86" phylum="Chordata" rank="species" species="tukano">
<emphasis box="[358,553,116,139]" italics="true" pageId="3" pageNumber="86">Corydoras tukano</emphasis>
</taxonomicName>
, a new species of corydoradine catfish from upper rio Negro basin
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4647111" ID-Zenodo-Dep="4647111" httpUri="https://zenodo.org/record/4647111/files/figure.png" pageId="3" pageNumber="86" startId="3.[253,292,718,742]" targetBox="[222,1417,191,691]" targetPageId="3">
<paragraph blockId="3.[253,1396,718,773]" pageId="3" pageNumber="86">
<emphasis bold="true" box="[253,327,718,742]" pageId="3" pageNumber="86">Fig. 4.</emphasis>
Dorsal (left) and ventral (right) views of head and pectoral fins of
<taxonomicName authorityName="Britto &amp; Lima" authorityYear="2003" box="[1082,1279,718,741]" class="Actinopterygii" family="Callichthyidae" genus="Corydoras" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="86" phylum="Chordata" rank="species" species="tukano">
<emphasis box="[1082,1279,718,741]" italics="true" pageId="3" pageNumber="86">Corydoras tukano</emphasis>
</taxonomicName>
, holotype, MZUSP 82100, 40.9 mm SL; Brazil, estado do Amazonas, rio Tiquié, comunidade de Caruru.
</paragraph>
</caption>
<paragraph blockId="3.[151,795,811,962]" lastBlockId="3.[832,1475,811,994]" pageId="3" pageNumber="86">
<taxonomicName authorityName="Britto &amp; Lima" authorityYear="2003" box="[189,408,811,834]" class="Actinopterygii" family="Callichthyidae" genus="Corydoras" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="86" phylum="Chordata" rank="species" species="tukano">
<emphasis box="[189,408,811,834]" italics="true" pageId="3" pageNumber="86">Corydoras tukano</emphasis>
</taxonomicName>
may be distinguished from
<taxonomicName authorityName="Nijssen" authorityYear="1971" class="Actinopterygii" family="Callichthyidae" genus="Corydoras" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="86" phylum="Chordata" rank="species" species="weitzmani">
<emphasis italics="true" pageId="3" pageNumber="86">C. weitzmani</emphasis>
</taxonomicName>
by the presence of a dark blotch or saddle on the dorsal portion of dorsolateral plates, just posterior to last dorsal-fin ray, that extends from the 9th to the 13th plate (
<emphasis box="[765,794,907,930]" italics="true" pageId="3" pageNumber="86">vs.</emphasis>
absent); by the second trunk blotch extending vertically from adipose- to anal-fin base and along ventral surface of ventrolateral body plates (
<emphasis box="[1116,1145,843,866]" italics="true" pageId="3" pageNumber="86">vs.</emphasis>
reaching neither anal-fin base nor ventral surface of ventrolateral body plates); and by the presence of four dark, vertical stripes, in the caudal fin (
<emphasis box="[1445,1474,907,930]" italics="true" pageId="3" pageNumber="86">vs.</emphasis>
stripes absent) (see
<bibRefCitation author="Nijssen, H" box="[1035,1183,939,962]" pageId="3" pageNumber="86" pagination="89 - 98" refId="ref7563" refString="Nijssen, H. 1971. Two new species and one new subspecies of the South American catfish genus Corydoras (Pisces, Siluriformes, Callichthyidae). Beaufortia, 19: 89 - 98." type="journal article" year="1971">Nijssen, 1971</bibRefCitation>
: fig. 1; Nijssen &amp; Isbrücker, 1986: fig. 27;
<bibRefCitation author="Burgess, W" box="[974,1126,971,994]" pageId="3" pageNumber="86" refId="ref6938" refString="Burgess, W. E. 1989. An atlas of freshwater and marine catfishes. A preliminary survey of the Siluriformes. T. F. H. Publications, Neptune City, NJ. 784 p." type="book" year="1989">Burgess, 1989</bibRefCitation>
: plate 180).
</paragraph>
</subSubSection>
<caption pageId="3" pageNumber="86">
<paragraph blockId="3.[151,794,1030,1214]" pageId="3" pageNumber="86">
<emphasis bold="true" box="[151,239,1030,1054]" pageId="3" pageNumber="86">Table 1.</emphasis>
Morphometric data of the holotype and paratypes of
<taxonomicName authorityName="Britto &amp; Lima" authorityYear="2003" box="[151,343,1063,1086]" class="Actinopterygii" family="Callichthyidae" genus="Corydoras" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="86" phylum="Chordata" rank="species" species="tukano">
<emphasis box="[151,343,1063,1086]" italics="true" pageId="3" pageNumber="86">Corydoras tukano</emphasis>
</taxonomicName>
(ANSP 179201, n = 1; CAS 217960, n = 1; INPA 21423, n = 2; MNRJ 25355, n = 1; MZUSP 64096, n = 4; MZUSP 65689, n = 3; MZUSP 81179, n = 2; MZUSP 81194, n = 5; MZUSP 81244, n = 9; MZUSP 81276, n = 2; MZUSP 81153, n = 3). SD = Standard deviation.
</paragraph>
</caption>
<subSubSection lastPageId="5" lastPageNumber="88" pageId="3" pageNumber="86" type="description">
<paragraph blockId="3.[832,1475,1034,1986]" pageId="3" pageNumber="86">
<emphasis bold="true" box="[832,971,1034,1058]" pageId="3" pageNumber="86">Description.</emphasis>
Morphometric data presented in
<tableCitation box="[1325,1405,1035,1058]" captionStart="Table 1" captionStartId="3.[151,214,1030,1054]" captionText="Table 1. Morphometric data of the holotype and paratypes of Corydoras tukano (ANSP 179201, n = 1; CAS 217960, n = 1; INPA 21423, n = 2; MNRJ 25355, n = 1; MZUSP 64096, n = 4; MZUSP 65689, n = 3; MZUSP 81179, n = 2; MZUSP 81194, n = 5; MZUSP 81244, n = 9; MZUSP 81276, n = 2; MZUSP 81153, n = 3). SD = Standard deviation." pageId="3" pageNumber="86">Table 1</tableCitation>
. Head compressed with slightly convex dorsal profile; roughly triangular in dorsal view (
<figureCitation box="[1098,1161,1099,1122]" captionStart="Fig" captionStartId="3.[253,292,718,742]" captionTargetBox="[222,1417,191,691]" captionTargetId="figure-488@3.[222,1417,191,691]" captionTargetPageId="3" captionText="Fig. 4. Dorsal (left) and ventral (right) views of head and pectoral fins of Corydoras tukano, holotype, MZUSP 82100, 40.9 mm SL; Brazil, estado do Amazonas, rio Tiquié, comunidade de Caruru." figureDoi="http://doi.org/10.5281/zenodo.4647111" httpUri="https://zenodo.org/record/4647111/files/figure.png" pageId="3" pageNumber="86">Fig. 4</figureCitation>
). Snout rounded. Head profile convex from upper lip to horizontal through the anterior nares; slightly convex from that point to tip of parieto-supraoccipital expansion. Dorsal profile of body slightly convex from tip of parieto-supraoccipital expansion to last dorsal-fin ray. Postdorsal-fin body profile slightly concave to adipose-fin spine; markedly concave from this point to caudal-fin base. Ventral profile of body slightly convex from isthmus to analfin origin. Profile markedly concave from first anal-fin ray to caudal-fin base. Body roughly cylindrical in cross section at pectoral girdle, gradually becoming more compressed toward caudal fin.
</paragraph>
<paragraph blockId="3.[832,1475,1034,1986]" lastBlockId="4.[159,802,195,1978]" lastPageId="4" lastPageNumber="87" pageId="3" pageNumber="86">
Eye round, located dorso-laterally on head; orbit delimited dorsally by frontal and sphenotic, ventrally by infraorbitals. Anterior and posterior nares proximal, only separated by flap of skin. Anterior naris tubular. Posterior naris close to anterodorsal margin of orbit, separated from it by distance equal to naris diameter. Mouth small, subterminal, width nearly equal to bony orbit diameter. Maxillary barbel elongate, usually reaching anteroventral limit of gill opening (
<figureCitation box="[1395,1462,1707,1730]" captionStart="Fig" captionStartId="3.[253,292,718,742]" captionTargetBox="[222,1417,191,691]" captionTargetId="figure-488@3.[222,1417,191,691]" captionTargetPageId="3" captionText="Fig. 4. Dorsal (left) and ventral (right) views of head and pectoral fins of Corydoras tukano, holotype, MZUSP 82100, 40.9 mm SL; Brazil, estado do Amazonas, rio Tiquié, comunidade de Caruru." figureDoi="http://doi.org/10.5281/zenodo.4647111" httpUri="https://zenodo.org/record/4647111/files/figure.png" pageId="3" pageNumber="86">Fig. 4</figureCitation>
). Maxillary barbel slightly longer than outer mental barbel. Inner mental barbel fleshy. Right inner mental barbel forked in one specimen (MZUSP 81194, 38.0 mm SL). Small rounded papillae covering entire surface of all barbels, upper and lower lips, and isthmus. Gill membranes united to isthmus. Four branchiostegal rays covered by thick layer of skin; distal two rays united at their tips by branchiostegal cartilage. Teeth on upper pharyngeal tooth plate 32 (2), 39 (1), 52 (1), 53 (1), or 54 (1), and on fifth ceratobranchial 32 (1), 35 (1), 42 (1), 46 (1), 49 (1), or 50 (1), increasing in number during ontogeny.
</paragraph>
<paragraph blockId="4.[159,802,195,1978]" pageId="4" pageNumber="87">Nasal, frontal, sphenotic, compound pterotic, and parietosupraoccipital visible externally, all covered by thin layer of skin and bearing minute scattered odontodes. Frontal fontanel elongate, ellipsoid, covered by thin layer of skin; posterior tip extending into parieto-supraoccipital. Nasal slender, slightly curved laterally, mesial border contacting frontal. Frontal quadrangular; anterior expansion in contact with nasal bone, posterior portion contacting sphenotic and parietosupraoccipital. Sphenotic trapezoid in shape, contacting parieto-supraoccipital dorsally, compound pterotic posteriorly, second infraorbital ventrally. Compound pterotic roughly pipe-shaped, with slender posterior expansion contacting first dorsal body plate and first lateral-line ossicle. Contact region between compound pterotic and first dorsal body plate covered by area of thick skin. Ventral margin of compound pterotic contacting infraorbital 2 and cleithrum. Parietosupraoccipital quadrangular with posterior expansion notched at its tip, sutured with nuchal plate.</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4647113" ID-Zenodo-Dep="4647113" httpUri="https://zenodo.org/record/4647113/files/figure.png" pageId="4" pageNumber="87" startId="4.[839,878,654,678]" targetBox="[851,1475,201,592]" targetPageId="4">
<paragraph blockId="4.[839,1480,654,742]" pageId="4" pageNumber="87">
<emphasis bold="true" box="[839,906,654,678]" pageId="4" pageNumber="87">Fig. 5.</emphasis>
Orbital region of
<taxonomicName authorityName="Britto &amp; Lima" authorityYear="2003" box="[1090,1279,655,678]" class="Actinopterygii" family="Callichthyidae" genus="Corydoras" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="87" phylum="Chordata" rank="species" species="tukano">
<emphasis box="[1090,1279,655,678]" italics="true" pageId="4" pageNumber="87">Corydoras tukano</emphasis>
</taxonomicName>
, paratype, MZUSP 81153, showing odontode-bearing platelets around the eye. Odontodes not depicted.
</paragraph>
</caption>
<paragraph blockId="4.[159,802,195,1978]" pageId="4" pageNumber="87">
Two infraorbital bones, externally visible, covered by thin layer of skin. First infraorbital with slender anterior expansion. Minute odontodes-bearing platelets anterodorsally on orbit (
<figureCitation box="[167,229,931,954]" captionStart="Fig" captionStartId="4.[839,878,654,678]" captionTargetBox="[851,1475,201,592]" captionTargetId="figure-790@4.[839,1483,191,602]" captionTargetPageId="4" captionText="Fig. 5. Orbital region of Corydoras tukano, paratype, MZUSP 81153, showing odontode-bearing platelets around the eye. Odontodes not depicted." figureDoi="http://doi.org/10.5281/zenodo.4647113" httpUri="https://zenodo.org/record/4647113/files/figure.png" pageId="4" pageNumber="87">Fig. 5</figureCitation>
). Presence of small, narrow, roughly triangular platelet between eye and anterior expansion of first infraorbital (
<figureCitation captionStart="Fig" captionStartId="4.[839,878,654,678]" captionTargetBox="[851,1475,201,592]" captionTargetId="figure-790@4.[839,1483,191,602]" captionTargetPageId="4" captionText="Fig. 5. Orbital region of Corydoras tukano, paratype, MZUSP 81153, showing odontode-bearing platelets around the eye. Odontodes not depicted." figureDoi="http://doi.org/10.5281/zenodo.4647113" httpUri="https://zenodo.org/record/4647113/files/figure.png" pageId="4" pageNumber="87">Fig. 5</figureCitation>
). Opercle exposed, compact in shape, with angular free border. Preopercle externally visible, slender and covered by thin layer of skin. Interopercle triangular, covered by thin layer of skin.
</paragraph>
<paragraph blockId="4.[159,802,195,1978]" pageId="4" pageNumber="87">Trunk lateral line with two laterosensory canals, reduced to small ossicles. Lateral-line canal entering neurocranium through compound pterotic, splitting into two branches before entering sphenotic: pterotic and preoperculomandibular, each with single pore. Sensory canal continuing through compound pterotic, entering sphenotic as temporal canal, which splits into two branches: one branch giving rise to infraorbital canal, other branch entering frontal through supraorbital canal. Supraorbital canal with two branches: epiphyseal, opening in frontal bone, and anterior, running through nasal bone. Nasal canal with single opening at each end. Infraorbital canal running through entire second infraorbital, extending to infraorbital 1 and opening into two pores. Preoperculomandibular branch giving rise to preoperculomandibular canal, which runs through entire preopercle with three openings, leading to pores 3, 4, and 5, respectively.</paragraph>
<paragraph blockId="4.[159,802,195,1978]" lastBlockId="4.[839,1483,835,1754]" pageId="4" pageNumber="87">
Body plates with minute odontodes restricted to posterior margins. Nuchal plate exposed. Cleithrum and mesial process of scapulocoracoid exposed. Minute odontodes scattered over area between scapulocoracoids. Body plates not touching counterparts in specimens up to
<quantity box="[585,677,1795,1818]" metricMagnitude="-2" metricUnit="m" metricValue="2.82" pageId="4" pageNumber="87" unit="mm" value="28.2">28.2 mm</quantity>
SL, leaving narrow naked area on medial dorsal and ventral surfaces. Dorsolateral body plates 23 (11), 24* (21), or 25 (1); ventrolateral body plates 20 (2), 21* (29), or 22 (2); dorsolateral body plates along dorsal-fin base 6 (12), 7* (20), or 8 (1); dorsolateral body plates from adipose fin to caudal-fin base 7 (13) or 8* (20); preadipose platelets 2 (4), 3* (8), 4 (18), or 5 (3). Precaudal vertebrae 8 (3), 9 (3); caudal vertebrae 13 (1), 14 (3), 15 (2); five pairs of ribs, first pair conspicuously larger than others.
</paragraph>
<paragraph blockId="4.[839,1483,835,1754]" pageId="4" pageNumber="87">
Dorsal fin roughly triangular; its origin just posterior to third dorsolateral body plate. Dorsal spine shorter than 1st or 2nd branched rays. Distal tip of spine with minute-segmented unossified portion. Anterior border of dorsal spine smooth; posterior border with minute serrations. First two branched dorsal-fin rays and unossified portion of dorsal spine markedly elongate in large males; first branched ray the longest (see
<figureCitation box="[839,898,1155,1178]" captionStart="Fig" captionStartId="2.[298,337,1411,1435]" captionTargetBox="[299,1359,803,1387]" captionTargetId="figure-366@2.[299,1359,803,1387]" captionTargetPageId="2" captionText="Fig. 3. Corydoras tukano, paratype, MZUSP 81179, presumably a mature male, 36.1 mm SL; Brazil," figureDoi="http://doi.org/10.5281/zenodo.4647109" httpUri="https://zenodo.org/record/4647109/files/figure.png" pageId="4" pageNumber="87">Fig. 3</figureCitation>
and “Sexual dimorphism”, below). Dorsal-fin rays I,
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all specimens examined. Adipose fin roughly triangular; its origin separated from base of last dorsal-fin ray by 7-9 dorsolateral body plates. Anal fin roughly triangular; its origin located just posterior to 13th to14th ventrolateral body plates, at vertical through anterior margin of adipose-fin spine. Analfin rays ii,
<quantity box="[945,984,1347,1370]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="4" pageNumber="87" unit="in" value="5.0">5 in</quantity>
all specimens. Pectoral fin triangular; its origin located just posterior to gill opening. Ossified portion of pectoral spine shorter than first branched rays. Distal tip of spine with minute-segmented unossified portion. Pectoral spine with well-developed serrations along entire posterior border. Pectoral-fin rays I,8, one specimen with I,7. Pelvic fin ellipsoid; its origin just below second ventrolateral body plate, at vertical through base of second branched dorsal-fin ray. Pelvic-fin rays i,5. Caudal fin bilobed; upper lobe slightly longer. Principal caudal-fin rays i,6/6,i, one specimen i,7/6,i; upper procurrent caudal-fin rays iii; lower procurrent caudalfin rays iii. All fins with minute odontodes scattered over all rays.
</paragraph>
<paragraph blockId="4.[839,1482,1795,1978]" lastBlockId="5.[151,795,195,794]" lastPageId="5" lastPageNumber="88" pageId="4" pageNumber="87">
<emphasis bold="true" box="[839,1029,1795,1819]" pageId="4" pageNumber="87">Color in alcohol.</emphasis>
Ground coloration of head gray to brown, light ventrally. Wide, slightly oblique dark vertical blotch (“mask”) from top of head throughout anterior and posterior margins of eye to lower anterior opercle corner. Dark v-shaped stripe between nares, with angle directed anteriorly. Barbels cream. Opercle and preopercle with narrow, dark, oblique, vertically-elongated blotch, formed by relatively large chromatophores.
</paragraph>
<paragraph blockId="5.[151,795,195,794]" pageId="5" pageNumber="88">Ground color of trunk cream to light brown. Nuchal plate gray to dark brown. Large, roughly rectangular, dorsolateral dark blotch (“first trunk blotch”) extending dorsally from posterior margin of second plate to fifth-eighth dorsal plate, and ventrally from posterior margin of cleithrum to fifth ventral plate. Dark blotch on dorsal portion of dorsolateral plates, just posterior to last dorsal-fin ray, extending from 9th to 13th plate; in some specimens connected to large, anterior dorsolateral blotch. Second large, roughly rectangular, dorsolateral dark blotch (“second trunk blotch”) below adipose fin, extending vertically from adipose- to anal-fin base. Scattered chromatophores on margins of plates, close to midline, between the two large dorsolateral blotches. Caudal fin with four vertical, narrow dark stripes. Dorsal- and pectoral-fin spines gray; remaining rays and membrane of both fins unpigmented. Base of anterior anal-fin rays dark, remaining rays and membrane unpigmented.</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4647115" ID-Zenodo-Dep="4647115" httpUri="https://zenodo.org/record/4647115/files/figure.png" pageId="5" pageNumber="88" startId="5.[832,871,754,778]" targetBox="[832,1475,191,722]" targetPageId="5">
<paragraph blockId="5.[832,1475,754,873]" pageId="5" pageNumber="88">
<emphasis bold="true" box="[832,906,754,778]" pageId="5" pageNumber="88">Fig. 6.</emphasis>
Upper rio Negro basin and adjoining river basins, showing the distribution of
<taxonomicName authorityName="Britto &amp; Lima" authorityYear="2003" box="[1135,1330,786,809]" class="Actinopterygii" family="Callichthyidae" genus="Corydoras" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="88" phylum="Chordata" rank="species" species="tukano">
<emphasis box="[1135,1330,786,809]" italics="true" pageId="5" pageNumber="88">Corydoras tukano</emphasis>
</taxonomicName>
(squares). 1- Type locality, rio Tiquié; 2- rio Uaupés; 3 rio Negro; 4- rio Içana; and 5 rio Japurá.
</paragraph>
</caption>
<paragraph blockId="5.[151,794,834,986]" pageId="5" pageNumber="88">
<emphasis bold="true" box="[151,298,834,858]" pageId="5" pageNumber="88">Color in life.</emphasis>
Based on photographs published by
<bibRefCitation author="Lucanus, O" pageId="5" pageNumber="88" pagination="10 - 11" refId="ref7430" refString="Lucanus, O. 1998. Interesting imports. Tropical Fish Hobbyist, 47: 10 - 11, 14." type="journal article" year="1998">Lucanus (1998: 14)</bibRefCitation>
,
<bibRefCitation author="Finley, L" box="[267,458,867,890]" pageId="5" pageNumber="88" pagination="128 - 130" refId="ref7203" refString="Finley, L. 1998. A couple on Corydoras. Corydoras cf. reynoldsi. Tropical Fish Hobbyist, 46: 128 - 130." type="journal article" year="1998">Finley (1998: 129)</bibRefCitation>
,
<bibRefCitation author="Anonymous" box="[466,717,867,890]" pageId="5" pageNumber="88" pagination="210" refId="ref6446" refString="Anonymous. 1998. Neu importiert: Panzerwelse aus Brasilien. Die Aquarien und Terrarien Zeitschrifft (DATZ), 51 (4): 210." type="journal article" year="1998">Anonymous (1998: 210)</bibRefCitation>
, and on field observations by FCTL. Very similar to preserved specimens, but ground color cream, never light brown; greenish metallic hue on opercle and cleithrum.
</paragraph>
<paragraph blockId="5.[151,795,1026,1370]" pageId="5" pageNumber="88">
<emphasis bold="true" box="[151,381,1026,1050]" pageId="5" pageNumber="88">Sexual dimorphism.</emphasis>
We have not dissected any specimens to confirm their sex, but aquarists who have bred the species in captivity (
<emphasis box="[287,326,1091,1114]" italics="true" pageId="5" pageNumber="88">e.g.</emphasis>
<bibRefCitation author="James, A" box="[332,479,1091,1114]" pageId="5" pageNumber="88" refId="ref7322" refString="James, A. 2003 a. Corydoras sp. cf. reynoldsi Myers &amp; Weitzman, 1960. http: // www. scotcat. com / factsheets / c _ cf _ reynoldsi _ asher. htm (updated March 2003)." type="journal volume" year="2003">James, 2003a</bibRefCitation>
, b) mentioned that specimens with elongated dorsal fins are mature males. Presumably mature males examined (MZUSP 64096, 1 of 4; MZUSP 81179, 2) possess the soft portion of the dorsal spine, and the first two dorsal-fin branched rays very elongate. The first branched ray is the longest, followed by the soft portion of the dorsal spine. This results in an anteriorly elongated dorsal fin (see
<figureCitation box="[151,215,1315,1338]" captionStart="Fig" captionStartId="2.[298,337,1411,1435]" captionTargetBox="[299,1359,803,1387]" captionTargetId="figure-366@2.[299,1359,803,1387]" captionTargetPageId="2" captionText="Fig. 3. Corydoras tukano, paratype, MZUSP 81179, presumably a mature male, 36.1 mm SL; Brazil," figureDoi="http://doi.org/10.5281/zenodo.4647109" httpUri="https://zenodo.org/record/4647109/files/figure.png" pageId="5" pageNumber="88">Fig. 3</figureCitation>
;
<tableCitation box="[227,306,1315,1338]" captionStart="Table 1" captionStartId="3.[151,214,1030,1054]" captionText="Table 1. Morphometric data of the holotype and paratypes of Corydoras tukano (ANSP 179201, n = 1; CAS 217960, n = 1; INPA 21423, n = 2; MNRJ 25355, n = 1; MZUSP 64096, n = 4; MZUSP 65689, n = 3; MZUSP 81179, n = 2; MZUSP 81194, n = 5; MZUSP 81244, n = 9; MZUSP 81276, n = 2; MZUSP 81153, n = 3). SD = Standard deviation." pageId="5" pageNumber="88">Table 1</tableCitation>
). See the “Discussion”, for comparisons with another
<taxonomicName authorityName="Lacepede" authorityYear="1803" box="[241,355,1347,1370]" class="Actinopterygii" family="Callichthyidae" genus="Corydoras" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="88" phylum="Chordata" rank="genus">
<emphasis box="[241,355,1347,1370]" italics="true" pageId="5" pageNumber="88">Corydoras</emphasis>
</taxonomicName>
that possess elongated dorsal fins.
</paragraph>
<paragraph blockId="5.[151,794,1410,1466]" pageId="5" pageNumber="88">
<emphasis bold="true" box="[151,328,1410,1434]" pageId="5" pageNumber="88">Popular name.</emphasis>
“Waipotá” (both in Tukano and Tuyuka languages).
</paragraph>
<paragraph blockId="5.[151,795,1506,1754]" pageId="5" pageNumber="88">
<emphasis bold="true" box="[151,470,1506,1530]" pageId="5" pageNumber="88">Habitat and ecological notes.</emphasis>
The rio Tiquié, type-locality of
<taxonomicName authorityName="Britto &amp; Lima" authorityYear="2003" box="[151,254,1539,1562]" class="Actinopterygii" family="Callichthyidae" genus="Corydoras" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="88" phylum="Chordata" rank="species" species="tukano">
<emphasis box="[151,254,1539,1562]" italics="true" pageId="5" pageNumber="88">C. tukano</emphasis>
</taxonomicName>
, is a blackwater river. At the exact
<typeStatus box="[623,669,1539,1562]" pageId="5" pageNumber="88">type</typeStatus>
locality the bottom is sandy, with a moderate amount of vegetal debris.
<taxonomicName authorityName="Britto &amp; Lima" authorityYear="2003" box="[151,351,1603,1626]" class="Actinopterygii" family="Callichthyidae" genus="Corydoras" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="88" phylum="Chordata" rank="species" species="tukano">
<emphasis box="[151,351,1603,1626]" italics="true" pageId="5" pageNumber="88">Corydoras tukano</emphasis>
</taxonomicName>
also was collected at sites with predominantly clay substrates.
<bibRefCitation author="James, A" box="[441,597,1635,1658]" pageId="5" pageNumber="88" refId="ref7322" refString="James, A. 2003 a. Corydoras sp. cf. reynoldsi Myers &amp; Weitzman, 1960. http: // www. scotcat. com / factsheets / c _ cf _ reynoldsi _ asher. htm (updated March 2003)." type="journal volume" year="2003">James (2003a</bibRefCitation>
,b) related that in captivity specimens of this species (identified by him as
<emphasis box="[151,440,1699,1722]" italics="true" pageId="5" pageNumber="88">
<taxonomicName authorityName="Lacepede" authorityYear="1803" box="[151,265,1699,1722]" class="Actinopterygii" family="Callichthyidae" genus="Corydoras" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="88" phylum="Chordata" rank="genus">Corydoras</taxonomicName>
sp.
<taxonomicName authorityName="Myers &amp; Weitzman" authorityYear="1960" box="[308,440,1699,1722]" class="Actinopterygii" family="Callichthyidae" genus="Corydoras" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="88" phylum="Chordata" rank="species" species="reynoldsi">cf. reynoldsi</taxonomicName>
</emphasis>
) lay several scattered clutches of large eggs (
<quantity box="[272,354,1731,1754]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="5" pageNumber="88" unit="mm" value="2.2">2.2 mm</quantity>
in size). See also “Discussion”, below.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="88" type="distribution">
<paragraph blockId="5.[151,794,1794,1978]" lastBlockId="5.[832,1475,930,1337]" pageId="5" pageNumber="88">
<emphasis bold="true" box="[151,307,1794,1818]" pageId="5" pageNumber="88">Distribution.</emphasis>
Known only from the rio Tiquié and its tributaries, as the igarapé Cabari. Rio Tiquié is a tributary of the rio Uaupés, in the upper rio Negro basin, Amazonas,
<collectingCountry box="[723,789,1859,1882]" name="Brazil" pageId="5" pageNumber="88">Brazil</collectingCountry>
, near the Brazilian/Colombian border (
<figureCitation box="[572,641,1891,1914]" captionStart="Fig" captionStartId="5.[832,871,754,778]" captionTargetBox="[832,1475,191,722]" captionTargetId="figure-753@5.[830,1475,191,722]" captionTargetPageId="5" captionText="Fig. 6. Upper rio Negro basin and adjoining river basins, showing the distribution of Corydoras tukano (squares). 1- Type locality, rio Tiquié; 2- rio Uaupés; 3 rio Negro; 4- rio Içana; and 5 rio Japurá." figureDoi="http://doi.org/10.5281/zenodo.4647115" httpUri="https://zenodo.org/record/4647115/files/figure.png" pageId="5" pageNumber="88">Fig. 6</figureCitation>
). Most of the specimens studied were collected in the upper rio Tiquié, in the area between Pari-Cachoeira (
<geoCoordinate box="[536,619,1955,1978]" degrees="0" direction="north" minutes="15" orientation="latitude" pageId="5" pageNumber="88" precision="925" value="0.25">0°15N</geoCoordinate>
,
<geoCoordinate box="[634,736,1955,1978]" degrees="69" direction="west" minutes="46" orientation="longitude" pageId="5" pageNumber="88" precision="925" value="-69.76667">69°46W</geoCoordinate>
) and Caruru (
<geoCoordinate box="[918,998,930,953]" degrees="0" direction="north" minutes="16" orientation="latitude" pageId="5" pageNumber="88" precision="925" value="0.26666668">0°16N</geoCoordinate>
,
<geoCoordinate box="[1007,1108,930,953]" degrees="69" direction="west" minutes="54" orientation="longitude" pageId="5" pageNumber="88" precision="925" value="-69.9">69°54W</geoCoordinate>
). One sample (MZUSP 65689) was collected near Cunuri (
<geoCoordinate box="[1094,1177,962,985]" degrees="0" direction="north" minutes="12" orientation="latitude" pageId="5" pageNumber="88" precision="925" value="0.2">0°12N</geoCoordinate>
,
<geoCoordinate box="[1192,1296,962,985]" degrees="69" direction="west" minutes="22" orientation="longitude" pageId="5" pageNumber="88" precision="925" value="-69.36667">69°22W</geoCoordinate>
), a downstream locality not intensively sampled by the second author or associates. According to Tukano and Tuyuka fishermen,
<taxonomicName authorityName="Britto &amp; Lima" authorityYear="2003" box="[832,1024,1058,1081]" class="Actinopterygii" family="Callichthyidae" genus="Corydoras" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="88" phylum="Chordata" rank="species" species="tukano">
<emphasis box="[832,1024,1058,1081]" italics="true" pageId="5" pageNumber="88">Corydoras tukano</emphasis>
</taxonomicName>
occurs thorough the rio Tiquié basin, from its lower course upstream to Caruru, where a major waterfall about
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high, the Cachoeira do Caruru, effectively separates the downstream ichthyofauna from the upper section of this river. Again according to Tukano and Tuyuka fishermen,
<taxonomicName authorityName="Britto &amp; Lima" authorityYear="2003" class="Actinopterygii" family="Callichthyidae" genus="Corydoras" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="88" phylum="Chordata" rank="species" species="tukano">
<emphasis italics="true" pageId="5" pageNumber="88">C. tukano</emphasis>
</taxonomicName>
is absent upstream of this waterfall. In fact, despite considerable time and effort spent in apparently suitable sites, the second author was unable to collect any
<taxonomicName authorityName="Britto &amp; Lima" authorityYear="2003" box="[1361,1474,1282,1305]" class="Actinopterygii" family="Callichthyidae" genus="Corydoras" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="88" phylum="Chordata" rank="species" species="tukano">
<emphasis box="[1361,1474,1282,1305]" italics="true" pageId="5" pageNumber="88">C. tukano</emphasis>
</taxonomicName>
upstream of the Cachoeira do Caruru.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="88" type="etymology">
<paragraph blockId="5.[832,1475,1377,1593]" pageId="5" pageNumber="88">
<emphasis bold="true" box="[832,960,1377,1401]" pageId="5" pageNumber="88">Etymology.</emphasis>
After the Tukano, an Amerindian group from the upper rio Negro and Japurá/Caquetá basins in
<collectingCountry box="[1396,1470,1410,1433]" name="Brazil" pageId="5" pageNumber="88">Brazil</collectingCountry>
,
<collectingCountry box="[832,940,1442,1465]" name="Colombia" pageId="5" pageNumber="88">Colombia</collectingCountry>
, and
<collectingCountry box="[999,1113,1442,1465]" name="Venezuela" pageId="5" pageNumber="88">Venezuela</collectingCountry>
. Main cultural features displayed by this ethnic group are the Tukanoan language, the patrilineal clans system, and a diet having as staple foods manioc and fish (
<bibRefCitation author="Chernella, J" box="[887,1063,1538,1561]" pageId="5" pageNumber="88" pagination="238 - 248" refId="ref7109" refString="Chernella, J. M. 1989. Managing rivers of hunger: the Tukano of Brazil. Advances in Economic Botany, 7: 238 - 248." type="journal article" year="1989">Chernella, 1989</bibRefCitation>
;
<bibRefCitation author="Ribeiro, B" box="[1075,1226,1538,1561]" pageId="5" pageNumber="88" refId="ref8018" refString="Ribeiro, B. G. 1995. Os indios das aguas pretas. EDUSP / Companhia das Letras, Sao Paulo. 270 p." type="book" year="1995">Ribeiro, 1995</bibRefCitation>
). The presently known range of
<taxonomicName authorityName="Britto &amp; Lima" authorityYear="2003" box="[927,1032,1570,1593]" class="Actinopterygii" family="Callichthyidae" genus="Corydoras" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="88" phylum="Chordata" rank="species" species="tukano">
<emphasis box="[927,1032,1570,1593]" italics="true" pageId="5" pageNumber="88">C. tukano</emphasis>
</taxonomicName>
is entirely within the Tukano territory.
</paragraph>
</subSubSection>
</treatment>
</document>