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<document id="DD59CE8C749B9FD4854D28C68B16A520" ID-CLB-Dataset="45578" ID-DOI="10.5281/zenodo.279882" ID-GBIF-Dataset="1877b029-3def-4acb-96d3-4351b84a59a5" ID-ISSN="1175-5326" ID-Zenodo-Dep="279882" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460432780646" checkinUser="plazi" docAuthor="Vargovitsh, Robert S." docDate="2012" docId="03CC7853487FFFD9FF65FCD7FCA19E73" docLanguage="en" docName="zt03174p021.pdf" docOrigin="Zootaxa 3174" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Arrhopalites macronyx Vargovitsh, 2012, sp. nov." docType="treatment" docVersion="12" lastPageNumber="10" masterDocId="FFF5002B487EFFD0FFF2FFE3FFAC9A23" masterDocTitle="New troglomorphic Arrhopalitidae (Collembola: Symphypleona) from the Western Caucasus" masterLastPageNumber="21" masterPageNumber="1" pageNumber="2" updateTime="1698331604264" updateUser="ExternalLinkService">
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<mods:title id="09F287768C2263C3B6888CAA39622C40">New troglomorphic Arrhopalitidae (Collembola: Symphypleona) from the Western Caucasus</mods:title>
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<mods:namePart id="5D5B62BCC04A741363614C46B1F72647">Vargovitsh, Robert S.</mods:namePart>
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<mods:date id="C0BC9C230F1D1BE23B712EDEACDF9F64">2012</mods:date>
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<mods:identifier id="A1AA0C6888E456F2B39D444117FE2CF4" type="DOI">10.5281/zenodo.279882</mods:identifier>
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<treatment id="03CC7853487FFFD9FF65FCD7FCA19E73" ID-DOI="http://doi.org/10.5281/zenodo.5696866" ID-GBIF-Taxon="119430729" ID-Zenodo-Dep="5696866" LSID="urn:lsid:zoobank.org:act:B7E7FA4C-C37B-4712-96E2-E4B559497A15" httpUri="http://treatment.plazi.org/id/03CC7853487FFFD9FF65FCD7FCA19E73" lastPageId="9" lastPageNumber="10" pageId="1" pageNumber="2">
<subSubSection id="C37F9ACE487FFFD1FF65FCD7FE22994D" pageId="1" pageNumber="2" type="nomenclature">
<paragraph id="8BDAC945487FFFD1FF65FCD7FDBE996E" blockId="1.[151,530,819,878]" box="[151,530,819,846]" pageId="1" pageNumber="2">
<heading id="D0927E29487FFFD1FF65FCD7FDBE996E" bold="true" box="[151,530,819,846]" fontSize="11" level="1" pageId="1" pageNumber="2" reason="1">
<emphasis id="B9111557487FFFD1FF65FCD7FDBE996E" bold="true" box="[151,530,819,846]" pageId="1" pageNumber="2">
<taxonomicName id="4C65B2C6487FFFD1FF65FCD7FE02996D" box="[151,430,820,846]" class="Entognatha" family="Arrhopalitidae" genus="Arrhopalites" kingdom="Animalia" order="Collembola" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="macronyx" status="sp. nov.">
<emphasis id="B9111557487FFFD1FF65FCD7FE02996D" bold="true" box="[151,430,820,846]" italics="true" pageId="1" pageNumber="2">Arrhopalites macronyx</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A222A82C487FFFD1FE47FCD0FDBE996E" box="[437,530,819,845]" pageId="1" pageNumber="2" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BDAC945487FFFD1FF65FCB5FE22994D" blockId="1.[151,530,819,878]" box="[151,398,854,878]" pageId="1" pageNumber="2">
<figureCitation id="135ED5C0487FFFD1FF65FCB5FEA5994D" box="[151,265,854,878]" captionStart-0="FIGURES 1 – 14" captionStart-1="FIGURES 15 – 18" captionStart-2="FIGURES 19 – 24" captionStart-3="FIGURES 25 – 31" captionStart-4="FIGURES 32 – 36" captionStartId-0="3.[151,264,1923,1946]" captionStartId-1="4.[151,264,1976,1999]" captionStartId-2="5.[151,264,1940,1963]" captionStartId-3="6.[151,264,1932,1955]" captionStartId-4="7.[151,264,1916,1939]" captionTargetBox-0="[186,1401,218,1903]" captionTargetBox-1="[199,1384,213,1938]" captionTargetBox-2="[247,1333,193,1912]" captionTargetBox-3="[252,1324,193,1910]" captionTargetBox-4="[168,1414,201,1892]" captionTargetId-0="figure@3.[174,1417,190,1909]" captionTargetId-1="figure@4.[186,1401,193,1955]" captionTargetId-2="figure@5.[237,1355,193,1924]" captionTargetId-3="figure@6.[249,1338,193,1911]" captionTargetId-4="figure@7.[151,1436,193,1895]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionText-0="FIGURES 1 – 14. Arrhopalites macronyx sp. nov.: 1, chaetotaxy of head, frontal view; 2, chaetotaxy of great abdomen, lateral view, male; 3 – 12, shape of setae: 3, ms 1 of Abd VI; 4, mps 3 of Abd VI; 5, seta of head vertex; 6, dorsal seta of mesothorax; 7, posterior dorsal seta of great abdomen; 8, dI- 1 of great abdomen; 9, Ia of dens; 10, Ipe of dens; 11, appendices anales, lateral view; 12, appendices anales, dorsal view; 13 – 14, tenaculum." captionText-1="FIGURES 15 – 18. Arrhopalites macronyx sp. nov.: 15, chaetotaxy of Ant I – III, anterior view; 16, Ant IV, anterior view; 17, chaetotaxy of furca, posterior view; 18, chaetotaxy of female Abd VI, lateral view." captionText-2="FIGURES 19 – 24. Arrhopalites macronyx sp. nov.: 19 – 21, chaetotaxy of trochanter and femur, anterior view: 19, foreleg; 20, mid leg; 21, hind leg; 22 – 24, chaetotaxy of tibiotarsus and foot complex: 22, foreleg, posterior view; 23, mid leg, posterior view; 24, hind leg, anterior view." captionText-3="FIGURES 25 – 31. Arrhopalites macronyx sp. nov.: 25, habitus and pigmentation, alcohol specimen; 26, habitus of holotype, on slide; 27, fragment of head dorsum with eye field; 28, chaetotaxy of trichobothrial complex; 29 – 30, Ant III: 29, entire segment; 30, fragment with antennal organ; 31, fragment of Ant IV with annulations between subsegments." captionText-4="FIGURES 32 – 36. Arrhopalites macronyx sp. nov.: 32, distal parts of legs (a—foreleg, b—mid leg, c—hind leg, d—the longest fore claw of type series); 33, dens base, anterior view; 34, mucro (a—lateral view, b—dorso-lateral view, c—dorsal view); 35, chaetotaxy of central dorsal complex of great abdomen; 36, fragment of female Abd VI with winged circumanal setae." httpUri-0="https://zenodo.org/record/279883/files/figure.png" httpUri-1="https://zenodo.org/record/279884/files/figure.png" httpUri-2="https://zenodo.org/record/279885/files/figure.png" httpUri-3="https://zenodo.org/record/279886/files/figure.png" httpUri-4="https://zenodo.org/record/279887/files/figure.png" pageId="1" pageNumber="2">Figs 1–36</figureCitation>
,
<tableCitation id="C6E7FCFE487FFFD1FEE6FCB5FEDE994D" box="[276,370,854,878]" captionStart="TABLE 1" captionStartId="2.[151,235,153,174]" captionTargetBox="[151,1436,275,1290]" captionTargetPageId="2" captionText="TABLE 1. Lengths (in µm) for body parts of types of Arrhopalites macronyx sp. nov. and Troglopalites stygios gen. nov. sp. nov. (all measurements, except total body length, are given only for females)." httpUri="http://table.plazi.org/id/DF1A99CD487CFFD2FF65FF7AFC559AEC" pageId="1" pageNumber="2" tableUuid="DF1A99CD487CFFD2FF65FF7AFC559AEC">Tables 1</tableCitation>
,
<tableCitation id="C6E7FCFE487FFFD1FE72FCB5FE22994D" box="[384,398,854,878]" captionStart="TABLE 2" captionStartId="9.[151,235,440,461]" captionText="TABLE 2. Differences between Troglopalites stygios gen. nov. sp. nov., Arrhopalites macronyx sp. nov. and similar species. Species ABC angle Ant: H Ant IV ss Ant IV setae whorls AO-III rods, µm H spiny setae H: tita I T. stygios ~ flat 3 13 20 5 – 8 - 1.2" pageId="1" pageNumber="2">2</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37F9ACE487FFFD1FF65FC7DFED39E91" pageId="1" pageNumber="2" type="diagnosis">
<paragraph id="8BDAC945487FFFD1FF65FC7DFED39E91" blockId="1.[151,1436,926,2033]" pageId="1" pageNumber="2">
<emphasis id="B9111557487FFFD1FF65FC7DFEBD9994" bold="true" box="[151,273,926,951]" pageId="1" pageNumber="2">Diagnosis.</emphasis>
Trichobothria ABC form about right angle and AB&gt;BC. Antenna about 2× of head; Ant IV with 14 whorls of setae and with 8 (pseudo)subsegments indistinctly separated from each other by several annuli. No spiny setae on head. All claws much elongated and thin, without tunica and inner tooth; fore and mid claw 2× shorter than tibiotarsus. Empodia much shorter than claws, only empodium III with small corner tooth. Manubrium with 5 + 5 setae; dens without distinct spines, with 3, 2, 1, 1, 1 thick anterior setae. Tip of mucro rounded but not broadened; posterior lamellae serrated with teeth often accreted together, anterior lamella well-developed. Small abdomen without cuticular spines; circumanal setae winged and serrated basally; appendices anales rod-like, curved, apically acuminated.
</paragraph>
</subSubSection>
<subSubSection id="C37F9ACE487FFFD1FF37FB5CFD4A9F45" pageId="1" pageNumber="2" type="materials_examined">
<paragraph id="8BDAC945487FFFD1FF37FB5CFBFD9F61" blockId="1.[151,1436,926,2033]" pageId="1" pageNumber="2">
<emphasis id="B9111557487FFFD1FF37FB5CFEDE9EF4" bold="true" box="[197,370,1214,1239]" pageId="1" pageNumber="2">
<typeStatus id="54DE77E7487FFFD1FF37FB5CFEAC9EF4" box="[197,256,1215,1239]" pageId="1" pageNumber="2">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54DE77E7487FFFD1FE8AFB5DFE4C9EF5" box="[376,480,1214,1238]" pageId="1" pageNumber="2" type="holotype">Holotype</typeStatus>
on slide “C-698-5”: female (
<figureCitation id="135ED5C0487FFFD1FCD6FB5DFCDB9EF5" box="[804,887,1214,1238]" captionStart="FIGURES 25 – 31" captionStartId="6.[151,264,1932,1955]" captionTargetBox="[252,1324,193,1910]" captionTargetId="figure@6.[249,1338,193,1911]" captionTargetPageId="6" captionText="FIGURES 25 – 31. Arrhopalites macronyx sp. nov.: 25, habitus and pigmentation, alcohol specimen; 26, habitus of holotype, on slide; 27, fragment of head dorsum with eye field; 28, chaetotaxy of trichobothrial complex; 29 – 30, Ant III: 29, entire segment; 30, fragment with antennal organ; 31, fragment of Ant IV with annulations between subsegments." httpUri="https://zenodo.org/record/279886/files/figure.png" pageId="1" pageNumber="2">Fig. 26</figureCitation>
), Western Caucasus, Abkhazia, Gudauta District, Gumishkha Massif, about
<quantity id="4C9D64A0487FFFD1FE32FB01FE569ED9" box="[448,506,1250,1274]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="2" unit="km" value="5.0">5 km</quantity>
north-east from Novy Afon, Anukhvinskaya Cave,
<quantity id="4C9D64A0487FFFD1FBB6FB01FB279ED9" box="[1092,1163,1250,1274]" metricMagnitude="2" metricUnit="m" metricValue="5.8" pageId="1" pageNumber="2" unit="m" value="580.0">580 m</quantity>
a.s.l.,
<date id="FFDBEF85487FFFD1FB24FB01FAF39ED9" box="[1238,1375,1250,1274]" pageId="1" pageNumber="2" value="2009-08-25">25.viii.2009</date>
, R.S. Vargovitsh leg.
<typeStatus id="54DE77E7487FFFD1FEBAFAE4FE199F3C" box="[328,437,1287,1311]" pageId="1" pageNumber="2" type="paratype">Paratypes</typeStatus>
on slides:
<specimenCount id="9D6302CC487FFFD1FDDDFAE5FD249F3D" box="[559,648,1286,1310]" pageId="1" pageNumber="2" type="male">2 males</specimenCount>
,
<specimenCount id="9D6302CC487FFFD1FD67FAE5FCBE9F3D" box="[661,786,1286,1310]" pageId="1" pageNumber="2" type="female">10 females</specimenCount>
, Anukhvinskaya Cave,
<date id="FFDBEF85487FFFD1FBEFFAE5FB0A9F3D" box="[1053,1190,1286,1310]" pageId="1" pageNumber="2" value="2009-08-25">25.viii.2009</date>
, R.S. Vargovitsh leg.; male, female, same cave and collector,
<date id="FFDBEF85487FFFD1FDBDFAC9FD769F61" box="[591,730,1322,1346]" pageId="1" pageNumber="2" value="2010-08-22">22.viii.2010</date>
.
<typeStatus id="54DE77E7487FFFD1FD16FAC8FC8B9F60" box="[740,807,1323,1347]" pageId="1" pageNumber="2">Types</typeStatus>
are preserved in SIZNAS.
</paragraph>
<paragraph id="8BDAC945487FFFD1FF37FAADFD4A9F45" blockId="1.[151,1436,926,2033]" box="[197,742,1358,1382]" pageId="1" pageNumber="2">
Besides slides,
<specimenCount id="9D6302CC487FFFD1FE83FAADFDA59F45" box="[369,521,1358,1382]" pageId="1" pageNumber="2" type="generic">62 specimens</specimenCount>
are kept in alcohol.
</paragraph>
</subSubSection>
<subSubSection id="C37F9ACE487FFFD8FF37FA91FA3A9CC6" lastPageId="8" lastPageNumber="9" pageId="1" pageNumber="2" type="description">
<paragraph id="8BDAC945487FFFD1FF37FA91FC929FD5" blockId="1.[151,1436,926,2033]" pageId="1" pageNumber="2">
<emphasis id="B9111557487FFFD1FF37FA91FE6C9FA8" bold="true" box="[197,448,1394,1419]" pageId="1" pageNumber="2">Description. Female:</emphasis>
body about
<quantity id="4C9D64A0487FFFD1FDA6FA91FD019FA9" box="[596,685,1394,1418]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="2" unit="mm" value="1.2">1.2 mm</quantity>
length (
<typeStatus id="54DE77E7487FFFD1FCFFFA91FCDD9FA9" box="[781,881,1394,1418]" pageId="1" pageNumber="2" type="holotype">holotype</typeStatus>
<quantity id="4C9D64A0487FFFD1FC89FA91FC4A9FA9" box="[891,998,1394,1418]" metricMagnitude="-3" metricUnit="m" metricValue="1.23" pageId="1" pageNumber="2" unit="mm" value="1.23">1.23 mm</quantity>
). Coloration (alcohol material): specimens collected in 2010 with usually very weak diffuse reddish-brownish pigment spots on dorsal side, a little more intensive on head dorsum and with very small eye spots (
<figureCitation id="135ED5C0487FFFD1FCD7FA59FCD69FF1" box="[805,890,1466,1490]" captionStart="FIGURES 25 – 31" captionStartId="6.[151,264,1932,1955]" captionTargetBox="[252,1324,193,1910]" captionTargetId="figure@6.[249,1338,193,1911]" captionTargetPageId="6" captionText="FIGURES 25 – 31. Arrhopalites macronyx sp. nov.: 25, habitus and pigmentation, alcohol specimen; 26, habitus of holotype, on slide; 27, fragment of head dorsum with eye field; 28, chaetotaxy of trichobothrial complex; 29 – 30, Ant III: 29, entire segment; 30, fragment with antennal organ; 31, fragment of Ant IV with annulations between subsegments." httpUri="https://zenodo.org/record/279886/files/figure.png" pageId="1" pageNumber="2">Fig. 25</figureCitation>
); the largest specimen is pigmented more intensively; specimens collected in 2009 are totally unpigmented.
</paragraph>
<paragraph id="8BDAC945487FFFD1FF37F9E1FEFC9C8E" blockId="1.[151,1436,926,2033]" pageId="1" pageNumber="2">
Head (
<figureCitation id="135ED5C0487FFFD1FEE2F9E1FEFD9C39" box="[272,337,1538,1562]" captionStart="FIGURES 1 – 14" captionStartId="3.[151,264,1923,1946]" captionTargetBox="[186,1401,218,1903]" captionTargetId="figure@3.[174,1417,190,1909]" captionTargetPageId="3" captionText="FIGURES 1 – 14. Arrhopalites macronyx sp. nov.: 1, chaetotaxy of head, frontal view; 2, chaetotaxy of great abdomen, lateral view, male; 3 – 12, shape of setae: 3, ms 1 of Abd VI; 4, mps 3 of Abd VI; 5, seta of head vertex; 6, dorsal seta of mesothorax; 7, posterior dorsal seta of great abdomen; 8, dI- 1 of great abdomen; 9, Ia of dens; 10, Ipe of dens; 11, appendices anales, lateral view; 12, appendices anales, dorsal view; 13 – 14, tenaculum." httpUri="https://zenodo.org/record/279883/files/figure.png" pageId="1" pageNumber="2">Fig. 1</figureCitation>
). Eyes uncertain to observe, possibly much reduced: 1 + 1 cuticular swellings and not granulated depressions near them are present but usual corneal lenses were not observed (
<figureCitation id="135ED5C0487FFFD1FBFEF9C5FBCD9C1D" box="[1036,1121,1574,1598]" captionStart="FIGURES 25 – 31" captionStartId="6.[151,264,1932,1955]" captionTargetBox="[252,1324,193,1910]" captionTargetId="figure@6.[249,1338,193,1911]" captionTargetPageId="6" captionText="FIGURES 25 – 31. Arrhopalites macronyx sp. nov.: 25, habitus and pigmentation, alcohol specimen; 26, habitus of holotype, on slide; 27, fragment of head dorsum with eye field; 28, chaetotaxy of trichobothrial complex; 29 – 30, Ant III: 29, entire segment; 30, fragment with antennal organ; 31, fragment of Ant IV with annulations between subsegments." httpUri="https://zenodo.org/record/279886/files/figure.png" pageId="1" pageNumber="2">Fig. 27</figureCitation>
). Labral chaetotaxy a: 4, m: 5, p: 5; prelabral: 6 (=6/554). Clypeal area with axial seta in row a. Interantennal area: 2 rows (α and β) with axial seta in row β. Dorsal area: 4 rows (from A to D) with 3 axial setae in rows A, B and C; no modified (spine-like) setae (
<figureCitation id="135ED5C0487FFFD1FF2CF976FEEE9C8E" box="[222,322,1685,1709]" captionStart="FIGURES 1 – 14" captionStartId="3.[151,264,1923,1946]" captionTargetBox="[186,1401,218,1903]" captionTargetId="figure@3.[174,1417,190,1909]" captionTargetPageId="3" captionText="FIGURES 1 – 14. Arrhopalites macronyx sp. nov.: 1, chaetotaxy of head, frontal view; 2, chaetotaxy of great abdomen, lateral view, male; 3 – 12, shape of setae: 3, ms 1 of Abd VI; 4, mps 3 of Abd VI; 5, seta of head vertex; 6, dorsal seta of mesothorax; 7, posterior dorsal seta of great abdomen; 8, dI- 1 of great abdomen; 9, Ia of dens; 10, Ipe of dens; 11, appendices anales, lateral view; 12, appendices anales, dorsal view; 13 – 14, tenaculum." httpUri="https://zenodo.org/record/279883/files/figure.png" pageId="1" pageNumber="2">Figs 1, 5</figureCitation>
).
</paragraph>
<paragraph id="8BDAC945487FFFD1FF37F959FE189DD2" blockId="1.[151,1436,926,2033]" pageId="1" pageNumber="2">
Antennae (
<figureCitation id="135ED5C0487FFFD1FECDF95AFE6C9CF2" box="[319,448,1721,1745]" captionStart="FIGURES 15 – 18" captionStartId="4.[151,264,1976,1999]" captionTargetBox="[199,1384,213,1938]" captionTargetId="figure@4.[186,1401,193,1955]" captionTargetPageId="4" captionText="FIGURES 15 – 18. Arrhopalites macronyx sp. nov.: 15, chaetotaxy of Ant I – III, anterior view; 16, Ant IV, anterior view; 17, chaetotaxy of furca, posterior view; 18, chaetotaxy of female Abd VI, lateral view." httpUri="https://zenodo.org/record/279884/files/figure.png" pageId="1" pageNumber="2">Figs 15, 16</figureCitation>
): length about 2× (1.8–2.3) of head (
<quantity id="4C9D64A0487FFFD1FCA9F95AFC309CF2" box="[859,924,1721,1745]" metricMagnitude="-2" metricUnit="m" metricValue="5.842" pageId="1" pageNumber="2" unit="in" value="2.3">2.3 in</quantity>
<typeStatus id="54DE77E7487FFFD1FC51F95AFBA69CF2" box="[931,1034,1721,1745]" pageId="1" pageNumber="2" type="holotype">holotype</typeStatus>
). Mean length ratio of antennal segments I: II: III: IV = 1: 2.2: 3.2: 9 (1: 2.3: 3.5:
<quantity id="4C9D64A0487FFFD1FD25F93EFCBB9CD6" box="[727,791,1757,1781]" metricMagnitude="-1" metricUnit="m" metricValue="2.4638" pageId="1" pageNumber="2" unit="in" value="9.7">9.7 in</quantity>
<typeStatus id="54DE77E7487FFFD1FCECF93EFC299CD6" box="[798,901,1757,1781]" pageId="1" pageNumber="2" type="holotype">holotype</typeStatus>
). Antennal segment I with 6 short setae, subapical one is very small. Ant II with 14 setae. Ant III without swelling, with 18 setae and 2 relatively long (11.6 µm in
<typeStatus id="54DE77E7487FFFD1FF65F8C6FF579D1E" box="[151,251,1829,1853]" pageId="1" pageNumber="2" type="holotype">holotype</typeStatus>
) and heavy sense rods inserted in a single pit (
<figureCitation id="135ED5C0487FFFD1FCF6F8C6FCF79D1E" box="[772,859,1829,1853]" captionStart="FIGURES 15 – 18" captionStartId="4.[151,264,1976,1999]" captionTargetBox="[199,1384,213,1938]" captionTargetId="figure@4.[186,1401,193,1955]" captionTargetPageId="4" captionText="FIGURES 15 – 18. Arrhopalites macronyx sp. nov.: 15, chaetotaxy of Ant I – III, anterior view; 16, Ant IV, anterior view; 17, chaetotaxy of furca, posterior view; 18, chaetotaxy of female Abd VI, lateral view." httpUri="https://zenodo.org/record/279884/files/figure.png" pageId="1" pageNumber="2">Figs 15</figureCitation>
,
<figureCitation id="135ED5C0487FFFD1FC95F8C6FC039D1E" box="[871,943,1829,1853]" captionStart="FIGURES 25 – 31" captionStartId="6.[151,264,1932,1955]" captionTargetBox="[252,1324,193,1910]" captionTargetId="figure@6.[249,1338,193,1911]" captionTargetPageId="6" captionText="FIGURES 25 – 31. Arrhopalites macronyx sp. nov.: 25, habitus and pigmentation, alcohol specimen; 26, habitus of holotype, on slide; 27, fragment of head dorsum with eye field; 28, chaetotaxy of trichobothrial complex; 29 – 30, Ant III: 29, entire segment; 30, fragment with antennal organ; 31, fragment of Ant IV with annulations between subsegments." httpUri="https://zenodo.org/record/279886/files/figure.png" pageId="1" pageNumber="2">29, 30</figureCitation>
,
<tableCitation id="C6E7FCFE487FFFD1FC49F8C6FBA19D1E" box="[955,1037,1829,1853]" captionStart="TABLE 1" captionStartId="2.[151,235,153,174]" captionTargetBox="[151,1436,275,1290]" captionTargetPageId="2" captionText="TABLE 1. Lengths (in µm) for body parts of types of Arrhopalites macronyx sp. nov. and Troglopalites stygios gen. nov. sp. nov. (all measurements, except total body length, are given only for females)." httpUri="http://table.plazi.org/id/DF1A99CD487CFFD2FF65FF7AFC559AEC" pageId="1" pageNumber="2" tableUuid="DF1A99CD487CFFD2FF65FF7AFC559AEC">Table 1</tableCitation>
); setae Api and Ape short and thin, seta Aai very small and blunt. Ant IV indistinctly subdivided into 7–8 subsegments or pseudosubsegments, often separated from each other by 0–6 weakly developed and hardly visible annuli which usually increase in number towards distal part of the segment (
<figureCitation id="135ED5C0487FFFD1FDD2F872FDDB9D8A" box="[544,631,1937,1961]" captionStart="FIGURES 15 – 18" captionStartId="4.[151,264,1976,1999]" captionTargetBox="[199,1384,213,1938]" captionTargetId="figure@4.[186,1401,193,1955]" captionTargetPageId="4" captionText="FIGURES 15 – 18. Arrhopalites macronyx sp. nov.: 15, chaetotaxy of Ant I – III, anterior view; 16, Ant IV, anterior view; 17, chaetotaxy of furca, posterior view; 18, chaetotaxy of female Abd VI, lateral view." httpUri="https://zenodo.org/record/279884/files/figure.png" pageId="1" pageNumber="2">Figs 16</figureCitation>
,
<figureCitation id="135ED5C0487FFFD1FD71F872FD0E9D8A" box="[643,674,1937,1961]" captionStart="FIGURES 25 – 31" captionStartId="6.[151,264,1932,1955]" captionTargetBox="[252,1324,193,1910]" captionTargetId="figure@6.[249,1338,193,1911]" captionTargetPageId="6" captionText="FIGURES 25 – 31. Arrhopalites macronyx sp. nov.: 25, habitus and pigmentation, alcohol specimen; 26, habitus of holotype, on slide; 27, fragment of head dorsum with eye field; 28, chaetotaxy of trichobothrial complex; 29 – 30, Ant III: 29, entire segment; 30, fragment with antennal organ; 31, fragment of Ant IV with annulations between subsegments." httpUri="https://zenodo.org/record/279886/files/figure.png" pageId="1" pageNumber="2">31</figureCitation>
). (Pseudo)subsegmental formula: 1 + 6 + 1; Ant IV bears 14 whorls of setae: 4 on apical subsegment (AI–AIII + M1), 6 on median subsegments (M2–M6 + BA) and 4 on basal subsegment (BM1–BM3 + BB).
</paragraph>
<caption id="DF1A99CD487CFFD2FF65FF7AFC559AEC" ID-Table-UUID="DF1A99CD487CFFD2FF65FF7AFC559AEC" httpUri="http://table.plazi.org/id/DF1A99CD487CFFD2FF65FF7AFC559AEC" pageId="2" pageNumber="3" targetBox="[151,1436,275,1290]" targetIsTable="true" targetPageId="2">
<paragraph id="8BDAC945487CFFD2FF65FF7AFC559AEC" blockId="2.[151,1436,151,256]" pageId="2" pageNumber="3">
<emphasis id="B9111557487CFFD2FF65FF7AFEA89A8D" bold="true" box="[151,260,153,174]" pageId="2" pageNumber="3">TABLE 1.</emphasis>
Lengths (in µm) for body parts of types of
<taxonomicName id="4C65B2C6487CFFD2FD23FF7BFC699A8C" box="[721,965,152,175]" class="Entognatha" family="Arrhopalitidae" genus="Arrhopalites" kingdom="Animalia" order="Collembola" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="macronyx" status="sp. nov.">
<emphasis id="B9111557487CFFD2FD23FF7BFC699A8C" box="[721,965,152,175]" italics="true" pageId="2" pageNumber="3">Arrhopalites macronyx</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A222A82C487CFFD2FC3EFF74FB8D9A8C" box="[972,1057,151,175]" pageId="2" pageNumber="3" rank="species">
<emphasis id="B9111557487CFFD2FC3EFF74FBB69A8C" bold="true" box="[972,1050,151,175]" pageId="2" pageNumber="3">sp. nov</emphasis>
.
</taxonomicNameLabel>
and
<taxonomicName id="4C65B2C6487CFFD2FBA6FF7BFA9E9A8C" box="[1108,1330,152,175]" class="Entognatha" family="Arrhopalitidae" genus="Troglopalites" kingdom="Animalia" order="Collembola" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="stygios" status="gen. nov.">
<emphasis id="B9111557487CFFD2FBA6FF7BFA9E9A8C" box="[1108,1330,152,175]" italics="true" pageId="2" pageNumber="3">Troglopalites stygios</emphasis>
</taxonomicName>
<emphasis id="B9111557487CFFD2FACAFF74FF409AEC" bold="true" pageId="2" pageNumber="3">
<taxonomicNameLabel id="A222A82C487CFFD2FACAFF74FA309A8C" box="[1336,1436,151,175]" pageId="2" pageNumber="3" rank="genus">gen. nov.</taxonomicNameLabel>
sp. nov.
</emphasis>
(all measurements, except total body length, are given only for females).
</paragraph>
</caption>
<paragraph id="8BDAC945487CFFD2FF6DFF09FB2A9B23" blockId="2.[151,1436,151,256]" box="[159,1158,234,256]" pageId="2" pageNumber="3">
Body part
<emphasis id="B9111557487CFFD2FE01FF09FB2A9B23" box="[499,1158,234,256]" italics="true" pageId="2" pageNumber="3">
<taxonomicName id="4C65B2C6487CFFD2FE01FF09FD779B23" box="[499,731,234,256]" class="Entognatha" family="Arrhopalitidae" genus="Arrhopalites" kingdom="Animalia" order="Collembola" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="macronyx">Arrhopalites macronyx</taxonomicName>
<taxonomicName id="4C65B2C6487CFFD2FC46FF09FB2A9B23" box="[948,1158,234,256]" class="Entognatha" family="Arrhopalitidae" genus="Troglopalites" kingdom="Animalia" order="Collembola" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="stygios">Troglopalites stygios</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8BDAC945487CFFD2FE01FEF0FA909F2A" pageId="2" pageNumber="3">
<table id="F9653BE5487C002FFF65FEF0FA309F29" box="[151,1436,275,1290]" gridcols="5" gridrows="25" pageId="2" pageNumber="3">
<tr id="3555CB07487C002FFF65FEF0FA309B09" box="[151,1436,275,298]" gridrow="0" pageId="2" pageNumber="3" rowspan-0="1">
<th id="7684A27B487C002FFE00FEF0FDDD9B09" box="[498,625,275,298]" gridcol="1" gridrow="0" pageId="2" pageNumber="3">min–max</th>
<th id="7684A27B487C002FFCE1FEF0FCE19B09" box="[787,845,275,298]" gridcol="2" gridrow="0" pageId="2" pageNumber="3">mean</th>
<th id="7684A27B487C002FFC46FEF0FB9E9B09" box="[948,1074,275,298]" gridcol="3" gridrow="0" pageId="2" pageNumber="3">min–max</th>
<th id="7684A27B487C002FFAF1FEF0FA309B09" box="[1283,1436,275,298]" gridcol="4" gridrow="0" pageId="2" pageNumber="3">mean</th>
</tr>
<tr id="3555CB07487C002FFF65FEDEFA309B70" box="[151,1436,317,339]" gridrow="1" pageId="2" pageNumber="3">
<th id="7684A27B487C002FFF65FEDEFE2A9B70" box="[151,390,317,339]" gridcol="0" gridrow="1" pageId="2" pageNumber="3">Body total, male</th>
<td id="7684A27B487C002FFE00FEDEFDDD9B70" box="[498,625,317,339]" gridcol="1" gridrow="1" pageId="2" pageNumber="3">831–1074</td>
<td id="7684A27B487C002FFCE1FEDEFCE19B70" box="[787,845,317,339]" gridcol="2" gridrow="1" pageId="2" pageNumber="3">927</td>
<td id="7684A27B487C002FFC46FEDEFB9E9B70" box="[948,1074,317,339]" gridcol="3" gridrow="1" pageId="2" pageNumber="3">1150–1257</td>
<td id="7684A27B487C002FFAF1FEDEFA309B70" box="[1283,1436,317,339]" gridcol="4" gridrow="1" pageId="2" pageNumber="3">1203</td>
</tr>
<tr id="3555CB07487C002FFF65FE85FA309B5F" box="[151,1436,358,380]" gridrow="2" pageId="2" pageNumber="3">
<th id="7684A27B487C002FFF65FE85FE2A9B5F" box="[151,390,358,380]" gridcol="0" gridrow="2" pageId="2" pageNumber="3">Body total, female</th>
<td id="7684A27B487C002FFE00FE85FDDD9B5F" box="[498,625,358,380]" gridcol="1" gridrow="2" pageId="2" pageNumber="3">1063–1283</td>
<td id="7684A27B487C002FFCE1FE85FCE19B5F" box="[787,845,358,380]" gridcol="2" gridrow="2" pageId="2" pageNumber="3">1187</td>
<td id="7684A27B487C002FFC46FE85FB9E9B5F" box="[948,1074,358,380]" gridcol="3" gridrow="2" pageId="2" pageNumber="3">1084–1578</td>
<td id="7684A27B487C002FFAF1FE85FA309B5F" box="[1283,1436,358,380]" gridcol="4" gridrow="2" pageId="2" pageNumber="3">1355</td>
</tr>
<tr id="3555CB07487C002FFF65FE73FA309B85" box="[151,1436,400,422]" gridrow="3" pageId="2" pageNumber="3">
<th id="7684A27B487C002FFF65FE73FE2A9B85" box="[151,390,400,422]" gridcol="0" gridrow="3" pageId="2" pageNumber="3">Head</th>
<td id="7684A27B487C002FFE00FE73FDDD9B85" box="[498,625,400,422]" gridcol="1" gridrow="3" pageId="2" pageNumber="3">335–415</td>
<td id="7684A27B487C002FFCE1FE73FCE19B85" box="[787,845,400,422]" gridcol="2" gridrow="3" pageId="2" pageNumber="3">387</td>
<td id="7684A27B487C002FFC46FE73FB9E9B85" box="[948,1074,400,422]" gridcol="3" gridrow="3" pageId="2" pageNumber="3">358–538</td>
<td id="7684A27B487C002FFAF1FE73FA309B85" box="[1283,1436,400,422]" gridcol="4" gridrow="3" pageId="2" pageNumber="3">418</td>
</tr>
<tr id="3555CB07487C002FFF65FE5AFA309BEC" box="[151,1436,441,463]" gridrow="4" pageId="2" pageNumber="3">
<th id="7684A27B487C002FFF65FE5AFE2A9BEC" box="[151,390,441,463]" gridcol="0" gridrow="4" pageId="2" pageNumber="3">Antenna</th>
<td id="7684A27B487C002FFE00FE5AFDDD9BEC" box="[498,625,441,463]" gridcol="1" gridrow="4" pageId="2" pageNumber="3">707–972</td>
<td id="7684A27B487C002FFCE1FE5AFCE19BEC" box="[787,845,441,463]" gridcol="2" gridrow="4" pageId="2" pageNumber="3">793</td>
<td id="7684A27B487C002FFC46FE5AFB9E9BEC" box="[948,1074,441,463]" gridcol="3" gridrow="4" pageId="2" pageNumber="3">967–1500</td>
<td id="7684A27B487C002FFAF1FE5AFA309BEC" box="[1283,1436,441,463]" gridcol="4" gridrow="4" pageId="2" pageNumber="3">1193</td>
</tr>
<tr id="3555CB07487C002FFF65FE01FA309BDB" box="[151,1436,482,504]" gridrow="5" pageId="2" pageNumber="3">
<th id="7684A27B487C002FFF65FE01FE2A9BDB" box="[151,390,482,504]" gridcol="0" gridrow="5" pageId="2" pageNumber="3">Ant I</th>
<td id="7684A27B487C002FFE00FE01FDDD9BDB" box="[498,625,482,504]" gridcol="1" gridrow="5" pageId="2" pageNumber="3">46.7–59</td>
<td id="7684A27B487C002FFCE1FE01FCE19BDB" box="[787,845,482,504]" gridcol="2" gridrow="5" pageId="2" pageNumber="3">53.4</td>
<td id="7684A27B487C002FFC46FE01FB9E9BDB" box="[948,1074,482,504]" gridcol="3" gridrow="5" pageId="2" pageNumber="3">67–86</td>
<td id="7684A27B487C002FFAF1FE01FA309BDB" box="[1283,1436,482,504]" gridcol="4" gridrow="5" pageId="2" pageNumber="3">73</td>
</tr>
<tr id="3555CB07487C002FFF65FDE8FA309801" box="[151,1436,523,546]" gridrow="6" pageId="2" pageNumber="3">
<th id="7684A27B487C002FFF65FDE8FE2A9801" box="[151,390,523,546]" gridcol="0" gridrow="6" pageId="2" pageNumber="3">Ant II</th>
<td id="7684A27B487C002FFE00FDE8FDDD9801" box="[498,625,523,546]" gridcol="1" gridrow="6" pageId="2" pageNumber="3">103.2–134</td>
<td id="7684A27B487C002FFCE1FDE8FCE19801" box="[787,845,523,546]" gridcol="2" gridrow="6" pageId="2" pageNumber="3">117</td>
<td id="7684A27B487C002FFC46FDE8FB9E9801" box="[948,1074,523,546]" gridcol="3" gridrow="6" pageId="2" pageNumber="3">170–270</td>
<td id="7684A27B487C002FFAF1FDE8FA309801" box="[1283,1436,523,546]" gridcol="4" gridrow="6" pageId="2" pageNumber="3">211.6</td>
</tr>
<tr id="3555CB07487C002FFF65FDD6FA309868" box="[151,1436,565,587]" gridrow="7" pageId="2" pageNumber="3">
<th id="7684A27B487C002FFF65FDD6FE2A9868" box="[151,390,565,587]" gridcol="0" gridrow="7" pageId="2" pageNumber="3">Ant III</th>
<td id="7684A27B487C002FFE00FDD6FDDD9868" box="[498,625,565,587]" gridcol="1" gridrow="7" pageId="2" pageNumber="3">141.7–206</td>
<td id="7684A27B487C002FFCE1FDD6FCE19868" box="[787,845,565,587]" gridcol="2" gridrow="7" pageId="2" pageNumber="3">168.7</td>
<td id="7684A27B487C002FFC46FDD6FB9E9868" box="[948,1074,565,587]" gridcol="3" gridrow="7" pageId="2" pageNumber="3">195.6–309</td>
<td id="7684A27B487C002FFAF1FDD6FA309868" box="[1283,1436,565,587]" gridcol="4" gridrow="7" pageId="2" pageNumber="3">240.4</td>
</tr>
<tr id="3555CB07487C002FFF65FDBDFA309857" box="[151,1436,606,628]" gridrow="8" pageId="2" pageNumber="3">
<th id="7684A27B487C002FFF65FDBDFE2A9857" box="[151,390,606,628]" gridcol="0" gridrow="8" pageId="2" pageNumber="3">Ant IV</th>
<td id="7684A27B487C002FFE00FDBDFDDD9857" box="[498,625,606,628]" gridcol="1" gridrow="8" pageId="2" pageNumber="3">419–572</td>
<td id="7684A27B487C002FFCE1FDBDFCE19857" box="[787,845,606,628]" gridcol="2" gridrow="8" pageId="2" pageNumber="3">477.5</td>
<td id="7684A27B487C002FFC46FDBDFB9E9857" box="[948,1074,606,628]" gridcol="3" gridrow="8" pageId="2" pageNumber="3">556.1–865</td>
<td id="7684A27B487C002FFAF1FDBDFA309857" box="[1283,1436,606,628]" gridcol="4" gridrow="8" pageId="2" pageNumber="3">704.6</td>
</tr>
<tr id="3555CB07487C002FFF65FD6BFA3098BD" box="[151,1436,648,670]" gridrow="9" pageId="2" pageNumber="3">
<th id="7684A27B487C002FFF65FD6BFE2A98BD" box="[151,390,648,670]" gridcol="0" gridrow="9" pageId="2" pageNumber="3">AO-III rods</th>
<td id="7684A27B487C002FFE00FD6BFDDD98BD" box="[498,625,648,670]" gridcol="1" gridrow="9" pageId="2" pageNumber="3">9.9–13.2</td>
<td id="7684A27B487C002FFCE1FD6BFCE198BD" box="[787,845,648,670]" gridcol="2" gridrow="9" pageId="2" pageNumber="3">11.5</td>
<td id="7684A27B487C002FFC46FD6BFB9E98BD" box="[948,1074,648,670]" gridcol="3" gridrow="9" pageId="2" pageNumber="3">5.1–8.2</td>
<td id="7684A27B487C002FFAF1FD6BFA3098BD" box="[1283,1436,648,670]" gridcol="4" gridrow="9" pageId="2" pageNumber="3">7.14</td>
</tr>
<tr id="3555CB07487C002FFF65FD52FA3098E4" box="[151,1436,689,711]" gridrow="10" pageId="2" pageNumber="3">
<th id="7684A27B487C002FFF65FD52FE2A98E4" box="[151,390,689,711]" gridcol="0" gridrow="10" pageId="2" pageNumber="3">Tibiotarsus I</th>
<td id="7684A27B487C002FFE00FD52FDDD98E4" box="[498,625,689,711]" gridcol="1" gridrow="10" pageId="2" pageNumber="3">218–284</td>
<td id="7684A27B487C002FFCE1FD52FCE198E4" box="[787,845,689,711]" gridcol="2" gridrow="10" pageId="2" pageNumber="3">253.6</td>
<td id="7684A27B487C002FFC46FD52FB9E98E4" box="[948,1074,689,711]" gridcol="3" gridrow="10" pageId="2" pageNumber="3">290.4–414.2</td>
<td id="7684A27B487C002FFAF1FD52FA3098E4" box="[1283,1436,689,711]" gridcol="4" gridrow="10" pageId="2" pageNumber="3">351.8</td>
</tr>
<tr id="3555CB07487C002FFF65FD39FA3098D3" box="[151,1436,730,752]" gridrow="11" pageId="2" pageNumber="3">
<th id="7684A27B487C002FFF65FD39FE2A98D3" box="[151,390,730,752]" gridcol="0" gridrow="11" pageId="2" pageNumber="3">Tibiotarsus II</th>
<td id="7684A27B487C002FFE00FD39FDDD98D3" box="[498,625,730,752]" gridcol="1" gridrow="11" pageId="2" pageNumber="3">218–270</td>
<td id="7684A27B487C002FFCE1FD39FCE198D3" box="[787,845,730,752]" gridcol="2" gridrow="11" pageId="2" pageNumber="3">245.8</td>
<td id="7684A27B487C002FFC46FD39FB9E98D3" box="[948,1074,730,752]" gridcol="3" gridrow="11" pageId="2" pageNumber="3">271.7–401</td>
<td id="7684A27B487C002FFAF1FD39FA3098D3" box="[1283,1436,730,752]" gridcol="4" gridrow="11" pageId="2" pageNumber="3">335</td>
</tr>
<tr id="3555CB07487C002FFF65FCE0FA309939" box="[151,1436,771,794]" gridrow="12" pageId="2" pageNumber="3">
<th id="7684A27B487C002FFF65FCE0FE2A9939" box="[151,390,771,794]" gridcol="0" gridrow="12" pageId="2" pageNumber="3">Tibiotarsus III</th>
<td id="7684A27B487C002FFE00FCE0FDDD9939" box="[498,625,771,794]" gridcol="1" gridrow="12" pageId="2" pageNumber="3">241–308</td>
<td id="7684A27B487C002FFCE1FCE0FCE19939" box="[787,845,771,794]" gridcol="2" gridrow="12" pageId="2" pageNumber="3">272.9</td>
<td id="7684A27B487C002FFC46FCE0FB9E9939" box="[948,1074,771,794]" gridcol="3" gridrow="12" pageId="2" pageNumber="3">321–486.3</td>
<td id="7684A27B487C002FFAF1FCE0FA309939" box="[1283,1436,771,794]" gridcol="4" gridrow="12" pageId="2" pageNumber="3">401.8</td>
</tr>
<tr id="3555CB07487C002FFF65FCCEFA309960" box="[151,1436,813,835]" gridrow="13" pageId="2" pageNumber="3">
<th id="7684A27B487C002FFF65FCCEFE2A9960" box="[151,390,813,835]" gridcol="0" gridrow="13" pageId="2" pageNumber="3">Claw I</th>
<td id="7684A27B487C002FFE00FCCEFDDD9960" box="[498,625,813,835]" gridcol="1" gridrow="13" pageId="2" pageNumber="3">113–154</td>
<td id="7684A27B487C002FFCE1FCCEFCE19960" box="[787,845,813,835]" gridcol="2" gridrow="13" pageId="2" pageNumber="3">130</td>
<td id="7684A27B487C002FFC46FCCEFB9E9960" box="[948,1074,813,835]" gridcol="3" gridrow="13" pageId="2" pageNumber="3">98–139</td>
<td id="7684A27B487C002FFAF1FCCEFA309960" box="[1283,1436,813,835]" gridcol="4" gridrow="13" pageId="2" pageNumber="3">120</td>
</tr>
<tr id="3555CB07487C002FFF65FCB5FA30994F" box="[151,1436,854,876]" gridrow="14" pageId="2" pageNumber="3">
<th id="7684A27B487C002FFF65FCB5FE2A994F" box="[151,390,854,876]" gridcol="0" gridrow="14" pageId="2" pageNumber="3">Claw II</th>
<td id="7684A27B487C002FFE00FCB5FDDD994F" box="[498,625,854,876]" gridcol="1" gridrow="14" pageId="2" pageNumber="3">111–148</td>
<td id="7684A27B487C002FFCE1FCB5FCE1994F" box="[787,845,854,876]" gridcol="2" gridrow="14" pageId="2" pageNumber="3">129</td>
<td id="7684A27B487C002FFC46FCB5FB9E994F" box="[948,1074,854,876]" gridcol="3" gridrow="14" pageId="2" pageNumber="3">96–133</td>
<td id="7684A27B487C002FFAF1FCB5FA30994F" box="[1283,1436,854,876]" gridcol="4" gridrow="14" pageId="2" pageNumber="3">117</td>
</tr>
<tr id="3555CB07487C002FFF65FC63FA3099B5" box="[151,1436,896,918]" gridrow="15" pageId="2" pageNumber="3">
<th id="7684A27B487C002FFF65FC63FE2A99B5" box="[151,390,896,918]" gridcol="0" gridrow="15" pageId="2" pageNumber="3">Claw III</th>
<td id="7684A27B487C002FFE00FC63FDDD99B5" box="[498,625,896,918]" gridcol="1" gridrow="15" pageId="2" pageNumber="3">100–131</td>
<td id="7684A27B487C002FFCE1FC63FCE199B5" box="[787,845,896,918]" gridcol="2" gridrow="15" pageId="2" pageNumber="3">113.3</td>
<td id="7684A27B487C002FFC46FC63FB9E99B5" box="[948,1074,896,918]" gridcol="3" gridrow="15" pageId="2" pageNumber="3">86–114</td>
<td id="7684A27B487C002FFAF1FC63FA3099B5" box="[1283,1436,896,918]" gridcol="4" gridrow="15" pageId="2" pageNumber="3">101.5</td>
</tr>
<tr id="3555CB07487C002FFF65FC4AFA30999C" box="[151,1436,937,959]" gridrow="16" pageId="2" pageNumber="3">
<th id="7684A27B487C002FFF65FC4AFE2A999C" box="[151,390,937,959]" gridcol="0" gridrow="16" pageId="2" pageNumber="3">Empodium I</th>
<td id="7684A27B487C002FFE00FC4AFDDD999C" box="[498,625,937,959]" gridcol="1" gridrow="16" pageId="2" pageNumber="3">55–82</td>
<td id="7684A27B487C002FFCE1FC4AFCE1999C" box="[787,845,937,959]" gridcol="2" gridrow="16" pageId="2" pageNumber="3">67.1</td>
<td id="7684A27B487C002FFC46FC4AFB9E999C" box="[948,1074,937,959]" gridcol="3" gridrow="16" pageId="2" pageNumber="3">42–61</td>
<td id="7684A27B487C002FFAF1FC4AFA30999C" box="[1283,1436,937,959]" gridcol="4" gridrow="16" pageId="2" pageNumber="3">55</td>
</tr>
<tr id="3555CB07487C002FFF65FC31FA3099CB" box="[151,1436,978,1000]" gridrow="17" pageId="2" pageNumber="3">
<th id="7684A27B487C002FFF65FC31FE2A99CB" box="[151,390,978,1000]" gridcol="0" gridrow="17" pageId="2" pageNumber="3">Empodium II</th>
<td id="7684A27B487C002FFE00FC31FDDD99CB" box="[498,625,978,1000]" gridcol="1" gridrow="17" pageId="2" pageNumber="3">55–82</td>
<td id="7684A27B487C002FFCE1FC31FCE199CB" box="[787,845,978,1000]" gridcol="2" gridrow="17" pageId="2" pageNumber="3">67.2</td>
<td id="7684A27B487C002FFC46FC31FB9E99CB" box="[948,1074,978,1000]" gridcol="3" gridrow="17" pageId="2" pageNumber="3">45–70.3</td>
<td id="7684A27B487C002FFAF1FC31FA3099CB" box="[1283,1436,978,1000]" gridcol="4" gridrow="17" pageId="2" pageNumber="3">55.6</td>
</tr>
<tr id="3555CB07487C002FFF65FC18FA309E31" box="[151,1436,1019,1042]" gridrow="18" pageId="2" pageNumber="3">
<th id="7684A27B487C002FFF65FC18FE2A9E31" box="[151,390,1019,1042]" gridcol="0" gridrow="18" pageId="2" pageNumber="3">Empodium III</th>
<td id="7684A27B487C002FFE00FC18FDDD9E31" box="[498,625,1019,1042]" gridcol="1" gridrow="18" pageId="2" pageNumber="3">74–95</td>
<td id="7684A27B487C002FFCE1FC18FCE19E31" box="[787,845,1019,1042]" gridcol="2" gridrow="18" pageId="2" pageNumber="3">78.2</td>
<td id="7684A27B487C002FFC46FC18FB9E9E31" box="[948,1074,1019,1042]" gridcol="3" gridrow="18" pageId="2" pageNumber="3">54–73</td>
<td id="7684A27B487C002FFAF1FC18FA309E31" box="[1283,1436,1019,1042]" gridcol="4" gridrow="18" pageId="2" pageNumber="3">64.6</td>
</tr>
<tr id="3555CB07487C002FFF65FBC6FA309E18" box="[151,1436,1061,1083]" gridrow="19" pageId="2" pageNumber="3">
<th id="7684A27B487C002FFF65FBC6FE2A9E18" box="[151,390,1061,1083]" gridcol="0" gridrow="19" pageId="2" pageNumber="3">Abd seta dI-1</th>
<td id="7684A27B487C002FFE00FBC6FDDD9E18" box="[498,625,1061,1083]" gridcol="1" gridrow="19" pageId="2" pageNumber="3">70–84</td>
<td id="7684A27B487C002FFCE1FBC6FCE19E18" box="[787,845,1061,1083]" gridcol="2" gridrow="19" pageId="2" pageNumber="3">74.5</td>
<td id="7684A27B487C002FFC46FBC6FB9E9E18" box="[948,1074,1061,1083]" gridcol="3" gridrow="19" pageId="2" pageNumber="3">90–105</td>
<td id="7684A27B487C002FFAF1FBC6FA309E18" box="[1283,1436,1061,1083]" gridcol="4" gridrow="19" pageId="2" pageNumber="3">98.7</td>
</tr>
<tr id="3555CB07487C002FFF65FBADFA309E47" box="[151,1436,1102,1124]" gridrow="20" pageId="2" pageNumber="3">
<th id="7684A27B487C002FFF65FBADFE2A9E47" box="[151,390,1102,1124]" gridcol="0" gridrow="20" pageId="2" pageNumber="3">Abd VI longest seta</th>
<td id="7684A27B487C002FFE00FBADFDDD9E47" box="[498,625,1102,1124]" gridcol="1" gridrow="20" pageId="2" pageNumber="3">45.5–65</td>
<td id="7684A27B487C002FFCE1FBADFCE19E47" box="[787,845,1102,1124]" gridcol="2" gridrow="20" pageId="2" pageNumber="3">55</td>
<td id="7684A27B487C002FFC46FBADFB9E9E47" box="[948,1074,1102,1124]" gridcol="3" gridrow="20" pageId="2" pageNumber="3">50–60</td>
<td id="7684A27B487C002FFAF1FBADFA309E47" box="[1283,1436,1102,1124]" gridcol="4" gridrow="20" pageId="2" pageNumber="3">52</td>
</tr>
<tr id="3555CB07487C002FFF65FB9BFA309EAD" box="[151,1436,1144,1166]" gridrow="21" pageId="2" pageNumber="3">
<th id="7684A27B487C002FFF65FB9BFE2A9EAD" box="[151,390,1144,1166]" gridcol="0" gridrow="21" pageId="2" pageNumber="3">Abd VI append. anales</th>
<td id="7684A27B487C002FFE00FB9BFDDD9EAD" box="[498,625,1144,1166]" gridcol="1" gridrow="21" pageId="2" pageNumber="3">30.5–44.8</td>
<td id="7684A27B487C002FFCE1FB9BFCE19EAD" box="[787,845,1144,1166]" gridcol="2" gridrow="21" pageId="2" pageNumber="3">37.5</td>
<td id="7684A27B487C002FFC46FB9BFB9E9EAD" box="[948,1074,1144,1166]" gridcol="3" gridrow="21" pageId="2" pageNumber="3">23.8–31.5</td>
<td id="7684A27B487C002FFAF1FB9BFA309EAD" box="[1283,1436,1144,1166]" gridcol="4" gridrow="21" pageId="2" pageNumber="3">27.8</td>
</tr>
<tr id="3555CB07487C002FFF65FB42FA309E94" box="[151,1436,1185,1207]" gridrow="22" pageId="2" pageNumber="3">
<th id="7684A27B487C002FFF65FB42FE2A9E94" box="[151,390,1185,1207]" gridcol="0" gridrow="22" pageId="2" pageNumber="3">Manubrium</th>
<td id="7684A27B487C002FFE00FB42FDDD9E94" box="[498,625,1185,1207]" gridcol="1" gridrow="22" pageId="2" pageNumber="3">180–253</td>
<td id="7684A27B487C002FFCE1FB42FCE19E94" box="[787,845,1185,1207]" gridcol="2" gridrow="22" pageId="2" pageNumber="3">220</td>
<td id="7684A27B487C002FFC46FB42FB9E9E94" box="[948,1074,1185,1207]" gridcol="3" gridrow="22" pageId="2" pageNumber="3">213–285</td>
<td id="7684A27B487C002FFAF1FB42FA309E94" box="[1283,1436,1185,1207]" gridcol="4" gridrow="22" pageId="2" pageNumber="3">250</td>
</tr>
<tr id="3555CB07487C002FFF65FB29FA309EC3" box="[151,1436,1226,1248]" gridrow="23" pageId="2" pageNumber="3">
<th id="7684A27B487C002FFF65FB29FE2A9EC3" box="[151,390,1226,1248]" gridcol="0" gridrow="23" pageId="2" pageNumber="3">Dens</th>
<td id="7684A27B487C002FFE00FB29FDDD9EC3" box="[498,625,1226,1248]" gridcol="1" gridrow="23" pageId="2" pageNumber="3">231.7–302</td>
<td id="7684A27B487C002FFCE1FB29FCE19EC3" box="[787,845,1226,1248]" gridcol="2" gridrow="23" pageId="2" pageNumber="3">267.3</td>
<td id="7684A27B487C002FFC46FB29FB9E9EC3" box="[948,1074,1226,1248]" gridcol="3" gridrow="23" pageId="2" pageNumber="3">295–470</td>
<td id="7684A27B487C002FFAF1FB29FA309EC3" box="[1283,1436,1226,1248]" gridcol="4" gridrow="23" pageId="2" pageNumber="3">371</td>
</tr>
<tr id="3555CB07487C002FFF65FB10FA309F29" box="[151,1436,1267,1290]" gridrow="24" pageId="2" pageNumber="3">
<th id="7684A27B487C002FFF65FB10FE2A9F29" box="[151,390,1267,1290]" gridcol="0" gridrow="24" pageId="2" pageNumber="3">Mucro</th>
<td id="7684A27B487C002FFE00FB10FDDD9F29" box="[498,625,1267,1290]" gridcol="1" gridrow="24" pageId="2" pageNumber="3">104.3–135.7</td>
<td id="7684A27B487C002FFCE1FB10FCE19F29" box="[787,845,1267,1290]" gridcol="2" gridrow="24" pageId="2" pageNumber="3">121.5</td>
<td id="7684A27B487C002FFC46FB10FB9E9F29" box="[948,1074,1267,1290]" gridcol="3" gridrow="24" pageId="2" pageNumber="3">140–210</td>
<td id="7684A27B487C002FFAF1FB10FA309F29" box="[1283,1436,1267,1290]" gridcol="4" gridrow="24" pageId="2" pageNumber="3">173.3</td>
</tr>
</table>
</paragraph>
<paragraph id="8BDAC945487CFFD2FF37FAADFDAE9C41" blockId="2.[151,1436,1358,2031]" pageId="2" pageNumber="3">
Foreleg: precoxae 1, 2 and coxa with 1, 0, 1 setae respectively (
<figureCitation id="135ED5C0487CFFD2FC7FFAADFC7E9F45" box="[909,978,1358,1382]" captionStart="FIGURES 1 – 14" captionStartId="3.[151,264,1923,1946]" captionTargetBox="[186,1401,218,1903]" captionTargetId="figure@3.[174,1417,190,1909]" captionTargetPageId="3" captionText="FIGURES 1 – 14. Arrhopalites macronyx sp. nov.: 1, chaetotaxy of head, frontal view; 2, chaetotaxy of great abdomen, lateral view, male; 3 – 12, shape of setae: 3, ms 1 of Abd VI; 4, mps 3 of Abd VI; 5, seta of head vertex; 6, dorsal seta of mesothorax; 7, posterior dorsal seta of great abdomen; 8, dI- 1 of great abdomen; 9, Ia of dens; 10, Ipe of dens; 11, appendices anales, lateral view; 12, appendices anales, dorsal view; 13 – 14, tenaculum." httpUri="https://zenodo.org/record/279883/files/figure.png" pageId="2" pageNumber="3">Fig. 2</figureCitation>
). Trochanter with 3 anterior and 1 posterior setae; femur with 11 setae, a4 turned perpendicularly to the longitudinal axis of the segment (
<figureCitation id="135ED5C0487CFFD2FB21FA91FA8A9FA9" box="[1235,1318,1394,1418]" captionStart="FIGURES 19 – 24" captionStartId="5.[151,264,1940,1963]" captionTargetBox="[247,1333,193,1912]" captionTargetId="figure@5.[237,1355,193,1924]" captionTargetPageId="5" captionText="FIGURES 19 – 24. Arrhopalites macronyx sp. nov.: 19 – 21, chaetotaxy of trochanter and femur, anterior view: 19, foreleg; 20, mid leg; 21, hind leg; 22 – 24, chaetotaxy of tibiotarsus and foot complex: 22, foreleg, posterior view; 23, mid leg, posterior view; 24, hind leg, anterior view." httpUri="https://zenodo.org/record/279885/files/figure.png" pageId="2" pageNumber="3">Fig. 19</figureCitation>
). Tibiotarsus (
<figureCitation id="135ED5C0487CFFD2FF3BFA75FEB19F8D" box="[201,285,1430,1454]" captionStart="FIGURES 19 – 24" captionStartId="5.[151,264,1940,1963]" captionTargetBox="[247,1333,193,1912]" captionTargetId="figure@5.[237,1355,193,1924]" captionTargetPageId="5" captionText="FIGURES 19 – 24. Arrhopalites macronyx sp. nov.: 19 – 21, chaetotaxy of trochanter and femur, anterior view: 19, foreleg; 20, mid leg; 21, hind leg; 22 – 24, chaetotaxy of tibiotarsus and foot complex: 22, foreleg, posterior view; 23, mid leg, posterior view; 24, hind leg, anterior view." httpUri="https://zenodo.org/record/279885/files/figure.png" pageId="2" pageNumber="3">Fig. 22</figureCitation>
): 3 setae FP present (e, ae, pe); secondary seta FS missing; whorl I with 9 setae among which Ja curved and spine-like; each of whorls II–V with 8 setae but whorl V sometimes incomplete (setae Vai and/or Vpi present or absent). Pretarsus with 1 anterior and 1 posterior setulae. Foot complex (
<figureCitation id="135ED5C0487CFFD2FC2CFA3DFB999FD5" box="[990,1077,1502,1526]" captionStart="FIGURES 19 – 24" captionStartId="5.[151,264,1940,1963]" captionTargetBox="[247,1333,193,1912]" captionTargetId="figure@5.[237,1355,193,1924]" captionTargetPageId="5" captionText="FIGURES 19 – 24. Arrhopalites macronyx sp. nov.: 19 – 21, chaetotaxy of trochanter and femur, anterior view: 19, foreleg; 20, mid leg; 21, hind leg; 22 – 24, chaetotaxy of tibiotarsus and foot complex: 22, foreleg, posterior view; 23, mid leg, posterior view; 24, hind leg, anterior view." httpUri="https://zenodo.org/record/279885/files/figure.png" pageId="2" pageNumber="3">Figs 22</figureCitation>
,
<figureCitation id="135ED5C0487CFFD2FBB2FA3DFBF39FD5" box="[1088,1119,1502,1526]" captionStart="FIGURES 32 – 36" captionStartId="7.[151,264,1916,1939]" captionTargetBox="[168,1414,201,1892]" captionTargetId="figure@7.[151,1436,193,1895]" captionTargetPageId="7" captionText="FIGURES 32 – 36. Arrhopalites macronyx sp. nov.: 32, distal parts of legs (a—foreleg, b—mid leg, c—hind leg, d—the longest fore claw of type series); 33, dens base, anterior view; 34, mucro (a—lateral view, b—dorso-lateral view, c—dorsal view); 35, chaetotaxy of central dorsal complex of great abdomen; 36, fragment of female Abd VI with winged circumanal setae." httpUri="https://zenodo.org/record/279887/files/figure.png" pageId="2" pageNumber="3">32</figureCitation>
a). Claw: very long and thin, only 1.8–2× shorter than tibiotarsus (
<quantity id="4C9D64A0487CFFD2FDC7F9E1FD299C39" box="[565,645,1538,1562]" metricMagnitude="-2" metricUnit="m" metricValue="4.9784" pageId="2" pageNumber="3" unit="in" value="1.96">1.96 in</quantity>
<typeStatus id="54DE77E7487CFFD2FD7EF9E1FD5F9C39" box="[652,755,1538,1562]" pageId="2" pageNumber="3" type="holotype">holotype</typeStatus>
); without inner tooth, with 2 pairs of lateral teeth: very small subbasal and sharp thin subapical; without tunica. Empodium: thin, somewhat tapered on basal part, 1.8–2.2× shorter than claw, without tooth.
</paragraph>
<paragraph id="8BDAC945487CFFD2FF37F98DFED49DA1" blockId="2.[151,1436,1358,2031]" pageId="2" pageNumber="3">
Mid leg: precoxae 1 and 2 with 1, 1 setae respectively, precoxal process present, coxa with 2 or 3 setae and microsensilla (
<figureCitation id="135ED5C0487CFFD2FEC9F971FE2C9C89" box="[315,384,1682,1706]" captionStart="FIGURES 1 – 14" captionStartId="3.[151,264,1923,1946]" captionTargetBox="[186,1401,218,1903]" captionTargetId="figure@3.[174,1417,190,1909]" captionTargetPageId="3" captionText="FIGURES 1 – 14. Arrhopalites macronyx sp. nov.: 1, chaetotaxy of head, frontal view; 2, chaetotaxy of great abdomen, lateral view, male; 3 – 12, shape of setae: 3, ms 1 of Abd VI; 4, mps 3 of Abd VI; 5, seta of head vertex; 6, dorsal seta of mesothorax; 7, posterior dorsal seta of great abdomen; 8, dI- 1 of great abdomen; 9, Ia of dens; 10, Ipe of dens; 11, appendices anales, lateral view; 12, appendices anales, dorsal view; 13 – 14, tenaculum." httpUri="https://zenodo.org/record/279883/files/figure.png" pageId="2" pageNumber="3">Fig. 2</figureCitation>
). Trochanter with 3 simple setae and anterior trochanteral organ; femur with 12 setae, 2 posterior ones are shorter and thinner than others (
<figureCitation id="135ED5C0487CFFD2FD6EF955FD5E9CED" box="[668,754,1718,1742]" captionStart="FIGURES 19 – 24" captionStartId="5.[151,264,1940,1963]" captionTargetBox="[247,1333,193,1912]" captionTargetId="figure@5.[237,1355,193,1924]" captionTargetPageId="5" captionText="FIGURES 19 – 24. Arrhopalites macronyx sp. nov.: 19 – 21, chaetotaxy of trochanter and femur, anterior view: 19, foreleg; 20, mid leg; 21, hind leg; 22 – 24, chaetotaxy of tibiotarsus and foot complex: 22, foreleg, posterior view; 23, mid leg, posterior view; 24, hind leg, anterior view." httpUri="https://zenodo.org/record/279885/files/figure.png" pageId="2" pageNumber="3">Fig. 20</figureCitation>
). Tibiotarsus (
<figureCitation id="135ED5C0487CFFD2FC67F955FC409CED" box="[917,1004,1718,1742]" captionStart="FIGURES 19 – 24" captionStartId="5.[151,264,1940,1963]" captionTargetBox="[247,1333,193,1912]" captionTargetId="figure@5.[237,1355,193,1924]" captionTargetPageId="5" captionText="FIGURES 19 – 24. Arrhopalites macronyx sp. nov.: 19 – 21, chaetotaxy of trochanter and femur, anterior view: 19, foreleg; 20, mid leg; 21, hind leg; 22 – 24, chaetotaxy of tibiotarsus and foot complex: 22, foreleg, posterior view; 23, mid leg, posterior view; 24, hind leg, anterior view." httpUri="https://zenodo.org/record/279885/files/figure.png" pageId="2" pageNumber="3">Fig. 23</figureCitation>
): 3 setae FP present, seta FS missing; whorl I with 9 setae among which Ja curved and spine-like, whorls II–IV with 8 setae, whorl V with 7 setae (Vp absent) but sometimes incomplete (setae Vai and/or Vpi present or absent). Foot complex (
<figureCitation id="135ED5C0487CFFD2FB5FF91DFAAB9D35" box="[1197,1287,1790,1814]" captionStart="FIGURES 19 – 24" captionStartId="5.[151,264,1940,1963]" captionTargetBox="[247,1333,193,1912]" captionTargetId="figure@5.[237,1355,193,1924]" captionTargetPageId="5" captionText="FIGURES 19 – 24. Arrhopalites macronyx sp. nov.: 19 – 21, chaetotaxy of trochanter and femur, anterior view: 19, foreleg; 20, mid leg; 21, hind leg; 22 – 24, chaetotaxy of tibiotarsus and foot complex: 22, foreleg, posterior view; 23, mid leg, posterior view; 24, hind leg, anterior view." httpUri="https://zenodo.org/record/279885/files/figure.png" pageId="2" pageNumber="3">Figs 23</figureCitation>
,
<figureCitation id="135ED5C0487CFFD2FAE7F91DFA989D35" box="[1301,1332,1790,1814]" captionStart="FIGURES 32 – 36" captionStartId="7.[151,264,1916,1939]" captionTargetBox="[168,1414,201,1892]" captionTargetId="figure@7.[151,1436,193,1895]" captionTargetPageId="7" captionText="FIGURES 32 – 36. Arrhopalites macronyx sp. nov.: 32, distal parts of legs (a—foreleg, b—mid leg, c—hind leg, d—the longest fore claw of type series); 33, dens base, anterior view; 34, mucro (a—lateral view, b—dorso-lateral view, c—dorsal view); 35, chaetotaxy of central dorsal complex of great abdomen; 36, fragment of female Abd VI with winged circumanal setae." httpUri="https://zenodo.org/record/279887/files/figure.png" pageId="2" pageNumber="3">32</figureCitation>
b). Claw: very long and thin, only 1.8–2× shorter than tibiotarsus (
<quantity id="4C9D64A0487CFFD2FCE6F8C1FCCF9D19" box="[788,867,1826,1850]" metricMagnitude="-2" metricUnit="m" metricValue="4.7244" pageId="2" pageNumber="3" unit="in" value="1.86">1.86 in</quantity>
<typeStatus id="54DE77E7487CFFD2FC99F8C1FC7E9D19" box="[875,978,1826,1850]" pageId="2" pageNumber="3" type="holotype">holotype</typeStatus>
); without inner tooth, with 2 pairs of lateral teeth: very small subbasal and sharp thin subapical; without tunica. Empodium: thin, 1.8–2.2× shorter than claw; without tooth.
</paragraph>
<paragraph id="8BDAC945487CFFD8FF37F86DFCAB9B1C" blockId="2.[151,1436,1358,2031]" lastBlockId="8.[151,1437,151,2017]" lastPageId="8" lastPageNumber="9" pageId="2" pageNumber="3">
<collectingCountry id="F37289D5487CFFD2FF37F86DFF519D85" box="[197,253,1934,1958]" name="India" pageId="2" pageNumber="3">Hind</collectingCountry>
leg: precoxae 1 and 2 with 1, 1 setae respectively, process on precoxa 1 present, coxa with 3 setae and microsensilla (
<figureCitation id="135ED5C0487CFFD2FECFF851FE289DE9" box="[317,388,1970,1994]" captionStart="FIGURES 1 – 14" captionStartId="3.[151,264,1923,1946]" captionTargetBox="[186,1401,218,1903]" captionTargetId="figure@3.[174,1417,190,1909]" captionTargetPageId="3" captionText="FIGURES 1 – 14. Arrhopalites macronyx sp. nov.: 1, chaetotaxy of head, frontal view; 2, chaetotaxy of great abdomen, lateral view, male; 3 – 12, shape of setae: 3, ms 1 of Abd VI; 4, mps 3 of Abd VI; 5, seta of head vertex; 6, dorsal seta of mesothorax; 7, posterior dorsal seta of great abdomen; 8, dI- 1 of great abdomen; 9, Ia of dens; 10, Ipe of dens; 11, appendices anales, lateral view; 12, appendices anales, dorsal view; 13 – 14, tenaculum." httpUri="https://zenodo.org/record/279883/files/figure.png" pageId="2" pageNumber="3">Fig. 2</figureCitation>
). Trochanter with anterior trochanteral organ, 3 anterior and 1 posterior simple setae; femur with 12 setae, among them 2 very small posterior setulae (
<figureCitation id="135ED5C0487CFFD2FCDDF835FC289DCD" box="[815,900,2006,2030]" captionStart="FIGURES 19 – 24" captionStartId="5.[151,264,1940,1963]" captionTargetBox="[247,1333,193,1912]" captionTargetId="figure@5.[237,1355,193,1924]" captionTargetPageId="5" captionText="FIGURES 19 – 24. Arrhopalites macronyx sp. nov.: 19 – 21, chaetotaxy of trochanter and femur, anterior view: 19, foreleg; 20, mid leg; 21, hind leg; 22 – 24, chaetotaxy of tibiotarsus and foot complex: 22, foreleg, posterior view; 23, mid leg, posterior view; 24, hind leg, anterior view." httpUri="https://zenodo.org/record/279885/files/figure.png" pageId="2" pageNumber="3">Fig. 21</figureCitation>
). Tibiotarsus (
<figureCitation id="135ED5C0487CFFD2FBDAF835FBD29DCD" box="[1064,1150,2006,2030]" captionStart="FIGURES 19 – 24" captionStartId="5.[151,264,1940,1963]" captionTargetBox="[247,1333,193,1912]" captionTargetId="figure@5.[237,1355,193,1924]" captionTargetPageId="5" captionText="FIGURES 19 – 24. Arrhopalites macronyx sp. nov.: 19 – 21, chaetotaxy of trochanter and femur, anterior view: 19, foreleg; 20, mid leg; 21, hind leg; 22 – 24, chaetotaxy of tibiotarsus and foot complex: 22, foreleg, posterior view; 23, mid leg, posterior view; 24, hind leg, anterior view." httpUri="https://zenodo.org/record/279885/files/figure.png" pageId="2" pageNumber="3">Fig. 24</figureCitation>
): 3 setae FP and seta FSa present; whorl I with 9 setae without spine-like ones, whorls II–IV with 8 setae, whorl V with 7 setae (Vp absent) but sometimes incomplete (setae Vai and/or Vpi present or absent). Foot complex (
<figureCitation id="135ED5C04876FFD8FBB5FF5FFB0C9AF7" box="[1095,1184,188,212]" captionStart="FIGURES 19 – 24" captionStartId="5.[151,264,1940,1963]" captionTargetBox="[247,1333,193,1912]" captionTargetId="figure@5.[237,1355,193,1924]" captionTargetPageId="5" captionText="FIGURES 19 – 24. Arrhopalites macronyx sp. nov.: 19 – 21, chaetotaxy of trochanter and femur, anterior view: 19, foreleg; 20, mid leg; 21, hind leg; 22 – 24, chaetotaxy of tibiotarsus and foot complex: 22, foreleg, posterior view; 23, mid leg, posterior view; 24, hind leg, anterior view." httpUri="https://zenodo.org/record/279885/files/figure.png" pageId="8" pageNumber="9">Figs 24</figureCitation>
,
<figureCitation id="135ED5C04876FFD8FB5FFF5FFB609AF7" box="[1197,1228,188,212]" captionStart="FIGURES 32 – 36" captionStartId="7.[151,264,1916,1939]" captionTargetBox="[168,1414,201,1892]" captionTargetId="figure@7.[151,1436,193,1895]" captionTargetPageId="7" captionText="FIGURES 32 – 36. Arrhopalites macronyx sp. nov.: 32, distal parts of legs (a—foreleg, b—mid leg, c—hind leg, d—the longest fore claw of type series); 33, dens base, anterior view; 34, mucro (a—lateral view, b—dorso-lateral view, c—dorsal view); 35, chaetotaxy of central dorsal complex of great abdomen; 36, fragment of female Abd VI with winged circumanal setae." httpUri="https://zenodo.org/record/279887/files/figure.png" pageId="8" pageNumber="9">32</figureCitation>
c). Claw: long and thin, but a little shorter than in fore- and mid leg, only 2.3–2.5× shorter than tibiotarsus (
<quantity id="4C9D64A04876FFD8FB87FF3CFB699AD4" box="[1141,1221,223,247]" metricMagnitude="-2" metricUnit="m" metricValue="6.2484" pageId="8" pageNumber="9" unit="in" value="2.46">2.46 in</quantity>
<typeStatus id="54DE77E74876FFD8FB3EFF3CFA9F9AD4" box="[1228,1331,223,247]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
); without inner tooth, with 2 pairs of lateral teeth: very small subbasal and sharp thin subapical; without tunica. Empodium: broad, 1.3–1.5× shorter than claw; corner tooth present.
</paragraph>
<caption id="DF1A99CD487DFFD3FF65F860FD529DD5" httpUri="https://zenodo.org/record/279883/files/figure.png" pageId="3" pageNumber="4" targetBox="[186,1401,218,1903]" targetPageId="3">
<paragraph id="8BDAC945487DFFD3FF65F860FD529DD5" blockId="3.[151,1436,1923,2038]" pageId="3" pageNumber="4">
<emphasis id="B9111557487DFFD3FF65F860FEED9DBA" bold="true" box="[151,321,1923,1946]" pageId="3" pageNumber="4">FIGURES 1–14</emphasis>
.
<taxonomicName id="4C65B2C6487DFFD3FEBCF867FD9B9DB9" box="[334,567,1924,1946]" class="Entognatha" family="Arrhopalitidae" genus="Arrhopalites" kingdom="Animalia" order="Collembola" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="macronyx" status="sp. nov.">
<emphasis id="B9111557487DFFD3FEBCF867FD9B9DB9" box="[334,567,1924,1946]" italics="true" pageId="3" pageNumber="4">Arrhopalites macronyx</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A222A82C487DFFD3FDCCF867FD239DBA" box="[574,655,1924,1945]" pageId="3" pageNumber="4" rank="species">
<emphasis id="B9111557487DFFD3FDCCF867FD239DBA" bold="true" box="[574,655,1924,1945]" pageId="3" pageNumber="4">sp. nov.</emphasis>
</taxonomicNameLabel>
: 1, chaetotaxy of head, frontal view; 2, chaetotaxy of great abdomen, lateral view, male; 3–12, shape of setae: 3, ms1 of Abd VI; 4, mps3 of Abd VI; 5, seta of head vertex; 6, dorsal seta of mesothorax; 7, posterior dorsal seta of great abdomen; 8, dI-1 of great abdomen; 9, Ia of dens; 10, Ipe of dens; 11, appendices anales, lateral view; 12, appendices anales, dorsal view; 13–14, tenaculum.
</paragraph>
</caption>
<caption id="DF1A99CD487AFFD4FF65F85BFC709DCE" httpUri="https://zenodo.org/record/279884/files/figure.png" pageId="4" pageNumber="5" targetBox="[199,1384,213,1938]" targetPageId="4">
<paragraph id="8BDAC945487AFFD4FF65F85BFC709DCE" blockId="4.[151,1435,1976,2029]" pageId="4" pageNumber="5">
<emphasis id="B9111557487AFFD4FF65F85BFEF99DED" bold="true" box="[151,341,1976,1999]" pageId="4" pageNumber="5">FIGURES 15–18.</emphasis>
<taxonomicName id="4C65B2C6487AFFD4FEAEF85BFDEB9DEC" box="[348,583,1976,1999]" class="Entognatha" family="Arrhopalitidae" genus="Arrhopalites" kingdom="Animalia" order="Collembola" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="macronyx" status="sp. nov.">
<emphasis id="B9111557487AFFD4FEAEF85BFDEB9DEC" box="[348,583,1976,1999]" italics="true" pageId="4" pageNumber="5">Arrhopalites macronyx</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A222A82C487AFFD4FDBDF85AFD0C9DED" box="[591,672,1977,1998]" pageId="4" pageNumber="5" rank="species">
<emphasis id="B9111557487AFFD4FDBDF85AFD0C9DED" bold="true" box="[591,672,1977,1998]" pageId="4" pageNumber="5">sp. nov.</emphasis>
</taxonomicNameLabel>
: 15, chaetotaxy of Ant I–III, anterior view; 16, Ant IV, anterior view; 17, chaetotaxy of furca, posterior view; 18, chaetotaxy of female Abd VI, lateral view.
</paragraph>
</caption>
<caption id="DF1A99CD487BFFD5FF65F877FE4A9DCB" httpUri="https://zenodo.org/record/279885/files/figure.png" pageId="5" pageNumber="6" targetBox="[247,1333,193,1912]" targetPageId="5">
<paragraph id="8BDAC945487BFFD5FF65F877FE4A9DCB" blockId="5.[151,1436,1940,2024]" pageId="5" pageNumber="6">
<emphasis id="B9111557487BFFD5FF65F877FEF89D89" bold="true" box="[151,340,1940,1963]" pageId="5" pageNumber="6">FIGURES 19–24.</emphasis>
<taxonomicName id="4C65B2C6487BFFD5FEA9F876FDE99D88" box="[347,581,1941,1963]" class="Entognatha" family="Arrhopalitidae" genus="Arrhopalites" kingdom="Animalia" order="Collembola" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="macronyx" status="sp. nov.">
<emphasis id="B9111557487BFFD5FEA9F876FDE99D88" box="[347,581,1941,1963]" italics="true" pageId="5" pageNumber="6">Arrhopalites macronyx</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A222A82C487BFFD5FDBEF876FD319D89" box="[588,669,1941,1962]" pageId="5" pageNumber="6" rank="species">
<emphasis id="B9111557487BFFD5FDBEF876FD319D89" bold="true" box="[588,669,1941,1962]" pageId="5" pageNumber="6">sp. nov.</emphasis>
</taxonomicNameLabel>
: 19–21, chaetotaxy of trochanter and femur, anterior view: 19, foreleg; 20, mid leg; 21, hind leg; 22–24, chaetotaxy of tibiotarsus and foot complex: 22, foreleg, posterior view; 23, mid leg, posterior view; 24, hind leg, anterior view.
</paragraph>
</caption>
<caption id="DF1A99CD4878FFD6FF65F86FFB1B9DC3" httpUri="https://zenodo.org/record/279886/files/figure.png" pageId="6" pageNumber="7" targetBox="[252,1324,193,1910]" targetPageId="6">
<paragraph id="8BDAC9454878FFD6FF65F86FFB1B9DC3" blockId="6.[151,1436,1932,2016]" pageId="6" pageNumber="7">
<emphasis id="B91115574878FFD6FF65F86FFEF99D81" bold="true" box="[151,341,1932,1955]" pageId="6" pageNumber="7">FIGURES 25–31.</emphasis>
<taxonomicName id="4C65B2C64878FFD6FEAEF86EFDEA9D80" box="[348,582,1933,1955]" class="Entognatha" family="Arrhopalitidae" genus="Arrhopalites" kingdom="Animalia" order="Collembola" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="macronyx" status="sp. nov.">
<emphasis id="B91115574878FFD6FEAEF86EFDEA9D80" box="[348,582,1933,1955]" italics="true" pageId="6" pageNumber="7">Arrhopalites macronyx</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A222A82C4878FFD6FDBCF86EFD0C9D81" box="[590,672,1933,1954]" pageId="6" pageNumber="7" rank="species">
<emphasis id="B91115574878FFD6FDBCF86EFD0C9D81" bold="true" box="[590,672,1933,1954]" pageId="6" pageNumber="7">sp. nov.</emphasis>
</taxonomicNameLabel>
: 25, habitus and pigmentation, alcohol specimen; 26, habitus of holotype, on slide; 27, fragment of head dorsum with eye field; 28, chaetotaxy of trichobothrial complex; 29–30, Ant III: 29, entire segment; 30, fragment with antennal organ; 31, fragment of Ant IV with annulations between subsegments.
</paragraph>
</caption>
<caption id="DF1A99CD4879FFD7FF65F89FFAE49DF3" httpUri="https://zenodo.org/record/279887/files/figure.png" pageId="7" pageNumber="8" targetBox="[168,1414,201,1892]" targetPageId="7">
<paragraph id="8BDAC9454879FFD7FF65F89FFAE49DF3" blockId="7.[151,1436,1916,2000]" pageId="7" pageNumber="8">
<emphasis id="B91115574879FFD7FF65F89FFEFF9DB1" bold="true" box="[151,339,1916,1939]" pageId="7" pageNumber="8">FIGURES 32–36.</emphasis>
<taxonomicName id="4C65B2C64879FFD7FEAAF89EFDEC9DB0" box="[344,576,1917,1939]" class="Entognatha" family="Arrhopalitidae" genus="Arrhopalites" kingdom="Animalia" order="Collembola" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="macronyx" status="sp. nov.">
<emphasis id="B91115574879FFD7FEAAF89EFDEC9DB0" box="[344,576,1917,1939]" italics="true" pageId="7" pageNumber="8">Arrhopalites macronyx</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A222A82C4879FFD7FDB4F89EFD3A9DB1" box="[582,662,1917,1938]" pageId="7" pageNumber="8" rank="species">
<emphasis id="B91115574879FFD7FDB4F89EFD3A9DB1" bold="true" box="[582,662,1917,1938]" pageId="7" pageNumber="8">sp. nov.</emphasis>
</taxonomicNameLabel>
: 32, distal parts of legs (a—foreleg, b—mid leg, c—hind leg, d—the longest fore claw of type series); 33, dens base, anterior view; 34, mucro (a—lateral view, b—dorso-lateral view, c—dorsal view); 35, chaetotaxy of central dorsal complex of great abdomen; 36, fragment of female Abd VI with winged circumanal setae.
</paragraph>
</caption>
<paragraph id="8BDAC9454876FFD8FF37FEAEFDA39BA4" blockId="8.[151,1437,151,2017]" pageId="8" pageNumber="9">
Mean lengths ratio of tibiotarsi I: II: III = 1: 0.97: 1.08 (in
<typeStatus id="54DE77E74876FFD8FC8CFEAFFC4C9B47" box="[894,992,332,356]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
1: 0.95: 1.08). Mean ratio head: tibiotarsus I = 1.5 (
<quantity id="4C9D64A04876FFD8FEB4FE8CFE3A9BA4" box="[326,406,367,391]" metricMagnitude="-2" metricUnit="m" metricValue="3.7592" pageId="8" pageNumber="9" unit="in" value="1.48">1.48 in</quantity>
<typeStatus id="54DE77E74876FFD8FE6EFE8CFDAF9BA4" box="[412,515,367,391]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
).
</paragraph>
<paragraph id="8BDAC9454876FFD8FF37FE77FCC89BD7" blockId="8.[151,1437,151,2017]" pageId="8" pageNumber="9">
Ventral tube with 1 + 1 subapical setulae. Tenaculum (
<figureCitation id="135ED5C04876FFD8FCDDFE77FC189B8F" box="[815,948,404,428]" captionStart="FIGURES 1 – 14" captionStartId="3.[151,264,1923,1946]" captionTargetBox="[186,1401,218,1903]" captionTargetId="figure@3.[174,1417,190,1909]" captionTargetPageId="3" captionText="FIGURES 1 – 14. Arrhopalites macronyx sp. nov.: 1, chaetotaxy of head, frontal view; 2, chaetotaxy of great abdomen, lateral view, male; 3 – 12, shape of setae: 3, ms 1 of Abd VI; 4, mps 3 of Abd VI; 5, seta of head vertex; 6, dorsal seta of mesothorax; 7, posterior dorsal seta of great abdomen; 8, dI- 1 of great abdomen; 9, Ia of dens; 10, Ipe of dens; 11, appendices anales, lateral view; 12, appendices anales, dorsal view; 13 – 14, tenaculum." httpUri="https://zenodo.org/record/279883/files/figure.png" pageId="8" pageNumber="9">Figs 13, 14</figureCitation>
): each ramus with 3 sclerotic teeth and not sclerotized basal process; anterior lobe usually with 1 (as in
<typeStatus id="54DE77E74876FFD8FCB5FE54FC079BEC" box="[839,939,439,463]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
) or 2 (in
<specimenCount id="9D6302CC4876FFD8FBEBFE54FB0A9BEC" box="[1049,1190,439,463]" pageId="8" pageNumber="9" type="generic">3 specimens</specimenCount>
) apical setulae; tip of posterior lobe is approximately on one level with tip of anterior.
</paragraph>
<paragraph id="8BDAC9454876FFD8FF37FDE3FCF698EF" blockId="8.[151,1437,151,2017]" pageId="8" pageNumber="9">
Furca (
<figureCitation id="135ED5C04876FFD8FEE6FE1CFECB9834" box="[276,359,511,535]" captionStart="FIGURES 15 – 18" captionStartId="4.[151,264,1976,1999]" captionTargetBox="[199,1384,213,1938]" captionTargetId="figure@4.[186,1401,193,1955]" captionTargetPageId="4" captionText="FIGURES 15 – 18. Arrhopalites macronyx sp. nov.: 15, chaetotaxy of Ant I – III, anterior view; 16, Ant IV, anterior view; 17, chaetotaxy of furca, posterior view; 18, chaetotaxy of female Abd VI, lateral view." httpUri="https://zenodo.org/record/279884/files/figure.png" pageId="8" pageNumber="9">Fig. 17</figureCitation>
): manubrium with 5 + 5 setae on posterior side, seta p2 thin sensilla-like, seta p3 thin and short. Dens: with distinct subbasal broadening on interior side (
<figureCitation id="135ED5C04876FFD8FCE8FDC7FCC2981F" box="[794,878,548,572]" captionStart="FIGURES 32 – 36" captionStartId="7.[151,264,1916,1939]" captionTargetBox="[168,1414,201,1892]" captionTargetId="figure@7.[151,1436,193,1895]" captionTargetPageId="7" captionText="FIGURES 32 – 36. Arrhopalites macronyx sp. nov.: 32, distal parts of legs (a—foreleg, b—mid leg, c—hind leg, d—the longest fore claw of type series); 33, dens base, anterior view; 34, mucro (a—lateral view, b—dorso-lateral view, c—dorsal view); 35, chaetotaxy of central dorsal complex of great abdomen; 36, fragment of female Abd VI with winged circumanal setae." httpUri="https://zenodo.org/record/279887/files/figure.png" pageId="8" pageNumber="9">Fig. 33</figureCitation>
); anterior side with 3, 2, 1, 1, 1 setae, Ia– IVa, Ba and IIae are heavy (
<figureCitation id="135ED5C04876FFD8FE8BFDA4FE6F987C" box="[377,451,583,607]" captionStart="FIGURES 1 – 14" captionStartId="3.[151,264,1923,1946]" captionTargetBox="[186,1401,218,1903]" captionTargetId="figure@3.[174,1417,190,1909]" captionTargetPageId="3" captionText="FIGURES 1 – 14. Arrhopalites macronyx sp. nov.: 1, chaetotaxy of head, frontal view; 2, chaetotaxy of great abdomen, lateral view, male; 3 – 12, shape of setae: 3, ms 1 of Abd VI; 4, mps 3 of Abd VI; 5, seta of head vertex; 6, dorsal seta of mesothorax; 7, posterior dorsal seta of great abdomen; 8, dI- 1 of great abdomen; 9, Ia of dens; 10, Ipe of dens; 11, appendices anales, lateral view; 12, appendices anales, dorsal view; 13 – 14, tenaculum." httpUri="https://zenodo.org/record/279883/files/figure.png" pageId="8" pageNumber="9">Figs 9</figureCitation>
,
<figureCitation id="135ED5C04876FFD8FE22FDA4FE43987C" box="[464,495,583,607]" captionStart="FIGURES 15 – 18" captionStartId="4.[151,264,1976,1999]" captionTargetBox="[199,1384,213,1938]" captionTargetId="figure@4.[186,1401,193,1955]" captionTargetPageId="4" captionText="FIGURES 15 – 18. Arrhopalites macronyx sp. nov.: 15, chaetotaxy of Ant I – III, anterior view; 16, Ant IV, anterior view; 17, chaetotaxy of furca, posterior view; 18, chaetotaxy of female Abd VI, lateral view." httpUri="https://zenodo.org/record/279884/files/figure.png" pageId="8" pageNumber="9">17</figureCitation>
). Posterior side without distinct spines, but setae Ie, Ipe and Ii are thickened at the basis and somewhat spine-like (
<figureCitation id="135ED5C04876FFD8FDF2FD8FFDF498A7" box="[512,600,620,644]" captionStart="FIGURES 1 – 14" captionStartId="3.[151,264,1923,1946]" captionTargetBox="[186,1401,218,1903]" captionTargetId="figure@3.[174,1417,190,1909]" captionTargetPageId="3" captionText="FIGURES 1 – 14. Arrhopalites macronyx sp. nov.: 1, chaetotaxy of head, frontal view; 2, chaetotaxy of great abdomen, lateral view, male; 3 – 12, shape of setae: 3, ms 1 of Abd VI; 4, mps 3 of Abd VI; 5, seta of head vertex; 6, dorsal seta of mesothorax; 7, posterior dorsal seta of great abdomen; 8, dI- 1 of great abdomen; 9, Ia of dens; 10, Ipe of dens; 11, appendices anales, lateral view; 12, appendices anales, dorsal view; 13 – 14, tenaculum." httpUri="https://zenodo.org/record/279883/files/figure.png" pageId="8" pageNumber="9">Figs 10</figureCitation>
,
<figureCitation id="135ED5C04876FFD8FD97FD8FFD2898A7" box="[613,644,620,644]" captionStart="FIGURES 15 – 18" captionStartId="4.[151,264,1976,1999]" captionTargetBox="[199,1384,213,1938]" captionTargetId="figure@4.[186,1401,193,1955]" captionTargetPageId="4" captionText="FIGURES 15 – 18. Arrhopalites macronyx sp. nov.: 15, chaetotaxy of Ant I – III, anterior view; 16, Ant IV, anterior view; 17, chaetotaxy of furca, posterior view; 18, chaetotaxy of female Abd VI, lateral view." httpUri="https://zenodo.org/record/279884/files/figure.png" pageId="8" pageNumber="9">17</figureCitation>
). Mucro (
<figureCitation id="135ED5C04876FFD8FD06FD8FFCE498A7" box="[756,840,620,644]" captionStart="FIGURES 32 – 36" captionStartId="7.[151,264,1916,1939]" captionTargetBox="[168,1414,201,1892]" captionTargetId="figure@7.[151,1436,193,1895]" captionTargetPageId="7" captionText="FIGURES 32 – 36. Arrhopalites macronyx sp. nov.: 32, distal parts of legs (a—foreleg, b—mid leg, c—hind leg, d—the longest fore claw of type series); 33, dens base, anterior view; 34, mucro (a—lateral view, b—dorso-lateral view, c—dorsal view); 35, chaetotaxy of central dorsal complex of great abdomen; 36, fragment of female Abd VI with winged circumanal setae." httpUri="https://zenodo.org/record/279887/files/figure.png" pageId="8" pageNumber="9">Fig. 34</figureCitation>
a–c): posterior lamellae serrated with teeth separated from each other or often looking accreted together; anterior lamella well-developed; tip rounded but not much broadened. Dens 2.1–2.3× as long as mucro (
<quantity id="4C9D64A04876FFD8FD63FD57FD4C98EF" box="[657,736,692,716]" metricMagnitude="-2" metricUnit="m" metricValue="5.6642" pageId="8" pageNumber="9" unit="in" value="2.23">2.23 in</quantity>
<typeStatus id="54DE77E74876FFD8FD15FD57FCE298EF" box="[743,846,692,716]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
).
</paragraph>
<paragraph id="8BDAC9454876FFD8FF37FD34FACF99E9" blockId="8.[151,1437,151,2017]" pageId="8" pageNumber="9">
Great abdomen (
<figureCitation id="135ED5C04876FFD8FE71FD34FE6598CC" box="[387,457,727,751]" captionStart="FIGURES 1 – 14" captionStartId="3.[151,264,1923,1946]" captionTargetBox="[186,1401,218,1903]" captionTargetId="figure@3.[174,1417,190,1909]" captionTargetPageId="3" captionText="FIGURES 1 – 14. Arrhopalites macronyx sp. nov.: 1, chaetotaxy of head, frontal view; 2, chaetotaxy of great abdomen, lateral view, male; 3 – 12, shape of setae: 3, ms 1 of Abd VI; 4, mps 3 of Abd VI; 5, seta of head vertex; 6, dorsal seta of mesothorax; 7, posterior dorsal seta of great abdomen; 8, dI- 1 of great abdomen; 9, Ia of dens; 10, Ipe of dens; 11, appendices anales, lateral view; 12, appendices anales, dorsal view; 13 – 14, tenaculum." httpUri="https://zenodo.org/record/279883/files/figure.png" pageId="8" pageNumber="9">Fig. 2</figureCitation>
): mesothorax with slender and curved or S-shaped dorsal setae (
<figureCitation id="135ED5C04876FFD8FB55FD34FB4098CC" box="[1191,1260,727,751]" captionStart="FIGURES 1 – 14" captionStartId="3.[151,264,1923,1946]" captionTargetBox="[186,1401,218,1903]" captionTargetId="figure@3.[174,1417,190,1909]" captionTargetPageId="3" captionText="FIGURES 1 – 14. Arrhopalites macronyx sp. nov.: 1, chaetotaxy of head, frontal view; 2, chaetotaxy of great abdomen, lateral view, male; 3 – 12, shape of setae: 3, ms 1 of Abd VI; 4, mps 3 of Abd VI; 5, seta of head vertex; 6, dorsal seta of mesothorax; 7, posterior dorsal seta of great abdomen; 8, dI- 1 of great abdomen; 9, Ia of dens; 10, Ipe of dens; 11, appendices anales, lateral view; 12, appendices anales, dorsal view; 13 – 14, tenaculum." httpUri="https://zenodo.org/record/279883/files/figure.png" pageId="8" pageNumber="9">Fig. 6</figureCitation>
). Trichobothria ABC form about right angle (85o–101o; 89o in
<typeStatus id="54DE77E74876FFD8FD58FD1CFCA29934" box="[682,782,767,791]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
) and AB is about 1.4× (1.2–1.6) longer than BC (
<quantity id="4C9D64A04876FFD8FAB9FD1CFA309934" box="[1355,1436,767,791]" metricMagnitude="-2" metricUnit="m" metricValue="3.175" pageId="8" pageNumber="9" unit="in" value="1.25">1.25 in</quantity>
<typeStatus id="54DE77E74876FFD8FF65FCC1FF529919" box="[151,254,802,826]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
). Single p seta of p-row of Abd I is located above the level of trichobothrium B; seta b1 lies on line BC, closer to C; seta c1 small (~20 µm) and lies on the level and seta c2—below the level of trichobothrium C (
<figureCitation id="135ED5C04876FFD8FACEFCA4FA23997C" box="[1340,1423,839,863]" captionStart="FIGURES 25 – 31" captionStartId="6.[151,264,1932,1955]" captionTargetBox="[252,1324,193,1910]" captionTargetId="figure@6.[249,1338,193,1911]" captionTargetPageId="6" captionText="FIGURES 25 – 31. Arrhopalites macronyx sp. nov.: 25, habitus and pigmentation, alcohol specimen; 26, habitus of holotype, on slide; 27, fragment of head dorsum with eye field; 28, chaetotaxy of trichobothrial complex; 29 – 30, Ant III: 29, entire segment; 30, fragment with antennal organ; 31, fragment of Ant IV with annulations between subsegments." httpUri="https://zenodo.org/record/279886/files/figure.png" pageId="8" pageNumber="9">Fig. 28</figureCitation>
). Posterior lateral complex with 2 + 3 and furca base complex with 8 setae. Central dorsal complex with three setae of which seta 3 is more than 3× shorter than two others (
<figureCitation id="135ED5C04876FFD8FCE3FC6CFCCA9984" box="[785,870,911,935]" captionStart="FIGURES 32 – 36" captionStartId="7.[151,264,1916,1939]" captionTargetBox="[168,1414,201,1892]" captionTargetId="figure@7.[151,1436,193,1895]" captionTargetPageId="7" captionText="FIGURES 32 – 36. Arrhopalites macronyx sp. nov.: 32, distal parts of legs (a—foreleg, b—mid leg, c—hind leg, d—the longest fore claw of type series); 33, dens base, anterior view; 34, mucro (a—lateral view, b—dorso-lateral view, c—dorsal view); 35, chaetotaxy of central dorsal complex of great abdomen; 36, fragment of female Abd VI with winged circumanal setae." httpUri="https://zenodo.org/record/279887/files/figure.png" pageId="8" pageNumber="9">Fig. 35</figureCitation>
). Posterior dorsal complex with about 15 setae the longest of which is about 1.5× shorter than hind claw (
<figureCitation id="135ED5C04876FFD8FD0BFC51FCF299E9" box="[761,862,946,970]" captionStart="FIGURES 1 – 14" captionStartId="3.[151,264,1923,1946]" captionTargetBox="[186,1401,218,1903]" captionTargetId="figure@3.[174,1417,190,1909]" captionTargetPageId="3" captionText="FIGURES 1 – 14. Arrhopalites macronyx sp. nov.: 1, chaetotaxy of head, frontal view; 2, chaetotaxy of great abdomen, lateral view, male; 3 – 12, shape of setae: 3, ms 1 of Abd VI; 4, mps 3 of Abd VI; 5, seta of head vertex; 6, dorsal seta of mesothorax; 7, posterior dorsal seta of great abdomen; 8, dI- 1 of great abdomen; 9, Ia of dens; 10, Ipe of dens; 11, appendices anales, lateral view; 12, appendices anales, dorsal view; 13 – 14, tenaculum." httpUri="https://zenodo.org/record/279883/files/figure.png" pageId="8" pageNumber="9">Figs 7, 8</figureCitation>
). Ventral complex with 1 or 2 (rarely 0) setae.
</paragraph>
<paragraph id="8BDAC9454876FFD8FF37FC34FC989E31" blockId="8.[151,1437,151,2017]" pageId="8" pageNumber="9">
Fifth abdominal segment with 2 setae and trichobothrium D in row a, and 2 setae in row p (
<figureCitation id="135ED5C04876FFD8FB4AFC34FB5099CC" box="[1208,1276,983,1007]" captionStart="FIGURES 1 – 14" captionStartId="3.[151,264,1923,1946]" captionTargetBox="[186,1401,218,1903]" captionTargetId="figure@3.[174,1417,190,1909]" captionTargetPageId="3" captionText="FIGURES 1 – 14. Arrhopalites macronyx sp. nov.: 1, chaetotaxy of head, frontal view; 2, chaetotaxy of great abdomen, lateral view, male; 3 – 12, shape of setae: 3, ms 1 of Abd VI; 4, mps 3 of Abd VI; 5, seta of head vertex; 6, dorsal seta of mesothorax; 7, posterior dorsal seta of great abdomen; 8, dI- 1 of great abdomen; 9, Ia of dens; 10, Ipe of dens; 11, appendices anales, lateral view; 12, appendices anales, dorsal view; 13 – 14, tenaculum." httpUri="https://zenodo.org/record/279883/files/figure.png" pageId="8" pageNumber="9">Fig. 2</figureCitation>
). Genital field with 2 + 2 setae close to anterior margin of genital opening.
</paragraph>
<paragraph id="8BDAC9454876FFD8FF37FBFCFB299EE4" blockId="8.[151,1437,151,2017]" pageId="8" pageNumber="9">
Sixth abdominal segment (
<figureCitation id="135ED5C04876FFD8FE04FBFCFDE79E14" box="[502,587,1055,1079]" captionStart="FIGURES 15 – 18" captionStartId="4.[151,264,1976,1999]" captionTargetBox="[199,1384,213,1938]" captionTargetId="figure@4.[186,1401,193,1955]" captionTargetPageId="4" captionText="FIGURES 15 – 18. Arrhopalites macronyx sp. nov.: 15, chaetotaxy of Ant I – III, anterior view; 16, Ant IV, anterior view; 17, chaetotaxy of furca, posterior view; 18, chaetotaxy of female Abd VI, lateral view." httpUri="https://zenodo.org/record/279884/files/figure.png" pageId="8" pageNumber="9">Fig. 18</figureCitation>
): without cuticular spines; circumanal setae (
<figureCitation id="135ED5C04876FFD8FBA2FBFCFB149E14" box="[1104,1208,1055,1079]" captionStart="FIGURES 1 – 14" captionStartId="3.[151,264,1923,1946]" captionTargetBox="[186,1401,218,1903]" captionTargetId="figure@3.[174,1417,190,1909]" captionTargetPageId="3" captionText="FIGURES 1 – 14. Arrhopalites macronyx sp. nov.: 1, chaetotaxy of head, frontal view; 2, chaetotaxy of great abdomen, lateral view, male; 3 – 12, shape of setae: 3, ms 1 of Abd VI; 4, mps 3 of Abd VI; 5, seta of head vertex; 6, dorsal seta of mesothorax; 7, posterior dorsal seta of great abdomen; 8, dI- 1 of great abdomen; 9, Ia of dens; 10, Ipe of dens; 11, appendices anales, lateral view; 12, appendices anales, dorsal view; 13 – 14, tenaculum." httpUri="https://zenodo.org/record/279883/files/figure.png" pageId="8" pageNumber="9">Figs 3, 4</figureCitation>
,
<figureCitation id="135ED5C04876FFD8FB34FBFCFB489E14" box="[1222,1252,1055,1079]" captionStart="FIGURES 15 – 18" captionStartId="4.[151,264,1976,1999]" captionTargetBox="[199,1384,213,1938]" captionTargetId="figure@4.[186,1401,193,1955]" captionTargetPageId="4" captionText="FIGURES 15 – 18. Arrhopalites macronyx sp. nov.: 15, chaetotaxy of Ant I – III, anterior view; 16, Ant IV, anterior view; 17, chaetotaxy of furca, posterior view; 18, chaetotaxy of female Abd VI, lateral view." httpUri="https://zenodo.org/record/279884/files/figure.png" pageId="8" pageNumber="9">18</figureCitation>
,
<figureCitation id="135ED5C04876FFD8FB00FBFCFABE9E14" box="[1266,1298,1055,1079]" captionStart="FIGURES 32 – 36" captionStartId="7.[151,264,1916,1939]" captionTargetBox="[168,1414,201,1892]" captionTargetId="figure@7.[151,1436,193,1895]" captionTargetPageId="7" captionText="FIGURES 32 – 36. Arrhopalites macronyx sp. nov.: 32, distal parts of legs (a—foreleg, b—mid leg, c—hind leg, d—the longest fore claw of type series); 33, dens base, anterior view; 34, mucro (a—lateral view, b—dorso-lateral view, c—dorsal view); 35, chaetotaxy of central dorsal complex of great abdomen; 36, fragment of female Abd VI with winged circumanal setae." httpUri="https://zenodo.org/record/279887/files/figure.png" pageId="8" pageNumber="9">36</figureCitation>
): broadened without serration (ms1, mpi3), winged and serrated subbasally (usually mps1–mps3 and mpi1–mpi2) or simple (mpi4). Seta ms5 on dorsal valve is absent. Dorsal valve bears 10 + 2 axial + 10 setae, each of lateral valve bears 17 setae (totally 56 setae on Abd VI). Appendices anales (44.8 µm in
<typeStatus id="54DE77E74876FFD8FC73FB69FC489E81" box="[897,996,1162,1186]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
) rod-like, smooth, gradually narrowed, curved in distal half, with acuminated tip, inserted into semiglobular papilla (
<figureCitation id="135ED5C04876FFD8FC05FB4CFBDB9EE4" box="[1015,1143,1199,1223]" captionStart="FIGURES 1 – 14" captionStartId="3.[151,264,1923,1946]" captionTargetBox="[186,1401,218,1903]" captionTargetId="figure@3.[174,1417,190,1909]" captionTargetPageId="3" captionText="FIGURES 1 – 14. Arrhopalites macronyx sp. nov.: 1, chaetotaxy of head, frontal view; 2, chaetotaxy of great abdomen, lateral view, male; 3 – 12, shape of setae: 3, ms 1 of Abd VI; 4, mps 3 of Abd VI; 5, seta of head vertex; 6, dorsal seta of mesothorax; 7, posterior dorsal seta of great abdomen; 8, dI- 1 of great abdomen; 9, Ia of dens; 10, Ipe of dens; 11, appendices anales, lateral view; 12, appendices anales, dorsal view; 13 – 14, tenaculum." httpUri="https://zenodo.org/record/279883/files/figure.png" pageId="8" pageNumber="9">Figs 11, 12</figureCitation>
).
</paragraph>
<paragraph id="8BDAC9454876FFD8FF37FB30FCB09F74" blockId="8.[151,1437,151,2017]" pageId="8" pageNumber="9">
<emphasis id="B91115574876FFD8FF37FB30FEA79EC8" bold="true" box="[197,267,1235,1259]" pageId="8" pageNumber="9">Male:</emphasis>
smaller than females, body length is about
<quantity id="4C9D64A04876FFD8FD01FB31FC989EC9" box="[755,820,1234,1258]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="9" unit="mm" value="1.0">1 mm</quantity>
or less (
<tableCitation id="C6E7FCFE4876FFD8FC60FB31FC4A9EC9" box="[914,998,1234,1258]" captionStart="TABLE 1" captionStartId="2.[151,235,153,174]" captionTargetBox="[151,1436,275,1290]" captionTargetPageId="2" captionText="TABLE 1. Lengths (in µm) for body parts of types of Arrhopalites macronyx sp. nov. and Troglopalites stygios gen. nov. sp. nov. (all measurements, except total body length, are given only for females)." httpUri="http://table.plazi.org/id/DF1A99CD487CFFD2FF65FF7AFC559AEC" pageId="8" pageNumber="9" tableUuid="DF1A99CD487CFFD2FF65FF7AFC559AEC">Table 1</tableCitation>
). Chaetotaxy is the same as in females except Abd VI (
<figureCitation id="135ED5C04876FFD8FEB9FB14FE239F2C" box="[331,399,1271,1295]" captionStart="FIGURES 1 – 14" captionStartId="3.[151,264,1923,1946]" captionTargetBox="[186,1401,218,1903]" captionTargetId="figure@3.[174,1417,190,1909]" captionTargetPageId="3" captionText="FIGURES 1 – 14. Arrhopalites macronyx sp. nov.: 1, chaetotaxy of head, frontal view; 2, chaetotaxy of great abdomen, lateral view, male; 3 – 12, shape of setae: 3, ms 1 of Abd VI; 4, mps 3 of Abd VI; 5, seta of head vertex; 6, dorsal seta of mesothorax; 7, posterior dorsal seta of great abdomen; 8, dI- 1 of great abdomen; 9, Ia of dens; 10, Ipe of dens; 11, appendices anales, lateral view; 12, appendices anales, dorsal view; 13 – 14, tenaculum." httpUri="https://zenodo.org/record/279883/files/figure.png" pageId="8" pageNumber="9">Fig. 2</figureCitation>
) with no broadened or winged setae, with 7 + 2 axial + 7 setae on upper anal valve, 13 setae on each of lateral valves (totally 42 setae on Abd VI versus
<quantity id="4C9D64A04876FFD8FCB3FAF9FCD79F11" box="[833,891,1306,1330]" metricMagnitude="0" metricUnit="m" metricValue="1.4224" pageId="8" pageNumber="9" unit="in" value="56.0">56 in</quantity>
female) and different chaetotaxy around genital opening on Abd V (about 10 setulae versus
<quantity id="4C9D64A04876FFD8FD70FADCFD019F74" box="[642,685,1343,1367]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="8" pageNumber="9" unit="in" value="4.0">4 in</quantity>
females).
</paragraph>
<paragraph id="8BDAC9454876FFD8FF37FA81FD849FC4" blockId="8.[151,1437,151,2017]" pageId="8" pageNumber="9">
<emphasis id="B91115574876FFD8FF37FA81FEE69F58" bold="true" box="[197,330,1378,1403]" pageId="8" pageNumber="9">Variability.</emphasis>
Ventral complex of great abdomen includes 1 or 2 (rarely asymmetrically 0) setae. Seta Vpe of dens present or absent. Tenaculum with 1 or 2 setae. Setae Vai and/or Vpi on tibiotarsi sometimes absent. Corner tooth of empodium III in
<specimenCount id="9D6302CC4876FFD8FE41FA49FD9E9FE1" box="[435,562,1450,1474]" pageId="8" pageNumber="9" type="generic">1 specimen</specimenCount>
absent. Two males possess 7 + 2 axial + 6 setae on dorsal anal valve with seta ms2 asymmetrically present/absent.
</paragraph>
<paragraph id="8BDAC9454876FFD8FF37FA11FA3A9CC6" blockId="8.[151,1437,151,2017]" pageId="8" pageNumber="9">
<emphasis id="B91115574876FFD8FF37FA11FE539C28" bold="true" box="[197,511,1522,1547]" pageId="8" pageNumber="9">Bionomy and distribution.</emphasis>
Specimens of the new species were sampled about
<quantity id="4C9D64A04876FFD8FBCCFA11FB289C29" box="[1086,1156,1522,1546]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="9" unit="m" value="100.0">100 m</quantity>
from the entrance exclusively on the water surface of small subterranean flow lake and on wet speleothems just near the lake. The water and air temperature was 9.9o C. Terrestrial hand sampling and Barber’s traps in the same cave provided another species of
<taxonomicName id="4C65B2C64876FFD8FF1EF982FE239C5A" box="[236,399,1633,1657]" class="Entognatha" family="Arrhopalitidae" kingdom="Animalia" order="Collembola" pageId="8" pageNumber="9" phylum="Arthropoda" rank="family">Arrhopalitidae</taxonomicName>
but not
<taxonomicName id="4C65B2C64876FFD8FE01F982FD2D9C5A" box="[499,641,1633,1657]" class="Entognatha" family="Arrhopalitidae" genus="Arrhopalites" kingdom="Animalia" order="Collembola" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="macronyx" status="sp. nov.">
<emphasis id="B91115574876FFD8FE01F982FD2D9C5A" box="[499,641,1633,1657]" italics="true" pageId="8" pageNumber="9">A. macronyx</emphasis>
</taxonomicName>
<emphasis id="B91115574876FFD8FD79F981FD4A9C59" bold="true" box="[651,742,1634,1658]" pageId="8" pageNumber="9">
<taxonomicNameLabel id="A222A82C4876FFD8FD79F981FD4A9C59" box="[651,742,1634,1658]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
This fact, together with morphological characters (long thin claws and mucro with partly accreted teeth and well-developed anterior lamella) supports an opinion about epineustonic and hygropetric mode of life of the new species in subterranean ecosystem. The species was found only in one of 45 sampled caves of the Western Caucasus and most likely represents a narrow troglobiont endemic.
</paragraph>
</subSubSection>
<subSubSection id="C37F9ACE4876FFD8FF37F911FAD89D0E" pageId="8" pageNumber="9" type="etymology">
<paragraph id="8BDAC9454876FFD8FF37F911FAD89D0E" blockId="8.[151,1437,151,2017]" pageId="8" pageNumber="9">
<emphasis id="B91115574876FFD8FF37F911FEE09D29" bold="true" box="[197,332,1778,1802]" pageId="8" pageNumber="9">Etymology.</emphasis>
“
<taxonomicName id="4C65B2C64876FFD8FE92F911FE619D2A" box="[352,461,1778,1801]" class="Entognatha" family="Arrhopalitidae" genus="Arrhopalites" kingdom="Animalia" order="Collembola" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="macronyx">
<emphasis id="B91115574876FFD8FE92F911FE619D2A" box="[352,461,1778,1801]" italics="true" pageId="8" pageNumber="9">macronyx</emphasis>
</taxonomicName>
” from Greek ‘μάĸρος’—big, long and ‘όνυξ’—claw. The name denotes extraordinary long claws (up to 154 µm:
<figureCitation id="135ED5C04876FFD8FE37F8F6FDBA9D0E" box="[453,534,1813,1837]" captionStart="FIGURES 32 – 36" captionStartId="7.[151,264,1916,1939]" captionTargetBox="[168,1414,201,1892]" captionTargetId="figure@7.[151,1436,193,1895]" captionTargetPageId="7" captionText="FIGURES 32 – 36. Arrhopalites macronyx sp. nov.: 32, distal parts of legs (a—foreleg, b—mid leg, c—hind leg, d—the longest fore claw of type series); 33, dens base, anterior view; 34, mucro (a—lateral view, b—dorso-lateral view, c—dorsal view); 35, chaetotaxy of central dorsal complex of great abdomen; 36, fragment of female Abd VI with winged circumanal setae." httpUri="https://zenodo.org/record/279887/files/figure.png" pageId="8" pageNumber="9">Fig. 32</figureCitation>
d), possibly the longest among the known species of the genus
<taxonomicName id="4C65B2C64876FFD8FB24F8F5FACF9D0D" box="[1238,1379,1814,1838]" class="Entognatha" family="Arrhopalitidae" genus="Arrhopalites" kingdom="Animalia" order="Collembola" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B91115574876FFD8FB24F8F5FACF9D0D" box="[1238,1379,1814,1838]" italics="true" pageId="8" pageNumber="9">Arrhopalites</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C37F9ACE4876FFD9FF37F8D9FCA19E73" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="discussion">
<paragraph id="8BDAC9454876FFD9FF37F8D9FA9E9BA4" blockId="8.[151,1437,151,2017]" lastBlockId="9.[151,1436,151,392]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
<emphasis id="B91115574876FFD8FF37F8D9FE959D71" bold="true" box="[197,313,1850,1874]" pageId="8" pageNumber="9">Remarks.</emphasis>
<taxonomicName id="4C65B2C64876FFD8FECDF8DAFE679D72" box="[319,459,1849,1873]" class="Entognatha" family="Arrhopalitidae" genus="Arrhopalites" kingdom="Animalia" order="Collembola" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="macronyx" status="sp. nov.">
<emphasis id="B91115574876FFD8FECDF8DAFE679D72" box="[319,459,1849,1873]" italics="true" pageId="8" pageNumber="9">A. macronyx</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A222A82C4876FFD8FE20F8D9FD879D71" box="[466,555,1850,1874]" pageId="8" pageNumber="9" rank="species">
<emphasis id="B91115574876FFD8FE20F8D9FD879D71" bold="true" box="[466,555,1850,1874]" pageId="8" pageNumber="9">sp. nov.</emphasis>
</taxonomicNameLabel>
can be compared with species of
<taxonomicName id="4C65B2C64876FFD8FC56F8DAFB9D9D72" box="[932,1073,1849,1873]" class="Entognatha" family="Arrhopalitidae" genus="Arrhopalites" kingdom="Animalia" order="Collembola" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B91115574876FFD8FC56F8DAFB9D9D72" box="[932,1073,1849,1873]" italics="true" pageId="8" pageNumber="9">Arrhopalites</emphasis>
</taxonomicName>
without cuticular spines on Abd VI, with anterior dens formula 3, 2, 1, 1, 1 and without strong anterior distal spine on dens:
<taxonomicName id="4C65B2C64876FFD8FB49F8BDFEB19DBA" authority="Park &amp; Kang, 2007" authorityName="Park &amp; Kang" authorityYear="2007" class="Entognatha" family="Arrhopalitidae" genus="Arrhopalites" kingdom="Animalia" order="Collembola" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="coreanus">
<emphasis id="B91115574876FFD8FB49F8BDFAED9D55" box="[1211,1345,1886,1910]" italics="true" pageId="8" pageNumber="9">A. coreanus</emphasis>
<bibRefCitation id="EFF4B4B44876FFD8FAB8F8BEFEB19DBA" author="Park" pageId="8" pageNumber="9" refString="Park, K. - H. &amp; Kang, J. - S. (2007) Three new species of Arrhopalites (Collembola, Symphypleona, Arrhopalitidae) from Korea. Entomological Research, 37, 157 - 162." type="journal article" year="2007">Park &amp; Kang, 2007</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C65B2C64876FFD8FEDAF862FDDD9DBA" authority="Park &amp; Kang, 2007" authorityName="Park &amp; Kang" authorityYear="2007" box="[296,625,1921,1946]" class="Entognatha" family="Arrhopalitidae" genus="Arrhopalites" kingdom="Animalia" order="Collembola" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="B91115574876FFD8FEDAF862FE279DBA" box="[296,395,1921,1945]" italics="true" pageId="8" pageNumber="9">A. minor</emphasis>
<bibRefCitation id="EFF4B4B44876FFD8FE61F862FDDD9DBA" author="Park" box="[403,625,1921,1946]" pageId="8" pageNumber="9" refString="Park, K. - H. &amp; Kang, J. - S. (2007) Three new species of Arrhopalites (Collembola, Symphypleona, Arrhopalitidae) from Korea. Entomological Research, 37, 157 - 162." type="journal article" year="2007">Park &amp; Kang, 2007</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C65B2C64876FFD8FD8EF862FB969DBA" authority="Wu &amp; Christiansen,1997" authorityName="Wu &amp; Christiansen" authorityYear="1997" box="[636,1082,1921,1945]" class="Entognatha" family="Arrhopalitidae" genus="Arrhopalites" kingdom="Animalia" order="Collembola" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="pukouensis">
<emphasis id="B91115574876FFD8FD8EF862FCB49DBA" box="[636,792,1921,1945]" italics="true" pageId="8" pageNumber="9">A. pukouensis</emphasis>
<bibRefCitation id="EFF4B4B44876FFD8FCD2F861FB969DBA" author="Wu" box="[800,1082,1921,1945]" pageId="8" pageNumber="9" refString="Wu, M. &amp; Christiansen, K. (1997) A new species of Arrhopalites from China (Collembola: Sminthuridae). Florida Entomologist, 80 (2), 266 - 269." type="journal article" year="1997">Wu &amp; Christiansen,1997</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C65B2C64876FFD8FBB7F862FA309DBA" authority="Vargovitsh, 2009" authorityName="Vargovitsh" authorityYear="2009" box="[1093,1436,1921,1945]" class="Entognatha" family="Arrhopalitidae" genus="Arrhopalites" kingdom="Animalia" order="Collembola" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="peculiaris">
<emphasis id="B91115574876FFD8FBB7F862FB799DBA" box="[1093,1237,1921,1945]" italics="true" pageId="8" pageNumber="9">A. peculiaris</emphasis>
<bibRefCitation id="EFF4B4B44876FFD8FB2EF862FA309DBA" author="Vargovitsh" box="[1244,1436,1921,1945]" pageId="8" pageNumber="9" refString="Vargovitsh, R. S. (2009) New Cave Arrhopalitidae (Collembola: Symphypleona) from the Crimea (Ukraine). Zootaxa, 2047, 1 - 47." type="journal article" year="2009">Vargovitsh, 2009</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C65B2C64876FFD8FF35F845FE359D9E" authority="Yosii, 1966" authorityName="Yosii" authorityYear="1966" box="[199,409,1957,1982]" class="Entognatha" family="Arrhopalitidae" genus="Arrhopalites" kingdom="Animalia" order="Collembola" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="gul">
<emphasis id="B91115574876FFD8FF35F845FEA09D9D" box="[199,268,1958,1982]" italics="true" pageId="8" pageNumber="9">A. gul</emphasis>
<bibRefCitation id="EFF4B4B44876FFD8FEE6F846FE359D9E" author="Yosii" box="[276,409,1957,1981]" pageId="8" pageNumber="9" refString="Yosii, R. (1966) Results of the speleological survey in South Korea 1966. IV. Cave Collembola of South Korea. Bulletin of the National Science Museum, Tokyo, 9 (4), 541 - 561." type="journal article" year="1966">Yosii, 1966</bibRefCitation>
</taxonomicName>
. The first three have short antennae with undivided Ant IV, strongly spine-like distolateral seta on dens and claws neither elongated nor thinned. The latest two share with the new species elongated antennae with annuli between subsegments of Ant IV and troglomorphic claws. Besides, as well as new species
<taxonomicName id="4C65B2C64877FFD9FAFEFF7BFAE29A93" box="[1292,1358,152,176]" class="Entognatha" family="Arrhopalitidae" genus="Arrhopalites" kingdom="Animalia" order="Collembola" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="gul">
<emphasis id="B91115574877FFD9FAFEFF7BFAE29A93" box="[1292,1358,152,176]" italics="true" pageId="9" pageNumber="10">A. gul</emphasis>
</taxonomicName>
has no strongly spine-like setae on dens and has winged circumanal setae. From
<taxonomicName id="4C65B2C64877FFD9FC3DFF5FFBBE9AF7" box="[975,1042,188,212]" class="Entognatha" family="Arrhopalitidae" genus="Arrhopalites" kingdom="Animalia" order="Collembola" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="gul">
<emphasis id="B91115574877FFD9FC3DFF5FFBBE9AF7" box="[975,1042,188,212]" italics="true" pageId="9" pageNumber="10">A. gul</emphasis>
</taxonomicName>
the new species differs by absence of spiny setae on head dorsum, presence of corner tooth on hind empodium, antennae: head ratio 2× (3× in
<taxonomicName id="4C65B2C64877FFD9FABBFF03FA209ADB" box="[1353,1420,224,248]" class="Entognatha" family="Arrhopalitidae" genus="Arrhopalites" kingdom="Animalia" order="Collembola" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="gul">
<emphasis id="B91115574877FFD9FABBFF03FA209ADB" box="[1353,1420,224,248]" italics="true" pageId="9" pageNumber="10">A. gul</emphasis>
</taxonomicName>
), and not broadened tip of mucro. Differences with
<taxonomicName id="4C65B2C64877FFD9FD33FEE7FCFC9B3F" box="[705,848,260,284]" class="Entognatha" family="Arrhopalitidae" genus="Arrhopalites" kingdom="Animalia" order="Collembola" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="peculiaris">
<emphasis id="B91115574877FFD9FD33FEE7FCFC9B3F" box="[705,848,260,284]" italics="true" pageId="9" pageNumber="10">A. peculiaris</emphasis>
</taxonomicName>
are: very long claws (thinned but not elongated in
<taxonomicName id="4C65B2C64877FFD9FA76FEE7FEAB9B63" class="Entognatha" family="Arrhopalitidae" genus="Arrhopalites" kingdom="Animalia" order="Collembola" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="peculiaris">
<emphasis id="B91115574877FFD9FA76FEE7FEAB9B63" italics="true" pageId="9" pageNumber="10">A. peculiaris</emphasis>
</taxonomicName>
), absence of cephalic spiny setae (5 spines in
<taxonomicName id="4C65B2C64877FFD9FCD2FECBFC1D9B63" box="[800,945,296,320]" class="Entognatha" family="Arrhopalitidae" genus="Arrhopalites" kingdom="Animalia" order="Collembola" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="peculiaris">
<emphasis id="B91115574877FFD9FCD2FECBFC1D9B63" box="[800,945,296,320]" italics="true" pageId="9" pageNumber="10">A. peculiaris</emphasis>
</taxonomicName>
), size of seta c1 of trichobothrial complex (about 20 µm in
<taxonomicName id="4C65B2C64877FFD9FEA4FEAEFE6F9B47" box="[342,451,333,356]" class="Entognatha" family="Arrhopalitidae" genus="Arrhopalites" kingdom="Animalia" order="Collembola" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="macronyx">
<emphasis id="B91115574877FFD9FEA4FEAEFE6F9B47" box="[342,451,333,356]" italics="true" pageId="9" pageNumber="10">macronyx</emphasis>
</taxonomicName>
versus about 8 µm in
<taxonomicName id="4C65B2C64877FFD9FD31FEAFFC9F9B47" box="[707,819,332,356]" class="Entognatha" family="Arrhopalitidae" genus="Arrhopalites" kingdom="Animalia" order="Collembola" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="peculiaris">
<emphasis id="B91115574877FFD9FD31FEAFFC9F9B47" box="[707,819,332,356]" italics="true" pageId="9" pageNumber="10">peculiaris</emphasis>
</taxonomicName>
), broadened and serrated circumanal setae, absence of strongly spine-like setae on dens, shape of mucro and antennal organ of third antennal segment (
<tableCitation id="C6E7FCFE4877FFD9FB3FFE8CFA899BA4" box="[1229,1317,367,391]" captionStart="TABLE 2" captionStartId="9.[151,235,440,461]" captionText="TABLE 2. Differences between Troglopalites stygios gen. nov. sp. nov., Arrhopalites macronyx sp. nov. and similar species. Species ABC angle Ant: H Ant IV ss Ant IV setae whorls AO-III rods, µm H spiny setae H: tita I T. stygios ~ flat 3 13 20 5 – 8 - 1.2" pageId="9" pageNumber="10">Table 2</tableCitation>
).
</paragraph>
<caption id="DF1A99CD4877FFD9FF65FE5BFAE1980B" pageId="9" pageNumber="10">
<paragraph id="8BDAC9454877FFD9FF65FE5BFAE1980B" blockId="9.[151,1419,440,677]" pageId="9" pageNumber="10">
<emphasis id="B91115574877FFD9FF65FE5BFEA89BEE" bold="true" box="[151,260,440,461]" pageId="9" pageNumber="10">TABLE 2.</emphasis>
Differences between
<taxonomicName id="4C65B2C64877FFD9FE13FE5BFD189BED" box="[481,692,440,462]" class="Entognatha" family="Arrhopalitidae" genus="Troglopalites" kingdom="Animalia" order="Collembola" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="stygios" status="gen. nov.">
<emphasis id="B91115574877FFD9FE13FE5BFD189BED" box="[481,692,440,462]" italics="true" pageId="9" pageNumber="10">Troglopalites stygios</emphasis>
</taxonomicName>
<emphasis id="B91115574877FFD9FD48FE5BFCDC9BEE" bold="true" box="[698,880,440,461]" pageId="9" pageNumber="10">
<taxonomicNameLabel id="A222A82C4877FFD9FD48FE5BFCB59BEE" box="[698,793,440,461]" pageId="9" pageNumber="10" rank="genus">gen. nov.</taxonomicNameLabel>
sp. nov.
</emphasis>
,
<taxonomicName id="4C65B2C64877FFD9FC8FFE5BFBC99BED" box="[893,1125,440,462]" class="Entognatha" family="Arrhopalitidae" genus="Arrhopalites" kingdom="Animalia" order="Collembola" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="macronyx" status="sp. nov.">
<emphasis id="B91115574877FFD9FC8FFE5BFBC99BED" box="[893,1125,440,462]" italics="true" pageId="9" pageNumber="10">Arrhopalites macronyx</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A222A82C4877FFD9FB9EFE5BFB109BEE" box="[1132,1212,440,461]" pageId="9" pageNumber="10" rank="species">
<emphasis id="B91115574877FFD9FB9EFE5BFB109BEE" bold="true" box="[1132,1212,440,461]" pageId="9" pageNumber="10">sp. nov.</emphasis>
</taxonomicNameLabel>
and similar species. Species ABC angle Ant: H Ant IV ss Ant IV setae whorls AO-III rods, µm H spiny setae H: tita I
<taxonomicName id="4C65B2C64877FFD9FF6DFDF0FF50980B" box="[159,252,531,552]" class="Entognatha" family="Arrhopalitidae" genus="Troglopalites" kingdom="Animalia" order="Collembola" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="stygios">
<emphasis id="B91115574877FFD9FF6DFDF0FF50980B" box="[159,252,531,552]" italics="true" pageId="9" pageNumber="10">T. stygios</emphasis>
</taxonomicName>
~ flat 3 13 20 5–8 - 1.2
</paragraph>
</caption>
<paragraph id="8BDAC9454877FFD9FF6DFDDFFAE19872" blockId="9.[151,1419,440,677]" box="[159,1357,571,594]" pageId="9" pageNumber="10">
<taxonomicName id="4C65B2C64877FFD9FF6DFDDFFEB29871" box="[159,286,572,594]" class="Entognatha" family="Arrhopalitidae" genus="Arrhopalites" kingdom="Animalia" order="Collembola" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="macronyx">
<emphasis id="B91115574877FFD9FF6DFDDFFEB29871" box="[159,286,572,594]" italics="true" pageId="9" pageNumber="10">A. macronyx</emphasis>
</taxonomicName>
~ right
<date id="FFDBEF854877FFD9FE0BFDDFFD569872" box="[505,762,572,593]" pageId="9" pageNumber="10" value="1914-07-02" valueMax="1914-08-02" valueMin="1914-07-02">2 7–8 14</date>
10–13 - 1.5
</paragraph>
<paragraph id="8BDAC9454877FFD9FF6DFD85FA949887" blockId="9.[151,1419,440,677]" pageId="9" pageNumber="10">
<taxonomicName id="4C65B2C64877FFD9FF6DFD85FE8C9858" box="[159,288,613,635]" class="Entognatha" family="Arrhopalitidae" genus="Arrhopalites" kingdom="Animalia" order="Collembola" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="peculiaris">
<emphasis id="B91115574877FFD9FF6DFD85FE8C9858" box="[159,288,613,635]" italics="true" pageId="9" pageNumber="10">A. peculiaris</emphasis>
</taxonomicName>
~ right 2–2.4 6–7
<date id="FFDBEF854877FFD9FD13FD86FB3F9858" box="[737,1171,613,635]" pageId="9" pageNumber="10" value="1915-07-05" valueMax="1915-09-05" valueMin="1915-07-05">15 7–9 5</date>
1.3–1.5
<taxonomicName id="4C65B2C64877FFD9FF6DFD6CFF709886" box="[159,220,655,677]" class="Entognatha" family="Arrhopalitidae" genus="Arrhopalites" kingdom="Animalia" order="Collembola" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="gul">
<emphasis id="B91115574877FFD9FF6DFD6CFF709886" box="[159,220,655,677]" italics="true" pageId="9" pageNumber="10">A. gul</emphasis>
</taxonomicName>
?
<date id="FFDBEF854877FFD9FE0BFD6CFD569887" box="[505,762,655,676]" pageId="9" pageNumber="10" value="1914-07-03">3 7 14</date>
? +?
</paragraph>
<paragraph id="8BDAC9454877FFD9FF65FD28FEAE98C2" blockId="9.[151,1345,715,955]" box="[151,258,715,737]" pageId="9" pageNumber="10">continued.</paragraph>
<paragraph id="8BDAC9454877FFD9FF6DFD1EFABE9998" blockId="9.[151,1345,715,955]" pageId="9" pageNumber="10">
Species Tita: Cl I, III Emp tooth I, II, III Tr III setae App anal Abd VI circ
<taxonomicName id="4C65B2C64877FFD9FF6DFCC9FF50991C" box="[159,252,810,831]" class="Entognatha" family="Arrhopalitidae" genus="Troglopalites" kingdom="Animalia" order="Collembola" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="stygios">
<emphasis id="B91115574877FFD9FF6DFCC9FF50991C" box="[159,252,810,831]" italics="true" pageId="9" pageNumber="10">T. stygios</emphasis>
</taxonomicName>
3, 4 0, 1, 1 4 br, ser simple
<taxonomicName id="4C65B2C64877FFD9FF6DFCB0FEB2994B" box="[159,286,851,872]" class="Entognatha" family="Arrhopalitidae" genus="Arrhopalites" kingdom="Animalia" order="Collembola" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="macronyx">
<emphasis id="B91115574877FFD9FF6DFCB0FEB2994B" box="[159,286,851,872]" italics="true" pageId="9" pageNumber="10">A. macronyx</emphasis>
</taxonomicName>
2, 2.5 0, 0, 1 5 ac, sm, cu L, b-ser
<taxonomicName id="4C65B2C64877FFD9FF6DFC9FFE8C99B1" box="[159,288,892,914]" class="Entognatha" family="Arrhopalitidae" genus="Arrhopalites" kingdom="Animalia" order="Collembola" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="peculiaris">
<emphasis id="B91115574877FFD9FF6DFC9FFE8C99B1" box="[159,288,892,914]" italics="true" pageId="9" pageNumber="10">A. peculiaris</emphasis>
</taxonomicName>
4.5, 6 1(0), 1(0), 1(0) 5 br, ser simple
<taxonomicName id="4C65B2C64877FFD9FF6DFC45FF709998" box="[159,220,933,955]" class="Entognatha" family="Arrhopalitidae" genus="Arrhopalites" kingdom="Animalia" order="Collembola" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="gul">
<emphasis id="B91115574877FFD9FF6DFC45FF709998" box="[159,220,933,955]" italics="true" pageId="9" pageNumber="10">A. gul</emphasis>
</taxonomicName>
? 0, 0, 0; (rarely 1)? rod, bl L, b-cil
</paragraph>
<paragraph id="8BDAC9454877FFD9FF65FC3DFCA19E73" blockId="9.[151,1436,990,1104]" pageId="9" pageNumber="10">ABC—trichobothria; H—head; ss—subsegments; AO-III—sense organ of third antennal segment, length is given for females only; Tr—trochanter; Tita—tibiotarsus; Cl—claw; Emp—empodium; App anal—female appendices anales: ac—acuminated, bl—blunt, br—gradually broadened, cu—curved, sm—smooth, ser—serrated, rod—rod-like; circ—circumanal setae: Llamellate, b-ser—serrated near base, b-cil—ciliated near base.</paragraph>
</subSubSection>
</treatment>
</document>
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