260 lines
38 KiB
XML
260 lines
38 KiB
XML
<document id="AC4A0961E8EAE62BB3AE9C3FE6DAB0B7" ID-CLB-Dataset="36382" ID-DOI="10.11646/zootaxa.3828.1.1" ID-GBIF-Dataset="8c1cda9b-0d82-41f3-a1c1-d8eb574a1ba7" ID-ISSN="1175-5326" ID-Zenodo-Dep="286513" ID-ZooBank="B2FEAE60-7E51-45FA-A38F-930A084A5AAA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460452080076" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2014" docId="03F68791FFC018108FFE3D20074FF944" docLanguage="en" docName="zt03828p076.pdf" docOrigin="Zootaxa 3828 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Leascydmus armstrongi Jałoszyński, 2014, sp. n." docType="treatment" docVersion="7" lastPageNumber="25" masterDocId="FFCFFFE9FFD718088F693A1C034CFC3C" masterDocTitle="' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae)" masterLastPageNumber="76" masterPageNumber="1" pageNumber="24" updateTime="1698338417320" updateUser="ExternalLinkService">
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<mods:titleInfo id="551399B7FD2C87B03C2DF479520C6706">
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<mods:title id="3E3592C4E5E35955104FDA502D394C41">' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae)</mods:title>
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<mods:namePart id="A1281F11D59852322EA9690EC21CC37A">Jałoszyński, Paweł</mods:namePart>
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<mods:title id="F1B0984797E7C54567C58C81D17AD249">Zootaxa</mods:title>
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<mods:part id="8A6F43208EA95738350F5FDF4D8B5CEB">
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<mods:date id="015A7E9BF7CC0CEB44C5A74176BE9A76">2014</mods:date>
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<mods:number id="4435E6D9A61A3D7BD17DD5C0661C4CEB">3828</mods:number>
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<mods:identifier id="F08E8493A526E0A4CF0F8DCE41AAA468" type="DOI">10.11646/zootaxa.3828.1.1</mods:identifier>
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<treatment id="03F68791FFC018108FFE3D20074FF944" ID-DOI="http://doi.org/10.5281/zenodo.6141060" ID-GBIF-Taxon="127711782" ID-Zenodo-Dep="6141060" LSID="urn:lsid:plazi:treatment:03F68791FFC018108FFE3D20074FF944" httpUri="http://treatment.plazi.org/id/03F68791FFC018108FFE3D20074FF944" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
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<subSubSection id="C345650CFFC0181F8FFE3D20010AFB44" pageId="23" pageNumber="24" type="nomenclature">
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<paragraph id="8BE03687FFC0181F8FFE3D20014DFB6A" blockId="23.[151,582,1852,1912]" box="[151,513,1852,1878]" pageId="23" pageNumber="24">
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<heading id="D0A881EBFFC0181F8FFE3D20014DFB6A" bold="true" box="[151,513,1852,1878]" fontSize="11" level="1" pageId="23" pageNumber="24" reason="1">
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<emphasis id="B92BEA95FFC0181F8FFE3D20014DFB6A" bold="true" box="[151,513,1852,1878]" pageId="23" pageNumber="24">
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<taxonomicName id="4C5F4D04FFC0181F8FFE3D2002FBFB6A" ID-CoL="3SPGG" box="[151,439,1852,1878]" class="Insecta" family="Staphylinidae" genus="Leascydmus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="armstrongi" status="sp. nov.">
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<emphasis id="B92BEA95FFC0181F8FFE3D2002FBFB6A" bold="true" box="[151,439,1852,1878]" italics="true" pageId="23" pageNumber="24">Leascydmus armstrongi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21857EEFFC0181F8ED63D20014DFB6A" box="[447,513,1852,1878]" pageId="23" pageNumber="24" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BE03687FFC0181F8FFE3D43010AFB44" blockId="23.[151,582,1852,1912]" box="[151,582,1887,1912]" pageId="23" pageNumber="24">
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(
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<figureCitation id="13642A02FFC0181F8FF63D43026BFB44" box="[159,295,1887,1912]" captionStart-0="FIGURES 37 – 41" captionStart-1="FIGURES 42 – 45" captionStartId-0="19.[151,264,1848,1871]" captionStartId-1="20.[151,264,1431,1454]" captionTargetBox-0="[192,1382,201,1809]" captionTargetBox-1="[151,1436,212,1404]" captionTargetId-0="figure@19.[187,1401,193,1836]" captionTargetId-1="figure@20.[151,1436,193,1411]" captionTargetPageId-0="19" captionTargetPageId-1="20" captionText-0="FIGURES 37 – 41. Morphology of Leascydmus armstrongi sp. n. Head in dorsal (37) and ventral (38) views, labrum in dorsal view (39), right mandible in dorsal view (40), and left maxilla in ventral view (41). Abbreviations: bst, basistipes; cd, cardo; cl, clypeus; fo, foramen occipitale; fr, frons; gal, galea; gp, gular plate; gs, gular suture; hr, hypomeral ridge; lac, lacinia; lbr, labrum; lg, ligula; md, mandible; mn, mentum; mst, mediostipes; mxp, maxillary palpus; mxp 1 – 4, maxillary palpomere 1 – 4; occ, occipital constriction; ocp, occiput; pmn, prementum; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; smn, submentum; tm, temple; vt, vertex." captionText-1="FIGURES 42 – 45. Morphology of Leascydmus armstrongi sp. n. Base of pronotum and elytra in dorsal view (42), prothorax in ventral view (43), meso- and metanotum in dorsal view (44), pterothorax in ventral view (45). Abbreviations: aest 3, metanepisternum; alc, alacrista; amvp, anterior metaventral process; ar, anterior ridge; bef, basal elytral fovea; bst, basisternal part of prosternum; cx 3, metacoxa; dlf, dorsolateral fovea; elb, elytral base; hd, humeral denticle; hy, hypomeron; hyr, hypomeral ridge; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; mma, median membranous area; mscc, mesocoxal cavity; mscs, mesocoxal socket; msf, mesofurca; msvp, mesoventral process; mtvp, metaventral intercoxal process; nss, pronotosternal suture; pcc, procoxal cavity; pcr, procoxal rest; pcs, procoxal socket; pff, profurcal fovea; pre, prepectus; psn 3, metapostnotum; sc 2, mesoscutellum; sc 3, metascutum; sss, mesoscutoscutellar suture; tr 3, metatrochanter; v 3, metaventrite; vlf, ventrolateral fovea; wb, wing base." httpUri-0="https://zenodo.org/record/286522/files/figure.png" httpUri-1="https://zenodo.org/record/286523/files/figure.png" pageId="23" pageNumber="24">Figs. 37–45</figureCitation>
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,
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<figureCitation id="13642A02FFC0181F8E5B3D43021DFB4B" box="[306,337,1887,1911]" captionStart="FIGURES 46 – 48" captionStartId="22.[151,264,1976,1999]" captionTargetBox="[162,1428,885,1944]" captionTargetId="figure@22.[151,1436,879,1955]" captionTargetPageId="22" captionText="FIGURES 46 – 48. Leascydmus spp. Dorsal habitus (46, 48) and original set of labels of holotype (47). Leascydmus simplicicornis (Lea) comb. n., female (46); Leascydmus armstrongi sp. n., male (48)." httpUri="https://zenodo.org/record/286524/files/figure.png" pageId="23" pageNumber="24">48</figureCitation>
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,
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<figureCitation id="13642A02FFC0181F8E343D4302E9FB44" box="[349,421,1887,1912]" captionStart="FIGURES 49 – 52" captionStartId="23.[151,264,891,914]" captionTargetBox="[167,1425,211,862]" captionTargetId="figure@23.[151,1436,193,870]" captionTargetPageId="23" captionText="FIGURES 49 – 52. Aedeagi of Leascydmus spp. in ventral (49, 51) and lateral (50, 52) views. Leascydmus simplicicornis (Lea) comb. n. (49 – 50), Leascydmus armstrongi sp. n. (51 – 52)." httpUri="https://zenodo.org/record/286525/files/figure.png" pageId="23" pageNumber="24">51–52</figureCitation>
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,
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<figureCitation id="13642A02FFC0181F8ED93D4302B5FB44" box="[432,505,1887,1912]" captionStart="FIGURES 53 – 56" captionStartId="23.[151,264,1710,1733]" captionTargetBox="[163,1428,1002,1677]" captionTargetId="figure@23.[151,1436,994,1689]" captionTargetPageId="23" captionText="FIGURES 53 – 56. Aedeagi of Leascydmus spp. in ventral (53, 55) and lateral (54, 56) views. Leascydmus simplicicornis (Lea) comb. n. (53 – 54), Leascydmus armstrongi sp. n. (55 – 56)." httpUri="https://zenodo.org/record/286526/files/figure.png" pageId="23" pageNumber="24">55–56</figureCitation>
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,
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<figureCitation id="13642A02FFC0181F8D6D3D430163FB44" box="[516,559,1887,1912]" captionStart="FIGURES 193 – 198" captionStartId="72.[151,264,1762,1785]" captionTargetBox="[195,1376,193,1707]" captionTargetId="figure@72.[191,1378,193,1739]" captionTargetPageId="72" captionText="FIGURES 193 – 198. Distribution of revised Australian species of Horaeomorphus Schaufuss (193), Leascydmus gen. n. (194), Scydmaenozila gen. n. (195), Spinosciacharis gen. n. (196), Sciacharoides gen. n. (197), and Sciacharis Broun. Horaeomorphus anthicoides (Lea) comb. n. (193 a), Horaeomorphus verus sp. n. (193 b), Leascydmus simplicicornis (Lea) comb. n. (194 a), Leascydmus armstrongi sp. n. (194 b), Scydmaenozila macrosticta (Lea) comb. n. (195 a), Scydmaenozila latipennis (Lea) comb. n. (195 b), Scydmaenozila mallacootana sp. n. (195 c), Scydmaenozila inexpectata sp. n. (195 d), Spinosciacharis puncticeps (Franz) comb. n. (196 a), Spinosciacharis sassy sp. n. (196 b), Spinosciacharis clydensis sp. n. (196 c), Spinosciacharis acheron sp. n. (196 d), Sciacharoides thompsoni (Franz) comb. n. (197 a), Sciacharoides eucalypti (Franz) comb. n. (197 b), Sciacharoides maipotonensis (Franz) comb. n. (197 c), Sciacharoides mantistamborinensis (Franz) comb. n. (197 d), and Sciacharis (Maorinus) tasmaniensis (Franz) comb. n. (198)." httpUri="https://zenodo.org/record/286555/files/figure.png" pageId="23" pageNumber="24">194</figureCitation>
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b)
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</paragraph>
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</subSubSection>
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<subSubSection id="C345650CFFC018108FFE3DBB0383FCE8" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24" type="materials_examined">
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<paragraph id="8BE03687FFC018108FFE3DBB0383FCE8" blockId="23.[151,1436,1959,2020]" lastBlockId="24.[151,1437,151,1400]" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
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<emphasis id="B92BEA95FFC0181F8FFE3DBB02D7FBFC" bold="true" box="[151,411,1959,1984]" pageId="23" pageNumber="24">
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<typeStatus id="54E48825FFC0181F8FFE3DBB039EFBFC" box="[151,210,1959,1984]" pageId="23" pageNumber="24">Type</typeStatus>
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material studied
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</emphasis>
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.
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<emphasis id="B92BEA95FFC0181F8EC03DBB0198FBFC" bold="true" box="[425,724,1959,1984]" pageId="23" pageNumber="24">
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<typeStatus id="54E48825FFC0181F8EC03DBB0156FBFC" box="[425,538,1959,1984]" pageId="23" pageNumber="24" type="holotype">Holotype</typeStatus>
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:
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<collectingCountry id="F3487617FFC0181F8D4F3DBB0183FBFC" box="[550,719,1959,1984]" name="Australia" pageId="23" pageNumber="24">AUSTRALIA</collectingCountry>
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:
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</emphasis>
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♂: two labels: "Megalong / N. S.
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<collectingCountry id="F3487617FFC0181F8B393DBB07D8FBFC" box="[1104,1172,1959,1984]" name="United Kingdom" pageId="23" pageNumber="24">Wales</collectingCountry>
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/ J. Armstrong" [white, printed and handwritten], "
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<emphasis id="B92BEA95FFC0181F8EBD3DD0006CFBDF" box="[468,800,1996,2019]" italics="true" pageId="23" pageNumber="24">
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<taxonomicName id="4C5F4D04FFC0181F8EBD3DD001C8FBDF" box="[468,644,1996,2019]" class="Insecta" family="Staphylinidae" genus="Leascydmus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="73" phylum="Arthropoda" rank="genus" status="gen. nov.">LEASCYDMUS</taxonomicName>
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/
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<taxonomicName id="4C5F4D04FFC0181F8DCD3DD0006CFBDF" box="[676,800,1996,2019]" class="Insecta" family="Staphylinidae" genus="Leascydmus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="armstrongi">armstrongi</taxonomicName>
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</emphasis>
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m. / det. P. Jałoszyński, 2014 /
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<typeStatus id="54E48825FFC0181F8BC33DD7061FFBDF" box="[1194,1363,1995,2019]" pageId="23" pageNumber="24" type="holotype">HOLOTYPUS</typeStatus>
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" [red, printed] (
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<collectionCode id="ED4EAE42FFCF18108E683A840200FC8C" LSID="urn:lsid:biocol.org:col:32981" box="[257,332,152,176]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="24" pageNumber="25">ANIC</collectionCode>
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).
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<typeStatus id="54E48825FFCF18108E093A84029DFC8C" box="[352,465,152,176]" pageId="24" pageNumber="25" type="paratype">Paratypes</typeStatus>
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: 2 ♂♂, 4 ♀♀, same data as for
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<typeStatus id="54E48825FFCF18108C2E3A8B00E0FC8C" box="[839,940,151,176]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
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, all with yellow "
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<typeStatus id="54E48825FFCF18108B113A8407A7FC8C" box="[1144,1259,152,176]" pageId="24" pageNumber="25" type="paratype">paratypus</typeStatus>
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" labels (
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<collectionCode id="ED4EAE42FFCF18108A263A8406D4FC8C" LSID="urn:lsid:biocol.org:col:32981" box="[1359,1432,152,176]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="24" pageNumber="25">ANIC</collectionCode>
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, cPJ).
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</paragraph>
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</subSubSection>
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<subSubSection id="C345650CFFCF18108FAE3AC302A6FD21" pageId="24" pageNumber="25" type="diagnosis">
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<paragraph id="8BE03687FFCF18108FAE3AC302A6FD21" blockId="24.[151,1437,151,1400]" pageId="24" pageNumber="25">
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<emphasis id="B92BEA95FFCF18108FAE3AC3020DFCC4" bold="true" box="[199,321,223,248]" pageId="24" pageNumber="25">Diagnosis.</emphasis>
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Male and female: body uniformly brown, only tarsi lighter. Male: aedeagus in lateral view with apex slightly curved ventrally.
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</paragraph>
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</subSubSection>
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<subSubSection id="C345650CFFCF18108FAE3B3B0242F89C" pageId="24" pageNumber="25" type="description">
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<paragraph id="8BE03687FFCF18108FAE3B3B0634FD59" blockId="24.[151,1437,151,1400]" pageId="24" pageNumber="25">
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<emphasis id="B92BEA95FFCF18108FAE3B3B0215FD7C" bold="true" box="[199,345,295,320]" pageId="24" pageNumber="25">Description.</emphasis>
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Body of male (
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<figureCitation id="13642A02FFCF18108D493B3B0134FD7C" box="[544,632,295,320]" captionStart="FIGURES 46 – 48" captionStartId="22.[151,264,1976,1999]" captionTargetBox="[162,1428,885,1944]" captionTargetId="figure@22.[151,1436,879,1955]" captionTargetPageId="22" captionText="FIGURES 46 – 48. Leascydmus spp. Dorsal habitus (46, 48) and original set of labels of holotype (47). Leascydmus simplicicornis (Lea) comb. n., female (46); Leascydmus armstrongi sp. n., male (48)." httpUri="https://zenodo.org/record/286524/files/figure.png" pageId="24" pageNumber="25">Fig. 48</figureCitation>
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) strongly convex, elongate and slender, with long appendages, BL
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<quantity id="4CA79B62FFCF18108FFE3B500276FD58" box="[151,314,332,357]" metricMagnitude="-3" metricUnit="m" metricValue="2.415" metricValueMax="2.48" metricValueMin="2.35" pageId="24" pageNumber="25" unit="mm" value="2.415" valueMax="2.48" valueMin="2.35">2.35–2.48 mm</quantity>
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(mean
|
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<quantity id="4CA79B62FFCF18108EE73B5002BAFD58" box="[398,502,332,356]" metricMagnitude="-3" metricUnit="m" metricValue="2.43" pageId="24" pageNumber="25" unit="mm" value="2.43">2.43 mm</quantity>
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); cuticle glossy, body brown, tarsi lighter, yellowish-brown, vestiture yellowish.
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</paragraph>
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<paragraph id="8BE03687FFCF18108FAE3B73061CFE24" blockId="24.[151,1437,151,1400]" pageId="24" pageNumber="25">
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Head (
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<figureCitation id="13642A02FFCF18108E7A3B7302D0FDB4" box="[275,412,367,392]" captionStart="FIGURES 37 – 41" captionStartId="19.[151,264,1848,1871]" captionTargetBox="[192,1382,201,1809]" captionTargetId="figure@19.[187,1401,193,1836]" captionTargetPageId="19" captionText="FIGURES 37 – 41. Morphology of Leascydmus armstrongi sp. n. Head in dorsal (37) and ventral (38) views, labrum in dorsal view (39), right mandible in dorsal view (40), and left maxilla in ventral view (41). Abbreviations: bst, basistipes; cd, cardo; cl, clypeus; fo, foramen occipitale; fr, frons; gal, galea; gp, gular plate; gs, gular suture; hr, hypomeral ridge; lac, lacinia; lbr, labrum; lg, ligula; md, mandible; mn, mentum; mst, mediostipes; mxp, maxillary palpus; mxp 1 – 4, maxillary palpomere 1 – 4; occ, occipital constriction; ocp, occiput; pmn, prementum; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; smn, submentum; tm, temple; vt, vertex." httpUri="https://zenodo.org/record/286522/files/figure.png" pageId="24" pageNumber="25">Figs. 37–38</figureCitation>
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,
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<figureCitation id="13642A02FFCF18108EC03B6C028AFDB4" box="[425,454,368,392]" captionStart="FIGURES 46 – 48" captionStartId="22.[151,264,1976,1999]" captionTargetBox="[162,1428,885,1944]" captionTargetId="figure@22.[151,1436,879,1955]" captionTargetPageId="22" captionText="FIGURES 46 – 48. Leascydmus spp. Dorsal habitus (46, 48) and original set of labels of holotype (47). Leascydmus simplicicornis (Lea) comb. n., female (46); Leascydmus armstrongi sp. n., male (48)." httpUri="https://zenodo.org/record/286524/files/figure.png" pageId="24" pageNumber="25">48</figureCitation>
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) broadest at eyes, HL
|
||
<quantity id="4CA79B62FFCF18108DA33B6C007EFDBB" box="[714,818,368,392]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="24" pageNumber="25" unit="mm" value="0.43">0.43 mm</quantity>
|
||
, HW
|
||
<quantity id="4CA79B62FFCF18108C1F3B730757FDBB" box="[886,1051,367,392]" metricMagnitude="-4" metricUnit="m" metricValue="4.9" metricValueMax="5.0" metricValueMin="4.8" pageId="24" pageNumber="25" unit="mm" value="0.49" valueMax="0.5" valueMin="0.48">0.48–0.50 mm</quantity>
|
||
(mean
|
||
<quantity id="4CA79B62FFCF18108B1B3B6C0797FDB4" box="[1138,1243,368,392]" metricMagnitude="-4" metricUnit="m" metricValue="4.9" pageId="24" pageNumber="25" unit="mm" value="0.49">0.49 mm</quantity>
|
||
); tempora longer than eyes, strongly curved posteromesally; vertex and frons confluent and evenly convex. Eyes small and weakly projecting laterally from the head silhouette, finely faceted. Punctures on head dorsum fine and sparse, separated by spaces 1.5–
|
||
<date id="FFE11047FFCF18108E543BC00207FDC9" box="[317,331,476,501]" pageId="24" pageNumber="25">2</date>
|
||
x as wide as diameters of punctures; setae short, sparse and suberect. Antennae (
|
||
<figureCitation id="13642A02FFCF18108B8B3BC00679FDC8" box="[1250,1333,476,501]" captionStart="FIGURES 46 – 48" captionStartId="22.[151,264,1976,1999]" captionTargetBox="[162,1428,885,1944]" captionTargetId="figure@22.[151,1436,879,1955]" captionTargetPageId="22" captionText="FIGURES 46 – 48. Leascydmus spp. Dorsal habitus (46, 48) and original set of labels of holotype (47). Leascydmus simplicicornis (Lea) comb. n., female (46); Leascydmus armstrongi sp. n., male (48)." httpUri="https://zenodo.org/record/286524/files/figure.png" pageId="24" pageNumber="25">Fig. 48</figureCitation>
|
||
) slender, AnL
|
||
<quantity id="4CA79B62FFCF18108FB83BE30238FE2B" box="[209,372,511,536]" metricMagnitude="-3" metricUnit="m" metricValue="1.84" metricValueMax="1.85" metricValueMin="1.83" pageId="24" pageNumber="25" unit="mm" value="1.84" valueMax="1.85" valueMin="1.83">1.83–1.85 mm</quantity>
|
||
(mean
|
||
<quantity id="4CA79B62FFCF18108EA1381C017CFE24" box="[456,560,512,536]" metricMagnitude="-3" metricUnit="m" metricValue="1.84" pageId="24" pageNumber="25" unit="mm" value="1.84">1.84 mm</quantity>
|
||
); all antennomeres elongate, antennomere XI as long as IX–X together.
|
||
</paragraph>
|
||
<paragraph id="8BE03687FFCF18108FAE383906C4FEF1" blockId="24.[151,1437,151,1400]" pageId="24" pageNumber="25">
|
||
Pronotum (
|
||
<figureCitation id="13642A02FFCF18108E2F383802E8FE00" box="[326,420,548,573]" captionStart="FIGURES 42 – 45" captionStartId="20.[151,264,1431,1454]" captionTargetBox="[151,1436,212,1404]" captionTargetId="figure@20.[151,1436,193,1411]" captionTargetPageId="20" captionText="FIGURES 42 – 45. Morphology of Leascydmus armstrongi sp. n. Base of pronotum and elytra in dorsal view (42), prothorax in ventral view (43), meso- and metanotum in dorsal view (44), pterothorax in ventral view (45). Abbreviations: aest 3, metanepisternum; alc, alacrista; amvp, anterior metaventral process; ar, anterior ridge; bef, basal elytral fovea; bst, basisternal part of prosternum; cx 3, metacoxa; dlf, dorsolateral fovea; elb, elytral base; hd, humeral denticle; hy, hypomeron; hyr, hypomeral ridge; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; mma, median membranous area; mscc, mesocoxal cavity; mscs, mesocoxal socket; msf, mesofurca; msvp, mesoventral process; mtvp, metaventral intercoxal process; nss, pronotosternal suture; pcc, procoxal cavity; pcr, procoxal rest; pcs, procoxal socket; pff, profurcal fovea; pre, prepectus; psn 3, metapostnotum; sc 2, mesoscutellum; sc 3, metascutum; sss, mesoscutoscutellar suture; tr 3, metatrochanter; v 3, metaventrite; vlf, ventrolateral fovea; wb, wing base." httpUri="https://zenodo.org/record/286523/files/figure.png" pageId="24" pageNumber="25">Figs. 42</figureCitation>
|
||
,
|
||
<figureCitation id="13642A02FFCF18108ED838380282FE00" box="[433,462,548,572]" captionStart="FIGURES 46 – 48" captionStartId="22.[151,264,1976,1999]" captionTargetBox="[162,1428,885,1944]" captionTargetId="figure@22.[151,1436,879,1955]" captionTargetPageId="22" captionText="FIGURES 46 – 48. Leascydmus spp. Dorsal habitus (46, 48) and original set of labels of holotype (47). Leascydmus simplicicornis (Lea) comb. n., female (46); Leascydmus armstrongi sp. n., male (48)." httpUri="https://zenodo.org/record/286524/files/figure.png" pageId="24" pageNumber="25">48</figureCitation>
|
||
) in dorsal view approximately subquadrate with rounded sides, broadest near anterior third, PL
|
||
<quantity id="4CA79B62FFCF18108E6C385B02E7FE63" box="[261,427,583,608]" metricMagnitude="-4" metricUnit="m" metricValue="5.8999999999999995" metricValueMax="6.0" metricValueMin="5.8" pageId="24" pageNumber="25" unit="mm" value="0.59" valueMax="0.6" valueMin="0.58">0.58–0.60 mm</quantity>
|
||
(mean
|
||
<quantity id="4CA79B62FFCF18108D6B385B0121FE5C" box="[514,621,583,608]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" pageId="24" pageNumber="25" unit="mm" value="0.58">0.58 mm</quantity>
|
||
), PW
|
||
<quantity id="4CA79B62FFCF18108DDC385B0052FE63" box="[693,798,583,608]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="24" pageNumber="25" unit="mm" value="0.55">0.55 mm</quantity>
|
||
; anterior margin nearly straight; anterior corners obtuse and blunt; lateral margins rounded; posterior corners strongly obtuse and indistinctly marked; posterior margin nearly straight; pronotal base with shallow and indistinct transverse impression. Punctures on pronotal disc similar to those on frons and vertex; setae sparse, short and suberect, near posterior corners distinctly denser and thicker.
|
||
</paragraph>
|
||
<paragraph id="8BE03687FFCF18108FAE38CB0193FFBC" blockId="24.[151,1437,151,1400]" pageId="24" pageNumber="25">
|
||
Elytra (
|
||
<figureCitation id="13642A02FFCF18108E7238CB0235FECC" box="[283,377,727,752]" captionStart="FIGURES 42 – 45" captionStartId="20.[151,264,1431,1454]" captionTargetBox="[151,1436,212,1404]" captionTargetId="figure@20.[151,1436,193,1411]" captionTargetPageId="20" captionText="FIGURES 42 – 45. Morphology of Leascydmus armstrongi sp. n. Base of pronotum and elytra in dorsal view (42), prothorax in ventral view (43), meso- and metanotum in dorsal view (44), pterothorax in ventral view (45). Abbreviations: aest 3, metanepisternum; alc, alacrista; amvp, anterior metaventral process; ar, anterior ridge; bef, basal elytral fovea; bst, basisternal part of prosternum; cx 3, metacoxa; dlf, dorsolateral fovea; elb, elytral base; hd, humeral denticle; hy, hypomeron; hyr, hypomeral ridge; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; mma, median membranous area; mscc, mesocoxal cavity; mscs, mesocoxal socket; msf, mesofurca; msvp, mesoventral process; mtvp, metaventral intercoxal process; nss, pronotosternal suture; pcc, procoxal cavity; pcr, procoxal rest; pcs, procoxal socket; pff, profurcal fovea; pre, prepectus; psn 3, metapostnotum; sc 2, mesoscutellum; sc 3, metascutum; sss, mesoscutoscutellar suture; tr 3, metatrochanter; v 3, metaventrite; vlf, ventrolateral fovea; wb, wing base." httpUri="https://zenodo.org/record/286523/files/figure.png" pageId="24" pageNumber="25">Figs. 44</figureCitation>
|
||
,
|
||
<figureCitation id="13642A02FFCF18108EED38C402EDFECC" box="[388,417,728,752]" captionStart="FIGURES 46 – 48" captionStartId="22.[151,264,1976,1999]" captionTargetBox="[162,1428,885,1944]" captionTargetId="figure@22.[151,1436,879,1955]" captionTargetPageId="22" captionText="FIGURES 46 – 48. Leascydmus spp. Dorsal habitus (46, 48) and original set of labels of holotype (47). Leascydmus simplicicornis (Lea) comb. n., female (46); Leascydmus armstrongi sp. n., male (48)." httpUri="https://zenodo.org/record/286524/files/figure.png" pageId="24" pageNumber="25">48</figureCitation>
|
||
) oval, more convex than pronotum, broadest slightly anterior to middle, EL
|
||
<quantity id="4CA79B62FFCF18108B9338CB06D1FED3" box="[1274,1437,727,752]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" metricValueMax="1.45" metricValueMin="1.35" pageId="24" pageNumber="25" unit="mm" value="1.4" valueMax="1.45" valueMin="1.35">1.35–1.45 mm</quantity>
|
||
(mean
|
||
<quantity id="4CA79B62FFCF18108F8C38E00202FF28" box="[229,334,764,788]" metricMagnitude="-3" metricUnit="m" metricValue="1.42" pageId="24" pageNumber="25" unit="mm" value="1.42">1.42 mm</quantity>
|
||
), EW
|
||
<quantity id="4CA79B62FFCF18108EFC38E00175FF28" box="[405,569,764,788]" metricMagnitude="-4" metricUnit="m" metricValue="9.05" metricValueMax="9.8" metricValueMin="8.299999999999999" pageId="24" pageNumber="25" unit="mm" value="0.905" valueMax="0.98" valueMin="0.83">0.83–0.98 mm</quantity>
|
||
(mean
|
||
<quantity id="4CA79B62FFCF18108DE638E001B4FF28" box="[655,760,764,788]" metricMagnitude="-4" metricUnit="m" metricValue="9.1" pageId="24" pageNumber="25" unit="mm" value="0.91">0.91 mm</quantity>
|
||
), EI 1.49–1.64; basal impressions absent; humeral denticles tiny but distinct; elytral apices separately rounded. Punctures on elytral disc fine but slightly more distinct than those on pronotum, separated by spaces 2–
|
||
<date id="FFE11047FFCF18108D1E395801CFFF60" box="[631,643,836,860]" pageId="24" pageNumber="25">3</date>
|
||
x as wide as diameters of punctures; setae short, sparse and erect.
|
||
<collectingCountry id="F3487617FFCF18108A0D395806D0FF61" box="[1380,1436,836,861]" name="India" pageId="24" pageNumber="25">Hind</collectingCountry>
|
||
wings well developed, about twice as long as elytra.
|
||
</paragraph>
|
||
<paragraph id="8BE03687FFCF18108FAE3991078FFF99" blockId="24.[151,1437,151,1400]" box="[199,1219,908,933]" pageId="24" pageNumber="25">
|
||
Legs (
|
||
<figureCitation id="13642A02FFCF18108E6439900212FF98" box="[269,350,908,933]" captionStart="FIGURES 46 – 48" captionStartId="22.[151,264,1976,1999]" captionTargetBox="[162,1428,885,1944]" captionTargetId="figure@22.[151,1436,879,1955]" captionTargetPageId="22" captionText="FIGURES 46 – 48. Leascydmus spp. Dorsal habitus (46, 48) and original set of labels of holotype (47). Leascydmus simplicicornis (Lea) comb. n., female (46); Leascydmus armstrongi sp. n., male (48)." httpUri="https://zenodo.org/record/286524/files/figure.png" pageId="24" pageNumber="25">Fig. 48</figureCitation>
|
||
) long and slender, without modifications; all tibiae broadest in proximal half.
|
||
</paragraph>
|
||
<paragraph id="8BE03687FFCF18108FAE39B30713F833" blockId="24.[151,1437,151,1400]" pageId="24" pageNumber="25">
|
||
Aedeagus (
|
||
<figureCitation id="13642A02FFCF18108E2E39B3029EFFF4" box="[327,466,943,968]" captionStart="FIGURES 49 – 52" captionStartId="23.[151,264,891,914]" captionTargetBox="[167,1425,211,862]" captionTargetId="figure@23.[151,1436,193,870]" captionTargetPageId="23" captionText="FIGURES 49 – 52. Aedeagi of Leascydmus spp. in ventral (49, 51) and lateral (50, 52) views. Leascydmus simplicicornis (Lea) comb. n. (49 – 50), Leascydmus armstrongi sp. n. (51 – 52)." httpUri="https://zenodo.org/record/286525/files/figure.png" pageId="24" pageNumber="25">Figs. 51–52</figureCitation>
|
||
,
|
||
<figureCitation id="13642A02FFCF18108E8939B30164FFF4" box="[480,552,943,968]" captionStart="FIGURES 53 – 56" captionStartId="23.[151,264,1710,1733]" captionTargetBox="[163,1428,1002,1677]" captionTargetId="figure@23.[151,1436,994,1689]" captionTargetPageId="23" captionText="FIGURES 53 – 56. Aedeagi of Leascydmus spp. in ventral (53, 55) and lateral (54, 56) views. Leascydmus simplicicornis (Lea) comb. n. (53 – 54), Leascydmus armstrongi sp. n. (55 – 56)." httpUri="https://zenodo.org/record/286526/files/figure.png" pageId="24" pageNumber="25">55–56</figureCitation>
|
||
) elongate, darkly sclerotized, AeL
|
||
<quantity id="4CA79B62FFCF18108CA939AC0765FFFB" box="[960,1065,944,968]" metricMagnitude="-4" metricUnit="m" metricValue="7.3" pageId="24" pageNumber="25" unit="mm" value="0.73">0.73 mm</quantity>
|
||
, in ventral view apex of ventral wall subtrapezoidal and apex of dorsal wall rapidly narrowing; internal armature composed of a complicated set of elongate sclerites; parameres slender, with moderately long apical and subapical setae.
|
||
</paragraph>
|
||
<paragraph id="8BE03687FFCF18108FAE3E000242F89C" blockId="24.[151,1437,151,1400]" pageId="24" pageNumber="25">
|
||
<emphasis id="B92BEA95FFCF18108FAE3E000252F809" bold="true" box="[199,286,1052,1077]" pageId="24" pageNumber="25">Female</emphasis>
|
||
. Similar to male; BL
|
||
<quantity id="4CA79B62FFCF18108D433E00019EF808" box="[554,722,1052,1076]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" metricValueMax="2.4" metricValueMin="2.2" pageId="24" pageNumber="25" unit="mm" value="2.3" valueMax="2.4" valueMin="2.2">2.20–2.40 mm</quantity>
|
||
(mean
|
||
<quantity id="4CA79B62FFCF18108C473E0000D7F808" box="[814,923,1052,1076]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="24" pageNumber="25" unit="mm" value="2.3">2.30 mm</quantity>
|
||
); HL
|
||
<quantity id="4CA79B62FFCF18108C8C3E0007C0F808" box="[997,1164,1052,1076]" metricMagnitude="-4" metricUnit="m" metricValue="4.15" metricValueMax="4.3" metricValueMin="4.0" pageId="24" pageNumber="25" unit="mm" value="0.415" valueMax="0.43" valueMin="0.4">0.40–0.43 mm</quantity>
|
||
(mean
|
||
<quantity id="4CA79B62FFCF18108B803E00061AF808" box="[1257,1366,1052,1076]" metricMagnitude="-4" metricUnit="m" metricValue="4.2" pageId="24" pageNumber="25" unit="mm" value="0.42">0.42 mm</quantity>
|
||
), HW
|
||
<quantity id="4CA79B62FFCF18108FFE3E230270F86B" box="[151,316,1087,1112]" metricMagnitude="-4" metricUnit="m" metricValue="4.9" metricValueMax="5.0" metricValueMin="4.8" pageId="24" pageNumber="25" unit="mm" value="0.49" valueMax="0.5" valueMin="0.48">0.48–0.50 mm</quantity>
|
||
(mean
|
||
<quantity id="4CA79B62FFCF18108EFD3E5C02B2F864" box="[404,510,1088,1112]" metricMagnitude="-4" metricUnit="m" metricValue="4.9" pageId="24" pageNumber="25" unit="mm" value="0.49">0.49 mm</quantity>
|
||
), AnL
|
||
<quantity id="4CA79B62FFCF18108D263E2301B8F86B" box="[591,756,1087,1112]" metricMagnitude="-3" metricUnit="m" metricValue="1.705" metricValueMax="1.78" metricValueMin="1.63" pageId="24" pageNumber="25" unit="mm" value="1.705" valueMax="1.78" valueMin="1.63">1.63–1.78 mm</quantity>
|
||
(mean
|
||
<quantity id="4CA79B62FFCF18108C253E2300FBF864" box="[844,951,1087,1112]" metricMagnitude="-3" metricUnit="m" metricValue="1.68" pageId="24" pageNumber="25" unit="mm" value="1.68">1.68 mm</quantity>
|
||
); PL
|
||
<quantity id="4CA79B62FFCF18108C9F3E2307D7F86B" box="[1014,1179,1087,1112]" metricMagnitude="-4" metricUnit="m" metricValue="5.4" metricValueMax="5.5" metricValueMin="5.3" pageId="24" pageNumber="25" unit="mm" value="0.54" valueMax="0.55" valueMin="0.53">0.53–0.55 mm</quantity>
|
||
(mean
|
||
<quantity id="4CA79B62FFCF18108B9A3E230611F864" box="[1267,1373,1087,1112]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" pageId="24" pageNumber="25" unit="mm" value="0.53">0.53 mm</quantity>
|
||
), PW
|
||
<quantity id="4CA79B62FFCF18108FFE3E780271F840" box="[151,317,1124,1149]" metricMagnitude="-4" metricUnit="m" metricValue="5.4" metricValueMax="5.5" metricValueMin="5.3" pageId="24" pageNumber="25" unit="mm" value="0.54" valueMax="0.55" valueMin="0.53">0.53–0.55 mm</quantity>
|
||
(mean
|
||
<quantity id="4CA79B62FFCF18108EF13E78014FF840" box="[408,515,1124,1149]" metricMagnitude="-4" metricUnit="m" metricValue="5.4" pageId="24" pageNumber="25" unit="mm" value="0.54">0.54 mm</quantity>
|
||
); EL
|
||
<quantity id="4CA79B62FFCF18108D2E3E7801A1F840" box="[583,749,1124,1149]" metricMagnitude="-3" metricUnit="m" metricValue="1.3399999999999999" metricValueMax="1.43" metricValueMin="1.25" pageId="24" pageNumber="25" unit="mm" value="1.3399999999999999" valueMax="1.43" valueMin="1.25">1.25–1.43 mm</quantity>
|
||
(mean
|
||
<quantity id="4CA79B62FFCF18108C213E7800FFF840" box="[840,947,1124,1149]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" pageId="24" pageNumber="25" unit="mm" value="1.35">1.35 mm</quantity>
|
||
), EW
|
||
<quantity id="4CA79B62FFCF18108C963E7807EAF840" box="[1023,1190,1124,1148]" metricMagnitude="-4" metricUnit="m" metricValue="9.8" metricValueMax="10.3" metricValueMin="9.3" pageId="24" pageNumber="25" unit="mm" value="0.98" valueMax="1.03" valueMin="0.93">0.93–1.03 mm</quantity>
|
||
(mean
|
||
<quantity id="4CA79B62FFCF18108A693E780620F840" box="[1280,1388,1124,1148]" metricMagnitude="-4" metricUnit="m" metricValue="9.8" pageId="24" pageNumber="25" unit="mm" value="0.98">0.98 mm</quantity>
|
||
), EI 1.22–1.49.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C345650CFFCF18108FAE3EB007F8F8F9" box="[199,1204,1196,1221]" pageId="24" pageNumber="25" type="distribution">
|
||
<paragraph id="8BE03687FFCF18108FAE3EB007F8F8F9" blockId="24.[151,1437,151,1400]" box="[199,1204,1196,1221]" pageId="24" pageNumber="25">
|
||
<emphasis id="B92BEA95FFCF18108FAE3EB00215F8F9" bold="true" box="[199,345,1196,1221]" pageId="24" pageNumber="25">Distribution</emphasis>
|
||
(
|
||
<figureCitation id="13642A02FFCF18108E013EB00286F8F9" box="[360,458,1196,1221]" captionStart="FIGURES 193 – 198" captionStartId="72.[151,264,1762,1785]" captionTargetBox="[195,1376,193,1707]" captionTargetId="figure@72.[191,1378,193,1739]" captionTargetPageId="72" captionText="FIGURES 193 – 198. Distribution of revised Australian species of Horaeomorphus Schaufuss (193), Leascydmus gen. n. (194), Scydmaenozila gen. n. (195), Spinosciacharis gen. n. (196), Sciacharoides gen. n. (197), and Sciacharis Broun. Horaeomorphus anthicoides (Lea) comb. n. (193 a), Horaeomorphus verus sp. n. (193 b), Leascydmus simplicicornis (Lea) comb. n. (194 a), Leascydmus armstrongi sp. n. (194 b), Scydmaenozila macrosticta (Lea) comb. n. (195 a), Scydmaenozila latipennis (Lea) comb. n. (195 b), Scydmaenozila mallacootana sp. n. (195 c), Scydmaenozila inexpectata sp. n. (195 d), Spinosciacharis puncticeps (Franz) comb. n. (196 a), Spinosciacharis sassy sp. n. (196 b), Spinosciacharis clydensis sp. n. (196 c), Spinosciacharis acheron sp. n. (196 d), Sciacharoides thompsoni (Franz) comb. n. (197 a), Sciacharoides eucalypti (Franz) comb. n. (197 b), Sciacharoides maipotonensis (Franz) comb. n. (197 c), Sciacharoides mantistamborinensis (Franz) comb. n. (197 d), and Sciacharis (Maorinus) tasmaniensis (Franz) comb. n. (198)." httpUri="https://zenodo.org/record/286555/files/figure.png" pageId="24" pageNumber="25">Fig. 194</figureCitation>
|
||
b). South-eastern
|
||
<collectingCountry id="F3487617FFCF18108DE43EB001B8F8F9" box="[653,756,1196,1221]" name="Australia" pageId="24" pageNumber="25">Australia</collectingCountry>
|
||
(Blue Mountains in New South
|
||
<collectingCountry id="F3487617FFCF18108B093EB007E5F8F9" box="[1120,1193,1196,1221]" name="United Kingdom" pageId="24" pageNumber="25">Wales</collectingCountry>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C345650CFFCF18108FAE3ED301FAF931" pageId="24" pageNumber="25" type="etymology">
|
||
<paragraph id="8BE03687FFCF18108FAE3ED301FAF931" blockId="24.[151,1437,151,1400]" pageId="24" pageNumber="25">
|
||
<emphasis id="B92BEA95FFCF18108FAE3ED30202F8D4" bold="true" box="[199,334,1231,1256]" pageId="24" pageNumber="25">Etymology.</emphasis>
|
||
<taxonomicName id="4C5F4D04FFCF18108E3D3ED3012CF8D4" box="[340,608,1231,1256]" class="Insecta" family="Staphylinidae" genus="Leascydmus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="armstrongi">
|
||
<emphasis id="B92BEA95FFCF18108E3D3ED3012CF8D4" box="[340,608,1231,1256]" italics="true" pageId="24" pageNumber="25">Leascydmus armstrongi</emphasis>
|
||
</taxonomicName>
|
||
is dedicated to J. W. T. Armstrong, who described many Australian beetle taxa and collected the
|
||
<typeStatus id="54E48825FFCF18108EF83EE9028DF931" box="[401,449,1269,1293]" pageId="24" pageNumber="25">type</typeStatus>
|
||
series of this species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C345650CFFCF18108FAE3F0B074FF944" pageId="24" pageNumber="25" type="discussion">
|
||
<paragraph id="8BE03687FFCF18108FAE3F0B074FF944" blockId="24.[151,1437,151,1400]" pageId="24" pageNumber="25">
|
||
<emphasis id="B92BEA95FFCF18108FAE3F0B0277F90C" bold="true" box="[199,315,1303,1328]" pageId="24" pageNumber="25">Remarks.</emphasis>
|
||
This species is highly similar to
|
||
<taxonomicName id="4C5F4D04FFCF18108DC63F050020F90C" box="[687,876,1303,1328]" class="Insecta" family="Staphylinidae" genus="Leascydmus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="simplicicornis">
|
||
<emphasis id="B92BEA95FFCF18108DC63F050020F90C" box="[687,876,1303,1328]" italics="true" pageId="24" pageNumber="25">L. simplicicornis</emphasis>
|
||
</taxonomicName>
|
||
in the body shape, proportions of body parts, and even in the aedeagus. The striking difference in pigmentation, smaller body and the antennae distinctly longer in relation to body allow for unambiguous discrimination from
|
||
<taxonomicName id="4C5F4D04FFCF18108C293F7D00B0F944" box="[832,1020,1375,1400]" class="Insecta" family="Staphylinidae" genus="Leascydmus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="simplicicornis">
|
||
<emphasis id="B92BEA95FFCF18108C293F7D00B0F944" box="[832,1020,1375,1400]" italics="true" pageId="24" pageNumber="25">L. simplicicornis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |