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<document id="B06554D9CF5322B1CEEFD3FDDA1F5ABA" ID-DOI="10.5281/zenodo.13203160" ID-ISSN="1407-8619" ID-Zenodo-Dep="13203160" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722657382176" checkinUser="felipe" docAuthor="Fanti, Fabrizio &amp; Müller, Patrick" docDate="2022" docId="03DAD52BFFB9FFF4FD32BC155E5FA546" docLanguage="en" docName="BalticJColeopterol.22.2.331-380.pdf" docOrigin="Baltic Journal of Coleopterology 22 (2)" docStyle="DocumentStyle:9720869F1FB170B2A509257D1FA394CD.2:BalticJColeop.2022.journal_article" docStyleId="9720869F1FB170B2A509257D1FA394CD" docStyleName="BalticJColeop.2022.journal_article" docStyleVersion="2" docTitle="Sanaungulus myitkyinaensis Fanti &amp; Müller, 2022, sp. nov." docType="treatment" docVersion="3" lastPageNumber="361" masterDocId="FFE3AD53FFA5FFEAFF9ABD325F46A170" masterDocTitle="Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information" masterLastPageNumber="380" masterPageNumber="331" pageNumber="359" updateTime="1722697113453" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="FB2344F04CD4BA29F72262F577A06F8D">Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information</mods:title>
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<mods:name id="E3F24F4A383C798C315EA6D8C1E634B8" type="personal">
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<mods:namePart id="0F392B7869A55641DA1FEFEFEB145D1C">Fanti, Fabrizio</mods:namePart>
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<mods:namePart id="4A42DE782189073E7730F84BD96B8FDC">Müller, Patrick</mods:namePart>
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<mods:title id="88410A5B079A87C1FF4A4B2DE4104E83">Baltic Journal of Coleopterology</mods:title>
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<mods:part id="83204CA2064A361E387DC5BE16EDADDD">
<mods:date id="CB506E748D331BF362BD6D9496DE8522">2022</mods:date>
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<mods:number id="3B366057BA542DD77A673CDA8D0A3437">22</mods:number>
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<treatment id="03DAD52BFFB9FFF4FD32BC155E5FA546" ID-DOI="http://doi.org/10.5281/zenodo.13199829" ID-Zenodo-Dep="13199829" LSID="urn:lsid:plazi:treatment:03DAD52BFFB9FFF4FD32BC155E5FA546" httpUri="http://treatment.plazi.org/id/03DAD52BFFB9FFF4FD32BC155E5FA546" lastPageId="30" lastPageNumber="361" pageId="28" pageNumber="359">
<subSubSection id="C36937B6FFB9FFF6FD32BC155B06A04F" box="[680,1088,295,319]" pageId="28" pageNumber="359" type="nomenclature">
<paragraph id="8BCC643DFFB9FFF6FD32BC155B06A04F" blockId="28.[680,1088,295,350]" box="[680,1088,295,319]" pageId="28" pageNumber="359">
<heading id="D084D351FFB9FFF6FD32BC155B06A04F" bold="true" box="[680,1088,295,319]" fontSize="10" level="2" pageId="28" pageNumber="359" reason="4">
<emphasis id="B907B82FFFB9FFF6FD32BC155B06A04F" bold="true" box="[680,1088,295,319]" pageId="28" pageNumber="359">
<taxonomicName id="4C731FBEFFB9FFF6FD32BC155CA3A04F" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[680,997,295,319]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="359" phylum="Arthropoda" rank="species" species="myitkyinaensis" status="sp. nov.">
<emphasis id="B907B82FFFB9FFF6FD32BC155CA3A04F" bold="true" box="[680,997,295,319]" italics="true" pageId="28" pageNumber="359">Sanaungulus myitkyinaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFB9FFF6FC71BC155B06A04F" box="[1003,1088,295,319]" pageId="28" pageNumber="359" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFB9FFF6FD32BC755C43A02E" box="[680,773,327,350]" pageId="28" pageNumber="359" type="description">
<paragraph id="8BCC643DFFB9FFF6FD32BC755C43A02E" blockId="28.[680,1088,295,350]" box="[680,773,327,350]" pageId="28" pageNumber="359">
(
<figureCitation id="134878B8FFB9FFF6FD2ABC755DBBA02E" box="[688,765,327,350]" captionStart="Fig" captionStartId="29.[113,152,1636,1659]" captionTargetBox="[113,1134,380,1605]" captionTargetId="figure-125@29.[113,1134,380,1605]" captionTargetPageId="29" captionText="Fig. 16. Sanaungulus myitkyinaensis sp. nov. in Burmese amber.A: Holotype, dorso­lateral view; B: Holotype, ventro­lateral view." figureDoi="http://doi.org/10.5281/zenodo.13203211" httpUri="https://zenodo.org/record/13203211/files/figure.png" pageId="28" pageNumber="359">Fig. 16</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFB9FFF6FD32BC4F5C81A3CF" pageId="28" pageNumber="359" type="materials_examined">
<paragraph id="8BCC643DFFB9FFF6FD32BC4F5CC2A363" blockId="28.[680,1172,381,531]" pageId="28" pageNumber="359">
<materialsCitation id="3B1B6E60FFB9FFF6FD32BC4F5CC2A363" collectionCode="BUB" county="Probable" location="Catalog" municipality="Burmese" pageId="28" pageNumber="359" specimenCode="BUB4462" specimenCount="1" stateProvince="Kachin" typeStatus="holotype">
<emphasis id="B907B82FFFB9FFF6FD32BC4F5C51A0E5" bold="true" box="[680,791,381,405]" pageId="28" pageNumber="359">
<typeStatus id="54C8DA9FFFB9FFF6FD32BC4F5C54A0E5" box="[680,786,381,405]" pageId="28" pageNumber="359" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCounty id="62AD1CB1FFB9FFF6FC84BC4F5C3BA0E4" box="[798,893,381,404]" pageId="28" pageNumber="359">Probable</collectingCounty>
male, adult specimen in a
<collectingMunicipality id="6BA8FE47FFB9FFF6FD32BCAF5C42A0C4" box="[680,772,413,436]" pageId="28" pageNumber="359">Burmese</collectingMunicipality>
(
<collectingRegion id="49B7AADFFFB9FFF6FC8ABCAF5C18A0C4" box="[784,862,413,436]" country="Myanmar" name="Kachin" pageId="28" pageNumber="359">Kachin</collectingRegion>
) amber piece:
<location id="8EAC32E6FFB9FFF6FC6ABCAF5B06A0C4" LSID="urn:lsid:plazi:treatment:03DAD52BFFB9FFF4FD32BC155E5FA546:8EAC32E6FFB9FFF6FC6ABCAF5B06A0C4" box="[1008,1088,413,436]" county="Probable" municipality="Burmese" name="Catalog" pageId="28" pageNumber="359" stateProvince="Kachin">Catalog</location>
number SNHM­
<quantity id="4C8BC9D8FFB9FFF6FD62BC8F5C01A0A4" box="[760,839,445,468]" metricMagnitude="2" metricUnit="m" metricValue="1.52654" pageId="28" pageNumber="359" unit="in" value="6010.0">6010 in</quantity>
the
<location id="8EAC32E6FFB9FFF6FCF5BC8F5CE7A084" LSID="urn:lsid:plazi:treatment:03DAD52BFFB9FFF4FD32BC155E5FA546:8EAC32E6FFB9FFF6FCF5BC8F5CE7A084" county="Probable" municipality="Burmese" name="Staatliches Naturhistorisches Museum Braunschweig" pageId="28" pageNumber="359" stateProvince="Kachin">Staatliches Naturhistorisches Museum Braunschweig</location>
(ex
<location id="8EAC32E6FFB9FFF6FC56BCEF5B60A084" LSID="urn:lsid:plazi:treatment:03DAD52BFFB9FFF4FD32BC155E5FA546:8EAC32E6FFB9FFF6FC56BCEF5B60A084" box="[972,1062,477,500]" county="Probable" municipality="Burmese" name="Muller's" pageId="28" pageNumber="359" stateProvince="Kachin">Müllers</location>
collection number:
<specimenCode id="DBD5CC46FFB9FFF6FC92BCCE5C3FA363" box="[776,889,508,531]" collectionCode="BUB" pageId="28" pageNumber="359">BUB4462</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BCC643DFFB9FFF6FD32BF005CBDA319" blockId="28.[680,1172,562,617]" pageId="28" pageNumber="359">
<emphasis id="B907B82FFFB9FFF6FD32BF005C07A33A" bold="true" box="[680,833,562,586]" pageId="28" pageNumber="359">Type locality.</emphasis>
<collectingCountry id="F36424ADFFB9FFF6FCD0BF015CF0A33A" box="[842,950,563,586]" name="Myanmar" pageId="28" pageNumber="359">Myanmar</collectingCountry>
:
<collectingRegion id="49B7AADFFFB9FFF6FC5BBF015B0DA33A" box="[961,1099,563,586]" country="Myanmar" name="Kachin" pageId="28" pageNumber="359">Kachin state</collectingRegion>
, Myitkyina District, Hukawng Valley.
</paragraph>
<paragraph id="8BCC643DFFB9FFF6FD32BFBA5C81A3CF" blockId="28.[680,1172,648,703]" pageId="28" pageNumber="359">
<emphasis id="B907B82FFFB9FFF6FD32BFBA5C7AA3D0" bold="true" box="[680,828,648,672]" pageId="28" pageNumber="359">Type horizon.</emphasis>
Lowermost Cenomanian (98.79 ± 0.62 Ma), mid­Cretaceous.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFB9FFF6FD32BFEC5CE1A2E4" pageId="28" pageNumber="359" type="diagnosis">
<paragraph id="8BCC643DFFB9FFF6FD32BFEC5CE1A2E4" blockId="28.[680,1172,734,916]" pageId="28" pageNumber="359">
<emphasis id="B907B82FFFB9FFF6FD32BFEC5CE2A386" bold="true" box="[680,932,734,758]" pageId="28" pageNumber="359">Differential diagnosis.</emphasis>
<taxonomicName id="4C731FBEFFB9FFF6FC36BFED5DBEA266" authorityName="Fanti &amp; Müller" authorityYear="2022" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="359" phylum="Arthropoda" rank="species" species="myitkyinaensis" status="sp. nov.">
<emphasis id="B907B82FFFB9FFF6FC36BFED5DBEA266" italics="true" pageId="28" pageNumber="359">Sanaungulus myitkyinaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFB9FFF6FD65BFCD5C17A266" box="[767,849,767,790]" pageId="28" pageNumber="359" rank="species">sp. nov.</taxonomicNameLabel>
is characterized by pectinated antennae with an antennal process in antennomeres VIX. The new species is easily distinguishable from other
<taxonomicName id="4C731FBEFFB9FFF6FC0DBE6C5B67A205" authorityName="Fanti, Damgaard &amp; Ellenberger" authorityYear="2018" box="[919,1057,862,885]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="359" phylum="Arthropoda" rank="genus">
<emphasis id="B907B82FFFB9FFF6FC0DBE6C5B67A205" box="[919,1057,862,885]" italics="true" pageId="28" pageNumber="359">Sanaungulus</emphasis>
</taxonomicName>
by its extremely elongated head.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFB9FFF4FD32BE865E05A382" lastPageId="30" lastPageNumber="361" pageId="28" pageNumber="359" type="description">
<paragraph id="8BCC643DFFB9FFF6FD32BE865B2EA57A" blockId="28.[680,1172,948,1034]" pageId="28" pageNumber="359">
<emphasis id="B907B82FFFB9FFF6FD32BE865C74A2BC" bold="true" box="[680,818,948,972]" pageId="28" pageNumber="359">Description.</emphasis>
Adult, winged, rather robust. Probable male, defined on the basis of long antennae. Body length
<quantity id="4C8BC9D8FFB9FFF6FCF9BEC15CF1A57A" box="[867,951,1011,1034]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="28" pageNumber="359" unit="mm" value="3.3">3.3 mm</quantity>
. Entirely brown.
</paragraph>
<paragraph id="8BCC643DFFB9FFF4FD32B9185E05A382" blockId="28.[680,1172,1066,1692]" lastBlockId="30.[151,643,155,754]" lastPageId="30" lastPageNumber="361" pageId="28" pageNumber="359">Head extremely long, curved, surface smooth equipped with only some long hairs. Eyes relatively small, very slightly sub­elliptical, inserted in the upper and lateral part of the head. Mandibles elongated, falciform, slender with a pointed apex. Maxillary palps 4­ segmented with the last palpomere securiform. Labial palps 3­ segmented, with the last palpomere securiform.Antennae 11­ segmented, pectinate, very long, almost reaching the last abdominal segment, antennal insertion in the eyes proximity; scape club­shaped; antennomere II short, slightly swollen apically; antennomere III elongated, filiform, about 1.4 times longer than second; antennomeres IVV extremely long, without antennal process, enlarged (in a shape of a draft) apically, antennomere V the longest; antennomere VI elongated, as long as antennomere IV, equipped near the base with a long antennal process which is very long and rounded apically; antennomere VII shorter than previous one, equipped with a long antennal process near the base; antennomeres VIIIX sub­equal, shorter and sturdier than previous one, each equipped near the base with an antennal process shorter and slightly sturdier than others; antennomere XI rather robust, with slightly squadrate and pointed apex; all the antennomeres pubescent. Pronotum elongated, slightly narrower than head, anterior margin straight and strongly bordered, posterior margin slightly undulate (expecially in the middle) and strongly bordered, sides straight and bordered, corners rounded, pronotum disc almost flat with short pubescence. Elytra short which reveals various abdominal segments, wider than pronotum, gibbous at humeri, parallel­sided, rounded apexes, surface pubescent with confused and impressed punctuation. Metathoracic wings longer than elytra, reaching the last abdominal segments. Sternites transverse, last sternite very wide. Legs thin, long, pubescent; coxae massive; trochanters elongated; femora almost straight, cylindrical, sturdier than tibiae; tibiae, thin, cylindrical, long, without a spur at apex, protibiae shorter than profemora, meso­ and metatibiae longer than meso­ and metafemora. Tarsal formula 5­ 5­ 5; first tarsomere very long, about 2.0 times longer than second; third tarsomere robust; fourth strongly bilobed; fifth tarsomere elongated, thin, curved; claws simple, without denticle at the base.</paragraph>
</subSubSection>
<caption id="DF0C34B5FFB8FFF7FFEBBB565E89A7EB" ID-DOI="http://doi.org/10.5281/zenodo.13203211" ID-Zenodo-Dep="13203211" httpUri="https://zenodo.org/record/13203211/files/figure.png" pageId="29" pageNumber="360" startId="29.[113,152,1636,1659]" targetBox="[113,1134,380,1605]" targetPageId="29" targetType="figure">
<paragraph id="8BCC643DFFB8FFF7FFEBBB565E89A7EB" blockId="29.[113,1134,1636,1691]" pageId="29" pageNumber="360">
Fig. 16.
<taxonomicName id="4C731FBEFFB8FFF7FF5DBB565EB2A70B" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[199,500,1636,1659]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="360" phylum="Arthropoda" rank="species" species="myitkyinaensis" status="sp. nov.">
<emphasis id="B907B82FFFB8FFF7FF5DBB565EB2A70B" box="[199,500,1636,1659]" italics="true" pageId="29" pageNumber="360">Sanaungulus myitkyinaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFB8FFF7FE63BB565D0CA70B" box="[505,586,1636,1659]" pageId="29" pageNumber="360" rank="species">sp. nov.</taxonomicNameLabel>
in Burmese amber. A: Holotype, dorso­lateral view; B: Holotype, ventro­lateral view.
</paragraph>
</caption>
<subSubSection id="C36937B6FFBBFFF4FF0DBE235D59A2EF" pageId="30" pageNumber="361" type="etymology">
<paragraph id="8BCC643DFFBBFFF4FF0DBE235E8DA218" blockId="30.[151,643,785,873]" pageId="30" pageNumber="361">
<emphasis id="B907B82FFFBBFFF4FF0DBE235E5EA259" bold="true" box="[151,280,785,809]" pageId="30" pageNumber="361">Etymology.</emphasis>
Derived from the toponym Myitkyina (district where the amber mines are located) + the Latin suffix ­Á
<emphasis id="B907B82FFFBBFFF4FE00BE605E82A219" box="[410,452,850,873]" italics="true" pageId="30" pageNumber="361">nsis</emphasis>
.
</paragraph>
<paragraph id="8BCC643DFFBBFFF4FF0DBEBA5D59A2EF" blockId="30.[151,543,904,928]" box="[151,543,904,928]" pageId="30" pageNumber="361">
<emphasis id="B907B82FFFBBFFF4FF0DBEBA5E7CA2D0" bold="true" box="[151,314,904,928]" pageId="30" pageNumber="361">Syninclusions.</emphasis>
Air bubbles, detritus.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFBBFFF4FF0DBE8C5E5FA546" pageId="30" pageNumber="361" type="discussion">
<paragraph id="8BCC643DFFBBFFF4FF0DBE8C5E5FA546" blockId="30.[151,643,958,1078]" pageId="30" pageNumber="361">
<emphasis id="B907B82FFFBBFFF4FF0DBE8C5E45A2A6" bold="true" box="[151,259,958,982]" pageId="30" pageNumber="361">Remarks.</emphasis>
Piece of clear, golden amber that measures 13x10x
<quantity id="4C8BC9D8FFBBFFF4FEBCBEED5E20A286" box="[294,358,991,1014]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="30" pageNumber="361" unit="mm" value="3.0">3 mm</quantity>
. The inclusion is complete but the head is bent and the left proleg is not well visible.
</paragraph>
</subSubSection>
</treatment>
</document>