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<document id="F21542035593BC05404409A512EAFDE5" ID-DOI="10.11646/zootaxa.3669.4.3" ID-GBIF-Dataset="13b37162-4203-4029-985f-3bd89ff3cb10" ID-ISSN="1175-5326" ID-Zenodo-Dep="222578" ID-ZooBank="CD30BE63-C779-45CD-A25E-D633BBC0B9FF" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460632612540" checkinUser="plazi" docAuthor="Frýda, Jiří, Ferrová, Lenka &amp; Frýdová, Barbora" docDate="2013" docId="03F687FAC540C25CFF3EFDFB0468F93C" docLanguage="en" docName="zt03669p489.pdf" docOrigin="Zootaxa 3669 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Palaeozygopleura lukesi Frýda, Ferrová &amp; Frýdová, 2013, sp. nov." docType="treatment" docVersion="8" lastPageNumber="481" masterDocId="FFCFFF82C54CC250FFA9FD6C054CFFA8" masterDocTitle="Review of palaeozygopleurid gastropods (Palaeozygopleuridae, Gastropoda) from Devonian strata of the Perunica microplate (Bohemia), with a re-evaluation of their stratigraphic distribution, notes on their ontogeny, and descriptions of new taxa" masterLastPageNumber="489" masterPageNumber="469" pageNumber="481" updateTime="1698334631360" updateUser="plazi">
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<mods:title id="ACB944E375C29E20246095A0E9681135">Review of palaeozygopleurid gastropods (Palaeozygopleuridae, Gastropoda) from Devonian strata of the Perunica microplate (Bohemia), with a re-evaluation of their stratigraphic distribution, notes on their ontogeny, and descriptions of new taxa</mods:title>
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<mods:namePart id="725F2B72ADC858374A4F366B48CB8B3F">Frýda, Jiří</mods:namePart>
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<mods:namePart id="64C19644E1058269B1123E12F4201D6B">Ferrová, Lenka</mods:namePart>
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<mods:namePart id="1505B237469F85D2058AB45F57F0F8A0">Frýdová, Barbora</mods:namePart>
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<mods:title id="A75B1F0C5944680D744B4DDFA894401C">Zootaxa</mods:title>
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<mods:part id="88352094D45A591464AB51DBF4285634">
<mods:date id="38914E0FE291475449675677576AD178">2013</mods:date>
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<mods:identifier id="8230078E8581AB36A51BB9CDD9395B9E" type="DOI">10.11646/zootaxa.3669.4.3</mods:identifier>
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<treatment id="03F687FAC540C25CFF3EFDFB0468F93C" ID-DOI="http://doi.org/10.5281/zenodo.6165362" ID-GBIF-Taxon="119570001" ID-Zenodo-Dep="6165362" LSID="urn:lsid:plazi:treatment:03F687FAC540C25CFF3EFDFB0468F93C" httpUri="http://treatment.plazi.org/id/03F687FAC540C25CFF3EFDFB0468F93C" lastPageNumber="481" pageId="12" pageNumber="481">
<subSubSection id="C3456567C540C25CFF3EFDFB0442FF7A" pageId="12" pageNumber="481" type="nomenclature">
<paragraph id="8BE036ECC540C25CFF3EFDFB076DFF19" blockId="12.[151,545,151,210]" box="[151,545,151,177]" pageId="12" pageNumber="481">
<heading id="D0A88180C540C25CFF3EFDFB076DFF19" bold="true" box="[151,545,151,177]" fontSize="11" level="1" pageId="12" pageNumber="481" reason="1">
<emphasis id="B92BEAFEC540C25CFF3EFDFB076DFF19" bold="true" box="[151,545,151,177]" pageId="12" pageNumber="481">
<taxonomicName id="4C5F4D6FC540C25CFF3EFDFB04F1FF19" box="[151,445,151,177]" class="Gastropoda" family="Palaeozygopleuridae" genus="Palaeozygopleura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Murchisoniina" pageId="12" pageNumber="481" phylum="Mollusca" rank="species" species="lukesi" status="sp. nov.">
<emphasis id="B92BEAFEC540C25CFF3EFDFB04F1FF19" bold="true" box="[151,445,151,177]" italics="true" pageId="12" pageNumber="481">Palaeozygopleura lukesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2185785C540C25CFE6DFDFB076DFF19" box="[452,545,151,177]" pageId="12" pageNumber="481" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE036ECC540C25CFF3EFDD60442FF7A" blockId="12.[151,545,151,210]" box="[151,270,186,210]" pageId="12" pageNumber="481">
<figureCitation id="13642A69C540C25CFF3EFDD6059AFF7A" box="[151,214,186,210]" captionStart="FIGURE 3" captionStartId="11.[151,250,1783,1806]" captionTargetBox="[201,1362,184,1763]" captionTargetId="figure@11.[201,1362,184,1764]" captionTargetPageId="11" captionText="FIGURE 3. Palaeozygopleura lukesi sp. nov. from middle Early Devonian (Pragian) age strata of the Konĕprusy Limestone (Praha Formation) in Barrandian, Czech Republic. A C, H, and I. Holotype (UUG JF 922), D. Paratype A (UUG JF 923), E, F. Paratype B (UUG JF 924), G. Paratype C (UUG JF 925), J, M. Paratype D (UUG JF 926), K. Paratype E (UUG JF 927), L, N. Paratype F (UUG JF 928). All samples are from the type locality. A— shell width 1.5 mm, shell height: B, C— 4.6 mm, D— 3.9 mm, E, F— 3.8 mm, K— 3.6 mm, L— 5 mm, figure height of shell detail: G— 1.1 mm, H— 1.5 mm, I— 1.6 mm, J— 5.6 mm, M— 2.8 mm, N— 2.1 mm." httpUri="https://zenodo.org/record/222581/files/figure.png" pageId="12" pageNumber="481">Fig. 3</figureCitation>
AN
</paragraph>
</subSubSection>
<subSubSection id="C3456567C540C25CFF3EFC6C04A1FDBC" pageId="12" pageNumber="481" type="materials_examined">
<paragraph id="8BE036ECC540C25CFF3EFC6C07ECFEBF" blockId="12.[151,1437,255,1684]" box="[151,672,255,280]" pageId="12" pageNumber="481">
<emphasis id="B92BEAFEC540C25CFF3EFC6C0447FEB0" bold="true" box="[151,267,256,280]" pageId="12" pageNumber="481">
<typeStatus id="54E4884EC540C25CFF3EFC6C044AFEB0" box="[151,262,256,280]" pageId="12" pageNumber="481" type="holotype">Holotype</typeStatus>
.
</emphasis>
UUG JF 922 (
<figureCitation id="13642A69C540C25CFE19FD9304BCFEBF" box="[432,496,255,279]" captionStart="FIGURE 3" captionStartId="11.[151,250,1783,1806]" captionTargetBox="[201,1362,184,1763]" captionTargetId="figure@11.[201,1362,184,1764]" captionTargetPageId="11" captionText="FIGURE 3. Palaeozygopleura lukesi sp. nov. from middle Early Devonian (Pragian) age strata of the Konĕprusy Limestone (Praha Formation) in Barrandian, Czech Republic. A C, H, and I. Holotype (UUG JF 922), D. Paratype A (UUG JF 923), E, F. Paratype B (UUG JF 924), G. Paratype C (UUG JF 925), J, M. Paratype D (UUG JF 926), K. Paratype E (UUG JF 927), L, N. Paratype F (UUG JF 928). All samples are from the type locality. A— shell width 1.5 mm, shell height: B, C— 4.6 mm, D— 3.9 mm, E, F— 3.8 mm, K— 3.6 mm, L— 5 mm, figure height of shell detail: G— 1.1 mm, H— 1.5 mm, I— 1.6 mm, J— 5.6 mm, M— 2.8 mm, N— 2.1 mm." httpUri="https://zenodo.org/record/222581/files/figure.png" pageId="12" pageNumber="481">Fig. 3</figureCitation>
AC, H, and I).
</paragraph>
<paragraph id="8BE036ECC540C25CFF6EFC480709FE0F" blockId="12.[151,1437,255,1684]" pageId="12" pageNumber="481">
<emphasis id="B92BEAFEC540C25CFF6EFC48040AFE94" bold="true" box="[199,326,292,316]" pageId="12" pageNumber="481">
<typeStatus id="54E4884EC540C25CFF6EFC48040DFE94" box="[199,321,292,316]" pageId="12" pageNumber="481" type="paratype">Paratypes</typeStatus>
.
</emphasis>
The shells figured herein as
<figureCitation id="13642A69C540C25CFD23FC4807A4FE94" box="[650,744,292,316]" captionStart="FIGURE 3" captionStartId="11.[151,250,1783,1806]" captionTargetBox="[201,1362,184,1763]" captionTargetId="figure@11.[201,1362,184,1764]" captionTargetPageId="11" captionText="FIGURE 3. Palaeozygopleura lukesi sp. nov. from middle Early Devonian (Pragian) age strata of the Konĕprusy Limestone (Praha Formation) in Barrandian, Czech Republic. A C, H, and I. Holotype (UUG JF 922), D. Paratype A (UUG JF 923), E, F. Paratype B (UUG JF 924), G. Paratype C (UUG JF 925), J, M. Paratype D (UUG JF 926), K. Paratype E (UUG JF 927), L, N. Paratype F (UUG JF 928). All samples are from the type locality. A— shell width 1.5 mm, shell height: B, C— 4.6 mm, D— 3.9 mm, E, F— 3.8 mm, K— 3.6 mm, L— 5 mm, figure height of shell detail: G— 1.1 mm, H— 1.5 mm, I— 1.6 mm, J— 5.6 mm, M— 2.8 mm, N— 2.1 mm." httpUri="https://zenodo.org/record/222581/files/figure.png" pageId="12" pageNumber="481">Figure 3</figureCitation>
D (
<typeStatus id="54E4884EC540C25CFCA5FC480622FE94" box="[780,878,292,316]" pageId="12" pageNumber="481" type="paratype">paratype</typeStatus>
A; UUG JF 923),
<figureCitation id="13642A69C540C25CFB96FC4801D1FE94" box="[1087,1181,292,316]" captionStart="FIGURE 3" captionStartId="11.[151,250,1783,1806]" captionTargetBox="[201,1362,184,1763]" captionTargetId="figure@11.[201,1362,184,1764]" captionTargetPageId="11" captionText="FIGURE 3. Palaeozygopleura lukesi sp. nov. from middle Early Devonian (Pragian) age strata of the Konĕprusy Limestone (Praha Formation) in Barrandian, Czech Republic. A C, H, and I. Holotype (UUG JF 922), D. Paratype A (UUG JF 923), E, F. Paratype B (UUG JF 924), G. Paratype C (UUG JF 925), J, M. Paratype D (UUG JF 926), K. Paratype E (UUG JF 927), L, N. Paratype F (UUG JF 928). All samples are from the type locality. A— shell width 1.5 mm, shell height: B, C— 4.6 mm, D— 3.9 mm, E, F— 3.8 mm, K— 3.6 mm, L— 5 mm, figure height of shell detail: G— 1.1 mm, H— 1.5 mm, I— 1.6 mm, J— 5.6 mm, M— 2.8 mm, N— 2.1 mm." httpUri="https://zenodo.org/record/222581/files/figure.png" pageId="12" pageNumber="481">Figure 3</figureCitation>
E,F (
<typeStatus id="54E4884EC540C25CFB7DFC48007AFE94" box="[1236,1334,292,316]" pageId="12" pageNumber="481" type="paratype">paratype</typeStatus>
B; UUG JF 924),
<figureCitation id="13642A69C540C25CFF54FC2B0417FEF7" box="[253,347,327,351]" captionStart="FIGURE 3" captionStartId="11.[151,250,1783,1806]" captionTargetBox="[201,1362,184,1763]" captionTargetId="figure@11.[201,1362,184,1764]" captionTargetPageId="11" captionText="FIGURE 3. Palaeozygopleura lukesi sp. nov. from middle Early Devonian (Pragian) age strata of the Konĕprusy Limestone (Praha Formation) in Barrandian, Czech Republic. A C, H, and I. Holotype (UUG JF 922), D. Paratype A (UUG JF 923), E, F. Paratype B (UUG JF 924), G. Paratype C (UUG JF 925), J, M. Paratype D (UUG JF 926), K. Paratype E (UUG JF 927), L, N. Paratype F (UUG JF 928). All samples are from the type locality. A— shell width 1.5 mm, shell height: B, C— 4.6 mm, D— 3.9 mm, E, F— 3.8 mm, K— 3.6 mm, L— 5 mm, figure height of shell detail: G— 1.1 mm, H— 1.5 mm, I— 1.6 mm, J— 5.6 mm, M— 2.8 mm, N— 2.1 mm." httpUri="https://zenodo.org/record/222581/files/figure.png" pageId="12" pageNumber="481">Figure 3</figureCitation>
G (
<typeStatus id="54E4884EC540C25CFE2BFC2B04A8FEF7" box="[386,484,327,351]" pageId="12" pageNumber="481" type="paratype">paratype</typeStatus>
C; UUG JF 925),
<figureCitation id="13642A69C540C25CFD15FC2B0657FEF7" box="[700,795,327,351]" captionStart="FIGURE 3" captionStartId="11.[151,250,1783,1806]" captionTargetBox="[201,1362,184,1763]" captionTargetId="figure@11.[201,1362,184,1764]" captionTargetPageId="11" captionText="FIGURE 3. Palaeozygopleura lukesi sp. nov. from middle Early Devonian (Pragian) age strata of the Konĕprusy Limestone (Praha Formation) in Barrandian, Czech Republic. A C, H, and I. Holotype (UUG JF 922), D. Paratype A (UUG JF 923), E, F. Paratype B (UUG JF 924), G. Paratype C (UUG JF 925), J, M. Paratype D (UUG JF 926), K. Paratype E (UUG JF 927), L, N. Paratype F (UUG JF 928). All samples are from the type locality. A— shell width 1.5 mm, shell height: B, C— 4.6 mm, D— 3.9 mm, E, F— 3.8 mm, K— 3.6 mm, L— 5 mm, figure height of shell detail: G— 1.1 mm, H— 1.5 mm, I— 1.6 mm, J— 5.6 mm, M— 2.8 mm, N— 2.1 mm." httpUri="https://zenodo.org/record/222581/files/figure.png" pageId="12" pageNumber="481">Figure 3</figureCitation>
J,M (
<typeStatus id="54E4884EC540C25CFCFEFC2B06F5FEF7" box="[855,953,327,351]" pageId="12" pageNumber="481" type="paratype">paratype</typeStatus>
D; UUG JF 926),
<figureCitation id="13642A69C540C25CFB3AFC2B01BDFEF7" box="[1171,1265,327,351]" captionStart="FIGURE 3" captionStartId="11.[151,250,1783,1806]" captionTargetBox="[201,1362,184,1763]" captionTargetId="figure@11.[201,1362,184,1764]" captionTargetPageId="11" captionText="FIGURE 3. Palaeozygopleura lukesi sp. nov. from middle Early Devonian (Pragian) age strata of the Konĕprusy Limestone (Praha Formation) in Barrandian, Czech Republic. A C, H, and I. Holotype (UUG JF 922), D. Paratype A (UUG JF 923), E, F. Paratype B (UUG JF 924), G. Paratype C (UUG JF 925), J, M. Paratype D (UUG JF 926), K. Paratype E (UUG JF 927), L, N. Paratype F (UUG JF 928). All samples are from the type locality. A— shell width 1.5 mm, shell height: B, C— 4.6 mm, D— 3.9 mm, E, F— 3.8 mm, K— 3.6 mm, L— 5 mm, figure height of shell detail: G— 1.1 mm, H— 1.5 mm, I— 1.6 mm, J— 5.6 mm, M— 2.8 mm, N— 2.1 mm." httpUri="https://zenodo.org/record/222581/files/figure.png" pageId="12" pageNumber="481">Figure 3</figureCitation>
K (
<typeStatus id="54E4884EC540C25CFAB1FC2B0036FEF7" box="[1304,1402,327,351]" pageId="12" pageNumber="481" type="paratype">paratype</typeStatus>
E; UUG JF 927),
<figureCitation id="13642A69C540C25CFEE3FC0004E0FE2C" box="[330,428,364,388]" captionStart="FIGURE 3" captionStartId="11.[151,250,1783,1806]" captionTargetBox="[201,1362,184,1763]" captionTargetId="figure@11.[201,1362,184,1764]" captionTargetPageId="11" captionText="FIGURE 3. Palaeozygopleura lukesi sp. nov. from middle Early Devonian (Pragian) age strata of the Konĕprusy Limestone (Praha Formation) in Barrandian, Czech Republic. A C, H, and I. Holotype (UUG JF 922), D. Paratype A (UUG JF 923), E, F. Paratype B (UUG JF 924), G. Paratype C (UUG JF 925), J, M. Paratype D (UUG JF 926), K. Paratype E (UUG JF 927), L, N. Paratype F (UUG JF 928). All samples are from the type locality. A— shell width 1.5 mm, shell height: B, C— 4.6 mm, D— 3.9 mm, E, F— 3.8 mm, K— 3.6 mm, L— 5 mm, figure height of shell detail: G— 1.1 mm, H— 1.5 mm, I— 1.6 mm, J— 5.6 mm, M— 2.8 mm, N— 2.1 mm." httpUri="https://zenodo.org/record/222581/files/figure.png" pageId="12" pageNumber="481">Figure 3</figureCitation>
L,N (
<typeStatus id="54E4884EC540C25CFE45FC000702FE2C" box="[492,590,364,388]" pageId="12" pageNumber="481" type="paratype">paratype</typeStatus>
F; UUG JF 928), and 32 unfigured specimens (UUG JF 929960). All
<typeStatus id="54E4884EC540C25CFF3EFCFC044EFE00" box="[151,258,400,424]" pageId="12" pageNumber="481" type="paratype">paratypes</typeStatus>
come from the
<typeStatus id="54E4884EC540C25CFE1AFCFC04AFFE00" box="[435,483,400,424]" pageId="12" pageNumber="481">type</typeStatus>
locality.
</paragraph>
<paragraph id="8BE036ECC540C25CFF6EFCD8006DFE64" blockId="12.[151,1437,255,1684]" box="[199,1313,435,461]" pageId="12" pageNumber="481">
<emphasis id="B92BEAFEC540C25CFF6EFCD80429FE64" bold="true" box="[199,357,435,460]" pageId="12" pageNumber="481">
<typeStatus id="54E4884EC540C25CFF6EFCD8044EFE64" box="[199,258,436,460]" pageId="12" pageNumber="481">Type</typeStatus>
locality.
</emphasis>
Císařský lom quarry, S of the village of Konĕprusy (
<geoCoordinate id="EE6B502BC540C25CFC1FFCD8012DFE64" box="[950,1121,436,460]" direction="north" orientation="latitude" pageId="12" pageNumber="481" precision="1" value="49.916943">49°55'00.99&quot;N</geoCoordinate>
,
<geoCoordinate id="EE6B502BC540C25CFBC4FCD80058FE64" box="[1133,1300,436,460]" direction="east" orientation="longitude" pageId="12" pageNumber="481" precision="1" value="14.060117">14°3'36.420&quot;E</geoCoordinate>
).
</paragraph>
<paragraph id="8BE036ECC540C25CFF6EFCB404A1FDBC" blockId="12.[151,1437,255,1684]" pageId="12" pageNumber="481">
<emphasis id="B92BEAFEC540C25CFF6EFCB40420FE58" bold="true" box="[199,364,471,496]" pageId="12" pageNumber="481">
<typeStatus id="54E4884EC540C25CFF6EFCB4044EFE58" box="[199,258,472,496]" pageId="12" pageNumber="481">Type</typeStatus>
horizon.
</emphasis>
Middle part of the Konĕprusy Limestone (Praha Formation), at the same level as
<emphasis id="B92BEAFEC540C25CFAACFCB40588FDBC" italics="true" pageId="12" pageNumber="481">Konepruselia zaki</emphasis>
Frýda and Ferrová, 2011.
</paragraph>
</subSubSection>
<subSubSection id="C3456567C540C25CFF6EFF4C065BFDF4" pageId="12" pageNumber="481" type="etymology">
<paragraph id="8BE036ECC540C25CFF6EFF4C065BFDF4" blockId="12.[151,1437,255,1684]" pageId="12" pageNumber="481">
<emphasis id="B92BEAFEC540C25CFF6EFF4C0402FD90" bold="true" box="[199,334,544,568]" pageId="12" pageNumber="481">Etymology.</emphasis>
After our friend Pavel Lukeš, who has contributed considerably to our understanding of the Devonian stratigraphy of the Barrandian area (
<collectingCountry id="F348767CC540C25CFD09FF280640FDF4" box="[672,780,580,604]" name="Czech Republic" pageId="12" pageNumber="481">Bohemia</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C3456567C540C25CFF6EFF0B01C2FCB8" pageId="12" pageNumber="481" type="diagnosis">
<paragraph id="8BE036ECC540C25CFF6EFF0B01C2FCB8" blockId="12.[151,1437,255,1684]" pageId="12" pageNumber="481">
<emphasis id="B92BEAFEC540C25CFF6EFF0B040DFD28" bold="true" box="[199,321,615,640]" pageId="12" pageNumber="481">Diagnosis.</emphasis>
<taxonomicName id="4C5F4D6FC540C25CFEE0FF040758FD28" box="[329,532,616,640]" class="Gastropoda" family="Palaeozygopleuridae" genus="Palaeozygopleura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Murchisoniina" pageId="12" pageNumber="481" phylum="Mollusca" rank="genus">
<emphasis id="B92BEAFEC540C25CFEE0FF040758FD28" box="[329,532,616,640]" italics="true" pageId="12" pageNumber="481">Palaeozygopleura</emphasis>
</taxonomicName>
with slightly convex sides, convex whorl profile; whorls symmetrically arched, ornamented by prominent, regularly spaced, asymmetrically curved opisthocyrt costae; costae covering lateral as well as basal surface of whorls (
<figureCitation id="13642A69C540C25CFDA7FFC3071FFD6F" box="[526,595,687,711]" captionStart="FIGURE 3" captionStartId="11.[151,250,1783,1806]" captionTargetBox="[201,1362,184,1763]" captionTargetId="figure@11.[201,1362,184,1764]" captionTargetPageId="11" captionText="FIGURE 3. Palaeozygopleura lukesi sp. nov. from middle Early Devonian (Pragian) age strata of the Konĕprusy Limestone (Praha Formation) in Barrandian, Czech Republic. A C, H, and I. Holotype (UUG JF 922), D. Paratype A (UUG JF 923), E, F. Paratype B (UUG JF 924), G. Paratype C (UUG JF 925), J, M. Paratype D (UUG JF 926), K. Paratype E (UUG JF 927), L, N. Paratype F (UUG JF 928). All samples are from the type locality. A— shell width 1.5 mm, shell height: B, C— 4.6 mm, D— 3.9 mm, E, F— 3.8 mm, K— 3.6 mm, L— 5 mm, figure height of shell detail: G— 1.1 mm, H— 1.5 mm, I— 1.6 mm, J— 5.6 mm, M— 2.8 mm, N— 2.1 mm." httpUri="https://zenodo.org/record/222581/files/figure.png" pageId="12" pageNumber="481">Fig. 3</figureCitation>
F); collabral costae culminating slightly above mid-whorl; angle between costae and lower as well as upper suture about 45 degrees; pleural angle of adult shell about 15 degrees; apical angle at first three whorls about 28 degrees; diameter of first half of whorl about
<quantity id="4CA79B09C540C25CFB8BFF9B01C5FCB8" box="[1058,1161,759,784]" metricMagnitude="-4" metricUnit="m" metricValue="2.7" pageId="12" pageNumber="481" unit="mm" value="0.27">0.27 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C3456567C540C25CFF6EFE770468F93C" pageId="12" pageNumber="481" type="description">
<paragraph id="8BE036ECC540C25CFF6EFE77044BFBFC" blockId="12.[151,1437,255,1684]" pageId="12" pageNumber="481">
<emphasis id="B92BEAFEC540C25CFF6EFE770415FC9C" bold="true" box="[199,345,795,820]" pageId="12" pageNumber="481">Description.</emphasis>
High-spired, multiwhorled dextrally coiled shell with up to 10 whorls; sides of shell slightly convex; pleural angle about 15 degrees, sutural slope angle about 17 degrees; lateral whorl sides between sutures very convex and symmetrically arched; sutures relatively deep; base of whorls rounded, anomphalous; lateral, basal parts of whorls ornamented by prominent, asymmetrical arched costae; costae forming wide asymmetrical arched sinus; sinus culminating above middle of whorl; angle between costae, lower as well as upper suture about 45 degrees; distance between costae about double that of costae; protruding protoconch relatively large; diameter of first half of whorl about
<quantity id="4CA79B09C540C25CFE6FFE980761FBA4" box="[454,557,1012,1036]" metricMagnitude="-4" metricUnit="m" metricValue="2.7" pageId="12" pageNumber="481" unit="mm" value="0.27">0.27 mm</quantity>
, that of embryonic chamber about
<quantity id="4CA79B09C540C25CFC18FE980154FBA4" box="[945,1048,1012,1036]" metricMagnitude="-4" metricUnit="m" metricValue="1.4" pageId="12" pageNumber="481" unit="mm" value="0.14">0.14 mm</quantity>
; first preserved costae from end of first whorl; preservation of shell not allowing for determination of exact position of protoconch/teleoconch boundary.
</paragraph>
<paragraph id="8BE036ECC540C25CFF6EF9330173FA30" blockId="12.[151,1437,255,1684]" pageId="12" pageNumber="481">
<emphasis id="B92BEAFEC540C25CFF6EF933042EFBD0" bold="true" box="[199,354,1119,1144]" pageId="12" pageNumber="481">Comparison.</emphasis>
The deep, asymmetrically arched costae differ
<taxonomicName id="4C5F4D6FC540C25CFCD8F90D0681FBD0" box="[881,973,1120,1144]" class="Gastropoda" family="Palaeozygopleuridae" genus="Palaeozygopleura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Murchisoniina" pageId="12" pageNumber="481" phylum="Mollusca" rank="species" species="lukesi">
<emphasis id="B92BEAFEC540C25CFCD8F90D0681FBD0" box="[881,973,1120,1144]" italics="true" pageId="12" pageNumber="481">P. lukesi</emphasis>
</taxonomicName>
from the majority of the species of genus
<taxonomicName id="4C5F4D6FC540C25CFF3EF9E8042FFB34" box="[151,355,1156,1180]" class="Gastropoda" family="Palaeozygopleuridae" genus="Palaeozygopleura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Murchisoniina" pageId="12" pageNumber="481" phylum="Mollusca" rank="genus">
<emphasis id="B92BEAFEC540C25CFF3EF9E8042FFB34" box="[151,355,1156,1180]" italics="true" pageId="12" pageNumber="481">Palaeozygopleura</emphasis>
</taxonomicName>
, which have either a straight or symmetrical arched costae. Only three Early Devonian palaeozygopleurid gastropods:
<taxonomicName id="4C5F4D6FC540C25CFDACF9C507EBFB68" box="[517,679,1192,1216]" class="Gastropoda" family="Palaeozygopleuridae" genus="Palaeozygopleura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Murchisoniina" pageId="12" pageNumber="481" phylum="Mollusca" rank="species" species="devonicans">
<emphasis id="B92BEAFEC540C25CFDACF9C507EBFB68" box="[517,679,1192,1216]" italics="true" pageId="12" pageNumber="481">P. devonicans</emphasis>
</taxonomicName>
,
<taxonomicName id="4C5F4D6FC540C25CFD14F9C50625FB68" box="[701,873,1193,1216]" class="Gastropoda" family="Palaeozygopleuridae" genus="Bojozyga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Murchisoniina" pageId="12" pageNumber="481" phylum="Mollusca" rank="species" species="mira">
<emphasis id="B92BEAFEC540C25CFD14F9C50625FB68" box="[701,873,1193,1216]" italics="true" pageId="12" pageNumber="481">Bojozyga mira</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C5F4D6FC540C25CFC1EF9C50191FB68" authority="Fryda, 1993" authorityName="Fryda" authorityYear="1993" box="[951,1245,1192,1216]" class="Gastropoda" family="Palaeozygopleuridae" genus="Palaeozygopleura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Murchisoniina" pageId="12" pageNumber="481" phylum="Mollusca" rank="species" species="chlupaci">
<emphasis id="B92BEAFEC540C25CFC1EF9C50170FB68" box="[951,1084,1192,1216]" italics="true" pageId="12" pageNumber="481">P. chlupaci</emphasis>
Frýda, 1993
</taxonomicName>
, have distinctly asymmetrically arched costae. Shells of
<taxonomicName id="4C5F4D6FC540C25CFDC0F9A10784FB4C" box="[617,712,1228,1252]" class="Gastropoda" family="Palaeozygopleuridae" genus="Palaeozygopleura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Murchisoniina" pageId="12" pageNumber="481" phylum="Mollusca" rank="species" species="lukesi">
<emphasis id="B92BEAFEC540C25CFDC0F9A10784FB4C" box="[617,712,1228,1252]" italics="true" pageId="12" pageNumber="481">P. lukesi</emphasis>
</taxonomicName>
have very convex lateral sides of the whorls, in contrast to
<taxonomicName id="4C5F4D6FC540C25CFA2EF9A10459FAA0" class="Gastropoda" family="Palaeozygopleuridae" genus="Palaeozygopleura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Murchisoniina" pageId="12" pageNumber="481" phylum="Mollusca" rank="species" species="devonicans">
<emphasis id="B92BEAFEC540C25CFA2EF9A10459FAA0" italics="true" pageId="12" pageNumber="481">P. devonicans</emphasis>
</taxonomicName>
, where the moderately convex whorls are flattened in their median region (Horný 1955: pl. 1, figs 1, 2). Shells of
<taxonomicName id="4C5F4D6FC540C25CFF56F8790417FA84" box="[255,347,1300,1324]" class="Gastropoda" family="Palaeozygopleuridae" genus="Palaeozygopleura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Murchisoniina" pageId="12" pageNumber="481" phylum="Mollusca" rank="species" species="lukesi">
<emphasis id="B92BEAFEC540C25CFF56F8790417FA84" box="[255,347,1300,1324]" italics="true" pageId="12" pageNumber="481">P. lukesi</emphasis>
</taxonomicName>
have a much wider pleural angle than
<taxonomicName id="4C5F4D6FC540C25CFCA0F8790610FA84" box="[777,860,1301,1324]" class="Gastropoda" family="Palaeozygopleuridae" genus="Bojozyga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Murchisoniina" pageId="12" pageNumber="481" phylum="Mollusca" rank="species" species="mira">
<emphasis id="B92BEAFEC540C25CFCA0F8790610FA84" box="[777,860,1301,1324]" italics="true" pageId="12" pageNumber="481">B. mira</emphasis>
</taxonomicName>
. In addition,
<taxonomicName id="4C5F4D6FC540C25CFC58F8790108FA84" box="[1009,1092,1301,1324]" class="Gastropoda" family="Palaeozygopleuridae" genus="Bojozyga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Murchisoniina" pageId="12" pageNumber="481" phylum="Mollusca" rank="species" species="mira">
<emphasis id="B92BEAFEC540C25CFC58F8790108FA84" box="[1009,1092,1301,1324]" italics="true" pageId="12" pageNumber="481">B. mira</emphasis>
</taxonomicName>
has less convex whorls, which are rather flat below the upper suture, and a much finer, and less arched costae. The Pragian aged
<taxonomicName id="4C5F4D6FC540C25CFA93F85500D0FAF8" box="[1338,1436,1336,1360]" class="Gastropoda" family="Palaeozygopleuridae" genus="Palaeozygopleura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Murchisoniina" pageId="12" pageNumber="481" phylum="Mollusca" rank="species" species="lukesi">
<emphasis id="B92BEAFEC540C25CFA93F85500D0FAF8" box="[1338,1436,1336,1360]" italics="true" pageId="12" pageNumber="481">P. lukesi</emphasis>
</taxonomicName>
resembles the Lochkovian
<taxonomicName id="4C5F4D6FC540C25CFE6BF8310772FADC" box="[450,574,1372,1396]" class="Gastropoda" family="Palaeozygopleuridae" genus="Palaeozygopleura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Murchisoniina" pageId="12" pageNumber="481" phylum="Mollusca" rank="species" species="chlupaci">
<emphasis id="B92BEAFEC540C25CFE6BF8310772FADC" box="[450,574,1372,1396]" italics="true" pageId="12" pageNumber="481">P. chlupaci</emphasis>
</taxonomicName>
by the shape of its costae, but it differs from the latter species by much lower whorls, a wider apical angle, and symmetrically arched whorl side between sutures.
</paragraph>
<paragraph id="8BE036ECC540C25CFF6EF8C80783F9E4" blockId="12.[151,1437,255,1684]" pageId="12" pageNumber="481">
<emphasis id="B92BEAFEC540C25CFF6EF8C80408FA14" bold="true" box="[199,324,1444,1468]" pageId="12" pageNumber="481">Pathology.</emphasis>
An anomalous development of the ornamentation was found on the lateral whorl in one specimen (UUG JF 928:
<figureCitation id="13642A69C540C25CFEE1F8A404C7FA48" box="[328,395,1480,1504]" captionStart="FIGURE 3" captionStartId="11.[151,250,1783,1806]" captionTargetBox="[201,1362,184,1763]" captionTargetId="figure@11.[201,1362,184,1764]" captionTargetPageId="11" captionText="FIGURE 3. Palaeozygopleura lukesi sp. nov. from middle Early Devonian (Pragian) age strata of the Konĕprusy Limestone (Praha Formation) in Barrandian, Czech Republic. A C, H, and I. Holotype (UUG JF 922), D. Paratype A (UUG JF 923), E, F. Paratype B (UUG JF 924), G. Paratype C (UUG JF 925), J, M. Paratype D (UUG JF 926), K. Paratype E (UUG JF 927), L, N. Paratype F (UUG JF 928). All samples are from the type locality. A— shell width 1.5 mm, shell height: B, C— 4.6 mm, D— 3.9 mm, E, F— 3.8 mm, K— 3.6 mm, L— 5 mm, figure height of shell detail: G— 1.1 mm, H— 1.5 mm, I— 1.6 mm, J— 5.6 mm, M— 2.8 mm, N— 2.1 mm." httpUri="https://zenodo.org/record/222581/files/figure.png" pageId="12" pageNumber="481">Fig. 3</figureCitation>
N), which was most likely caused by shell repair after a failed predation attack. Similarly damaged shells of other species of
<taxonomicName id="4C5F4D6FC540C25CFD8BF88007A2F9AC" box="[546,750,1516,1540]" class="Gastropoda" family="Palaeozygopleuridae" genus="Palaeozygopleura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Murchisoniina" pageId="12" pageNumber="481" phylum="Mollusca" rank="genus">
<emphasis id="B92BEAFEC540C25CFD8BF88007A2F9AC" box="[546,750,1516,1540]" italics="true" pageId="12" pageNumber="481">Palaeozygopleura</emphasis>
</taxonomicName>
were described in the Pragian
<emphasis id="B92BEAFEC540C25CFBE3F88101E5F9AC" box="[1098,1193,1516,1540]" italics="true" pageId="12" pageNumber="481">P. a li na e</emphasis>
(see Horný 1955: pl. III, fig. 4), in the Lochkovian
<taxonomicName id="4C5F4D6FC540C25CFE40FB7D072AF980" box="[489,614,1552,1576]" class="Gastropoda" family="Palaeozygopleuridae" genus="Palaeozygopleura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Murchisoniina" pageId="12" pageNumber="481" phylum="Mollusca" rank="species" species="chlupaci">
<emphasis id="B92BEAFEC540C25CFE40FB7D072AF980" box="[489,614,1552,1576]" italics="true" pageId="12" pageNumber="481">P. chlupaci</emphasis>
</taxonomicName>
(see Frýda 1993: fig. 2.6), and also occur in the Emsian
<taxonomicName id="4C5F4D6FC540C25CFB4EFB7D070EF9E4" authority="Fryda, Ferrova, Berkyova and Frydova, 2008" authorityName="Fryda, Ferrova, Berkyova and Frydova" authorityYear="2008" class="Gastropoda" family="Palaeozygopleuridae" genus="Palaeozygopleura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Murchisoniina" pageId="12" pageNumber="481" phylum="Mollusca" rank="species" species="vaneki">
<emphasis id="B92BEAFEC540C25CFB4EFB7D0000F980" box="[1255,1356,1552,1576]" italics="true" pageId="12" pageNumber="481">P. vaneki</emphasis>
Frýda, Ferrová, Berkyová and Frýdová, 2008
</taxonomicName>
(this study).
</paragraph>
<paragraph id="8BE036ECC540C25CFF6EFB3B0468F93C" blockId="12.[151,1437,255,1684]" pageId="12" pageNumber="481">
<emphasis id="B92BEAFEC540C25CFF6EFB3B079FF9D8" bold="true" box="[199,723,1623,1648]" pageId="12" pageNumber="481">Stratigraphic and geographic distributions.</emphasis>
<taxonomicName id="4C5F4D6FC540C25CFD75FB3406BFF9D8" box="[732,1011,1624,1648]" class="Gastropoda" family="Palaeozygopleuridae" genus="Palaeozygopleura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Murchisoniina" pageId="12" pageNumber="481" phylum="Mollusca" rank="species" species="lukesi">
<emphasis id="B92BEAFEC540C25CFD75FB3406BFF9D8" box="[732,1011,1624,1648]" italics="true" pageId="12" pageNumber="481">Palaeozygopleura lukesi</emphasis>
</taxonomicName>
is only known from its
<typeStatus id="54E4884EC540C25CFAA7FB350072F9D9" box="[1294,1342,1625,1649]" pageId="12" pageNumber="481">type</typeStatus>
locality and horizon.
</paragraph>
</subSubSection>
</treatment>
</document>