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<document id="03FE4D2627708B5A98037C2E8904D286" ID-CLB-Dataset="3084" ID-DOI="10.11646/zootaxa.5076.1.6" ID-GBIF-Dataset="ab615a5f-4aa3-4d62-88dd-a05e7c613727" ID-ISSN="1175-5326" ID-Zenodo-Dep="5763298" ID-ZooBank="043A7E1F-451B-4B8E-B1B9-6B69DBA6C3D5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638862753845" checkinUser="plazi" docAuthor="Orlando, Thales Yann, Salles, Frederico Falcão, Boldrini, Rafael &amp; Krolow, Tiago Kütter" docDate="2021" docId="03F2013D0313FF9876AEFDE0FAC4F9D0" docLanguage="en" docName="zootaxa.5076.1.6.pdf" docOrigin="Zootaxa 5076 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Thraulodes solstitium Orlando &amp; Salles &amp; Boldrini &amp; Krolow 2021, sp. nov." docType="treatment" docVersion="5" lastPageNumber="48" masterDocId="FFCB7945031BFF917639FFC2FFC3FFD9" masterDocTitle="Updated records for Leptophlebiidae (Ephemeroptera) and a new species of Thraulodes Ulmer, 1920 from Tocantins State, Northern Brazil" masterLastPageNumber="55" masterPageNumber="39" pageNumber="47" updateTime="1699472034857" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="07560A2E8E63631803BEC43705C12A14">Updated records for Leptophlebiidae (Ephemeroptera) and a new species of Thraulodes Ulmer, 1920 from Tocantins State, Northern Brazil</mods:title>
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<mods:name id="A9F8A5AFFCB844D6D2938E4FB3118E24" type="personal">
<mods:role id="58BBE75427A64D87E8AFB9D000069011">
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<mods:namePart id="DE435CA8F3417407DC77A9CB7C9E6214">Orlando, Thales Yann</mods:namePart>
<mods:nameIdentifier id="F39CE8B57D3012CAF2FC3BD6296FE765" type="ORCID">0000-0002-0438-3796</mods:nameIdentifier>
<mods:affiliation id="AA687B3F91CCF152225C6E86E7B97F3A">Museu de Entomologia, Departamento de Entomologia, Universidade Federal de Viçosa, Av. P. H. Rolfs, s / n, Campus Universitário, CEP 36570 - 900, Viçosa, Minas Gerais, Brazil.</mods:affiliation>
<mods:nameIdentifier id="E34CC18995056D9EE8D528E4E417063A" type="email">thalesyann@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="3CB671858345D771CF1DCD1F2DD29103" type="personal">
<mods:role id="7C633CD7CB7C88AAF614812CB7376EB3">
<mods:roleTerm id="84490398EFC4044EFEFF288C54DA596B">Author</mods:roleTerm>
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<mods:namePart id="8BDAD8C8E26872A95582A48BFCCBE7DB">Salles, Frederico Falcão</mods:namePart>
<mods:affiliation id="B4EA0B733D6F7439E4B765C5F2E6EA28">Museu de Entomologia, Departamento de Entomologia, Universidade Federal de Viçosa, Av. P. H. Rolfs, s / n, Campus Universitário, CEP 36570 - 900, Viçosa, Minas Gerais, Brazil.</mods:affiliation>
</mods:name>
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<mods:role id="DFCD9DF34CB37754B2456DFCEB607A2D">
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<mods:namePart id="80FFB1A673BF6F059CA4FBF91BBECE50">Boldrini, Rafael</mods:namePart>
<mods:affiliation id="B2A4C5E1CA091DE8C4E71B1FDBEED9B4">Universidade Federal de Roraima (UFRR), Campus Paricarana, Centro de Estudos da Biodiversidade (CBio), Laboratório de Entomologia, 69310 - 000, Boa Vista, RR, Brazil.</mods:affiliation>
</mods:name>
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<mods:role id="B0345C5CEDE53D77BBA7455382CF47F4">
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<mods:namePart id="00B70CD6F628C1922633FF7D7DB2C229">Krolow, Tiago Kütter</mods:namePart>
<mods:affiliation id="C49705B60C81BEAAEA44FD93AAB2C86F">Universidade Federal do Tocantins - UFT, Programa de Pós Graduação em Biodiversidade, Ecologia e Conservação, Cx. Postal 136, 77500 - 000, Porto Nacional, TO, Brazil.</mods:affiliation>
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<mods:title id="F1E151823E97D4BA5D8DD5700F768007">Zootaxa</mods:title>
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<mods:date id="BBEFC4E8F51E1F608F71DBAD31683027">2021</mods:date>
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<mods:number id="91D83AD288B35E2639DC9A161678CB70">2021-12-07</mods:number>
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<mods:number id="FF86EABB3D048BA694546AD3D5DA10D5">5076</mods:number>
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<treatment id="03F2013D0313FF9876AEFDE0FAC4F9D0" ID-DOI="http://doi.org/10.5281/zenodo.5763319" ID-GBIF-Taxon="191197910" ID-Zenodo-Dep="5763319" LSID="urn:lsid:plazi:treatment:03F2013D0313FF9876AEFDE0FAC4F9D0" httpUri="http://treatment.plazi.org/id/03F2013D0313FF9876AEFDE0FAC4F9D0" lastPageId="9" lastPageNumber="48" pageId="8" pageNumber="47">
<subSubSection id="C341E3A00313FF9976AEFDE0FD4EFDE5" box="[151,653,546,573]" pageId="8" pageNumber="47" type="nomenclature">
<paragraph id="8BE4B02B0313FF9976AEFDE0FD4EFDE5" blockId="8.[151,653,546,573]" box="[151,653,546,573]" pageId="8" pageNumber="47">
<heading id="D0AC07470313FF9976AEFDE0FD4EFDE5" bold="true" box="[151,653,546,573]" fontSize="11" level="1" pageId="8" pageNumber="47" reason="1">
<emphasis id="B92F6C390313FF9976AEFDE0FD4EFDE5" bold="true" box="[151,653,546,573]" pageId="8" pageNumber="47">
<taxonomicName id="4C5BCBA80313FF9976AEFDE0FE5DFDE4" authority="Orlando &amp; Salles &amp; Boldrini &amp; Krolow, 2021" authorityName="Orlando &amp; Salles &amp; Boldrini &amp; Krolow" authorityYear="2021" box="[151,414,546,573]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="47" phylum="Arthropoda" rank="species" species="solstitium" status="sp. nov.">
<emphasis id="B92F6C390313FF9976AEFDE0FE5DFDE4" bold="true" box="[151,414,546,573]" italics="true" pageId="8" pageNumber="47">Thraulodes solstitium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21CD1420313FF99779FFDE0FDC0FDE5" box="[422,515,546,572]" pageId="8" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="1360ACAE0313FF99742AFDE0FD46FDE5" box="[531,645,546,572]" captionStart-0="FIGURES 2–4" captionStart-1="FIGURES 5–8" captionStartId-0="10.[151,275,1941,1967]" captionStartId-1="11.[151,275,1942,1968]" captionTargetBox-0="[260,1326,321,1896]" captionTargetBox-1="[239,1351,193,1910]" captionTargetId-0="figure-25@10.[259,1328,179,1918]" captionTargetId-1="figure-19@11.[221,1366,179,1918]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="FIGURES 2–4. Thraulodes solstitium sp. nov., male imago: 2) habitus, (dorsal view); 3) head and thorax (lateral view); 4) abdomen (lateral view)." captionText-1="FIGURES 5–8. Thraulodes solstitium sp. nov., male imago: 5) forewing; 6a) hind wing; 6b) hind wing (enlarged); 7) genitalia, ventral view; 8) genitalia, dorsal view." figureDoi-0="http://doi.org/10.5281/zenodo.5763306" figureDoi-1="http://doi.org/10.5281/zenodo.5763310" httpUri-0="https://zenodo.org/record/5763306/files/figure.png" httpUri-1="https://zenodo.org/record/5763310/files/figure.png" pageId="8" pageNumber="47">Figs. 2–8</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C341E3A00313FF9976AEFDA8FCF4FC84" pageId="8" pageNumber="47" type="diagnosis">
<paragraph id="8BE4B02B0313FF9976AEFDA8FCF4FC84" blockId="8.[151,1437,618,861]" pageId="8" pageNumber="47">
<emphasis id="B92F6C390313FF9976AEFDA8FEC9FD5D" bold="true" box="[151,266,618,644]" pageId="8" pageNumber="47">Diagnosis</emphasis>
. Male imago: 1) costal cross veins basal to bulla either absent or colorless and very thin and 8 cross-veins distal (
<figureCitation id="1360ACAE0313FF9976DAFD4CFEEBFD70" box="[227,296,654,681]" captionStart="FIGURES 5–8" captionStartId="11.[151,275,1942,1968]" captionTargetBox="[239,1351,193,1910]" captionTargetId="figure-19@11.[221,1366,179,1918]" captionTargetPageId="11" captionText="FIGURES 5–8. Thraulodes solstitium sp. nov., male imago: 5) forewing; 6a) hind wing; 6b) hind wing (enlarged); 7) genitalia, ventral view; 8) genitalia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5763310" httpUri="https://zenodo.org/record/5763310/files/figure.png" pageId="8" pageNumber="47">Fig. 5</figureCitation>
); 2) area between C and Sc yellowish brown; 3) terga II–V yellowish white, with a pair of submedian blackish dots and one pair of sublateral blackish dots, terga VI anteriorly yellowish-white and posteriorly light brown, terga VII–X light brown (
<figureCitation id="1360ACAE0313FF99742AFD14FD9AFD29" box="[531,601,726,752]" captionStart="FIGURES 2–4" captionStartId="10.[151,275,1941,1967]" captionTargetBox="[260,1326,321,1896]" captionTargetId="figure-25@10.[259,1328,179,1918]" captionTargetPageId="10" captionText="FIGURES 2–4. Thraulodes solstitium sp. nov., male imago: 2) habitus, (dorsal view); 3) head and thorax (lateral view); 4) abdomen (lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5763306" httpUri="https://zenodo.org/record/5763306/files/figure.png" pageId="8" pageNumber="47">Fig. 2</figureCitation>
); 4) penes light-brown with blackish pigmentation on “ear” area (
<figureCitation id="1360ACAE0313FF99737EFD14FA4FFD29" box="[1351,1420,726,752]" captionStart="FIGURES 5–8" captionStartId="11.[151,275,1942,1968]" captionTargetBox="[239,1351,193,1910]" captionTargetId="figure-19@11.[221,1366,179,1918]" captionTargetPageId="11" captionText="FIGURES 5–8. Thraulodes solstitium sp. nov., male imago: 5) forewing; 6a) hind wing; 6b) hind wing (enlarged); 7) genitalia, ventral view; 8) genitalia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5763310" httpUri="https://zenodo.org/record/5763310/files/figure.png" pageId="8" pageNumber="47">Fig. 8</figureCitation>
); 5) styliger plate triangular, with a median rounded projection covering 2/3 of penes (
<figureCitation id="1360ACAE0313FF997272FD38FB4DFCCD" box="[1099,1166,762,788]" captionStart="FIGURES 5–8" captionStartId="11.[151,275,1942,1968]" captionTargetBox="[239,1351,193,1910]" captionTargetId="figure-19@11.[221,1366,179,1918]" captionTargetPageId="11" captionText="FIGURES 5–8. Thraulodes solstitium sp. nov., male imago: 5) forewing; 6a) hind wing; 6b) hind wing (enlarged); 7) genitalia, ventral view; 8) genitalia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5763310" httpUri="https://zenodo.org/record/5763310/files/figure.png" pageId="8" pageNumber="47">Fig. 8</figureCitation>
); 6) telopenes short and slender, 1.7 times the width of the medial region of the penes, lanceolate, narrowing towards the apex, arising on dorsal side of penes and directed median-anteriorly (
<figureCitation id="1360ACAE0313FF9974DDFC80FCEAFC84" box="[740,809,834,861]" captionStart="FIGURES 5–8" captionStartId="11.[151,275,1942,1968]" captionTargetBox="[239,1351,193,1910]" captionTargetId="figure-19@11.[221,1366,179,1918]" captionTargetPageId="11" captionText="FIGURES 5–8. Thraulodes solstitium sp. nov., male imago: 5) forewing; 6a) hind wing; 6b) hind wing (enlarged); 7) genitalia, ventral view; 8) genitalia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5763310" httpUri="https://zenodo.org/record/5763310/files/figure.png" pageId="8" pageNumber="47">Fig. 7</figureCitation>
).
</paragraph>
</subSubSection>
<subSection id="E2D4A9C00313FF9876AEFC6CFAC4F9D0" lastPageId="9" lastPageNumber="48" pageId="8" pageNumber="47" type="multiple">
<subSubSection id="C341E3A00313FF9976AEFC6CFEDDFBC9" pageId="8" pageNumber="47" type="multiple">
<paragraph id="8BE4B02B0313FF9976AEFC6CFEECFC11" blockId="8.[151,303,942,968]" box="[151,303,942,968]" pageId="8" pageNumber="47">
<heading id="D0AC07470313FF9976AEFC6CFEECFC11" bold="true" box="[151,303,942,968]" fontSize="11" level="1" pageId="8" pageNumber="47" reason="1">
<emphasis id="B92F6C390313FF9976AEFC6CFEECFC11" bold="true" box="[151,303,942,968]" pageId="8" pageNumber="47">Description.</emphasis>
</heading>
</paragraph>
<paragraph id="8BE4B02B0313FF9976AEFC34FEDDFBC9" blockId="8.[151,1437,1014,2013]" box="[151,286,1014,1040]" pageId="8" pageNumber="47">
<emphasis id="B92F6C390313FF9976AEFC34FEDDFBC9" box="[151,286,1014,1040]" italics="true" pageId="8" pageNumber="47">Male imago</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C341E3A00313FF9876FEFBD8FDDDFE74" lastPageId="9" lastPageNumber="48" pageId="8" pageNumber="47" type="description">
<paragraph id="8BE4B02B0313FF9976FEFBD8FE10FB80" blockId="8.[151,1437,1014,2013]" pageId="8" pageNumber="47">
Length of body:
<quantity id="4CA31DCE0313FF9977B8FBD8FDC9FBEC" box="[385,522,1050,1077]" metricMagnitude="-3" metricUnit="m" metricValue="6.05" metricValueMax="6.6" metricValueMin="5.5" pageId="8" pageNumber="47" unit="mm" value="6.05" valueMax="6.6" valueMin="5.5">5.5–6.6 mm</quantity>
; forewings:
<quantity id="4CA31DCE0313FF9974AAFBD8FCDEFBEC" box="[659,797,1050,1077]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" metricValueMax="6.4" metricValueMin="5.2" pageId="8" pageNumber="47" unit="mm" value="5.8" valueMax="6.4" valueMin="5.2">5.2–6.4 mm</quantity>
; hind wings:
<quantity id="4CA31DCE0313FF997588FBD8FBF9FBEC" box="[945,1082,1050,1077]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" metricValueMax="1.2" metricValueMin="0.9" pageId="8" pageNumber="47" unit="mm" value="1.05" valueMax="1.2" valueMin="0.9">0.9–1.2 mm</quantity>
. General coloration light brown with yellowish brown areas.
</paragraph>
<paragraph id="8BE4B02B0313FF9976FEFBA0FF3CFB1D" blockId="8.[151,1437,1014,2013]" pageId="8" pageNumber="47">
Head (
<figureCitation id="1360ACAE0313FF99772DFBA0FE46FBA5" box="[276,389,1122,1148]" captionStart="FIGURES 2–4" captionStartId="10.[151,275,1941,1967]" captionTargetBox="[260,1326,321,1896]" captionTargetId="figure-25@10.[259,1328,179,1918]" captionTargetPageId="10" captionText="FIGURES 2–4. Thraulodes solstitium sp. nov., male imago: 2) habitus, (dorsal view); 3) head and thorax (lateral view); 4) abdomen (lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5763306" httpUri="https://zenodo.org/record/5763306/files/figure.png" pageId="8" pageNumber="47">Figs. 2, 3</figureCitation>
). General coloration brown. Antenna with scapus light brown, pedicellus brown, flagellum light brown. Ocelli white, surrounded by black at base. Eyes black, turbinate portion orangish brown, contiguous medially.
</paragraph>
<paragraph id="8BE4B02B0313FF9976FEFB0CFC1CFAE9" blockId="8.[151,1437,1014,2013]" pageId="8" pageNumber="47">
Thorax (
<figureCitation id="1360ACAE0313FF997711FB0CFE57FB31" box="[296,404,1230,1256]" captionStart="FIGURES 2–4" captionStartId="10.[151,275,1941,1967]" captionTargetBox="[260,1326,321,1896]" captionTargetId="figure-25@10.[259,1328,179,1918]" captionTargetPageId="10" captionText="FIGURES 2–4. Thraulodes solstitium sp. nov., male imago: 2) habitus, (dorsal view); 3) head and thorax (lateral view); 4) abdomen (lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5763306" httpUri="https://zenodo.org/record/5763306/files/figure.png" pageId="8" pageNumber="47">Figs. 2, 3</figureCitation>
). General coloration brown. Prothorax whitish brown with pair of dark brown stripes transversal to coxa. Meso- and metanotum brown; prescutum and scutum light brown, scutellum dark brown. Pleura brown, membranous parts whitish. Sterna brown; basisternum light brown.
</paragraph>
<paragraph id="8BE4B02B0313FF9976FEFAF8FABEFA19" blockId="8.[151,1437,1014,2013]" pageId="8" pageNumber="47">
Wings (
<figureCitation id="1360ACAE0313FF997719FAF8FE5FFA8C" box="[288,412,1338,1365]" captionStart="FIGURES 5–8" captionStartId="11.[151,275,1942,1968]" captionTargetBox="[239,1351,193,1910]" captionTargetId="figure-19@11.[221,1366,179,1918]" captionTargetPageId="11" captionText="FIGURES 5–8. Thraulodes solstitium sp. nov., male imago: 5) forewing; 6a) hind wing; 6b) hind wing (enlarged); 7) genitalia, ventral view; 8) genitalia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5763310" httpUri="https://zenodo.org/record/5763310/files/figure.png" pageId="8" pageNumber="47">Figs. 5–6b</figureCitation>
). Forewing (
<figureCitation id="1360ACAE0313FF997414FAF8FDB3FA8C" box="[557,624,1338,1365]" captionStart="FIGURES 5–8" captionStartId="11.[151,275,1942,1968]" captionTargetBox="[239,1351,193,1910]" captionTargetId="figure-19@11.[221,1366,179,1918]" captionTargetPageId="11" captionText="FIGURES 5–8. Thraulodes solstitium sp. nov., male imago: 5) forewing; 6a) hind wing; 6b) hind wing (enlarged); 7) genitalia, ventral view; 8) genitalia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5763310" httpUri="https://zenodo.org/record/5763310/files/figure.png" pageId="8" pageNumber="47">Fig. 5</figureCitation>
) with veins yellowish brown; wing membrane hyaline; area between C and Sc yellowish brown. Fore and hind wings each with dark brown macula at base; costal cross veins basal to bulla either absent or colorless and very thin and 8 cross-veins distally. Trajectory line that connects bullae barely crossing MA fork. Hind wing (
<figureCitation id="1360ACAE0313FF9977B4FA64FDC5FA18" box="[397,518,1446,1473]" captionStart="FIGURES 5–8" captionStartId="11.[151,275,1942,1968]" captionTargetBox="[239,1351,193,1910]" captionTargetId="figure-19@11.[221,1366,179,1918]" captionTargetPageId="11" captionText="FIGURES 5–8. Thraulodes solstitium sp. nov., male imago: 5) forewing; 6a) hind wing; 6b) hind wing (enlarged); 7) genitalia, ventral view; 8) genitalia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5763310" httpUri="https://zenodo.org/record/5763310/files/figure.png" pageId="8" pageNumber="47">Fig. 6a–6b</figureCitation>
) with blunt costal projection; wing membrane hyaline; veins yellowish-brown.
</paragraph>
<paragraph id="8BE4B02B0313FF9976FEFA09FCCEF941" blockId="8.[151,1437,1014,2013]" pageId="8" pageNumber="47">Legs. Coxae and trochanters yellowish-brown. Cuticle of legs mostly yellowish-brown. Fore femur light brown with brown macula and blackish stripe on distal half; light brown, wide macula sometimes present on proximal half. Fore tibia yellowish brown, proximal region light brown. Fore tarsus yellowish brown with pigmented articulations. Middle and hind femora yellowish brown, distal 1/3 of femur brown with pre-apical, blackish-brown band; light brown, wide spot sometimes present on proximal half. Mid- and hind tibiae yellowish brown, proximal region light brown. Tarsus light brown with pigmented articulations.</paragraph>
<paragraph id="8BE4B02B0313FF9976FEF960FEC4F894" blockId="8.[151,1437,1014,2013]" pageId="8" pageNumber="47">
Abdomen (
<figureCitation id="1360ACAE0313FF99777CF960FE72F965" box="[325,433,1698,1724]" captionStart="FIGURES 2–4" captionStartId="10.[151,275,1941,1967]" captionTargetBox="[260,1326,321,1896]" captionTargetId="figure-25@10.[259,1328,179,1918]" captionTargetPageId="10" captionText="FIGURES 2–4. Thraulodes solstitium sp. nov., male imago: 2) habitus, (dorsal view); 3) head and thorax (lateral view); 4) abdomen (lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5763306" httpUri="https://zenodo.org/record/5763306/files/figure.png" pageId="8" pageNumber="47">Figs. 2, 4</figureCitation>
). Tergum I brownish, with pair of submedian blackish spots. Terga II–VI yellowish translucent with pair of submedian blackish dots and one pair of sublateral blackish dots; terga VI yellowish translucent anteriorly and light brown posteriorly; terga VII–X light brown. Abdominal sternum I-VI yellowish translucent, remaining light brown. Caudal filaments mostly yellowish brown, with black maculae on proximal ½ of alternated segments.
</paragraph>
<paragraph id="8BE4B02B0313FF9876FEF894FE87FEC4" blockId="8.[151,1437,1014,2013]" lastBlockId="9.[151,1437,151,1545]" lastPageId="9" lastPageNumber="48" pageId="8" pageNumber="47">
Genitalia (
<figureCitation id="1360ACAE0313FF997779F894FE6CF8A9" box="[320,431,1878,1904]" captionStart="FIGURES 5–8" captionStartId="11.[151,275,1942,1968]" captionTargetBox="[239,1351,193,1910]" captionTargetId="figure-19@11.[221,1366,179,1918]" captionTargetPageId="11" captionText="FIGURES 5–8. Thraulodes solstitium sp. nov., male imago: 5) forewing; 6a) hind wing; 6b) hind wing (enlarged); 7) genitalia, ventral view; 8) genitalia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5763310" httpUri="https://zenodo.org/record/5763310/files/figure.png" pageId="8" pageNumber="47">Figs. 7–8</figureCitation>
). Styliger plate yellowish translucent. Forceps mostly yellowish brown with pigmented articulations; middle of segment I strongly curved, proximal ½ of inner margin and distal ⅔ of outer margin light brown. Penes light brown with blackish pigmentation on “ear” area. Styliger plate triangular, median projection distally rounded and covering 2/3 of penes. Penes lobes short and distally divergent; each lobe gradually widened towards apex; lateral pouch present; apico-lateral area with ear-like projection. Medio-ventral ridge projected and distally integrated with oblique preapical ridge, forming “lapel.” Telopenes short and slender, 1.7 times width of penes on its median portion, lanceolate, narrowing towards apex, arising on dorsal side of penes and directed median-anteriorly.
</paragraph>
<paragraph id="8BE4B02B0312FF9876FEFEE4FDDDFE74" blockId="9.[151,1437,151,1545]" pageId="9" pageNumber="48">
<emphasis id="B92F6C390312FF9876FEFEE4FE81FE99" bold="true" box="[199,322,294,320]" pageId="9" pageNumber="48">Variations</emphasis>
. In some male
<typeStatus id="54E00E890312FF9877CDFEE5FDA1FE98" box="[500,610,295,321]" pageId="9" pageNumber="48" type="paratype">paratypes</typeStatus>
, the abdominal segment VIII is lighter than the segments VII, IX and X. Another important variation is that specimens may not have the maculae on the proximal half of the femora. Depending on the presence or absence of that characteristic, different comparisons can be drawn out when analyzing the key proposed by
<bibRefCitation id="EFCACDDA0312FF9877B9FE51FDD9FE74" author="Kluge, N. J." box="[384,538,403,429]" pageId="9" pageNumber="48" pagination="1 - 142" refId="ref11541" refString="Kluge, N. J. (2020) Systematic position of Thraulodes Ulmer 1920 (Ephemeroptera: Leptophlebiidae) and descriptions of new and little-known species. Zootaxa, 4756 (1), 1 - 142. https: // doi. org / 10.11646 / zootaxa. 4756.1.1" type="journal article" year="2020">Kluge (2020)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C341E3A00312FF9876FEFE75FBAFFDC0" pageId="9" pageNumber="48" type="etymology">
<paragraph id="8BE4B02B0312FF9876FEFE75FBAFFDC0" blockId="9.[151,1437,151,1545]" pageId="9" pageNumber="48">
<emphasis id="B92F6C390312FF9876FEFE75FE8BFE08" bold="true" box="[199,328,439,465]" pageId="9" pageNumber="48">Etymology</emphasis>
. From the Latin word
<taxonomicName id="4C5BCBA80312FF987406FE75FD6FFE08" authorityName="Orlando &amp; Salles &amp; Boldrini &amp; Krolow" authorityYear="2021" box="[575,684,439,465]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="solstitium">
<emphasis id="B92F6C390312FF987406FE75FD6FFE08" box="[575,684,439,465]" italics="true" pageId="9" pageNumber="48">solstitium</emphasis>
</taxonomicName>
, solstice. In reference to the event in which a planet’s poles are most extremely inclined toward or away from the star it orbits. This Earth phenomenon occurs in June and December, the same months in which some specimens were captured. This name is used in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C341E3A00312FF9876FEFDE0FDAFFDE4" box="[199,620,546,573]" pageId="9" pageNumber="48" type="distribution">
<paragraph id="8BE4B02B0312FF9876FEFDE0FDAFFDE4" blockId="9.[151,1437,151,1545]" box="[199,620,546,573]" pageId="9" pageNumber="48">
<emphasis id="B92F6C390312FF9876FEFDE0FE9AFDE5" bold="true" box="[199,345,546,572]" pageId="9" pageNumber="48">Distribution</emphasis>
.
<collectingRegion id="499F7EC90312FF98775EFDE1FDD6FDE4" box="[359,533,547,573]" country="Brazil" name="Tocantins" pageId="9" pageNumber="48">Tocantins State</collectingRegion>
,
<collectingCountry id="F34CF0BB0312FF987419FDE1FDABFDE4" box="[544,616,547,573]" name="Brazil" pageId="9" pageNumber="48">Brazil</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C341E3A00312FF9876FEFD84FE40FC58" pageId="9" pageNumber="48" type="materials_examined">
<paragraph id="8BE4B02B0312FF9876FEFD84FE40FC58" blockId="9.[151,1437,151,1545]" pageId="9" pageNumber="48">
<emphasis id="B92F6C390312FF9876FEFD84FE6DFDB9" bold="true" box="[199,430,582,608]" pageId="9" pageNumber="48">Material Examined</emphasis>
.
<materialsCitation id="3B33BA760312FF987782FD85FC89FD70" ID-GBIF-Occurrence="3415111333" collectingDate="2019-08-27" collectingDateMax="2019-08-30" collectingDateMin="2019-08-27" collectionCode="INPA" collectorName="Fernandes, A. S. &amp; Montanuci, P. S. B. &amp; Oliveira, L. P. &amp; Orlando, T. Y. S." country="Brazil" latitude="-10.366917" location="Taquarucu do Porto" longLatPrecision="15" longitude="-48.133614" municipality="Palmas" pageId="9" pageNumber="48" specimenCount="1" specimenCount-male="1" stateProvince="Tocantins state" typeStatus="holotype">
<typeStatus id="54E00E890312FF987782FD85FDEBFDB8" box="[443,552,583,609]" pageId="9" pageNumber="48" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D5D7BA20312FF987408FD85FD97FDB8" box="[561,596,583,609]" pageId="9" pageNumber="48" type="male">1♂</specimenCount>
(preserved in alcohol)
<collectingCountry id="F34CF0BB0312FF98756BFD85FC5AFDB8" box="[850,921,583,609]" name="Brazil" pageId="9" pageNumber="48">Brazil</collectingCountry>
,
<collectingRegion id="499F7EC90312FF98759BFD85FB89FDB8" box="[930,1098,583,609]" country="Brazil" name="Tocantins" pageId="9" pageNumber="48">Tocantins state</collectingRegion>
,
<collectingMunicipality id="6B802A510312FF98726DFD85FB6BFDB8" box="[1108,1192,583,609]" pageId="9" pageNumber="48">Palmas</collectingMunicipality>
, distrito de
<location id="8E84E6F00312FF98731FFD85FF39FD5C" LSID="urn:lsid:plazi:treatment:03F2013D0313FF9876AEFDE0FAC4F9D0:8E84E6F00312FF98731FFD85FF39FD5C" country="Brazil" latitude="-10.366917" longLatPrecision="15" longitude="-48.133614" municipality="Palmas" name="Taquarucu do Porto" pageId="9" pageNumber="48" stateProvince="Tocantins state">Taquaruçu do Porto</location>
, vale do
<location id="8E84E6F00312FF987767FDA9FDD3FD5C" LSID="urn:lsid:plazi:treatment:03F2013D0313FF9876AEFDE0FAC4F9D0:8E84E6F00312FF987767FDA9FDD3FD5C" box="[350,528,619,645]" country="Brazil" latitude="-10.366917" longLatPrecision="15" longitude="-48.133614" municipality="Palmas" name="Vai Quem Quer" pageId="9" pageNumber="48" stateProvince="Tocantins state">Vai Quem Quer</location>
,
<geoCoordinate id="EE6FD6EC0312FF987425FDA9FD6AFD5C" box="[540,681,619,645]" degrees="10" direction="south" minutes="22" orientation="latitude" pageId="9" pageNumber="48" precision="1" seconds="0.9" value="-10.366917">10°22’0.9”S</geoCoordinate>
/
<geoCoordinate id="EE6FD6EC0312FF98748AFDA9FCE1FD5C" box="[691,802,619,645]" degrees="48" direction="west" minutes="8" orientation="longitude" pageId="9" pageNumber="48" precision="15" seconds="1" value="-48.133614">48°8’1”W</geoCoordinate>
,
<date id="FFE596EB0312FF987517FDA9FC1EFD5C" box="[814,989,619,645]" pageId="9" pageNumber="48" value="2019-08-27" valueMax="2019-08-30" valueMin="2019-08-27">
<collectingDate id="EFA16F030312FF987517FDA9FC1EFD5C" box="[814,989,619,645]" pageId="9" pageNumber="48" value="2019-08-27" valueMax="2019-08-30" valueMin="2019-08-27">27-30.viii.2019</collectingDate>
</date>
, white sheet light trap,
<collectorName id="26AED5FD0312FF9872DDFDA9FA54FD5C" box="[1252,1431,619,645]" pageId="9" pageNumber="48">Fernandes, A.S.</collectorName>
,
<collectorName id="26AED5FD0312FF9876AEFD4DFEA0FD70" box="[151,355,655,681]" pageId="9" pageNumber="48">Montanuci, P.S.B.</collectorName>
,
<collectorName id="26AED5FD0312FF987757FD4DFDC6FD70" box="[366,517,655,681]" pageId="9" pageNumber="48">Oliveira, L.P.</collectorName>
,
<collectorName id="26AED5FD0312FF987436FD4DFD7FFD70" box="[527,700,655,681]" pageId="9" pageNumber="48">Orlando, T.Y.S.</collectorName>
cols (
<collectionCode id="ED4A28EE0312FF9874C5FD4DFC81FD70" box="[764,834,655,681]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="9" pageNumber="48">INPA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B33BA760312FF98756AFD4DFC70FD28" ID-GBIF-Occurrence="3415111335" collectingDate="2017-11-13" collectingDateMax="2017-11-14" collectingDateMin="2017-11-13" collectionCode="UFRR, CEUFT" collectorName="Krolow, T. K." country="Brazil" latitude="-10.366917" location="Taquarucu do Porto" longLatPrecision="15" longitude="-48.133614" municipality="Palmas" pageId="9" pageNumber="48" specimenCount="91" specimenCount-male="91" stateProvince="Tocantins state" typeStatus="paratype">
<typeStatus id="54E00E890312FF98756AFD4DFC07FD70" box="[851,964,655,681]" pageId="9" pageNumber="48" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D5D7BA20312FF9875F7FD4DFC3CFD70" box="[974,1023,655,681]" pageId="9" pageNumber="48" type="male">87♂</specimenCount>
(preserved in alcohol)
<collectingCountry id="F34CF0BB0312FF9872C7FD4DFA86FD70" box="[1278,1349,655,681]" name="Brazil" pageId="9" pageNumber="48">Brazil</collectingCountry>
,
<collectingRegion id="499F7EC90312FF987376FD4DFF3EFD14" pageId="9" pageNumber="48">Tocantins state</collectingRegion>
,
<collectingMunicipality id="6B802A510312FF987730FD71FE9EFD14" box="[265,349,691,717]" pageId="9" pageNumber="48">Palmas</collectingMunicipality>
, distrito de
<location id="8E84E6F00312FF9877D8FD71FD01FD14" LSID="urn:lsid:plazi:treatment:03F2013D0313FF9876AEFDE0FAC4F9D0:8E84E6F00312FF9877D8FD71FD01FD14" box="[481,706,691,717]" country="Brazil" latitude="-10.366917" longLatPrecision="15" longitude="-48.133614" municipality="Palmas" name="Taquarucu do Porto" pageId="9" pageNumber="48" stateProvince="Tocantins state">Taquaruçu do Porto</location>
, vale do
<location id="8E84E6F00312FF98751EFD71FC19FD14" LSID="urn:lsid:plazi:treatment:03F2013D0313FF9876AEFDE0FAC4F9D0:8E84E6F00312FF98751EFD71FC19FD14" box="[807,986,691,717]" country="Brazil" latitude="-10.366917" longLatPrecision="15" longitude="-48.133614" municipality="Palmas" name="Vai Quem Quer" pageId="9" pageNumber="48" stateProvince="Tocantins state">Vai Quem Quer</location>
,
<geoCoordinate id="EE6FD6EC0312FF9875DCFD71FBB0FD14" box="[997,1139,691,717]" degrees="10" direction="south" minutes="22" orientation="latitude" pageId="9" pageNumber="48" precision="1" seconds="0.9" value="-10.366917">10°22’0.9”S</geoCoordinate>
/
<geoCoordinate id="EE6FD6EC0312FF987244FD71FB28FD14" box="[1149,1259,691,717]" degrees="48" direction="west" minutes="8" orientation="longitude" pageId="9" pageNumber="48" precision="15" seconds="1" value="-48.133614">48°8’1”W</geoCoordinate>
,
<date id="FFE596EB0312FF9872C1FD71FA5BFD14" box="[1272,1432,691,717]" pageId="9" pageNumber="48" value="2017-11-13" valueMax="2017-11-14" valueMin="2017-11-13">
<collectingDate id="EFA16F030312FF9872C1FD71FA5BFD14" box="[1272,1432,691,717]" pageId="9" pageNumber="48" value="2017-11-13" valueMax="2017-11-14" valueMin="2017-11-13">13-14.xi.2017</collectingDate>
</date>
, white sheet light trap,
<collectorName id="26AED5FD0312FF987798FD15FDF4FD28" box="[417,567,727,753]" pageId="9" pageNumber="48">Krolow, T.K.</collectorName>
coll. (
<specimenCount id="9D5D7BA20312FF9874BEFD15FD6FFD28" box="[647,684,727,753]" pageId="9" pageNumber="48" type="male">2♂</specimenCount>
,
<collectionCode id="ED4A28EE0312FF987485FD15FCCAFD28" box="[700,777,727,753]" pageId="9" pageNumber="48">UFRR</collectionCode>
;
<specimenCount id="9D5D7BA20312FF987521FD15FCFEFD28" box="[792,829,727,753]" pageId="9" pageNumber="48" type="male">2♂</specimenCount>
,
<collectionCode id="ED4A28EE0312FF987574FD15FC68FD28" box="[845,939,727,753]" pageId="9" pageNumber="48">CEUFT</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B33BA760312FF9875F8FD15FDC5FCCC" ID-GBIF-Occurrence="3415111341" collectingDate="2017-11-15" collectingDateMax="2017-11-16" collectingDateMin="2017-11-15" collectionCode="CEUFT" collectorName="Krolow, T. K." country="Brazil" latitude="-10.366917" location="Taquarucu do Porto" longLatPrecision="15" longitude="-48.133614" municipality="Palmas" pageId="9" pageNumber="48" specimenCount="1" specimenCount-male="1" stateProvince="Tocantins state">
idem,
<date id="FFE596EB0312FF987232FD14FB68FD28" box="[1035,1195,726,753]" pageId="9" pageNumber="48" value="2017-11-15" valueMax="2017-11-16" valueMin="2017-11-15">
<collectingDate id="EFA16F030312FF987232FD14FB68FD28" box="[1035,1195,726,753]" pageId="9" pageNumber="48" value="2017-11-15" valueMax="2017-11-16" valueMin="2017-11-15">15-16.xi.2017</collectingDate>
</date>
, U.V. light pan trap,
<collectorName id="26AED5FD0312FF9876AEFD39FEEAFCCC" box="[151,297,763,789]" pageId="9" pageNumber="48">Krolow, T.K.</collectorName>
coll. (
<specimenCount id="9D5D7BA20312FF987748FD39FE56FCCC" box="[369,405,763,789]" pageId="9" pageNumber="48" type="male">1♂</specimenCount>
,
<collectionCode id="ED4A28EE0312FF987799FD39FE3DFCCC" box="[416,510,763,789]" pageId="9" pageNumber="48">CEUFT</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B33BA760312FF987429FD39FF3EFCE0" ID-GBIF-Occurrence="3415111355" collectingDate="2018-12-04" collectingDateMax="2018-12-06" collectingDateMin="2018-12-04" collectionCode="CEUFT" collectorName="Fernandes, A. S." country="Brazil" latitude="-10.366917" location="Taquarucu do Porto" longLatPrecision="15" longitude="-48.133614" municipality="Palmas" pageId="9" pageNumber="48" specimenCount="10" specimenCount-male="10" stateProvince="Tocantins state">
idem,
<date id="FFE596EB0312FF98746CFD39FD3EFCCC" box="[597,765,763,789]" pageId="9" pageNumber="48" value="2018-12-04" valueMax="2018-12-06" valueMin="2018-12-04">
<collectingDate id="EFA16F030312FF98746CFD39FD3EFCCC" box="[597,765,763,789]" pageId="9" pageNumber="48" value="2018-12-04" valueMax="2018-12-06" valueMin="2018-12-04">04-06.xii.2018</collectingDate>
</date>
, white sheet light trap,
<collectorName id="26AED5FD0312FF9875C7FD39FB6CFCCC" box="[1022,1199,763,789]" pageId="9" pageNumber="48">Fernandes, A.S.</collectorName>
and team cols. (
<specimenCount id="9D5D7BA20312FF98735DFD39FA5BFCCC" box="[1380,1432,763,789]" pageId="9" pageNumber="48" type="male">10♂</specimenCount>
,
<collectionCode id="ED4A28EE0312FF9876AEFCDDFF36FCE0" box="[151,245,799,825]" pageId="9" pageNumber="48">CEUFT</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B33BA760312FF98773EFCDDFA5BFCE0" ID-GBIF-Occurrence="3415111343" box="[263,1432,799,825]" collectingDate="2019-08-27" collectingDateMax="2019-08-30" collectingDateMin="2019-08-27" collectionCode="CEUFT, UFVB" collectorName="Fernandes, A. S." country="Brazil" latitude="-10.366917" location="Taquarucu do Porto" longLatPrecision="15" longitude="-48.133614" municipality="Palmas" pageId="9" pageNumber="48" specimenCount="61" specimenCount-male="61" stateProvince="Tocantins state">
idem,
<date id="FFE596EB0312FF987774FCDDFE3FFCE0" box="[333,508,799,825]" pageId="9" pageNumber="48" value="2019-08-27" valueMax="2019-08-30" valueMin="2019-08-27">
<collectingDate id="EFA16F030312FF987774FCDDFE3FFCE0" box="[333,508,799,825]" pageId="9" pageNumber="48" value="2019-08-27" valueMax="2019-08-30" valueMin="2019-08-27">27-30.viii.2019</collectingDate>
</date>
, white sheet light trap,
<collectorName id="26AED5FD0312FF9874C7FCDDFC6CFCE0" box="[766,943,799,825]" pageId="9" pageNumber="48">Fernandes, A.S.</collectorName>
and team cols. (
<specimenCount id="9D5D7BA20312FF98725CFCDDFB5BFCE0" box="[1125,1176,799,825]" pageId="9" pageNumber="48" type="male">41♂</specimenCount>
,
<collectionCode id="ED4A28EE0312FF98729DFCDDFB3CFCE0" box="[1188,1279,799,825]" pageId="9" pageNumber="48">CEUFT</collectionCode>
;
<specimenCount id="9D5D7BA20312FF987330FCDDFAF9FCE0" box="[1289,1338,799,825]" pageId="9" pageNumber="48" type="male">20♂</specimenCount>
<collectionCode id="ED4A28EE0312FF987379FCDDFA53FCE0" box="[1344,1424,799,825]" pageId="9" pageNumber="48">UFVB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B33BA760312FF9876AEFC81FE40FC58" ID-GBIF-Occurrence="3415111306" collectingDate="2018-07-13" collectingDateMax="2019-08-30" collectingDateMin="2019-08-27" collectionCode="CEUFT" collectorName="Fernandes, A. S." country="Brazil" latitude="-10.233389" location="cachoeira do Evilson" longLatPrecision="15" longitude="-48.121387" municipality="Palmas" pageId="9" pageNumber="48" specimenCount="11" specimenCount-male="11" stateProvince="Tocantins state">
idem,
<location id="8E84E6F00312FF9876E4FC81FE0AFC84" LSID="urn:lsid:plazi:treatment:03F2013D0313FF9876AEFDE0FAC4F9D0:8E84E6F00312FF9876E4FC81FE0AFC84" box="[221,457,835,861]" latitude="-10.233389" longLatPrecision="15" longitude="-48.121387" name="cachoeira do Evilson" pageId="9" pageNumber="48">cachoeira do Evilson</location>
,
<geoCoordinate id="EE6FD6EC0312FF9877EDFC81FD9DFC84" box="[468,606,835,861]" degrees="10" direction="south" minutes="14" orientation="latitude" pageId="9" pageNumber="48" precision="1" seconds="0.2" value="-10.233389">10°14’0.2”S</geoCoordinate>
/
<geoCoordinate id="EE6FD6EC0312FF987457FC81FD2CFC84" box="[622,751,835,861]" degrees="48" direction="west" minutes="7" orientation="longitude" pageId="9" pageNumber="48" precision="15" seconds="17" value="-48.121387">48°7’17”W</geoCoordinate>
,
<date id="FFE596EB0312FF9874C3FC81FCBFFC84" box="[762,892,835,861]" pageId="9" pageNumber="48" value="2018-07-13">
<collectingDate id="EFA16F030312FF9874C3FC81FCBFFC84" box="[762,892,835,861]" pageId="9" pageNumber="48" value="2018-07-13">13.vii.2018</collectingDate>
</date>
, white sheet light trap,
<collectorName id="26AED5FD0312FF987246FC81FAF3FC84" box="[1151,1328,835,861]" pageId="9" pageNumber="48">Fernandes, A.S.</collectorName>
and team cols. (
<specimenCount id="9D5D7BA20312FF9876E2FCA5FECEFC58" box="[219,269,871,897]" pageId="9" pageNumber="48" type="male">11♂</specimenCount>
,
<collectionCode id="ED4A28EE0312FF987723FCA5FEBBFC58" box="[282,376,871,897]" pageId="9" pageNumber="48">CEUFT</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C341E3A00312FF9876FEFC48FAC4F9D0" pageId="9" pageNumber="48" type="discussion">
<paragraph id="8BE4B02B0312FF9876FEFC48FD72FBEC" blockId="9.[151,1437,151,1545]" pageId="9" pageNumber="48">
<emphasis id="B92F6C390312FF9876FEFC48FE8CFC7D" bold="true" box="[199,335,906,932]" pageId="9" pageNumber="48">Comments.</emphasis>
One of the most critical characteristics for the identification of the species of
<taxonomicName id="4C5BCBA80312FF987286FC49FAFCFC7C" authorityName="Ulmer" authorityYear="1920" box="[1215,1343,907,933]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="B92F6C390312FF987286FC49FAFCFC7C" box="[1215,1343,907,933]" italics="true" pageId="9" pageNumber="48">Thraulodes</emphasis>
</taxonomicName>
, at least when using taxonomic keys (
<bibRefCitation id="EFCACDDA0312FF9877DBFC6DFCC3FC10" author="Traver, J. R. &amp; Edmunds, G. F. Jr." box="[482,768,943,969]" pageId="9" pageNumber="48" pagination="349 - 395" refId="ref14380" refString="Traver, J. R. &amp; Edmunds, G. F. Jr. (1967) A revision of the genus Thraulodes (Ephemeroptera: Leptophlebiidae). Miscellaneous Publications of the Entomological Society of America, 5, 349 - 395." type="journal article" year="1967">Traver &amp; Edmunds 1967</bibRefCitation>
;
<bibRefCitation id="EFCACDDA0312FF987534FC6DFBD6FC10" author="Dominguez, E. &amp; Molineri, C. &amp; Pescador, M. L. &amp; Hubbard, M. D. &amp; Nieto, C." box="[781,1045,943,969]" pageId="9" pageNumber="48" refId="ref10712" refString="Dominguez, E., Molineri, C., Pescador, M. L., Hubbard, M. D. &amp; Nieto, C. (2006) Ephemeroptera of South America, in Aquatic Biodiversity in Latin America. Vol. 2. Pensoft, Sofia-Moscow, 646 pp." type="book" year="2006">
Domínguez
<emphasis id="B92F6C390312FF9875A1FC6DFC09FC10" box="[920,970,943,969]" italics="true" pageId="9" pageNumber="48">et al</emphasis>
. 2006
</bibRefCitation>
;
<bibRefCitation id="EFCACDDA0312FF987218FC6DFB23FC10" author="Lima, L. R. C. &amp; Mariano, R. &amp; Pinheiro, U." box="[1057,1248,943,969]" pageId="9" pageNumber="48" pagination="230 - 242" refId="ref11757" refString="Lima, L. R. C., Mariano, R. &amp; Pinheiro, U. (2013) New species for Thraulodes Ulmer, 1920 (Ephemeroptera: Leptophlebiidae: Atalophlebiinae) and the first key to adults from Brazil. Zootaxa, 3709 (3), 230 - 242. https: // doi. org / 10.11646 / zootaxa. 3709.3.2" type="journal article" year="2013">
<collectingRegion id="499F7EC90312FF987218FC6DFB9FFC10" box="[1057,1116,943,969]" country="Peru" name="Lima" pageId="9" pageNumber="48">Lima</collectingRegion>
<emphasis id="B92F6C390312FF98725DFC6DFB55FC10" box="[1124,1174,943,969]" italics="true" pageId="9" pageNumber="48">et al</emphasis>
. 2013
</bibRefCitation>
;
<bibRefCitation id="EFCACDDA0312FF9872D4FC6DFAB6FC10" author="Kluge, N. J." box="[1261,1397,943,969]" pageId="9" pageNumber="48" pagination="1 - 142" refId="ref11541" refString="Kluge, N. J. (2020) Systematic position of Thraulodes Ulmer 1920 (Ephemeroptera: Leptophlebiidae) and descriptions of new and little-known species. Zootaxa, 4756 (1), 1 - 142. https: // doi. org / 10.11646 / zootaxa. 4756.1.1" type="journal article" year="2020">Kluge 2020</bibRefCitation>
), is the presence or absence of a macula on the proximal half of the mid and hind femora. However, among the material examined of the new species, we found some specimens with a very light macula and others without it, which can be due to preservation or intraspecific variation.
</paragraph>
<paragraph id="8BE4B02B0312FF9876FEFBFDFE04FA8C" blockId="9.[151,1437,151,1545]" pageId="9" pageNumber="48">
Among the species of the genus with the maculae on the mid and hind femora, the new species is morphologically similar to
<taxonomicName id="4C5BCBA80312FF98777CFBA1FE36FBA4" authorityName="Traver" authorityYear="1964" box="[325,501,1123,1149]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="paysandensis">
<emphasis id="B92F6C390312FF98777CFBA1FE36FBA4" box="[325,501,1123,1149]" italics="true" pageId="9" pageNumber="48">T. paysandensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5BCBA80312FF987415FBA1FD4CFBA4" box="[556,655,1123,1149]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="pinhoi">
<emphasis id="B92F6C390312FF987415FBA1FD4CFBA4" box="[556,655,1123,1149]" italics="true" pageId="9" pageNumber="48">T. pinhoi</emphasis>
</taxonomicName>
, sharing the abdominal terga II to VI which are yellowish translucent. The new species can be distinguished from them based on the morphology of the genitalia: in
<taxonomicName id="4C5BCBA80312FF9872F8FB45FAB1FB78" authorityName="Traver" authorityYear="1964" box="[1217,1394,1159,1185]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="paysandensis">
<emphasis id="B92F6C390312FF9872F8FB45FAB1FB78" box="[1217,1394,1159,1185]" italics="true" pageId="9" pageNumber="48">T. paysandensis</emphasis>
</taxonomicName>
the apicolateral area of each penes lobe is not ear-like (
<bibRefCitation id="EFCACDDA0312FF9874D1FB69FC36FB1C" author="Dominguez, E. &amp; Molineri, C. &amp; Pescador, M. L. &amp; Hubbard, M. D. &amp; Nieto, C." box="[744,1013,1195,1221]" pageId="9" pageNumber="48" refId="ref10712" refString="Dominguez, E., Molineri, C., Pescador, M. L., Hubbard, M. D. &amp; Nieto, C. (2006) Ephemeroptera of South America, in Aquatic Biodiversity in Latin America. Vol. 2. Pensoft, Sofia-Moscow, 646 pp." type="book" year="2006">
Domínguez
<emphasis id="B92F6C390312FF98754FFB69FC6AFB1C" box="[886,937,1195,1221]" italics="true" pageId="9" pageNumber="48">et al</emphasis>
. 2006
</bibRefCitation>
: pg 500, Fig. 189B), in
<taxonomicName id="4C5BCBA80312FF987335FB69FAB2FB1C" box="[1292,1393,1195,1221]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="pinhoi">
<emphasis id="B92F6C390312FF987335FB69FAB2FB1C" box="[1292,1393,1195,1221]" italics="true" pageId="9" pageNumber="48">T. pinhoi</emphasis>
</taxonomicName>
the posterior margin of the subgenital plate is almost straight, with medial rounded projection, and in both species the telopenes are long, slender, and medially directed (Fig. 16 from
<bibRefCitation id="EFCACDDA0312FF987555FB31FBE8FAD4" author="Lima, L. R. C. &amp; Mariano, R. &amp; Pinheiro, U." box="[876,1067,1267,1293]" pageId="9" pageNumber="48" pagination="230 - 242" refId="ref11757" refString="Lima, L. R. C., Mariano, R. &amp; Pinheiro, U. (2013) New species for Thraulodes Ulmer, 1920 (Ephemeroptera: Leptophlebiidae: Atalophlebiinae) and the first key to adults from Brazil. Zootaxa, 3709 (3), 230 - 242. https: // doi. org / 10.11646 / zootaxa. 3709.3.2" type="journal article" year="2013">
<collectingRegion id="499F7EC90312FF987555FB31FC64FAD4" box="[876,935,1267,1293]" country="Peru" name="Lima" pageId="9" pageNumber="48">Lima</collectingRegion>
<emphasis id="B92F6C390312FF987596FB31FC22FAD4" box="[943,993,1267,1293]" italics="true" pageId="9" pageNumber="48">et al</emphasis>
. 2013
</bibRefCitation>
: pg, Fig. 16). On the other hand, in
<taxonomicName id="4C5BCBA80312FF98768FFAD5FE82FAE8" authority="Orlando &amp; Salles &amp; Boldrini &amp; Krolow, 2021" authorityName="Orlando &amp; Salles &amp; Boldrini &amp; Krolow" authorityYear="2021" box="[182,321,1303,1329]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="solstitium" status="sp. nov.">
<emphasis id="B92F6C390312FF98768FFAD5FE82FAE8" box="[182,321,1303,1329]" italics="true" pageId="9" pageNumber="48">T. solstitium</emphasis>
</taxonomicName>
<emphasis id="B92F6C390312FF987773FAD5FE65FAE8" bold="true" box="[330,422,1303,1329]" pageId="9" pageNumber="48">
<taxonomicNameLabel id="A21CD1420312FF987773FAD5FE65FAE8" box="[330,422,1303,1329]" pageId="9" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, an ear-like area is present apicolaterally on the penes lobe, and the telopenes are short, directed median-anteriorly.
</paragraph>
<paragraph id="8BE4B02B0312FF9876FEFA9DFAC4F9D0" blockId="9.[151,1437,151,1545]" pageId="9" pageNumber="48">
When considering the maculae on the proximal half of femora absent, the new species is morphologically similar to
<taxonomicName id="4C5BCBA80312FF9876EEFA41FE44FA44" authorityName="Flowers" authorityYear="2009" box="[215,391,1411,1437]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="quevedoensis">
<emphasis id="B92F6C390312FF9876EEFA41FE44FA44" box="[215,391,1411,1437]" italics="true" pageId="9" pageNumber="48">T. quevedoensis</emphasis>
</taxonomicName>
by sharing the abdominal terga II–V light; VI–VII almost entirely colored and penes widened apically. The new species can be distinguished from it based on the morphology of the genitalia: in
<taxonomicName id="4C5BCBA80312FF9872D5FA65FA5FFA18" authorityName="Flowers" authorityYear="2009" box="[1260,1436,1447,1473]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="quevedoensis">
<emphasis id="B92F6C390312FF9872D5FA65FA5FFA18" box="[1260,1436,1447,1473]" italics="true" pageId="9" pageNumber="48">T. quevedoensis</emphasis>
</taxonomicName>
the lateral pouch is absent and the telopenes is long and directed medially (
<bibRefCitation id="EFCACDDA0312FF9875E7FA09FBBEFA3C" author="Flowers, R. W." box="[990,1149,1483,1509]" pageId="9" pageNumber="48" pagination="55 - 61" refId="ref10867" refString="Flowers, R. W. (2009) A new species of Thraulodes (Ephemeroptera: Leptophlebiidae, Atalophlebiinae) from a highly altered river in western Ecuador. Zootaxa, 2052 (1), 55 - 61. https: // doi. org / 10.11646 / zootaxa. 2052.1.4" type="journal article" year="2009">Flowers 2009</bibRefCitation>
: pg 57,
<figureCitation id="1360ACAE0312FF9872EDFA09FAD5FA3C" box="[1236,1302,1483,1509]" captionStart="FIGURES 2–4" captionStartId="10.[151,275,1941,1967]" captionTargetBox="[260,1326,321,1896]" captionTargetId="figure-25@10.[259,1328,179,1918]" captionTargetPageId="10" captionText="FIGURES 2–4. Thraulodes solstitium sp. nov., male imago: 2) habitus, (dorsal view); 3) head and thorax (lateral view); 4) abdomen (lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5763306" httpUri="https://zenodo.org/record/5763306/files/figure.png" pageId="9" pageNumber="48">Fig. 3</figureCitation>
) while in
<taxonomicName id="4C5BCBA80312FF9873B1FA09FEC7F9D0" authority="Orlando &amp; Salles &amp; Boldrini &amp; Krolow, 2021" authorityName="Orlando &amp; Salles &amp; Boldrini &amp; Krolow" authorityYear="2021" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="solstitium" status="sp. nov.">
<emphasis id="B92F6C390312FF9873B1FA09FEC7F9D0" italics="true" pageId="9" pageNumber="48">T. solstitium</emphasis>
</taxonomicName>
<emphasis id="B92F6C390312FF987732FA2DFEA6F9D0" bold="true" box="[267,357,1519,1545]" pageId="9" pageNumber="48">
<taxonomicNameLabel id="A21CD1420312FF987732FA2DFEA6F9D0" box="[267,357,1519,1545]" pageId="9" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
the lateral pouch is present and the telopenes are short, directed median-anteriorly.
</paragraph>
</subSubSection>
</subSection>
</treatment>
</document>
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