146 lines
15 KiB
XML
146 lines
15 KiB
XML
<document id="3FB4AA4FF31C03490673919760112126" ID-DOI="10.5281/zenodo.182004" ID-GBIF-Dataset="23b80268-81c4-4a9b-b53b-b59ff4eca084" ID-ISSN="1175-5326" ID-Zenodo-Dep="182004" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459847869286" checkinUser="plazi" docAuthor="Shear, William A." docDate="2008" docId="03CC5E1CFFC4B90D10A0FE3BFB0219BB" docLanguage="en" docName="zt01764p065.pdf" docOrigin="Zootaxa 1764" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Pseudotremia fongi Shear, 2008, n. sp." docType="treatment" docVersion="9" lastPageNumber="61" masterDocId="FFF52664FFC3B9051037FFECFFDD1D51" masterDocTitle="Cave millipeds of the United States. VII. New species and records of the genus Pseudotremia Cope. I. Species from West Virginia, USA (Diplopoda, Chordeumatida, Cleidogonidae)" masterLastPageNumber="65" masterPageNumber="53" pageNumber="60" updateTime="1698224999803" updateUser="plazi">
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<mods:title id="38D190440627FF39AA07C8594499A89C">Cave millipeds of the United States. VII. New species and records of the genus Pseudotremia Cope. I. Species from West Virginia, USA (Diplopoda, Chordeumatida, Cleidogonidae)</mods:title>
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<mods:name id="F9997AF76E25C6048B9CE02F8306A4FB" type="personal">
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<mods:namePart id="88842ED336BD6608D19058E5C19FFFA0">Shear, William A.</mods:namePart>
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<mods:title id="07EE1CD00D82D33042D4D455037589CF">Zootaxa</mods:title>
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<mods:part id="C3F607EF5EFAA89D5F9CB02909A1D95D">
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<mods:date id="3756CB37522C312B883E5BC348EA6EC9">2008</mods:date>
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<mods:detail id="53F10014458232F5F3E11E0F05F64601" type="volume">
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<mods:number id="F9C1B1C8E85693DBA338C97B4D1C78D2">1764</mods:number>
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<mods:start id="4789FD03C7A4E8B00330F32EB12FD08B">53</mods:start>
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<mods:end id="D1E256EB3F81EE75D9B315421E35EB3F">65</mods:end>
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<mods:classification id="A3713FE75B7947BB3C6F7266DF642EB8">journal article</mods:classification>
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<mods:identifier id="7E48986898363E1BE6137C1A0A667B6C" type="DOI">10.5281/zenodo.182004</mods:identifier>
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<mods:identifier id="1E1AD2B7692D6B239DEDE2EA9B7A0373" type="GBIF-Dataset">23b80268-81c4-4a9b-b53b-b59ff4eca084</mods:identifier>
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<mods:identifier id="2E30855F3D39F56924B08FF75CAA701F" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="5BCBA78132B91BED1AB5AF16A2BA04F9" type="Zenodo-Dep">182004</mods:identifier>
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<treatment id="03CC5E1CFFC4B90D10A0FE3BFB0219BB" ID-DOI="http://doi.org/10.5281/zenodo.6227852" ID-GBIF-Taxon="119361845" ID-Zenodo-Dep="6227852" LSID="urn:lsid:plazi:treatment:03CC5E1CFFC4B90D10A0FE3BFB0219BB" httpUri="http://treatment.plazi.org/id/03CC5E1CFFC4B90D10A0FE3BFB0219BB" lastPageId="8" lastPageNumber="61" pageId="7" pageNumber="60">
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<subSubSection id="C37FBC81FFC4B90210A0FE3BFED91F45" pageId="7" pageNumber="60" type="nomenclature">
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<paragraph id="8BDAEF0AFFC4B90210A0FE3BFE081CA0" blockId="7.[151,469,471,532]" box="[151,469,471,497]" pageId="7" pageNumber="60">
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<heading id="D0925866FFC4B90210A0FE3BFE081CA0" bold="true" box="[151,469,471,497]" fontSize="11" level="1" pageId="7" pageNumber="60" reason="1">
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<emphasis id="B9113318FFC4B90210A0FE3BFE081CA0" bold="true" box="[151,469,471,497]" pageId="7" pageNumber="60">
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<emphasis id="B9113318FFC4B90210A0FE3BFE561CA0" bold="true" box="[151,395,471,497]" italics="true" pageId="7" pageNumber="60">
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<taxonomicName id="4C659489FFC4B90210A0FE3BFE581CA0" box="[151,389,471,497]" class="Diplopoda" family="Cleidogonidae" genus="Pseudotremia" kingdom="Animalia" order="Chordeumatida" pageId="7" pageNumber="60" phylum="Arthropoda" rank="species" species="fongi" status="sp. nov.">Pseudotremia fongi</taxonomicName>
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,
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</emphasis>
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<taxonomicNameLabel id="A2228E63FFC4B90211A5FE3BFE081CA0" box="[402,469,471,497]" pageId="7" pageNumber="60" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BDAEF0AFFC4B90210A0FE16FED91F45" blockId="7.[151,469,471,532]" box="[151,260,506,532]" pageId="7" pageNumber="60">
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<figureCitation id="135EF38FFFC4B90210A0FE16FED91F45" box="[151,260,506,532]" captionStart="FIGURES 5 – 8" captionStartId="7.[151,270,1969,1993]" captionTargetBox="[165,1404,699,1943]" captionTargetId="figure@7.[151,1416,673,1956]" captionTargetPageId="7" captionText="FIGURES 5 – 8. Pseudotremia fongi, n. sp. 5. Gonopods, anterior view. 6. Gonopods, lateral view. 7. Right leg 9 of male, anterior view. 8. Right cyphopod, ventral view." httpUri="https://zenodo.org/record/182006/files/figure.png" pageId="7" pageNumber="60">Figs. 5–8</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37FBC81FFC4B90210A0FDA6FB7B1FDA" pageId="7" pageNumber="60" type="description">
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<paragraph id="8BDAEF0AFFC4B90210A0FDA6FB7B1FDA" blockId="7.[151,1436,586,652]" pageId="7" pageNumber="60">
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<emphasis id="B9113318FFC4B90210A0FDA6FF3A1F35" bold="true" box="[151,231,586,612]" pageId="7" pageNumber="60">
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<typeStatus id="54DE51A8FFC4B90210A0FDA6FF3F1F35" box="[151,226,586,612]" pageId="7" pageNumber="60">Types</typeStatus>
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:
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</emphasis>
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Male
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<typeStatus id="54DE51A8FFC4B902110FFDA6FE421F35" box="[312,415,586,612]" pageId="7" pageNumber="60" type="holotype">holotype</typeStatus>
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and female
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<typeStatus id="54DE51A8FFC4B9021202FDA6FD471F35" box="[565,666,586,612]" pageId="7" pageNumber="60" type="paratype">paratype</typeStatus>
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(VMNH) from Falling Springs Cavern,
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<quantity id="4C9D42EFFFC4B90214B7FDA6FB331F35" box="[1152,1262,586,612]" metricMagnitude="3" metricUnit="m" metricValue="7.242048" pageId="7" pageNumber="60" unit="mi" value="4.5">4.5 miles</quantity>
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north of Slaty Fork,
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<emphasis id="B9113318FFC4B90210EAFD9EFEA51FDA" box="[221,376,626,651]" italics="true" pageId="7" pageNumber="60">Randolph Co</emphasis>
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., WEST VIRGINIA, collected
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<date id="FFDBC9CAFFC4B90212DCFD9EFC5B1FDD" box="[747,902,626,652]" pageId="7" pageNumber="60" value="1977-06-11">11 June 1977</date>
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by John Holsinger
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<emphasis id="B9113318FFC4B902145CFD9EFB7B1FDA" box="[1131,1190,626,651]" italics="true" pageId="7" pageNumber="60">et al.</emphasis>
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</paragraph>
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</subSubSection>
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<caption id="DF1ABF82FFC4B90210A0F85DFD501ABA" httpUri="https://zenodo.org/record/182006/files/figure.png" pageId="7" pageNumber="60" targetBox="[165,1404,699,1943]" targetPageId="7">
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<paragraph id="8BDAEF0AFFC4B90210A0F85DFD501ABA" blockId="7.[151,1435,1969,2027]" pageId="7" pageNumber="60">
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<emphasis id="B9113318FFC4B90210A0F85DFE9F1A98" bold="true" box="[151,322,1969,1993]" pageId="7" pageNumber="60">FIGURES 5–8.</emphasis>
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<emphasis id="B9113318FFC4B902117FF85EFDC21A98" box="[328,543,1970,1993]" italics="true" pageId="7" pageNumber="60">
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<taxonomicName id="4C659489FFC4B902117FF85EFDC61A98" box="[328,539,1970,1993]" class="Diplopoda" family="Cleidogonidae" genus="Pseudotremia" kingdom="Animalia" order="Chordeumatida" pageId="7" pageNumber="60" phylum="Arthropoda" rank="species" species="fongi" status="sp. nov.">Pseudotremia fongi</taxonomicName>
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,
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</emphasis>
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<taxonomicNameLabel id="A2228E63FFC4B9021212F85DFDBD1A98" box="[549,608,1969,1993]" pageId="7" pageNumber="60" rank="species">
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<emphasis id="B9113318FFC4B9021212F85DFDBD1A98" bold="true" box="[549,608,1969,1993]" pageId="7" pageNumber="60">n. sp.</emphasis>
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</taxonomicNameLabel>
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5. Gonopods, anterior view. 6. Gonopods, lateral view. 7. Right leg 9 of male, anterior view. 8. Right cyphopod, ventral view.
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</paragraph>
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</caption>
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<subSubSection id="C37FBC81FFCBB90D10F1FF7BFC321C53" pageId="8" pageNumber="61" type="diagnosis">
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<paragraph id="8BDAEF0AFFCBB90D10F1FF7BFC321C53" blockId="8.[151,1437,151,1258]" pageId="8" pageNumber="61">
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<emphasis id="B9113318FFCBB90D10F1FF7BFE951DE0" bold="true" box="[198,328,151,177]" pageId="8" pageNumber="61">Diagnosis:</emphasis>
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<taxonomicName id="4C659489FFCBB90D1165FF74FDE41DE0" box="[338,569,152,177]" class="Diplopoda" family="Cleidogonidae" genus="Pseudotremia" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="fongi">
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<emphasis id="B9113318FFCBB90D1165FF74FDE41DE0" box="[338,569,152,177]" italics="true" pageId="8" pageNumber="61">Pseudotremia fongi</emphasis>
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</taxonomicName>
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has a massive ventral colpocoxite process that is unique in the genus; it protrudes far anterior of the angiocoxites and curves ventrally. In addition, the ventral branch of the lateral colpocoxite is deflexed dorsally and bears two short, acute sub-branches.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37FBC81FFCBB90D10F1FEE3FC791C03" pageId="8" pageNumber="61" type="etymology">
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<paragraph id="8BDAEF0AFFCBB90D10F1FEE3FC791C03" blockId="8.[151,1437,151,1258]" pageId="8" pageNumber="61">
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<emphasis id="B9113318FFCBB90D10F1FEE3FE8A1C78" bold="true" box="[198,343,271,297]" pageId="8" pageNumber="61">Etymology:</emphasis>
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The specific epithet honors Daniel Fong, in recognition of his leadership in the compilation of
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<emphasis id="B9113318FFCBB90D1081FED4FC401C00" box="[182,925,312,337]" italics="true" pageId="8" pageNumber="61">The Invertebrate Cave Fauna of West Viriginia, Second Edition</emphasis>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37FBC81FFCBB90D10F1FEB3FB0219BB" pageId="8" pageNumber="61" type="description">
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<paragraph id="8BDAEF0AFFCBB90D10F1FEB3FA4F1F4B" blockId="8.[151,1437,151,1258]" pageId="8" pageNumber="61">
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<emphasis id="B9113318FFCBB90D10F1FEB3FEBC1C28" bold="true" box="[198,353,351,377]" pageId="8" pageNumber="61">Description:</emphasis>
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Male
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<typeStatus id="54DE51A8FFCBB90D119DFE8CFDCC1C2B" box="[426,529,352,378]" pageId="8" pageNumber="61" type="holotype">holotype</typeStatus>
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about
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<quantity id="4C9D42EFFFCBB90D1257FE8CFD6F1C2B" box="[608,690,352,378]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="8" pageNumber="61" unit="mm" value="35.0">35 mm</quantity>
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long,
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<quantity id="4C9D42EFFFCBB90D12CDFE8CFC891C2B" box="[762,852,352,378]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="8" pageNumber="61" unit="mm" value="2.8">2.8 mm</quantity>
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wide, third antennal segment
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<quantity id="4C9D42EFFFCBB90D1485FE8CFAD61C2B" box="[1202,1291,352,378]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="61" unit="mm" value="1.5">1.5 mm</quantity>
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long, Ocelli 22 on each side, well-formed, contiguous and darkly pigmented. Segmental shoulders large, protruding, persisting posteriorly to segment 25. Metazonites anteriorly nearly smooth, posteriorly with 10–12 rugae strongest along posterior margin of each segment, anteriorly on the posterior segments breaking up into low, oblong tubercles; 8–12 obvious, ridge-like lateral striations. Color dark purplish gray-brown, mottled lighter.
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</paragraph>
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<paragraph id="8BDAEF0AFFCBB90D10F1FDC4FA8F1FB3" blockId="8.[151,1437,151,1258]" pageId="8" pageNumber="61">Anterior gonopods (figs. 5, 6) large, robust; median angiocoxites relatively short but very broad in lateral view, with subapical spine directed toward midline; lateral angiocoxites divided, ventral branch decurved behind dorsal branch, trifid, with median prong longest, ventral branch entire, narrow, acute. Colpocoxites not deeply divided; ventral colpocoxite process massive, extending anterior to angiocoxites, curved ventrally, apically bifid; ventral process of moderate size, evenly curved, acute. Ninth legs (fig. 7) typically reduced.</paragraph>
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<paragraph id="8BDAEF0AFFCBB90D10F1FD1CFEBE1E63" blockId="8.[151,1437,151,1258]" pageId="8" pageNumber="61">
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Female similar to male, but
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<quantity id="4C9D42EFFFCBB90D122DFD1CFDB21E5B" box="[538,623,752,778]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" pageId="8" pageNumber="61" unit="mm" value="31.0">31 mm</quantity>
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long,
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<quantity id="4C9D42EFFFCBB90D128AFD1CFCC71E5B" box="[701,794,752,778]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="8" pageNumber="61" unit="mm" value="2.6">2.6 mm</quantity>
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wide. Cyphopods (fig. 8) with median valve longest, posteriorly acute.
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</paragraph>
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<paragraph id="8BDAEF0AFFCBB90D10F1FCD3FC9C1ED3" blockId="8.[151,1437,151,1258]" pageId="8" pageNumber="61">
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<emphasis id="B9113318FFCBB90D10F1FCD3FE721E08" bold="true" box="[198,431,831,857]" pageId="8" pageNumber="61">Additional record:</emphasis>
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WEST VIRGINIA: Greenbrier Co., Dry Cave,
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<quantity id="4C9D42EFFFCBB90D13EAFCACFBFD1E0B" box="[989,1056,832,858]" metricMagnitude="4" metricUnit="m" metricValue="1.609344" pageId="8" pageNumber="61" unit="mi" value="10.0">10 mi</quantity>
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NNE of White Sulphur Springs,
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<date id="FFDBC9CAFFCBB90D10A0FC84FEE31ED3" box="[151,318,872,898]" pageId="8" pageNumber="61" value="1969-04-26">26 April 1969</date>
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, J. Holsinger & R. Baroody, 1ɗ. 3Ψ, 3 juvs.
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</paragraph>
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<paragraph id="8BDAEF0AFFCBB90D10F1FC63FB0219BB" blockId="8.[151,1437,151,1258]" pageId="8" pageNumber="61">
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<emphasis id="B9113318FFCBB90D10F1FC63FECB1EF8" bold="true" box="[198,278,911,937]" pageId="8" pageNumber="61">Notes:</emphasis>
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The unique gonopods of this very large species nevertheless conform to the
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<taxonomicName id="4C659489FFCBB90D14ABFC7CFB2C1EFB" box="[1180,1265,912,938]" class="Diplopoda" family="Cleidogonidae" genus="Pseudotremia" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="57" phylum="Arthropoda" rank="species" species="hobbsi">Hobbsi</taxonomicName>
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Group model, while also suggesting the seemingly unrelated
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<taxonomicName id="4C659489FFCBB90D128BFC54FCF81E80" box="[700,805,952,977]" class="Diplopoda" family="Cleidogonidae" genus="Pseudotremia" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="kerboi">
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<emphasis id="B9113318FFCBB90D128BFC54FCF81E80" box="[700,805,952,977]" italics="true" pageId="8" pageNumber="61">P. kerboi</emphasis>
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</taxonomicName>
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(described below); however, that species lacks a dorsal colpocoxite process entirely. While collected in a cave, this must be primarily an epigean species because of its well-developed eyes and dark pigmentation. The male from Dry Cave differs slightly from the
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<typeStatus id="54DE51A8FFCBB90D157DFBE4FAA01973" box="[1354,1405,1032,1058]" pageId="8" pageNumber="61">type</typeStatus>
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in not having the shallow division at the tip of the ventral colpocoxite process, and lacking teeth on the ventral branch of the lateral angiocoxite. The two localities are widely separated and suggest that this species may be a broadly distributed one, known so far just from two distant caves, but present in poorly collected intervening habitats. Alternatively the differences between the two may signify separate species status, but more evidence is needed. Because of this ambiguity, the Dry Cave specimens have not been made
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<typeStatus id="54DE51A8FFCBB90D1450FB3CFB0619BB" box="[1127,1243,1232,1258]" pageId="8" pageNumber="61" type="paratype">paratypes</typeStatus>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |