treatments-xml/data/03/D1/87/03D187F1FFA9FFB92089FA5DFAD959A7.xml

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<mods:title id="27B9B5B1E54B9F3FA192B0928E09F627">Taxonomy and phylogeny of the Japanese species of the genus Dasycerus Brongniart (Coleoptera: Staphylinidae: Dasycerinae) with description of a new species and notes on D. poseidon Hu &amp; Liang</mods:title>
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<mods:namePart id="9126C42BDBD7C79A4A5768656744A07C">Hashizume, Takuto</mods:namePart>
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<mods:affiliation id="541DB78DC1083556EAD8182D0E407256">Entomological Laboratory, Graduate School of Bioresource and Bioenvironmental Sciences, Kyushu University, Fukuoka, 819 - 0395 Japan. 0214 sakana @ gmail. com; https: // orcid. org / 0000 - 0002 - 4910 - 1836</mods:affiliation>
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<mods:namePart id="E9FDF69A987E2DE6ECDDCA3F7D0B83D3">Maruyama, Munetoshi</mods:namePart>
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<mods:affiliation id="4555FF8B1A54059354B7ADC150C377F1">The Kyushu University Museum, Fukuoka, 812 - 8581 Japan. dendrolasius @ gmail. com; https: // orcid. org / 0000 - 0003 - 4531 - 1008</mods:affiliation>
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<emphasis id="B90CEAF5FFA9FFBC2089FA5DFCDA5D07" bold="true" box="[151,813,1518,1545]" pageId="1" pageNumber="234">
<taxonomicName id="4C784D64FFA9FFBC2089FA5DFD3F5D07" ID-CoL="9GDLD" authority="Hashizume &amp; Maruyama" authorityName="Hashizume &amp; Maruyama" authorityYear="2022" box="[151,712,1518,1545]" class="Insecta" family="Staphylinidae" genus="Dasycerus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="234" phylum="Arthropoda" rank="species" species="occultus" status="sp. nov.">
<emphasis id="B90CEAF5FFA9FFBC2089FA5DFE765D07" bold="true" box="[151,385,1518,1545]" italics="true" pageId="1" pageNumber="234">Dasycerus occultus</emphasis>
Hashizume &amp; Maruyama
</taxonomicName>
<taxonomicNameLabel id="A23F578EFFA9FFBC22D1FA5DFCDA5D07" box="[719,813,1519,1545]" pageId="1" pageNumber="234" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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(
<figureCitation id="13432A62FFA9FFBC2081F9A1FF175D23" box="[159,224,1555,1581]" captionStart="FIGURE 1" captionStartId="3.[151,250,1855,1879]" captionTargetBox="[151,1435,298,1813]" captionTargetId="figure-59@3.[151,1436,270,1829]" captionTargetPageId="3" captionText="FIGURE 1. Dasycerus occultus sp. nov. (A) Habitus in dorsal view. (B) SEM image of pronotum in dorsal view. (C–D) Prothorax in ventral view (black triangles indicate the protergosternal keel). (E–F) Right elytron in dorsolateral view. (G) Aedeagus in lateral view. (H) Female genital chamber in ventral view. Scale bars: 1.0 mm for A; 0.5 mm for C &amp; E; 0.2 mm for G–H." figureDoi="http://doi.org/10.5281/zenodo.6986249" httpUri="https://zenodo.org/record/6986249/files/figure.png" pageId="1" pageNumber="234">Fig. 1</figureCitation>
)
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<treatmentCitationGroup id="AB6811C9FFA9FFBC2089F9CEFD1E5D9D" box="[151,745,1659,1683]" pageId="1" pageNumber="234">
<treatmentCitation id="0AD910F6FFA9FFBC2089F9CEFD7E5D9D" author="Lobl, I. &amp; Calame, F. G." box="[151,649,1659,1683]" page="278" pageId="1" pageNumber="234" year="1996">
<taxonomicName id="4C784D64FFA9FFBC2089F9CEFD7E5D9D" ID-CoL="347NJ" authority=": Lobl &amp; Calame, 1996: 278" authorityName=": Lobl &amp; Calame" authorityPageNumber="278" authorityYear="1996" box="[151,649,1659,1683]" class="Insecta" family="Staphylinidae" genus="Dasycerus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="234" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B90CEAF5FFA9FFBC2089F9CEFE9C5D9D" box="[151,363,1660,1683]" italics="true" pageId="1" pageNumber="234">Dasycerus japonicus</emphasis>
: 
<bibRefCitation id="EFE94B16FFA9FFBC2166F9C9FD7E5D9D" author="Lobl &amp; Calame" box="[376,649,1659,1683]" firstAuthor="Lobl" page="278" pageId="1" pageNumber="234" pagination="247 - 291" refId="ref9214" refString="Lobl, I. &amp; Calame, F. G. (1996) Taxonomy and phylogeny of the Dasycerinae (Coleoptera: Staphylinidae). Journal of Natural History, 30, 247 - 291. https: // doi. org / 10.1080 / 00222939600771151" type="journal article" year="1996">Löbl &amp; Calame, 1996: 278</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(in part).
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</paragraph>
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<emphasis id="B90CEAF5FFA9FFBC2089F90EFDDE5DD8" bold="true" box="[151,553,1724,1750]" pageId="1" pageNumber="234">Specimen examined. Type series.</emphasis>
<materialsCitation id="3B103CBAFFA9FFBC222AF90EFF195C4C" ID-GBIF-Occurrence="3887777323" collectingDate="2020-06-18" collectionCode="CBM-ZI" collectorName="Akiko Saito" country="Japan" determinerName="Hashizume" location="Honshu" pageId="1" pageNumber="234" specimenCode="CBM-ZI / 224407" specimenCount="2" stateProvince="Chiba" typeStatus="holotype">
<typeStatus id="54C38845FFA9FFBC222AF90EFD565DD8" box="[564,673,1724,1750]" pageId="1" pageNumber="234" type="holotype">Holotype</typeStatus>
: male (
<collectionCode id="ED69AE22FFA9FFBC22E7F90EFC965DD8" box="[761,865,1724,1750]" pageId="1" pageNumber="234">CBM-ZI</collectionCode>
), 
<collectingCountry id="F36F7677FFA9FFBC2369F90EFC275DD8" box="[887,976,1724,1750]" name="Japan" pageId="1" pageNumber="234">JAPAN</collectingCountry>
: 
<location id="8EA7603CFFA9FFBC23C1F90EFBCC5DD8" LSID="urn:lsid:plazi:treatment:03D187F1FFA9FFB92089FA5DFAD959A7:8EA7603CFFA9FFBC23C1F90EFBCC5DD8" box="[991,1083,1724,1750]" country="Japan" name="Honshu" pageId="1" pageNumber="234" stateProvince="Chiba">Honshu</location>
: “Honzawa Rindo / Sakamoto / Kamogawa-shi / 
<collectingRegion id="49BCF805FFA9FFBC2171F952FE025DF4" box="[367,501,1760,1786]" country="Japan" name="Chiba" pageId="1" pageNumber="234">Chiba Pref.</collectingRegion>
, 
<date id="FFC61027FFA9FFBC221DF952FD745DF4" box="[515,643,1760,1786]" pageId="1" pageNumber="234" value="2020-06-18">
<collectingDate id="EF82E9CFFFA9FFBC221DF952FD745DF4" box="[515,643,1760,1786]" pageId="1" pageNumber="234" value="2020-06-18">18.VI.2020</collectingDate>
</date>
/ 
<collectorName id="268D5331FFA9FFBC2280F952FCDF5DF4" box="[670,808,1760,1786]" pageId="1" pageNumber="234">Akiko Saito</collectorName>
leg. // 
<specimenCode id="DBDE9E9CFFA9FFBC2361F952FBA55DF4" box="[895,1106,1760,1786]" collectionCode="CBM-ZI" pageId="1" pageNumber="234">CBM-ZI / 224407</specimenCode>
// 
<typeStatus id="54C38845FFA9FFBC2468F952FAFD5DF4" box="[1142,1290,1760,1786]" pageId="1" pageNumber="234" type="holotype">HOLOTYPE</typeStatus>
/ 
<taxonomicName id="4C784D64FFA9FFBC2538F953FEF15C10" ID-CoL="9GDLD" authority="Hashizume &amp; Maruyama, 2022" authorityName="Hashizume &amp; Maruyama" authorityYear="2022" class="Insecta" family="Staphylinidae" genus="Dasycerus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="234" phylum="Arthropoda" rank="species" species="occultus" status="sp. nov.">
<emphasis id="B90CEAF5FFA9FFBC2538F953FA6B5DF4" box="[1318,1436,1761,1786]" italics="true" pageId="1" pageNumber="234">Dasycerus</emphasis>
/ 
<emphasis id="B90CEAF5FFA9FFBC20B7F8B6FEF15C10" box="[169,262,1796,1822]" italics="true" pageId="1" pageNumber="234">occultus</emphasis>
</taxonomicName>
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/ det. 
<determinerName id="7E1BDA23FFA9FFBC2186F8B6FDE15C10" box="[408,534,1796,1822]" pageId="1" pageNumber="234">Hashizume</determinerName>
&amp; Maruyama 2022” (recorded as 
<taxonomicName id="4C784D64FFA9FFBC23B7F8B7FB605C10" ID-CoL="347NJ" authorityName=": Lobl &amp; Calame" authorityYear="1996" box="[937,1175,1797,1822]" class="Insecta" family="Staphylinidae" genus="Dasycerus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="234" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B90CEAF5FFA9FFBC23B7F8B7FB605C10" box="[937,1175,1797,1822]" italics="true" pageId="1" pageNumber="234">Dasycerus japonicus</emphasis>
</taxonomicName>
in 
<bibRefCitation id="EFE94B16FFA9FFBC24DFF8B6FF195C4C" author="Saito &amp; Murakawa" firstAuthor="Saito" pageId="1" pageNumber="234" pagination="82 - 83" refId="ref9314" refString="Saito, A. &amp; Murakawa, I. (2020) Additions and Corrections to &quot; Insect Fauna of the University of Tokyo Chiba Forest &quot; No. 5. Boso no Konchu, 67, 82 - 83. [in Japanese]" type="journal article" year="2020">Saitô &amp; Murakawa (2020))</bibRefCitation>
</materialsCitation>
. 
<materialsCitation id="3B103CBAFFA9FFBC20E1F89BFA765C68" ID-GBIF-Occurrence="3887777337" collectingDate="2020-06-18" collectionCode="CBM-ZI" collectorName="Honshu &amp; Chiba-ken" country="Japan" county="Toyota-shi" determinerName="Hashizume" location="Aichi-ken" municipality="Mennoki-toge" pageId="1" pageNumber="234" specimenCount="9" stateProvince="Chiba" typeStatus="paratype">
<typeStatus id="54C38845FFA9FFBC20E1F89BFE875C4D" box="[255,368,1833,1859]" pageId="1" pageNumber="234" type="paratype">Paratypes</typeStatus>
: 
<emphasis id="B90CEAF5FFA9FFBC219EF89AFE2A5C4C" bold="true" box="[384,477,1832,1858]" pageId="1" pageNumber="234">
<collectorName id="268D5331FFA9FFBC219EF89AFE2A5C4C" box="[384,477,1832,1858]" pageId="1" pageNumber="234">Honshu</collectorName>
</emphasis>
: 
<collectorName id="268D5331FFA9FFBC21ECF89AFD9C5C4C" box="[498,619,1832,1858]" pageId="1" pageNumber="234">Chiba-ken</collectorName>
: 
<specimenCount id="9D7EFD6EFFA9FFBC2265F89AFD335C4D" box="[635,708,1832,1859]" pageId="1" pageNumber="234" type="generic">8 exs.</specimenCount>
, same data as holotype. (
<collectionCode id="ED69AE22FFA9FFBC23E7F89AFB965C4C" box="[1017,1121,1832,1858]" pageId="1" pageNumber="234">CBM-ZI</collectionCode>
: 224405–224406, 224408– 224413) (recorded as 
<taxonomicName id="4C784D64FFA9FFBC218CF8FFFDD45C68" ID-CoL="347NJ" authorityName=": Lobl &amp; Calame" authorityYear="1996" box="[402,547,1869,1895]" class="Insecta" family="Staphylinidae" genus="Dasycerus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="234" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B90CEAF5FFA9FFBC218CF8FFFE515C68" box="[402,422,1869,1894]" italics="true" pageId="1" pageNumber="234">D</emphasis>
. 
<emphasis id="B90CEAF5FFA9FFBC21A8F8FFFDD45C68" box="[438,547,1869,1894]" italics="true" pageId="1" pageNumber="234">japonicus</emphasis>
</taxonomicName>
in 
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; 
<location id="8EA7603CFFA9FFBC238EF8FEFBF25C68" LSID="urn:lsid:plazi:treatment:03D187F1FFA9FFB92089FA5DFAD959A7:8EA7603CFFA9FFBC238EF8FEFBF25C68" box="[912,1029,1868,1894]" county="Toyota-shi" municipality="Mennoki-toge" name="Aichi-ken" pageId="1" pageNumber="234">Aichi-ken</location>
: 
<specimenCount id="9D7EFD6EFFA9FFBC240EF8FEFBBC5C69" box="[1040,1099,1868,1895]" pageId="1" pageNumber="234" type="generic">1 ex.</specimenCount>
, 
<collectingMunicipality id="6BA3AC9DFFA9FFBC2447F8FEFB0C5C68" box="[1113,1275,1868,1894]" pageId="1" pageNumber="234">Mennoki-tôge</collectingMunicipality>
, 
<collectingCounty id="62A64E6BFFA9FFBC2519F8FEFA765C68" box="[1287,1409,1868,1894]" pageId="1" pageNumber="234">Toyota-shi</collectingCounty>
</materialsCitation>
, 
<date id="FFC61027FFA9FFBC2590F8FEFF195C84" pageId="1" pageNumber="234" value="1989-05-02">
2 V 
<materialsCitation id="3B103CBAFFA9FFBC20ADF8C2FF195C84" ID-GBIF-Occurrence="3887777325" box="[179,238,1904,1930]" collectionCode="KUM" collectorName="N. Kanie" country="Japan" county="Shitara-cho" elevation="930" location="Dandouradani" municipality="Tamine" pageId="1" pageNumber="234" specimenCount="1" stateProvince="Chiba" typeStatus="paratype">1989</materialsCitation>
</date>
, 
<collectorName id="268D5331FFA9FFBC20E4F8C3FE975C84" box="[250,352,1904,1931]" pageId="1" pageNumber="234">N. Kanie</collectorName>
leg. (KUM); 
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, alt. 
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</quantity>
, 
<location id="8EA7603CFFA9FFBC22DAF8C2FC915C84" LSID="urn:lsid:plazi:treatment:03D187F1FFA9FFB92089FA5DFAD959A7:8EA7603CFFA9FFBC22DAF8C2FC915C84" box="[708,870,1904,1930]" country="Japan" county="Shitara-cho" municipality="Tamine" name="Dandouradani" pageId="1" pageNumber="234" stateProvince="Chiba">Dandouradani</location>
, 
<collectingMunicipality id="6BA3AC9DFFA9FFBC236FF8C2FC3F5C84" box="[881,968,1904,1930]" pageId="1" pageNumber="234">Tamine</collectingMunicipality>
, 
<collectingCounty id="62A64E6BFFA9FFBC23CDF8C2FBA05C84" box="[979,1111,1904,1930]" pageId="1" pageNumber="234">Shitara-chô</collectingCounty>
, 
<date id="FFC61027FFA9FFBC247DF8C3FB1B5C84" box="[1123,1260,1904,1931]" pageId="1" pageNumber="234" value="2020-08-07">
7 VIII 
<materialsCitation id="3B103CBAFFA9FFBC24AFF8C2FB1B5C84" ID-GBIF-Occurrence="3887777403" box="[1201,1260,1904,1930]" collectionCode="KUM" collectorName="F. Nakano" country="Japan" location="Tottori-ken" municipality="Dai-sen" pageId="1" pageNumber="234" specimenCount="1" stateProvince="Chiba" typeStatus="paratype">2020</materialsCitation>
</date>
, 
<collectorName id="268D5331FFA9FFBC24E6F8C3FA9C5C84" box="[1272,1387,1904,1931]" pageId="1" pageNumber="234">F. Nakano</collectorName>
leg. (KUM); 
<location id="8EA7603CFFA9FFBC20E7F826FE8C5CA0" LSID="urn:lsid:plazi:treatment:03D187F1FFA9FFB92089FA5DFAD959A7:8EA7603CFFA9FFBC20E7F826FE8C5CA0" box="[249,379,1940,1966]" country="Japan" municipality="Dai-sen" name="Tottori-ken" pageId="1" pageNumber="234" stateProvince="Chiba">Tottori-ken</location>
: 
<specimenCount id="9D7EFD6EFFA9FFBC219BF826FE495CA1" box="[389,446,1940,1967]" pageId="1" pageNumber="234" type="generic">1 ex.</specimenCount>
, 
<collectingMunicipality id="6BA3AC9DFFA9FFBC21D7F826FDD45CA0" box="[457,547,1940,1966]" pageId="1" pageNumber="234">Dai-sen</collectingMunicipality>
, 
<materialsCitation id="3B103CBAFFA9FFBC2230F826FE725CDC" ID-GBIF-Occurrence="3887777400" collectingDate="2008-07-06" collectionCode="KUM" collectorName="I. Tanaka &amp; Kyushu" country="Japan" county="Yame-shi" location="Fukuoka-ken" municipality="Yabemura" pageId="1" pageNumber="234" specimenCount="2" stateProvince="Chiba" typeStatus="paratype">
<date id="FFC61027FFA9FFBC2230F826FD5C5CA0" box="[558,683,1940,1967]" pageId="1" pageNumber="234" value="2008-07-06">
<collectingDate id="EF82E9CFFFA9FFBC2230F826FD5C5CA0" box="[558,683,1940,1967]" pageId="1" pageNumber="234" value="2008-07-06">6 VII 2008</collectingDate>
</date>
, 
<collectorName id="268D5331FFA9FFBC22ABF827FCEB5CA0" box="[693,796,1940,1967]" pageId="1" pageNumber="234">I. Tanaka</collectorName>
leg. (
<collectionCode id="ED69AE22FFA9FFBC2345F826FC555CA0" box="[859,930,1940,1966]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="1" pageNumber="234" type="Herbarium">KUM</collectionCode>
); 
<emphasis id="B90CEAF5FFA9FFBC23A8F826FBE45CA0" bold="true" box="[950,1043,1940,1966]" pageId="1" pageNumber="234">
<collectorName id="268D5331FFA9FFBC23A8F826FBE45CA0" box="[950,1043,1940,1966]" pageId="1" pageNumber="234">Kyushu</collectorName>
</emphasis>
: 
<location id="8EA7603CFFA9FFBC243CF826FB4D5CA0" LSID="urn:lsid:plazi:treatment:03D187F1FFA9FFB92089FA5DFAD959A7:8EA7603CFFA9FFBC243CF826FB4D5CA0" box="[1058,1210,1940,1966]" country="Japan" county="Yame-shi" municipality="Yabemura" name="Fukuoka-ken" pageId="1" pageNumber="234" stateProvince="Chiba">Fukuoka-ken</location>
: 
<specimenCount id="9D7EFD6EFFA9FFBC24DAF826FAF05CA1" box="[1220,1287,1940,1967]" pageId="1" pageNumber="234" type="generic">2 exs.</specimenCount>
, 
<location id="8EA7603CFFA9FFBC250DF826FA6E5CA0" LSID="urn:lsid:plazi:treatment:03D187F1FFA9FFB92089FA5DFAD959A7:8EA7603CFFA9FFBC250DF826FA6E5CA0" box="[1299,1433,1940,1966]" country="Japan" county="Yame-shi" municipality="Yabemura" name="Shaka-dake" pageId="1" pageNumber="234" stateProvince="Chiba">Shaka-dake</location>
, 
<collectingMunicipality id="6BA3AC9DFFA9FFBC2089F80AFEFB5CDC" box="[151,268,1976,2002]" pageId="1" pageNumber="234">Yabemura</collectingMunicipality>
, 
<collectingCounty id="62A64E6BFFA9FFBC2106F80AFE725CDC" box="[280,389,1976,2002]" pageId="1" pageNumber="234">Yame-shi</collectingCounty>
</materialsCitation>
, 
<date id="FFC61027FFA9FFBC218FF80AFDE05CDC" box="[401,535,1976,2003]" pageId="1" pageNumber="234" value="2017-06-23">
23 VI 
<materialsCitation id="3B103CBAFFA9FFBC21C2F80AFDE05CDC" ID-GBIF-Occurrence="3887777362" box="[476,535,1976,2002]" collectingDate="2008-07-06" collectionCode="KUM" collectorName="S. Imasaka" country="Japan" county="Yame-shi" location="Fukuoka-ken" municipality="Yabemura" pageId="1" pageNumber="234" specimenCount="1" stateProvince="Chiba" typeStatus="paratype">2017</materialsCitation>
</date>
, 
<collectorName id="268D5331FFA9FFBC223DF80AFD575CDC" box="[547,672,1976,2002]" pageId="1" pageNumber="234">S. Imasaka</collectorName>
leg. (KUM); 
<specimenCount id="9D7EFD6EFFA9FFBC2321F80AFC8E5CDD" box="[831,889,1976,2003]" pageId="1" pageNumber="234" type="generic">1 ex.</specimenCount>
, same data, but 
<materialsCitation id="3B103CBAFFA9FFBC242DF80AFDD25CF8" ID-GBIF-Occurrence="3887777333" collectingDate="2017-07-26" collectionCode="KUM" collectorName="Saga-ken" country="Japan" location="Hiroseyama - Ko" municipality="Arita-cho" pageId="1" pageNumber="234" specimenCount="7" stateProvince="Chiba" typeStatus="paratype">
<date id="FFC61027FFA9FFBC242DF80AFB485CDC" box="[1075,1215,1976,2003]" pageId="1" pageNumber="234" value="2017-07-26">
<collectingDate id="EF82E9CFFFA9FFBC242DF80AFB485CDC" box="[1075,1215,1976,2003]" pageId="1" pageNumber="234" value="2017-07-26">26 VII 2017</collectingDate>
</date>
(
<collectionCode id="ED69AE22FFA9FFBC24CEF80AFAE05CDC" box="[1232,1303,1976,2002]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="1" pageNumber="234" type="Herbarium">KUM</collectionCode>
); 
<collectorName id="268D5331FFA9FFBC2532F80AFA6E5CDC" box="[1324,1433,1976,2002]" pageId="1" pageNumber="234">Saga-ken</collectorName>
: 
<specimenCount id="9D7EFD6EFFA9FFBC2089F86FFF2C5CF9" box="[151,219,2013,2039]" pageId="1" pageNumber="234" type="generic">7 exs.</specimenCount>
, 
<location id="8EA7603CFFA9FFBC20F6F86EFE5A5CF8" LSID="urn:lsid:plazi:treatment:03D187F1FFA9FFB92089FA5DFAD959A7:8EA7603CFFA9FFBC20F6F86EFE5A5CF8" box="[232,429,2012,2039]" country="Japan" municipality="Arita-cho" name="Hiroseyama - Ko" pageId="1" pageNumber="234" stateProvince="Chiba">Hiroseyama - Kô</location>
, 
<collectingMunicipality id="6BA3AC9DFFA9FFBC21A9F86EFDD25CF8" box="[439,549,2012,2038]" pageId="1" pageNumber="234">Arita-chô</collectingMunicipality>
</materialsCitation>
, 
<date id="FFC61027FFA9FFBC222FF86EFD5D5CF8" box="[561,682,2012,2039]" pageId="1" pageNumber="234" value="2021-05-29">29 V 2021</date>
, T. 
<materialsCitation id="3B103CBAFFA9FFBC22CDF86EFA835CF8" ID-GBIF-Occurrence="3887777387" box="[723,1396,2012,2039]" collectionCode="KUM" collectorName="Hashizume" country="Japan" location="Kurokami-yama" municipality="Takeo-shi" pageId="1" pageNumber="234" specimenCount="2" stateProvince="Chiba" typeStatus="paratype">
<collectorName id="268D5331FFA9FFBC22CDF86EFCA65CF8" box="[723,849,2012,2038]" pageId="1" pageNumber="234">Hashizume</collectorName>
leg. (
<collectionCode id="ED69AE22FFA9FFBC238CF86EFC2E5CF8" box="[914,985,2012,2038]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="1" pageNumber="234" type="Herbarium">KUM</collectionCode>
); 
<specimenCount id="9D7EFD6EFFA9FFBC23F3F86EFBC65CF9" box="[1005,1073,2012,2039]" pageId="1" pageNumber="234" type="generic">2 exs.</specimenCount>
, 
<location id="8EA7603CFFA9FFBC2420F86EFB0E5CF8" LSID="urn:lsid:plazi:treatment:03D187F1FFA9FFB92089FA5DFAD959A7:8EA7603CFFA9FFBC2420F86EFB0E5CF8" box="[1086,1273,2012,2038]" country="Japan" municipality="Takeo-shi" name="Kurokami-yama" pageId="1" pageNumber="234" stateProvince="Chiba">Kurokami-yama</location>
, 
<collectingMunicipality id="6BA3AC9DFFA9FFBC251DF86EFA835CF8" box="[1283,1396,2012,2038]" pageId="1" pageNumber="234">Takeo-shi</collectingMunicipality>
</materialsCitation>
, 
<materialsCitation id="3B103CBAFFA9FFBF259EF86EFC515BBF" ID-GBIF-Occurrence="3887777330" collectingDate="2016-09-21" collectionCode="KUM" collectorName="M. Nishida" country="Japan" lastPageId="2" lastPageNumber="235" location="Kurokami-yama" municipality="Arita-cho" pageId="1" pageNumber="234" specimenCount="1" stateProvince="Chiba" typeStatus="paratype">
<date id="FFC61027FFA9FFBF259EF86EFF0E5BBF" lastPageId="2" lastPageNumber="235" pageId="1" pageNumber="234" value="2016-09-21">
<collectingDate id="EF82E9CFFFA9FFBF259EF86EFF0E5BBF" lastPageId="2" lastPageNumber="235" pageId="1" pageNumber="234" value="2016-09-21">21 IX 2016</collectingDate>
</date>
, 
<collectorName id="268D5331FFAAFFBF2119FF25FE705BBF" box="[263,391,151,177]" pageId="2" pageNumber="235">M. Nishida</collectorName>
leg. (
<collectionCode id="ED69AE22FFAAFFBF21D5FF25FDE55BBF" box="[459,530,151,177]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="2" pageNumber="235" type="Herbarium">KUM</collectionCode>
); 
<specimenCount id="9D7EFD6EFFAAFFBF2236FF25FD945BBF" box="[552,611,151,177]" pageId="2" pageNumber="235" type="generic">1 ex.</specimenCount>
, 
<location id="8EA7603CFFAAFFBF226FFF25FCDB5BBF" LSID="urn:lsid:plazi:treatment:03D187F1FFA9FFB92089FA5DFAD959A7:8EA7603CFFAAFFBF226FFF25FCDB5BBF" box="[625,812,151,177]" country="Japan" municipality="Arita-cho" name="Kurokami-yama" pageId="2" pageNumber="235" stateProvince="Chiba">Kurokami-yama</location>
, 
<collectingMunicipality id="6BA3AC9DFFAAFFBF2326FF25FC515BBF" box="[824,934,151,177]" pageId="2" pageNumber="235">Arita-chô</collectingMunicipality>
</materialsCitation>
, 
<date id="FFC61027FFAAFFBF23ADFF24FBD55BBF" box="[947,1058,150,177]" pageId="2" pageNumber="235" value="2016-05-05">
5 V 
<materialsCitation id="3B103CBAFFAAFFBF23F9FF25FBD55BBF" ID-GBIF-Occurrence="3887777346" box="[999,1058,151,177]" collectionCode="KUM" collectorName="M. Nishida" country="Japan" location="Kurokami-yama" municipality="Arita-cho" pageId="2" pageNumber="235" specimenCount="2" stateProvince="Chiba" typeStatus="paratype">2016</materialsCitation>
</date>
, 
<collectorName id="268D5331FFAAFFBF242EFF25FB475BBF" box="[1072,1200,151,177]" pageId="2" pageNumber="235">M. Nishida</collectorName>
leg. (KUM); 
<specimenCount id="9D7EFD6EFFAAFFBF254FFF25FA6F5BBF" box="[1361,1432,151,177]" pageId="2" pageNumber="235" type="generic">2 exs.</specimenCount>
, 
<location id="8EA7603CFFAAFFBF2089FF09FEA55BDB" LSID="urn:lsid:plazi:treatment:03D187F1FFA9FFB92089FA5DFAD959A7:8EA7603CFFAAFFBF2089FF09FEA55BDB" box="[151,338,187,213]" country="Japan" municipality="Arita-cho" name="Kurokami-yama" pageId="2" pageNumber="235" stateProvince="Chiba">Kurokami-yama</location>
, 
<collectingMunicipality id="6BA3AC9DFFAAFFBF2141FF09FE3A5BDB" box="[351,461,187,213]" pageId="2" pageNumber="235">Arita-chô</collectingMunicipality>
, 
<date id="FFC61027FFAAFFBF21C5FF09FDAD5BDB" box="[475,602,187,213]" pageId="2" pageNumber="235" value="2021-05-29">
29 V 
<materialsCitation id="3B103CBAFFAAFFBF2201FF09FDAD5BDB" ID-GBIF-Occurrence="3887777350" box="[543,602,187,213]" collectionCode="KUM" collectorName="Y. Hisasue" country="Japan" location="Tosen-zan" municipality="Ureshino-shi" pageId="2" pageNumber="235" specimenCount="1" stateProvince="Chiba" typeStatus="paratype">2021</materialsCitation>
</date>
, 
<collectorName id="268D5331FFAAFFBF2276FF09FD145BDB" box="[616,739,187,213]" pageId="2" pageNumber="235">Y. Hisasue</collectorName>
leg. (KUM); 
<specimenCount id="9D7EFD6EFFAAFFBF239FFF09FC4F5BDB" box="[897,952,187,213]" pageId="2" pageNumber="235" type="generic">1 ex.</specimenCount>
, 
<location id="8EA7603CFFAAFFBF23D9FF09FBC85BDB" LSID="urn:lsid:plazi:treatment:03D187F1FFA9FFB92089FA5DFAD959A7:8EA7603CFFAAFFBF23D9FF09FBC85BDB" box="[967,1087,187,213]" country="Japan" municipality="Ureshino-shi" name="Tosen-zan" pageId="2" pageNumber="235" stateProvince="Chiba">Tôsen-zan</location>
, 
<collectingMunicipality id="6BA3AC9DFFAAFFBF2453FF09FB165BDB" box="[1101,1249,187,213]" pageId="2" pageNumber="235">Ureshino-shi</collectingMunicipality>
, 
<date id="FFC61027FFAAFFBF24F1FF09FA995BDB" box="[1263,1390,187,213]" pageId="2" pageNumber="235" value="2016-05-26">
26 V 
<materialsCitation id="3B103CBAFFAAFFBF252DFF09FA995BDB" ID-GBIF-Occurrence="3887777401" box="[1331,1390,187,213]" collectionCode="KUM" collectorName="M. Nishida" country="Japan" location="Tosen-zan" municipality="Ureshino-shi" pageId="2" pageNumber="235" specimenCount="2" stateProvince="Chiba" typeStatus="paratype">2016</materialsCitation>
</date>
, 
<collectorName id="268D5331FFAAFFBF2562FF09FF195BF7" pageId="2" pageNumber="235">M. Nishida</collectorName>
leg. (KUM); 
<specimenCount id="9D7EFD6EFFAAFFBF2190FF6DFE245BF7" box="[398,467,223,249]" pageId="2" pageNumber="235" type="generic">2 exs.</specimenCount>
, same data, but 
<materialsCitation id="3B103CBAFFAAFFBF2290FF6DFAF95BF7" ID-GBIF-Occurrence="3887777321" box="[654,1294,223,249]" collectingDate="2017-07-13" collectionCode="KUM" collectorName="Nagasaki-ken" country="Japan" location="Unzen-dake" pageId="2" pageNumber="235" specimenCount="2" stateProvince="Chiba" typeStatus="paratype">
<date id="FFC61027FFAAFFBF2290FF6DFCED5BF7" box="[654,794,223,249]" pageId="2" pageNumber="235" value="2017-07-13">
<collectingDate id="EF82E9CFFFAAFFBF2290FF6DFCED5BF7" box="[654,794,223,249]" pageId="2" pageNumber="235" value="2017-07-13">13 VII 2017</collectingDate>
</date>
(
<collectionCode id="ED69AE22FFAAFFBF2334FF6DFC865BF7" box="[810,881,223,249]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="2" pageNumber="235" type="Herbarium">KUM</collectionCode>
); 
<collectorName id="268D5331FFAAFFBF2399FF6DFBD25BF7" box="[903,1061,223,249]" pageId="2" pageNumber="235">Nagasaki-ken</collectorName>
: 
<specimenCount id="9D7EFD6EFFAAFFBF242FFF6DFB805BF7" box="[1073,1143,223,249]" pageId="2" pageNumber="235" type="generic">2 exs.</specimenCount>
, 
<location id="8EA7603CFFAAFFBF249AFF6DFAF95BF7" LSID="urn:lsid:plazi:treatment:03D187F1FFA9FFB92089FA5DFAD959A7:8EA7603CFFAAFFBF249AFF6DFAF95BF7" box="[1156,1294,223,249]" country="Japan" name="Unzen-dake" pageId="2" pageNumber="235" stateProvince="Chiba">Unzen-dake</location>
</materialsCitation>
, 
<materialsCitation id="3B103CBAFFAAFFBF2505FF6DFCD45A13" ID-GBIF-Occurrence="3887777312" collectingDate="1978-10-24" collectionCode="KUM" collectorName="S. Imasaka" country="Japan" location="Gokahara-dake" municipality="Tara-dake" pageId="2" pageNumber="235" specimenCount="1" stateProvince="Chiba" typeStatus="paratype">
<date id="FFC61027FFAAFFBF2505FF6DFA6F5BF7" box="[1307,1432,223,249]" pageId="2" pageNumber="235" value="1978-10-24">
<collectingDate id="EF82E9CFFFAAFFBF2505FF6DFA6F5BF7" box="[1307,1432,223,249]" pageId="2" pageNumber="235" value="1978-10-24">24 X 1978</collectingDate>
</date>
, 
<collectorName id="268D5331FFAAFFBF2089FEB1FEE45A13" box="[151,275,259,285]" pageId="2" pageNumber="235">S. Imasaka</collectorName>
leg. (
<collectionCode id="ED69AE22FFAAFFBF214AFEB1FE6C5A13" box="[340,411,259,285]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="2" pageNumber="235" type="Herbarium">KUM</collectionCode>
); 
<specimenCount id="9D7EFD6EFFAAFFBF21AEFEB1FE1D5A13" box="[432,490,259,285]" pageId="2" pageNumber="235" type="generic">1 ex.</specimenCount>
, 
<location id="8EA7603CFFAAFFBF21E8FEB1FD535A13" LSID="urn:lsid:plazi:treatment:03D187F1FFA9FFB92089FA5DFAD959A7:8EA7603CFFAAFFBF21E8FEB1FD535A13" box="[502,676,259,285]" country="Japan" municipality="Tara-dake" name="Gokahara-dake" pageId="2" pageNumber="235" stateProvince="Chiba">Gokahara-dake</location>
, 
<collectingMunicipality id="6BA3AC9DFFAAFFBF22AEFEB1FCD45A13" box="[688,803,259,285]" pageId="2" pageNumber="235">Tara-dake</collectingMunicipality>
</materialsCitation>
, 
<materialsCitation id="3B103CBAFFAAFFBF2330FEB1FE485A4F" ID-GBIF-Occurrence="3887777332" collectingDate="1978-07-11" collectionCode="KUM" collectorName="S. Imasaka" country="Japan" location="Onobarukogen" municipality="Higashisonogi-cho" pageId="2" pageNumber="235" specimenCount="2" stateProvince="Chiba" typeStatus="paratype">
<date id="FFC61027FFAAFFBF2330FEB1FC4C5A13" box="[814,955,259,285]" pageId="2" pageNumber="235" value="1978-07-11">
<collectingDate id="EF82E9CFFFAAFFBF2330FEB1FC4C5A13" box="[814,955,259,285]" pageId="2" pageNumber="235" value="1978-07-11">11 VII 1978</collectingDate>
</date>
, 
<collectorName id="268D5331FFAAFFBF23D9FEB1FBB55A13" box="[967,1090,259,285]" pageId="2" pageNumber="235">S. Imasaka</collectorName>
leg. (
<collectionCode id="ED69AE22FFAAFFBF249AFEB1FB3C5A13" box="[1156,1227,259,285]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="2" pageNumber="235" type="Herbarium">KUM</collectionCode>
); 
<specimenCount id="9D7EFD6EFFAAFFBF24FEFEB1FAD35A13" box="[1248,1316,259,285]" pageId="2" pageNumber="235" type="generic">2 exs.</specimenCount>
, 
<location id="8EA7603CFFAAFFBF252FFEB6FF295A4F" LSID="urn:lsid:plazi:treatment:03D187F1FFA9FFB92089FA5DFAD959A7:8EA7603CFFAAFFBF252FFEB6FF295A4F" country="Japan" municipality="Higashisonogi-cho" name="Onobarukogen" pageId="2" pageNumber="235" stateProvince="Chiba">Ônobarukôgen</location>
, 
<collectingMunicipality id="6BA3AC9DFFAAFFBF20F7FE95FE485A4F" box="[233,447,295,321]" pageId="2" pageNumber="235">Higashisonogi-chô</collectingMunicipality>
</materialsCitation>
, 
<materialsCitation id="3B103CBAFFAAFFBF21D7FE95FBBC5A4F" ID-GBIF-Occurrence="3887777318" box="[457,1099,295,321]" collectingDate="2002-11-17" collectionCode="KUM" collectorName="M. Nishida" country="Japan" location="Onobarukogen" municipality="Higashisonogi-cho" pageId="2" pageNumber="235" specimenCount="1" stateProvince="Chiba" typeStatus="paratype">
<date id="FFC61027FFAAFFBF21D7FE95FDBD5A4F" box="[457,586,295,321]" pageId="2" pageNumber="235" value="2002-11-17">
<collectingDate id="EF82E9CFFFAAFFBF21D7FE95FDBD5A4F" box="[457,586,295,321]" pageId="2" pageNumber="235" value="2002-11-17">17 XI 2002</collectingDate>
</date>
, 
<collectorName id="268D5331FFAAFFBF224AFE95FD255A4F" box="[596,722,295,321]" pageId="2" pageNumber="235">M. Nishida</collectorName>
leg. (
<collectionCode id="ED69AE22FFAAFFBF2311FE95FCA15A4F" box="[783,854,295,321]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="2" pageNumber="235" type="Herbarium">KUM</collectionCode>
); 
<specimenCount id="9D7EFD6EFFAAFFBF2377FE95FC565A4F" box="[873,929,295,321]" pageId="2" pageNumber="235" type="generic">1 ex.</specimenCount>
, same data, but
</materialsCitation>
<materialsCitation id="3B103CBAFFAAFFBF244FFE95FDBC5A6B" ID-GBIF-Occurrence="3887777345" collectingDate="2002-12-07" collectionCode="KUM" collectorName="Oita-ken" country="Japan" county="Hita-shi" location="Shaka-dake" municipality="Maetsue" pageId="2" pageNumber="235" specimenCount="4" stateProvince="Chiba" typeStatus="paratype">
<date id="FFC61027FFAAFFBF244FFE95FB3D5A4F" box="[1105,1226,295,321]" pageId="2" pageNumber="235" value="2002-12-07">
<collectingDate id="EF82E9CFFFAAFFBF244FFE95FB3D5A4F" box="[1105,1226,295,321]" pageId="2" pageNumber="235" value="2002-12-07">7 XII 2002</collectingDate>
</date>
(
<collectionCode id="ED69AE22FFAAFFBF24C7FE95FAD75A4F" box="[1241,1312,295,321]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="2" pageNumber="235" type="Herbarium">KUM</collectionCode>
); 
<collectorName id="268D5331FFAAFFBF252DFE9AFA6E5A4D" box="[1331,1433,296,323]" pageId="2" pageNumber="235">Ôita-ken</collectorName>
: 
<specimenCount id="9D7EFD6EFFAAFFBF2089FEF9FF2A5A6B" box="[151,221,331,357]" pageId="2" pageNumber="235" type="generic">4 exs.</specimenCount>
, 
<location id="8EA7603CFFAAFFBF20F5FEF9FE865A6B" LSID="urn:lsid:plazi:treatment:03D187F1FFA9FFB92089FA5DFAD959A7:8EA7603CFFAAFFBF20F5FEF9FE865A6B" box="[235,369,331,357]" country="Japan" county="Hita-shi" municipality="Maetsue" name="Shaka-dake" pageId="2" pageNumber="235" stateProvince="Chiba">Shaka-dake</location>
, 
<collectingMunicipality id="6BA3AC9DFFAAFFBF2160FEF9FE175A6B" box="[382,480,331,357]" pageId="2" pageNumber="235">Maetsue</collectingMunicipality>
, 
<collectingCounty id="62A64E6BFFAAFFBF21F0FEF9FDBC5A6B" box="[494,587,331,357]" pageId="2" pageNumber="235">Hita-shi</collectingCounty>
</materialsCitation>
, 
<date id="FFC61027FFAAFFBF2246FEF9FD285A6B" box="[600,735,331,357]" pageId="2" pageNumber="235" value="2017-06-23">
23 VI 
<materialsCitation id="3B103CBAFFAAFFBF22BAFEF9FD285A6B" ID-GBIF-Occurrence="3887777406" box="[676,735,331,357]" collectingDate="2002-12-07" collectionCode="KUM" collectorName="S. Imasaka" country="Japan" county="Hita-shi" location="Shaka-dake" municipality="Maetsue" pageId="2" pageNumber="235" specimenCount="2" stateProvince="Chiba" typeStatus="paratype">2017</materialsCitation>
</date>
, 
<collectorName id="268D5331FFAAFFBF22F3FEF9FC9D5A6B" box="[749,874,331,357]" pageId="2" pageNumber="235">S. Imasaka</collectorName>
leg. (KUM); 
<specimenCount id="9D7EFD6EFFAAFFBF2415FEF9FBA55A6B" box="[1035,1106,331,357]" pageId="2" pageNumber="235" type="generic">2 exs.</specimenCount>
, same data, but 
<materialsCitation id="3B103CBAFFAAFFBF2511FEF9FE1B5A87" ID-GBIF-Occurrence="3887777348" collectingDate="2017-07-14" collectionCode="KUM" collectorName="S. Imasaka" country="Japan" county="Hita-shi" location="Shaka-dake" municipality="Maetsue" pageId="2" pageNumber="235" specimenCount="2" stateProvince="Chiba" typeStatus="paratype">
<date id="FFC61027FFAAFFBF2511FEF9FA6B5A6B" box="[1295,1436,331,357]" pageId="2" pageNumber="235" value="2017-07-14">
<collectingDate id="EF82E9CFFFAAFFBF2511FEF9FA6B5A6B" box="[1295,1436,331,357]" pageId="2" pageNumber="235" value="2017-07-14">14 VII 2017</collectingDate>
</date>
(
<collectionCode id="ED69AE22FFAAFFBF2081FEDDFF115A87" box="[159,230,367,393]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="2" pageNumber="235" type="Herbarium">KUM</collectionCode>
); 
<specimenCount id="9D7EFD6EFFAAFFBF20E4FEDDFEC95A87" box="[250,318,367,393]" pageId="2" pageNumber="235" type="generic">2 exs.</specimenCount>
, same data, but
</materialsCitation>
<date id="FFC61027FFAAFFBF21EDFEDDFD805A87" box="[499,631,367,393]" pageId="2" pageNumber="235" value="2017-08-02">
2 VIII 
<materialsCitation id="3B103CBAFFAAFFBF2221FEDDFD805A87" ID-GBIF-Occurrence="3887777354" box="[575,631,367,393]" collectingDate="2017-07-14" collectionCode="KUM" collectorName="S. Imasaka" country="Japan" county="Hita-shi" location="Shaka-dake" municipality="Maetsue" pageId="2" pageNumber="235" specimenCount="3" stateProvince="Chiba" typeStatus="paratype">2017</materialsCitation>
</date>
(KUM); 
<specimenCount id="9D7EFD6EFFAAFFBF22FFFEDDFCD25A87" box="[737,805,367,393]" pageId="2" pageNumber="235" type="generic">3 exs.</specimenCount>
, same data, but 
<date id="FFC61027FFAAFFBF23C4FEDDFB9B5A87" box="[986,1132,367,393]" pageId="2" pageNumber="235" value="2017-08-23">
23 VIII 
<materialsCitation id="3B103CBAFFAAFFBF242AFEDDFB9B5A87" ID-GBIF-Occurrence="3887777364" box="[1076,1132,367,393]" collectingDate="2017-07-14" collectionCode="KUM" collectorName="S. Imasaka" country="Japan" county="Hita-shi" location="Shaka-dake" municipality="Maetsue" pageId="2" pageNumber="235" specimenCount="2" stateProvince="Chiba" typeStatus="paratype">2017</materialsCitation>
</date>
(KUM); 
<specimenCount id="9D7EFD6EFFAAFFBF24C8FEDDFAED5A87" box="[1238,1306,367,393]" pageId="2" pageNumber="235" type="generic">2 exs.</specimenCount>
, same data, but 
<date id="FFC61027FFAAFFBF20DAFE21FEB15AA3" box="[196,326,403,429]" pageId="2" pageNumber="235" value="2018-06-13">
13 VI 
<materialsCitation id="3B103CBAFFAAFFBF2110FE21FEB15AA3" ID-GBIF-Occurrence="3887777383" box="[270,326,403,429]" collectingDate="2017-07-14" collectionCode="KUM" collectorName="S. Imasaka" country="Japan" county="Hita-shi" location="Shaka-dake" municipality="Maetsue" pageId="2" pageNumber="235" specimenCount="1" stateProvince="Chiba" typeStatus="paratype">2018</materialsCitation>
</date>
(KUM); 
<specimenCount id="9D7EFD6EFFAAFFBF21ADFE21FE1A5AA3" box="[435,493,403,429]" pageId="2" pageNumber="235" type="generic">1 ex.</specimenCount>
, same data, but 
<materialsCitation id="3B103CBAFFAAFFBF22B6FE21FB7A5AA3" ID-GBIF-Occurrence="3887777363" box="[680,1165,403,429]" collectingDate="2018-07-10" collectionCode="KUM" collectorName="S. Imasaka" country="Japan" county="Hita-shi" location="Shaka-dake" municipality="Maetsue" pageId="2" pageNumber="235" specimenCount="1" stateProvince="Chiba" typeStatus="paratype">
<date id="FFC61027FFAAFFBF22B6FE21FCC35AA3" box="[680,820,403,429]" pageId="2" pageNumber="235" value="2018-07-10">
<collectingDate id="EF82E9CFFFAAFFBF22B6FE21FCC35AA3" box="[680,820,403,429]" pageId="2" pageNumber="235" value="2018-07-10">10 VII 2018</collectingDate>
</date>
(
<collectionCode id="ED69AE22FFAAFFBF235AFE21FC7C5AA3" box="[836,907,403,429]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="2" pageNumber="235" type="Herbarium">KUM</collectionCode>
); 
<specimenCount id="9D7EFD6EFFAAFFBF23BFFE21FC2C5AA3" box="[929,987,403,429]" pageId="2" pageNumber="235" type="generic">1 ex.</specimenCount>
, same data, but
</materialsCitation>
<materialsCitation id="3B103CBAFFAAFFBF248BFE21FE915ADF" ID-GBIF-Occurrence="3887777309" collectingDate="2018-07-21" collectionCode="KUM" collectorName="S. Imasaka" country="Japan" county="Hita-shi" location="Shaka-dake" municipality="Maetsue" pageId="2" pageNumber="235" specimenCount="1" stateProvince="Chiba" typeStatus="paratype">
<date id="FFC61027FFAAFFBF248BFE21FAD65AA3" box="[1173,1313,403,429]" pageId="2" pageNumber="235" value="2018-07-21">
<collectingDate id="EF82E9CFFFAAFFBF248BFE21FAD65AA3" box="[1173,1313,403,429]" pageId="2" pageNumber="235" value="2018-07-21">21 VII 2018</collectingDate>
</date>
(
<collectionCode id="ED69AE22FFAAFFBF252CFE21FA8E5AA3" box="[1330,1401,403,429]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="2" pageNumber="235" type="Herbarium">KUM</collectionCode>
); 
<specimenCount id="9D7EFD6EFFAAFFBF2590FE21FF4C5ADF" pageId="2" pageNumber="235" type="generic">1 ex.</specimenCount>
, same data, but
</materialsCitation>
<date id="FFC61027FFAAFFBF2172FE05FE0B5ADF" box="[364,508,439,465]" pageId="2" pageNumber="235" value="2018-08-11">
11 VIII 
<materialsCitation id="3B103CBAFFAAFFBF21DAFE05FE0B5ADF" ID-GBIF-Occurrence="3887777367" box="[452,508,439,465]" collectionCode="KUM" country="Japan" county="Beppu-shi" location="Zenigame-toge" municipality="Takasaki-yama" pageId="2" pageNumber="235" specimenCount="1" stateProvince="Chiba" typeStatus="paratype">2018</materialsCitation>
</date>
(KUM); 
<specimenCount id="9D7EFD6EFFAAFFBF227BFE05FD6A5ADF" box="[613,669,439,465]" pageId="2" pageNumber="235" type="generic">1 ex.</specimenCount>
, 
<location id="8EA7603CFFAAFFBF22B6FE05FCA35ADF" LSID="urn:lsid:plazi:treatment:03D187F1FFA9FFB92089FA5DFAD959A7:8EA7603CFFAAFFBF22B6FE05FCA35ADF" box="[680,852,439,465]" country="Japan" county="Beppu-shi" municipality="Takasaki-yama" name="Zenigame-toge" pageId="2" pageNumber="235" stateProvince="Chiba">Zenigame-tôge</location>
, 
<collectingMunicipality id="6BA3AC9DFFAAFFBF2341FE05FBFC5ADF" box="[863,1035,439,465]" pageId="2" pageNumber="235">Takasaki-yama</collectingMunicipality>
, 
<collectingCounty id="62A64E6BFFAAFFBF240BFE05FB7C5ADF" box="[1045,1163,439,465]" pageId="2" pageNumber="235">Beppu-shi</collectingCounty>
, 
<materialsCitation id="3B103CBAFFAAFFBF2488FE05FACB5ADF" ID-GBIF-Occurrence="3887777335" box="[1174,1340,438,465]" collectingDate="2015-07-26" collectionCode="T" country="Japan" pageId="2" pageNumber="235" specimenCount="1" stateProvince="Chiba" typeStatus="paratype">
<date id="FFC61027FFAAFFBF2488FE05FAD75ADF" box="[1174,1312,438,465]" pageId="2" pageNumber="235" value="2015-07-26">
<collectingDate id="EF82E9CFFFAAFFBF2488FE05FAD75ADF" box="[1174,1312,438,465]" pageId="2" pageNumber="235" value="2015-07-26">26 VII 2015</collectingDate>
</date>
, 
<collectionCode id="ED69AE22FFAAFFBF2534FE05FACB5ADF" box="[1322,1340,439,465]" name="Tavera, Department of Geology and Geophysics" pageId="2" pageNumber="235">T</collectionCode>
</materialsCitation>
. 
<materialsCitation id="3B103CBAFFAAFFBF2559FE05FBB05AFB" ID-GBIF-Occurrence="3887777380" collectionCode="KUM" collectorName="Miyake &amp; Miyazaki-ken" country="Japan" county="Miyakonojo-shi" elevation="385" location="Miike" municipality="Natsuo-cho" pageId="2" pageNumber="235" specimenCount="6" stateProvince="Chiba" typeStatus="paratype">
<collectorName id="268D5331FFAAFFBF2559FE05FA6A5ADF" box="[1351,1437,439,465]" pageId="2" pageNumber="235">Miyake</collectorName>
leg. (
<collectionCode id="ED69AE22FFAAFFBF20CFFE69FEEF5AFB" box="[209,280,475,501]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="2" pageNumber="235" type="Herbarium">KUM</collectionCode>
); 
<collectorName id="268D5331FFAAFFBF2133FE69FE3A5AFB" box="[301,461,475,501]" pageId="2" pageNumber="235">Miyazaki-ken</collectorName>
: 
<specimenCount id="9D7EFD6EFFAAFFBF21C7FE69FDE95AFB" box="[473,542,475,501]" pageId="2" pageNumber="235" type="generic">6 exs.</specimenCount>
, alt. 
<quantity id="4C809B02FFAAFFBF2248FE68FD565AFB" box="[598,673,474,501]" metricMagnitude="2" metricUnit="m" metricValue="3.85" pageId="2" pageNumber="235" unit="m" value="385.0">
<elevation id="0055D1D4FFAAFFBF2248FE68FD565AFB" box="[598,673,474,501]" metricMagnitude="2" metricUnit="m" metricValue="3.85" pageId="2" pageNumber="235" unit="m" value="385.0">385 m</elevation>
</quantity>
, 
<location id="8EA7603CFFAAFFBF22B0FE69FD035AFB" LSID="urn:lsid:plazi:treatment:03D187F1FFA9FFB92089FA5DFAD959A7:8EA7603CFFAAFFBF22B0FE69FD035AFB" box="[686,756,475,501]" country="Japan" county="Miyakonojo-shi" municipality="Natsuo-cho" name="Miike" pageId="2" pageNumber="235" stateProvince="Chiba">Miike</location>
, 
<collectingMunicipality id="6BA3AC9DFFAAFFBF231EFE69FC735AFB" box="[768,900,475,501]" pageId="2" pageNumber="235">Natsuo-chô</collectingMunicipality>
, 
<collectingCounty id="62A64E6BFFAAFFBF238FFE69FBB05AFB" box="[913,1095,475,501]" pageId="2" pageNumber="235">Miyakonojô-shi</collectingCounty>
</materialsCitation>
, 
<date id="FFC61027FFAAFFBF244DFE69FB1D5AFB" box="[1107,1258,475,501]" pageId="2" pageNumber="235" value="2019-08-17">
17 VIII 
<materialsCitation id="3B103CBAFFAAFFBF24B1FE69FB1D5AFB" ID-GBIF-Occurrence="3887777344" box="[1199,1258,475,501]" collectionCode="KUM" collectorName="F. Nakano" country="Japan" location="Gokasho" municipality="Takachiho-cho" pageId="2" pageNumber="235" specimenCount="1" stateProvince="Chiba" typeStatus="paratype">2019</materialsCitation>
</date>
, 
<collectorName id="268D5331FFAAFFBF24E9FE69FA9D5AFB" box="[1271,1386,475,501]" pageId="2" pageNumber="235">F. Nakano</collectorName>
leg. (KUM); 
<specimenCount id="9D7EFD6EFFAAFFBF20E5FE4DFEC25917" box="[251,309,511,537]" pageId="2" pageNumber="235" type="generic">1 ex.</specimenCount>
, 
<location id="8EA7603CFFAAFFBF215CFE4DFE5F5917" LSID="urn:lsid:plazi:treatment:03D187F1FFA9FFB92089FA5DFAD959A7:8EA7603CFFAAFFBF215CFE4DFE5F5917" box="[322,424,511,537]" country="Japan" municipality="Takachiho-cho" name="Gokasho" pageId="2" pageNumber="235" stateProvince="Chiba">Gokasho</location>
, 
<collectingMunicipality id="6BA3AC9DFFAAFFBF21ABFE4DFDA95917" box="[437,606,511,537]" pageId="2" pageNumber="235">Takachiho-chô</collectingMunicipality>
, 
<materialsCitation id="3B103CBAFFAAFFBF2274FE4CFCE15917" ID-GBIF-Occurrence="3887777326" box="[618,790,510,537]" collectingDate="2020-07-25" collectionCode="T" country="Japan" pageId="2" pageNumber="235" specimenCount="1" stateProvince="Chiba" typeStatus="paratype">
<date id="FFC61027FFAAFFBF2274FE4CFD0F5917" box="[618,760,510,537]" pageId="2" pageNumber="235" value="2020-07-25">
<collectingDate id="EF82E9CFFFAAFFBF2274FE4CFD0F5917" box="[618,760,510,537]" pageId="2" pageNumber="235" value="2020-07-25">25 VII 2020</collectingDate>
</date>
, 
<collectionCode id="ED69AE22FFAAFFBF231AFE4DFCE15917" box="[772,790,511,537]" name="Tavera, Department of Geology and Geophysics" pageId="2" pageNumber="235">T</collectionCode>
</materialsCitation>
. 
<materialsCitation id="3B103CBAFFAAFFBF233CFE4DFD625933" ID-GBIF-Occurrence="3887777415" collectionCode="KUM" collectorName="Hashizume &amp; Kagoshima-ken" country="Japan" county="Kirishima-shi" location="Maruo Scenic Nature Path" municipality="Makizono" pageId="2" pageNumber="235" specimenCount="2" stateProvince="Chiba" typeStatus="paratype">
<collectorName id="268D5331FFAAFFBF233CFE4DFC575917" box="[802,928,511,537]" pageId="2" pageNumber="235">Hashizume</collectorName>
leg. (
<collectionCode id="ED69AE22FFAAFFBF23FCFE4DFBDE5917" box="[994,1065,511,537]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="2" pageNumber="235" type="Herbarium">KUM</collectionCode>
); 
<collectorName id="268D5331FFAAFFBF2420FE4DFB035917" box="[1086,1268,511,537]" pageId="2" pageNumber="235">Kagoshima-ken</collectorName>
: 
<specimenCount id="9D7EFD6EFFAAFFBF251EFE4DFAB25917" box="[1280,1349,511,537]" pageId="2" pageNumber="235" type="generic">2 exs.</specimenCount>
, 
<location id="8EA7603CFFAAFFBF254CFE4DFE985933" LSID="urn:lsid:plazi:treatment:03D187F1FFA9FFB92089FA5DFAD959A7:8EA7603CFFAAFFBF254CFE4DFE985933" country="Japan" county="Kirishima-shi" municipality="Makizono" name="Maruo Scenic Nature Path" pageId="2" pageNumber="235" stateProvince="Chiba">Maruo Scenic Nature Path</location>
, 
<collectingMunicipality id="6BA3AC9DFFAAFFBF2167FD91FE1A5933" box="[377,493,547,573]" pageId="2" pageNumber="235">Makizono</collectingMunicipality>
, 
<collectingCounty id="62A64E6BFFAAFFBF21E6FD91FD625933" box="[504,661,547,573]" pageId="2" pageNumber="235">Kirishima-shi</collectingCounty>
</materialsCitation>
, 
<materialsCitation id="3B103CBAFFAAFFBF2281FD91FC3D5933" ID-GBIF-Occurrence="3887777371" box="[671,970,547,573]" collectingDate="2021-10-09" collectingDateMax="2021-10-10" collectingDateMin="2021-10-09" collectionCode="T" country="Japan" location="Tsuji" pageId="2" pageNumber="235" specimenCount="1" stateProvince="Chiba" typeStatus="paratype">
<date id="FFC61027FFAAFFBF2281FD91FCC45933" box="[671,819,547,573]" pageId="2" pageNumber="235" value="2021-10-09" valueMax="2021-10-10" valueMin="2021-10-09">
<collectingDate id="EF82E9CFFFAAFFBF2281FD91FCC45933" box="[671,819,547,573]" pageId="2" pageNumber="235" value="2021-10-09" valueMax="2021-10-10" valueMin="2021-10-09">9–10 X 2021</collectingDate>
</date>
, N. 
<location id="8EA7603CFFAAFFBF2340FD91FC615933" LSID="urn:lsid:plazi:treatment:03D187F1FFA9FFB92089FA5DFAD959A7:8EA7603CFFAAFFBF2340FD91FC615933" box="[862,918,547,573]" country="Japan" name="Tsuji" pageId="2" pageNumber="235" stateProvince="Chiba">Tsuji</location>
&amp; 
<collectionCode id="ED69AE22FFAAFFBF23A6FD91FC3D5933" box="[952,970,547,573]" name="Tavera, Department of Geology and Geophysics" pageId="2" pageNumber="235">T</collectionCode>
</materialsCitation>
. 
<materialsCitation id="3B103CBAFFAAFFBF23CAFD91FE9E596F" ID-GBIF-Occurrence="3887777303" collectionCode="KUM" collectorName="Nozaki" country="Japan" county="Minamiosumi-cho" location="Izashiki" municipality="Sata" pageId="2" pageNumber="235" specimenCount="1" stateProvince="Chiba" typeStatus="paratype">
<collectorName id="268D5331FFAAFFBF23CAFD91FBD25933" box="[980,1061,547,573]" pageId="2" pageNumber="235">Nozaki</collectorName>
leg. (
<collectionCode id="ED69AE22FFAAFFBF247CFD91FB5E5933" box="[1122,1193,547,573]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="2" pageNumber="235" type="Herbarium">KUM</collectionCode>
); 
<specimenCount id="9D7EFD6EFFAAFFBF24A2FD91FB035933" box="[1212,1268,547,573]" pageId="2" pageNumber="235" type="generic">1 ex.</specimenCount>
, 
<location id="8EA7603CFFAAFFBF24E1FD91FAAC5933" LSID="urn:lsid:plazi:treatment:03D187F1FFA9FFB92089FA5DFAD959A7:8EA7603CFFAAFFBF24E1FD91FAAC5933" box="[1279,1371,547,573]" country="Japan" county="Minamiosumi-cho" municipality="Sata" name="Izashiki" pageId="2" pageNumber="235" stateProvince="Chiba">Izashiki</location>
, 
<collectingMunicipality id="6BA3AC9DFFAAFFBF257BFD91FA6F5933" box="[1381,1432,547,573]" pageId="2" pageNumber="235">Sata</collectingMunicipality>
, 
<collectingCounty id="62A64E6BFFAAFFBF2089FDF5FE9E596F" box="[151,361,583,609]" pageId="2" pageNumber="235">Minamiôsumi-chô</collectingCounty>
</materialsCitation>
, 
<date id="FFC61027FFAAFFBF2168FDF5FE0C596F" box="[374,507,583,609]" pageId="2" pageNumber="235" value="2021-06-22">
22 VI 
<materialsCitation id="3B103CBAFFAAFFBF21DEFDF5FE0C596F" ID-GBIF-Occurrence="3887777372" box="[448,507,583,609]" collectionCode="KUM" collectorName="S. Inoue" country="Japan" county="Satsumasendai-shi" location="Teuchi" municipality="Shimokoshiki-shima" pageId="2" pageNumber="235" specimenCount="1" stateProvince="Chiba" typeStatus="paratype">2021</materialsCitation>
</date>
, 
<collectorName id="268D5331FFAAFFBF2216FDF5FD90596F" box="[520,615,583,609]" pageId="2" pageNumber="235">S. Inoue</collectorName>
leg. (KUM); 
<specimenCount id="9D7EFD6EFFAAFFBF2318FDF5FCB7596F" box="[774,832,583,609]" pageId="2" pageNumber="235" type="generic">1 ex.</specimenCount>
, 
<location id="8EA7603CFFAAFFBF2353FDF5FC6B596F" LSID="urn:lsid:plazi:treatment:03D187F1FFA9FFB92089FA5DFAD959A7:8EA7603CFFAAFFBF2353FDF5FC6B596F" box="[845,924,583,609]" country="Japan" county="Satsumasendai-shi" municipality="Shimokoshiki-shima" name="Teuchi" pageId="2" pageNumber="235" stateProvince="Chiba">Teuchi</location>
, 
<collectingMunicipality id="6BA3AC9DFFAAFFBF23B7FDF5FB63596F" box="[937,1172,583,609]" pageId="2" pageNumber="235">Shimokoshiki-shima</collectingMunicipality>
, 
<collectingCounty id="62A64E6BFFAAFFBF24BEFDF5FA84596F" box="[1184,1395,583,609]" pageId="2" pageNumber="235">Satsumasendai-shi</collectingCounty>
, 
<materialsCitation id="3B103CBAFFAAFFBF259EFDF5FED6598B" ID-GBIF-Occurrence="3887777338" collectingDate="2020-07-22" collectionCode="R" country="Japan" pageId="2" pageNumber="235" specimenCount="1" stateProvince="Chiba" typeStatus="paratype">
<date id="FFC61027FFAAFFBF259EFDF5FEF7598B" pageId="2" pageNumber="235" value="2020-07-22">
<collectingDate id="EF82E9CFFFAAFFBF259EFDF5FEF7598B" pageId="2" pageNumber="235" value="2020-07-22">22 VII 2020</collectingDate>
</date>
, 
<collectionCode id="ED69AE22FFAAFFBF2115FDD9FED6598B" box="[267,289,619,645]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="235">R</collectionCode>
</materialsCitation>
. 
<materialsCitation id="3B103CBAFFAAFFBF2132FDD9FC05598B" ID-GBIF-Occurrence="3887777368" box="[300,1010,619,645]" collectionCode="KUM" collectorName="Ito" country="Japan" location="Shimokoshiki-shima" municipality="Satsumasendai-shi" pageId="2" pageNumber="235" specimenCount="1" stateProvince="Chiba" typeStatus="paratype">
<collectorName id="268D5331FFAAFFBF2132FDD9FEBC598B" box="[300,331,619,645]" pageId="2" pageNumber="235">Ito</collectorName>
leg. (
<collectionCode id="ED69AE22FFAAFFBF2194FDD9FE26598B" box="[394,465,619,645]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="2" pageNumber="235" type="Herbarium">KUM</collectionCode>
); 
<specimenCount id="9D7EFD6EFFAAFFBF21FBFDD9FDEA598B" box="[485,541,619,645]" pageId="2" pageNumber="235" type="generic">1 ex.</specimenCount>
, 
<location id="8EA7603CFFAAFFBF2237FDD9FCE3598B" LSID="urn:lsid:plazi:treatment:03D187F1FFA9FFB92089FA5DFAD959A7:8EA7603CFFAAFFBF2237FDD9FCE3598B" box="[553,788,619,645]" country="Japan" municipality="Satsumasendai-shi" name="Shimokoshiki-shima" pageId="2" pageNumber="235" stateProvince="Chiba">Shimokoshiki-shima</location>
, 
<collectingMunicipality id="6BA3AC9DFFAAFFBF2301FDD9FC05598B" box="[799,1010,619,645]" pageId="2" pageNumber="235">Satsumasendai-shi</collectingMunicipality>
</materialsCitation>
, 
<materialsCitation id="3B103CBAFFAAFFBF23E3FDD8FB5D598B" ID-GBIF-Occurrence="3887777324" box="[1021,1194,618,645]" collectingDate="2020-07-25" collectionCode="R" country="Japan" pageId="2" pageNumber="235" specimenCount="1" stateProvince="Chiba" typeStatus="paratype">
<date id="FFC61027FFAAFFBF23E3FDD8FB7E598B" box="[1021,1161,618,645]" pageId="2" pageNumber="235" value="2020-07-25">
<collectingDate id="EF82E9CFFFAAFFBF23E3FDD8FB7E598B" box="[1021,1161,618,645]" pageId="2" pageNumber="235" value="2020-07-25">25 VII 2020</collectingDate>
</date>
, 
<collectionCode id="ED69AE22FFAAFFBF248AFDD9FB5D598B" box="[1172,1194,619,645]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="235">R</collectionCode>
</materialsCitation>
. 
<collectorName id="268D5331FFAAFFBF24AAFDD9FB24598B" box="[1204,1235,619,645]" pageId="2" pageNumber="235">Ito</collectorName>
leg. (KUM).
</paragraph>
</subSubSection>
<subSubSection id="C362656CFFAAFFBF20D9FD3CFE3B58C7" pageId="2" pageNumber="235" type="diagnosis">
<paragraph id="8BC736E7FFAAFFBF20D9FD3CFE3B58C7" blockId="2.[151,1437,150,2040]" pageId="2" pageNumber="235">
<emphasis id="B90CEAF5FFAAFFBF20D9FD3CFEB559A6" bold="true" box="[199,322,654,680]" pageId="2" pageNumber="235">Diagnosis.</emphasis>
This species is similar to 
<taxonomicName id="4C784D64FFAAFFBF2297FD3DFC5459A7" authority="Lobl, 1977" authorityName="Lobl" authorityYear="1977" box="[649,931,655,681]" class="Insecta" family="Staphylinidae" genus="Dasycerus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="235" phylum="Arthropoda" rank="species" species="fasciatus">
<emphasis id="B90CEAF5FFAAFFBF2297FD3DFCE259A7" box="[649,789,655,681]" italics="true" pageId="2" pageNumber="235">D. fasciatus</emphasis>
<bibRefCitation id="EFE94B16FFAAFFBF233CFD3DFC5459A7" author="Lobl" box="[802,931,655,681]" firstAuthor="Lobl" pageId="2" pageNumber="235" pagination="95 - 106" refId="ref9175" refString="Lobl, I. (1977) Beitrag zur Kenntnis der Gattung Dasycerus Brongniart (Coleoptera, Dasyceridae). Mitteilungen der Schweizerischen Entomologischen Gesellschaft = Bulletin de la Societe Entomologique Suisse, 50, 95 - 106." type="journal article" year="1977">Löbl, 1977</bibRefCitation>
</taxonomicName>
from 
<collectingCountry id="F36F7677FFAAFFBF23EDFD3DFBC759A7" box="[1011,1072,655,681]" name="India" pageId="2" pageNumber="235">India</collectingCountry>
. However, this species can be distinguished from 
<taxonomicName id="4C784D64FFAAFFBF2171FD01FE0259C3" authorityName="Lobl" authorityYear="1977" box="[367,501,691,717]" class="Insecta" family="Staphylinidae" genus="Dasycerus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="235" phylum="Arthropoda" rank="species" species="fasciatus">
<emphasis id="B90CEAF5FFAAFFBF2171FD01FE0259C3" box="[367,501,691,717]" italics="true" pageId="2" pageNumber="235">D. fasciatus</emphasis>
</taxonomicName>
by its larger body, and by the shape of the median lobe of the aedeagus. The number of rows of punctures between first and second internal discal costae is also different (
<taxonomicName id="4C784D64FFAAFFBF2466FD65FB0D59FF" authority="Hashizume &amp; Maruyama, 2022" authorityName="Hashizume &amp; Maruyama" authorityYear="2022" box="[1144,1274,727,753]" class="Insecta" family="Staphylinidae" genus="Dasycerus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="235" phylum="Arthropoda" rank="species" species="occultus" status="sp. nov.">
<emphasis id="B90CEAF5FFAAFFBF2466FD65FB7B59FE" box="[1144,1164,727,752]" italics="true" pageId="2" pageNumber="235">D</emphasis>
. 
<emphasis id="B90CEAF5FFAAFFBF2483FD65FB0D59FF" box="[1181,1274,727,753]" italics="true" pageId="2" pageNumber="235">occultus</emphasis>
</taxonomicName>
<emphasis id="B90CEAF5FFAAFFBF251AFD65FA9659FF" bold="true" box="[1284,1377,727,753]" pageId="2" pageNumber="235">
<taxonomicNameLabel id="A23F578EFFAAFFBF251AFD65FA9659FF" box="[1284,1377,727,753]" pageId="2" pageNumber="235" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
with two rows of punctures; 
<taxonomicName id="4C784D64FFAAFFBF21B9FD49FDD8581B" authorityName="Lobl" authorityYear="1977" box="[423,559,763,789]" class="Insecta" family="Staphylinidae" genus="Dasycerus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="235" phylum="Arthropoda" rank="species" species="fasciatus">
<emphasis id="B90CEAF5FFAAFFBF21B9FD49FDD8581B" box="[423,559,763,789]" italics="true" pageId="2" pageNumber="235">D. fasciatus</emphasis>
</taxonomicName>
with two rows of punctures and additional short row). In contrast to another Japanese species, 
<taxonomicName id="4C784D64FFAAFFBF2179FCADFE0E5836" authorityName=": Lobl &amp; Calame" authorityYear="1996" box="[359,505,799,825]" class="Insecta" family="Staphylinidae" genus="Dasycerus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="235" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B90CEAF5FFAAFFBF2179FCADFE8C5836" box="[359,379,799,824]" italics="true" pageId="2" pageNumber="235">D</emphasis>
. 
<emphasis id="B90CEAF5FFAAFFBF2192FCADFE0E5836" box="[396,505,799,824]" italics="true" pageId="2" pageNumber="235">japonicus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C784D64FFAAFFBF2214FCADFD7B5837" authority="Hashizume &amp; Maruyama, 2022" authorityName="Hashizume &amp; Maruyama" authorityYear="2022" box="[522,652,799,825]" class="Insecta" family="Staphylinidae" genus="Dasycerus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="235" phylum="Arthropoda" rank="species" species="occultus" status="sp. nov.">
<emphasis id="B90CEAF5FFAAFFBF2214FCADFD7B5837" box="[522,652,799,825]" italics="true" pageId="2" pageNumber="235">D. occultus</emphasis>
</taxonomicName>
<emphasis id="B90CEAF5FFAAFFBF2288FCADFD045837" bold="true" box="[662,755,799,825]" pageId="2" pageNumber="235">
<taxonomicNameLabel id="A23F578EFFAAFFBF2288FCADFD045837" box="[662,755,799,825]" pageId="2" pageNumber="235" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has two pairs of dark brown spots on their elytra, and the female genital chamber of 
<taxonomicName id="4C784D64FFAAFFBF21D8FCF1FDB25853" authority="Hashizume &amp; Maruyama, 2022" authorityName="Hashizume &amp; Maruyama" authorityYear="2022" box="[454,581,835,861]" class="Insecta" family="Staphylinidae" genus="Dasycerus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="235" phylum="Arthropoda" rank="species" species="occultus" status="sp. nov.">
<emphasis id="B90CEAF5FFAAFFBF21D8FCF1FDB25853" box="[454,581,835,861]" italics="true" pageId="2" pageNumber="235">D. occultus</emphasis>
</taxonomicName>
<emphasis id="B90CEAF5FFAAFFBF2253FCF1FD505853" bold="true" box="[589,679,835,861]" pageId="2" pageNumber="235">
<taxonomicNameLabel id="A23F578EFFAAFFBF2253FCF1FD505853" box="[589,679,835,861]" pageId="2" pageNumber="235" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="13432A62FFAAFFBF22A9FCF1FCF95853" box="[695,782,835,861]" captionStart="FIGURE 1" captionStartId="3.[151,250,1855,1879]" captionTargetBox="[151,1435,298,1813]" captionTargetId="figure-59@3.[151,1436,270,1829]" captionTargetPageId="3" captionText="FIGURE 1. Dasycerus occultus sp. nov. (A) Habitus in dorsal view. (B) SEM image of pronotum in dorsal view. (C–D) Prothorax in ventral view (black triangles indicate the protergosternal keel). (E–F) Right elytron in dorsolateral view. (G) Aedeagus in lateral view. (H) Female genital chamber in ventral view. Scale bars: 1.0 mm for A; 0.5 mm for C &amp; E; 0.2 mm for G–H." figureDoi="http://doi.org/10.5281/zenodo.6986249" httpUri="https://zenodo.org/record/6986249/files/figure.png" pageId="2" pageNumber="235">Fig. 1H</figureCitation>
) is weakly sclerotized and has a number of fine spines on its surface. The median groove of the pronotum is relatively long (
<taxonomicName id="4C784D64FFAAFFBF239CFCD5FBF5588F" authority="Hashizume &amp; Maruyama, 2022" authorityName="Hashizume &amp; Maruyama" authorityYear="2022" box="[898,1026,871,897]" class="Insecta" family="Staphylinidae" genus="Dasycerus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="235" phylum="Arthropoda" rank="species" species="occultus" status="sp. nov.">
<emphasis id="B90CEAF5FFAAFFBF239CFCD5FBF5588F" box="[898,1026,871,897]" italics="true" pageId="2" pageNumber="235">D. occultus</emphasis>
</taxonomicName>
<emphasis id="B90CEAF5FFAAFFBF2417FCD5FB94588F" bold="true" box="[1033,1123,871,897]" pageId="2" pageNumber="235">
<taxonomicNameLabel id="A23F578EFFAAFFBF2417FCD5FB94588F" box="[1033,1123,871,897]" pageId="2" pageNumber="235" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: 0.52–0.54 times as long as pronotum; 
<taxonomicName id="4C784D64FFAAFFBF210BFC39FE5158AA" authorityName=": Lobl &amp; Calame" authorityYear="1996" box="[277,422,907,932]" class="Insecta" family="Staphylinidae" genus="Dasycerus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="235" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B90CEAF5FFAAFFBF210BFC39FE5158AA" box="[277,422,907,932]" italics="true" pageId="2" pageNumber="235">D. japonicus</emphasis>
</taxonomicName>
: 0.42–0.43 times as long as pronotum) (
<figureCitation id="13432A62FFAAFFBF2368FC39FC3858AB" box="[886,975,907,933]" captionStart="FIGURE 1" captionStartId="3.[151,250,1855,1879]" captionTargetBox="[151,1435,298,1813]" captionTargetId="figure-59@3.[151,1436,270,1829]" captionTargetPageId="3" captionText="FIGURE 1. Dasycerus occultus sp. nov. (A) Habitus in dorsal view. (B) SEM image of pronotum in dorsal view. (C–D) Prothorax in ventral view (black triangles indicate the protergosternal keel). (E–F) Right elytron in dorsolateral view. (G) Aedeagus in lateral view. (H) Female genital chamber in ventral view. Scale bars: 1.0 mm for A; 0.5 mm for C &amp; E; 0.2 mm for G–H." figureDoi="http://doi.org/10.5281/zenodo.6986249" httpUri="https://zenodo.org/record/6986249/files/figure.png" pageId="2" pageNumber="235">Fig. 1B</figureCitation>
). The punctures of elytron are relatively large and circular (
<figureCitation id="13432A62FFAAFFBF2174FC1DFE4958C7" box="[362,446,943,969]" captionStart="FIGURE 1" captionStartId="3.[151,250,1855,1879]" captionTargetBox="[151,1435,298,1813]" captionTargetId="figure-59@3.[151,1436,270,1829]" captionTargetPageId="3" captionText="FIGURE 1. Dasycerus occultus sp. nov. (A) Habitus in dorsal view. (B) SEM image of pronotum in dorsal view. (C–D) Prothorax in ventral view (black triangles indicate the protergosternal keel). (E–F) Right elytron in dorsolateral view. (G) Aedeagus in lateral view. (H) Female genital chamber in ventral view. Scale bars: 1.0 mm for A; 0.5 mm for C &amp; E; 0.2 mm for G–H." figureDoi="http://doi.org/10.5281/zenodo.6986249" httpUri="https://zenodo.org/record/6986249/files/figure.png" pageId="2" pageNumber="235">Fig. 1F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C362656CFFAAFFB920D9FC60FEF85AA3" lastPageId="4" lastPageNumber="237" pageId="2" pageNumber="235" type="description">
<paragraph id="8BC736E7FFAAFFBF20D9FC60FCA45F1F" blockId="2.[151,1437,150,2040]" pageId="2" pageNumber="235">
<emphasis id="B90CEAF5FFAAFFBF20D9FC60FEAE58E2" bold="true" box="[199,345,978,1004]" pageId="2" pageNumber="235">Description.</emphasis>
Measurements. (n=8): BL: 1.82 (1.72–1.92); HL: 0.34 (0.33–0.36); HW: 0.46 (0.45–0.48); EL: 0.14 (0.13–0.14); PL: 0.42 (0.41–0.44); PW: 0.56 (0.54–0.58).
</paragraph>
<paragraph id="8BC736E7FFAAFFBF20D9FBA9FAE55F3B" blockId="2.[151,1437,150,2040]" box="[199,1298,1051,1077]" pageId="2" pageNumber="235">Coloration. Body, legs, and antennae reddish brown, with two pairs of dark brown spots on elytra.</paragraph>
<paragraph id="8BC736E7FFAAFFBF20D9FB8DFF185E5B" blockId="2.[151,1437,150,2040]" pageId="2" pageNumber="235">Head. Apex of frontoclypeus arcuate; vertexal grooves indistinct; pair of rows of tubercles present on mesal vertex extending to frontoclypeus, each row with 8–9 tubercles each with one microseta on apex; group of tubercles present on external margin of frontoclypeus beside the mesal vertex row of tubercles on each side. Antenna 11- segmented, inserted above anterior part of eye; long and slender, sparsely setose on antennomeres III–VII, densely setose on antennomeres I–II and VIII–XI; antennomere I and II globular, enlarged, with dorsal setiferous tubercles; antennomere II slightly smaller than I; antennomeres III–VII extremely slender, filiform, slightly dilated at the apex; antennomeres VIII–XI globular; antennomere XI wider than all other antennomeres. Eyes large; eye pedestal present.</paragraph>
<paragraph id="8BC736E7FFAAFFBF20D9FAEDFDB05CF9" blockId="2.[151,1437,150,2040]" pageId="2" pageNumber="235">
Thorax. Pronotum hexagonal (
<figureCitation id="13432A62FFAAFFBF223BFAEDFD8A5E77" box="[549,637,1375,1401]" captionStart="FIGURE 1" captionStartId="3.[151,250,1855,1879]" captionTargetBox="[151,1435,298,1813]" captionTargetId="figure-59@3.[151,1436,270,1829]" captionTargetPageId="3" captionText="FIGURE 1. Dasycerus occultus sp. nov. (A) Habitus in dorsal view. (B) SEM image of pronotum in dorsal view. (C–D) Prothorax in ventral view (black triangles indicate the protergosternal keel). (E–F) Right elytron in dorsolateral view. (G) Aedeagus in lateral view. (H) Female genital chamber in ventral view. Scale bars: 1.0 mm for A; 0.5 mm for C &amp; E; 0.2 mm for G–H." figureDoi="http://doi.org/10.5281/zenodo.6986249" httpUri="https://zenodo.org/record/6986249/files/figure.png" pageId="2" pageNumber="235">Fig. 1B</figureCitation>
), with rows of tubercles on the lateral margin and disc, each tubercle with one microseta on apex. Apicolateral angle distinct, not prominent. Disc with a pair of swellings in front of antebasal transverse groove. Median groove distinct, 0.52–0.54 times as long as length of pronotum, extending from antebasal transverse groove. Oblique tubercle row absent. Lateral tubercle row parallel to margin. Middle of antebasal transverse groove strongly arcuate; basal area with five groups of irregular foveae separated by tubercles. Hypomeron with anterior transverse groove. Protergosternal keel entire (
<figureCitation id="13432A62FFAAFFBF23DCF9A2FBEC5D24" box="[962,1051,1552,1578]" captionStart="FIGURE 1" captionStartId="3.[151,250,1855,1879]" captionTargetBox="[151,1435,298,1813]" captionTargetId="figure-59@3.[151,1436,270,1829]" captionTargetPageId="3" captionText="FIGURE 1. Dasycerus occultus sp. nov. (A) Habitus in dorsal view. (B) SEM image of pronotum in dorsal view. (C–D) Prothorax in ventral view (black triangles indicate the protergosternal keel). (E–F) Right elytron in dorsolateral view. (G) Aedeagus in lateral view. (H) Female genital chamber in ventral view. Scale bars: 1.0 mm for A; 0.5 mm for C &amp; E; 0.2 mm for G–H." figureDoi="http://doi.org/10.5281/zenodo.6986249" httpUri="https://zenodo.org/record/6986249/files/figure.png" pageId="2" pageNumber="235">Fig. 1D</figureCitation>
). Keel of procoxal process weekly arcuate; some prosternal foveae on prosternum of each side. Elytra rounded laterally and posterolaterally, widest near middle, covering whole abdomen; rows of microsetae along sutural margin and costae; elytra humeral area rounded and raised. Elytral disc with three costae on each elytron, two internal discal costae extending from base to apex of elytron; one external discal costa shorter than internal discal costae, extending from base to 4/5 point of elytron; posterior-lateral margin of elytron with few tubercles which are similar to those on costae. Each elytron with ten rows of circular punctures (
<figureCitation id="13432A62FFAAFFBF222CF957FD7E5DF1" box="[562,649,1765,1791]" captionStart="FIGURE 1" captionStartId="3.[151,250,1855,1879]" captionTargetBox="[151,1435,298,1813]" captionTargetId="figure-59@3.[151,1436,270,1829]" captionTargetPageId="3" captionText="FIGURE 1. Dasycerus occultus sp. nov. (A) Habitus in dorsal view. (B) SEM image of pronotum in dorsal view. (C–D) Prothorax in ventral view (black triangles indicate the protergosternal keel). (E–F) Right elytron in dorsolateral view. (G) Aedeagus in lateral view. (H) Female genital chamber in ventral view. Scale bars: 1.0 mm for A; 0.5 mm for C &amp; E; 0.2 mm for G–H." figureDoi="http://doi.org/10.5281/zenodo.6986249" httpUri="https://zenodo.org/record/6986249/files/figure.png" pageId="2" pageNumber="235">Fig. 1E</figureCitation>
): two rows between sutural margin and first internal discal costa; two rows between first and second internal discal costae; two rows between second and external discal costae; three rows between external discal costa and the edge of elytron (punctures strongly reduced on humeral area, the three rows gradually fused to one toward apex), and one row between marginal costa and margin of elytron. Each discal row with 20–28 punctures; rows 1–4 each with about 26 punctures, rows 5–10 with fewer. Hind wings well developed. Scutellum glossy, oblique. Mesoventral median keel extending from anterior margin of prepectus to apex of internal process. Prepectus as long as distance from its posterior margin to procoxae. Posterior margin of mesovaxal cavity with a distinct projection on each side.
</paragraph>
<paragraph id="8BC736E7FFABFFBE20D9FF25FD7F5BF7" blockId="3.[151,1436,151,249]" pageId="3" pageNumber="236">Abdomen gradually narrowed apically, not visible in dorsal view. Abdominal tergites V–VIII sclerotized, IV– VI with median furrows. Sternite III with three pairs of coxal foveae and five pairs of microsetae on basal margin; intercoxal process depressed at anterior end.</paragraph>
<caption id="DF07666FFFABFFBE2089F88DFF035CCD" ID-DOI="http://doi.org/10.5281/zenodo.6986249" ID-Zenodo-Dep="6986249" httpUri="https://zenodo.org/record/6986249/files/figure.png" pageId="3" pageNumber="236" startId="3.[151,250,1855,1879]" targetBox="[151,1435,298,1813]" targetPageId="3">
<paragraph id="8BC736E7FFABFFBE2089F88DFF035CCD" blockId="3.[151,1437,1855,1987]" pageId="3" pageNumber="236">
<emphasis id="B90CEAF5FFABFFBE2089F88DFEE05C59" bold="true" box="[151,279,1855,1879]" pageId="3" pageNumber="236">FIGURE 1.</emphasis>
<taxonomicName id="4C784D64FFABFFBE213EF8F2FE1E5C59" authority="Hashizume &amp; Maruyama, 2022" authorityName="Hashizume &amp; Maruyama" authorityYear="2022" box="[288,489,1855,1879]" class="Insecta" family="Staphylinidae" genus="Dasycerus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="236" phylum="Arthropoda" rank="species" species="occultus" status="sp. nov.">
<emphasis id="B90CEAF5FFABFFBE213EF8F2FE1E5C59" box="[288,489,1855,1879]" italics="true" pageId="3" pageNumber="236">Dasycerus occultus</emphasis>
</taxonomicName>
<emphasis id="B90CEAF5FFABFFBE21EDF88DFDB05C59" bold="true" box="[499,583,1855,1879]" pageId="3" pageNumber="236">
<taxonomicNameLabel id="A23F578EFFABFFBE21EDF88DFDB05C59" box="[499,583,1855,1879]" pageId="3" pageNumber="236" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A) Habitus in dorsal view. (B) SEM image of pronotum in dorsal view. (C–D) Prothorax in ventral view (black triangles indicate the protergosternal keel). (E–F) Right elytron in dorsolateral view. (G) Aedeagus in lateral view. (H) Female genital chamber in ventral view. Scale bars: 1.0 mm for A; 0.5 mm for C &amp; E; 0.2 mm for G–H.
</paragraph>
</caption>
<paragraph id="8BC736E7FFACFFB920D9FF25FC8F5BF7" blockId="4.[151,1437,151,681]" pageId="4" pageNumber="237">
Male. Aedeagus (
<figureCitation id="13432A62FFACFFB92195FF25FE155BBF" box="[395,482,151,177]" captionStart="FIGURE 1" captionStartId="3.[151,250,1855,1879]" captionTargetBox="[151,1435,298,1813]" captionTargetId="figure-59@3.[151,1436,270,1829]" captionTargetPageId="3" captionText="FIGURE 1. Dasycerus occultus sp. nov. (A) Habitus in dorsal view. (B) SEM image of pronotum in dorsal view. (C–D) Prothorax in ventral view (black triangles indicate the protergosternal keel). (E–F) Right elytron in dorsolateral view. (G) Aedeagus in lateral view. (H) Female genital chamber in ventral view. Scale bars: 1.0 mm for A; 0.5 mm for C &amp; E; 0.2 mm for G–H." figureDoi="http://doi.org/10.5281/zenodo.6986249" httpUri="https://zenodo.org/record/6986249/files/figure.png" pageId="4" pageNumber="237">Fig. 1G</figureCitation>
) symmetrical. Median lobe with basal bulb moderately large, about as long as apical portion. Apical portion strongly arcuate. Paramere weakly arcuate, narrow in middle, widened at apex; base of paramere triangular; four setae on apex of apex of each paramere.
</paragraph>
<paragraph id="8BC736E7FFACFFB920D9FEB1FE3E5A6B" blockId="4.[151,1437,151,681]" pageId="4" pageNumber="237">
Female. Female genital chamber (
<figureCitation id="13432A62FFACFFB92256FEB1FD575A13" box="[584,672,259,285]" captionStart="FIGURE 1" captionStartId="3.[151,250,1855,1879]" captionTargetBox="[151,1435,298,1813]" captionTargetId="figure-59@3.[151,1436,270,1829]" captionTargetPageId="3" captionText="FIGURE 1. Dasycerus occultus sp. nov. (A) Habitus in dorsal view. (B) SEM image of pronotum in dorsal view. (C–D) Prothorax in ventral view (black triangles indicate the protergosternal keel). (E–F) Right elytron in dorsolateral view. (G) Aedeagus in lateral view. (H) Female genital chamber in ventral view. Scale bars: 1.0 mm for A; 0.5 mm for C &amp; E; 0.2 mm for G–H." figureDoi="http://doi.org/10.5281/zenodo.6986249" httpUri="https://zenodo.org/record/6986249/files/figure.png" pageId="4" pageNumber="237">Fig. 1H</figureCitation>
) weakly sclerotized and has a number of fine spines on its surface. Distal gonocoxite has about 10 preapical setae. Gonostylus has one seta slightly shorter than gonostylus at the apex and shorter preapical setae.
</paragraph>
<paragraph id="8BC736E7FFACFFB920D9FEDCFEF85AA3" blockId="4.[151,1437,151,681]" pageId="4" pageNumber="237">
<emphasis id="B90CEAF5FFACFFB920D9FEDCFEBC5A86" bold="true" box="[199,331,366,392]" pageId="4" pageNumber="237">Bionomics.</emphasis>
They appeared to be feed on fruiting bodies of basidiomcetous fungi that arose on the surfaces of dead trees.
</paragraph>
</subSubSection>
<subSubSection id="C362656CFFACFFB920D9FE04FC395ADF" box="[199,974,438,465]" pageId="4" pageNumber="237" type="distribution">
<paragraph id="8BC736E7FFACFFB920D9FE04FC395ADF" blockId="4.[151,1437,151,681]" box="[199,974,438,465]" pageId="4" pageNumber="237">
<emphasis id="B90CEAF5FFACFFB920D9FE04FE965ADE" bold="true" box="[199,353,438,464]" pageId="4" pageNumber="237">Distribution.</emphasis>
<collectingCountry id="F36F7677FFACFFB92179FE05FE505ADF" box="[359,423,439,465]" name="Japan" pageId="4" pageNumber="237">Japan</collectingCountry>
(Honshu, Kyushu (incl. Shimo-Koshiki-shima)).
</paragraph>
</subSubSection>
<subSubSection id="C362656CFFACFFB920D9FE69FC2B5917" pageId="4" pageNumber="237" type="etymology">
<paragraph id="8BC736E7FFACFFB920D9FE69FC2B5917" blockId="4.[151,1437,151,681]" pageId="4" pageNumber="237">
<emphasis id="B90CEAF5FFACFFB920D9FE69FEB95AFB" bold="true" box="[199,334,475,501]" pageId="4" pageNumber="237">Etymology.</emphasis>
The epithet of this new species is derived from the Latin adjective “ 
<taxonomicName id="4C784D64FFACFFB92496FE69FB125AFB" authorityName="Hashizume &amp; Maruyama" authorityYear="2022" box="[1160,1253,475,501]" class="Insecta" family="Staphylinidae" genus="Dasycerus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="237" phylum="Arthropoda" rank="species" species="occultus">
<emphasis id="B90CEAF5FFACFFB92496FE69FB125AFB" box="[1160,1253,475,501]" italics="true" pageId="4" pageNumber="237">occultus</emphasis>
</taxonomicName>
”, which means “hidden”, because this species was previously confused with 
<taxonomicName id="4C784D64FFACFFB92358FE4DFC225916" authorityName=": Lobl &amp; Calame" authorityYear="1996" box="[838,981,511,537]" class="Insecta" family="Staphylinidae" genus="Dasycerus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="237" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B90CEAF5FFACFFB92358FE4DFCAD5916" box="[838,858,511,536]" italics="true" pageId="4" pageNumber="237">D</emphasis>
. 
<emphasis id="B90CEAF5FFACFFB92376FE4DFC225916" box="[872,981,511,536]" italics="true" pageId="4" pageNumber="237">japonicus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C362656CFFACFFB920D9FD91FAD959A7" pageId="4" pageNumber="237" type="discussion">
<paragraph id="8BC736E7FFACFFB920D9FD91FAD959A7" blockId="4.[151,1437,151,681]" pageId="4" pageNumber="237">
<emphasis id="B90CEAF5FFACFFB920D9FD91FECC5933" bold="true" box="[199,315,547,573]" pageId="4" pageNumber="237">Remarks.</emphasis>
<bibRefCitation id="EFE94B16FFACFFB9215DFD91FDB35933" author="Lobl &amp; Calame" box="[323,580,547,573]" firstAuthor="Lobl" pageId="4" pageNumber="237" pagination="247 - 291" refId="ref9214" refString="Lobl, I. &amp; Calame, F. G. (1996) Taxonomy and phylogeny of the Dasycerinae (Coleoptera: Staphylinidae). Journal of Natural History, 30, 247 - 291. https: // doi. org / 10.1080 / 00222939600771151" type="journal article" year="1996">Löbl &amp; Calame (1996)</bibRefCitation>
redescribed “ 
<taxonomicName id="4C784D64FFACFFB922FEFD91FC875932" authorityName=": Lobl &amp; Calame" authorityYear="1996" box="[736,880,547,573]" class="Insecta" family="Staphylinidae" genus="Dasycerus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="237" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B90CEAF5FFACFFB922FEFD91FD035932" box="[736,756,547,572]" italics="true" pageId="4" pageNumber="237">D</emphasis>
. 
<emphasis id="B90CEAF5FFACFFB9231DFD91FC875932" box="[771,880,547,572]" italics="true" pageId="4" pageNumber="237">japonicus</emphasis>
</taxonomicName>
” based on the 
<typeStatus id="54C38845FFACFFB92406FD91FB8F5933" box="[1048,1144,547,573]" pageId="4" pageNumber="237" type="paratype">paratype</typeStatus>
and additional specimens in the revision of 
<taxonomicName id="4C784D64FFACFFB92141FDF5FE22596E" authorityName="Brongniart" authorityYear="1800" box="[351,469,583,608]" class="Insecta" family="Staphylinidae" genus="Dasycerus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="237" phylum="Arthropoda" rank="genus">
<emphasis id="B90CEAF5FFACFFB92141FDF5FE22596E" box="[351,469,583,608]" italics="true" pageId="4" pageNumber="237">Dasycerus</emphasis>
</taxonomicName>
but illustrated the male genitalia of this species. Because it was initially believed that there was only one species of 
<taxonomicName id="4C784D64FFACFFB921F6FDD9FD80598A" authorityName=": Lobl &amp; Calame" authorityYear="1996" box="[488,631,619,645]" class="Insecta" family="Staphylinidae" genus="Dasycerus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="237" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B90CEAF5FFACFFB921F6FDD9FE0B598A" box="[488,508,619,644]" italics="true" pageId="4" pageNumber="237">D</emphasis>
. 
<emphasis id="B90CEAF5FFACFFB92214FDD9FD80598A" box="[522,631,619,644]" italics="true" pageId="4" pageNumber="237">japonicus</emphasis>
</taxonomicName>
in 
<collectingCountry id="F36F7677FFACFFB92282FDD9FD28598B" box="[668,735,619,645]" name="Japan" pageId="4" pageNumber="237">Japan</collectingCountry>
, all previous records of 
<taxonomicName id="4C784D64FFACFFB923F2FDD9FB8B598A" authorityName=": Lobl &amp; Calame" authorityYear="1996" box="[1004,1148,619,645]" class="Insecta" family="Staphylinidae" genus="Dasycerus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="237" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B90CEAF5FFACFFB923F2FDD9FBF7598A" box="[1004,1024,619,644]" italics="true" pageId="4" pageNumber="237">D</emphasis>
. 
<emphasis id="B90CEAF5FFACFFB92411FDD9FB8B598A" box="[1039,1148,619,644]" italics="true" pageId="4" pageNumber="237">japonicus</emphasis>
</taxonomicName>
should be reviewed. This species has been collected from lowland laurel forest to the montane zone at elevations of up to 
<quantity id="4C809B02FFACFFB924D5FD3DFADD59A7" box="[1227,1322,655,681]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="237" unit="m" value="1200.0">1,200 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>