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<document id="2AF73064A2E1DB3891753C5D6A27FF1B" ID-DOI="10.11646/phytotaxa.515.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="8061143" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="jonas" IM.materialsCitations_approvedBy="jonas" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="jonas" IM.taxonomicNames_approvedBy="jonas" IM.treatmentCitations_approvedBy="jonas" IM.treatments_approvedBy="jonas" checkinTime="1687282820830" checkinUser="plazi" docAuthor="Beek, Abraham Van De" docDate="2021" docId="03DE3646FF99FFCBFF43FC73FEB4FA5A" docLanguage="en" docName="Phytotaxa.515.1.1.pdf" docOrigin="Phytotaxa 515 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.515.1.1" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.3:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="3" docTitle="Rubus exsuccus Steudel ex Richard 1847" docType="treatment" docVersion="2" lastPageNumber="23" masterDocId="FFE74E3EFF8FFFDDFFCBFFE1FF96FF93" masterDocTitle="Rubi Capenses: a further contribution to the knowledge of the genus Rubus (Rosaceae) in South Africa" masterLastPageNumber="71" masterPageNumber="1" pageNumber="23" updateTime="1687379631607" updateUser="ExternalLinkService">
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<mods:title id="406574E106C4BF62CBBC96741BC5D3D2">Rubi Capenses: a further contribution to the knowledge of the genus Rubus (Rosaceae) in South Africa</mods:title>
</mods:titleInfo>
<mods:name id="FF7B52C130E4057D9FF8049DF9D979A7" type="personal">
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<mods:namePart id="1A6733B1F923F016232A9495638B8EE0">Beek, Abraham Van De</mods:namePart>
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<mods:title id="1021DDF47EC232047CB692D9137DBEC7">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="279F73AB4BE481F028586AF4E1489FE2">
<mods:date id="8B4E348E4F59EA8EE609A8FD56BA79A1">2021</mods:date>
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<mods:number id="21A88CECEB6DFBD710AB7AB144E9176B">2021-08-19</mods:number>
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<mods:number id="B30D98F3038A2959986CBBAB315751F0">515</mods:number>
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<mods:url id="7942B336B830D8C0FFB111523824E179">http://dx.doi.org/10.11646/phytotaxa.515.1.1</mods:url>
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<mods:classification id="D5795E18ACAFF1BFAC7B04582DB364F2">journal article</mods:classification>
<mods:identifier id="DB2E2D62719975425FB227A187DACB78" type="DOI">10.11646/phytotaxa.515.1.1</mods:identifier>
<mods:identifier id="D4A3281C90C5D8059047FC97B38485F1" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="2AB5FF26871CF420E59D07E1704AE4C4" type="Zenodo-Dep">8061143</mods:identifier>
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<treatment id="03DE3646FF99FFCBFF43FC73FEB4FA5A" ID-DOI="http://doi.org/10.5281/zenodo.8067178" ID-Zenodo-Dep="8067178" LSID="urn:lsid:plazi:treatment:03DE3646FF99FFCBFF43FC73FEB4FA5A" httpUri="http://treatment.plazi.org/id/03DE3646FF99FFCBFF43FC73FEB4FA5A" lastPageNumber="23" pageId="22" pageNumber="23">
<subSubSection id="C36DD4DBFF99FFCBFF43FC73FD41FC3E" box="[136,727,914,941]" pageId="22" pageNumber="23" type="nomenclature">
<paragraph id="8BC88750FF99FFCBFF43FC73FD41FC3E" blockId="22.[136,1452,914,1085]" box="[136,727,914,941]" pageId="22" pageNumber="23">
<emphasis id="B9035B42FF99FFCBFF43FC73FF0EFC3E" bold="true" box="[136,152,914,941]" pageId="22" pageNumber="23">f.</emphasis>
<taxonomicName id="4C77FCD3FF99FFCBFF6FFC73FD44FC3E" ID-CoL="4TK4C" authority="Steud. ex Richard (1847: 256)" authorityName="Steudel ex Richard" authorityPageNumber="256" authorityYear="1847" box="[164,722,914,941]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="22" pageNumber="23" phylum="Tracheophyta" rank="species" species="exsuccus">
<emphasis id="B9035B42FF99FFCBFF6FFC73FEF7FC3E" bold="true" box="[164,353,914,941]" italics="true" pageId="22" pageNumber="23">Rubus exsuccus</emphasis>
<materialsCitation id="3B1F8D0DFF99FFCBFEA6FC73FD44FC3E" box="[365,722,914,941]" collectingDate="1839-12-28" collectionCode="P, A, BM, BR, GH, K, L, LG, MPU, S" country="Ethiopia" county="Dsche-nausa'" location="In" municipality="Aber" pageId="22" pageNumber="23" specimenCode="P00374023, A 00056042, BM000622346, BR 842, GH 00056041, K000310009, K000310011, K000310012, L 0019729, LG0000090026836, MPU024530, P00374017, P00374018, S12-13510" specimenCount="2" typeStatus="lectotype">
Steud. ex
<bibRefCitation id="EFE6FAA1FF99FFCBFE2FFC73FD44FC3E" author="Richard, A." box="[484,722,914,941]" pageId="22" pageNumber="23" refId="ref49050" refString="Richard, A. (1847), Vol. 1. Arthus Bertrand, Parisiis, xi + 472 pp. https: // doi. org / 10.5962 / bhl. title. 334" type="book" year="1847">Richard (1847: 256)</bibRefCitation>
</materialsCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C36DD4DBFF99FFCBFD2AFC72FB80FBAE" pageId="22" pageNumber="23" type="materials_examined">
<paragraph id="8BC88750FF99FFCBFD2AFC72FB80FBAE" blockId="22.[136,1452,914,1085]" pageId="22" pageNumber="23">
<materialsCitation id="3B1F8D0DFF99FFCBFD2AFC72FC3DFC67" collectingDate="1839-12-28" collectionCode="P" collectorName="Schimper" country="Ethiopia" county="Dsche-nausa'" location="Dsche-nausa'" pageId="22" pageNumber="23" specimenCode="P00374023" specimenCount="2" typeStatus="lectotype">
<typeStatus id="54CC39F2FF99FFCBFD2AFC72FCC7FC3E" box="[737,849,915,941]" pageId="22" pageNumber="23">Lectotype</typeStatus>
(designated here):—ABYSSINIA [
<collectingCountry id="F360C7C0FF99FFCBFB20FC73FAE0FC3F" box="[1259,1398,914,940]" name="Ethiopia" pageId="22" pageNumber="23">ETHIOPIA</collectingCountry>
]. In umbrosis lateris borealis montis Aber prope
<collectingCounty id="62A9FFDCFF99FFCBFD4BFC57FC8AFC43" box="[640,796,950,976]" pageId="22" pageNumber="23">Dsche-nausa</collectingCounty>
.
<date id="FFC9A190FF99FFCBFCE3FC57FB95FC43" box="[808,1027,950,976]" pageId="22" pageNumber="23" value="1839-12-28">
<collectingDate id="EF8D5878FF99FFCBFCE3FC57FB95FC43" box="[808,1027,950,976]" pageId="22" pageNumber="23" value="1839-12-28">28 December 1839</collectingDate>
</date>
,
<emphasis id="B9035B42FF99FFCBFBC4FC57FB3DFC43" box="[1039,1195,950,976]" italics="true" pageId="22" pageNumber="23">
<collectorName id="2682E286FF99FFCBFBC4FC57FBEFFC43" box="[1039,1145,950,976]" pageId="22" pageNumber="23">Schimper</collectorName>
867
</emphasis>
(
<typeStatus id="54CC39F2FF99FFCBFB70FC57FAB2FC43" box="[1211,1316,950,976]" pageId="22" pageNumber="23">lectotype</typeStatus>
<specimenCode id="DBD12F2BFF99FFCBFAE7FC57FA3DFC43" box="[1324,1451,950,976]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="22" pageNumber="23" type="Herbarium">P00374023</specimenCode>
[https://science.mnhn.fr/institution/mnhn/collection/p/item/p00374023]
</materialsCitation>
; isolectotypes
<specimenCode id="DBD12F2BFF99FFCBFB92FC3BFB7CFC66" box="[1113,1258,986,1013]" collectionCode="A" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="22" pageNumber="23" type="Herbarium">A 00056042</specimenCode>
,
<specimenCode id="DBD12F2BFF99FFCBFB30FC3BFA31FC67" box="[1275,1447,986,1012]" collectionCode="BM" country="United Kingdom" name="Bristol Museum" pageId="22" pageNumber="23">BM000622346</specimenCode>
,
<specimenCode id="DBD12F2BFF99FFCBFF43FC1EFF74FB8B" box="[136,226,1022,1049]" collectionCode="BR" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection" pageId="22" pageNumber="23">BR 842</specimenCode>
202,
<specimenCode id="DBD12F2BFF99FFCBFEE3FC1FFE5DFB8A" box="[296,459,1022,1049]" collectionCode="GH" country="USA" lsid="urn:lsid:biocol.org:col:15631" name="Harvard University - Gray Herbarium" pageId="22" pageNumber="23" type="Herbarium">GH 00056041</specimenCode>
(strong sample),
<specimenCode id="DBD12F2BFF99FFCBFD51FC1FFCBBFB8B" box="[666,813,1022,1048]" collectionCode="K" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="22" pageNumber="23" type="Herbarium">K000310009</specimenCode>
(the right hand branch),
<specimenCode id="DBD12F2BFF99FFCBFB9EFC1FFB7FFB8B" box="[1109,1257,1022,1048]" collectionCode="K" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="22" pageNumber="23" type="Herbarium">K000310011</specimenCode>
,
<specimenCode id="DBD12F2BFF99FFCBFB3CFC1FFA1AFB8B" box="[1271,1420,1022,1048]" collectionCode="K" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="22" pageNumber="23" type="Herbarium">K000310012</specimenCode>
,
<specimenCode id="DBD12F2BFF99FFCBFA50FC1EFF7BFBAF" collectionCode="L" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="22" pageNumber="23" type="Herbarium">L 0019729</specimenCode>
,
<specimenCode id="DBD12F2BFF99FFCBFF33FBC3FE41FBAF" box="[248,471,1058,1084]" collectionCode="LG" country="Belgium" name="Université de Liège" pageId="22" pageNumber="23" type="Herbarium">LG0000090026836</specimenCode>
,
<specimenCode id="DBD12F2BFF99FFCBFE2AFBC3FDE3FBAE" box="[481,629,1058,1085]" collectionCode="MPU" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15519" lsid="urn:lsid:biocol.org:col:15519" name="Université Montpellier 2" pageId="22" pageNumber="23" type="Herbarium">MPU024530</specimenCode>
,
<specimenCode id="DBD12F2BFF99FFCBFD4BFBC3FC94FBAF" box="[640,770,1058,1084]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="22" pageNumber="23" type="Herbarium">P00374017</specimenCode>
,
<specimenCode id="DBD12F2BFF99FFCBFCC5FBC3FC06FBAF" box="[782,912,1058,1084]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="22" pageNumber="23" type="Herbarium">P00374018</specimenCode>
,
<specimenCode id="DBD12F2BFF99FFCBFC50FBC3FB80FBAE" box="[923,1046,1058,1085]" collectionCode="S" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="22" pageNumber="23" type="Herbarium">S12-13510</specimenCode>
</paragraph>
</subSubSection>
<subSubSection id="C36DD4DBFF99FFCBFF43FB8BFAB1FAAB" pageId="22" pageNumber="23" type="discussion">
<paragraph id="8BC88750FF99FFCBFF43FB8BFD0CFB5F" blockId="22.[136,1451,1130,1481]" pageId="22" pageNumber="23">
<emphasis id="B9035B42FF99FFCBFF43FB8BFF42FB17" bold="true" box="[136,212,1130,1156]" pageId="22" pageNumber="23">Notes:</emphasis>
—The specimen
<specimenCode id="DBD12F2BFF99FFCBFE5FFB8BFDB4FB17" box="[404,546,1130,1156]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="22" pageNumber="23" type="Herbarium">P003740122</specimenCode>
does not have a printed label.
<collectorName id="2682E286FF99FFCBFCB0FB8AFC1AFB16" box="[891,908,1131,1157]" pageId="22" pageNumber="23">It</collectorName>
is similar to the
<typeStatus id="54CC39F2FF99FFCBFB82FB8AFBEFFB16" box="[1097,1145,1131,1157]" pageId="22" pageNumber="23">type</typeStatus>
collected by
<collectorName id="2682E286FF99FFCBFADBFB8BFAEDFB17" box="[1296,1403,1130,1156]" pageId="22" pageNumber="23">Schimper</collectorName>
and conserved by
<collectorName id="2682E286FF99FFCBFEEAFB6FFE7CFB3A" box="[289,490,1166,1193]" pageId="22" pageNumber="23">Steudel. However</collectorName>
, the number on the label is different (807).
<collectorName id="2682E286FF99FFCBFC0DFB6FFC6EFB3B" box="[966,1016,1166,1192]" pageId="22" pageNumber="23">This</collectorName>
might be a mistake, but the uncertainty makes it unsuitable for selection as a
<typeStatus id="54CC39F2FF99FFCBFDE0FB53FD03FB5F" box="[555,661,1202,1228]" pageId="22" pageNumber="23">lectotype</typeStatus>
.
</paragraph>
<paragraph id="8BC88750FF99FFCBFF76FB37FAB1FAAB" blockId="22.[136,1451,1130,1481]" pageId="22" pageNumber="23">
This species has long patent reddish brown hairs on the stem and in the inflorescence; the serrature is rather obtuse, rounded or triangular, with short mucros; prickles from
<quantity id="4C8F2AB5FF99FFCBFC8FFB1BFC36FA86" box="[836,928,1274,1301]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="22" pageNumber="23" unit="mm" value="2.0" valueMax="3.0" valueMin="1.0">13 mm</quantity>
broad base are compressed, strongly declining, sometimes somewhat curved, up to
<quantity id="4C8F2AB5FF99FFCBFDD2FAFFFDE0FAAA" box="[537,630,1310,1337]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="22" pageNumber="23" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">45 mm</quantity>
long, stipules
<quantity id="4C8F2AB5FF99FFCBFCD2FAFFFC05FAAA" box="[793,915,1310,1337]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="1.5" metricValueMin="1.1" pageId="22" pageNumber="23" unit="mm" value="13.0" valueMax="15.0" valueMin="11.0">1115 mm</quantity>
, linear-lanceolate; ovaries glabrous.
</paragraph>
</subSubSection>
<subSubSection id="C36DD4DBFF99FFCBFF76FAA3FEB4FA5A" pageId="22" pageNumber="23" type="materials_examined">
<paragraph id="8BC88750FF99FFCBFF76FAA3FEA1FA12" blockId="22.[136,1451,1130,1481]" pageId="22" pageNumber="23">
<emphasis id="B9035B42FF99FFCBFF76FAA3FEF7FACE" bold="true" box="[189,353,1346,1373]" pageId="22" pageNumber="23">Specimens:—</emphasis>
<materialsCitation id="3B1F8D0DFF99FFCBFEAAFAA3FEA5FA12" collectingDate="1852-10-27" collectionCode="P" country="Ethiopia" elevation="1829" location="Gomienque" municipality="Sabra" pageId="22" pageNumber="23" specimenCode="P02550711" specimenCount="1">
<emphasis id="B9035B42FF99FFCBFEAAFAA3FD38FACF" bold="true" box="[353,686,1346,1372]" pageId="22" pageNumber="23">
<collectingCountry id="F360C7C0FF99FFCBFEAAFAA3FE6EFACF" box="[353,504,1346,1372]" name="Ethiopia" pageId="22" pageNumber="23">ABYSSINIA</collectingCountry>
[
<collectingCountry id="F360C7C0FF99FFCBFDC4FAA3FD34FACF" box="[527,674,1346,1372]" name="Ethiopia" pageId="22" pageNumber="23">ETHIOPIA</collectingCountry>
].
</emphasis>
<location id="8EA8D18BFF99FFCBFD75FAA3FCD2FACF" LSID="urn:lsid:plazi:treatment:03DE3646FF99FFCBFF43FC73FEB4FA5A:8EA8D18BFF99FFCBFD75FAA3FCD2FACF" box="[702,836,1346,1372]" country="Ethiopia" municipality="Sabra" name="Gomienque" pageId="22" pageNumber="23">Gomiénqué</location>
,
<collectingMunicipality id="6BAC1D2AFF99FFCBFC93FAA3FC0DFACF" box="[856,923,1346,1372]" pageId="22" pageNumber="23">Sabra</collectingMunicipality>
,
<quantity id="4C8F2AB5FF99FFCBFC7BFAA3FB9AFACF" box="[944,1036,1346,1372]" metricMagnitude="3" metricUnit="m" metricValue="1.8288" pageId="22" pageNumber="23" unit="ft" value="6000.0">
<elevation id="005A6063FF99FFCBFC7BFAA3FB9AFACF" box="[944,1036,1346,1372]" metricMagnitude="3" metricUnit="m" metricValue="1.8288" pageId="22" pageNumber="23" unit="ft" value="6000.0">6000 ft</elevation>
</quantity>
,
<date id="FFC9A190FF99FFCBFBEAFAA3FB65FACE" box="[1057,1267,1346,1373]" pageId="22" pageNumber="23" value="1852-10-27">
<collectingDate id="EF8D5878FF99FFCBFBEAFAA3FB65FACE" box="[1057,1267,1346,1373]" pageId="22" pageNumber="23" value="1852-10-27">27 October 1852</collectingDate>
</date>
,
<emphasis id="B9035B42FF99FFCBFACCFAA3FA3DFACF" box="[1287,1451,1346,1372]" italics="true" pageId="22" pageNumber="23">Schimper 796</emphasis>
(
<specimenCode id="DBD12F2BFF99FFCBFF5BFA87FE83FA12" box="[144,277,1382,1409]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="22" pageNumber="23" type="Herbarium">
<collectionCode id="ED661F95FF99FFCBFF5BFA87FF36FA12" box="[144,160,1382,1409]" pageId="22" pageNumber="23">P</collectionCode>
02550711
</specimenCode>
/4)
</materialsCitation>
.
</paragraph>
<paragraph id="8BC88750FF99FFCBFF76FA6AFEB4FA5A" blockId="22.[136,1451,1130,1481]" pageId="22" pageNumber="23">
<materialsCitation id="3B1F8D0DFF99FFCBFF12FA6BFE88FA5A" collectingDate="1961-01-30" collectionCode="P" country="Tanzania" elevation="1800" location="Milo Mission" pageId="22" pageNumber="23" specimenCode="P02550730" specimenCount="1" stateProvince="Njombe">
<emphasis id="B9035B42FF99FFCBFF12FA6BFEE2FA37" bold="true" box="[217,372,1418,1444]" pageId="22" pageNumber="23">
<collectingCountry id="F360C7C0FF99FFCBFF12FA6BFEF8FA37" box="[217,366,1418,1444]" name="Tanzania" pageId="22" pageNumber="23">TANZANIA</collectingCountry>
.
</emphasis>
<collectingRegion id="49B349B2FF99FFCBFE4AFA6BFDD6FA37" box="[385,576,1418,1444]" country="Tanzania" name="Njombe" pageId="22" pageNumber="23">Njombe District</collectingRegion>
, valley below
<location id="8EA8D18BFF99FFCBFD3DFA6BFC01FA37" LSID="urn:lsid:plazi:treatment:03DE3646FF99FFCBFF43FC73FEB4FA5A:8EA8D18BFF99FFCBFD3DFA6BFC01FA37" box="[758,919,1418,1444]" country="Tanzania" name="Milo Mission" pageId="22" pageNumber="23" stateProvince="Njombe">Milo Mission</location>
,
<quantity id="4C8F2AB5FF99FFCBFC62FA6BFB90FA36" box="[937,1030,1418,1445]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="22" pageNumber="23" unit="m" value="1800.0">
<elevation id="005A6063FF99FFCBFC62FA6BFB90FA36" box="[937,1030,1418,1445]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="22" pageNumber="23" unit="m" value="1800.0">1800 m</elevation>
</quantity>
,
<date id="FFC9A190FF99FFCBFBD2FA6BFB77FA37" box="[1049,1249,1418,1445]" pageId="22" pageNumber="23" value="1961-01-30">
<collectingDate id="EF8D5878FF99FFCBFBD2FA6BFB77FA37" box="[1049,1249,1418,1445]" pageId="22" pageNumber="23" value="1961-01-30">30 January 1961</collectingDate>
</date>
,
<emphasis id="B9035B42FF99FFCBFB3FFA6BFA3DFA37" box="[1268,1451,1418,1444]" italics="true" pageId="22" pageNumber="23">Richards 14053</emphasis>
(
<specimenCode id="DBD12F2BFF99FFCBFF5BFA4FFE81FA5A" box="[144,279,1454,1481]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="22" pageNumber="23" type="Herbarium">
<collectionCode id="ED661F95FF99FFCBFF5BFA4FFF36FA5A" box="[144,160,1454,1481]" pageId="22" pageNumber="23">P</collectionCode>
02550730
</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>