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<document id="EAC61C4F88EAC6013C2D36C8C88752EA" ID-DOI="10.11646/zootaxa.4500.3.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="5297723" ID-ZooBank="A0081D71-D208-40E7-88FA-86CA929E4E80" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630085678920" checkinUser="felipe" docAuthor="Grebennikov, Vasily V." docDate="2018" docId="03D42902EE473844FF47F9DAFD675FA2" docLanguage="en" docName="zootaxa.4500.3.5.pdf" docOrigin="Zootaxa 4500 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nephius argus Grebennikov 2018, sp. nov." docType="treatment" docVersion="4" lastPageNumber="386" masterDocId="FFED517AEE433841FFD0FF9DFFFF5F5A" masterDocTitle="Dryophthorinae weevils (Coleoptera: Curculionidae) of the forest floor in Southeast Asia: DNA analysis of two new Nephius from Vietnam and Taiwan suggests non-monophyly of Stromboscerini" masterLastPageNumber="387" masterPageNumber="381" pageNumber="385" updateTime="1699141661482" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="BE82753940C766785E31422C3ED4A654" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="5397AD533FA061FAFF93874B5144310E">
<mods:title id="281E3F0AD299595DB54CB5A349630599">Dryophthorinae weevils (Coleoptera: Curculionidae) of the forest floor in Southeast Asia: DNA analysis of two new Nephius from Vietnam and Taiwan suggests non-monophyly of Stromboscerini</mods:title>
</mods:titleInfo>
<mods:name id="64375B4AD66FCC3C2C99DF33FD638B39" type="personal">
<mods:role id="BE9768BB24E0D963ABC335DF587E2DDE">
<mods:roleTerm id="97A1FF9379DCD0F77CC631FB8E42E48E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6A85CBB6241C8CE8900325E5AC02F9F9">Grebennikov, Vasily V.</mods:namePart>
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<mods:typeOfResource id="5B3D7535EDEEC48B80641CD6A8E9873A">text</mods:typeOfResource>
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<mods:title id="6B837DFE04257796803F2A517E53934A">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="0B5BA33411F2CAEB4C084F7AB006FB3C">
<mods:date id="466ACA7BFF06C404FCD43637C54A52EF">2018</mods:date>
<mods:detail id="D43D91DCFF7B43873A3F93896BFDCABE" type="pubDate">
<mods:number id="BB3E11826530C1A8BD5E81AB7C666026">2018-10-17</mods:number>
</mods:detail>
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<mods:number id="3BA5ED7A7CC0CAF14CEC856E182D3324">4500</mods:number>
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<mods:number id="E86E4408D1E9F50650757E5EF47495B7">3</mods:number>
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<mods:classification id="8A227C7F55041112B0B5DBA51253CEB0">journal article</mods:classification>
<mods:identifier id="16F4698E576651DFA943B9DDE6C376E5" type="DOI">10.11646/zootaxa.4500.3.5</mods:identifier>
<mods:identifier id="164DF0CA76D660DE6A02ADA75451E75D" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="4672EAD88AD19C98183BF6CB9B57A6C0" type="Zenodo-Dep">5297723</mods:identifier>
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</mods:mods>
<treatment id="03D42902EE473844FF47F9DAFD675FA2" ID-DOI="http://doi.org/10.5281/zenodo.5777717" ID-Zenodo-Dep="5777717" LSID="urn:lsid:plazi:treatment:03D42902EE473844FF47F9DAFD675FA2" httpUri="http://treatment.plazi.org/id/03D42902EE473844FF47F9DAFD675FA2" lastPageId="5" lastPageNumber="386" pageId="4" pageNumber="385">
<subSubSection id="C367CB9FEE473845FF47F9DAFE535938" box="[151,428,1607,1634]" pageId="4" pageNumber="385" type="nomenclature">
<paragraph id="8BC29814EE473845FF47F9DAFE535938" blockId="4.[151,428,1607,1667]" box="[151,428,1607,1634]" pageId="4" pageNumber="385">
<heading id="D08A2F78EE473845FF47F9DAFE535938" bold="true" box="[151,428,1607,1634]" fontSize="11" level="1" pageId="4" pageNumber="385" reason="1">
<emphasis id="B9094406EE473845FF47F9DAFE535938" bold="true" box="[151,428,1607,1634]" pageId="4" pageNumber="385">
<taxonomicName id="4C7DE397EE473845FF47F9DAFEB8593B" authority="Grebennikov, 2018" authorityName="Grebennikov" authorityYear="2018" box="[151,327,1607,1633]" class="Insecta" family="Dryophthoridae" genus="Nephius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="385" phylum="Arthropoda" rank="species" species="argus" status="sp. nov.">
<emphasis id="B9094406EE473845FF47F9DAFEB8593B" bold="true" box="[151,327,1607,1633]" italics="true" pageId="4" pageNumber="385">Nephius argus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23AF97DEE473845FE9FF9D5FE535938" box="[335,428,1608,1634]" pageId="4" pageNumber="385" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C367CB9FEE473845FF47F9F7FCB558F9" pageId="4" pageNumber="385" type="description">
<paragraph id="8BC29814EE473845FF47F9F7FEED59D9" blockId="4.[151,428,1607,1667]" box="[151,274,1642,1667]" pageId="4" pageNumber="385">
<figureCitation id="13468491EE473845FF47F9F7FEED59D9" box="[151,274,1642,1667]" captionStart="FIGURE 2" captionStartId="4.[151,250,1474,1496]" captionTargetBox="[189,1434,292,1452]" captionTargetId="figure-250@4.[151,1436,273,1453]" captionTargetPageId="4" captionText="FIGURE 2. Nephius argus sp. nov., holotype. A–D: habitus, dorsal (A), ventral (B), lateral (C), fronto-lateral (D); E: eyes, ventral; F: antenna, G: tarsus; H–J: aedeagus, dorsal (H), ventral (I), lateral (J); K: sternite 9." figureDoi="http://doi.org/10.5281/zenodo.5297727" httpUri="https://zenodo.org/record/5297727/files/figure.png" pageId="4" pageNumber="385">Figs 2A–K</figureCitation>
</paragraph>
<paragraph id="8BC29814EE473845FF47F92FFCB558F9" blockId="4.[151,1437,1714,2027]" pageId="4" pageNumber="385">
<emphasis id="B9094406EE473845FF47F92FFE555991" bold="true" box="[151,426,1714,1739]" inLineHeading="true" pageId="4" pageNumber="385" reason="1">Diagnostic description.</emphasis>
<typeStatus id="54C626B6EE473845FE62F92FFDE55991" box="[434,538,1714,1739]" pageId="4" pageNumber="385" type="holotype">Holotype</typeStatus>
male (
<figureCitation id="13468491EE473845FDBAF92FFD4F5991" box="[618,688,1714,1739]" captionStart="FIGURE 2" captionStartId="4.[151,250,1474,1496]" captionTargetBox="[189,1434,292,1452]" captionTargetId="figure-250@4.[151,1436,273,1453]" captionTargetPageId="4" captionText="FIGURE 2. Nephius argus sp. nov., holotype. A–D: habitus, dorsal (A), ventral (B), lateral (C), fronto-lateral (D); E: eyes, ventral; F: antenna, G: tarsus; H–J: aedeagus, dorsal (H), ventral (I), lateral (J); K: sternite 9." figureDoi="http://doi.org/10.5281/zenodo.5297727" httpUri="https://zenodo.org/record/5297727/files/figure.png" pageId="4" pageNumber="385">Fig. 2</figureCitation>
); length between anterior edge of pronotum and elytral apex
<quantity id="4C8535F1EE473845FABBF92FFF3959B4" metricMagnitude="-3" metricUnit="m" metricValue="5.95" pageId="4" pageNumber="385" unit="mm" value="5.95">5.95 mm</quantity>
; body background color (not considering whitish pilosity) black (
<figureCitation id="13468491EE473845FC69F94BFBC259B5" box="[953,1085,1750,1775]" captionStart="FIGURE 2" captionStartId="4.[151,250,1474,1496]" captionTargetBox="[189,1434,292,1452]" captionTargetId="figure-250@4.[151,1436,273,1453]" captionTargetPageId="4" captionText="FIGURE 2. Nephius argus sp. nov., holotype. A–D: habitus, dorsal (A), ventral (B), lateral (C), fronto-lateral (D); E: eyes, ventral; F: antenna, G: tarsus; H–J: aedeagus, dorsal (H), ventral (I), lateral (J); K: sternite 9." figureDoi="http://doi.org/10.5281/zenodo.5297727" httpUri="https://zenodo.org/record/5297727/files/figure.png" pageId="4" pageNumber="385">Figs 2A–D</figureCitation>
); rostrum in lateral view more curved (
<figureCitation id="13468491EE473845FF23F967FEB25849" box="[243,333,1786,1811]" captionStart="FIGURE 2" captionStartId="4.[151,250,1474,1496]" captionTargetBox="[189,1434,292,1452]" captionTargetId="figure-250@4.[151,1436,273,1453]" captionTargetPageId="4" captionText="FIGURE 2. Nephius argus sp. nov., holotype. A–D: habitus, dorsal (A), ventral (B), lateral (C), fronto-lateral (D); E: eyes, ventral; F: antenna, G: tarsus; H–J: aedeagus, dorsal (H), ventral (I), lateral (J); K: sternite 9." figureDoi="http://doi.org/10.5281/zenodo.5297727" httpUri="https://zenodo.org/record/5297727/files/figure.png" pageId="4" pageNumber="385">Fig. 2C</figureCitation>
); eyes ventrally clearly separated (
<figureCitation id="13468491EE473845FD07F967FCD05849" box="[727,815,1786,1811]" captionStart="FIGURE 2" captionStartId="4.[151,250,1474,1496]" captionTargetBox="[189,1434,292,1452]" captionTargetId="figure-250@4.[151,1436,273,1453]" captionTargetPageId="4" captionText="FIGURE 2. Nephius argus sp. nov., holotype. A–D: habitus, dorsal (A), ventral (B), lateral (C), fronto-lateral (D); E: eyes, ventral; F: antenna, G: tarsus; H–J: aedeagus, dorsal (H), ventral (I), lateral (J); K: sternite 9." figureDoi="http://doi.org/10.5281/zenodo.5297727" httpUri="https://zenodo.org/record/5297727/files/figure.png" pageId="4" pageNumber="385">Fig. 2E</figureCitation>
); central longitudinal groove on pronotum absent (
<figureCitation id="13468491EE473845FABFF967FF42586D" captionStart="FIGURE 2" captionStartId="4.[151,250,1474,1496]" captionTargetBox="[189,1434,292,1452]" captionTargetId="figure-250@4.[151,1436,273,1453]" captionTargetPageId="4" captionText="FIGURE 2. Nephius argus sp. nov., holotype. A–D: habitus, dorsal (A), ventral (B), lateral (C), fronto-lateral (D); E: eyes, ventral; F: antenna, G: tarsus; H–J: aedeagus, dorsal (H), ventral (I), lateral (J); K: sternite 9." figureDoi="http://doi.org/10.5281/zenodo.5297727" httpUri="https://zenodo.org/record/5297727/files/figure.png" pageId="4" pageNumber="385">Fig. 2A</figureCitation>
); disk of pronotum nearly smooth, with only occasional shallow and small punctures separated by smooth areas 2–3 times their diameter (
<figureCitation id="13468491EE473845FE68F8DFFDED5801" box="[440,530,1858,1883]" captionStart="FIGURE 2" captionStartId="4.[151,250,1474,1496]" captionTargetBox="[189,1434,292,1452]" captionTargetId="figure-250@4.[151,1436,273,1453]" captionTargetPageId="4" captionText="FIGURE 2. Nephius argus sp. nov., holotype. A–D: habitus, dorsal (A), ventral (B), lateral (C), fronto-lateral (D); E: eyes, ventral; F: antenna, G: tarsus; H–J: aedeagus, dorsal (H), ventral (I), lateral (J); K: sternite 9." figureDoi="http://doi.org/10.5281/zenodo.5297727" httpUri="https://zenodo.org/record/5297727/files/figure.png" pageId="4" pageNumber="385">Fig. 2A</figureCitation>
); elytra widest at middle (
<figureCitation id="13468491EE473845FCE7F8DFFC6E5801" box="[823,913,1858,1883]" captionStart="FIGURE 2" captionStartId="4.[151,250,1474,1496]" captionTargetBox="[189,1434,292,1452]" captionTargetId="figure-250@4.[151,1436,273,1453]" captionTargetPageId="4" captionText="FIGURE 2. Nephius argus sp. nov., holotype. A–D: habitus, dorsal (A), ventral (B), lateral (C), fronto-lateral (D); E: eyes, ventral; F: antenna, G: tarsus; H–J: aedeagus, dorsal (H), ventral (I), lateral (J); K: sternite 9." figureDoi="http://doi.org/10.5281/zenodo.5297727" httpUri="https://zenodo.org/record/5297727/files/figure.png" pageId="4" pageNumber="385">Fig. 2A</figureCitation>
); odd interstriae about twice as wide and high, as even interstriae (
<figureCitation id="13468491EE473845FEA8F8FBFE2A5825" box="[376,469,1894,1919]" captionStart="FIGURE 2" captionStartId="4.[151,250,1474,1496]" captionTargetBox="[189,1434,292,1452]" captionTargetId="figure-250@4.[151,1436,273,1453]" captionTargetPageId="4" captionText="FIGURE 2. Nephius argus sp. nov., holotype. A–D: habitus, dorsal (A), ventral (B), lateral (C), fronto-lateral (D); E: eyes, ventral; F: antenna, G: tarsus; H–J: aedeagus, dorsal (H), ventral (I), lateral (J); K: sternite 9." figureDoi="http://doi.org/10.5281/zenodo.5297727" httpUri="https://zenodo.org/record/5297727/files/figure.png" pageId="4" pageNumber="385">Fig. 2A</figureCitation>
); interstriae 2 and 3 at elytral apex each with sizable hemispherical projection (
<figureCitation id="13468491EE473845FABFF8FBFF4258F9" captionStart="FIGURE 2" captionStartId="4.[151,250,1474,1496]" captionTargetBox="[189,1434,292,1452]" captionTargetId="figure-250@4.[151,1436,273,1453]" captionTargetPageId="4" captionText="FIGURE 2. Nephius argus sp. nov., holotype. A–D: habitus, dorsal (A), ventral (B), lateral (C), fronto-lateral (D); E: eyes, ventral; F: antenna, G: tarsus; H–J: aedeagus, dorsal (H), ventral (I), lateral (J); K: sternite 9." figureDoi="http://doi.org/10.5281/zenodo.5297727" httpUri="https://zenodo.org/record/5297727/files/figure.png" pageId="4" pageNumber="385">Fig. 2A</figureCitation>
); male genitalia as in
<figureCitation id="13468491EE473845FE61F817FDCF58F9" box="[433,560,1930,1955]" captionStart="FIGURE 2" captionStartId="4.[151,250,1474,1496]" captionTargetBox="[189,1434,292,1452]" captionTargetId="figure-250@4.[151,1436,273,1453]" captionTargetPageId="4" captionText="FIGURE 2. Nephius argus sp. nov., holotype. A–D: habitus, dorsal (A), ventral (B), lateral (C), fronto-lateral (D); E: eyes, ventral; F: antenna, G: tarsus; H–J: aedeagus, dorsal (H), ventral (I), lateral (J); K: sternite 9." figureDoi="http://doi.org/10.5281/zenodo.5297727" httpUri="https://zenodo.org/record/5297727/files/figure.png" pageId="4" pageNumber="385">Figs 2H–K</figureCitation>
; DNA data as in
<tableCitation id="C6FFADAFEE473845FD20F817FCB958F9" box="[752,838,1930,1955]" captionStart="TABLE 2" captionStartId="2.[151,239,652,675]" captionText="TABLE 2. GenBank accession numbers of Nephius weevils; for those of the remaining 45 herein analysed terminals and for sample codes see Table 2 and Table S1 in Grebennikov (2018b), respectively" pageId="4" pageNumber="385">Table 2</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C367CB9FEE473845FF17F833FCED58B1" pageId="4" pageNumber="385" type="materials_examined">
<paragraph id="8BC29814EE473845FF17F833FCED58B1" blockId="4.[151,1437,1714,2027]" pageId="4" pageNumber="385">
<emphasis id="B9094406EE473845FF17F833FE48589D" bold="true" box="[199,439,1966,1991]" inLineHeading="true" pageId="4" pageNumber="385" reason="1">Material examined.</emphasis>
<materialsCitation id="3B159249EE473845FE15F833FCED58B1" accessionNumber="E10538" collectingDate="2012-05-16" collectionCode="CNC" collectorName="Tam Dao, N &amp; V. Grebennikov" country="Vietnam" location="VIETNAM" pageId="4" pageNumber="385" specimenCount="1" typeStatus="holotype">
<typeStatus id="54C626B6EE473845FE15F833FDD2589D" box="[453,557,1966,1991]" pageId="4" pageNumber="385" type="holotype">Holotype</typeStatus>
male (
<collectionCode id="ED6C00D1EE473845FD58F833FD37589D" box="[648,712,1966,1991]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="4" pageNumber="385">CNC</collectionCode>
): “
<collectingCountry id="F36AD884EE473845FD3CF833FC8D589D" box="[748,882,1966,1991]" name="Vietnam" pageId="4" pageNumber="385">VIETNAM</collectingCountry>
,
<collectorName id="2688FDC2EE473845FC55F832FBE2589D" box="[901,1053,1966,1991]" pageId="4" pageNumber="385">Tam Dao, N</collectorName>
2127'48&quot; E10538'46&quot;,
<date id="FFC3BED4EE473845FAF7F833FA67589D" box="[1319,1432,1966,1991]" pageId="4" pageNumber="385" value="2012-05-16">
<collectingDate id="EF87473CEE473845FAF7F833FA67589D" box="[1319,1432,1966,1991]" pageId="4" pageNumber="385" value="2012-05-16">16.v.2012</collectingDate>
</date>
, 1229m, sift06,
<collectorName id="2688FDC2EE473845FE90F84FFE1458B1" box="[320,491,2002,2027]" pageId="4" pageNumber="385">V.Grebennikov</collectorName>
, “CNCCOLVG00004402.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C367CB9FEE463844FF17FF0AFAFB5FF5" box="[199,1284,151,176]" pageId="5" pageNumber="386" type="distribution">
<paragraph id="8BC29814EE463844FF17FF0AFAFB5FF5" blockId="5.[151,1436,151,248]" box="[199,1284,151,176]" pageId="5" pageNumber="386">
<emphasis id="B9094406EE463844FF17FF0AFE9F5FEA" bold="true" box="[199,352,151,176]" inLineHeading="true" pageId="5" pageNumber="386" reason="1">Distribution.</emphasis>
Known only from the Tam Dao Mountains, northern
<collectingCountry id="F36AD884EE463844FC6AFF0AFBE35FEA" box="[954,1052,151,176]" name="Vietnam" pageId="5" pageNumber="386">Vietnam</collectingCountry>
. Elevation:
<quantity id="4C8535F1EE463844FB71FF0AFAFF5FF5" box="[1185,1280,151,176]" metricMagnitude="3" metricUnit="m" metricValue="1.229" pageId="5" pageNumber="386" unit="m" value="1229.0">1,229 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C367CB9FEE463844FF17FF21FD675FA2" pageId="5" pageNumber="386" type="etymology">
<paragraph id="8BC29814EE463844FF17FF21FD675FA2" blockId="5.[151,1436,151,248]" pageId="5" pageNumber="386">
<emphasis id="B9094406EE463844FF17FF21FEB15F8F" bold="true" box="[199,334,188,213]" inLineHeading="true" pageId="5" pageNumber="386" reason="1">Etymology.</emphasis>
The species epithet is a Latinized Greek mythical name of an Argonaut, co-builder (with the goddess Athena) of Argo; noun in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>
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