429 lines
73 KiB
XML
429 lines
73 KiB
XML
<document id="E2965A989A63A4E789B12DA4DEA108BF" ID-CLB-Dataset="45567" ID-DOI="10.11646/zootaxa.3814.1.7" ID-GBIF-Dataset="008ff01b-111d-4ae7-916f-5354c87b2146" ID-ISSN="1175-5326" ID-Zenodo-Dep="231251" ID-ZooBank="78219981-7610-4584-B9BE-2D226A87FA55" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460433448350" checkinUser="plazi" docAuthor="Botero-Trujillo, Ricardo" docDate="2014" docId="03CB49558C535C77FF6AFC1D4ED2FC54" docLanguage="en" docName="zt03814p132.pdf" docOrigin="Zootaxa 3814 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cryptocellus sofiae Botero-Trujillo, 2014, sp. nov." docType="treatment" docVersion="9" lastPageNumber="129" masterDocId="FFF2312D8C515C7FFFFDFFBB4B44FF92" masterDocTitle="A new Colombian species of Cryptocellus (Arachnida, Ricinulei), with notes on the taxonomy of the genus" masterLastPageNumber="132" masterPageNumber="121" pageNumber="123" updateTime="1698331689057" updateUser="ExternalLinkService">
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<mods:mods id="CDB2F68922C878B6C365FE6D992D5E08" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="A427C8319602A49D3F4E48492937CC58">
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<mods:title id="252B675C4655F5558AE6CBB5196A891F">A new Colombian species of Cryptocellus (Arachnida, Ricinulei), with notes on the taxonomy of the genus</mods:title>
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<mods:name id="878670B1A755175B430E9310EE37F45E" type="personal">
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<mods:namePart id="656222FEB7E95BABCE891B356AEB3CFE">Botero-Trujillo, Ricardo</mods:namePart>
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<mods:title id="306DABBB11A6A3A07A9F53B96BD88EBD">Zootaxa</mods:title>
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<mods:part id="CDD41159B36532E33CFDA0B1BB22FE25">
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<mods:date id="6427342FFFE88FA88C3E422054F2CB10">2014</mods:date>
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<mods:number id="38DCB21E15AD21E0A70CD7FA18626B38">3814</mods:number>
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<mods:classification id="ED80FF7AF61EA62C400548708FD2DF98">journal article</mods:classification>
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<mods:identifier id="87C825D621BA0D289B78243AE881D8F5" type="CLB-Dataset">45567</mods:identifier>
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<mods:identifier id="58AF3CE8541A63EF6930186AFB1C1078" type="DOI">10.11646/zootaxa.3814.1.7</mods:identifier>
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<treatment id="03CB49558C535C77FF6AFC1D4ED2FC54" ID-DOI="http://doi.org/10.5281/zenodo.6136158" ID-GBIF-Taxon="119430945" ID-Zenodo-Dep="6136158" LSID="urn:lsid:plazi:treatment:03CB49558C535C77FF6AFC1D4ED2FC54" httpUri="http://treatment.plazi.org/id/03CB49558C535C77FF6AFC1D4ED2FC54" lastPageId="8" lastPageNumber="129" pageId="2" pageNumber="123">
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<subSubSection id="C378ABC88C535C7DFF6AFC1D4A6EFC70" pageId="2" pageNumber="123" type="nomenclature">
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<paragraph id="8BDDF8438C535C7DFF6AFC1D4AA6FC52" blockId="2.[151,482,934,994]" box="[151,482,934,960]" pageId="2" pageNumber="123">
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<heading id="D0954F2F8C535C7DFF6AFC1D4AA6FC52" bold="true" box="[151,482,934,960]" fontSize="11" level="1" pageId="2" pageNumber="123" reason="1">
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<emphasis id="B91624518C535C7DFF6AFC1D4AA6FC52" bold="true" box="[151,482,934,960]" pageId="2" pageNumber="123">
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<taxonomicName id="4C6283C08C535C7DFF6AFC1D4A38FC52" ID-CoL="ZVQL" box="[151,380,934,960]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="2" pageNumber="123" phylum="Arthropoda" rank="species" species="sofiae" status="sp. nov.">
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<emphasis id="B91624518C535C7DFF6AFC1D4A38FC52" bold="true" box="[151,380,934,960]" italics="true" pageId="2" pageNumber="123">Cryptocellus sofiae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A225992A8C535C7DFE79FC1D4AA6FC52" box="[388,482,934,960]" pageId="2" pageNumber="123" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BDDF8438C535C7DFF6AFC724A6EFC70" blockId="2.[151,482,934,994]" box="[151,298,969,994]" pageId="2" pageNumber="123">
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<figureCitation id="1359E4C68C535C7DFF6AFC724A6EFC70" box="[151,298,969,994]" captionStart-0="FIGURE 1" captionStart-1="FIGURES 2 – 7" captionStart-2="FIGURES 8 – 13" captionStart-3="FIGURES 14 – 20" captionStart-4="FIGURES 21 – 22" captionStartId-0="4.[151,250,1974,1996]" captionStartId-1="5.[151,264,1825,1847]" captionStartId-2="6.[151,264,1828,1850]" captionStartId-3="7.[151,264,1672,1694]" captionStartId-4="8.[151,264,700,722]" captionTargetBox-0="[230,1358,841,1944]" captionTargetBox-1="[151,1436,193,1802]" captionTargetBox-2="[151,1436,193,1806]" captionTargetBox-3="[151,1436,194,1645]" captionTargetBox-4="[151,1435,197,673]" captionTargetId-0="figure@4.[224,1362,841,1953]" captionTargetId-1="figure@5.[151,1436,193,1804]" captionTargetId-2="figure@6.[151,1436,193,1807]" captionTargetId-3="figure@7.[151,1436,193,1651]" captionTargetId-4="figure@8.[151,1436,193,678]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionText-0="FIGURE 1. Known distribution records of the genus Cryptocellus in Colombia." captionText-1="FIGURES 2 – 7. Cryptocellus sofiae sp. nov. 2, holotype male (ICN-Ari- 009), dorsal; 3, holotype male, ventral; 4, paratype female (ICN-Ari- 010), dorsal; 5, paratype female, ventral (some setae accidentally removed from the opisthosoma); 6, paratype female (ICN-Ari- 012), frontal aspect (median longitudinal furrow of carapace indicated by arrow); 7, holotype male, polygonal setae on tergites XI and XII." captionText-2="FIGURES 8 – 13. Cryptocellus sofiae sp. nov. 8, holotype male (ICN-Ari- 009), sternal region, pedipalps and sternites X – XI; 9, paratype female (ICN-Ari- 012), sternal region, pedipalps and sternites X – XI; 10, holotype male, pygidium, dorsal posterior aspect; 11, paratype male (ICN-Ari- 011), dorsal aspect of left chelicera; 12, paratype female (ICN-Ari- 012), genital lip and spermathecae, anterior aspect; 13, paratype female, genital lip and spermathecae, posterior aspect." captionText-3="FIGURES 14 – 20. Cryptocellus sofiae sp. nov., SEM micrographs. 14 – 18, paratype male (ICN-Ari- 011), right leg III; 14, prolateral aspect; 15, copulatory apparatus, prolateral aspect; 16, prolaterodorsal aspect; 17, apex of tarsal process and accessory piece; 18, level of junction between accessory piece and tarsal process; 19 – 20, paratype female (ICN-Ari- 012), spermathecae; 19, posterior aspect; 20, inclined posterior aspect. Abbreviations: ac, accessory piece (sperm transfer groove indicated by arrowheads, beginning of bifurcation indicated by arrow); Lc, lamina cyathiformis; L 1 – 2, lobes of tarsal process; L ′, L ″, lobes of accessory piece; mc, medial constriction of spermathecae; mP, metatarsal process; MT, metatarsus; pe, prolateral expansion of tarsal process; tP, tarsal process." captionText-4="FIGURES 21 – 22. Cryptocellus sofiae sp. nov., SEM micrographs, different setal types. 21, paratype female (ICN-Ari- 012), part of tergite XIII; 22, paratype male (ICN-Ari- 011), part of cucullus." httpUri-0="https://zenodo.org/record/231252/files/figure.png" httpUri-1="https://zenodo.org/record/231253/files/figure.png" httpUri-2="https://zenodo.org/record/231254/files/figure.png" httpUri-3="https://zenodo.org/record/231255/files/figure.png" httpUri-4="https://zenodo.org/record/231256/files/figure.png" pageId="2" pageNumber="123">Figures 1–22</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C378ABC88C535C7DFF6AFBAA4AFDFB4C" pageId="2" pageNumber="123" type="materials_examined">
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<paragraph id="8BDDF8438C535C7DFF6AFBAA4AFDFB4C" blockId="2.[151,1436,1041,2038]" pageId="2" pageNumber="123">
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<emphasis id="B91624518C535C7DFF6AFBAA4A09FBB8" bold="true" box="[151,333,1041,1066]" pageId="2" pageNumber="123">
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<typeStatus id="54D946E18C535C7DFF6AFBAA4B96FBB8" box="[151,210,1041,1066]" pageId="2" pageNumber="123">Type</typeStatus>
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material.
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</emphasis>
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<emphasis id="B91624518C535C7DFEA1FBAA492AFBBB" box="[348,622,1041,1065]" italics="true" pageId="2" pageNumber="123">
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<typeStatus id="54D946E18C535C7DFEA1FBAA4A87FBBB" box="[348,451,1041,1065]" pageId="2" pageNumber="123" type="holotype">Holotype</typeStatus>
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male (adult):
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</emphasis>
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<collectingCountry id="F375B8D38C535C7DFD83FBA9485CFBB8" box="[638,792,1042,1066]" name="Colombia" pageId="2" pageNumber="123">COLOMBIA</collectingCountry>
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: Vichada Department, Cumaribo, El Tuparro Natural National Park, Administrative Center, 0.5º21′15.1′′
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<geoCoordinate id="EE569E848C535C7DFD22FB8D48C4FBDC" box="[735,896,1077,1102]" direction="north" orientation="latitude" pageId="2" pageNumber="123" precision="1" value="67.86214">
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N
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<geoCoordinate id="EE569E848C535C7DFD06FB8E48C4FBDC" box="[763,896,1077,1102]" direction="west" orientation="longitude" pageId="2" pageNumber="123" precision="1" value="-67.86214">67º51′43.7′′</geoCoordinate>
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</geoCoordinate>
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W,
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<quantity id="4C9A55A68C535C7DFC52FB8D48ADFBDF" box="[943,1001,1078,1102]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="123" unit="m" value="80.0">80 m</quantity>
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elev.,
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<date id="FFDCDE838C535C7DFBC9FB8D4FA1FBDC" box="[1076,1253,1077,1102]" pageId="2" pageNumber="123" value="2012-03-21">21 March 2012</date>
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, in litter sample from forest associated with rocky outcrop, D. A. Luna & C. Romero (ICN-Ari-009).
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<emphasis id="B91624518C535C7DFB99FBE14F9BFBE3" box="[1124,1247,1114,1137]" italics="true" pageId="2" pageNumber="123">
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<typeStatus id="54D946E18C535C7DFB99FBE14F9DFBE3" box="[1124,1241,1114,1137]" pageId="2" pageNumber="123" type="paratype">Paratypes</typeStatus>
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:
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</emphasis>
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<collectingCountry id="F375B8D38C535C7DFB15FBE14EC6FBE0" box="[1256,1410,1114,1138]" name="Colombia" pageId="2" pageNumber="123">COLOMBIA</collectingCountry>
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:
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<specimenCount id="9D6433CA8C535C7DFA73FBE14B8BFB04" pageId="2" pageNumber="123" type="adult">2 adult</specimenCount>
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females (ICN-Ari-010 and ICN-Ari-012) and
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<specimenCount id="9D6433CA8C535C7DFD24FBC54862FB04" box="[729,806,1149,1174]" pageId="2" pageNumber="123" type="adult">1 adult</specimenCount>
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male (ICN-Ari-011), same data as
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<typeStatus id="54D946E18C535C7DFB4FFBC64E5CFB04" box="[1202,1304,1149,1174]" pageId="2" pageNumber="123" type="holotype">holotype</typeStatus>
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. Specimens ICN-Ari-009 and 0 10 preserved in 70% ethanol; specimens ICN-Ari-011 and 0 12 dehydrated and fixed to aluminum stubs for SEM.
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</paragraph>
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</subSubSection>
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<subSubSection id="C378ABC88C535C7DFF3AFB524809FA90" box="[199,845,1257,1282]" pageId="2" pageNumber="123" type="distribution">
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<paragraph id="8BDDF8438C535C7DFF3AFB524809FA90" blockId="2.[151,1436,1041,2038]" box="[199,845,1257,1282]" pageId="2" pageNumber="123">
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<emphasis id="B91624518C535C7DFF3AFB524A24FA90" bold="true" box="[199,352,1257,1282]" pageId="2" pageNumber="123">Distribution.</emphasis>
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Known only from the
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<typeStatus id="54D946E18C535C7DFD9DFB5149D4FA90" box="[608,656,1258,1282]" pageId="2" pageNumber="123">type</typeStatus>
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locality (
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<figureCitation id="1359E4C68C535C7DFD07FB524878FA90" box="[762,828,1257,1282]" captionStart="FIGURE 1" captionStartId="4.[151,250,1974,1996]" captionTargetBox="[230,1358,841,1944]" captionTargetId="figure@4.[224,1362,841,1953]" captionTargetPageId="4" captionText="FIGURE 1. Known distribution records of the genus Cryptocellus in Colombia." httpUri="https://zenodo.org/record/231252/files/figure.png" pageId="2" pageNumber="123">Fig. 1</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C378ABC88C535C7DFF3AFAB64A2DFAFC" pageId="2" pageNumber="123" type="etymology">
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<paragraph id="8BDDF8438C535C7DFF3AFAB64A2DFAFC" blockId="2.[151,1436,1041,2038]" pageId="2" pageNumber="123">
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<emphasis id="B91624518C535C7DFF3AFAB64A0AFAB4" bold="true" box="[199,334,1293,1318]" pageId="2" pageNumber="123">Etymology.</emphasis>
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The species is named after my lovely 7 year old niece, Sofia Carlosama Botero, who always kneels on the chair to take a curious glance at the “cute bugs” (as she says) I work with. It is a celebration of her precious company.
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</paragraph>
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</subSubSection>
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<subSubSection id="C378ABC88C535C7DFF3AFAC248EDF9B0" pageId="2" pageNumber="123" type="diagnosis">
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<paragraph id="8BDDF8438C535C7DFF3AFAC248EDF9B0" blockId="2.[151,1436,1041,2038]" pageId="2" pageNumber="123">
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<emphasis id="B91624518C535C7DFF3AFAC24A05FA00" bold="true" box="[199,321,1401,1426]" pageId="2" pageNumber="123">Diagnosis.</emphasis>
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A member of the
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<taxonomicName id="4C6283C08C535C7DFDE5FAC2490AFA03" box="[536,590,1401,1425]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="2" pageNumber="123" phylum="Arthropoda" rank="species" species="adisi">
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<emphasis id="B91624518C535C7DFDE5FAC2490AFA03" box="[536,590,1401,1425]" italics="true" pageId="2" pageNumber="123">adisi</emphasis>
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</taxonomicName>
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species-group, as here defined, based on the morphology of male leg III, which exhibits a relatively thin and curved tarsal process of the copulatory apparatus (
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<figureCitation id="1359E4C68C535C7DFB45FA264E56FA24" box="[1208,1298,1437,1462]" captionStart="FIGURES 14 – 20" captionStartId="7.[151,264,1672,1694]" captionTargetBox="[151,1436,194,1645]" captionTargetId="figure@7.[151,1436,193,1651]" captionTargetPageId="7" captionText="FIGURES 14 – 20. Cryptocellus sofiae sp. nov., SEM micrographs. 14 – 18, paratype male (ICN-Ari- 011), right leg III; 14, prolateral aspect; 15, copulatory apparatus, prolateral aspect; 16, prolaterodorsal aspect; 17, apex of tarsal process and accessory piece; 18, level of junction between accessory piece and tarsal process; 19 – 20, paratype female (ICN-Ari- 012), spermathecae; 19, posterior aspect; 20, inclined posterior aspect. Abbreviations: ac, accessory piece (sperm transfer groove indicated by arrowheads, beginning of bifurcation indicated by arrow); Lc, lamina cyathiformis; L 1 – 2, lobes of tarsal process; L ′, L ″, lobes of accessory piece; mc, medial constriction of spermathecae; mP, metatarsal process; MT, metatarsus; pe, prolateral expansion of tarsal process; tP, tarsal process." httpUri="https://zenodo.org/record/231255/files/figure.png" pageId="2" pageNumber="123">Fig. 15</figureCitation>
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). It can be distinguished from other species in the group in having the median plates of tergites XI and XII wider than long (
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<figureCitation id="1359E4C68C535C7DFF62FA5E4A77FA6C" box="[159,307,1509,1534]" captionStart="FIGURES 2 – 7" captionStartId="5.[151,264,1825,1847]" captionTargetBox="[151,1436,193,1802]" captionTargetId="figure@5.[151,1436,193,1804]" captionTargetPageId="5" captionText="FIGURES 2 – 7. Cryptocellus sofiae sp. nov. 2, holotype male (ICN-Ari- 009), dorsal; 3, holotype male, ventral; 4, paratype female (ICN-Ari- 010), dorsal; 5, paratype female, ventral (some setae accidentally removed from the opisthosoma); 6, paratype female (ICN-Ari- 012), frontal aspect (median longitudinal furrow of carapace indicated by arrow); 7, holotype male, polygonal setae on tergites XI and XII." httpUri="https://zenodo.org/record/231253/files/figure.png" pageId="2" pageNumber="123">Figs. 2, 4, 7</figureCitation>
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), a moderate ventral expansion on distal half of male tarsal process (=spoon-like shaped) (
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<figureCitation id="1359E4C68C535C7DFA99FA5E4BA4F9B0" captionStart="FIGURES 14 – 20" captionStartId="7.[151,264,1672,1694]" captionTargetBox="[151,1436,194,1645]" captionTargetId="figure@7.[151,1436,193,1651]" captionTargetPageId="7" captionText="FIGURES 14 – 20. Cryptocellus sofiae sp. nov., SEM micrographs. 14 – 18, paratype male (ICN-Ari- 011), right leg III; 14, prolateral aspect; 15, copulatory apparatus, prolateral aspect; 16, prolaterodorsal aspect; 17, apex of tarsal process and accessory piece; 18, level of junction between accessory piece and tarsal process; 19 – 20, paratype female (ICN-Ari- 012), spermathecae; 19, posterior aspect; 20, inclined posterior aspect. Abbreviations: ac, accessory piece (sperm transfer groove indicated by arrowheads, beginning of bifurcation indicated by arrow); Lc, lamina cyathiformis; L 1 – 2, lobes of tarsal process; L ′, L ″, lobes of accessory piece; mc, medial constriction of spermathecae; mP, metatarsal process; MT, metatarsus; pe, prolateral expansion of tarsal process; tP, tarsal process." httpUri="https://zenodo.org/record/231255/files/figure.png" pageId="2" pageNumber="123">Figs. 14–16</figureCitation>
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), and the female spermathecae touching together (
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<figureCitation id="1359E4C68C535C7DFCEEF9B248D9F9B0" box="[787,925,1545,1570]" captionStart="FIGURES 14 – 20" captionStartId="7.[151,264,1672,1694]" captionTargetBox="[151,1436,194,1645]" captionTargetId="figure@7.[151,1436,193,1651]" captionTargetPageId="7" captionText="FIGURES 14 – 20. Cryptocellus sofiae sp. nov., SEM micrographs. 14 – 18, paratype male (ICN-Ari- 011), right leg III; 14, prolateral aspect; 15, copulatory apparatus, prolateral aspect; 16, prolaterodorsal aspect; 17, apex of tarsal process and accessory piece; 18, level of junction between accessory piece and tarsal process; 19 – 20, paratype female (ICN-Ari- 012), spermathecae; 19, posterior aspect; 20, inclined posterior aspect. Abbreviations: ac, accessory piece (sperm transfer groove indicated by arrowheads, beginning of bifurcation indicated by arrow); Lc, lamina cyathiformis; L 1 – 2, lobes of tarsal process; L ′, L ″, lobes of accessory piece; mc, medial constriction of spermathecae; mP, metatarsal process; MT, metatarsus; pe, prolateral expansion of tarsal process; tP, tarsal process." httpUri="https://zenodo.org/record/231255/files/figure.png" pageId="2" pageNumber="123">Figs. 19–20</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C378ABC88C535C7DFF3AF9964FFAF9D4" box="[199,1214,1581,1606]" pageId="2" pageNumber="123" type="description">
|
||
<paragraph id="8BDDF8438C535C7DFF3AF9964FFAF9D4" blockId="2.[151,1436,1041,2038]" box="[199,1214,1581,1606]" pageId="2" pageNumber="123">
|
||
<emphasis id="B91624518C535C7DFF3AF9964972F9D4" bold="true" box="[199,566,1581,1606]" pageId="2" pageNumber="123">Description (male and female).</emphasis>
|
||
The description is based on the four specimens available.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C378ABC88C535C7CFF6AF9EA4FFEFA77" lastPageId="3" lastPageNumber="124" pageId="2" pageNumber="123" type="materials_examined">
|
||
<paragraph id="8BDDF8438C535C7DFF6AF9EA4E1EF920" blockId="2.[151,1436,1041,2038]" pageId="2" pageNumber="123">
|
||
<emphasis id="B91624518C535C7DFF6AF9EA4972F9FB" box="[151,566,1617,1641]" italics="true" pageId="2" pageNumber="123">Coloration (in preserved specimens).</emphasis>
|
||
Body and appendages with cuticle uniformly chestnut to light reddish, except for medial region of sternite XI in male, which is dark-red; carapace without pale or yellowish areas in ocular region; opisthosomal membranes yellow. Polygonal (navicular or calyx-like) setae whitish (
|
||
<figureCitation id="1359E4C68C535C7DFB67F9224E42F920" box="[1178,1286,1689,1714]" captionStart="FIGURES 2 – 7" captionStartId="5.[151,264,1825,1847]" captionTargetBox="[151,1436,193,1802]" captionTargetId="figure@5.[151,1436,193,1804]" captionTargetPageId="5" captionText="FIGURES 2 – 7. Cryptocellus sofiae sp. nov. 2, holotype male (ICN-Ari- 009), dorsal; 3, holotype male, ventral; 4, paratype female (ICN-Ari- 010), dorsal; 5, paratype female, ventral (some setae accidentally removed from the opisthosoma); 6, paratype female (ICN-Ari- 012), frontal aspect (median longitudinal furrow of carapace indicated by arrow); 7, holotype male, polygonal setae on tergites XI and XII." httpUri="https://zenodo.org/record/231253/files/figure.png" pageId="2" pageNumber="123">Figs. 2–7</figureCitation>
|
||
,
|
||
<figureCitation id="1359E4C68C535C7DFAEFF9214E09F920" box="[1298,1357,1690,1714]" captionStart="FIGURES 8 – 13" captionStartId="6.[151,264,1828,1850]" captionTargetBox="[151,1436,193,1806]" captionTargetId="figure@6.[151,1436,193,1807]" captionTargetPageId="6" captionText="FIGURES 8 – 13. Cryptocellus sofiae sp. nov. 8, holotype male (ICN-Ari- 009), sternal region, pedipalps and sternites X – XI; 9, paratype female (ICN-Ari- 012), sternal region, pedipalps and sternites X – XI; 10, holotype male, pygidium, dorsal posterior aspect; 11, paratype male (ICN-Ari- 011), dorsal aspect of left chelicera; 12, paratype female (ICN-Ari- 012), genital lip and spermathecae, anterior aspect; 13, paratype female, genital lip and spermathecae, posterior aspect." httpUri="https://zenodo.org/record/231254/files/figure.png" pageId="2" pageNumber="123">8–10</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDDF8438C535C7CFF3AF9054B9BFF46" blockId="2.[151,1436,1041,2038]" lastBlockId="3.[151,1437,151,2013]" lastPageId="3" lastPageNumber="124" pageId="2" pageNumber="123">
|
||
<emphasis id="B91624518C535C7DFF3AF9054A6EF947" box="[199,298,1726,1749]" italics="true" pageId="2" pageNumber="123">Setation.</emphasis>
|
||
Body entirely covered with setae. Most conspicuous are the large whitish polygonal (navicular or calyx-like) setae, that vary in shape (
|
||
<figureCitation id="1359E4C68C535C7DFDB9F95A49F6F968" box="[580,690,1761,1786]" captionStart="FIGURES 2 – 7" captionStartId="5.[151,264,1825,1847]" captionTargetBox="[151,1436,193,1802]" captionTargetId="figure@5.[151,1436,193,1804]" captionTargetPageId="5" captionText="FIGURES 2 – 7. Cryptocellus sofiae sp. nov. 2, holotype male (ICN-Ari- 009), dorsal; 3, holotype male, ventral; 4, paratype female (ICN-Ari- 010), dorsal; 5, paratype female, ventral (some setae accidentally removed from the opisthosoma); 6, paratype female (ICN-Ari- 012), frontal aspect (median longitudinal furrow of carapace indicated by arrow); 7, holotype male, polygonal setae on tergites XI and XII." httpUri="https://zenodo.org/record/231253/files/figure.png" pageId="2" pageNumber="123">Figs. 2–5</figureCitation>
|
||
,
|
||
<figureCitation id="1359E4C68C535C7DFD3CF959484EF968" box="[705,778,1762,1786]" captionStart="FIGURES 21 – 22" captionStartId="8.[151,264,700,722]" captionTargetBox="[151,1435,197,673]" captionTargetId="figure@8.[151,1436,193,678]" captionTargetPageId="8" captionText="FIGURES 21 – 22. Cryptocellus sofiae sp. nov., SEM micrographs, different setal types. 21, paratype female (ICN-Ari- 012), part of tergite XIII; 22, paratype male (ICN-Ari- 011), part of cucullus." httpUri="https://zenodo.org/record/231256/files/figure.png" pageId="2" pageNumber="123">21–22</figureCitation>
|
||
), on external surface of cucullus (except for distal fifth), carapace (including ocular areas and median longitudinal furrow), opisthosoma (except for medial region of sternites X and XI), pygidium, dorsal surfaces of pedipalp trochanter II and femur, leg coxae (only distally), leg trochanters I and II, femora, patellae, tibiae and metatarsi. These setae are large and not very numerous (e.g. about 60–70 setae on median plate of tergite XII;
|
||
<figureCitation id="1359E4C68C535C7DFD6DF8CA4992F818" box="[656,726,1905,1930]" captionStart="FIGURES 2 – 7" captionStartId="5.[151,264,1825,1847]" captionTargetBox="[151,1436,193,1802]" captionTargetId="figure@5.[151,1436,193,1804]" captionTargetPageId="5" captionText="FIGURES 2 – 7. Cryptocellus sofiae sp. nov. 2, holotype male (ICN-Ari- 009), dorsal; 3, holotype male, ventral; 4, paratype female (ICN-Ari- 010), dorsal; 5, paratype female, ventral (some setae accidentally removed from the opisthosoma); 6, paratype female (ICN-Ari- 012), frontal aspect (median longitudinal furrow of carapace indicated by arrow); 7, holotype male, polygonal setae on tergites XI and XII." httpUri="https://zenodo.org/record/231253/files/figure.png" pageId="2" pageNumber="123">Fig. 7</figureCitation>
|
||
). Distal fifth of external surface of cucullus, medial region of sternites X and XI, tritosternum, pedipalp coxae, pedipalp trochanter I, prolateral, retrolateral and ventral surfaces of pedipalp trochanter II and femur, pedipalp tibia, most of leg coxae and leg tarsomeres lack these large setae, being covered instead with smaller and thinner setae (
|
||
<figureCitation id="1359E4C68C535C7DFCFBF86648D6F864" box="[774,914,2013,2038]" captionStart="FIGURES 2 – 7" captionStartId="5.[151,264,1825,1847]" captionTargetBox="[151,1436,193,1802]" captionTargetId="figure@5.[151,1436,193,1804]" captionTargetPageId="5" captionText="FIGURES 2 – 7. Cryptocellus sofiae sp. nov. 2, holotype male (ICN-Ari- 009), dorsal; 3, holotype male, ventral; 4, paratype female (ICN-Ari- 010), dorsal; 5, paratype female, ventral (some setae accidentally removed from the opisthosoma); 6, paratype female (ICN-Ari- 012), frontal aspect (median longitudinal furrow of carapace indicated by arrow); 7, holotype male, polygonal setae on tergites XI and XII." httpUri="https://zenodo.org/record/231253/files/figure.png" pageId="2" pageNumber="123">Figs. 3, 5–6</figureCitation>
|
||
,
|
||
<figureCitation id="1359E4C68C535C7DFC5DF865488AF864" box="[928,974,2014,2038]" captionStart="FIGURES 8 – 13" captionStartId="6.[151,264,1828,1850]" captionTargetBox="[151,1436,193,1806]" captionTargetId="figure@6.[151,1436,193,1807]" captionTargetPageId="6" captionText="FIGURES 8 – 13. Cryptocellus sofiae sp. nov. 8, holotype male (ICN-Ari- 009), sternal region, pedipalps and sternites X – XI; 9, paratype female (ICN-Ari- 012), sternal region, pedipalps and sternites X – XI; 10, holotype male, pygidium, dorsal posterior aspect; 11, paratype male (ICN-Ari- 011), dorsal aspect of left chelicera; 12, paratype female (ICN-Ari- 012), genital lip and spermathecae, anterior aspect; 13, paratype female, genital lip and spermathecae, posterior aspect." httpUri="https://zenodo.org/record/231254/files/figure.png" pageId="2" pageNumber="123">8–9</figureCitation>
|
||
). SEM observations revealed a range of setal
|
||
<typeStatus id="54D946E18C525C7CFF2FFF234A49FF22" box="[210,269,152,176]" pageId="3" pageNumber="124">types</typeStatus>
|
||
on the cucullus, carapace and opisthosoma (
|
||
<figureCitation id="1359E4C68C525C7CFCFBFF2C48CBFF22" box="[774,911,151,176]" captionStart="FIGURES 21 – 22" captionStartId="8.[151,264,700,722]" captionTargetBox="[151,1435,197,673]" captionTargetId="figure@8.[151,1436,193,678]" captionTargetPageId="8" captionText="FIGURES 21 – 22. Cryptocellus sofiae sp. nov., SEM micrographs, different setal types. 21, paratype female (ICN-Ari- 012), part of tergite XIII; 22, paratype male (ICN-Ari- 011), part of cucullus." httpUri="https://zenodo.org/record/231256/files/figure.png" pageId="3" pageNumber="124">Figs. 21–22</figureCitation>
|
||
), similar to that present in
|
||
<taxonomicName id="4C6283C08C525C7CFB43FF234B9FFF46" authority="Adis et al. 1999" authorityName="Adis et al." authorityYear="1999" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="3" pageNumber="124" phylum="Arthropoda" rank="species" species="adisi">
|
||
<emphasis id="B91624518C525C7CFB43FF234E51FF22" box="[1214,1301,152,176]" italics="true" pageId="3" pageNumber="124">C. adisi</emphasis>
|
||
(
|
||
<bibRefCitation id="EFF385B28C525C7CFAD8FF2C4B97FF46" author="Adis" pageId="3" pageNumber="124" refString="Adis, J., Messner, B. & Platnick, N. (1999) Morphological structures and vertical distribution in the soil indicate facultative plastron respiration in Cryptocellus adisi (Arachnida, Ricinulei) from Central Amazonia. Studies on Neotropical Fauna and Environment, 34, 1 - 9. http: // dx. doi. org / 10.1076 / snfe. 34.1.1.8915" type="journal article" year="1999">
|
||
Adis
|
||
<emphasis id="B91624518C525C7CFA9EFF224ED8FF22" box="[1379,1436,152,176]" italics="true" pageId="3" pageNumber="124">et al.</emphasis>
|
||
1999
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BDDF8438C525C7CFF3AFF5B4A89FE3E" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="124">
|
||
<emphasis id="B91624518C525C7CFF3AFF5B4A79FF65" box="[199,317,224,247]" italics="true" pageId="3" pageNumber="124">Carapace.</emphasis>
|
||
Lateral margins not parallel (carapace narrowing anteriorly); anterior margin slightly concave, posterior margin convex; carapace approximately as long as wide, widest at level of leg III (
|
||
<figureCitation id="1359E4C68C525C7CFB2FFEBF4E01FE8E" box="[1234,1349,260,285]" captionStart="FIGURES 2 – 7" captionStartId="5.[151,264,1825,1847]" captionTargetBox="[151,1436,193,1802]" captionTargetId="figure@5.[151,1436,193,1804]" captionTargetPageId="5" captionText="FIGURES 2 – 7. Cryptocellus sofiae sp. nov. 2, holotype male (ICN-Ari- 009), dorsal; 3, holotype male, ventral; 4, paratype female (ICN-Ari- 010), dorsal; 5, paratype female, ventral (some setae accidentally removed from the opisthosoma); 6, paratype female (ICN-Ari- 012), frontal aspect (median longitudinal furrow of carapace indicated by arrow); 7, holotype male, polygonal setae on tergites XI and XII." httpUri="https://zenodo.org/record/231253/files/figure.png" pageId="3" pageNumber="124">Figs. 2, 4</figureCitation>
|
||
); ocelli completely absent, but corresponding ocular regions noticeably elevated, with an evident depression behind them; abundant cuticular pits of variable size, randomly distributed (most containing tubercles); very few tubercles present outside the pits; a shallow, median, longitudinal furrow is present (
|
||
<figureCitation id="1359E4C68C525C7CFC2BFED44F5EFE1A" box="[982,1050,367,392]" captionStart="FIGURES 2 – 7" captionStartId="5.[151,264,1825,1847]" captionTargetBox="[151,1436,193,1802]" captionTargetId="figure@5.[151,1436,193,1804]" captionTargetPageId="5" captionText="FIGURES 2 – 7. Cryptocellus sofiae sp. nov. 2, holotype male (ICN-Ari- 009), dorsal; 3, holotype male, ventral; 4, paratype female (ICN-Ari- 010), dorsal; 5, paratype female, ventral (some setae accidentally removed from the opisthosoma); 6, paratype female (ICN-Ari- 012), frontal aspect (median longitudinal furrow of carapace indicated by arrow); 7, holotype male, polygonal setae on tergites XI and XII." httpUri="https://zenodo.org/record/231253/files/figure.png" pageId="3" pageNumber="124">Fig. 6</figureCitation>
|
||
), beginning at level of coxae I and ending at level of coxae III.
|
||
</paragraph>
|
||
<paragraph id="8BDDF8438C525C7CFF3AFE034930FD8A" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="124">
|
||
<emphasis id="B91624518C525C7CFF3AFE034A76FE42" box="[199,306,440,464]" italics="true" pageId="3" pageNumber="124">Cucullus.</emphasis>
|
||
Wider than long, with lateral margins not parallel (cucullus narrowing posteriorly) (
|
||
<figureCitation id="1359E4C68C525C7CFAF4FE0C4E15FE42" box="[1289,1361,439,464]" captionStart="FIGURES 2 – 7" captionStartId="5.[151,264,1825,1847]" captionTargetBox="[151,1436,193,1802]" captionTargetId="figure@5.[151,1436,193,1804]" captionTargetPageId="5" captionText="FIGURES 2 – 7. Cryptocellus sofiae sp. nov. 2, holotype male (ICN-Ari- 009), dorsal; 3, holotype male, ventral; 4, paratype female (ICN-Ari- 010), dorsal; 5, paratype female, ventral (some setae accidentally removed from the opisthosoma); 6, paratype female (ICN-Ari- 012), frontal aspect (median longitudinal furrow of carapace indicated by arrow); 7, holotype male, polygonal setae on tergites XI and XII." httpUri="https://zenodo.org/record/231253/files/figure.png" pageId="3" pageNumber="124">Fig. 6</figureCitation>
|
||
); with abundant cuticular pits of variable size, randomly distributed (most of them containing tubercles); very few tubercles present outside the pits (
|
||
<figureCitation id="1359E4C68C525C7CFDE9FE44492CFD8A" box="[532,616,511,536]" captionStart="FIGURES 21 – 22" captionStartId="8.[151,264,700,722]" captionTargetBox="[151,1435,197,673]" captionTargetId="figure@8.[151,1436,193,678]" captionTargetPageId="8" captionText="FIGURES 21 – 22. Cryptocellus sofiae sp. nov., SEM micrographs, different setal types. 21, paratype female (ICN-Ari- 012), part of tergite XIII; 22, paratype male (ICN-Ari- 011), part of cucullus." httpUri="https://zenodo.org/record/231256/files/figure.png" pageId="3" pageNumber="124">Fig. 22</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDDF8438C525C7CFF3AFD9F486EFD5F" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="124">
|
||
<emphasis id="B91624518C525C7CFF3AFD9F4A79FDAE" box="[199,317,548,572]" italics="true" pageId="3" pageNumber="124">Chelicera.</emphasis>
|
||
Description based on left chelicera of
|
||
<typeStatus id="54D946E18C525C7CFCFDFD9E4824FDAF" box="[768,864,549,573]" pageId="3" pageNumber="124" type="paratype">paratype</typeStatus>
|
||
male (
|
||
<figureCitation id="1359E4C68C525C7CFC4FFD9F4F42FDAE" box="[946,1030,548,573]" captionStart="FIGURES 8 – 13" captionStartId="6.[151,264,1828,1850]" captionTargetBox="[151,1436,193,1806]" captionTargetId="figure@6.[151,1436,193,1807]" captionTargetPageId="6" captionText="FIGURES 8 – 13. Cryptocellus sofiae sp. nov. 8, holotype male (ICN-Ari- 009), sternal region, pedipalps and sternites X – XI; 9, paratype female (ICN-Ari- 012), sternal region, pedipalps and sternites X – XI; 10, holotype male, pygidium, dorsal posterior aspect; 11, paratype male (ICN-Ari- 011), dorsal aspect of left chelicera; 12, paratype female (ICN-Ari- 012), genital lip and spermathecae, anterior aspect; 13, paratype female, genital lip and spermathecae, posterior aspect." httpUri="https://zenodo.org/record/231254/files/figure.png" pageId="3" pageNumber="124">Fig. 11</figureCitation>
|
||
). Movable finger longer than fixed finger; movable finger armed with ten separate small teeth, sub-equal in size except for third basalmost tooth and the two distalmost teeth, which are twice the size of the others; fixed finger with markedly enlarged distal tooth, followed by two separate smaller teeth (slightly larger than the largest of the movable finger) and a small basal tooth, latter similar in size to most teeth of movable finger.
|
||
</paragraph>
|
||
<paragraph id="8BDDF8438C525C7CFF3AFD634F75FC12" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="124">
|
||
<emphasis id="B91624518C525C7CFF3AFD634A34FD7D" box="[199,368,728,752]" italics="true" pageId="3" pageNumber="124">Sternal region.</emphasis>
|
||
Without cuticular pits; coxae I not meeting tritosternum; coxae II–IV meeting entirely, subrectangular, progressively decreasing in length; coxae IV pear-shaped, with posterior margin slightly curved (female) or almost straight (male); suture line of coxae II about three fifths length of that of coxae III (coxae III being larger) (
|
||
<figureCitation id="1359E4C68C525C7CFEC5FCFF4AE2FCCE" box="[312,422,836,861]" captionStart="FIGURES 8 – 13" captionStartId="6.[151,264,1828,1850]" captionTargetBox="[151,1436,193,1806]" captionTargetId="figure@6.[151,1436,193,1807]" captionTargetPageId="6" captionText="FIGURES 8 – 13. Cryptocellus sofiae sp. nov. 8, holotype male (ICN-Ari- 009), sternal region, pedipalps and sternites X – XI; 9, paratype female (ICN-Ari- 012), sternal region, pedipalps and sternites X – XI; 10, holotype male, pygidium, dorsal posterior aspect; 11, paratype male (ICN-Ari- 011), dorsal aspect of left chelicera; 12, paratype female (ICN-Ari- 012), genital lip and spermathecae, anterior aspect; 13, paratype female, genital lip and spermathecae, posterior aspect." httpUri="https://zenodo.org/record/231254/files/figure.png" pageId="3" pageNumber="124">Figs. 8–9</figureCitation>
|
||
); coxae I with minute tubercles along posterior margin; coxae II–III with tubercles along the distal two thirds of anterior and posterior margins; coxae IV without tubercles.
|
||
</paragraph>
|
||
<paragraph id="8BDDF8438C525C7CFF3AFC374B87FB56" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="124">
|
||
<emphasis id="B91624518C525C7CFF3AFC374A26FC36" box="[199,354,908,932]" italics="true" pageId="3" pageNumber="124">Opisthosoma.</emphasis>
|
||
Longer than wide, widest at level of junction between tergites XI and XII (
|
||
<figureCitation id="1359E4C68C525C7CFB1CFC374E16FC37" box="[1249,1362,908,933]" captionStart="FIGURES 2 – 7" captionStartId="5.[151,264,1825,1847]" captionTargetBox="[151,1436,193,1802]" captionTargetId="figure@5.[151,1436,193,1804]" captionTargetPageId="5" captionText="FIGURES 2 – 7. Cryptocellus sofiae sp. nov. 2, holotype male (ICN-Ari- 009), dorsal; 3, holotype male, ventral; 4, paratype female (ICN-Ari- 010), dorsal; 5, paratype female, ventral (some setae accidentally removed from the opisthosoma); 6, paratype female (ICN-Ari- 012), frontal aspect (median longitudinal furrow of carapace indicated by arrow); 7, holotype male, polygonal setae on tergites XI and XII." httpUri="https://zenodo.org/record/231253/files/figure.png" pageId="3" pageNumber="124">Figs. 2–5</figureCitation>
|
||
); with abundant cuticular pits of variable size, randomly distributed on dorsal surface (most containing tubercles); very few tubercles present outside pits (
|
||
<figureCitation id="1359E4C68C525C7CFDC3FC6F49DCFC7E" box="[574,664,980,1005]" captionStart="FIGURES 21 – 22" captionStartId="8.[151,264,700,722]" captionTargetBox="[151,1435,197,673]" captionTargetId="figure@8.[151,1436,193,678]" captionTargetPageId="8" captionText="FIGURES 21 – 22. Cryptocellus sofiae sp. nov., SEM micrographs, different setal types. 21, paratype female (ICN-Ari- 012), part of tergite XIII; 22, paratype male (ICN-Ari- 011), part of cucullus." httpUri="https://zenodo.org/record/231256/files/figure.png" pageId="3" pageNumber="124">Fig. 21</figureCitation>
|
||
); median plates of tergites XI–XIII with paired, shallow, lateral depressions beside median elevation; median plates of tergites XI–XIII progressively decreasing in width, lateral margins not parallel on XI–XII (narrowing posteriorly); median plate of tergites XI–XII wider than long, median plate of tergite XIII approximately as wide as long (
|
||
<figureCitation id="1359E4C68C525C7CFCFDFB844830FBCA" box="[768,884,1087,1112]" captionStart="FIGURES 2 – 7" captionStartId="5.[151,264,1825,1847]" captionTargetBox="[151,1436,193,1802]" captionTargetId="figure@5.[151,1436,193,1804]" captionTargetPageId="5" captionText="FIGURES 2 – 7. Cryptocellus sofiae sp. nov. 2, holotype male (ICN-Ari- 009), dorsal; 3, holotype male, ventral; 4, paratype female (ICN-Ari- 010), dorsal; 5, paratype female, ventral (some setae accidentally removed from the opisthosoma); 6, paratype female (ICN-Ari- 012), frontal aspect (median longitudinal furrow of carapace indicated by arrow); 7, holotype male, polygonal setae on tergites XI and XII." httpUri="https://zenodo.org/record/231253/files/figure.png" pageId="3" pageNumber="124">Figs. 2, 4</figureCitation>
|
||
). Sternites without cuticular pits and devoid of tubercles; male with median region of sternite XI slightly protuberant and more sclerotized (unlike that of female, which is not elevated) (
|
||
<figureCitation id="1359E4C68C525C7CFE5DFB3C4949FB32" box="[416,525,1159,1184]" captionStart="FIGURES 2 – 7" captionStartId="5.[151,264,1825,1847]" captionTargetBox="[151,1436,193,1802]" captionTargetId="figure@5.[151,1436,193,1804]" captionTargetPageId="5" captionText="FIGURES 2 – 7. Cryptocellus sofiae sp. nov. 2, holotype male (ICN-Ari- 009), dorsal; 3, holotype male, ventral; 4, paratype female (ICN-Ari- 010), dorsal; 5, paratype female, ventral (some setae accidentally removed from the opisthosoma); 6, paratype female (ICN-Ari- 012), frontal aspect (median longitudinal furrow of carapace indicated by arrow); 7, holotype male, polygonal setae on tergites XI and XII." httpUri="https://zenodo.org/record/231253/files/figure.png" pageId="3" pageNumber="124">Figs. 3, 5</figureCitation>
|
||
). Basal segment of pygidium unnotched on either dorsal or ventral borders (
|
||
<figureCitation id="1359E4C68C525C7CFA92FB3C4BF2FB56" captionStart="FIGURES 8 – 13" captionStartId="6.[151,264,1828,1850]" captionTargetBox="[151,1436,193,1806]" captionTargetId="figure@6.[151,1436,193,1807]" captionTargetPageId="6" captionText="FIGURES 8 – 13. Cryptocellus sofiae sp. nov. 8, holotype male (ICN-Ari- 009), sternal region, pedipalps and sternites X – XI; 9, paratype female (ICN-Ari- 012), sternal region, pedipalps and sternites X – XI; 10, holotype male, pygidium, dorsal posterior aspect; 11, paratype male (ICN-Ari- 011), dorsal aspect of left chelicera; 12, paratype female (ICN-Ari- 012), genital lip and spermathecae, anterior aspect; 13, paratype female, genital lip and spermathecae, posterior aspect." httpUri="https://zenodo.org/record/231254/files/figure.png" pageId="3" pageNumber="124">Fig. 10</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDDF8438C525C7CFF3AFB6B49F4FAEA" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="124">
|
||
<emphasis id="B91624518C525C7CFF3AFB6B4A7AFB7A" box="[199,318,1232,1256]" italics="true" pageId="3" pageNumber="124">Pedipalps.</emphasis>
|
||
Without cuticular pits; with some minute tubercles on ventral surface of trochanters I and II; femur curved and widened in basal half; approximately the same shape in male and female (no distinct sexual dimorphism); femur without any trace of basal tubercle in either sex; tibia longer than femur, slightly widened in basal third (
|
||
<figureCitation id="1359E4C68C525C7CFED8FA874AD2FAC6" box="[293,406,1340,1365]" captionStart="FIGURES 8 – 13" captionStartId="6.[151,264,1828,1850]" captionTargetBox="[151,1436,193,1806]" captionTargetId="figure@6.[151,1436,193,1807]" captionTargetPageId="6" captionText="FIGURES 8 – 13. Cryptocellus sofiae sp. nov. 8, holotype male (ICN-Ari- 009), sternal region, pedipalps and sternites X – XI; 9, paratype female (ICN-Ari- 012), sternal region, pedipalps and sternites X – XI; 10, holotype male, pygidium, dorsal posterior aspect; 11, paratype male (ICN-Ari- 011), dorsal aspect of left chelicera; 12, paratype female (ICN-Ari- 012), genital lip and spermathecae, anterior aspect; 13, paratype female, genital lip and spermathecae, posterior aspect." httpUri="https://zenodo.org/record/231254/files/figure.png" pageId="3" pageNumber="124">Figs. 8–9</figureCitation>
|
||
); both movable and fixed claws armed with minute teeth, more evident on fixed claw; movable claw noticeably larger than fixed claw.
|
||
</paragraph>
|
||
<paragraph id="8BDDF8438C525C7CFF3AFA3E4FFEFA77" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="124">
|
||
<emphasis id="B91624518C525C7CFF3AFA3E4A47FA0E" box="[199,259,1413,1436]" italics="true" pageId="3" pageNumber="124">Legs.</emphasis>
|
||
Legs I–IV similar in shape in male and female (
|
||
<figureCitation id="1359E4C68C525C7CFCDDFA3F48C9FA0F" box="[800,909,1412,1437]" captionStart="FIGURES 2 – 7" captionStartId="5.[151,264,1825,1847]" captionTargetBox="[151,1436,193,1802]" captionTargetId="figure@5.[151,1436,193,1804]" captionTargetPageId="5" captionText="FIGURES 2 – 7. Cryptocellus sofiae sp. nov. 2, holotype male (ICN-Ari- 009), dorsal; 3, holotype male, ventral; 4, paratype female (ICN-Ari- 010), dorsal; 5, paratype female, ventral (some setae accidentally removed from the opisthosoma); 6, paratype female (ICN-Ari- 012), frontal aspect (median longitudinal furrow of carapace indicated by arrow); 7, holotype male, polygonal setae on tergites XI and XII." httpUri="https://zenodo.org/record/231253/files/figure.png" pageId="3" pageNumber="124">Figs. 2–5</figureCitation>
|
||
); without cuticular pits and with few tubercles; completely devoid of any spine-like projection or similar structures; leg II not widened; leg III metatarsus not inflated (noticeably longer than wide), trochanter IV unmodified; tarsal claws strongly curved.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C378ABC88C525C77FF3AFA4B4ED2FC54" lastPageId="8" lastPageNumber="129" pageId="3" pageNumber="124" type="description">
|
||
<paragraph id="8BDDF8438C525C7CFF3AFA4B48D5F8BA" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="124">
|
||
<emphasis id="B91624518C525C7CFF3AFA4B494DF99A" box="[199,521,1520,1544]" italics="true" pageId="3" pageNumber="124">Male copulatory apparatus.</emphasis>
|
||
Tarsal process connected basally to a bone-shaped (phalanx-like) base. Tarsal process bulkier than accessory piece, spoon-like, with a moderate ventral expansion in distal half forming a curved cavity along the prolateral surface where the accessory piece rests; apically with two distinct lobes (L1 and L2) forming a single canal. Accessory piece movably jointed with basal prolateral aspect of tarsal process; progressively becoming thinner and bifurcated into two prominent, sharp-pointed lobes (L ′ and L″) of similar length (snake-tongue-shaped); the accessory piece is rolled up, leaving visible the sperm transfer groove as a foldlike gap running longitudinally from the base towards the tip of L″. The bifurcation of the piece into L ′ and L″ starts at about one third of the piece from the base (its beginning is indicated by an arrow in
|
||
<figureCitation id="1359E4C68C525C7CFB47F9574E00F897" box="[1210,1348,1772,1797]" captionStart="FIGURES 14 – 20" captionStartId="7.[151,264,1672,1694]" captionTargetBox="[151,1436,194,1645]" captionTargetId="figure@7.[151,1436,193,1651]" captionTargetPageId="7" captionText="FIGURES 14 – 20. Cryptocellus sofiae sp. nov., SEM micrographs. 14 – 18, paratype male (ICN-Ari- 011), right leg III; 14, prolateral aspect; 15, copulatory apparatus, prolateral aspect; 16, prolaterodorsal aspect; 17, apex of tarsal process and accessory piece; 18, level of junction between accessory piece and tarsal process; 19 – 20, paratype female (ICN-Ari- 012), spermathecae; 19, posterior aspect; 20, inclined posterior aspect. Abbreviations: ac, accessory piece (sperm transfer groove indicated by arrowheads, beginning of bifurcation indicated by arrow); Lc, lamina cyathiformis; L 1 – 2, lobes of tarsal process; L ′, L ″, lobes of accessory piece; mc, medial constriction of spermathecae; mP, metatarsal process; MT, metatarsus; pe, prolateral expansion of tarsal process; tP, tarsal process." httpUri="https://zenodo.org/record/231255/files/figure.png" pageId="3" pageNumber="124">Figs. 15–17</figureCitation>
|
||
). Tarsal process and accessory piece strongly curved upwards (
|
||
<figureCitation id="1359E4C68C525C7CFD06F8B448C0F8BA" box="[763,900,1807,1832]" captionStart="FIGURES 14 – 20" captionStartId="7.[151,264,1672,1694]" captionTargetBox="[151,1436,194,1645]" captionTargetId="figure@7.[151,1436,193,1651]" captionTargetPageId="7" captionText="FIGURES 14 – 20. Cryptocellus sofiae sp. nov., SEM micrographs. 14 – 18, paratype male (ICN-Ari- 011), right leg III; 14, prolateral aspect; 15, copulatory apparatus, prolateral aspect; 16, prolaterodorsal aspect; 17, apex of tarsal process and accessory piece; 18, level of junction between accessory piece and tarsal process; 19 – 20, paratype female (ICN-Ari- 012), spermathecae; 19, posterior aspect; 20, inclined posterior aspect. Abbreviations: ac, accessory piece (sperm transfer groove indicated by arrowheads, beginning of bifurcation indicated by arrow); Lc, lamina cyathiformis; L 1 – 2, lobes of tarsal process; L ′, L ″, lobes of accessory piece; mc, medial constriction of spermathecae; mP, metatarsal process; MT, metatarsus; pe, prolateral expansion of tarsal process; tP, tarsal process." httpUri="https://zenodo.org/record/231255/files/figure.png" pageId="3" pageNumber="124">Figs. 14–18</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDDF8438C525C7CFF3AF88F4AA4F8E2" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="124">
|
||
<emphasis id="B91624518C525C7CFF3AF88F4A29F8DE" box="[199,365,1844,1868]" italics="true" pageId="3" pageNumber="124">Spermathecae.</emphasis>
|
||
Touching together, slightly wider than long, constricted medially to give a rounded protruding section (
|
||
<figureCitation id="1359E4C68C525C7CFF08F8EC4A39F8E2" box="[245,381,1879,1904]" captionStart="FIGURES 8 – 13" captionStartId="6.[151,264,1828,1850]" captionTargetBox="[151,1436,193,1806]" captionTargetId="figure@6.[151,1436,193,1807]" captionTargetPageId="6" captionText="FIGURES 8 – 13. Cryptocellus sofiae sp. nov. 8, holotype male (ICN-Ari- 009), sternal region, pedipalps and sternites X – XI; 9, paratype female (ICN-Ari- 012), sternal region, pedipalps and sternites X – XI; 10, holotype male, pygidium, dorsal posterior aspect; 11, paratype male (ICN-Ari- 011), dorsal aspect of left chelicera; 12, paratype female (ICN-Ari- 012), genital lip and spermathecae, anterior aspect; 13, paratype female, genital lip and spermathecae, posterior aspect." httpUri="https://zenodo.org/record/231254/files/figure.png" pageId="3" pageNumber="124">Figs. 12–13</figureCitation>
|
||
,
|
||
<figureCitation id="1359E4C68C525C7CFE74F8E34A97F8E2" box="[393,467,1880,1904]" captionStart="FIGURES 14 – 20" captionStartId="7.[151,264,1672,1694]" captionTargetBox="[151,1436,194,1645]" captionTargetId="figure@7.[151,1436,193,1651]" captionTargetPageId="7" captionText="FIGURES 14 – 20. Cryptocellus sofiae sp. nov., SEM micrographs. 14 – 18, paratype male (ICN-Ari- 011), right leg III; 14, prolateral aspect; 15, copulatory apparatus, prolateral aspect; 16, prolaterodorsal aspect; 17, apex of tarsal process and accessory piece; 18, level of junction between accessory piece and tarsal process; 19 – 20, paratype female (ICN-Ari- 012), spermathecae; 19, posterior aspect; 20, inclined posterior aspect. Abbreviations: ac, accessory piece (sperm transfer groove indicated by arrowheads, beginning of bifurcation indicated by arrow); Lc, lamina cyathiformis; L 1 – 2, lobes of tarsal process; L ′, L ″, lobes of accessory piece; mc, medial constriction of spermathecae; mP, metatarsal process; MT, metatarsus; pe, prolateral expansion of tarsal process; tP, tarsal process." httpUri="https://zenodo.org/record/231255/files/figure.png" pageId="3" pageNumber="124">19–20</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDDF8438C525C7BFF3AF8C64BBBFEF7" blockId="3.[151,1437,151,2013]" lastBlockId="4.[151,1437,151,789]" lastPageId="4" lastPageNumber="125" pageId="3" pageNumber="124">
|
||
<emphasis id="B91624518C525C7CFF3AF8C64A9CF801" box="[199,472,1916,1940]" italics="true" pageId="3" pageNumber="124">Measurements (in mm).</emphasis>
|
||
Male
|
||
<typeStatus id="54D946E18C525C7CFDDAF8C749CDF807" box="[551,649,1916,1941]" pageId="3" pageNumber="124" type="holotype">holotype</typeStatus>
|
||
(ICN-Ari-009) and female
|
||
<typeStatus id="54D946E18C525C7CFC33F8C64F6AF807" box="[974,1070,1917,1941]" pageId="3" pageNumber="124" type="paratype">paratype</typeStatus>
|
||
(ICN-Ari-010, in parentheses): body length, excluding pygidium 3.06 (3.50); cucullus 0.50 (0.58) long, greatest width 0.66 (0.74); carapace 1.03 (1.17) long, 1.06 (1.15) wide at level of leg III (where widest); opisthosoma 1.56 (1.75) long (excluding pygidium), 1.39 (1.57) wide at level of junction between tergites XI and XII (where widest); median plate of tergite XI 0.44 (0.49) long, 0.83 (0.97) wide (where widest); median plate of tergite XII 0.39 (0.43) long, 0.67 (0.74) wide; median plate of tergite XIII 0.53 (0.60) long, 0.56 (0.62) wide; suture line of coxae II 0.08 (0.08) long, of coxae III 0.14 (0.13) long; pedipalp femur 0.49 (0.52) long, greatest diameter 0.22 (0.24); pedipalp tibia 0.71 (0.87) long, greatest diameter 0.10 (0.12); femur I 0.47 (0.49) long, 0.21 (0.23) in diameter; femur II 0.69 (0.78) long, 0.21 (0.23) in diameter.
|
||
</paragraph>
|
||
<paragraph id="8BDDF8438C555C7BFF3AFED4484CFD8A" blockId="4.[151,1437,151,789]" pageId="4" pageNumber="125">
|
||
<emphasis id="B91624518C555C7BFF3AFED449CFFE1A" bold="true" box="[199,651,367,392]" pageId="4" pageNumber="125">Generic placement of the new species.</emphasis>
|
||
<taxonomicName id="4C6283C08C555C7BFD6FFECB482FFE1A" box="[658,875,368,392]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="4" pageNumber="125" phylum="Arthropoda" rank="species" species="sofiae" status="sp. nov.">
|
||
<emphasis id="B91624518C555C7BFD6FFECB482FFE1A" box="[658,875,368,392]" italics="true" pageId="4" pageNumber="125">Cryptocellus sofiae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A225992A8C555C7BFC8FFED44888FE1A" box="[882,972,367,392]" pageId="4" pageNumber="125" rank="species">
|
||
<emphasis id="B91624518C555C7BFC8FFED44888FE1A" bold="true" box="[882,972,367,392]" pageId="4" pageNumber="125">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, is herein assigned to
|
||
<taxonomicName id="4C6283C08C555C7BFB3BFECB4E11FE1A" box="[1222,1365,368,392]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="4" pageNumber="125" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91624518C555C7BFB3BFECB4E11FE1A" box="[1222,1365,368,392]" italics="true" pageId="4" pageNumber="125">Cryptocellus</emphasis>
|
||
</taxonomicName>
|
||
based on the presence of morphological features of this genus as defined by
|
||
<bibRefCitation id="EFF385B28C555C7BFC1EFE2F4FDDFE3E" author="Platnick" box="[995,1177,404,429]" pageId="4" pageNumber="125" refString="Platnick, N. I. (1980) On the phylogeny of Ricinulei. In: Gruber, J. (Ed), Verhandlungen des 8. Internationalen Arachnologen- Kongress, Wien. H. Egermann, Wien, pp. 349 - 353." type="journal article" year="1980">Platnick (1980)</bibRefCitation>
|
||
: body short and wide (median plate of tergite XII wider than long); and ocular areas small in size and directed laterally at the edge of the carapace near the second legs. Some aspects of its morphology do not completely match the features described for
|
||
<taxonomicName id="4C6283C08C555C7BFF6AFDBB4A62FD8A" box="[151,294,512,536]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="4" pageNumber="125" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91624518C555C7BFF6AFDBB4A62FD8A" box="[151,294,512,536]" italics="true" pageId="4" pageNumber="125">Cryptocellus</emphasis>
|
||
</taxonomicName>
|
||
, however, and are briefly discussed below.
|
||
</paragraph>
|
||
<paragraph id="8BDDF8438C555C7BFF3AFD9F4901FC86" blockId="4.[151,1437,151,789]" pageId="4" pageNumber="125">
|
||
The coloration pattern of the cuticle of
|
||
<taxonomicName id="4C6283C08C555C7BFD86FD9F499AFDAE" box="[635,734,548,572]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="4" pageNumber="125" phylum="Arthropoda" rank="species" species="sofiae" status="sp. nov.">
|
||
<emphasis id="B91624518C555C7BFD86FD9F499AFDAE" box="[635,734,548,572]" italics="true" pageId="4" pageNumber="125">C. sofiae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A225992A8C555C7BFD18FD9F487AFDAF" box="[741,830,548,573]" pageId="4" pageNumber="125" rank="species">
|
||
<emphasis id="B91624518C555C7BFD18FD9F487AFDAF" bold="true" box="[741,830,548,573]" pageId="4" pageNumber="125">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is chestnut to light reddish (
|
||
<figureCitation id="1359E4C68C555C7BFB84FD9F4FA0FDAE" box="[1145,1252,548,573]" captionStart="FIGURES 2 – 7" captionStartId="5.[151,264,1825,1847]" captionTargetBox="[151,1436,193,1802]" captionTargetId="figure@5.[151,1436,193,1804]" captionTargetPageId="5" captionText="FIGURES 2 – 7. Cryptocellus sofiae sp. nov. 2, holotype male (ICN-Ari- 009), dorsal; 3, holotype male, ventral; 4, paratype female (ICN-Ari- 010), dorsal; 5, paratype female, ventral (some setae accidentally removed from the opisthosoma); 6, paratype female (ICN-Ari- 012), frontal aspect (median longitudinal furrow of carapace indicated by arrow); 7, holotype male, polygonal setae on tergites XI and XII." httpUri="https://zenodo.org/record/231253/files/figure.png" pageId="4" pageNumber="125">Figs. 2–7</figureCitation>
|
||
,
|
||
<figureCitation id="1359E4C68C555C7BFB12FD9F4E6EFDAE" box="[1263,1322,548,572]" captionStart="FIGURES 8 – 13" captionStartId="6.[151,264,1828,1850]" captionTargetBox="[151,1436,193,1806]" captionTargetId="figure@6.[151,1436,193,1807]" captionTargetPageId="6" captionText="FIGURES 8 – 13. Cryptocellus sofiae sp. nov. 8, holotype male (ICN-Ari- 009), sternal region, pedipalps and sternites X – XI; 9, paratype female (ICN-Ari- 012), sternal region, pedipalps and sternites X – XI; 10, holotype male, pygidium, dorsal posterior aspect; 11, paratype male (ICN-Ari- 011), dorsal aspect of left chelicera; 12, paratype female (ICN-Ari- 012), genital lip and spermathecae, anterior aspect; 13, paratype female, genital lip and spermathecae, posterior aspect." httpUri="https://zenodo.org/record/231254/files/figure.png" pageId="4" pageNumber="125">8–10</figureCitation>
|
||
), which is a condition deemed diagnostic for
|
||
<taxonomicName id="4C6283C08C555C7BFDE1FDF349F4FDF2" box="[540,688,584,608]" class="Arachnida" family="Ricinoididae" genus="Pseudocellus" kingdom="Animalia" order="Ricinulei" pageId="4" pageNumber="125" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91624518C555C7BFDE1FDF349F4FDF2" box="[540,688,584,608]" italics="true" pageId="4" pageNumber="125">Pseudocellus</emphasis>
|
||
</taxonomicName>
|
||
species, as opposed to
|
||
<taxonomicName id="4C6283C08C555C7BFC4AFDF34F02FDF2" box="[951,1094,584,608]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="4" pageNumber="125" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91624518C555C7BFC4AFDF34F02FDF2" box="[951,1094,584,608]" italics="true" pageId="4" pageNumber="125">Cryptocellus</emphasis>
|
||
</taxonomicName>
|
||
whose members are supposed to display a dark reddish-black coloration (
|
||
<bibRefCitation id="EFF385B28C555C7BFD88FDD74851FD16" author="Platnick" box="[629,789,620,645]" pageId="4" pageNumber="125" refString="Platnick, N. I. (1980) On the phylogeny of Ricinulei. In: Gruber, J. (Ed), Verhandlungen des 8. Internationalen Arachnologen- Kongress, Wien. H. Egermann, Wien, pp. 349 - 353." type="journal article" year="1980">Platnick 1980</bibRefCitation>
|
||
). The light coloration of the new species is not considered to link it to
|
||
<taxonomicName id="4C6283C08C555C7BFED1FD2B4A84FD3A" box="[300,448,656,680]" class="Arachnida" family="Ricinoididae" genus="Pseudocellus" kingdom="Animalia" order="Ricinulei" pageId="4" pageNumber="125" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91624518C555C7BFED1FD2B4A84FD3A" box="[300,448,656,680]" italics="true" pageId="4" pageNumber="125">Pseudocellus</emphasis>
|
||
</taxonomicName>
|
||
, however, based on the above-referred features shared with other
|
||
<taxonomicName id="4C6283C08C555C7BFB2FFD2B4E25FD3A" box="[1234,1377,656,680]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="4" pageNumber="125" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91624518C555C7BFB2FFD2B4E25FD3A" box="[1234,1377,656,680]" italics="true" pageId="4" pageNumber="125">Cryptocellus</emphasis>
|
||
</taxonomicName>
|
||
, and because adults of other species in the later genus also display a color that is not particularly dark but rather chestnut-reddish [e.g.
|
||
<taxonomicName id="4C6283C08C555C7BFE6AFD634F12FD62" authority="Tourinho, Lo Man-Hung & Bonaldo, 2010" authorityName="Tourinho, Lo Man-Hung & Bonaldo" authorityYear="2010" box="[407,1110,727,752]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="4" pageNumber="125" phylum="Arthropoda" rank="species" species="iaci">
|
||
<emphasis id="B91624518C555C7BFE6AFD63491EFD7D" box="[407,602,728,752]" italics="true" pageId="4" pageNumber="125">Cryptocellus iaci</emphasis>
|
||
Tourinho, Lo Man-Hung &
|
||
<bibRefCitation id="EFF385B28C555C7BFC50FD6C4F12FD62" author="Bonaldo" box="[941,1110,727,752]" pageId="4" pageNumber="125" refString="Tourinho, A. L., Lo-Man-Hung, N. F. & Bonaldo, A. B. (2010) A new species of Ricinulei of the genus Cryptocellus Westwood (Arachnida) from northern Brazil. Zootaxa, 2684, 63 - 68." type="journal article" year="2010">Bonaldo, 2010</bibRefCitation>
|
||
</taxonomicName>
|
||
(Tourinho
|
||
<emphasis id="B91624518C555C7BFB27FD624E52FD62" box="[1242,1302,728,752]" italics="true" pageId="4" pageNumber="125">et al.</emphasis>
|
||
2010: figs. 7–12;
|
||
<bibRefCitation id="EFF385B28C555C7BFF20FD474A9DFC86" author="Salvatierra" box="[221,473,764,789]" pageId="4" pageNumber="125" refString="Salvatierra, L., Tourinho, A. L. & Giribet, G. (2013) Description of the male, larva and nymphal stages of Cryptocellus iaci (Arachnida, Ricinulei), with an overview of tarsal sensilla and other integumental structures. Zootaxa, 3709 (2), 149 - 161. http: // dx. doi. org / 10.11646 / zootaxa. 3709.2.3" type="journal article" year="2013">
|
||
Salvatierra
|
||
<emphasis id="B91624518C555C7BFEA3FD464AD2FC86" box="[350,406,764,788]" italics="true" pageId="4" pageNumber="125">et al.</emphasis>
|
||
2013
|
||
</bibRefCitation>
|
||
: fig. 2c)].
|
||
</paragraph>
|
||
<caption id="DF1DA8CB8C555C7BFF6AF80D4881F85E" box="[151,965,1974,1996]" httpUri="https://zenodo.org/record/231252/files/figure.png" pageId="4" pageNumber="125" targetBox="[230,1358,841,1944]" targetPageId="4">
|
||
<paragraph id="8BDDF8438C555C7BFF6AF80D4881F85E" blockId="4.[151,965,1974,1996]" box="[151,965,1974,1996]" pageId="4" pageNumber="125">
|
||
<emphasis id="B91624518C555C7BFF6AF80D4A57F85E" bold="true" box="[151,275,1974,1996]" pageId="4" pageNumber="125">FIGURE 1.</emphasis>
|
||
Known distribution records of the genus
|
||
<taxonomicName id="4C6283C08C555C7BFD45F80D487DF85E" box="[696,825,1974,1996]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="4" pageNumber="125" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91624518C555C7BFD45F80D487DF85E" box="[696,825,1974,1996]" italics="true" pageId="4" pageNumber="125">Cryptocellus</emphasis>
|
||
</taxonomicName>
|
||
in Colombia.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF1DA8CB8C545C7AFF6AF89A4AF2F801" httpUri="https://zenodo.org/record/231253/files/figure.png" pageId="5" pageNumber="126" targetBox="[151,1436,193,1802]" targetPageId="5">
|
||
<paragraph id="8BDDF8438C545C7AFF6AF89A4AF2F801" blockId="5.[151,1435,1825,1939]" pageId="5" pageNumber="126">
|
||
<emphasis id="B91624518C545C7AFF6AF89A4A78F8A5" bold="true" box="[151,316,1825,1847]" pageId="5" pageNumber="126">FIGURES 2–7.</emphasis>
|
||
<taxonomicName id="4C6283C08C545C7AFEB9F89A494EF8A5" box="[324,522,1825,1847]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="5" pageNumber="126" phylum="Arthropoda" rank="species" species="sofiae" status="sp. nov.">
|
||
<emphasis id="B91624518C545C7AFEB9F89A494EF8A5" box="[324,522,1825,1847]" italics="true" pageId="5" pageNumber="126">Cryptocellus sofiae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A225992A8C545C7AFDE9F89A4922F8A5" box="[532,614,1825,1847]" pageId="5" pageNumber="126" rank="species">
|
||
<emphasis id="B91624518C545C7AFDE9F89A4922F8A5" bold="true" box="[532,614,1825,1847]" pageId="5" pageNumber="126">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
2, holotype male (ICN-Ari-009), dorsal; 3, holotype male, ventral; 4, paratype female (ICN-Ari-010), dorsal; 5, paratype female, ventral (some setae accidentally removed from the opisthosoma); 6, paratype female (ICN-Ari-012), frontal aspect (median longitudinal furrow of carapace indicated by arrow); 7, holotype male, polygonal setae on tergites XI and XII.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF1DA8CB8C575C79FF6AF89F4F3CF804" httpUri="https://zenodo.org/record/231254/files/figure.png" pageId="6" pageNumber="127" targetBox="[151,1436,193,1806]" targetPageId="6">
|
||
<paragraph id="8BDDF8438C575C79FF6AF89F4F3CF804" blockId="6.[151,1436,1828,1942]" pageId="6" pageNumber="127">
|
||
<emphasis id="B91624518C575C79FF6AF89F4A03F8A8" bold="true" box="[151,327,1828,1850]" pageId="6" pageNumber="127">FIGURES 8–13.</emphasis>
|
||
<taxonomicName id="4C6283C08C575C79FEB1F89F4954F8A8" box="[332,528,1828,1850]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="6" pageNumber="127" phylum="Arthropoda" rank="species" species="sofiae" status="sp. nov.">
|
||
<emphasis id="B91624518C575C79FEB1F89F4954F8A8" box="[332,528,1828,1850]" italics="true" pageId="6" pageNumber="127">Cryptocellus sofiae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A225992A8C575C79FDEAF89F4923F8A8" box="[535,615,1828,1850]" pageId="6" pageNumber="127" rank="species">
|
||
<emphasis id="B91624518C575C79FDEAF89F4923F8A8" bold="true" box="[535,615,1828,1850]" pageId="6" pageNumber="127">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
8, holotype male (ICN-Ari-009), sternal region, pedipalps and sternites X–XI; 9, paratype female (ICN-Ari-012), sternal region, pedipalps and sternites X–XI; 10, holotype male, pygidium, dorsal posterior aspect; 11, paratype male (ICN-Ari-011), dorsal aspect of left chelicera; 12, paratype female (ICN-Ari-012), genital lip and spermathecae, anterior aspect; 13, paratype female, genital lip and spermathecae, posterior aspect.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF1DA8CB8C565C78FF6AF9334922F8C4" httpUri="https://zenodo.org/record/231255/files/figure.png" pageId="7" pageNumber="128" targetBox="[151,1436,194,1645]" targetPageId="7">
|
||
<paragraph id="8BDDF8438C565C78FF6AF9334922F8C4" blockId="7.[151,1436,1672,1879]" pageId="7" pageNumber="128">
|
||
<emphasis id="B91624518C565C78FF6AF9334A13F90C" bold="true" box="[151,343,1672,1694]" pageId="7" pageNumber="128">FIGURES 14–20.</emphasis>
|
||
<taxonomicName id="4C6283C08C565C78FE9DF9334963F90C" box="[352,551,1672,1694]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="7" pageNumber="128" phylum="Arthropoda" rank="species" species="sofiae" status="sp. nov.">
|
||
<emphasis id="B91624518C565C78FE9DF9334963F90C" box="[352,551,1672,1694]" italics="true" pageId="7" pageNumber="128">Cryptocellus sofiae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A225992A8C565C78FDCCF93349C1F90C" box="[561,645,1672,1694]" pageId="7" pageNumber="128" rank="species">
|
||
<emphasis id="B91624518C565C78FDCCF93349C1F90C" bold="true" box="[561,645,1672,1694]" pageId="7" pageNumber="128">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, SEM micrographs. 14–18, paratype male (ICN-Ari-011), right leg III; 14, prolateral aspect; 15, copulatory apparatus, prolateral aspect; 16, prolaterodorsal aspect; 17, apex of tarsal process and accessory piece; 18, level of junction between accessory piece and tarsal process; 19–20, paratype female (ICN-Ari-012), spermathecae; 19, posterior aspect; 20, inclined posterior aspect. Abbreviations:
|
||
<emphasis id="B91624518C565C78FC22F95E48B2F968" box="[991,1014,1765,1786]" italics="true" pageId="7" pageNumber="128">ac</emphasis>
|
||
, accessory piece (sperm transfer groove indicated by arrowheads, beginning of bifurcation indicated by arrow);
|
||
<emphasis id="B91624518C565C78FC8CF8BF48CEF88B" box="[881,906,1796,1817]" italics="true" pageId="7" pageNumber="128">Lc</emphasis>
|
||
, lamina cyathiformis;
|
||
<emphasis id="B91624518C565C78FB8CF8B84FE1F88B" box="[1137,1189,1795,1817]" italics="true" pageId="7" pageNumber="128">L1–2</emphasis>
|
||
, lobes of tarsal process;
|
||
<emphasis id="B91624518C565C78FF6AF8984BE1F8AA" box="[151,165,1827,1848]" italics="true" pageId="7" pageNumber="128">L</emphasis>
|
||
′,
|
||
<emphasis id="B91624518C565C78FF4BF8984B80F8AA" box="[182,196,1827,1848]" italics="true" pageId="7" pageNumber="128">L</emphasis>
|
||
″, lobes of accessory piece;
|
||
<emphasis id="B91624518C565C78FE21F8984ABDF8AA" box="[476,505,1827,1848]" italics="true" pageId="7" pageNumber="128">mc</emphasis>
|
||
, medial constriction of spermathecae;
|
||
<emphasis id="B91624518C565C78FC82F89848DAF8AA" box="[895,926,1827,1848]" italics="true" pageId="7" pageNumber="128">mP</emphasis>
|
||
, metatarsal process;
|
||
<emphasis id="B91624518C565C78FB8DF8984FD5F8AA" box="[1136,1169,1827,1848]" italics="true" pageId="7" pageNumber="128">MT</emphasis>
|
||
, metatarsus;
|
||
<emphasis id="B91624518C565C78FAEAF8984E6BF8AA" box="[1303,1327,1827,1848]" italics="true" pageId="7" pageNumber="128">pe</emphasis>
|
||
, prolateral expansion of tarsal process;
|
||
<emphasis id="B91624518C565C78FE48F8F94A8DF8C5" box="[437,457,1858,1879]" italics="true" pageId="7" pageNumber="128">tP</emphasis>
|
||
, tarsal process.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF1DA8CB8C595C77FF6AFD074824FD62" httpUri="https://zenodo.org/record/231256/files/figure.png" pageId="8" pageNumber="129" targetBox="[151,1435,197,673]" targetPageId="8">
|
||
<paragraph id="8BDDF8438C595C77FF6AFD074824FD62" blockId="8.[151,1436,700,752]" pageId="8" pageNumber="129">
|
||
<emphasis id="B91624518C595C77FF6AFD074A11FD40" bold="true" box="[151,341,700,722]" pageId="8" pageNumber="129">FIGURES 21–22.</emphasis>
|
||
<taxonomicName id="4C6283C08C595C77FEA0FD074966FD40" box="[349,546,700,722]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="8" pageNumber="129" phylum="Arthropoda" rank="species" species="sofiae" status="sp. nov.">
|
||
<emphasis id="B91624518C595C77FEA0FD074966FD40" box="[349,546,700,722]" italics="true" pageId="8" pageNumber="129">Cryptocellus sofiae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A225992A8C595C77FDD6FD074939FD40" box="[555,637,700,722]" pageId="8" pageNumber="129" rank="species">
|
||
<emphasis id="B91624518C595C77FDD6FD074939FD40" bold="true" box="[555,637,700,722]" pageId="8" pageNumber="129">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, SEM micrographs, different setal types. 21, paratype female (ICN-Ari-012), part of tergite XIII; 22, paratype male (ICN-Ari-011), part of cucullus.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDDF8438C595C77FF3AFCA7498FFCEF" blockId="8.[151,1436,796,966]" pageId="8" pageNumber="129">
|
||
Although the bifurcate accessory piece of the copulatory apparatus in the male
|
||
<taxonomicName id="4C6283C08C595C77FBC6FCA64FDBFCA7" box="[1083,1183,797,821]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="8" pageNumber="129" phylum="Arthropoda" rank="species" species="sofiae" status="sp. nov.">
|
||
<emphasis id="B91624518C595C77FBC6FCA64FDBFCA7" box="[1083,1183,797,821]" italics="true" pageId="8" pageNumber="129">C. sofiae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A225992A8C595C77FB58FCA74FBAFCA7" box="[1189,1278,796,821]" pageId="8" pageNumber="129" rank="species">
|
||
<emphasis id="B91624518C595C77FB58FCA74FBAFCA7" bold="true" box="[1189,1278,796,821]" pageId="8" pageNumber="129">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="1359E4C68C595C77FAF0FCA74ED7FCA7" box="[1293,1427,796,821]" captionStart="FIGURES 14 – 20" captionStartId="7.[151,264,1672,1694]" captionTargetBox="[151,1436,194,1645]" captionTargetId="figure@7.[151,1436,193,1651]" captionTargetPageId="7" captionText="FIGURES 14 – 20. Cryptocellus sofiae sp. nov., SEM micrographs. 14 – 18, paratype male (ICN-Ari- 011), right leg III; 14, prolateral aspect; 15, copulatory apparatus, prolateral aspect; 16, prolaterodorsal aspect; 17, apex of tarsal process and accessory piece; 18, level of junction between accessory piece and tarsal process; 19 – 20, paratype female (ICN-Ari- 012), spermathecae; 19, posterior aspect; 20, inclined posterior aspect. Abbreviations: ac, accessory piece (sperm transfer groove indicated by arrowheads, beginning of bifurcation indicated by arrow); Lc, lamina cyathiformis; L 1 – 2, lobes of tarsal process; L ′, L ″, lobes of accessory piece; mc, medial constriction of spermathecae; mP, metatarsal process; MT, metatarsus; pe, prolateral expansion of tarsal process; tP, tarsal process." httpUri="https://zenodo.org/record/231255/files/figure.png" pageId="8" pageNumber="129">Figs. 14–17</figureCitation>
|
||
) resembles that of some
|
||
<taxonomicName id="4C6283C08C595C77FE5DFCFA4970FCCB" box="[416,564,833,857]" class="Arachnida" family="Ricinoididae" genus="Pseudocellus" kingdom="Animalia" order="Ricinulei" pageId="8" pageNumber="129" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91624518C595C77FE5DFCFA4970FCCB" box="[416,564,833,857]" italics="true" pageId="8" pageNumber="129">Pseudocellus</emphasis>
|
||
</taxonomicName>
|
||
species, this condition is now also known to occur in some other members of
|
||
<taxonomicName id="4C6283C08C595C77FF6AFCDE4A62FCEF" box="[151,294,869,893]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="8" pageNumber="129" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91624518C595C77FF6AFCDE4A62FCEF" box="[151,294,869,893]" italics="true" pageId="8" pageNumber="129">Cryptocellus</emphasis>
|
||
</taxonomicName>
|
||
, as discussed at the end of this paper.
|
||
</paragraph>
|
||
<paragraph id="8BDDF8438C595C77FF3AFC324ED2FC54" blockId="8.[151,1436,796,966]" pageId="8" pageNumber="129">
|
||
Based on these considerations, and taking into account the fact that no species of
|
||
<taxonomicName id="4C6283C08C595C77FBA8FC324FADFC33" box="[1109,1257,905,929]" class="Arachnida" family="Ricinoididae" genus="Pseudocellus" kingdom="Animalia" order="Ricinulei" pageId="8" pageNumber="129" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91624518C595C77FBA8FC324FADFC33" box="[1109,1257,905,929]" italics="true" pageId="8" pageNumber="129">Pseudocellus</emphasis>
|
||
</taxonomicName>
|
||
is so far known to inhabit South
|
||
<collectingCountry id="F375B8D38C595C77FEACFC164AF0FC54" box="[337,436,941,966]" name="United States of America" pageId="8" pageNumber="129">America</collectingCountry>
|
||
, the new species is here considered to be a member of
|
||
<taxonomicName id="4C6283C08C595C77FBE7FC164FEDFC57" box="[1050,1193,941,965]" class="Arachnida" family="Ricinoididae" genus="Cryptocellus" kingdom="Animalia" order="Ricinulei" pageId="8" pageNumber="129" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91624518C595C77FBE7FC164FEDFC57" box="[1050,1193,941,965]" italics="true" pageId="8" pageNumber="129">Cryptocellus</emphasis>
|
||
</taxonomicName>
|
||
as currently defined.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |