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<document id="4D53A83C5F893F12CAD7CFEDABF1151C" ID-CLB-Dataset="50790" ID-DOI="10.5281/zenodo.172376" ID-GBIF-Dataset="54e48000-e28d-42b8-993e-d39b4923683f" ID-ISSN="1175­5326" ID-Zenodo-Dep="172376" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459755737607" checkinUser="plazi" docAuthor="Johanson, Kjell Arne &amp; Oláh, János" docDate="2006" docId="03F512008347FFFDFED2E6CBFCF4AC8F" docLanguage="en" docName="zt01205p030.pdf" docOrigin="Zootaxa 1205" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Helicopsyche (Petrotrichia) ninakosha Johanson &amp; Oláh, 2006, new species" docType="treatment" docVersion="11" lastPageNumber="9" masterDocId="FFCC6A788340FFF5FFDAE437FFE5AE11" masterDocTitle="Eleven new species of Sericostomatoidea from Madagascar (Trichoptera: Helicopsychidae, Petrothrincidae, Sericostomatidae)" masterLastPageNumber="30" masterPageNumber="1" pageNumber="8" updateTime="1698217669661" updateUser="ExternalLinkService">
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<mods:title id="EB43CB46C3BC1AEDCBAE63ED86F39DC8">Eleven new species of Sericostomatoidea from Madagascar (Trichoptera: Helicopsychidae, Petrothrincidae, Sericostomatidae)</mods:title>
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<mods:name id="223D736C220A89A412F76589F67E25A4" type="personal">
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<mods:namePart id="D34F0E458DFB079CAB56E9B00AB5E01D">Johanson, Kjell Arne</mods:namePart>
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<mods:namePart id="10354E53727B331F744DD47C7D9BDFBF">Oláh, János</mods:namePart>
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<mods:date id="2D5A5D26403DB95D9B3B740A33EDFDC5">2006</mods:date>
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<treatment id="03F512008347FFFDFED2E6CBFCF4AC8F" ID-DOI="http://doi.org/10.5281/zenodo.5049122" ID-GBIF-Taxon="119343616" ID-Zenodo-Dep="5049122" LSID="urn:lsid:plazi:treatment:03F512008347FFFDFED2E6CBFCF4AC8F" httpUri="http://treatment.plazi.org/id/03F512008347FFFDFED2E6CBFCF4AC8F" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
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<paragraph id="8BE3A3168347FFF2FED2E6CBFC9FAD07" blockId="7.[264,890,764,825]" box="[264,890,764,790]" pageId="7" pageNumber="8">
<heading id="D0AB147A8347FFF2FED2E6CBFC9FAD07" bold="true" box="[264,890,764,790]" fontSize="11" level="1" pageId="7" pageNumber="8" reason="1">
<emphasis id="B9287F048347FFF2FED2E6CBFC9FAD07" bold="true" box="[264,890,764,790]" pageId="7" pageNumber="8">
<taxonomicName id="4C5CD8958347FFF2FED2E6CBFD3EAD07" box="[264,731,764,790]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="ninakosha" status="sp. nov." subGenus="Petrotrichia">
<emphasis id="B9287F048347FFF2FED2E6CBFD3EAD07" bold="true" box="[264,731,764,790]" italics="true" pageId="7" pageNumber="8">Helicopsyche (Petrotrichia) ninakosha</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A21BC27F8347FFF2FD30E6CBFC9FAD07" box="[746,890,764,790]" pageId="7" pageNumber="8" rank="species">new species</taxonomicNameLabel>
</emphasis>
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<figureCitation id="1367BF938347FFF2FED2E728FE68AD28" box="[264,397,799,825]" captionStart="FIGURES 12 16" captionStartId="8.[264,383,1693,1717]" captionTargetBox="[288,1308,728,1672]" captionTargetId="figure@8.[268,1327,704,1690]" captionTargetPageId="8" captionText="FIGURES 12 16. Helicopsyche (Petrotrichia) ninakosha, new species, holotype. 12 — head and thorax, dorsal; 13 — right wing pair; 14 — genitalia, lateral; 15 — genitalia, dorsal; 16 — genitalia, ventral." httpUri="https://zenodo.org/record/172379/files/figure.png" pageId="7" pageNumber="8">Figs 1216</figureCitation>
,
<figureCitation id="1367BF938347FFF2FE40E728FE52AD28" box="[410,439,799,825]" captionStart="FIGURE 58" captionStartId="23.[264,368,1765,1789]" captionTargetBox="[418,1164,324,1727]" captionTargetId="figure@23.[396,1191,307,1742]" captionTargetPageId="23" captionText="FIGURE 58. Map of Madagascar showing distributions of Helicopsyche spp. (Helicopsychidae): H. ambodiva (circle); H. ninakosha and H. sahadika (rectangle); H. giboni (diamond); and H. anomana (triangle)." httpUri="https://zenodo.org/record/172389/files/figure.png" pageId="7" pageNumber="8">58</figureCitation>
</paragraph>
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<subSubSection id="C346F09D8347FFFDFED2E758FCF4AC8F" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="description">
<paragraph id="8BE3A3168347FFF2FED2E758FE58AAB0" blockId="7.[264,1323,879,1905]" pageId="7" pageNumber="8">
This new species is morphologically close to
<taxonomicName id="4C5CD8958347FFF2FCC6E758FC4CAD99" box="[796,937,879,904]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="sahadika" status="sp. nov.">
<emphasis id="B9287F048347FFF2FCC6E758FC4CAD99" box="[796,937,879,904]" italics="true" pageId="7" pageNumber="8">H. sahadika</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A21BC27F8347FFF2FC62E758FBA0AD98" box="[952,1093,879,905]" pageId="7" pageNumber="8" rank="species">new species</taxonomicNameLabel>
in that the pronotal setal warts are similarly modified into a single, wide wart. In genitalic features, the two species are very similar: both have segment IX highly reduced, forming a very narrow plate ventrally; they have Xth tergum simple, a strong and very long spine apically on the gonocoxite, and the ventral branch of the gonocoxite modified. The last similarity is, however, very differently expressed in the two species and they are easily distinguished from each other by the shape of the ventral branch of the gonocoxite in both lateral and ventral aspects.
</paragraph>
<paragraph id="8BE3A3168347FFF2FEE2E098FD8FAB80" blockId="7.[264,1323,879,1905]" pageId="7" pageNumber="8">
Male head and thorax (
<figureCitation id="1367BF938347FFF2FD97E098FD43AAD8" box="[589,678,1199,1225]" captionStart="FIGURES 12 16" captionStartId="8.[264,383,1693,1717]" captionTargetBox="[288,1308,728,1672]" captionTargetId="figure@8.[268,1327,704,1690]" captionTargetPageId="8" captionText="FIGURES 12 16. Helicopsyche (Petrotrichia) ninakosha, new species, holotype. 12 — head and thorax, dorsal; 13 — right wing pair; 14 — genitalia, lateral; 15 — genitalia, dorsal; 16 — genitalia, ventral." httpUri="https://zenodo.org/record/172379/files/figure.png" pageId="7" pageNumber="8">Fig. 12</figureCitation>
). Scape about 2.5x longer than wide. Second segment of maxillary palp about
<date id="FFE285D68347FFF2FDE7E0E0FDBEAAE0" box="[573,603,1239,1265]" pageId="7" pageNumber="8">2x</date>
longer than basal segment (
<figureCitation id="1367BF938347FFF2FC1CE0E0FBC0AAE0" box="[966,1061,1239,1265]" captionStart="FIGURES 12 16" captionStartId="8.[264,383,1693,1717]" captionTargetBox="[288,1308,728,1672]" captionTargetId="figure@8.[268,1327,704,1690]" captionTargetPageId="8" captionText="FIGURES 12 16. Helicopsyche (Petrotrichia) ninakosha, new species, holotype. 12 — head and thorax, dorsal; 13 — right wing pair; 14 — genitalia, lateral; 15 — genitalia, dorsal; 16 — genitalia, ventral." httpUri="https://zenodo.org/record/172379/files/figure.png" pageId="7" pageNumber="8">Fig. 12</figureCitation>
). Interantennal warts slightly separate from each other, cephalic warts nearly triangular, one pair of postantennal warts small and rounded, and postocular warts not conspicuous. Pronotum with paired setal warts completely fused medially into a single transverse wart. One pair of oval setal warts present on mesoscutum.
</paragraph>
<paragraph id="8BE3A3168347FFF2FEE2E1A8FBC4A848" blockId="7.[264,1323,879,1905]" pageId="7" pageNumber="8">
Male wings (
<figureCitation id="1367BF938347FFF2FE0CE1A8FDCAABA8" box="[470,559,1439,1465]" captionStart="FIGURES 12 16" captionStartId="8.[264,383,1693,1717]" captionTargetBox="[288,1308,728,1672]" captionTargetId="figure@8.[268,1327,704,1690]" captionTargetPageId="8" captionText="FIGURES 12 16. Helicopsyche (Petrotrichia) ninakosha, new species, holotype. 12 — head and thorax, dorsal; 13 — right wing pair; 14 — genitalia, lateral; 15 — genitalia, dorsal; 16 — genitalia, ventral." httpUri="https://zenodo.org/record/172379/files/figure.png" pageId="7" pageNumber="8">Fig. 13</figureCitation>
). Forewing length 2.7 mm, hind wing length 2.2 mm. Denuded wing membrane pale flavous with veins pale helvus. Venation similar to
<taxonomicName id="4C5CD8958347FFF2FB4EE1F0FE90A818" authority="Johanson" authorityName="Johanson" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="anomana">
<emphasis id="B9287F048347FFF2FB4EE1F0FACEABF1" box="[1172,1323,1479,1504]" italics="true" pageId="7" pageNumber="8">H. anomana</emphasis>
Johanson
</taxonomicName>
and
<taxonomicName id="4C5CD8958347FFF2FE6DE1D8FD3CA818" authority="Johanson. On" authorityName="Johanson. On" box="[439,729,1519,1545]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="giboni">
<emphasis id="B9287F048347FFF2FE6DE1D8FDCFA819" box="[439,554,1519,1544]" italics="true" pageId="7" pageNumber="8">H. giboni</emphasis>
Johanson. On
</taxonomicName>
forewing R5 originating immediately distally to crossvein R3­R4. M and Cu separate a longitudinal groove containing sensilla and reaching the wing margin. Discoidal cell about 1/5 the wing length.
</paragraph>
<paragraph id="8BE3A3168347FFFDFEE2E250FAE1AFEF" blockId="7.[264,1323,879,1905]" lastBlockId="8.[264,1323,284,670]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
Male abdomen and genitalia (
<figureCitation id="1367BF938347FFF2FD4FE250FCFFA890" box="[661,794,1639,1665]" captionStart="FIGURES 12 16" captionStartId="8.[264,383,1693,1717]" captionTargetBox="[288,1308,728,1672]" captionTargetId="figure@8.[268,1327,704,1690]" captionTargetPageId="8" captionText="FIGURES 12 16. Helicopsyche (Petrotrichia) ninakosha, new species, holotype. 12 — head and thorax, dorsal; 13 — right wing pair; 14 — genitalia, lateral; 15 — genitalia, dorsal; 16 — genitalia, ventral." httpUri="https://zenodo.org/record/172379/files/figure.png" pageId="7" pageNumber="8">Fig. 1416</figureCitation>
). Abdominal reticulation absent. VIth sternal process well developed. Segment IX strongly narrowed anteroposteriorly and slightly thicker at dorsal half (
<figureCitation id="1367BF938347FFF2FDD7E280FD81A8C0" box="[525,612,1719,1745]" captionStart="FIGURES 12 16" captionStartId="8.[264,383,1693,1717]" captionTargetBox="[288,1308,728,1672]" captionTargetId="figure@8.[268,1327,704,1690]" captionTargetPageId="8" captionText="FIGURES 12 16. Helicopsyche (Petrotrichia) ninakosha, new species, holotype. 12 — head and thorax, dorsal; 13 — right wing pair; 14 — genitalia, lateral; 15 — genitalia, dorsal; 16 — genitalia, ventral." httpUri="https://zenodo.org/record/172379/files/figure.png" pageId="7" pageNumber="8">Fig. 14</figureCitation>
); anterior margin with short, slightly pointed dorsal anterior lobe; posterior margin slightly produced into triangle opposite to anterior lobe; narrow in ventral (
<figureCitation id="1367BF938347FFF2FEB0E330FE5AA930" box="[362,447,1799,1825]" captionStart="FIGURES 12 16" captionStartId="8.[264,383,1693,1717]" captionTargetBox="[288,1308,728,1672]" captionTargetId="figure@8.[268,1327,704,1690]" captionTargetPageId="8" captionText="FIGURES 12 16. Helicopsyche (Petrotrichia) ninakosha, new species, holotype. 12 — head and thorax, dorsal; 13 — right wing pair; 14 — genitalia, lateral; 15 — genitalia, dorsal; 16 — genitalia, ventral." httpUri="https://zenodo.org/record/172379/files/figure.png" pageId="7" pageNumber="8">Fig. 16</figureCitation>
) and dorsal view (
<figureCitation id="1367BF938347FFF2FD46E330FD16A930" box="[668,755,1799,1825]" captionStart="FIGURES 12 16" captionStartId="8.[264,383,1693,1717]" captionTargetBox="[288,1308,728,1672]" captionTargetId="figure@8.[268,1327,704,1690]" captionTargetPageId="8" captionText="FIGURES 12 16. Helicopsyche (Petrotrichia) ninakosha, new species, holotype. 12 — head and thorax, dorsal; 13 — right wing pair; 14 — genitalia, lateral; 15 — genitalia, dorsal; 16 — genitalia, ventral." httpUri="https://zenodo.org/record/172379/files/figure.png" pageId="7" pageNumber="8">Fig. 15</figureCitation>
). Superior appendage fused with tergum IX and originating at posterodorsal corner of segment IX, finger­like, straight and oriented posterodorsad. Tergum X forms a pair of diverging, falciform plates, well separated in dorsal view (
<figureCitation id="1367BF938348FFFDFE7EE52BFE1EAF27" box="[420,507,284,310]" captionStart="FIGURES 12 16" captionStartId="8.[264,383,1693,1717]" captionTargetBox="[288,1308,728,1672]" captionTargetId="figure@8.[268,1327,704,1690]" captionTargetPageId="8" captionText="FIGURES 12 16. Helicopsyche (Petrotrichia) ninakosha, new species, holotype. 12 — head and thorax, dorsal; 13 — right wing pair; 14 — genitalia, lateral; 15 — genitalia, dorsal; 16 — genitalia, ventral." httpUri="https://zenodo.org/record/172379/files/figure.png" pageId="8" pageNumber="9">Fig. 15</figureCitation>
), inner lobe tapering in lateral aspect (
<figureCitation id="1367BF938348FFFDFC11E52BFBC0AF27" box="[971,1061,284,310]" captionStart="FIGURES 12 16" captionStartId="8.[264,383,1693,1717]" captionTargetBox="[288,1308,728,1672]" captionTargetId="figure@8.[268,1327,704,1690]" captionTargetPageId="8" captionText="FIGURES 12 16. Helicopsyche (Petrotrichia) ninakosha, new species, holotype. 12 — head and thorax, dorsal; 13 — right wing pair; 14 — genitalia, lateral; 15 — genitalia, dorsal; 16 — genitalia, ventral." httpUri="https://zenodo.org/record/172379/files/figure.png" pageId="8" pageNumber="9">Fig. 14</figureCitation>
). Gonocoxite primary branch forms a simple, weakly curving rod, its distal end armed with a subapical strong and long spine. The ventral branch of the gonocoxite forms a long, parallel­sided process, spiralling strongly towards and above phallus (
<figureCitation id="1367BF938348FFFDFCF6E5A3FC50AFBF" box="[812,949,404,430]" captionStart="FIGURES 12 16" captionStartId="8.[264,383,1693,1717]" captionTargetBox="[288,1308,728,1672]" captionTargetId="figure@8.[268,1327,704,1690]" captionTargetPageId="8" captionText="FIGURES 12 16. Helicopsyche (Petrotrichia) ninakosha, new species, holotype. 12 — head and thorax, dorsal; 13 — right wing pair; 14 — genitalia, lateral; 15 — genitalia, dorsal; 16 — genitalia, ventral." httpUri="https://zenodo.org/record/172379/files/figure.png" pageId="8" pageNumber="9">Figs 14, 16</figureCitation>
). Phallus basally broad, divided into a nearly uniformly thick ventral process and a dorsal branch that is about
<date id="FFE285D68348FFFDFB71E58BFB2DAFC7" box="[1195,1224,444,470]" pageId="8" pageNumber="9">2x</date>
broader than the ventral process and has three subapical setae below the spinose apex (
<figureCitation id="1367BF938348FFFDFB45E5D3FB12AFEF" box="[1183,1271,484,510]" captionStart="FIGURES 12 16" captionStartId="8.[264,383,1693,1717]" captionTargetBox="[288,1308,728,1672]" captionTargetId="figure@8.[268,1327,704,1690]" captionTargetPageId="8" captionText="FIGURES 12 16. Helicopsyche (Petrotrichia) ninakosha, new species, holotype. 12 — head and thorax, dorsal; 13 — right wing pair; 14 — genitalia, lateral; 15 — genitalia, dorsal; 16 — genitalia, ventral." httpUri="https://zenodo.org/record/172379/files/figure.png" pageId="8" pageNumber="9">Fig. 14</figureCitation>
).
</paragraph>
<paragraph id="8BE3A3168348FFFDFEE2E63BFCBCAC5F" blockId="8.[264,1323,284,670]" pageId="8" pageNumber="9">
Etymology. Named after the very specialised, spiralling shape of the ventral branch of gonocoxite. In Sanscrit, “
<taxonomicName id="4C5CD8958348FFFDFDEAE603FD48AC5F" box="[560,685,564,590]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="ninakosha">ninakosha</taxonomicName>
” means spiral.
</paragraph>
<paragraph id="8BE3A3168348FFFDFEE2E66BFCF4AC8F" blockId="8.[264,1323,284,670]" pageId="8" pageNumber="9">
<typeStatus id="54E71DB48348FFFDFEE2E66BFE40AC67" box="[312,421,604,630]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
male:
<collectingCountry id="F34BE3868348FFFDFE24E66BFD36AC67" box="[510,723,604,630]" name="Madagascar" pageId="8" pageNumber="9">MADAGASCAR</collectingCountry>
: Demongloy, Mananteina Fort, Dauphin District.
<date id="FFE285D68348FFFDFED2E6B3FE92AC8F" box="[264,375,644,670]" pageId="8" pageNumber="9" value="1948-12">Xii. 1948</date>
[Hoogstraal] (MNHN, in alcohol).
</paragraph>
</subSubSection>
</treatment>
</document>