272 lines
37 KiB
XML
272 lines
37 KiB
XML
<document id="4BD770E1852A5D094C9764DCAF4762D7" ID-DOI="10.5281/zenodo.5170203" ID-ISSN="1942-1354" ID-Zenodo-Dep="5170203" ID-ZooBank="AB1A57F0-7CB4-4830-920B-DF219740A596" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628281492031" checkinUser="felipe" docAuthor="Blahnik, Roger J. & Holzenthal, Ralph W." docDate="2017" docId="03F687A7FFCDF830FF01BB0641D7FC0F" docLanguage="en" docName="InsectaMundi.0602.1-251.pdf" docOrigin="Insecta Mundi 2017 (602)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Mortoniella (Mortoniella) spatulata Blahnik & Holzenthal 2017, new species" docType="treatment" docVersion="3" lastPageNumber="54" masterDocId="FFCFFFDFFFF8F807FFC1B865400BFFB4" masterDocTitle="Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *" masterLastPageNumber="251" masterPageNumber="1" pageNumber="52" updateTime="1699039969677" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:mods id="1FECA443955A1CC5AE97B6F70519F830" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="BD6384FBB4DB90E8DA03DC469EA23635">
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<mods:title id="71CDFA03CEDAF2FC0E78CFB50621E30A">Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *</mods:title>
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<mods:name id="359A942D872BFD580B078623A78631BD" type="personal">
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<mods:namePart id="ECCB5E08C7A33BF5B2B34BD92CA71D2F">Blahnik, Roger J.</mods:namePart>
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<mods:name id="DD9AB54F764B0E55247852FDD8009A61" type="personal">
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<mods:roleTerm id="5CBA33FB806C6A0140644BCDCD55D5A1">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="FDDA449D7F4791C0EB0A351DFCC58DD9">Holzenthal, Ralph W.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="8F04F27E02453E54B96286613FE45B56">text</mods:typeOfResource>
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<mods:title id="3BFA844F9C09BDABD8AF33F7A97F87F5">Insecta Mundi</mods:title>
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</mods:titleInfo>
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<mods:part id="1FCE56593068E59E32B7988502F9B9DF">
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<mods:date id="F6D0B50C0BBFE2C8F68BD43F7473210B">2017</mods:date>
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<mods:detail id="7C3354BBC03A29BC4C1E93A8B7617D81" type="pubDate">
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<mods:number id="0773095FB709011D5C47B28000F2801E">2017-12-29</mods:number>
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</mods:detail>
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<mods:detail id="8B03056F87EBC4C2F6D4027EC1969360" type="volume">
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<mods:number id="E6A0887FA753EC94F91769E303D09418">2017</mods:number>
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</mods:detail>
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<mods:detail id="37BD9667C1C912B2FE0D7CDE507A9CD3" type="issue">
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<mods:number id="D9E3E604945E54AA1C976ED2CC10EAD2">602</mods:number>
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<mods:start id="E747D269F929C9910D8CAD6F0D691815">1</mods:start>
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<mods:classification id="6345ABAF702E98222A2DE667935A1B9C">journal article</mods:classification>
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<mods:identifier id="2F01493656AF231DD4572B6B54787536" type="DOI">10.5281/zenodo.5170203</mods:identifier>
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<mods:identifier id="0DE1DF22D3ECB584D6CE2B2829DB8F1C" type="ISSN">1942-1354</mods:identifier>
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<mods:identifier id="2029F7A8820EF1A8820F147261B889A3" type="Zenodo-Dep">5170203</mods:identifier>
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<mods:identifier id="F3FC8D4BE0CD33A5C00C1A814898A54E" type="ZooBank">AB1A57F0-7CB4-4830-920B-DF219740A596</mods:identifier>
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</mods:mods>
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<treatment id="03F687A7FFCDF830FF01BB0641D7FC0F" LSID="urn:lsid:plazi:treatment:03F687A7FFCDF830FF01BB0641D7FC0F" httpUri="http://treatment.plazi.org/id/03F687A7FFCDF830FF01BB0641D7FC0F" lastPageId="55" lastPageNumber="54" pageId="53" pageNumber="52">
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<subSubSection id="C345653AFFCDF832FF01BB064362FCCF" box="[192,873,867,891]" pageId="53" pageNumber="52" type="nomenclature">
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<paragraph id="8BE036B1FFCDF832FF01BB064362FCCF" blockId="53.[192,873,867,923]" box="[192,873,867,891]" pageId="53" pageNumber="52">
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<heading id="D0A881DDFFCDF832FF01BB064362FCCF" bold="true" box="[192,873,867,891]" fontSize="10" level="2" pageId="53" pageNumber="52" reason="3">
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<emphasis id="B92BEAA3FFCDF832FF01BB064362FCCF" bold="true" box="[192,873,867,891]" pageId="53" pageNumber="52">
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<taxonomicName id="4C5F4D32FFCDF832FF01BB0642B8FCCF" authority="Blahnik & Holzenthal, 2017" authorityName="Blahnik & Holzenthal" authorityYear="2017" box="[192,691,867,891]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="53" pageNumber="52" phylum="Tracheophyta" rank="species" species="spatulata" status="sp. nov." subGenus="Mortoniella">
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<emphasis id="B92BEAA3FFCDF832FF01BB064168FCCF" bold="true" box="[192,355,867,891]" italics="true" pageId="53" pageNumber="52">Mortoniella</emphasis>
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(
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<emphasis id="B92BEAA3FFCDF832FEB7BB064212FCCF" bold="true" box="[374,537,867,891]" italics="true" pageId="53" pageNumber="52">Mortoniella</emphasis>
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)
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<emphasis id="B92BEAA3FFCDF832FDEDBB0642B8FCCF" bold="true" box="[556,691,867,891]" italics="true" pageId="53" pageNumber="52">spatulata</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A21857D8FFCDF832FD03BB064362FCCF" box="[706,873,867,891]" pageId="53" pageNumber="52" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C345653AFFCDF832FF01BBE64461F8AF" pageId="53" pageNumber="52" type="description">
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<paragraph id="8BE036B1FFCDF832FF01BBE6411DFC2F" blockId="53.[192,873,867,923]" box="[192,278,899,923]" pageId="53" pageNumber="52">
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<figureCitation id="13642A34FFCDF832FF01BBE6411DFC2F" box="[192,278,899,923]" captionStart="Figure 34" captionStartId="175.[192,276,1865,1887]" captionTargetBox="[347,1288,266,1743]" captionTargetId="figure-47@175.[347,1288,266,1743]" captionTargetPageId="175" captionText="Figure 34. Mortoniella (Mortoniella) spatulata, new species, male genitalia. A—lateral; B—segment IX and tergum X, dorsal; C—phallic ensemble, ventral; D—dorsal phallic spine, dorsal." figureDoi="http://doi.org/10.5281/zenodo.5170279" httpUri="https://zenodo.org/record/5170279/files/figure.png" pageId="53" pageNumber="52">Fig. 34</figureCitation>
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</paragraph>
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<paragraph id="8BE036B1FFCDF832FF31BBA64149FB2F" blockId="53.[192,1440,963,1179]" pageId="53" pageNumber="52">
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<taxonomicName id="4C5F4D32FFCDF832FF31BBA641F5FC6F" authorityName="Blahnik & Holzenthal" authorityYear="2017" box="[240,510,963,987]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="53" pageNumber="52" phylum="Tracheophyta" rank="species" species="spatulata">
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<emphasis id="B92BEAA3FFCDF832FF31BBA641F5FC6F" box="[240,510,963,987]" italics="true" pageId="53" pageNumber="52">Mortoniella spatulata</emphasis>
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</taxonomicName>
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bears a general similarity to the other species of the bolivica subgroup, but the dorsolateral lobes of the inferior appendages are shorter and not as strongly flexed. Also, the dorsal phallic spine, while being widened and spatulate apically, is more nearly symmetrical, the ventrolateral lobes of tergum X are acutely developed (separated from the apicolateral lobes by a distinct notch), and the ventral spines of the endophallic membrane are very weakly developed. The
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<typeStatus id="54E48813FFCDF832FF01BC064121FBCF" box="[192,298,1123,1147]" pageId="53" pageNumber="52" type="holotype">holotype</typeStatus>
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specimen is over cleared and only the genital capsule (segments IX-X and phallic apparatus) is present.
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</paragraph>
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<paragraph id="8BE036B1FFCDF832FF31BCA64455FA8F" blockId="53.[192,1441,1219,1915]" pageId="53" pageNumber="52">
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<emphasis id="B92BEAA3FFCDF832FF31BCA6413DFB6F" box="[240,310,1219,1243]" italics="true" pageId="53" pageNumber="52">Adult</emphasis>
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—Length of forewing: male
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<quantity id="4CA79B54FFCDF832FD56BCA642F3FB6F" box="[663,760,1219,1243]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="53" pageNumber="52" unit="mm" value="3.8">3.8 mm</quantity>
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. Forewing with forks I, II, and III present, hind wing with forks II and III. Spur formula 0:4:4. Overall color light brown, apices of tarsi and basal segments of antennae slightly paler. Tibial spurs about same color as legs, not contrasting in color. Wing bar at anastamosis relatively indistinct, interrupted, marked with whitish setae.
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</paragraph>
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<paragraph id="8BE036B1FFCDF832FF31BD264461F8AF" blockId="53.[192,1441,1219,1915]" pageId="53" pageNumber="52">
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<emphasis id="B92BEAA3FFCDF832FF31BD2641B0FAEF" box="[240,443,1347,1371]" italics="true" pageId="53" pageNumber="52">Male genitalia</emphasis>
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—Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin convexly rounded dorsally, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by much less than ½ width of segment. Tergum X with excision between lateral lobes slightly concave; apicolateral lobes acute, curved inward from lateral margins, mesal margins subparallel. Inferior appendages only moderately large, setose, incompletely divided ventromesally, apicolateral angles short and acute, dorsolateral lobes narrow and slightly posteriorly recurved, apices acute. Mesal pockets of inferior appendage with apical processes relatively short, dorsally curved. Paramere appendages elongate, narrow, extending about as far as dorsal phallic spine. Dorsal phallic spine, as viewed laterally, distinctly undulate in contour, nearly uniform in width, narrowing apically, base distinctly curved, apex very weakly and obtusely upturned in about apical ¼; in dorsal view, with apex slightly widened and spatulate in shape (dorsoventrally flattened). Phallicata with dorsal margin weakly sclerotized, apparently to accommodate dorsal phallic spine. Endophallic membrane moderately elongate and relatively simple, ventrally with pair of very short, weakly sclerotized spines; phallotremal spines (apparently) present, very small.
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</paragraph>
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</subSubSection>
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<subSubSection id="C345653AFFCDF830FF31BF4641D7FC0F" lastPageId="55" lastPageNumber="54" pageId="53" pageNumber="52" type="materials_examined">
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<paragraph id="8BE036B1FFCDF832FF31BF464127F8CF" blockId="53.[192,1441,1219,1915]" pageId="53" pageNumber="52">
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<materialsCitation id="3B373CECFFCDF832FF31BF464127F8CF" collectingDate="1997-03-21" collectionCode="UMSP" country="Venezuela" elevation="1000" latitude="8.4645" location="Quebrada San Juan in Santa Rosa" longLatPrecision="1" longitude="-70.84867" municipality="Parque Nacional Sierra Nevada" pageId="53" pageNumber="52" specimenCode="UMSP000041337" specimenCount="1" stateProvince="Barinas" typeStatus="holotype">
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<emphasis id="B92BEAA3FFCDF832FF31BF4641BDF88F" bold="true" box="[240,438,1827,1851]" italics="true" pageId="53" pageNumber="52">
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<typeStatus id="54E48813FFCDF832FF31BF464162F88F" box="[240,361,1827,1851]" pageId="53" pageNumber="52" type="holotype">Holotype</typeStatus>
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male
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</emphasis>
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(pinned)—
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<emphasis id="B92BEAA3FFCDF832FD81BF464375F88F" bold="true" box="[576,894,1827,1851]" pageId="53" pageNumber="52">
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<collectingCountry id="F3487621FFCDF832FD81BF4642F1F88F" box="[576,762,1827,1851]" name="Venezuela" pageId="53" pageNumber="52">VENEZUELA</collectingCountry>
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:
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<collectingRegion id="499BF853FFCDF832FCC8BF464373F88F" box="[777,888,1827,1851]" country="Venezuela" name="Barinas" pageId="53" pageNumber="52">Barinas</collectingRegion>
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:
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</emphasis>
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<collectingMunicipality id="6B84ACCBFFCDF832FC47BF414513F88F" box="[902,1304,1827,1851]" pageId="53" pageNumber="52">Parque Nacional Sierra Nevada</collectingMunicipality>
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,
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<location id="8E80606AFFCDF832FAE4BF4641E3F8EF" LSID="urn:lsid:plazi:treatment:03F687A7FFCDF830FF01BB0641D7FC0F:8E80606AFFCDF832FAE4BF4641E3F8EF" country="Venezuela" latitude="8.4645" longLatPrecision="1" longitude="-70.84867" municipality="Parque Nacional Sierra Nevada" name="Quebrada San Juan in Santa Rosa" pageId="53" pageNumber="52" stateProvince="Barinas">Quebrada San Juan in Santa Rosa</location>
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,
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<geoCoordinate id="EE6B5076FFCDF832FE33BF264277F8EF" box="[498,636,1859,1883]" degrees="8.46450" direction="north" orientation="latitude" pageId="53" pageNumber="52" precision="1" value="8.4645">8.46450° N</geoCoordinate>
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,
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<geoCoordinate id="EE6B5076FFCDF832FD46BF26432EF8EF" box="[647,805,1859,1883]" degrees="70.84867" direction="west" orientation="longitude" pageId="53" pageNumber="52" precision="1" value="-70.84867">70.84867° W</geoCoordinate>
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,
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<quantity id="4CA79B54FFCDF832FCF1BF264380F8EF" box="[816,907,1859,1883]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="53" pageNumber="52" unit="m" value="1000.0">
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<elevation id="0072D182FFCDF832FCF1BF264380F8EF" box="[816,907,1859,1883]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="53" pageNumber="52" unit="m" value="1000.0">1000 m</elevation>
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</quantity>
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,
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<date id="FFE11071FFCDF832FC57BF264410F8EF" box="[918,1051,1859,1883]" pageId="53" pageNumber="52" value="1997-03-21">
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<collectingDate id="EFA5E999FFCDF832FC57BF264410F8EF" box="[918,1051,1859,1883]" pageId="53" pageNumber="52" value="1997-03-21">21.iii.1997</collectingDate>
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</date>
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, Holzenthal (
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<specimenCode id="DBF99ECAFFCDF832FB7CBF264593F8EF" box="[1213,1432,1859,1883]" collectionCode="UMSP" country="USA" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="53" pageNumber="52" type="University or college">UMSP000041337</specimenCode>
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) (
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<collectionCode id="ED4EAE74FFCDF832FF09BF064114F8CF" box="[200,287,1891,1915]" country="USA" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="53" pageNumber="52" type="University or college">UMSP</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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<paragraph id="8BE036B1FFCEF831FF31B8A6434FFF4F" blockId="54.[192,1440,195,251]" pageId="54" pageNumber="53">
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<emphasis id="B92BEAA3FFCEF831FF31B8A64179FF6F" box="[240,370,195,219]" italics="true" pageId="54" pageNumber="53">Etymology</emphasis>
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—This species is named
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<taxonomicName id="4C5F4D32FFCEF831FD5FB8A14330FF6F" authorityName="Blahnik & Holzenthal" authorityYear="2017" box="[670,827,195,219]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="54" pageNumber="53" phylum="Tracheophyta" rank="species" species="spatulata">
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<emphasis id="B92BEAA3FFCEF831FD5FB8A14330FF6F" box="[670,827,195,219]" italics="true" pageId="54" pageNumber="53">M. spatulata</emphasis>
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</taxonomicName>
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for the dorsal phallic spine of the male, which has its apex flattened and distinctly spatulate in shape.
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</paragraph>
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<paragraph id="8BE036B1FFCEF831FF01B9214295FE6F" blockId="54.[192,562,323,347]" lastBlockId="54.[192,1440,387,475]" pageId="54" pageNumber="53">
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—
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<emphasis id="B92BEAA3FFCEF831FF1AB9264239FEEF" bold="true" box="[219,562,323,347]" pageId="54" pageNumber="53">florica/leroda subgroups</emphasis>
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included species (South America):
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<taxonomicName id="4C5F4D32FFCEF831FDA6B9E64362FE2F" authority="Blahnik & Holzenthal, 2017" authorityName="Blahnik & Holzenthal" authorityYear="2017" box="[615,873,387,411]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="54" pageNumber="53" phylum="Tracheophyta" rank="species" species="bothrops" status="sp. nov.">
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<emphasis id="B92BEAA3FFCEF831FDA6B9E64362FE2F" box="[615,873,387,411]" italics="true" pageId="54" pageNumber="53">Mortoniella bothrops</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A21857D8FFCEF831FCB6B9E143B0FE2F" box="[887,955,388,411]" pageId="54" pageNumber="53" rank="species">n. sp.</taxonomicNameLabel>
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;
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<taxonomicName id="4C5F4D32FFCEF831FC07B9E1444EFE2F" authority="Blahnik & Holzenthal, 2017" authorityName="Blahnik & Holzenthal" authorityYear="2017" box="[966,1093,388,411]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="54" pageNumber="53" phylum="Tracheophyta" rank="species" species="cressae" status="sp. nov.">
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<emphasis id="B92BEAA3FFCEF831FC07B9E1444EFE2F" box="[966,1093,388,411]" italics="true" pageId="54" pageNumber="53">M. cressae</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A21857D8FFCEF831FB92B9E1449CFE2F" box="[1107,1175,388,411]" pageId="54" pageNumber="53" rank="species">n. sp.</taxonomicNameLabel>
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;
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<taxonomicName id="4C5F4D32FFCEF831FB62B9E14542FE2F" authority="Blahnik & Holzenthal, 2017" authorityName="Blahnik & Holzenthal" authorityYear="2017" box="[1187,1353,388,411]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="54" pageNumber="53" phylum="Tracheophyta" rank="species" species="curtispina" status="sp. nov.">
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<emphasis id="B92BEAA3FFCEF831FB62B9E14542FE2F" box="[1187,1353,388,411]" italics="true" pageId="54" pageNumber="53">M. curtispina</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A21857D8FFCEF831FA96B9E14590FE2F" box="[1367,1435,388,411]" pageId="54" pageNumber="53" rank="species">n. sp.</taxonomicNameLabel>
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;
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<taxonomicName id="4C5F4D32FFCEF831FF01B9C1415FFE0F" authority="Blahnik & Holzenthal, 2017" authorityName="Blahnik & Holzenthal" authorityYear="2017" box="[192,340,419,443]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="54" pageNumber="53" phylum="Tracheophyta" rank="species" species="draconis" status="sp. nov.">
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<emphasis id="B92BEAA3FFCEF831FF01B9C1415FFE0F" box="[192,340,419,443]" italics="true" pageId="54" pageNumber="53">M. draconis</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A21857D8FFCEF831FEA2B9C141A2FE0F" box="[355,425,420,443]" pageId="54" pageNumber="53" rank="species">n. sp.</taxonomicNameLabel>
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;
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<taxonomicName id="4C5F4D32FFCEF831FE74B9C142A9FE0F" authority="(Flint)" baseAuthorityName="Flint" baseAuthorityYear="1963" box="[437,674,419,443]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="54" pageNumber="53" phylum="Tracheophyta" rank="species" species="elongata">
|
|
<emphasis id="B92BEAA3FFCEF831FE74B9C14243FE0F" box="[437,584,419,443]" italics="true" pageId="54" pageNumber="53">M. elongata</emphasis>
|
|
(Flint)
|
|
</taxonomicName>
|
|
;
|
|
<taxonomicName id="4C5F4D32FFCEF831FD6EB9C14339FE0F" authority="Blahnik & Holzenthal, 2017" authorityName="Blahnik & Holzenthal" authorityYear="2017" box="[687,818,419,443]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="54" pageNumber="53" phylum="Tracheophyta" rank="species" species="furcula" status="sp. nov.">
|
|
<emphasis id="B92BEAA3FFCEF831FD6EB9C14339FE0F" box="[687,818,419,443]" italics="true" pageId="54" pageNumber="53">M. furcula</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicNameLabel id="A21857D8FFCEF831FC80B9C1438CFE0F" box="[833,903,420,443]" pageId="54" pageNumber="53" rank="species">n. sp.</taxonomicNameLabel>
|
|
;
|
|
<taxonomicName id="4C5F4D32FFCEF831FC55B9C1444DFE0F" authority="Blahnik & Holzenthal, 2017" authorityName="Blahnik & Holzenthal" authorityYear="2017" box="[916,1094,419,443]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="54" pageNumber="53" phylum="Tracheophyta" rank="species" species="grandiloba" status="sp. nov.">
|
|
<emphasis id="B92BEAA3FFCEF831FC55B9C1444DFE0F" box="[916,1094,419,443]" italics="true" pageId="54" pageNumber="53">M. grandiloba</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="A21857D8FFCEF831FB8CB9C14498FE0F" box="[1101,1171,420,443]" pageId="54" pageNumber="53" rank="species">n. sp.</taxonomicNameLabel>
|
|
,
|
|
<taxonomicName id="4C5F4D32FFCEF831FB61B9C14516FE0F" authority="Blahnik & Holzenthal, 2017" authorityName="Blahnik & Holzenthal" authorityYear="2017" box="[1184,1309,419,443]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="54" pageNumber="53" phylum="Tracheophyta" rank="species" species="ruedae" status="sp. nov.">
|
|
<emphasis id="B92BEAA3FFCEF831FB61B9C14516FE0F" box="[1184,1309,419,443]" italics="true" pageId="54" pageNumber="53">M. ruedae</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicNameLabel id="A21857D8FFCEF831FAEDB9C14579FE0F" box="[1324,1394,420,443]" pageId="54" pageNumber="53" rank="species">n. sp.</taxonomicNameLabel>
|
|
;
|
|
<taxonomicName id="4C5F4D32FFCEF831FABEB9C14131FE6F" authority="Blahnik & Holzenthal, 2017" authorityName="Blahnik & Holzenthal" authorityYear="2017" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="54" pageNumber="53" phylum="Tracheophyta" rank="species" species="schlingeri" status="sp. nov.">
|
|
<emphasis id="B92BEAA3FFCEF831FABEB9C14131FE6F" italics="true" pageId="54" pageNumber="53">M. schlingeri</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicNameLabel id="A21857D8FFCEF831FE88B9A14184FE6F" box="[329,399,452,475]" pageId="54" pageNumber="53" rank="species">n. sp.</taxonomicNameLabel>
|
|
; and
|
|
<taxonomicName id="4C5F4D32FFCEF831FE13B9A14291FE6F" authority="(Flint)" baseAuthorityName="Flint" baseAuthorityYear="1974" box="[466,666,451,475]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="54" pageNumber="53" phylum="Tracheophyta" rank="species" species="simla">
|
|
<emphasis id="B92BEAA3FFCEF831FE13B9A14234FE6F" box="[466,575,451,475]" italics="true" pageId="54" pageNumber="53">M. simla</emphasis>
|
|
(Flint)
|
|
</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
<paragraph id="8BE036B1FFCEF831FF31BA66414EFACF" blockId="54.[192,1441,515,1915]" pageId="54" pageNumber="53">
|
|
<bibRefCitation id="EFCE4B40FFCEF831FF31BA664273FDAF" author="Blahnik, R. J. & R. W. Holzenthal" box="[240,632,515,539]" pageId="54" pageNumber="53" pagination="1 - 72" refId="ref106255" refString="Blahnik, R. J., and R. W. Holzenthal. 2008. Revision of the Mexican and Central American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae). Zootaxa 1711: 1 - 72." type="journal article" year="2008">Blahnik and Holzenthal (2008)</bibRefCitation>
|
|
defined the leroda and florica subgroups based on species from Central America and commented that the two groups were probably related. As in the majority of the subgroups of the leroda group treated in the current work, the hind wings in both subgroups have both forks
|
|
<collectionCode id="ED4EAE74FFCEF831FE87BA014157FDCF" box="[326,348,612,635]" pageId="54" pageNumber="53">II</collectionCode>
|
|
and
|
|
<collectionCode id="ED4EAE74FFCEF831FE5ABA0141B7FDCF" box="[411,444,612,635]" pageId="54" pageNumber="53">III</collectionCode>
|
|
present. Also, the dorsolateral projections of the inferior appendages are either absent or greatly reduced in both subgroups, usually appearing as short rounded projections. The latter is considered an apomorphy for the combined subgroups, since a projecting dorsal lobe is probably the plesiomorphic state for the subgenus
|
|
<taxonomicName id="4C5F4D32FFCEF831FD0FBAA64354FD6F" authorityName="Blahnik & Holzenthal" authorityYear="2017" box="[718,863,707,731]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="54" pageNumber="53" phylum="Tracheophyta" rank="genus">
|
|
<emphasis id="B92BEAA3FFCEF831FD0FBAA64354FD6F" box="[718,863,707,731]" italics="true" pageId="54" pageNumber="53">Mortoniella</emphasis>
|
|
</taxonomicName>
|
|
. Many of the species, especially of the leroda subgroup, have a rather characteristically formed ventral process on segment
|
|
<collectionCode id="ED4EAE74FFCEF831FB70BA8144D9FD4F" box="[1201,1234,740,763]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="54" pageNumber="53">VI</collectionCode>
|
|
, which is short, ventrally projecting, and with the apex more or less blunt or bluntly rounded. The anterobasal margin of the ventral process is characteristically somewhat retracted (
|
|
<figureCitation id="13642A34FFCEF831FC1BBB464443FC8F" box="[986,1096,803,827]" captionStart="Figure 38" captionStartId="179.[192,276,1836,1858]" captionTargetBox="[336,1293,229,1763]" captionTargetId="figure-58@179.[336,1294,229,1763]" captionTargetPageId="179" captionText="Figure 38. Mortoniella (Mortoniella) bothrops, new species, male genitalia. A—lateral; B—segment IX and tergum X, dorsal; C—phallic ensemble, ventral; D—dorsal phallic spine and paramere appendages, dorsal; E— ventral process of segment VI, lateral." figureDoi="http://doi.org/10.5281/zenodo.5170289" httpUri="https://zenodo.org/record/5170289/files/figure.png" pageId="54" pageNumber="53">Fig. 38E</figureCitation>
|
|
). However, many species in the florica subgroup from Central America have the ventral process more angular apically, and not as evidently retracted anteriorly, though always distinctly ventrally deflected.
|
|
<collectionCode id="ED4EAE74FFCEF831FBA0BB06447FFCCF" box="[1121,1140,867,891]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="54" pageNumber="53" type="Herbarium">A</collectionCode>
|
|
bluntly rounded ventral process otherwise occurs only in the atenuata subgroup, which is apomorphically distinct, and also in
|
|
<emphasis id="B92BEAA3FFCEF831FF01BBC1417EFC0F" box="[192,373,932,955]" italics="true" pageId="54" pageNumber="53">
|
|
<taxonomicName id="4C5F4D32FFCEF831FF01BBC1417BFC0F" authority="Blahnik & Holzenthal, 2017" authorityName="Blahnik & Holzenthal" authorityYear="2017" box="[192,368,932,955]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="54" pageNumber="53" phylum="Tracheophyta" rank="species" species="guyanensis" status="sp. nov.">M. guyanensis</taxonomicName>
|
|
,
|
|
</emphasis>
|
|
<taxonomicNameLabel id="A21857D8FFCEF831FEBBBBC141B0FC0F" box="[378,443,932,955]" pageId="54" pageNumber="53" rank="species">n. sp.</taxonomicNameLabel>
|
|
, in the limona subgroup. An additional, although more difficult to define, character uniting the two subgroups is the general structure of the dorsal phallic spine, which has a characteristic undulate or sinuous inflection, probably due to the tendency for the ventral margin of the spine to have a rounded deflection that articulates with a corresponding contour in the dorsal margin of the phallicata. However, this may be a plesiomorphic character for the leroda group as a whole, since a similar character state also occurs in the akantha and bolivica subgroups. Another character, which seems to characterize most of the species of the two subgroups, is that the paramere appendages are displaced ventrally on elongate membranous lobes, with the base of the appendages curved inward. This is already evident in
|
|
<taxonomicName id="4C5F4D32FFCEF831FDC1BCC1427EFB0F" baseAuthorityName="Mosely" baseAuthorityYear="1937" box="[512,629,1187,1211]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="54" pageNumber="53" phylum="Tracheophyta" rank="species" species="leroda">
|
|
<emphasis id="B92BEAA3FFCEF831FDC1BCC1427EFB0F" box="[512,629,1187,1211]" italics="true" pageId="54" pageNumber="53">M. leroda</emphasis>
|
|
</taxonomicName>
|
|
, which seems to be a basal, or near basal, species in the combined subgroups. The character is more developed in species with paired dorsolateral processes on the phallicata, especially those of the florica subgroup (eg.
|
|
<figureCitation id="13642A34FFCEF831FC8DBC8643BEFB4F" box="[844,949,1251,1275]" captionStart="Figure 35" captionStartId="176.[192,276,1865,1887]" captionTargetBox="[301,1329,264,1769]" captionTargetId="figure-48@176.[301,1330,264,1793]" captionTargetPageId="176" captionText="Figure 35. Mortoniella (Mortoniella) elongata (Flint), male genitalia. A—lateral; B—segment IX and tergum X, dorsal; C—phallic ensemble, ventral; D—dorsal phallic spine, dorsal." figureDoi="http://doi.org/10.5281/zenodo.5170281" httpUri="https://zenodo.org/record/5170281/files/figure.png" pageId="54" pageNumber="53">Fig. 35C</figureCitation>
|
|
,
|
|
<figureCitation id="13642A34FFCEF831FC7EBC8643F8FB4F" box="[959,1011,1251,1275]" captionStart="Figure 36" captionStartId="177.[192,276,1836,1858]" captionTargetBox="[342,1283,250,1753]" captionTargetId="figure-58@177.[342,1283,250,1761]" captionTargetPageId="177" captionText="Figure 36. Mortoniella (Mortoniella) grandiloba, new species, male genitalia. A—lateral; B—segment IX and tergum X, dorsal; C—phallic ensemble, ventral; D—dorsal phallic spine and dorsolateral projections of phallicata, dorsal; E—ventral process of segment VI, lateral." figureDoi="http://doi.org/10.5281/zenodo.5170283" httpUri="https://zenodo.org/record/5170283/files/figure.png" pageId="54" pageNumber="53">36C</figureCitation>
|
|
). An additional character similarity of the two subgroups is the structure tergum X, which has apicolateral lobes that are distinctly sclerotized and projecting and have a very reduced or nearly absent ventrolateral lobes. There is also some tendency for the lobes to be flattened and converge mesally, and for the dorsomesal margin to be projecting.
|
|
</paragraph>
|
|
<paragraph id="8BE036B1FFCEF830FF31BDE64240FCEF" blockId="54.[192,1441,515,1915]" lastBlockId="55.[192,1441,195,859]" lastPageId="55" lastPageNumber="54" pageId="54" pageNumber="53">
|
|
Although the majority of species from Central America seemed to fall into 2 groups, based on a combination of characters, in treating the species from South America the distinctness of the two groups is less evident. It therefore seems prudent to combine them here. The primary character difference used to define the two groups was the structure of the inferior appendages; members of florica subgroup have paired symmetrical apical processes on the inferior appendages (long or short), and those of the leroda subgroup have a single asymmetrical apicomesal process. Because the names were used previously, and the primary defining character is diagnostically useful, we have continued to group the species from South America in the two subgroups, while recognizing that they may not be mutually monophyletic. Many of the species of the leroda subgroup from South America have upright dorsal processes on the phallicata, a ventromesal spine on the endophallic membrane, and scabrous apices of the paramere appendages (characters generally found in the florica subgroup of Central America, but absent in the species assigned to the leroda subgroup). These characters occur in addition to the presence of a prominent asymmetric ventromesal process on the inferior appendages, used to characterize the leroda subgroup. Most of the South American species of the leroda subgroup also have the apex of this mesal process scabrous (or sensillate), as in
|
|
<taxonomicName id="4C5F4D32FFCEF831FBEABF214533F8EF" authority="(Mosely)" baseAuthorityName="Mosely" baseAuthorityYear="1954" box="[1067,1336,1859,1883]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="54" pageNumber="53" phylum="Tracheophyta" rank="species" species="meralda">
|
|
<emphasis id="B92BEAA3FFCEF831FBEABF2144CAF8EF" box="[1067,1217,1859,1883]" italics="true" pageId="54" pageNumber="53">M. meralda</emphasis>
|
|
(Mosely)
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="4C5F4D32FFCEF831FABEBF214292F8CF" authority="Blahnik and Holzenthal" authorityName="Blahnik and Holzenthal. All" authorityYear="2008" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="54" pageNumber="53" phylum="Tracheophyta" rank="species" species="panamensis">
|
|
<emphasis id="B92BEAA3FFCEF831FABEBF21415DF8CF" italics="true" pageId="54" pageNumber="53">M. panamensis</emphasis>
|
|
Blahnik and Holzenthal
|
|
</taxonomicName>
|
|
from Central America. It seems likely that an asymmetric ventromesal process on the inferior appendages is a plesiomorphic character that has been subsequently lost in the florica subgroup. This may also explain the presence (retention) of this character in a number of species of the albolineata subgroup from
|
|
<collectingCountry id="F3487621FFCFF830FC92B96643A9FEAF" box="[851,930,259,283]" name="Brazil" pageId="55" pageNumber="54">Brazil</collectingCountry>
|
|
, and an indication of the relationship of these subgroups. The most likely explanation for the presence of paired rounded or upright basodorsal processes on the phallicata in the majority of species of the florica subgroup is that they represent a lineage from within the leroda subgroup that derived this apomorphy, followed by the subsequent loss of the ventromesal process and gain of apicolateral projections on the inferior appendages. The new species described below as
|
|
<taxonomicName id="4C5F4D32FFCFF830FDCEB9C14284FE0F" authorityName="Blahnik & Holzenthal" authorityYear="2017" box="[527,655,420,443]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="55" pageNumber="54" phylum="Tracheophyta" rank="species" species="cressae">
|
|
<emphasis id="B92BEAA3FFCFF830FDCEB9C14284FE0F" box="[527,655,420,443]" italics="true" pageId="55" pageNumber="54">M. cressae</emphasis>
|
|
</taxonomicName>
|
|
(
|
|
<figureCitation id="13642A34FFCFF830FD61B9C642FDFE0F" box="[672,758,419,443]" captionStart="Figure 39" captionStartId="180.[192,276,1865,1887]" captionText="Figure 39. Mortoniella (Mortoniella) cressae, new species, male genitalia. A—lateral; B—segment IX and tergum X, dorsal; C—phallic ensemble, ventral." pageId="55" pageNumber="54">Fig. 39</figureCitation>
|
|
) possibly represents a transitional species (or one that has converged on similar character modifications), with apicolateral projections on the inferior appendages present, but with the ventromesal process much reduced. Absence of a ventromesal spine on the endophallic membrane in various species of the leroda subgroup, especially those from Central America, probably represents a character loss, judging by its general presence (retention) in the florica subgroup and in a number of other subgroups. However, there seems to be no parsimonious solution for its loss, which must have occurred repeatedly. Possibly, the plesiomorphic state was for the spine to be very small, as in
|
|
<taxonomicName id="4C5F4D32FFCFF830FE25BAE14256FD2F" baseAuthorityName="Mosely" baseAuthorityYear="1937" box="[484,605,643,667]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="55" pageNumber="54" phylum="Tracheophyta" rank="species" species="leroda">
|
|
<emphasis id="B92BEAA3FFCFF830FE25BAE14256FD2F" box="[484,605,643,667]" italics="true" pageId="55" pageNumber="54">M. leroda</emphasis>
|
|
</taxonomicName>
|
|
, or in the species of the atenuata subgroup, making its loss more probable. If redefined in a combined sense to include both members of the leroda and florica subgroups, the composite subgroup would include all of the species from Central America, in addition to the species described below, with the exception of the three species of the akantha subgroup and
|
|
<taxonomicName id="4C5F4D32FFCFF830FABEBA81418BFCAF" authority="(Flint)" baseAuthorityName="Flint" baseAuthorityYear="1974" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="55" pageNumber="54" phylum="Tracheophyta" rank="species" species="pacuara">
|
|
<emphasis id="B92BEAA3FFCFF830FABEBA81412DFCAF" italics="true" pageId="55" pageNumber="54">M. pacuara</emphasis>
|
|
(Flint)
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="4C5F4D32FFCFF830FE03BB614399FCAF" authority="Blahnik and Holzenthal" authorityName="Blahnik and Holzenthal. The" authorityYear="2008" box="[450,914,771,795]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="55" pageNumber="54" phylum="Tracheophyta" rank="species" species="rodmani">
|
|
<emphasis id="B92BEAA3FFCFF830FE03BB614253FCAF" box="[450,600,771,795]" italics="true" pageId="55" pageNumber="54">M. rodmani</emphasis>
|
|
Blahnik and Holzenthal
|
|
</taxonomicName>
|
|
of the subgenus
|
|
<taxonomicName id="4C5F4D32FFCFF830FBADBB664508FCAF" authorityName="Blahnik & Holzenthal" authorityYear="2017" box="[1132,1283,771,795]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="55" pageNumber="54" phylum="Tracheophyta" rank="subGenus" subGenus="Nanotrichia">
|
|
<emphasis id="B92BEAA3FFCFF830FBADBB664508FCAF" box="[1132,1283,771,795]" italics="true" pageId="55" pageNumber="54">Nanotrichia</emphasis>
|
|
</taxonomicName>
|
|
. This entire group constitutes a revised definition of the leroda subgroup, and will be referred to subsequently as the leroda subgroup,
|
|
<emphasis id="B92BEAA3FFCFF830FE09BB21424FFCEF" box="[456,580,835,859]" italics="true" pageId="55" pageNumber="54">sensu lato</emphasis>
|
|
.
|
|
</paragraph>
|
|
<paragraph id="8BE036B1FFCFF830FF01BBC141D7FC0F" blockId="55.[192,476,931,955]" box="[192,476,931,955]" pageId="55" pageNumber="54">
|
|
—
|
|
<emphasis id="B92BEAA3FFCFF830FF1ABBC641D7FC0F" bold="true" box="[219,476,931,955]" pageId="55" pageNumber="54">“florica” subgroup</emphasis>
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |