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<document id="D78093561B3B4DBFB10F09032AA84D32" ID-DOI="10.5281/zenodo.199386" ID-GBIF-Dataset="4c4060cd-6045-477e-9f36-e7c653aebe29" ID-ISSN="1175-5326" ID-Zenodo-Dep="199386" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460334800555" checkinUser="plazi" docAuthor="Wei, Lianmeng &amp; Zhang, Lili" docDate="2010" docId="03D187CDAA32FF8CFF08AF2CCB9EFF74" docLanguage="en" docName="zt02683p022.pdf" docOrigin="Zootaxa 2683" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Chrysotus xinanus Wei &amp; Zhang, 2010, sp. nov." docType="treatment" docVersion="9" lastPageNumber="22" masterDocId="FFE8FFB5AA21FF99FF9FAE0EC877FFA3" masterDocTitle="A taxonomic study on Chrysotus Meigen (Diptera: Dolichopodidae) from southwest China: descriptions of eleven new species belonging to the redefined C. laesus - group" masterLastPageNumber="22" masterPageNumber="1" pageNumber="20" updateTime="1698259528598" updateUser="plazi">
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<mods:title id="68766676A197FD78EEF200516BA0D0E9">A taxonomic study on Chrysotus Meigen (Diptera: Dolichopodidae) from southwest China: descriptions of eleven new species belonging to the redefined C. laesus - group</mods:title>
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<mods:namePart id="009AE5F78B8B8E89EAEE12918A235FEA">Wei, Lianmeng</mods:namePart>
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<mods:namePart id="4D7E3C409256A287624B43569A13B098">Zhang, Lili</mods:namePart>
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<mods:date id="55343D3CEB4A036ABA2B7B60AD35BB68">2010</mods:date>
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<treatment id="03D187CDAA32FF8CFF08AF2CCB9EFF74" ID-DOI="http://doi.org/10.5281/zenodo.6209000" ID-GBIF-Taxon="119405127" ID-Zenodo-Dep="6209000" LSID="urn:lsid:plazi:treatment:03D187CDAA32FF8CFF08AF2CCB9EFF74" httpUri="http://treatment.plazi.org/id/03D187CDAA32FF8CFF08AF2CCB9EFF74" lastPageId="21" lastPageNumber="22" pageId="19" pageNumber="20">
<subSubSection id="C3626550AA32FF8AFF08AF2CCA7EFE9F" box="[151,521,290,316]" pageId="19" pageNumber="20" type="nomenclature">
<paragraph id="8BC736DBAA32FF8AFF08AF2CCA7EFE9F" blockId="19.[151,521,290,351]" box="[151,521,290,316]" pageId="19" pageNumber="20">
<heading id="D08F81B7AA32FF8AFF08AF2CCA7EFE9F" bold="true" box="[151,521,290,316]" fontSize="11" level="1" pageId="19" pageNumber="20" reason="1">
<emphasis id="B90CEAC9AA32FF8AFF08AF2CCA7EFE9F" bold="true" box="[151,521,290,316]" pageId="19" pageNumber="20">
11.
<taxonomicName id="4C784D58AA32FF8AFF5EAF2CC9D3FE9F" ID-CoL="92NDF" box="[193,420,290,316]" class="Insecta" family="Dolichopodidae" genus="Chrysotus" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="xinanus" status="sp. nov.">
<emphasis id="B90CEAC9AA32FF8AFF5EAF2CC9D3FE9F" bold="true" box="[193,420,290,316]" italics="true" pageId="19" pageNumber="20">Chrysotus xinanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23F57B2AA32FF8AFE33AF2CCA7EFE9F" box="[428,521,290,316]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3626550AA32FF8AFF08AF4BC88BFEFC" box="[151,252,325,351]" pageId="19" pageNumber="20" type="description">
<paragraph id="8BC736DBAA32FF8AFF08AF4BC88BFEFC" blockId="19.[151,521,290,351]" box="[151,252,325,351]" pageId="19" pageNumber="20">
(
<figureCitation id="13432A5EAA32FF8AFF00AF4BC884FEFC" box="[159,243,325,351]" captionStart="FIGURE 11" captionStartId="20.[151,255,1964,1988]" captionTargetBox="[279,1268,209,1922]" captionTargetId="figure@20.[266,1280,194,1940]" captionTargetPageId="20" captionText="FIGURE 11. Chrysotus xinanus sp. nov. (A) male terminalia, left lateral view; (B C) female terminalia (B, dorsal view; C. lateral view)." httpUri="https://zenodo.org/record/199396/files/figure.png" pageId="19" pageNumber="20">Fig. 11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3626550AA32FF8AFF08AF81C9B7FE57" pageId="19" pageNumber="20" type="diagnosis">
<paragraph id="8BC736DBAA32FF8AFF08AF81C9B7FE57" blockId="19.[151,1437,399,2034]" pageId="19" pageNumber="20">
<emphasis id="B90CEAC9AA32FF8AFF08AF81C960FE0A" bold="true" box="[151,279,399,425]" pageId="19" pageNumber="20">Diagnosis.</emphasis>
Eyes contiguous on face. Proboscis black, palpus black. All coxae blackish; femora blackish; apex of hind tibiae brownish. Squama pale with brown setulae. Halter pale. Male terminalia with hypandrial ventral process rounded apically.
</paragraph>
</subSubSection>
<subSubSection id="C3626550AA32FF8AFF5AAFF1CAB9F9DF" pageId="19" pageNumber="20" type="description">
<paragraph id="8BC736DBAA32FF8AFF5AAFF1CAB9F9DF" blockId="19.[151,1437,399,2034]" pageId="19" pageNumber="20">
<emphasis id="B90CEAC9AA32FF8AFF5AAFF1C9D8FDBA" bold="true" box="[197,431,511,537]" pageId="19" pageNumber="20">Description. Male</emphasis>
: Body length
<quantity id="4C809B3EAA32FF8AFDFFAC0ECAB6FDB9" box="[608,705,512,538]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="19" pageNumber="20" unit="mm" value="2.5">2.5 mm</quantity>
, wing length
<quantity id="4C809B3EAA32FF8AFCF5AC0ECBBBFDB9" box="[874,972,512,538]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="19" pageNumber="20" unit="mm" value="2.2">2.2 mm</quantity>
.
<emphasis id="B90CEAC9AA32FF8AFC47AFF1CC6AFDBA" bold="true" box="[984,1053,511,537]" pageId="19" pageNumber="20">Head</emphasis>
: Metallic green with pale gray pollinosity; eyes contiguous on face. Setulae and setae on head black, postocular setae (including posteroventral setulae) black. Antenna black; first flagellomere subtriangular, 0.7 times as long as wide; arista black, short pubescence. Proboscis black with black setulae; palpus black with 1 black apical seta.
<emphasis id="B90CEAC9AA32FF8AFB6EAC61CD38FD2A" bold="true" box="[1265,1359,623,649]" pageId="19" pageNumber="20">Thorax</emphasis>
: Dark metallic green with pale gray pollinosity. Setulae and setae on thorax black; 6 strong dc, 78 irregularly paired acr short and setula-like; scutellum with 2 pairs of setae, basal pair short and setula-like; propleuron with 1 black setula and 1 black seta on lower part.
<emphasis id="B90CEAC9AA32FF8AFD0EACD1CABCFD5A" bold="true" box="[657,715,735,761]" pageId="19" pageNumber="20">Legs</emphasis>
: Brown, all coxae blackish; femora blackish; tibiae yellowish brown; fore and mid tarsi from tip of tarsomere 1 onwards brownish, hind tarsus brown. Setulae and setae on legs black. Mid and hind coxae each with 1 outer seta. Fore femur with 2 pv apically; mid femur with 2 av apically; hind femur with 23 av apically. Fore tibia with 1 ad, apically 3 short setae; mid tibia with 2 ad and 3 pd, apically with 4 setae; hind tibia with 2 ad and 4 pd, apically with 4 setae. Relative lengths of tibia and 5 tarsomeres of legs LI 43: 24: 10: 7: 5: 5; LII 50: 30: 14: 10: 6: 5; LIII 64: 20: 16: 10: 5: 5.
<emphasis id="B90CEAC9AA32FF8AFAD0AD94CDE4FC17" bold="true" box="[1359,1427,922,948]" pageId="19" pageNumber="20">Wing</emphasis>
: Hyaline, veins black; R4+5 and M1+2 parallel apically, CuAx ratio 0.3. Squama pale with brown setulae. Halter pale.
<emphasis id="B90CEAC9AA32FF8AFF47ADE6C923FBA1" bold="true" box="[216,340,1000,1026]" pageId="19" pageNumber="20">Abdomen</emphasis>
: Dark metallic green with pale gray pollinosity; setulae and setae on abdomen black.
<emphasis id="B90CEAC9AA32FF8AFAC0ADE6C966FB84" italics="true" pageId="19" pageNumber="20">Male terminalia</emphasis>
(
<figureCitation id="13432A5EAA32FF8AFEBEAA03C903FB84" box="[289,372,1037,1063]" captionStart="FIGURE 11" captionStartId="20.[151,255,1964,1988]" captionTargetBox="[279,1268,209,1922]" captionTargetId="figure@20.[266,1280,194,1940]" captionTargetPageId="20" captionText="FIGURE 11. Chrysotus xinanus sp. nov. (A) male terminalia, left lateral view; (B C) female terminalia (B, dorsal view; C. lateral view)." httpUri="https://zenodo.org/record/199396/files/figure.png" pageId="19" pageNumber="20">Fig. 11</figureCitation>
A,
<typeStatus id="54C38879AA32FF8AFE07AA03C98AFB84" box="[408,509,1037,1063]" pageId="19" pageNumber="20" type="paratype">paratype</typeStatus>
from Sichuan, Cifeng). Epandrium with angular projection posteriorly. Ventral projection of epandrium with end pointed, stands at level of distal margin of lateral epandrial lobe. Lateral epandrial lobe short and wide, flattened apically, bearing 2 setae. Surstylus curved. Cercus triangular, with seta. Hypandrium with ventral process rounded apically. Proximal portion of hypandrium with projection. Postgonite with two strip-like arms and anterior process, latter projects from epandrium and longer than arms.
<emphasis id="B90CEAC9AA32FF8AFF08AAC6C881FB41" bold="true" box="[151,246,1224,1250]" pageId="19" pageNumber="20">Female</emphasis>
: Body length
<quantity id="4C809B3EAA32FF8AFE37AAC6CA7EFB41" box="[424,521,1224,1250]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="19" pageNumber="20" unit="mm" value="2.7">2.7 mm</quantity>
, wing length
<quantity id="4C809B3EAA32FF8AFD2DAAC6CB63FB41" box="[690,788,1224,1250]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="19" pageNumber="20" unit="mm" value="2.2">2.2 mm</quantity>
. Similar to male, except terminalia feature. Relative lengths of tibia and 5 tarsomeres of legs LI 78: 40: 20: 15: 9: 9; LII 100: 50: 22: 18: 10: 10; LIII 115: 34: 25: 16: 10: 10.
<emphasis id="B90CEAC9AA32FF8AFE1CAB1DCA7DFA8F" box="[387,522,1299,1324]" italics="true" pageId="19" pageNumber="20">Terminalia</emphasis>
(
<figureCitation id="13432A5EAA32FF8AFD83AB1CCAF7FA8F" box="[540,640,1298,1324]" captionStart="FIGURE 11" captionStartId="20.[151,255,1964,1988]" captionTargetBox="[279,1268,209,1922]" captionTargetId="figure@20.[266,1280,194,1940]" captionTargetPageId="20" captionText="FIGURE 11. Chrysotus xinanus sp. nov. (A) male terminalia, left lateral view; (B C) female terminalia (B, dorsal view; C. lateral view)." httpUri="https://zenodo.org/record/199396/files/figure.png" pageId="19" pageNumber="20">Figs. 11</figureCitation>
A-B). Ventral view profile: Tg67 with deep inverted U-shaped incisions; Tg8 divided into pair of rod-like sclerites, each with large triangular sclerite on inner margin mesally, weakly sclerotized; Tg9 retrogressive; acanthophorites (Tg10) divided into two sclerites, dorsal surface of each with crest bearing 5 acute spines; posterior margin of St5 with shallow U-shaped incision; St6 rectangular, with length shorter than width, anterior margin with shallow U-shaped incision; St7 rectangular, weakly sclerotized, with length longer than width, anterior margin with shallow inverted U-shaped incisions; St8 divided into pair of rod-like sclerites, connected anteriorly by sclerotized bridge; St10 wider strip-like, with apex dental and setose. In profile: both Tg8 and St8 widely separated from each other basally; St8 with fork-like end; St9 comma-shaped, apically long; St10 long, bent, stick-like, expanded apically, with group of setae apically; cercus longer diamond-like, bare.
</paragraph>
</subSubSection>
<subSubSection id="C3626550AA32FF8AFF5AA886C91AF8B1" pageId="19" pageNumber="20" type="materials_examined">
<paragraph id="8BC736DBAA32FF8AFF5AA886C91AF8B1" blockId="19.[151,1437,399,2034]" pageId="19" pageNumber="20">
<emphasis id="B90CEAC9AA32FF8AFF5AA886CA59F901" bold="true" box="[197,558,1672,1698]" pageId="19" pageNumber="20">
<typeStatus id="54C38879AA32FF8AFF5AA886C976F901" box="[197,257,1672,1698]" pageId="19" pageNumber="20">Type</typeStatus>
material.
<typeStatus id="54C38879AA32FF8AFE19A886CA59F901" box="[390,558,1672,1698]" pageId="19" pageNumber="20" type="holotype">HOLOTYPE</typeStatus>
</emphasis>
ɗ, Yunnan: Lijing Yuquan Park,
<quantity id="4C809B3EAA32FF8AFC20A886CC6AF901" box="[959,1053,1672,1698]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="19" pageNumber="20" unit="m" value="2200.0">2200 m</quantity>
,
<date id="FFC6101BAA32FF8AFBB5A886CCB2F901" box="[1066,1221,1672,1698]" pageId="19" pageNumber="20" value="1993-08-14">14.VIII.1993</date>
LM Wei and MH Liu.
<emphasis id="B90CEAC9AA32FF8AFF51A8A3C90EF964" bold="true" box="[206,377,1709,1735]" pageId="19" pageNumber="20">
<typeStatus id="54C38879AA32FF8AFF51A8A3C90EF964" box="[206,377,1709,1735]" pageId="19" pageNumber="20" type="paratype">PARATYPES</typeStatus>
</emphasis>
: 386 ɗ and 419Ψ were collected from Guizhou, Guangxi, Sichuan and Yunnan,
<quantity id="4C809B3EAA32FF8AFAB4A8A3C8C6F94F" metricMagnitude="3" metricUnit="m" metricValue="1.25" metricValueMax="1.8" metricValueMin="0.7" pageId="19" pageNumber="20" unit="m" value="1250.0" valueMax="1800.0" valueMin="700.0">700-1800 m</quantity>
,
<date id="FFC6101BAA32FF8AFF22A8DCC936F94F" box="[189,321,1746,1772]" pageId="19" pageNumber="20" value="1993" valueMax="2010">1993-2010</date>
, LM Wei, SG Jiang, WP Chao, DX Wang, MH Liu, B Long, ZH Yang, JL Wu, ZW Song, M Yan and QL Xiao.
</paragraph>
</subSubSection>
<subSubSection id="C3626550AA32FF8AFF5AA913CD39F894" box="[197,1358,1821,1847]" pageId="19" pageNumber="20" type="etymology">
<paragraph id="8BC736DBAA32FF8AFF5AA913CD39F894" blockId="19.[151,1437,399,2034]" box="[197,1358,1821,1847]" pageId="19" pageNumber="20">
<emphasis id="B90CEAC9AA32FF8AFF5AA913C926F894" bold="true" box="[197,337,1821,1847]" pageId="19" pageNumber="20">Etymology.</emphasis>
The specific name refers to the
<typeStatus id="54C38879AA32FF8AFD55A913CA8AF894" box="[714,765,1821,1847]" pageId="19" pageNumber="20">type</typeStatus>
localities of Xinan area,
<collectingCountry id="F36F764BAA32FF8AFBBCA913CC1EF894" box="[1059,1129,1821,1847]" name="China" pageId="19" pageNumber="20">China</collectingCountry>
(southwest
<collectingCountry id="F36F764BAA32FF8AFB68A913CD35F894" box="[1271,1346,1821,1847]" name="China" pageId="19" pageNumber="20">China</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C3626550AA32FF8CFF5AA94CCB9EFF74" lastPageId="21" lastPageNumber="22" pageId="19" pageNumber="20" type="discussion">
<paragraph id="8BC736DBAA32FF8AFF5AA94CCAAEF821" blockId="19.[151,1437,399,2034]" pageId="19" pageNumber="20">
<emphasis id="B90CEAC9AA32FF8AFF5AA94CC948F8FF" bold="true" box="[197,319,1858,1884]" pageId="19" pageNumber="20">Remarks.</emphasis>
The new species is related to both
<taxonomicName id="4C784D58AA32FF8AFD7BA94DCBD4F8FF" box="[740,931,1859,1884]" class="Insecta" family="Dolichopodidae" genus="Chrysotus" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="biprojicienus" status="sp. nov.">
<emphasis id="B90CEAC9AA32FF8AFD7BA94DCBD4F8FF" box="[740,931,1859,1884]" italics="true" pageId="19" pageNumber="20">C. biprojicienus</emphasis>
</taxonomicName>
,
<emphasis id="B90CEAC9AA32FF8AFC25A94CCC6EF8FF" bold="true" box="[954,1049,1858,1884]" pageId="19" pageNumber="20">
<taxonomicNameLabel id="A23F57B2AA32FF8AFC25A94CCC6EF8FF" box="[954,1049,1858,1884]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C784D58AA32FF8AFBCBA94DCD69F8FF" box="[1108,1310,1859,1884]" class="Insecta" family="Dolichopodidae" genus="Chrysotus" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="apicicaudatus" status="sp. nov.">
<emphasis id="B90CEAC9AA32FF8AFBCBA94DCD69F8FF" box="[1108,1310,1859,1884]" italics="true" pageId="19" pageNumber="20">C. apicicaudatus</emphasis>
</taxonomicName>
,
<emphasis id="B90CEAC9AA32FF8AFAAAA94CCDE3F8FF" bold="true" box="[1333,1428,1858,1884]" pageId="19" pageNumber="20">
<taxonomicNameLabel id="A23F57B2AA32FF8AFAAAA94CCDE3F8FF" box="[1333,1428,1858,1884]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, but distinguished by features provided in the key.
</paragraph>
<paragraph id="8BC736DBAA32FF8AFF5AA983C9ADF851" blockId="19.[151,1437,399,2034]" pageId="19" pageNumber="20">
According to the key of
<bibRefCitation id="EFE94B2AAA32FF8AFE70A983CB7DF804" box="[495,778,1933,1959]" pageId="19" pageNumber="20" refString="Negrobov, O. P., Tsurikov, M. N. &amp; Maslova, O. O. (2000) Revision of the Palaearctic species of the genus Chrysotus Meigen. (Diptera, Dolichopodidae), III. Entomologicheskoe Obozrenie, 79, 227 - 238." type="journal article">
Negrobov
<emphasis id="B90CEAC9AA32FF8AFDEFA980CAD2F804" box="[624,677,1934,1959]" italics="true" pageId="19" pageNumber="20">et al</emphasis>
. (2000)
</bibRefCitation>
, the new species will run to
<taxonomicName id="4C784D58AA32FF8AFBF8A980C93DF86F" authority="Negrobov et Maslova, 1995" authorityName="Negrobov et Maslova" authorityYear="1995" class="Insecta" family="Dolichopodidae" genus="Chrysotus" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="nudisetus">
<emphasis id="B90CEAC9AA32FF8AFBF8A980CC8AF804" box="[1127,1277,1934,1959]" italics="true" pageId="19" pageNumber="20">C. nudisetus</emphasis>
<bibRefCitation id="EFE94B2AAA32FF8AFA9AA983C93DF86F" author="Negrobov" pageId="19" pageNumber="20" refString="Negrobov, O. P. &amp; Maslova, O. O. (1995) Revision of Palaearctic species of the genus Chrysotus Meigen (Diptera, Dolichopodidae), II. Entomologicheskoe Obozrenie, 74, 456 - 466." type="journal article" year="1995">
Negrobov
<emphasis id="B90CEAC9AA32FF8AFA19A980CDEBF804" box="[1414,1436,1934,1959]" italics="true" pageId="19" pageNumber="20">et</emphasis>
Maslova, 1995
</bibRefCitation>
</taxonomicName>
, but can be separated from the latter by surstylus more curved than in the latter; hypandrium with only a ventral process.
</paragraph>
<caption id="DF076653AA35FF8DFF08A9A2C934F84B" httpUri="https://zenodo.org/record/199396/files/figure.png" pageId="20" pageNumber="21" targetBox="[279,1268,209,1922]" targetPageId="20">
<paragraph id="8BC736DBAA35FF8DFF08A9A2C934F84B" blockId="20.[151,1436,1964,2024]" pageId="20" pageNumber="21">
<emphasis id="B90CEAC9AA35FF8DFF08A9A2C952F867" bold="true" box="[151,293,1964,1988]" pageId="20" pageNumber="21">FIGURE 11.</emphasis>
<taxonomicName id="4C784D58AA35FF8DFEB4A9A3C986F867" box="[299,497,1965,1988]" class="Insecta" family="Dolichopodidae" genus="Chrysotus" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="xinanus" status="sp. nov.">
<emphasis id="B90CEAC9AA35FF8DFEB4A9A3C986F867" box="[299,497,1965,1988]" italics="true" pageId="20" pageNumber="21">Chrysotus xinanus</emphasis>
</taxonomicName>
<emphasis id="B90CEAC9AA35FF8DFE68A9A2CA3BF867" bold="true" box="[503,588,1964,1988]" pageId="20" pageNumber="21">
<taxonomicNameLabel id="A23F57B2AA35FF8DFE68A9A2CA3BF867" box="[503,588,1964,1988]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A) male terminalia, left lateral view; (BC) female terminalia (B, dorsal view; C. lateral view).
</paragraph>
</caption>
<paragraph id="8BC736DBAA34FF8CFF5AAE96CB9EFF74" blockId="21.[151,1436,152,215]" pageId="21" pageNumber="22">
Based on the key of
<bibRefCitation id="EFE94B2AAA34FF8CFE5EAE96CA04FF11" author="Becker" box="[449,627,152,178]" pageId="21" pageNumber="22" refString="Becker, T. (1922 b) Dipterologische Studien. Dolichopodidae der Indo-Australischen Region. Capita Zoologica, 1, 1 - 247." type="journal article" year="1922">Becker (1922)</bibRefCitation>
, the new species will run to couplet 3, but differs from all species under the couplet by blackish brown femora and yellowish brown tibiae.
</paragraph>
</subSubSection>
</treatment>
</document>