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<document id="473B16B4B911C5E030A3E7F1F5E65475" ID-DOI="10.11646/zootaxa.3686.3.9" ID-GBIF-Dataset="59be20e6-35ff-4a96-86a0-9c97e9a5387e" ID-ISSN="1175-5326" ID-Zenodo-Dep="223476" ID-ZooBank="1C6C4EC6-C43A-4A34-9242-3040C8565496" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460648343225" checkinUser="plazi" docAuthor="Fate, Caitlin, Perrichot, Vincent &amp; Nel, Andre" docDate="2013" docId="03D187B6A36EFFA8FF46037FFC78C4D4" docLanguage="en" docName="zt03686p400.pdf" docOrigin="Zootaxa 3686 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ectaetia capdoliensis Fate, Perrichot &amp; Nel, 2013, sp. n." docType="treatment" docVersion="7" lastPageNumber="399" masterDocId="FFE8FFCEA36FFFABFFD10022FF96C074" masterDocTitle="A Mid Cretaceous representative of the modern scatopsid genus Ectaetia (Diptera: Scatopsidae: Ectaetiinae)" masterLastPageNumber="400" masterPageNumber="396" pageNumber="397" updateTime="1698337350798" updateUser="plazi">
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<mods:title id="C87D629BB9107526350C215E3EDAE790">A Mid Cretaceous representative of the modern scatopsid genus Ectaetia (Diptera: Scatopsidae: Ectaetiinae)</mods:title>
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<mods:namePart id="791A8D87ECA40577C8F1540D27E42BCD">Fate, Caitlin</mods:namePart>
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<mods:namePart id="890641F8ABF7B8BB1AB1862B4155D991">Perrichot, Vincent</mods:namePart>
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<mods:namePart id="36A3EDFCA06552F4EF07AB9E5D179FF5">Nel, Andre</mods:namePart>
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<mods:date id="AD37287641A207CE65ED705972043879">2013</mods:date>
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<treatment id="03D187B6A36EFFA8FF46037FFC78C4D4" ID-DOI="http://doi.org/10.5281/zenodo.6159474" ID-GBIF-Taxon="119572351" ID-Zenodo-Dep="6159474" LSID="urn:lsid:plazi:treatment:03D187B6A36EFFA8FF46037FFC78C4D4" httpUri="http://treatment.plazi.org/id/03D187B6A36EFFA8FF46037FFC78C4D4" lastPageId="3" lastPageNumber="399" pageId="1" pageNumber="397">
<subSubSection id="C362652BA36EFFAAFF46037FFF7DC3E3" pageId="1" pageNumber="397" type="nomenclature">
<paragraph id="8BC736A0A36EFFAAFF46037FFE49C303" blockId="1.[151,479,861,919]" box="[151,479,861,887]" pageId="1" pageNumber="397">
<heading id="D08F81CCA36EFFAAFF46037FFE49C303" bold="true" box="[151,479,861,887]" fontSize="11" level="1" pageId="1" pageNumber="397" reason="1">
<emphasis id="B90CEAB2A36EFFAAFF46037FFE49C303" bold="true" box="[151,479,861,887]" pageId="1" pageNumber="397">
<taxonomicName id="4C784D23A36EFFAAFF46037FFE03C303" box="[151,405,861,887]" class="Insecta" family="Scatopsidae" genus="Ectaetia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="397" phylum="Arthropoda" rank="species" species="capdoliensis" status="sp. nov.">
<emphasis id="B90CEAB2A36EFFAAFF46037FFE03C303" bold="true" box="[151,405,861,887]" italics="true" pageId="1" pageNumber="397">Ectaetia capdoliensis</emphasis>
</taxonomicName>
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<paragraph id="8BC736A0A36EFFAAFF46035DFF7DC3E3" blockId="1.[151,479,861,919]" box="[151,235,895,919]" pageId="1" pageNumber="397">
(
<figureCitation id="13432A25A36EFFAAFF4E035DFF76C3E3" box="[159,224,895,919]" captionStart="FIGURE 1" captionStartId="2.[151,250,1765,1788]" captionTargetBox="[151,1436,193,1742]" captionTargetId="figure@2.[151,1436,193,1744]" captionTargetPageId="2" captionText="FIGURE 1. Microphotographs of Ectaetia capdoliensis sp. n., holotype IGR. CDL- 7, in Cretaceous French amber. A. Habitus (PL: prothoracic leg; mL: mesothoracic leg; ML: metathoracic leg); B. Detail of head; C. Wing with venation nomenclature; D. Inner side of right metatibia and metatarsi. Scale bars represent 0.25 mm." httpUri="https://zenodo.org/record/223477/files/figure.png" pageId="1" pageNumber="397">Fig. 1</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C362652BA36EFFAAFF4603EAFB11C470" pageId="1" pageNumber="397" type="materials_examined">
<paragraph id="8BC736A0A36EFFAAFF4603EAFB11C470" blockId="1.[151,1437,967,1928]" pageId="1" pageNumber="397">
<emphasis id="B90CEAB2A36EFFAAFF4603EAFE9DC394" bold="true" box="[151,267,968,992]" pageId="1" pageNumber="397">
<typeStatus id="54C38802A36EFFAAFF4603EAFE90C394" box="[151,262,968,992]" pageId="1" pageNumber="397" type="holotype">Holotype</typeStatus>
.
</emphasis>
Specimen IGR.CDL-7, in amber from Cadeuil, Charente-Maritime, SW
<collectingCountry id="F36F7630A36EFFAAFB9003EAFB07C3AB" box="[1089,1169,968,991]" name="France" pageId="1" pageNumber="397">France</collectingCountry>
; deposited in the amber collection of the Geological Department and Museum of the University Rennes 1,
<collectingCountry id="F36F7630A36EFFAAFBE203CFFB15C470" box="[1075,1155,1005,1028]" name="France" pageId="1" pageNumber="397">France</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C362652BA36EFFAAFF16042DFB17C404" pageId="1" pageNumber="397" type="diagnosis">
<paragraph id="8BC736A0A36EFFAAFF16042DFB9FC453" blockId="1.[151,1437,967,1928]" box="[199,1033,1039,1064]" pageId="1" pageNumber="397">
<emphasis id="B90CEAB2A36EFFAAFF16042DFED7C45C" bold="true" box="[199,321,1039,1064]" pageId="1" pageNumber="397">Diagnosis.</emphasis>
<collectingCountry id="F36F7630A36EFFAAFE99042DFE16C453" box="[328,384,1039,1063]" name="India" pageId="1" pageNumber="397">Hind</collectingCountry>
tibia swollen; no apical comb of short setae on hind tibia.
</paragraph>
<paragraph id="8BC736A0A36EFFAAFF160411FB17C404" blockId="1.[151,1437,967,1928]" pageId="1" pageNumber="397">
<emphasis id="B90CEAB2A36EFFAAFF160411FE0CC438" bold="true" box="[199,410,1075,1100]" pageId="1" pageNumber="397">Age and outcrop.</emphasis>
Mid-Cretaceous, latest Albian or earliest Cenomanian, lithological subunit A1sl-A
<emphasis id="B90CEAB2A36EFFAAFA8F0417FA0AC438" box="[1374,1436,1077,1100]" italics="true" pageId="1" pageNumber="397">sensu</emphasis>
<bibRefCitation id="EFE94B51A36EFFAAFF460475FE18C41B" box="[151,398,1111,1136]" pageId="1" pageNumber="397" refString="Perrichot, V., Neraudeau, D. &amp; Tafforeau, P. (2010) Charentese amber. pp. 192 - 207. In: Penney D. (Ed.). Biodiversity of fossils in amber from the major world deposits. Siri Scientific Press, Manchester." type="book chapter">
Perrichot
<emphasis id="B90CEAB2A36EFFAAFED5047BFEA0C404" box="[260,310,1112,1136]" italics="true" pageId="1" pageNumber="397">et al</emphasis>
. (2010)
</bibRefCitation>
= A1a/b
<emphasis id="B90CEAB2A36EFFAAFE26047BFDA3C404" box="[503,565,1113,1136]" italics="true" pageId="1" pageNumber="397">sensu</emphasis>
<bibRefCitation id="EFE94B51A36EFFAAFDED0475FCDFC41B" box="[572,841,1111,1136]" pageId="1" pageNumber="397" refString="Neraudeau, D., Perrichot, V., Colin, J. - P., Girard, V., Gomez, B., Guillocheau, F., Masure, E., Peyrot, D., Tostain, F., Videt, B. &amp; Vullo, R. (2008) A new amber deposit from the Cretaceous (Uppermost Albian-Lowermost Cenomanian) of southwestern France. Cretaceous Research, 29 (5 - 6), 925 - 929. http: // dx. doi. org / 10.1016 / j. cretres. 2008.05.009" type="journal article">
Néraudeau
<emphasis id="B90CEAB2A36EFFAAFD6C047BFD78C404" box="[701,750,1112,1136]" italics="true" pageId="1" pageNumber="397">et al</emphasis>
. (2008)
</bibRefCitation>
; Cadeuil, Charente,
<collectingCountry id="F36F7630A36EFFAAFBFC047BFBEBC404" box="[1069,1149,1113,1136]" name="France" pageId="1" pageNumber="397">France</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C362652BA36EFFAAFF16045EFBCFC4E0" box="[199,1113,1148,1173]" pageId="1" pageNumber="397" type="etymology">
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<emphasis id="B90CEAB2A36EFFAAFF16045EFED8C4E0" bold="true" box="[199,334,1148,1172]" pageId="1" pageNumber="397">Etymology.</emphasis>
Named after Capdolium, the Latin name of the
<typeStatus id="54C38802A36EFFAAFCB9045FFC0EC4E1" box="[872,920,1149,1173]" pageId="1" pageNumber="397">type</typeStatus>
locality Cadeuil.
</paragraph>
</subSubSection>
<subSubSection id="C362652BA36EFFAAFF1604BDFD96C7FC" pageId="1" pageNumber="397" type="description">
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<emphasis id="B90CEAB2A36EFFAAFF1604BDFECFC4CC" bold="true" box="[199,345,1183,1208]" pageId="1" pageNumber="397">Description.</emphasis>
Head
<quantity id="4C809B45A36EFFAAFE7404BDFD9DC4CC" box="[421,523,1183,1208]" metricMagnitude="-4" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="397" unit="mm" value="0.17">0.17 mm</quantity>
long,
<quantity id="4C809B45A36EFFAAFD8404BDFD2DC4CC" box="[597,699,1183,1208]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="397" unit="mm" value="0.25">0.25 mm</quantity>
wide,
<quantity id="4C809B45A36EFFAAFCDB04BDFCE6C4CC" box="[778,880,1183,1208]" metricMagnitude="-4" metricUnit="m" metricValue="2.9" pageId="1" pageNumber="397" unit="mm" value="0.29">0.29 mm</quantity>
depth; maxillary palpus small; 12 antennomeres (
<figureCitation id="13432A25A36EFFAAFF4E04E6FF76C4A8" box="[159,224,1220,1244]" captionStart="FIGURE 1" captionStartId="2.[151,250,1765,1788]" captionTargetBox="[151,1436,193,1742]" captionTargetId="figure@2.[151,1436,193,1744]" captionTargetPageId="2" captionText="FIGURE 1. Microphotographs of Ectaetia capdoliensis sp. n., holotype IGR. CDL- 7, in Cretaceous French amber. A. Habitus (PL: prothoracic leg; mL: mesothoracic leg; ML: metathoracic leg); B. Detail of head; C. Wing with venation nomenclature; D. Inner side of right metatibia and metatarsi. Scale bars represent 0.25 mm." httpUri="https://zenodo.org/record/223477/files/figure.png" pageId="1" pageNumber="397">Fig. 1</figureCitation>
A, 1B); antennal flagellum short, equal to or shorter than depth of head,
<quantity id="4C809B45A36EFFAAFBFB04E6FB06C4A8" box="[1066,1168,1220,1244]" metricMagnitude="-4" metricUnit="m" metricValue="2.9" pageId="1" pageNumber="397" unit="mm" value="0.29">0.29 mm</quantity>
long, all antennomeres with setae, short and wider than long,
<quantity id="4C809B45A36EFFAAFD9D04C5FD27C574" box="[588,689,1255,1280]" metricMagnitude="-5" metricUnit="m" metricValue="7" pageId="1" pageNumber="397" unit="mm" value="0.07">0.07 mm</quantity>
long; no ocelli visible due to very small bubbles around the head (?); eyes widely separated dorsally and below antennae.
</paragraph>
<paragraph id="8BC736A0A36EFFAAFF160512FDBEC518" blockId="1.[151,1437,967,1928]" pageId="1" pageNumber="397">
Thorax longer than wide,
<quantity id="4C809B45A36EFFAAFE3A0512FDC6C53C" box="[491,592,1328,1352]" metricMagnitude="-4" metricUnit="m" metricValue="4.2" pageId="1" pageNumber="397" unit="mm" value="0.42">0.42 mm</quantity>
long,
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wide; no sign of U-shaped ridge though scutum not clearly visible; moderate pilosity of thorax.
</paragraph>
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Legs. All tibiae without apical spurs; protibia slightly swollen apically; profemur
<quantity id="4C809B45A36EFFAAFBB1055AFB52C5E4" box="[1120,1220,1400,1424]" metricMagnitude="-4" metricUnit="m" metricValue="3.5999999999999996" pageId="1" pageNumber="397" unit="mm" value="0.36">0.36 mm</quantity>
long, protibia
<quantity id="4C809B45A36EFFAAFABA055AFF55C5C0" metricMagnitude="-4" metricUnit="m" metricValue="2.9" pageId="1" pageNumber="397" unit="mm" value="0.29">0.29 mm</quantity>
long, protarsus
<quantity id="4C809B45A36EFFAAFEA605BEFE4CC5C0" box="[375,474,1436,1460]" metricMagnitude="-4" metricUnit="m" metricValue="2.6" pageId="1" pageNumber="397" unit="mm" value="0.26">0.26 mm</quantity>
long; metafemur
<quantity id="4C809B45A36EFFAAFD7305BEFC93C5C0" box="[674,773,1436,1460]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" pageId="1" pageNumber="397" unit="mm" value="0.37">0.37 mm</quantity>
long, metatibia
<quantity id="4C809B45A36EFFAAFC6A05BEFB89C5C0" box="[955,1055,1436,1460]" metricMagnitude="-4" metricUnit="m" metricValue="3.4" pageId="1" pageNumber="397" unit="mm" value="0.34">0.34 mm</quantity>
long,
<quantity id="4C809B45A36EFFAAFBB405BEFB5FC5C0" box="[1125,1225,1436,1460]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" pageId="1" pageNumber="397" unit="mm" value="0.09">0.09 mm</quantity>
wide at apex,
<quantity id="4C809B45A36EFFAAFABA05BEFF55C5AC" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="397" unit="mm" value="0.04">0.04 mm</quantity>
at base; metatibia narrow, moderately swollen in distal third, but without any comb of short bristles on inner side (
<figureCitation id="13432A25A36EFFAAFF0205C6FE85C588" box="[211,275,1508,1532]" captionStart="FIGURE 1" captionStartId="2.[151,250,1765,1788]" captionTargetBox="[151,1436,193,1742]" captionTargetId="figure@2.[151,1436,193,1744]" captionTargetPageId="2" captionText="FIGURE 1. Microphotographs of Ectaetia capdoliensis sp. n., holotype IGR. CDL- 7, in Cretaceous French amber. A. Habitus (PL: prothoracic leg; mL: mesothoracic leg; ML: metathoracic leg); B. Detail of head; C. Wing with venation nomenclature; D. Inner side of right metatibia and metatarsi. Scale bars represent 0.25 mm." httpUri="https://zenodo.org/record/223477/files/figure.png" pageId="1" pageNumber="397">Fig. 1</figureCitation>
D).
</paragraph>
<paragraph id="8BC736A0A36EFFAAFF16062AFBDFC734" blockId="1.[151,1437,967,1928]" pageId="1" pageNumber="397">
Wing with reduced venation (
<figureCitation id="13432A25A36EFFAAFDC2062AFDC4C654" box="[531,594,1544,1568]" captionStart="FIGURE 1" captionStartId="2.[151,250,1765,1788]" captionTargetBox="[151,1436,193,1742]" captionTargetId="figure@2.[151,1436,193,1744]" captionTargetPageId="2" captionText="FIGURE 1. Microphotographs of Ectaetia capdoliensis sp. n., holotype IGR. CDL- 7, in Cretaceous French amber. A. Habitus (PL: prothoracic leg; mL: mesothoracic leg; ML: metathoracic leg); B. Detail of head; C. Wing with venation nomenclature; D. Inner side of right metatibia and metatarsi. Scale bars represent 0.25 mm." httpUri="https://zenodo.org/record/223477/files/figure.png" pageId="1" pageNumber="397">Fig. 1</figureCitation>
C),
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long,
<quantity id="4C809B45A36EFFAAFCF8062AFC1BC654" box="[809,909,1544,1568]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="397" unit="mm" value="0.5">0.50 mm</quantity>
wide, veins with setae; membrane without any clear microtrichia or macrotrichia; anal lobe present; anal vein A1 present; veins C, R and Rs darkly pigmented, posterior veins weak; Costa ending at apex of Rs, well before wing apex,
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from wing base and
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from wing apex, C very slightly swollen at junction with Rs; Rs simple, extending to approximate middle of wing, distance between wing base and apex of Rs
<quantity id="4C809B45A36EFFAAFD7206BAFC9BC6C4" box="[675,781,1688,1712]" metricMagnitude="-4" metricUnit="m" metricValue="7.1" pageId="1" pageNumber="397" unit="mm" value="0.71">0.71 mm</quantity>
, between apex of Rs and wing apex
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; distance between the apices of R1 and Rs
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; no cell d; Rs and M fused and slightly swollen for a short but distinct distance (
<quantity id="4C809B45A36EFFAAFED306C2FEFBC68C" box="[258,365,1760,1784]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="397" unit="mm" value="0.05">0.05 mm</quantity>
), stem of M arising from Rs; M distinctly two-branched; length of M1+2 basal of its fork
<quantity id="4C809B45A36EFFAAFABA06C2FF53C768" metricMagnitude="-4" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="397" unit="mm" value="0.12">0.12 mm</quantity>
, length of M1 ca.
<quantity id="4C809B45A36EFFAAFE4F0726FE6FC768" box="[414,505,1796,1820]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="397" unit="mm" value="0.6">0.6 mm</quantity>
, of M2 ca.
<quantity id="4C809B45A36EFFAAFD530726FD48C768" box="[642,734,1796,1820]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="397" unit="mm" value="0.6">0.6 mm</quantity>
; no false, phantom-like vein in area between M2 and CuA1; CuA2 not sharply bent along its course and with no setae; no distinct setae on halter.
</paragraph>
<paragraph id="8BC736A0A36EFFAAFF16076EFD96C7FC" blockId="1.[151,1437,967,1928]" pageId="1" pageNumber="397">
Abdomen apparently bare or with very short microtrichia, broad (maybe female specimen),
<quantity id="4C809B45A36EFFAAFB2F076EFACEC710" box="[1278,1368,1868,1892]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="397" unit="mm" value="1.1">1.1 mm</quantity>
long; genital structures poorly visible.
</paragraph>
</subSubSection>
<caption id="DF076628A36DFFA9FF4606C7FCE8C74C" httpUri="https://zenodo.org/record/223477/files/figure.png" pageId="2" pageNumber="398" targetBox="[151,1436,193,1742]" targetPageId="2">
<paragraph id="8BC736A0A36DFFA9FF4606C7FCE8C74C" blockId="2.[151,1436,1765,1848]" pageId="2" pageNumber="398">
<emphasis id="B90CEAB2A36DFFA9FF4606C7FE85C68E" bold="true" box="[151,275,1765,1788]" pageId="2" pageNumber="398">FIGURE 1.</emphasis>
Microphotographs of
<taxonomicName id="4C784D23A36DFFA9FE2606C4FD47C68F" box="[503,721,1765,1787]" class="Insecta" family="Scatopsidae" genus="Ectaetia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="398" phylum="Arthropoda" rank="species" species="capdoliensis" status="sp. nov.">
<emphasis id="B90CEAB2A36DFFA9FE2606C4FD47C68F" box="[503,721,1765,1787]" italics="true" pageId="2" pageNumber="398">Ectaetia capdoliensis</emphasis>
</taxonomicName>
<emphasis id="B90CEAB2A36DFFA9FD0606C4FC87C68F" bold="true" box="[727,785,1766,1787]" pageId="2" pageNumber="398">
<taxonomicNameLabel id="A23F57C9A36DFFA9FD0606C4FC87C68F" box="[727,785,1766,1787]" pageId="2" pageNumber="398" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, holotype IGR.CDL-7, in Cretaceous French amber. A. Habitus (PL: prothoracic leg; mL: mesothoracic leg; ML: metathoracic leg); B. Detail of head; C. Wing with venation nomenclature; D. Inner side of right metatibia and metatarsi. Scale bars represent 0.25 mm.
</paragraph>
</caption>
<subSubSection id="C362652BA36DFFA8FF160747FC78C4D4" lastPageId="3" lastPageNumber="399" pageId="2" pageNumber="398" type="discussion">
<paragraph id="8BC736A0A36DFFA8FF160747FD53C1F3" blockId="2.[151,1436,1893,2025]" lastBlockId="3.[151,1436,151,1184]" lastPageId="3" lastPageNumber="399" pageId="2" pageNumber="398">
<emphasis id="B90CEAB2A36DFFA9FF160747FEDAC70A" bold="true" box="[199,332,1893,1918]" pageId="2" pageNumber="398">Discussion.</emphasis>
After Amorim &amp; Grimaldi (2006),
<taxonomicName id="4C784D23A36DFFA9FD0C0744FC58C709" box="[733,974,1893,1917]" class="Insecta" family="Scatopsidae" genus="Ectaetia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="398" phylum="Arthropoda" rank="species" species="capdoliensis" status="sp. nov.">
<emphasis id="B90CEAB2A36DFFA9FD0C0744FC58C709" box="[733,974,1893,1917]" italics="true" pageId="2" pageNumber="398">Ectaetia capdoliensis</emphasis>
</taxonomicName>
<emphasis id="B90CEAB2A36DFFA9FC040747FB83C709" bold="true" box="[981,1045,1893,1917]" pageId="2" pageNumber="398">
<taxonomicNameLabel id="A23F57C9A36DFFA9FC040747FB83C709" box="[981,1045,1893,1917]" pageId="2" pageNumber="398" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
has the character states “maxillary palpomeres short, stout, reduced in length”, maybe one-segmented (?), “tibial spurs lost”, “Sc incomplete, not reaching C at wing margin”, “R2+3 lost”, “basal cell completely lost”, “R4+5 ending at least about one-third the distance between wing apex and R1”, “R4+5 simple, R4 lost”, “A1 faint, not reaching the wing margin (but as developed as in recent
<taxonomicName id="4C784D23A36CFFA8FE7500BBFD97C0C4" box="[420,513,153,176]" class="Insecta" family="Scatopsidae" genus="Ectaetia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="399" phylum="Arthropoda" rank="genus">
<emphasis id="B90CEAB2A36CFFA8FE7500BBFD97C0C4" box="[420,513,153,176]" italics="true" pageId="3" pageNumber="399">Ectaetia</emphasis>
</taxonomicName>
”, all synapomorphies of the clade (
<taxonomicName id="4C784D23A36CFFA8FC7100B5FBB9C0DB" box="[928,1071,151,175]" class="Insecta" family="Axymyiidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="399" phylum="Arthropoda" rank="family">Axymyiidae</taxonomicName>
,
<taxonomicName id="4C784D23A36CFFA8FBEC00B5FA97C0DB" box="[1085,1281,151,175]" class="Insecta" family="Canthyloscelidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="399" phylum="Arthropoda" rank="family">Canthyloscelidae</taxonomicName>
,
<taxonomicName id="4C784D23A36CFFA8FADE00B5FA0EC0DB" box="[1295,1432,151,175]" class="Insecta" family="Scatopsidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="399" phylum="Arthropoda" rank="family">Scatopsidae</taxonomicName>
,
<taxonomicName id="4C784D23A36CFFA8FF46009EFEADC0A0" box="[151,315,188,212]" class="Insecta" family="Valeseguyidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="399" phylum="Arthropoda" rank="family">Valeseguyidae</taxonomicName>
) or (Cantyloscelidae,
<taxonomicName id="4C784D23A36CFFA8FD93009EFD5DC0A0" box="[578,715,188,212]" class="Insecta" family="Scatopsidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="399" phylum="Arthropoda" rank="family">Scatopsidae</taxonomicName>
,
<taxonomicName id="4C784D23A36CFFA8FD0A009EFC14C0A0" box="[731,898,188,212]" class="Insecta" family="Valeseguyidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="399" phylum="Arthropoda" rank="family">Valeseguyidae</taxonomicName>
). It has also the character states “medial and cubital wing veins very lightly sclerotized, significantly less than radial veins” and “antennal flagellum short, compact, length shorter or equal to head depth; flagellomeres with stems nestled in concave surface of proximal flagellomere”, synapomorphies of the clade (
<taxonomicName id="4C784D23A36CFFA8FD490105FCCDC14B" box="[664,859,295,319]" class="Insecta" family="Canthyloscelidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="399" phylum="Arthropoda" rank="family">Canthyloscelidae</taxonomicName>
+
<taxonomicName id="4C784D23A36CFFA8FCAC0105FB91C14B" box="[893,1031,295,319]" class="Insecta" family="Scatopsidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="399" phylum="Arthropoda" rank="family">Scatopsidae</taxonomicName>
), and the character state “flagellum with at most 10 flagellomeres” as a synapomorphy of the
<taxonomicName id="4C784D23A36CFFA8FCCB016EFC35C110" box="[794,931,332,356]" class="Insecta" family="Scatopsidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="399" phylum="Arthropoda" rank="family">Scatopsidae</taxonomicName>
. The simple R4+5 is a character absent in the
<taxonomicName id="4C784D23A36CFFA8FF46014DFECDC1F3" box="[151,347,367,391]" class="Insecta" family="Canthyloscelidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="399" phylum="Arthropoda" rank="family">Canthyloscelidae</taxonomicName>
, the sister group of
<taxonomicName id="4C784D23A36CFFA8FDE9014DFD57C1F3" box="[568,705,367,391]" class="Insecta" family="Scatopsidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="399" phylum="Arthropoda" rank="family">Scatopsidae</taxonomicName>
.
</paragraph>
<paragraph id="8BC736A0A36CFFA8FF1601B6FD8FC22B" blockId="3.[151,1436,151,1184]" pageId="3" pageNumber="399">
Following the keys to subfamilies and genera of
<bibRefCitation id="EFE94B51A36CFFA8FD3F01B6FCEBC1D8" author="Cook" box="[750,893,404,428]" pageId="3" pageNumber="399" refString="Cook, E. F. (1981) Scatopsidae. pp. 313 - 319. In: McAlpine, J. F., Peterson, B. V., Shewell, G. E., Teskey, H. J., Vockeroth, J. R. &amp; Wood, D. M. (coordinators). Manual of Nearctic Diptera, 1, Research Branch, Agricultural Canada Monograph, Ottawa, Ontario, 27, 674 pp." type="book" year="1981">Cook (1981)</bibRefCitation>
or
<bibRefCitation id="EFE94B51A36CFFA8FC7501B6FBDFC1D8" author="Haenni" box="[932,1097,404,428]" pageId="3" pageNumber="399" refString="Haenni, J. - P. (1997) Chapter 2.12. Family Scatopsidae. In: Papp, L. &amp; Darvas, B. (eds). Contributions to a manual of Palaearctic Diptera (with special reference to flies of economic importance). Volume 2, Science Herald, Budapest, 1 - 592. pp. 255 - 272." type="journal article" year="1997">Haenni (1997)</bibRefCitation>
,
<taxonomicName id="4C784D23A36CFFA8FB8401B7FAD1C1D8" box="[1109,1351,404,428]" class="Insecta" family="Scatopsidae" genus="Ectaetia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="399" phylum="Arthropoda" rank="species" species="capdoliensis">
<emphasis id="B90CEAB2A36CFFA8FB8401B7FAD1C1D8" box="[1109,1351,404,428]" italics="true" pageId="3" pageNumber="399">Ectaetia capdoliensis</emphasis>
</taxonomicName>
falls in the
<taxonomicName id="4C784D23A36CFFA8FF150195FED6C1BB" box="[196,320,439,463]" class="Insecta" family="Scatopsidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="399" phylum="Arthropoda" rank="subFamily" subFamily="Ectaetiinae">Ectaetiinae</taxonomicName>
because of the following characters: scutum without a U-shaped ridge, fore tibia not produced apically, C not swollen at junction with R4+5; wing with false vein absent; wing membrane without setae, R4+5 and M1+2 fused for a distance, cross-vein R-M obliterated, stem of M1+2 arising distal to base of R4+5. The anterior thoracic spiracle is on a dorsal extension of epimeron I as in the scatopsid subfamilies
<taxonomicName id="4C784D23A36CFFA8FB220206FAFFC248" box="[1267,1385,548,572]" class="Insecta" family="Scatopsidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="399" phylum="Arthropoda" rank="subFamily" subFamily="Aspistinae">Aspistinae</taxonomicName>
and
<taxonomicName id="4C784D23A36CFFA8FF460265FD83C22B" authority="(Amorim 2000: 1071)" baseAuthorityName="Amorim" baseAuthorityPageNumber="1071" baseAuthorityYear="2000" box="[151,533,583,607]" class="Insecta" family="Scatopsidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="399" phylum="Arthropoda" rank="subFamily" subFamily="Ectaetiinae">Ectaetiinae (Amorim 2000: 1071)</taxonomicName>
.
</paragraph>
<paragraph id="8BC736A0A36CFFA8FF16024EFCA6C2B8" blockId="3.[151,1436,151,1184]" pageId="3" pageNumber="399">
Amorim (1994) proposed a classification of the
<taxonomicName id="4C784D23A36CFFA8FD25024EFCEBC2F0" box="[756,893,620,644]" class="Insecta" family="Scatopsidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="399" phylum="Arthropoda" rank="family">Scatopsidae</taxonomicName>
, based on a phylogenetic analysis in which the subfamily
<taxonomicName id="4C784D23A36CFFA8FEC602ADFE05C2D3" box="[279,403,655,679]" class="Insecta" family="Scatopsidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="399" phylum="Arthropoda" rank="subFamily" subFamily="Ectaetiinae">Ectaetiinae</taxonomicName>
is respected and includes only the genus
<taxonomicName id="4C784D23A36CFFA8FC7C02B3FB9CC2DC" box="[941,1034,657,680]" class="Insecta" family="Scatopsidae" genus="Ectaetia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="399" phylum="Arthropoda" rank="genus">
<emphasis id="B90CEAB2A36CFFA8FC7C02B3FB9CC2DC" box="[941,1034,657,680]" italics="true" pageId="3" pageNumber="399">Ectaetia</emphasis>
</taxonomicName>
. The wing venation of
<taxonomicName id="4C784D23A36CFFA8FAEF02B3FEB5C2B8" class="Insecta" family="Scatopsidae" genus="Ectaetia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="399" phylum="Arthropoda" rank="species" species="capdoliensis">
<emphasis id="B90CEAB2A36CFFA8FAEF02B3FEB5C2B8" italics="true" pageId="3" pageNumber="399">Ectaetia capdoliensis</emphasis>
</taxonomicName>
is identical to that of modern
<taxonomicName id="4C784D23A36CFFA8FDA50297FD47C2B8" box="[628,721,693,716]" class="Insecta" family="Scatopsidae" genus="Ectaetia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="399" phylum="Arthropoda" rank="genus">
<emphasis id="B90CEAB2A36CFFA8FDA50297FD47C2B8" box="[628,721,693,716]" italics="true" pageId="3" pageNumber="399">Ectaetia</emphasis>
</taxonomicName>
species.
</paragraph>
<paragraph id="8BC736A0A36CFFA8FF1602F5FEFDC398" blockId="3.[151,1436,151,1184]" pageId="3" pageNumber="399">
<bibRefCitation id="EFE94B51A36CFFA8FF1602F5FE2CC29B" author="Nel" box="[199,442,727,751]" pageId="3" pageNumber="399" refString="Nel, A. &amp; Prokop, J. (2004) New Paleogene Scatopsidae (Diptera: Nematocera) in amber from Czech Republic and France. Acta Societatis Zoologicae Bohemicae, 68, 91 - 98." type="journal article" year="2004">Nel &amp; Prokop (2004)</bibRefCitation>
described the Paleocene-Eocene genus
<emphasis id="B90CEAB2A36CFFA8FCAE02FBFB86C284" box="[895,1040,729,752]" italics="true" pageId="3" pageNumber="399">Paraectaetia</emphasis>
, currently considered as a putative
<taxonomicName id="4C784D23A36CFFA8FF4602DEFE81C360" box="[151,279,764,788]" class="Insecta" family="Scatopsidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="399" phylum="Arthropoda" rank="subFamily" subFamily="Ectaetiinae">Ectaetiinae</taxonomicName>
. This taxon differs from all
<taxonomicName id="4C784D23A36CFFA8FDB702DFFD55C360" box="[614,707,765,788]" class="Insecta" family="Scatopsidae" genus="Ectaetia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="399" phylum="Arthropoda" rank="genus">
<emphasis id="B90CEAB2A36CFFA8FDB702DFFD55C360" box="[614,707,765,788]" italics="true" pageId="3" pageNumber="399">Ectaetia</emphasis>
</taxonomicName>
, including
<taxonomicName id="4C784D23A36CFFA8FC9B02DFFBA8C360" box="[842,1086,764,788]" class="Insecta" family="Scatopsidae" genus="Ectaetia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="399" phylum="Arthropoda" rank="species" species="capdoliensis">
<emphasis id="B90CEAB2A36CFFA8FC9B02DFFBA8C360" box="[842,1086,764,788]" italics="true" pageId="3" pageNumber="399">Ectaetia capdoliensis</emphasis>
</taxonomicName>
, in its hind tibia not swollen.
<taxonomicName id="4C784D23A36CFFA8FF460303FE10C34C" box="[151,390,800,824]" class="Insecta" family="Scatopsidae" genus="Ectaetia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="399" phylum="Arthropoda" rank="species" species="capdoliensis">
<emphasis id="B90CEAB2A36CFFA8FF460303FE10C34C" box="[151,390,800,824]" italics="true" pageId="3" pageNumber="399">Ectaetia capdoliensis</emphasis>
</taxonomicName>
differs from the modern
<taxonomicName id="4C784D23A36CFFA8FD4F0303FD6AC34C" box="[670,764,801,824]" class="Insecta" family="Scatopsidae" genus="Ectaetia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="399" phylum="Arthropoda" rank="genus">
<emphasis id="B90CEAB2A36CFFA8FD4F0303FD6AC34C" box="[670,764,801,824]" italics="true" pageId="3" pageNumber="399">Ectaetia</emphasis>
</taxonomicName>
in the absence of an apical comb of short setae on hind tibia (
<bibRefCitation id="EFE94B51A36CFFA8FF4F0366FEDAC328" author="Freeman" box="[158,332,836,860]" pageId="3" pageNumber="399" refString="Freeman, P. (1985) Family Scatopsidae. pp. 20 - 74. In: Freeman, P. and Lane, R. P. (eds). Bibionid and scatopsid flies (Diptera: Bibionidae and Scatopsidae). Handbook for the Identification of British Insects, 9, 1 - 74." type="journal article" year="1985">Freeman, 1985</bibRefCitation>
). The only fossil representative of the genus,
<taxonomicName id="4C784D23A36CFFA8FC990367FB54C328" authority="Meunier, 1907" authorityName="Meunier" authorityYear="1907" box="[840,1218,836,860]" class="Insecta" family="Scatopsidae" genus="Ectaetia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="399" phylum="Arthropoda" rank="species" species="fasciola">
<emphasis id="B90CEAB2A36CFFA8FC990367FB93C328" box="[840,1029,836,860]" italics="true" pageId="3" pageNumber="399">Ectaetia fasciola</emphasis>
(
<bibRefCitation id="EFE94B51A36CFFA8FBC20366FB2FC328" author="Meunier" box="[1043,1209,836,860]" pageId="3" pageNumber="399" refString="Meunier, F. (1907) Beitrag zur Fauna der Bibioniden, Simuliden, und Rhyphiden des Bernsteins. Jahrbuch der Koniglich- Preussischen Geologischen Landesanstalt und Bergakademie zu Berlin, 24, 391 - 404." type="journal article" year="1907">Meunier, 1907</bibRefCitation>
)
</taxonomicName>
from Baltic amber, has wing
<quantity id="4C809B45A36CFFA8FED40345FEFDC3F4" box="[261,363,871,896]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" pageId="3" pageNumber="399" unit="mm" value="1.75">1.75 mm</quantity>
long, distinctly longer than in
<taxonomicName id="4C784D23A36CFFA8FD1A034BFC2BC3F4" box="[715,957,872,896]" class="Insecta" family="Scatopsidae" genus="Ectaetia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="399" phylum="Arthropoda" rank="species" species="capdoliensis">
<emphasis id="B90CEAB2A36CFFA8FD1A034BFC2BC3F4" box="[715,957,872,896]" italics="true" pageId="3" pageNumber="399">Ectaetia capdoliensis</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFE94B51A36CFFA8FC1C0345FBE7C30B" author="Meunier" box="[973,1137,871,895]" pageId="3" pageNumber="399" refString="Meunier, F. (1907) Beitrag zur Fauna der Bibioniden, Simuliden, und Rhyphiden des Bernsteins. Jahrbuch der Koniglich- Preussischen Geologischen Landesanstalt und Bergakademie zu Berlin, 24, 391 - 404." type="journal article" year="1907">Meunier 1907</bibRefCitation>
; Amorim 1998). After the direct study of the
<typeStatus id="54C38802A36CFFA8FEA903AFFE3EC3D1" box="[376,424,909,933]" pageId="3" pageNumber="399">type</typeStatus>
of
<taxonomicName id="4C784D23A36CFFA8FE0403AFFCC7C3D0" authority="Meunier, 1907" authorityName="Meunier" authorityYear="1907" box="[469,849,908,932]" class="Insecta" family="Scatopsidae" genus="Scatopse" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="399" phylum="Arthropoda" rank="species" species="fasciola">
<emphasis id="B90CEAB2A36CFFA8FE0403AFFD0AC3D0" box="[469,668,908,932]" italics="true" pageId="3" pageNumber="399">Scatopse fasciola</emphasis>
<bibRefCitation id="EFE94B51A36CFFA8FD7603AEFCC7C3D0" author="Meunier" box="[679,849,908,932]" pageId="3" pageNumber="399" refString="Meunier, F. (1907) Beitrag zur Fauna der Bibioniden, Simuliden, und Rhyphiden des Bernsteins. Jahrbuch der Koniglich- Preussischen Geologischen Landesanstalt und Bergakademie zu Berlin, 24, 391 - 404." type="journal article" year="1907">Meunier, 1907</bibRefCitation>
</taxonomicName>
, Haenni (pers. comm.) indicated that this fossil is clearly an
<taxonomicName id="4C784D23A36CFFA8FEDA0393FEFEC3BC" box="[267,360,945,968]" class="Insecta" family="Scatopsidae" genus="Ectaetia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="399" phylum="Arthropoda" rank="genus">
<emphasis id="B90CEAB2A36CFFA8FEDA0393FEFEC3BC" box="[267,360,945,968]" italics="true" pageId="3" pageNumber="399">Ectaetia</emphasis>
</taxonomicName>
for its wing venation, hind tibia swollen in apical third, and presence of the apical comb of short setae on hind tibia.
</paragraph>
<paragraph id="8BC736A0A36CFFA8FF1603D5FC78C4D4" blockId="3.[151,1436,151,1184]" pageId="3" pageNumber="399">
The larvae of the modern representatives of the genus
<taxonomicName id="4C784D23A36CFFA8FCFA03DBFC1EC464" box="[811,904,1017,1040]" class="Insecta" family="Scatopsidae" genus="Ectaetia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="399" phylum="Arthropoda" rank="genus">
<emphasis id="B90CEAB2A36CFFA8FCFA03DBFC1EC464" box="[811,904,1017,1040]" italics="true" pageId="3" pageNumber="399">Ectaetia</emphasis>
</taxonomicName>
live in decaying wood, under bark of decaying stumps, or tree trunks lying on the ground, under wet environments (
<bibRefCitation id="EFE94B51A36CFFA8FC11043EFA92C440" author="Rotheray" box="[960,1284,1052,1076]" pageId="3" pageNumber="399" refString="Rotheray, G. &amp; Horsfield, D. (1997) Ectaetia christii sp. n., a Scottish species similar to Ectaetia clavipes (Diptera, Scatopsidae). Dipterists Digest, 4, 41 - 44." type="journal article" year="1997">Rotheray &amp; Horsfield 1997</bibRefCitation>
;
<bibRefCitation id="EFE94B51A36CFFA8FAC2043EFF42C423" author="Krivosheina" pageId="3" pageNumber="399" refString="Krivosheina, N. P. (2002) [A review of Palaearctic species of the genus Ectaetia Enderlein, 1912 (Diptera, Scatopsidae) with description of new species.] Entomologicheskoe Obozrenie, 81, 726 - 736. [in Russian] (English translation in Entomological Review, 82, 1010 - 1019)" type="journal article" year="2002">Krivosheina 2002</bibRefCitation>
). A similar ecology can be assumed with a good accuracy for
<taxonomicName id="4C784D23A36CFFA8FC040463FACDC423" authority="Nel 1997" authorityName="Nel" authorityYear="1997" box="[981,1371,1087,1112]" class="Insecta" family="Scatopsidae" genus="Ectaetia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="399" phylum="Arthropoda" rank="species" species="capdoliensis">
<emphasis id="B90CEAB2A36CFFA8FC040463FB5DC42C" box="[981,1227,1088,1112]" italics="true" pageId="3" pageNumber="399">Ectaetia capdoliensis</emphasis>
(
<bibRefCitation id="EFE94B51A36CFFA8FB31041DFAC5C423" author="Nel" box="[1248,1363,1087,1111]" pageId="3" pageNumber="399" refString="Nel, A. (1997) The probabilistic inference of unknown data in phylogenetic analysis. In: Grandcolas, P. (ed.). The origin of biodiversity in insects: phylogenetic tests of evolutionary scenarios. Memoires du Museum National d'Histoire Naturelle, Paris, 173, 305 - 327." type="journal article" year="1997">Nel 1997</bibRefCitation>
)
</taxonomicName>
, thus complementing the paleoenvironment already suggested by previous studies for the mid-Cretaceous amber forest of Cadeuil (
<bibRefCitation id="EFE94B51A36CFFA8FECA04AAFD81C4D4" author="Neraudeau" box="[283,535,1160,1184]" pageId="3" pageNumber="399" refString="Neraudeau, D., Perrichot, V., Colin, J. - P., Girard, V., Gomez, B., Guillocheau, F., Masure, E., Peyrot, D., Tostain, F., Videt, B. &amp; Vullo, R. (2008) A new amber deposit from the Cretaceous (Uppermost Albian-Lowermost Cenomanian) of southwestern France. Cretaceous Research, 29 (5 - 6), 925 - 929. http: // dx. doi. org / 10.1016 / j. cretres. 2008.05.009" type="journal article" year="2008">
Néraudeau
<emphasis id="B90CEAB2A36CFFA8FE4C04ABFE58C4D4" box="[413,462,1160,1184]" italics="true" pageId="3" pageNumber="399">et al</emphasis>
. 2008
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;
<bibRefCitation id="EFE94B51A36CFFA8FDF304AAFD7BC4D4" author="Girard" box="[546,749,1160,1184]" pageId="3" pageNumber="399" refString="Girard, V., Schmidt, A. R., Struwe, S., Perrichot, V., Breton, G. &amp; Neraudeau, D. (2009) Taphonomy and palaeoecology of mid- Cretaceous amber-preserved microorganisms from southwestern France. Geodiversitas, 31, 153 - 162. http: // dx. doi. org / 10.5252 / g 2009 n 1 a 14" type="journal article" year="2009">
Girard
<emphasis id="B90CEAB2A36CFFA8FDA204ABFD32C4D4" box="[627,676,1160,1184]" italics="true" pageId="3" pageNumber="399">et al</emphasis>
. 2009
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;
<bibRefCitation id="EFE94B51A36CFFA8FD2904AAFC74C4D4" author="Perrichot" box="[760,994,1160,1184]" pageId="3" pageNumber="399" refString="Perrichot, V., Neraudeau, D. &amp; Tafforeau, P. (2010) Charentese amber. pp. 192 - 207. In: Penney D. (Ed.). Biodiversity of fossils in amber from the major world deposits. Siri Scientific Press, Manchester." type="book chapter" year="2010">
Perrichot
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. 2010
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).
</paragraph>
</subSubSection>
</treatment>
</document>