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<document ID-DOI="http://doi.org/10.5281/zenodo.3803484" ID-GBIF-Dataset="fe76f860-39b1-4520-a13c-d01a47dfc130" ID-GBIF-Taxon="163738939" ID-ISSN="1175-5334" ID-Zenodo-Dep="3803484" ID-ZooBank="urn:lsid:zoobank.org:pub:BA570D47-FE77-446C-B000-9208C9A6E957" checkinTime="1587753273724" checkinUser="carolina" docAuthor="Ríos-Tamayo, Duniesky &amp; Lyle, Robin" docDate="2020" docId="03F11B20FFC1FFAA8EE83EBFFBF0D3B9" docLanguage="en" docName="zootaxa.4766.2.2.pdf.imf" docOrigin="Zootaxa 4766 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lepthercus haddadi Ríos-Tamayo &amp; Lyle 2020, sp. nov." docType="treatment" docVersion="5" lastPageId="34" lastPageNumber="295" masterDocId="FFC86358FFDEFF888E7F3E26FFD2D232" masterDocTitle="The South African genus Lepthercus Purcell, 1902 (Araneae: Mygalomorphae) phylogeny and taxonomy" masterLastPageNumber="305" masterPageNumber="261" pageId="31" pageNumber="292" updateTime="1636538462881" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo>
<mods:title>The South African genus Lepthercus Purcell, 1902 (Araneae: Mygalomorphae) phylogeny and taxonomy</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ríos-Tamayo, Duniesky</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lyle, Robin</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="pubDate">
<mods:number>2020-04-20</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>4766</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>261</mods:start>
<mods:end>305</mods:end>
</mods:extent>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.4766.2.2</mods:identifier>
<mods:identifier type="GBIF-Dataset">fe76f860-39b1-4520-a13c-d01a47dfc130</mods:identifier>
<mods:identifier type="ISSN">1175-5334</mods:identifier>
<mods:identifier type="Zenodo-Dep">3765467</mods:identifier>
<mods:identifier type="ZooBank">urn:lsid:zoobank.org:pub:BA570D47-FE77-446C-B000-9208C9A6E957</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3803484" ID-GBIF-Taxon="163738939" ID-Zenodo-Dep="3803484" LSID="urn:lsid:plazi:treatment:03F11B20FFC1FFAA8EE83EBFFBF0D3B9" httpUri="http://treatment.plazi.org/id/03F11B20FFC1FFAA8EE83EBFFBF0D3B9" lastPageId="34" lastPageNumber="295" pageId="31" pageNumber="292">
<subSubSection box="[151,492,153,180]" pageId="31" pageNumber="292" type="nomenclature">
<paragraph blockId="31.[151,1155,153,251]" box="[151,492,153,180]" pageId="31" pageNumber="292">
<heading bold="true" box="[151,492,153,180]" fontSize="10" level="2" pageId="31" pageNumber="292" reason="2">
<emphasis bold="true" box="[151,492,153,180]" pageId="31" pageNumber="292">
<taxonomicName authority="Ríos-Tamayo &amp; Lyle, 2020" authorityName="Ríos-Tamayo &amp; Lyle" authorityYear="2020" box="[151,392,153,180]" class="Arachnida" family="Nemesiidae" genus="Lepthercus" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="292" phylum="Arthropoda" rank="species" species="haddadi" status="sp. nov.">
<emphasis bold="true" box="[151,392,153,180]" italics="true" pageId="31" pageNumber="292">Lepthercus haddadi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[399,492,154,180]" pageId="31" pageNumber="292" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="31" pageNumber="292" type="description">
<paragraph blockId="31.[151,1155,153,251]" pageId="31" pageNumber="292">
(
<figureCitation box="[159,321,189,216]" captionStart="FIGURE" captionStartId="7.[151,251,1647,1672]" captionTargetBox="[151,1436,381,1622]" captionTargetId="figure@7.[151,1436,381,1622]" captionTargetPageId="7" captionText="FIGURE. 2. Preening comb. A Lepthercus haddadi sp. nov. (female paratype, NCA 2007/3847). Maxillary cuspules (males). B L. dippenaarae. C L. kwazuluensis. Serrula. D L. mandelai sp. nov. (female, NCA 2014/415A); E L. filmeri sp. nov. (female, NCA 91/1437); F L. haddadi sp. nov. (male, NCA 2007/1486); G L. haddadi sp. nov. (female, NCA 2007/3847); H L. engelbrechti sp. nov. (female, NCA 2018/385); I L. engelbrechti sp. nov. (male, NCA 2018/384); J L. dregei Purcell, 1902 (female, NCA 2012/2333); K L. sofiae sp. nov. (male, NCA 97/395)." figureDoi="http://doi.org/10.5281/zenodo.3765471" httpUri="https://zenodo.org/record/3765471/files/figure.png" pageId="31" pageNumber="292">Figs. 2A, F, G</figureCitation>
,
<figureCitation box="[333,359,189,215]" captionStart="FIGURE" captionStartId="31.[151,251,1721,1746]" captionTargetBox="[151,1436,561,1696]" captionTargetId="figure@31.[151,1436,561,1696]" captionTargetPageId="31" captionText="FIGURE. 19. Lepthercus haddadi sp. nov. (male holotype, NCA 2007/1486). A cephalothorax, dorsal view; B abdomen, dorsal view; C sternum, ventral view; D chelicerae, arrow indicates the intercheliceral tumescence with setae, prolateral view; E right pedipalp, arrows indicates the palpal tibia base dorsally very convex and the development of the anterior side of the retrolateral side of the ventral excavation, retrolateral view, F right pedipalp, arrows indicates the few proventral setae and the development of the anterior side of the retrolateral side of the ventral excavation, prolateral view; GI copulatory bulb, G retrolateral view, H ventral view, I prolateral view; J tibia-metatarsus I, showing the very robust tibia with a robust spur, arrow indicates the patch of numerous black blunt spinules, prolateral view; K tibia I, arrow indicates the strong curved spine near of the base of the megaspine, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3765509" httpUri="https://zenodo.org/record/3765509/files/figure.png" pageId="31" pageNumber="292">19</figureCitation>
AK, 20AC, 24) http://zoobank.org/
<uri box="[363,1155,227,251]" pageId="31" pageNumber="292">
urn:lsid:zoobank.org:act:
<uuid box="[625,1155,227,251]" pageId="31" pageNumber="292">4ADBA1C9-2B18-41FA-B2AC-2A597B8D8B38</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection pageId="31" pageNumber="292" type="materials_examined">
<paragraph blockId="31.[151,1436,297,540]" pageId="31" pageNumber="292">
<emphasis bold="true" box="[151,554,297,324]" pageId="31" pageNumber="292">Material examined. Type material</emphasis>
:
<materialsCitation ID-GBIF-Occurrence="2608670254" collectingDate="2005-03-25" collectionCode="NCA" collectorName="De Hoop Vlei, S &amp; Haddad, C." country="South Africa" latitude="-34.283333" location="Koppie Alleen" longLatPrecision="1200" longitude="20.283333" municipality="De Hoop Nature Reserve" pageId="31" pageNumber="292" specimenCode="NCA 2007" specimenCount="1" specimenCount-male="1" stateProvince="Western Cape Province" typeStatus="holotype">
<typeStatus box="[568,672,297,323]" pageId="31" pageNumber="292">Holotype</typeStatus>
<specimenCount box="[678,702,298,324]" pageId="31" pageNumber="292" type="male"></specimenCount>
.
<collectingCountry box="[712,865,297,324]" name="South Africa" pageId="31" pageNumber="292">SOUTH AFRICA</collectingCountry>
:
<collectingRegion box="[877,1140,297,324]" country="South Africa" name="Western Cape" pageId="31" pageNumber="292">Western Cape Province</collectingRegion>
,
<collectingMunicipality box="[1150,1432,298,324]" pageId="31" pageNumber="292">De Hoop Nature Reserve</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:03F11B20FFC1FFAA8EE83EBFFBF0D3B9:8E87FCEDFFC1FF978EE83F6BFEEAD355" box="[151,312,333,359]" country="South Africa" latitude="-34.283333" longLatPrecision="1200" longitude="20.283333" municipality="De Hoop Nature Reserve" name="Koppie Alleen" pageId="31" pageNumber="292" stateProvince="Western Cape Province">Koppie Alleen</location>
cottage,
<collectorName box="[409,590,333,360]" pageId="31" pageNumber="292">De Hoop Vlei, S</collectorName>
shore (
<geoCoordinate box="[669,758,333,360]" degrees="34" direction="south" minutes="17" orientation="latitude" pageId="31" pageNumber="292" precision="925" value="-34.283333">34°17S</geoCoordinate>
<geoCoordinate box="[765,859,333,360]" degrees="20" direction="east" minutes="17" orientation="longitude" pageId="31" pageNumber="292" precision="925" value="20.283333">20°17E</geoCoordinate>
),
<date box="[877,1001,333,360]" pageId="31" pageNumber="292" value="2005-03-25">
<collectingDate box="[877,1001,333,360]" pageId="31" pageNumber="292" value="2005-03-25">25.iii.2005</collectingDate>
</date>
,
<collectorName box="[1010,1135,333,359]" pageId="31" pageNumber="292">Haddad, C.</collectorName>
<emphasis box="[1140,1181,333,359]" italics="true" pageId="31" pageNumber="292">leg.</emphasis>
,. (
<specimenCode box="[1208,1330,333,360]" collectionCode="NCA" pageId="31" pageNumber="292">NCA 2007</specimenCode>
/1486)
</materialsCitation>
.—
<typeStatus box="[151,249,370,396]" pageId="31" pageNumber="292">Paratype</typeStatus>
.
<collectingCountry box="[262,415,369,396]" name="South Africa" pageId="31" pageNumber="292">SOUTH AFRICA</collectingCountry>
: De Hoop Nature Reserve, Koppie Alleen cottage (
<geoCoordinate box="[984,1074,369,396]" degrees="34" direction="south" minutes="17" orientation="latitude" pageId="31" pageNumber="292" precision="925" value="-34.283333">34°17S</geoCoordinate>
<geoCoordinate box="[1081,1175,369,396]" degrees="20" direction="east" minutes="17" orientation="longitude" pageId="31" pageNumber="292" precision="925" value="20.283333">20°17E</geoCoordinate>
),
<date box="[1194,1332,369,395]" pageId="31" pageNumber="292" value="2007-08-26">26.viii.2007</date>
. Haddad, C.
<emphasis box="[184,225,405,431]" italics="true" pageId="31" pageNumber="292">leg.</emphasis>
,
<specimenCount box="[239,274,405,431]" pageId="31" pageNumber="292" type="female">1♀</specimenCount>
(
<collectionCode box="[288,348,405,431]" pageId="31" pageNumber="292">NCA</collectionCode>
2007/3847): same data as the male
<typeStatus box="[750,851,405,431]" pageId="31" pageNumber="292">holotype</typeStatus>
,
<specimenCount box="[862,904,405,432]" pageId="31" pageNumber="292" type="female">1 ♀</specimenCount>
(subadult),
<specimenCount box="[1039,1083,405,432]" pageId="31" pageNumber="292" type="male">1 ♂</specimenCount>
,
<specimenCount box="[1095,1175,405,431]" pageId="31" pageNumber="292" type="generic">1 imm.</specimenCount>
(
<collectionCode box="[1188,1248,405,431]" pageId="31" pageNumber="292">NCA</collectionCode>
2007/1486).
</paragraph>
<paragraph blockId="31.[151,1436,297,540]" pageId="31" pageNumber="292">
<emphasis bold="true" box="[199,499,441,468]" pageId="31" pageNumber="292">Other material examined</emphasis>
:
<collectingCountry box="[513,666,441,468]" name="South Africa" pageId="31" pageNumber="292">SOUTH AFRICA</collectingCountry>
:
<collectingRegion box="[679,944,441,468]" country="South Africa" name="Western Cape" pageId="31" pageNumber="292">Western Cape Province</collectingRegion>
: Bredasdorp Distr, Duinehuisie Guest Farm, Farm Rondekop; ca.
<quantity box="[396,470,477,503]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="31" pageNumber="292" unit="km" value="12.0">12 km</quantity>
SE Bredasdorp; east of farm buildings (
<geoCoordinate box="[944,1109,477,504]" degrees="34" direction="south" minutes="35" orientation="latitude" pageId="31" pageNumber="292" precision="1" seconds="37,12" value="-34.593643">34°3537,12″S</geoCoordinate>
<geoCoordinate box="[1116,1271,477,504]" degrees="20" direction="east" minutes="9" orientation="longitude" pageId="31" pageNumber="292" precision="1" seconds="15.31" value="20.154253">20°915.31″E</geoCoordinate>
),
<date box="[1294,1432,477,503]" pageId="31" pageNumber="292" value="2017-08-23">23.viii.2017</date>
, Engelbrecht, I., N. Calver, D. Calver, P. Bester, A. Cilliers.
<emphasis box="[814,856,513,539]" italics="true" pageId="31" pageNumber="292">leg.</emphasis>
,
<specimenCount box="[869,911,513,540]" pageId="31" pageNumber="292" type="male">1 ♂</specimenCount>
(
<collectionCode box="[926,986,513,539]" pageId="31" pageNumber="292">NCA</collectionCode>
2019/40).
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3765509" ID-Zenodo-Dep="3765509" httpUri="https://zenodo.org/record/3765509/files/figure.png" pageId="31" pageNumber="292" startId="31.[151,251,1721,1746]" targetBox="[151,1436,561,1696]" targetPageId="31">
<paragraph blockId="31.[151,1437,1721,1998]" pageId="31" pageNumber="292">
<emphasis bold="true" box="[151,293,1721,1746]" pageId="31" pageNumber="292">FIGURE. 19.</emphasis>
<taxonomicName authority="Ríos-Tamayo &amp; Lyle, 2020" authorityName="Ríos-Tamayo &amp; Lyle" authorityYear="2020" box="[299,499,1722,1746]" class="Arachnida" family="Nemesiidae" genus="Lepthercus" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="292" phylum="Arthropoda" rank="species" species="haddadi" status="sp. nov.">
<emphasis box="[299,499,1722,1746]" italics="true" pageId="31" pageNumber="292">Lepthercus haddadi</emphasis>
</taxonomicName>
<emphasis bold="true" box="[504,584,1722,1746]" pageId="31" pageNumber="292">
<taxonomicNameLabel box="[504,584,1722,1746]" pageId="31" pageNumber="292" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(male
<typeStatus box="[653,744,1721,1746]" pageId="31" pageNumber="292">holotype</typeStatus>
,
<collectionCode box="[753,805,1722,1746]" pageId="31" pageNumber="292">NCA</collectionCode>
2007/1486).
<emphasis bold="true" box="[939,957,1721,1746]" pageId="31" pageNumber="292">A</emphasis>
cephalothorax, dorsal view;
<emphasis bold="true" box="[1247,1264,1722,1746]" pageId="31" pageNumber="292">B</emphasis>
abdomen, dorsal view;
<emphasis bold="true" box="[214,232,1757,1782]" pageId="31" pageNumber="292">C</emphasis>
sternum, ventral view;
<emphasis bold="true" box="[471,489,1758,1782]" pageId="31" pageNumber="292">D</emphasis>
chelicerae, arrow indicates the intercheliceral tumescence with setae, prolateral view;
<emphasis bold="true" box="[1365,1382,1758,1782]" pageId="31" pageNumber="292">E</emphasis>
right pedipalp, arrows indicates the palpal tibia base dorsally very convex and the development of the anterior side of the retrolateral side of the ventral excavation, retrolateral view,
<emphasis bold="true" box="[652,667,1830,1854]" pageId="31" pageNumber="292">F</emphasis>
right pedipalp, arrows indicates the few proventral setae and the development of the anterior side of the retrolateral side of the ventral excavation, prolateral view;
<emphasis bold="true" box="[1075,1118,1865,1890]" pageId="31" pageNumber="292">GI</emphasis>
copulatory bulb,
<emphasis bold="true" box="[1298,1318,1865,1890]" pageId="31" pageNumber="292">G</emphasis>
retrolateral view,
<emphasis bold="true" box="[212,232,1902,1926]" pageId="31" pageNumber="292">H</emphasis>
ventral view,
<emphasis bold="true" box="[378,388,1902,1926]" pageId="31" pageNumber="292">I</emphasis>
prolateral view;
<emphasis bold="true" box="[564,577,1902,1926]" pageId="31" pageNumber="292">J</emphasis>
tibia-metatarsus I, showing the very robust tibia with a robust spur, arrow indicates the patch of numerous black blunt spinules, prolateral view;
<emphasis bold="true" box="[769,789,1938,1962]" pageId="31" pageNumber="292">K</emphasis>
tibia I, arrow indicates the strong curved spine near of the base of the megaspine, ventral view.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3765511" ID-Zenodo-Dep="3765511" httpUri="https://zenodo.org/record/3765511/files/figure.png" pageId="32" pageNumber="293" startId="32.[151,251,1919,1944]" targetBox="[151,1436,207,1894]" targetPageId="32">
<paragraph blockId="32.[151,1436,1919,1980]" pageId="32" pageNumber="293">
<emphasis bold="true" box="[151,297,1919,1944]" pageId="32" pageNumber="293">FIGURE. 20.</emphasis>
<taxonomicName authority="Ríos-Tamayo &amp; Lyle, 2020" authorityName="Ríos-Tamayo &amp; Lyle" authorityYear="2020" box="[305,509,1920,1944]" class="Arachnida" family="Nemesiidae" genus="Lepthercus" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="293" phylum="Arthropoda" rank="species" species="haddadi" status="sp. nov.">
<emphasis box="[305,509,1920,1944]" italics="true" pageId="32" pageNumber="293">Lepthercus haddadi</emphasis>
</taxonomicName>
<emphasis bold="true" box="[518,601,1920,1944]" pageId="32" pageNumber="293">
<taxonomicNameLabel box="[518,601,1920,1944]" pageId="32" pageNumber="293" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(female
<typeStatus box="[696,786,1920,1944]" pageId="32" pageNumber="293">paratype</typeStatus>
,
<collectionCode box="[799,851,1920,1944]" pageId="32" pageNumber="293">NCA</collectionCode>
2007/3847).
<emphasis bold="true" box="[992,1010,1919,1944]" pageId="32" pageNumber="293">A</emphasis>
habitus, dorsal view;
<emphasis bold="true" box="[1243,1260,1920,1944]" pageId="32" pageNumber="293">B</emphasis>
sternum, ventral view;
<emphasis bold="true" box="[214,232,1955,1980]" pageId="32" pageNumber="293">C</emphasis>
chelicerae, prolateral view;
<emphasis bold="true" box="[520,538,1956,1980]" pageId="32" pageNumber="293">D</emphasis>
spermathecae
</paragraph>
</caption>
<paragraph blockId="32.[151,1436,1919,1980]" box="[686,690,1955,1980]" pageId="32" pageNumber="293">.</paragraph>
<subSubSection pageId="33" pageNumber="294" type="etymology">
<paragraph blockId="33.[151,1437,153,2015]" pageId="33" pageNumber="294">
<emphasis bold="true" box="[199,334,153,179]" pageId="33" pageNumber="294">Etymology.</emphasis>
The specific epithet is a patronymic in honor of Dr Charles Haddad who collected the
<typeStatus box="[1314,1362,153,179]" pageId="33" pageNumber="294">type</typeStatus>
specimens and for his contribution to arachnology in
<collectingCountry box="[691,837,189,215]" name="South Africa" pageId="33" pageNumber="294">South Africa</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection pageId="33" pageNumber="294" type="diagnosis">
<paragraph blockId="33.[151,1437,153,2015]" pageId="33" pageNumber="294">
<emphasis bold="true" box="[199,319,224,251]" pageId="33" pageNumber="294">Diagnosis.</emphasis>
Males of
<taxonomicName authority="Ríos-Tamayo &amp; Lyle, 2020" authorityName="Ríos-Tamayo &amp; Lyle" authorityYear="2020" box="[426,543,225,251]" class="Arachnida" family="Nemesiidae" genus="Lepthercus" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="294" phylum="Arthropoda" rank="species" species="haddadi" status="sp. nov.">
<emphasis box="[426,543,225,251]" italics="true" pageId="33" pageNumber="294">L. haddadi</emphasis>
</taxonomicName>
<emphasis bold="true" box="[549,635,225,251]" pageId="33" pageNumber="294">
<taxonomicNameLabel box="[549,635,225,251]" pageId="33" pageNumber="294" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, can be recognized inside of the “Group haddadi” by a very robust, darkly coloured tibia I (
<figureCitation box="[330,451,261,287]" captionStart="FIGURE" captionStartId="31.[151,251,1721,1746]" captionTargetBox="[151,1436,561,1696]" captionTargetId="figure@31.[151,1436,561,1696]" captionTargetPageId="31" captionText="FIGURE. 19. Lepthercus haddadi sp. nov. (male holotype, NCA 2007/1486). A cephalothorax, dorsal view; B abdomen, dorsal view; C sternum, ventral view; D chelicerae, arrow indicates the intercheliceral tumescence with setae, prolateral view; E right pedipalp, arrows indicates the palpal tibia base dorsally very convex and the development of the anterior side of the retrolateral side of the ventral excavation, retrolateral view, F right pedipalp, arrows indicates the few proventral setae and the development of the anterior side of the retrolateral side of the ventral excavation, prolateral view; GI copulatory bulb, G retrolateral view, H ventral view, I prolateral view; J tibia-metatarsus I, showing the very robust tibia with a robust spur, arrow indicates the patch of numerous black blunt spinules, prolateral view; K tibia I, arrow indicates the strong curved spine near of the base of the megaspine, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3765509" httpUri="https://zenodo.org/record/3765509/files/figure.png" pageId="33" pageNumber="294">Fig. 19J, K</figureCitation>
) with a robust spur; and by a more elongated copulatory bulb (
<figureCitation box="[1128,1205,261,287]" captionStart="FIGURE" captionStartId="31.[151,251,1721,1746]" captionTargetBox="[151,1436,561,1696]" captionTargetId="figure@31.[151,1436,561,1696]" captionTargetPageId="31" captionText="FIGURE. 19. Lepthercus haddadi sp. nov. (male holotype, NCA 2007/1486). A cephalothorax, dorsal view; B abdomen, dorsal view; C sternum, ventral view; D chelicerae, arrow indicates the intercheliceral tumescence with setae, prolateral view; E right pedipalp, arrows indicates the palpal tibia base dorsally very convex and the development of the anterior side of the retrolateral side of the ventral excavation, retrolateral view, F right pedipalp, arrows indicates the few proventral setae and the development of the anterior side of the retrolateral side of the ventral excavation, prolateral view; GI copulatory bulb, G retrolateral view, H ventral view, I prolateral view; J tibia-metatarsus I, showing the very robust tibia with a robust spur, arrow indicates the patch of numerous black blunt spinules, prolateral view; K tibia I, arrow indicates the strong curved spine near of the base of the megaspine, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3765509" httpUri="https://zenodo.org/record/3765509/files/figure.png" pageId="33" pageNumber="294">Fig. 19</figureCitation>
GI). Females can be distinguished from
<taxonomicName authority="Ríos-Tamayo &amp; Lyle, 2020" authorityName="Ríos-Tamayo &amp; Lyle" authorityYear="2020" box="[364,528,297,323]" class="Arachnida" family="Nemesiidae" genus="Lepthercus" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="294" phylum="Arthropoda" rank="species" species="engelbrechti" status="sp. nov.">
<emphasis box="[364,528,297,323]" italics="true" pageId="33" pageNumber="294">L. engelbrechti</emphasis>
</taxonomicName>
<emphasis bold="true" box="[535,622,297,323]" pageId="33" pageNumber="294">
<taxonomicNameLabel box="[535,622,297,323]" pageId="33" pageNumber="294" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, by the shape of their spermathecae with a longer stalk (
<figureCitation box="[1238,1340,297,323]" captionStart="FIGURE" captionStartId="32.[151,251,1919,1944]" captionTargetBox="[151,1436,207,1894]" captionTargetId="figure@32.[151,1436,181,1894]" captionTargetPageId="32" captionText="FIGURE. 20. Lepthercus haddadi sp. nov. (female paratype, NCA 2007/3847). A habitus, dorsal view; B sternum, ventral view; C chelicerae, prolateral view; D spermathecae" figureDoi="http://doi.org/10.5281/zenodo.3765511" httpUri="https://zenodo.org/record/3765511/files/figure.png" pageId="33" pageNumber="294">Fig. 20D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="34" lastPageNumber="295" pageId="33" pageNumber="294" type="description">
<paragraph blockId="33.[151,1437,153,2015]" pageId="33" pageNumber="294">
<emphasis bold="true" box="[199,342,332,359]" pageId="33" pageNumber="294">Description.</emphasis>
Male
<typeStatus box="[408,504,333,359]" pageId="33" pageNumber="294">holotype</typeStatus>
(NCA 2007/1486,
<figureCitation box="[709,784,333,359]" captionStart="FIGURE" captionStartId="31.[151,251,1721,1746]" captionTargetBox="[151,1436,561,1696]" captionTargetId="figure@31.[151,1436,561,1696]" captionTargetPageId="31" captionText="FIGURE. 19. Lepthercus haddadi sp. nov. (male holotype, NCA 2007/1486). A cephalothorax, dorsal view; B abdomen, dorsal view; C sternum, ventral view; D chelicerae, arrow indicates the intercheliceral tumescence with setae, prolateral view; E right pedipalp, arrows indicates the palpal tibia base dorsally very convex and the development of the anterior side of the retrolateral side of the ventral excavation, retrolateral view, F right pedipalp, arrows indicates the few proventral setae and the development of the anterior side of the retrolateral side of the ventral excavation, prolateral view; GI copulatory bulb, G retrolateral view, H ventral view, I prolateral view; J tibia-metatarsus I, showing the very robust tibia with a robust spur, arrow indicates the patch of numerous black blunt spinules, prolateral view; K tibia I, arrow indicates the strong curved spine near of the base of the megaspine, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3765509" httpUri="https://zenodo.org/record/3765509/files/figure.png" pageId="33" pageNumber="294">Fig. 19</figureCitation>
AK). Total length 15.95.
<emphasis box="[1068,1176,333,359]" italics="true" pageId="33" pageNumber="294">Carapace</emphasis>
(
<figureCitation box="[1189,1290,333,359]" captionStart="FIGURE" captionStartId="31.[151,251,1721,1746]" captionTargetBox="[151,1436,561,1696]" captionTargetId="figure@31.[151,1436,561,1696]" captionTargetPageId="31" captionText="FIGURE. 19. Lepthercus haddadi sp. nov. (male holotype, NCA 2007/1486). A cephalothorax, dorsal view; B abdomen, dorsal view; C sternum, ventral view; D chelicerae, arrow indicates the intercheliceral tumescence with setae, prolateral view; E right pedipalp, arrows indicates the palpal tibia base dorsally very convex and the development of the anterior side of the retrolateral side of the ventral excavation, retrolateral view, F right pedipalp, arrows indicates the few proventral setae and the development of the anterior side of the retrolateral side of the ventral excavation, prolateral view; GI copulatory bulb, G retrolateral view, H ventral view, I prolateral view; J tibia-metatarsus I, showing the very robust tibia with a robust spur, arrow indicates the patch of numerous black blunt spinules, prolateral view; K tibia I, arrow indicates the strong curved spine near of the base of the megaspine, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3765509" httpUri="https://zenodo.org/record/3765509/files/figure.png" pageId="33" pageNumber="294">Fig. 19A</figureCitation>
): length 6.09, width 4.71, with lateral dark marginal bristles, with a little pubescence.
<emphasis box="[941,1118,369,395]" italics="true" pageId="33" pageNumber="294">Cephalic region</emphasis>
: length 3.80; clypeus almost absent, with 6 bristles on the edge, 2 bristles in front of the OQ and 4 between the PME, line of bristles until de fovea.
<emphasis box="[151,220,442,467]" italics="true" pageId="33" pageNumber="294">Fovea</emphasis>
: width 0.45, almost straight (slightly recurved).
<emphasis box="[754,995,441,467]" italics="true" pageId="33" pageNumber="294">Ocular measurements</emphasis>
: AME 0.16, ALE 0.29, PME 0.20, PLE 0.26, OQ length 0.57, 1.03 width; AMEALE 0.07, PMEPLE 0.04, AMEPME 0.09, ALEPLE 0.08, AMEAME 0.10, OMPOMP 0.39.
<emphasis box="[415,535,513,539]" italics="true" pageId="33" pageNumber="294">Chelicerae</emphasis>
: length 3.53, width 1.93, with dark dorsalanterior bristles, intercheliceral tumescence small, well-marked with setae (
<figureCitation box="[571,674,549,575]" captionStart="FIGURE" captionStartId="31.[151,251,1721,1746]" captionTargetBox="[151,1436,561,1696]" captionTargetId="figure@31.[151,1436,561,1696]" captionTargetPageId="31" captionText="FIGURE. 19. Lepthercus haddadi sp. nov. (male holotype, NCA 2007/1486). A cephalothorax, dorsal view; B abdomen, dorsal view; C sternum, ventral view; D chelicerae, arrow indicates the intercheliceral tumescence with setae, prolateral view; E right pedipalp, arrows indicates the palpal tibia base dorsally very convex and the development of the anterior side of the retrolateral side of the ventral excavation, retrolateral view, F right pedipalp, arrows indicates the few proventral setae and the development of the anterior side of the retrolateral side of the ventral excavation, prolateral view; GI copulatory bulb, G retrolateral view, H ventral view, I prolateral view; J tibia-metatarsus I, showing the very robust tibia with a robust spur, arrow indicates the patch of numerous black blunt spinules, prolateral view; K tibia I, arrow indicates the strong curved spine near of the base of the megaspine, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3765509" httpUri="https://zenodo.org/record/3765509/files/figure.png" pageId="33" pageNumber="294">Fig. 19D</figureCitation>
). Fang groove with 8 promarginal teeth and 14 mesobasal denticles.
<emphasis box="[151,235,585,611]" italics="true" pageId="33" pageNumber="294">Labium</emphasis>
: length 0.45, width 0.81.
<emphasis box="[521,618,585,611]" italics="true" pageId="33" pageNumber="294">Maxillae</emphasis>
: length 1.98, width 0.85, with 70 cuspules on inner corner, prolateral face slightly curved, without soft area, with long uniformly distributed hairs, serrula present (
<figureCitation box="[1115,1198,621,647]" captionStart="FIGURE" captionStartId="7.[151,251,1647,1672]" captionTargetBox="[151,1436,381,1622]" captionTargetId="figure@7.[151,1436,381,1622]" captionTargetPageId="7" captionText="FIGURE. 2. Preening comb. A Lepthercus haddadi sp. nov. (female paratype, NCA 2007/3847). Maxillary cuspules (males). B L. dippenaarae. C L. kwazuluensis. Serrula. D L. mandelai sp. nov. (female, NCA 2014/415A); E L. filmeri sp. nov. (female, NCA 91/1437); F L. haddadi sp. nov. (male, NCA 2007/1486); G L. haddadi sp. nov. (female, NCA 2007/3847); H L. engelbrechti sp. nov. (female, NCA 2018/385); I L. engelbrechti sp. nov. (male, NCA 2018/384); J L. dregei Purcell, 1902 (female, NCA 2012/2333); K L. sofiae sp. nov. (male, NCA 97/395)." figureDoi="http://doi.org/10.5281/zenodo.3765471" httpUri="https://zenodo.org/record/3765471/files/figure.png" pageId="33" pageNumber="294">Fig. 2F</figureCitation>
).
<emphasis box="[1217,1308,621,647]" italics="true" pageId="33" pageNumber="294">Sternum</emphasis>
(
<figureCitation box="[1323,1424,621,647]" captionStart="FIGURE" captionStartId="31.[151,251,1721,1746]" captionTargetBox="[151,1436,561,1696]" captionTargetId="figure@31.[151,1436,561,1696]" captionTargetPageId="31" captionText="FIGURE. 19. Lepthercus haddadi sp. nov. (male holotype, NCA 2007/1486). A cephalothorax, dorsal view; B abdomen, dorsal view; C sternum, ventral view; D chelicerae, arrow indicates the intercheliceral tumescence with setae, prolateral view; E right pedipalp, arrows indicates the palpal tibia base dorsally very convex and the development of the anterior side of the retrolateral side of the ventral excavation, retrolateral view, F right pedipalp, arrows indicates the few proventral setae and the development of the anterior side of the retrolateral side of the ventral excavation, prolateral view; GI copulatory bulb, G retrolateral view, H ventral view, I prolateral view; J tibia-metatarsus I, showing the very robust tibia with a robust spur, arrow indicates the patch of numerous black blunt spinules, prolateral view; K tibia I, arrow indicates the strong curved spine near of the base of the megaspine, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3765509" httpUri="https://zenodo.org/record/3765509/files/figure.png" pageId="33" pageNumber="294">Fig. 19C</figureCitation>
): length 3.20, maximum width 2.56.
<emphasis box="[541,645,657,683]" italics="true" pageId="33" pageNumber="294">Abdomen</emphasis>
(
<figureCitation box="[661,762,657,683]" captionStart="FIGURE" captionStartId="31.[151,251,1721,1746]" captionTargetBox="[151,1436,561,1696]" captionTargetId="figure@31.[151,1436,561,1696]" captionTargetPageId="31" captionText="FIGURE. 19. Lepthercus haddadi sp. nov. (male holotype, NCA 2007/1486). A cephalothorax, dorsal view; B abdomen, dorsal view; C sternum, ventral view; D chelicerae, arrow indicates the intercheliceral tumescence with setae, prolateral view; E right pedipalp, arrows indicates the palpal tibia base dorsally very convex and the development of the anterior side of the retrolateral side of the ventral excavation, retrolateral view, F right pedipalp, arrows indicates the few proventral setae and the development of the anterior side of the retrolateral side of the ventral excavation, prolateral view; GI copulatory bulb, G retrolateral view, H ventral view, I prolateral view; J tibia-metatarsus I, showing the very robust tibia with a robust spur, arrow indicates the patch of numerous black blunt spinules, prolateral view; K tibia I, arrow indicates the strong curved spine near of the base of the megaspine, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3765509" httpUri="https://zenodo.org/record/3765509/files/figure.png" pageId="33" pageNumber="294">Fig. 19B</figureCitation>
): length 6.32, covered by dark hair and bristles. PMS: length 0.69; PLS: length of basal:medial:apical segments 1.33:1.04:1.64; total length 4.01.
<emphasis box="[1073,1344,693,719]" italics="true" pageId="33" pageNumber="294">Lengths of legs and palp</emphasis>
: I: 4.20, 2.68, 2.85 (width/length: 0.69), 2.44, 1.70, 13.87. II: 4.09, 2.47, 2.52, 2.57, 1.66, 13.31. III: 3.68, 1.93, 2.18, 3.06, 1.59, 12.44. IV: 4.44, 2.24, 3.05, 3.85, 1.77, 15.35. Palp: 2.28, 1.20, 1.69, —, 0.91, 6.08.
</paragraph>
<paragraph blockId="33.[151,1437,153,2015]" pageId="33" pageNumber="294">
<emphasis box="[199,327,801,827]" italics="true" pageId="33" pageNumber="294">Chaetotaxy</emphasis>
: Leg I: femur, 1-1-1-1 d (centrals), 1 P A; patella, 1 P A; tibia, 1-1 P, 1-2-1 V, very swollen, a strong cuticular spur, with an apical curved megaspine and a strong curved spine near of the base of the megaspine (
<figureCitation captionStart="FIGURE" captionStartId="31.[151,251,1721,1746]" captionTargetBox="[151,1436,561,1696]" captionTargetId="figure@31.[151,1436,561,1696]" captionTargetPageId="31" captionText="FIGURE. 19. Lepthercus haddadi sp. nov. (male holotype, NCA 2007/1486). A cephalothorax, dorsal view; B abdomen, dorsal view; C sternum, ventral view; D chelicerae, arrow indicates the intercheliceral tumescence with setae, prolateral view; E right pedipalp, arrows indicates the palpal tibia base dorsally very convex and the development of the anterior side of the retrolateral side of the ventral excavation, retrolateral view, F right pedipalp, arrows indicates the few proventral setae and the development of the anterior side of the retrolateral side of the ventral excavation, prolateral view; GI copulatory bulb, G retrolateral view, H ventral view, I prolateral view; J tibia-metatarsus I, showing the very robust tibia with a robust spur, arrow indicates the patch of numerous black blunt spinules, prolateral view; K tibia I, arrow indicates the strong curved spine near of the base of the megaspine, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3765509" httpUri="https://zenodo.org/record/3765509/files/figure.png" pageId="33" pageNumber="294">Fig. 19K</figureCitation>
); metatarsus, 1-1-2 V, presence of a protuberance (nearly twothirds of the segment from its base), covered with numerous black blunt spinules (
<figureCitation box="[567,663,909,935]" captionStart="FIGURE" captionStartId="31.[151,251,1721,1746]" captionTargetBox="[151,1436,561,1696]" captionTargetId="figure@31.[151,1436,561,1696]" captionTargetPageId="31" captionText="FIGURE. 19. Lepthercus haddadi sp. nov. (male holotype, NCA 2007/1486). A cephalothorax, dorsal view; B abdomen, dorsal view; C sternum, ventral view; D chelicerae, arrow indicates the intercheliceral tumescence with setae, prolateral view; E right pedipalp, arrows indicates the palpal tibia base dorsally very convex and the development of the anterior side of the retrolateral side of the ventral excavation, retrolateral view, F right pedipalp, arrows indicates the few proventral setae and the development of the anterior side of the retrolateral side of the ventral excavation, prolateral view; GI copulatory bulb, G retrolateral view, H ventral view, I prolateral view; J tibia-metatarsus I, showing the very robust tibia with a robust spur, arrow indicates the patch of numerous black blunt spinules, prolateral view; K tibia I, arrow indicates the strong curved spine near of the base of the megaspine, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3765509" httpUri="https://zenodo.org/record/3765509/files/figure.png" pageId="33" pageNumber="294">Fig. 19J</figureCitation>
); tarsus, 0. Leg II: femur, 1-1-1-1 d, 1-1 PA; patella, 1-1 P; tibia, 1-1 P, 1-1-3 V; metatarsus, 1-1-1 P, 2-3-3 V; tarsus, 0. Leg III: femur, 1-3-2-3 D; patella, 1-1 P, 1 R; tibia, 1-1 P, 1-1 D, 1-1 R, 2-2-2 V; metatarsus, 1-1-1-1 P, 1-1-1 D-P, 1-1-1-1 D-R, 2-3-3 V; tarsus, 0. Leg IV: femur, 1-1-1-1 D; patella, 1 R; tibia, 1-1 P, 1-1 R, 2-2-2 V; metatarsus, 1-1-1-1 P, 1-1-1 D-P, 1-2-1-1 D-R, 2-1-2-3 V; tarsus, 0. Palp (
<figureCitation captionStart="FIGURE" captionStartId="31.[151,251,1721,1746]" captionTargetBox="[151,1436,561,1696]" captionTargetId="figure@31.[151,1436,561,1696]" captionTargetPageId="31" captionText="FIGURE. 19. Lepthercus haddadi sp. nov. (male holotype, NCA 2007/1486). A cephalothorax, dorsal view; B abdomen, dorsal view; C sternum, ventral view; D chelicerae, arrow indicates the intercheliceral tumescence with setae, prolateral view; E right pedipalp, arrows indicates the palpal tibia base dorsally very convex and the development of the anterior side of the retrolateral side of the ventral excavation, retrolateral view, F right pedipalp, arrows indicates the few proventral setae and the development of the anterior side of the retrolateral side of the ventral excavation, prolateral view; GI copulatory bulb, G retrolateral view, H ventral view, I prolateral view; J tibia-metatarsus I, showing the very robust tibia with a robust spur, arrow indicates the patch of numerous black blunt spinules, prolateral view; K tibia I, arrow indicates the strong curved spine near of the base of the megaspine, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3765509" httpUri="https://zenodo.org/record/3765509/files/figure.png" pageId="33" pageNumber="294">Fig. 19E, F</figureCitation>
): femur, 1-1-1-1 d, 1 PA; patella, 1 pa; tibia, 1 R, (width/length: 0.68); tarsus, 0.
</paragraph>
<paragraph blockId="33.[151,1437,153,2015]" pageId="33" pageNumber="294">
<emphasis box="[199,302,1089,1115]" italics="true" pageId="33" pageNumber="294">Scopulae</emphasis>
: Metatarsi: I, sparse, around the black blunt spinules; II, dense (1:2 A), more abundant in the anterior part, divided by fine setae; IIIIV, 0. Tarsi: III, dense, uniformly distributed throughout the segment; III, dense, uniformly distributed and divided by a narrow band of setae; IV, sparse, uniformly distributed and divided by a wide band of setae.
<emphasis box="[433,589,1197,1223]" italics="true" pageId="33" pageNumber="294">Trichobothria</emphasis>
: Tibiae: I 10-11; II 10-10; III 10-11; IV 12-11. Metatarsi: I (6)1(2)1(4)1; II (3)1(2)1(4)1; III (4)1(1)1(3)1(6)1; IV (4)1(3)1(6)1. Tarsi: I 13; II 12; III 11; IV 13.
<emphasis box="[1082,1262,1233,1259]" italics="true" pageId="33" pageNumber="294">Preening combs</emphasis>
: metatarsus: II, 4 PV; III, 2 PV-3 RV; IV, 2 PV-3 RV.
<emphasis box="[560,744,1269,1295]" italics="true" pageId="33" pageNumber="294">Copulatory bulb</emphasis>
: long, with a curved embolus tapering gradually (
<figureCitation box="[1297,1375,1269,1295]" captionStart="FIGURE" captionStartId="31.[151,251,1721,1746]" captionTargetBox="[151,1436,561,1696]" captionTargetId="figure@31.[151,1436,561,1696]" captionTargetPageId="31" captionText="FIGURE. 19. Lepthercus haddadi sp. nov. (male holotype, NCA 2007/1486). A cephalothorax, dorsal view; B abdomen, dorsal view; C sternum, ventral view; D chelicerae, arrow indicates the intercheliceral tumescence with setae, prolateral view; E right pedipalp, arrows indicates the palpal tibia base dorsally very convex and the development of the anterior side of the retrolateral side of the ventral excavation, retrolateral view, F right pedipalp, arrows indicates the few proventral setae and the development of the anterior side of the retrolateral side of the ventral excavation, prolateral view; GI copulatory bulb, G retrolateral view, H ventral view, I prolateral view; J tibia-metatarsus I, showing the very robust tibia with a robust spur, arrow indicates the patch of numerous black blunt spinules, prolateral view; K tibia I, arrow indicates the strong curved spine near of the base of the megaspine, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3765509" httpUri="https://zenodo.org/record/3765509/files/figure.png" pageId="33" pageNumber="294">Fig. 19</figureCitation>
GI), spermatic duct slightly curved.
<emphasis box="[502,567,1305,1331]" italics="true" pageId="33" pageNumber="294">Color</emphasis>
: Overall dark brownreddish (chelicerae, MetatarsusTarsus I darker). Abdomen mottled with light spots distributed in all the dorsum; spinnerets orange, ventrally dark.
</paragraph>
<paragraph blockId="33.[151,1437,153,2015]" pageId="33" pageNumber="294">
Female
<typeStatus box="[289,385,1377,1403]" pageId="33" pageNumber="294">paratype</typeStatus>
(NCA 2007/3847,
<figureCitation box="[602,683,1377,1403]" captionStart="FIGURE" captionStartId="32.[151,251,1919,1944]" captionTargetBox="[151,1436,207,1894]" captionTargetId="figure@32.[151,1436,181,1894]" captionTargetPageId="32" captionText="FIGURE. 20. Lepthercus haddadi sp. nov. (female paratype, NCA 2007/3847). A habitus, dorsal view; B sternum, ventral view; C chelicerae, prolateral view; D spermathecae" figureDoi="http://doi.org/10.5281/zenodo.3765511" httpUri="https://zenodo.org/record/3765511/files/figure.png" pageId="33" pageNumber="294">Fig. 20</figureCitation>
AD). Total length 15.46.
<emphasis box="[979,1089,1377,1403]" italics="true" pageId="33" pageNumber="294">Carapace</emphasis>
(
<figureCitation box="[1106,1212,1377,1403]" captionStart="FIGURE" captionStartId="32.[151,251,1919,1944]" captionTargetBox="[151,1436,207,1894]" captionTargetId="figure@32.[151,1436,181,1894]" captionTargetPageId="32" captionText="FIGURE. 20. Lepthercus haddadi sp. nov. (female paratype, NCA 2007/3847). A habitus, dorsal view; B sternum, ventral view; C chelicerae, prolateral view; D spermathecae" figureDoi="http://doi.org/10.5281/zenodo.3765511" httpUri="https://zenodo.org/record/3765511/files/figure.png" pageId="33" pageNumber="294">Fig. 20A</figureCitation>
): length 5.18, width 3.61.
<emphasis box="[214,395,1413,1439]" italics="true" pageId="33" pageNumber="294">Cephalic region</emphasis>
: length 3.24, clypeus small (almost absent) with 6 marginal bristles; 7 bristles in front of the OQ and 6 between the PME.
<emphasis box="[482,561,1450,1475]" italics="true" pageId="33" pageNumber="294">Fovea:</emphasis>
width 0.28, almost straight (slightly procurved).
<emphasis box="[1115,1360,1449,1475]" italics="true" pageId="33" pageNumber="294">Ocular measurements</emphasis>
: AME 0.11, ALE 0.29, PME 0.19, PLE 0.22, OQ length 0.50, width 0.87; AMEALE 0.06, PMEPLE 0.04, AMEPME 0.07, ALEPLE 0.06, AMEAME 0.10, PMEPME 0.29.
<emphasis box="[829,952,1521,1547]" italics="true" pageId="33" pageNumber="294">Chelicerae</emphasis>
(
<figureCitation box="[970,1077,1521,1547]" captionStart="FIGURE" captionStartId="32.[151,251,1919,1944]" captionTargetBox="[151,1436,207,1894]" captionTargetId="figure@32.[151,1436,181,1894]" captionTargetPageId="32" captionText="FIGURE. 20. Lepthercus haddadi sp. nov. (female paratype, NCA 2007/3847). A habitus, dorsal view; B sternum, ventral view; C chelicerae, prolateral view; D spermathecae" figureDoi="http://doi.org/10.5281/zenodo.3765511" httpUri="https://zenodo.org/record/3765511/files/figure.png" pageId="33" pageNumber="294">Fig. 20C</figureCitation>
): length 3.08, width 1.95; with dorsalretrolateral dark bristles. Fang groove with 8 promarginal teeth and 12 mesobasal denticles.
<emphasis box="[1266,1352,1557,1583]" italics="true" pageId="33" pageNumber="294">Labium</emphasis>
: length 0.41, width 0.80 with one cuspule.
<emphasis box="[551,651,1593,1619]" italics="true" pageId="33" pageNumber="294">Maxillae</emphasis>
: length 1.87, width 0.91, with 86 cuspules on inner corner, prolateral face slightly curved, soft area long, with long uniformly distributed hairs, serrula present on anterior inner corner (
<figureCitation box="[160,252,1665,1691]" captionStart="FIGURE" captionStartId="7.[151,251,1647,1672]" captionTargetBox="[151,1436,381,1622]" captionTargetId="figure@7.[151,1436,381,1622]" captionTargetPageId="7" captionText="FIGURE. 2. Preening comb. A Lepthercus haddadi sp. nov. (female paratype, NCA 2007/3847). Maxillary cuspules (males). B L. dippenaarae. C L. kwazuluensis. Serrula. D L. mandelai sp. nov. (female, NCA 2014/415A); E L. filmeri sp. nov. (female, NCA 91/1437); F L. haddadi sp. nov. (male, NCA 2007/1486); G L. haddadi sp. nov. (female, NCA 2007/3847); H L. engelbrechti sp. nov. (female, NCA 2018/385); I L. engelbrechti sp. nov. (male, NCA 2018/384); J L. dregei Purcell, 1902 (female, NCA 2012/2333); K L. sofiae sp. nov. (male, NCA 97/395)." figureDoi="http://doi.org/10.5281/zenodo.3765471" httpUri="https://zenodo.org/record/3765471/files/figure.png" pageId="33" pageNumber="294">Fig. 2G</figureCitation>
).
<emphasis box="[274,367,1665,1691]" italics="true" pageId="33" pageNumber="294">Sternum</emphasis>
(
<figureCitation box="[385,491,1665,1691]" captionStart="FIGURE" captionStartId="32.[151,251,1919,1944]" captionTargetBox="[151,1436,207,1894]" captionTargetId="figure@32.[151,1436,181,1894]" captionTargetPageId="32" captionText="FIGURE. 20. Lepthercus haddadi sp. nov. (female paratype, NCA 2007/3847). A habitus, dorsal view; B sternum, ventral view; C chelicerae, prolateral view; D spermathecae" figureDoi="http://doi.org/10.5281/zenodo.3765511" httpUri="https://zenodo.org/record/3765511/files/figure.png" pageId="33" pageNumber="294">Fig. 20B</figureCitation>
): length 2.47, maximum width 2.17.
<emphasis box="[919,1025,1665,1691]" italics="true" pageId="33" pageNumber="294">Abdomen</emphasis>
(
<figureCitation box="[1042,1151,1665,1691]" captionStart="FIGURE" captionStartId="32.[151,251,1919,1944]" captionTargetBox="[151,1436,207,1894]" captionTargetId="figure@32.[151,1436,181,1894]" captionTargetPageId="32" captionText="FIGURE. 20. Lepthercus haddadi sp. nov. (female paratype, NCA 2007/3847). A habitus, dorsal view; B sternum, ventral view; C chelicerae, prolateral view; D spermathecae" figureDoi="http://doi.org/10.5281/zenodo.3765511" httpUri="https://zenodo.org/record/3765511/files/figure.png" pageId="33" pageNumber="294">Fig. 20A</figureCitation>
): length 7.42, with small bristles in the dorsum. PMS: length 0.77; PLS: length of basal:medial:apical segments 1.26:0.79:1.45; total length 3.50.
<emphasis box="[214,491,1737,1763]" italics="true" pageId="33" pageNumber="294">Lengths of legs and palp</emphasis>
: I: 3.42, 2.32, 2.26, 1.74, 1.21, 10.95. II: 3.05, 1.99, 1.88, 1,63, 1.14, 9.69. III: 2.71, 1.59, 1.50, 2.12, 1.10, 9.02. IV: 3.37, 2.05, 2.43, 2.92, 1.21, 11.98. Palp: 2.56, 1.55, 1.44, —, 1.53, 7.08.
</paragraph>
<paragraph blockId="33.[151,1437,153,2015]" pageId="33" pageNumber="294">
<emphasis box="[199,327,1809,1835]" italics="true" pageId="33" pageNumber="294">Chaetotaxy</emphasis>
: Leg I: femur, 1-1-1-1-1-1-1 d, 1 PA; patella, 0; tibia, 1-3 v; metatarsus, 2-1-2 V; tarsus, 0. Leg II: femur, 1-1-1-1-1-1-1-1 d, 1-1 PA; patella, 1 pa; tibia, 1-1 P, 1-2 V; metatarsus, 2-2-3 V; tarsus, 0. Leg III: femur, 1-1-1-1 d, 1-1 RA; patella, 1-1 P, 1 R; tibia, 1-1 P, 1-1 D, 1 R, 2-2-3 V; metatarsus, 1-1-1 P, 1-1-1 D-P, 1-1-1 D-R, 2-3-3 V; tarsus, 0. Leg IV: femur, 1-1-1-1-1-1 d; patella, 1 P; tibia, 1-1 P, 2-2-2 V; metatarsus, 1-1-1 P, 1-1-1 D-P, 1-1-1 D-R, 1-1-1 R, 2-3-3 V; tarsus, 0. Palp: femur, 1 pa; patella, 0; tibia, 1p, 2-2v, 3 VA; tarsus, 2 VB.
</paragraph>
<paragraph blockId="33.[151,1437,153,2015]" lastBlockId="34.[151,1437,153,395]" lastPageId="34" lastPageNumber="295" pageId="33" pageNumber="294">
<emphasis box="[199,302,1989,2015]" italics="true" pageId="33" pageNumber="294">Scopulae</emphasis>
: Metatarsi: I, dense, uniformly distributed, divided by setae; II, dense (1:2A), divided by setae; IIIIV, without scopula. Tarsi: I, dense, uniformly distributed; II dense, uniformly distributed, divided by setae; IIIIV, sparse, uniformly distributed and divided by a wide band of setae (IV wider).
<emphasis box="[1054,1210,189,215]" italics="true" pageId="34" pageNumber="295">Trichobothria</emphasis>
: Tibiae: palp 9-9; I 11-10; II 10-10; III 9-10; IV 10-11. Metatarsi: I (6)1(2)1(3)1; II (6)1(2)1(5)1; III (4)1(2)1(7)1; IV (5)1(1)1(4)1(6)1. Tarsi: palp 11; I 13; II 12; III 12; IV 13.
<emphasis box="[603,783,261,287]" italics="true" pageId="34" pageNumber="295">Preening combs</emphasis>
(
<figureCitation box="[798,890,261,287]" captionStart="FIGURE" captionStartId="7.[151,251,1647,1672]" captionTargetBox="[151,1436,381,1622]" captionTargetId="figure@7.[151,1436,381,1622]" captionTargetPageId="7" captionText="FIGURE. 2. Preening comb. A Lepthercus haddadi sp. nov. (female paratype, NCA 2007/3847). Maxillary cuspules (males). B L. dippenaarae. C L. kwazuluensis. Serrula. D L. mandelai sp. nov. (female, NCA 2014/415A); E L. filmeri sp. nov. (female, NCA 91/1437); F L. haddadi sp. nov. (male, NCA 2007/1486); G L. haddadi sp. nov. (female, NCA 2007/3847); H L. engelbrechti sp. nov. (female, NCA 2018/385); I L. engelbrechti sp. nov. (male, NCA 2018/384); J L. dregei Purcell, 1902 (female, NCA 2012/2333); K L. sofiae sp. nov. (male, NCA 97/395)." figureDoi="http://doi.org/10.5281/zenodo.3765471" httpUri="https://zenodo.org/record/3765471/files/figure.png" pageId="34" pageNumber="295">Fig. 2A</figureCitation>
): metatarsus: II 4 PV; III 4 PV3 RV; IV 3 PV2 RV.
<emphasis box="[201,360,297,323]" italics="true" pageId="34" pageNumber="295">Spermathecae</emphasis>
: small, tubulars and curved (
<figureCitation box="[700,808,297,323]" captionStart="FIGURE" captionStartId="32.[151,251,1919,1944]" captionTargetBox="[151,1436,207,1894]" captionTargetId="figure@32.[151,1436,181,1894]" captionTargetPageId="32" captionText="FIGURE. 20. Lepthercus haddadi sp. nov. (female paratype, NCA 2007/3847). A habitus, dorsal view; B sternum, ventral view; C chelicerae, prolateral view; D spermathecae" figureDoi="http://doi.org/10.5281/zenodo.3765511" httpUri="https://zenodo.org/record/3765511/files/figure.png" pageId="34" pageNumber="295">Fig. 20D</figureCitation>
).
<emphasis box="[830,895,297,323]" italics="true" pageId="34" pageNumber="295">Color</emphasis>
: Overall orangeyellowish (chelicerae darker). Abdomen mottled with light spots distributed in all the dorsum; spinnerets orange, ventrally dark.
</paragraph>
</subSubSection>
<subSubSection box="[199,1058,368,395]" pageId="34" pageNumber="295" type="distribution">
<paragraph blockId="34.[151,1437,153,395]" box="[199,1058,368,395]" pageId="34" pageNumber="295">
<emphasis bold="true" box="[199,353,368,395]" pageId="34" pageNumber="295">Distribution.</emphasis>
Eastern and
<collectingRegion box="[497,653,369,395]" country="South Africa" name="Western Cape" pageId="34" pageNumber="295">Western Cape</collectingRegion>
Provinces,
<collectingCountry box="[784,927,369,395]" name="South Africa" pageId="34" pageNumber="295">South Africa</collectingCountry>
(
<figureCitation box="[942,1045,369,395]" captionStart="FIGURE" captionStartId="35.[151,251,1982,2007]" captionTargetBox="[177,1411,472,1957]" captionTargetId="figure@35.[177,1411,417,1957]" captionText="FIGURE. 22. Lepthercus rattrayi Hewitt, 1917. (female paratype, NM 16647). A habitus, dorsal view; B sternum, ventral view; C chelicerae, prolateral view; D spermathecae." figureDoi="http://doi.org/10.5281/zenodo.3765515" httpUri="https://zenodo.org/record/3765515/files/figure.png" pageId="34" pageNumber="295">Fig. 22B</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>