treatments-xml/data/03/C5/EA/03C5EA23AE43FFB554C2F9E0A42A6C40.xml
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<document ID-DOI="http://doi.org/10.5281/zenodo.5698050" ID-GBIF-Dataset="44b12099-59c0-4c00-b34c-c4f14d37ec56" ID-GBIF-Taxon="135476927" ID-Zenodo-Dep="5698050" checkinTime="1509096787603" checkinUser="plazi" docAuthor="Jiří Šmíd, Jiří Moravec, Václav Gvoždík, Jan Štundl, Daniel Frynta, Petros Lymberakis, Paschalia Kapli, Thomas Wilms, Andreas Schmitz, Mohammed Shobrak, Saeed Hosseinian Yousefkhani, Eskandar Rastegar-Pouyani, Aurora M. Castilla, Johannes Els &amp; Werner Mayer" docDate="2017" docId="03C5EA23AE43FFB554C2F9E0A42A6C40" docLanguage="en" docName="Smid, J. et al. 2017.pdf" docOrigin="Zoologica Scripta 46 (6)" docStyle="DocumentStyle{}" docTitle="Mesalina saudiarabica Moravec, Smid, Schmitz, Shobrak, Wilms" docType="treatment" docVersion="7" lastPageId="19" lastPageNumber="20" masterDocId="FFFC925BAE52FFA6547EFFDBA00A6849" masterDocTitle="Cutting the Gordian Knot: Phylogenetic and ecological diversification of the Mesalina brevirostris species complex (Squamata, Lacertidae)" masterLastPageNumber="30" masterPageNumber="1" pageId="17" pageNumber="18" updateTime="1636832576005" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Cutting the Gordian Knot: Phylogenetic and ecological diversification of the Mesalina brevirostris species complex (Squamata, Lacertidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jiří Šmíd</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jiří Moravec</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Václav Gvoždík</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jan Štundl</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Daniel Frynta</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Petros Lymberakis</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Paschalia Kapli</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Thomas Wilms</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Andreas Schmitz</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mohammed Shobrak</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Saeed Hosseinian Yousefkhani</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Eskandar Rastegar-Pouyani</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Aurora M. Castilla</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Johannes Els</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Werner Mayer</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zoologica Scripta</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2017</mods:date>
<mods:detail type="pubDate">
<mods:number>2017-10-31</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>46</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>6</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>30</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1111/zsc.12254</mods:identifier>
<mods:identifier type="GBIF-Dataset">44b12099-59c0-4c00-b34c-c4f14d37ec56</mods:identifier>
<mods:identifier type="Zenodo-Dep">1037920</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.5698050" ID-GBIF-Taxon="135476927" ID-Zenodo-Dep="5698050" LSID="urn:lsid:plazi:treatment:03C5EA23AE43FFB554C2F9E0A42A6C40" httpUri="http://treatment.plazi.org/id/03C5EA23AE43FFB554C2F9E0A42A6C40" lastPageId="19" lastPageNumber="20" pageId="17" pageNumber="18">
<subSubSection box="[188,1174,1594,1623]" pageId="17" pageNumber="18" type="nomenclature">
<paragraph blockId="17.[188,1174,1594,1696]" box="[188,1174,1594,1623]" pageId="17" pageNumber="18">
<heading bold="true" box="[188,1174,1594,1623]" fontSize="12" level="4" pageId="17" pageNumber="18" reason="0">
<emphasis bold="true" box="[188,1174,1594,1623]" pageId="17" pageNumber="18">
<taxonomicName authority="Moravec, Smid, Schmitz, Shobrak, Wilms" authorityName="Moravec, Smid, Schmitz, Shobrak, Wilms" box="[188,1070,1594,1623]" class="Reptilia" family="Lacertidae" genus="Mesalina" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="saudiarabica">
<taxonomicName authority="Moravec, Smid, Schmitz, Shobrak" authorityName="Moravec, Smid, Schmitz, Shobrak" box="[188,968,1594,1623]" class="Reptilia" family="Lacertidae" genus="Mesalina" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="saudiarabica">
<taxonomicName authority="Moravec, Smid, Schmitz" authorityName="Moravec, Smid, Schmitz" box="[188,833,1594,1623]" class="Reptilia" family="Lacertidae" genus="Mesalina" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="saudiarabica">
<taxonomicName authority="Moravec, Smid" authorityName="Moravec, Smid" box="[188,707,1594,1623]" class="Reptilia" family="Lacertidae" genus="Mesalina" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="saudiarabica">
<taxonomicName authority="Moravec" authorityName="Moravec" box="[188,621,1594,1623]" class="Reptilia" family="Lacertidae" genus="Mesalina" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="saudiarabica">
<taxonomicName box="[188,498,1595,1623]" class="Reptilia" family="Lacertidae" genus="Mesalina" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="saudiarabica">
<emphasis bold="true" box="[188,498,1595,1623]" italics="true" pageId="17" pageNumber="18">
<taxonomicName box="[188,311,1595,1623]" class="Reptilia" family="Lacertidae" genus="Mesalina" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="18" phylum="Chordata" rank="genus">Mesalina</taxonomicName>
saudiarabica
</emphasis>
</taxonomicName>
Moravec
</taxonomicName>
, Šmíd
</taxonomicName>
, Schmitz
</taxonomicName>
, Shobrak
</taxonomicName>
, Wilms
</taxonomicName>
sp. n.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="17" pageNumber="18" type="reference_group">
<paragraph blockId="17.[188,1174,1594,1696]" box="[188,733,1631,1659]" pageId="17" pageNumber="18">
<heading box="[188,733,1631,1659]" fontSize="12" level="6" pageId="17" pageNumber="18" reason="1">
<taxonomicName authorityName="Jiří Šmíd &amp; Jiří Moravec &amp; Václav Gvoždík &amp; Jan Štundl &amp; Daniel Frynta &amp; Petros Lymberakis &amp; Paschalia Kapli &amp; Thomas Wilms &amp; Andreas Schmitz &amp; Mohammed Shobrak &amp; Saeed Hosseinian Yousefkhani &amp; Eskandar Rastegar-Pouyani &amp; Aurora M. Castilla &amp; Johannes Els &amp; Werner Mayer" authorityYear="2017" box="[188,464,1631,1659]" class="Reptilia" family="Lacertidae" genus="Mesalina" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="brevirostris">
<emphasis box="[188,464,1631,1659]" italics="true" pageId="17" pageNumber="18">
<taxonomicName box="[188,307,1631,1659]" class="Reptilia" family="Lacertidae" genus="Mesalina" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="18" phylum="Chordata" rank="genus">Mesalina</taxonomicName>
brevirostris
</emphasis>
</taxonomicName>
Kapli
<emphasis box="[576,639,1631,1659]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
(2015)
</heading>
</paragraph>
<paragraph blockId="17.[188,1174,1594,1696]" box="[188,620,1668,1696]" pageId="17" pageNumber="18">
<heading box="[188,620,1668,1696]" fontSize="12" level="6" pageId="17" pageNumber="18" reason="1">
<taxonomicName authorityName="Gray" authorityYear="1838" box="[188,307,1668,1696]" class="Reptilia" family="Lacertidae" genus="Mesalina" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="18" phylum="Chordata" rank="genus">
<emphasis box="[188,307,1668,1696]" italics="true" pageId="17" pageNumber="18">Mesalina</emphasis>
</taxonomicName>
sp. Kapli
<emphasis box="[463,519,1668,1696]" italics="true" pageId="17" pageNumber="18">et al</emphasis>
. (
<date box="[544,611,1668,1696]" pageId="17" pageNumber="18" value="2015">2015</date>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="materials_examined">
<paragraph blockId="17.[188,1407,1741,1843]" pageId="17" pageNumber="18">
<materialsCitation ID-GBIF-Occurrence="1676080761" collectingDate="2006-10" collectionCode="ZFMK" collectorName="T. Wilms. MorphoBank" country="Saudi Arabia" elevation="1000" latitude="22.237" location="Mahazat" longLatPrecision="75" longitude="41.843" pageId="17" pageNumber="18" specimenCode="ZFMK 91912" specimenCount="1" stateProvince="Makkah Province" typeStatus="holotype">
<typeStatus box="[188,311,1741,1769]" pageId="17" pageNumber="18">Holotype</typeStatus>
.
<specimenCode box="[327,511,1741,1769]" collectionCode="ZFMK" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="17" pageNumber="18">ZFMK 91912</specimenCode>
, subadult male,
<location LSID="urn:lsid:plazi:treatment:03C5EA23AE43FFB554C2F9E0A42A6C40:8EB30DEEAE43FFB756A6F916A34C6EA0" box="[728,838,1741,1769]" country="Saudi Arabia" latitude="22.237" longLatPrecision="75" longitude="41.843" name="Mahazat" pageId="17" pageNumber="18" stateProvince="Makkah Province">Mahazat</location>
as­Sayd,
<collectingRegion box="[970,1204,1741,1769]" country="Saudi Arabia" name="Makkah" pageId="17" pageNumber="18">Makkah Province</collectingRegion>
,
<collectingCountry box="[1220,1402,1741,1769]" name="Saudi Arabia" pageId="17" pageNumber="18">Saudi Arabia</collectingCountry>
,
<geoCoordinate box="[188,316,1778,1806]" direction="north" orientation="latitude" pageId="17" pageNumber="18" precision="55" value="22.237">22.237 N</geoCoordinate>
,
<geoCoordinate box="[335,459,1778,1806]" direction="east" orientation="longitude" pageId="17" pageNumber="18" precision="55" value="41.843">41.843 E</geoCoordinate>
,
<elevation box="[473,650,1778,1806]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="17" pageNumber="18" unit="m" value="1000.0">
<quantity box="[473,576,1778,1806]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="17" pageNumber="18" unit="m" value="1000.0">1000 m</quantity>
a.s.l.
</elevation>
, collected in
<date box="[839,1020,1778,1806]" pageId="17" pageNumber="18" value="2006-10">
<collectingDate box="[839,1020,1778,1806]" pageId="17" pageNumber="18" value="2006-10">October 2006</collectingDate>
</date>
by
<collectorName box="[1080,1407,1778,1806]" pageId="17" pageNumber="18">T. Wilms. MorphoBank</collectorName>
pictures: M407306M407312.
</materialsCitation>
</paragraph>
<paragraph blockId="17.[188,1408,1888,1990]" pageId="17" pageNumber="18">
<materialsCitation ID-GBIF-Occurrence="1676080762" collectingDate="2006-10" collectionCode="ZFMK" collectorName="T. Wilms. MorphoBank" country="Saudi Arabia" elevation="960" latitude="22.395" location="Mahazat" longLatPrecision="75" longitude="41.753" municipality="Al Muwayh" pageId="17" pageNumber="18" specimenCode="ZFMK 86583" specimenCount="1" stateProvince="Makkah Province" typeStatus="paratype">
<typeStatus box="[188,304,1888,1916]" pageId="17" pageNumber="18">Paratype</typeStatus>
.
<specimenCode box="[328,520,1888,1916]" collectionCode="ZFMK" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="17" pageNumber="18">ZFMK 86583</specimenCode>
, subadult male,
<location LSID="urn:lsid:plazi:treatment:03C5EA23AE43FFB554C2F9E0A42A6C40:8EB30DEEAE43FFB75682F8BBA3606F35" box="[764,874,1888,1916]" country="Saudi Arabia" latitude="22.395" longLatPrecision="75" longitude="41.753" municipality="Al Muwayh" name="Mahazat" pageId="17" pageNumber="18" stateProvince="Makkah Province">Mahazat</location>
as­Sayd, near
<collectingMunicipality box="[1100,1269,1888,1916]" pageId="17" pageNumber="18">Al Muwayh</collectingMunicipality>
,
<collectingRegion country="Saudi Arabia" name="Makkah" pageId="17" pageNumber="18">Makkah Province</collectingRegion>
,
<collectingCountry box="[322,498,1925,1953]" name="Saudi Arabia" pageId="17" pageNumber="18">Saudi Arabia</collectingCountry>
,
<geoCoordinate box="[516,639,1925,1953]" direction="north" orientation="latitude" pageId="17" pageNumber="18" precision="55" value="22.395">22.395 N</geoCoordinate>
<geoCoordinate box="[652,774,1925,1953]" direction="east" orientation="longitude" pageId="17" pageNumber="18" precision="55" value="41.753">41.753 E</geoCoordinate>
,
<elevation box="[792,951,1925,1953]" metricMagnitude="2" metricUnit="m" metricValue="9.6" pageId="17" pageNumber="18" unit="m" value="960.0">
<quantity box="[792,878,1925,1953]" metricMagnitude="2" metricUnit="m" metricValue="9.6" pageId="17" pageNumber="18" unit="m" value="960.0">960 m</quantity>
a.s.l.
</elevation>
, collected in
<date box="[1135,1315,1925,1953]" pageId="17" pageNumber="18" value="2006-10">
<collectingDate box="[1135,1315,1925,1953]" pageId="17" pageNumber="18" value="2006-10">October 2006</collectingDate>
</date>
by
<collectorName pageId="17" pageNumber="18">
<collectionCode box="[1372,1402,1925,1953]" country="0" httpUri="http://grbio.org/cool/ey0i-ifzt" name="Tavera, Department of Geology and Geophysics" pageId="17" pageNumber="18">T</collectionCode>
. Wilms. MorphoBank
</collectorName>
picture: M410851.
</materialsCitation>
</paragraph>
<paragraph blockId="18.[188,1407,196,261]" pageId="18" pageNumber="19">
Referred material not included in the
<typeStatus box="[683,738,196,224]" pageId="18" pageNumber="19">type</typeStatus>
series.
<collectionCode box="[840,911,196,224]" country="South Africa" httpUri="http://grbio.org/cool/fx6k-enrh" name="Natal Museum" pageId="18" pageNumber="19">NMP</collectionCode>
6
<collectionCode box="[926,944,196,224]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15707" name="Field Museum of Natural History, Botany Department" pageId="18" pageNumber="19">F</collectionCode>
29­30 (photovoucher), adult male, observed on the
<typeStatus box="[398,453,233,261]" pageId="18" pageNumber="19">type</typeStatus>
locality in
<date box="[598,773,233,261]" pageId="18" pageNumber="19" value="2006-10">October 2006</date>
by
<collectionCode box="[821,844,233,261]" country="0" httpUri="http://grbio.org/cool/ey0i-ifzt" name="Tavera, Department of Geology and Geophysics" pageId="18" pageNumber="19">T</collectionCode>
. Wilms.
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="19" type="diagnosis">
<paragraph blockId="18.[188,1408,306,813]" pageId="18" pageNumber="19">
Diagnosis.
<collectionCode box="[336,359,306,334]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="18" pageNumber="19">A</collectionCode>
species of
<taxonomicName authorityName="Gray" authorityYear="1838" box="[515,634,306,334]" class="Reptilia" family="Lacertidae" genus="Mesalina" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis box="[515,634,306,334]" italics="true" pageId="18" pageNumber="19">Mesalina</emphasis>
</taxonomicName>
and a member of the
<taxonomicName authorityName="Jiří Šmíd &amp; Jiří Moravec &amp; Václav Gvoždík &amp; Jan Štundl &amp; Daniel Frynta &amp; Petros Lymberakis &amp; Paschalia Kapli &amp; Thomas Wilms &amp; Andreas Schmitz &amp; Mohammed Shobrak &amp; Saeed Hosseinian Yousefkhani &amp; Eskandar Rastegar-Pouyani &amp; Aurora M. Castilla &amp; Johannes Els &amp; Werner Mayer" authorityYear="2017" box="[938,1133,306,334]" class="Reptilia" family="Lacertidae" genus="Mesalina" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="brevirostris">
<emphasis box="[938,1133,306,334]" italics="true" pageId="18" pageNumber="19">
<collectionCode box="[938,969,306,334]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung M<>nchen" pageId="18" pageNumber="19">M</collectionCode>
. brevirostris
</emphasis>
</taxonomicName>
species complex as revealed by the genetic analyses and characterized by the following combination of characters: (1) genetic (uncorrected) distance of 2.0% from
<taxonomicName authorityName="Jiří Šmíd &amp; Jiří Moravec &amp; Václav Gvoždík &amp; Jan Štundl &amp; Daniel Frynta &amp; Petros Lymberakis &amp; Paschalia Kapli &amp; Thomas Wilms &amp; Andreas Schmitz &amp; Mohammed Shobrak &amp; Saeed Hosseinian Yousefkhani &amp; Eskandar Rastegar-Pouyani &amp; Aurora M. Castilla &amp; Johannes Els &amp; Werner Mayer" authorityYear="2017" box="[992,1190,380,408]" class="Reptilia" family="Lacertidae" genus="Mesalina" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="brevirostris">
<emphasis box="[992,1190,380,408]" italics="true" pageId="18" pageNumber="19">
<collectionCode box="[992,1023,380,408]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung M<>nchen" pageId="18" pageNumber="19">M</collectionCode>
. brevirostris
</emphasis>
</taxonomicName>
, 2.7% from
<taxonomicName authorityName="Schenkel" authorityYear="1901" class="Reptilia" family="Lacertidae" genus="Eremias" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="bernoullii">
<emphasis italics="true" pageId="18" pageNumber="19">
<collectionCodeEnum box="[1365,1403,380,408]" pageId="18" pageNumber="19">
<collectionCode box="[1365,1403,380,408]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung M<>nchen" pageId="18" pageNumber="19">M</collectionCode>
</collectionCodeEnum>
. bernoullii
</emphasis>
</taxonomicName>
, and 2.8% from
<taxonomicName authorityName="Jiří Šmíd &amp; Jiří Moravec &amp; Václav Gvoždík &amp; Jan Štundl &amp; Daniel Frynta &amp; Petros Lymberakis &amp; Paschalia Kapli &amp; Thomas Wilms &amp; Andreas Schmitz &amp; Mohammed Shobrak &amp; Saeed Hosseinian Yousefkhani &amp; Eskandar Rastegar-Pouyani &amp; Aurora M. Castilla &amp; Johannes Els &amp; Werner Mayer" authorityYear="2017" baseAuthorityName="Angel" baseAuthorityYear="1936" box="[574,763,417,445]" class="Reptilia" family="Lacertidae" genus="Mesalina" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="microlepis">
<emphasis box="[574,763,417,445]" italics="true" pageId="18" pageNumber="19">
<collectionCodeEnum box="[574,605,417,445]" pageId="18" pageNumber="19">
<collectionCode box="[574,605,417,445]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung M<>nchen" pageId="18" pageNumber="19">M</collectionCode>
</collectionCodeEnum>
. microlepis
</emphasis>
</taxonomicName>
for the
<emphasis box="[897,945,417,445]" italics="true" pageId="18" pageNumber="19">
12
<collectionCodeEnum box="[928,945,417,445]" pageId="18" pageNumber="19">
<collectionCode box="[928,945,417,445]" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="18" pageNumber="19">S</collectionCode>
</collectionCodeEnum>
</emphasis>
(after Gblocks); 2.9% from
<taxonomicName authorityName="Jiří Šmíd &amp; Jiří Moravec &amp; Václav Gvoždík &amp; Jan Štundl &amp; Daniel Frynta &amp; Petros Lymberakis &amp; Paschalia Kapli &amp; Thomas Wilms &amp; Andreas Schmitz &amp; Mohammed Shobrak &amp; Saeed Hosseinian Yousefkhani &amp; Eskandar Rastegar-Pouyani &amp; Aurora M. Castilla &amp; Johannes Els &amp; Werner Mayer" authorityYear="2017" class="Reptilia" family="Lacertidae" genus="Mesalina" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="brevirostris">
<emphasis italics="true" pageId="18" pageNumber="19">
<collectionCode box="[1364,1402,417,445]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung M<>nchen" pageId="18" pageNumber="19">M</collectionCode>
. brevirostris
</emphasis>
</taxonomicName>
, 3.3% from
<taxonomicName authorityName="Schenkel" authorityYear="1901" box="[524,702,454,482]" class="Reptilia" family="Lacertidae" genus="Eremias" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="bernoullii">
<emphasis box="[524,702,454,482]" italics="true" pageId="18" pageNumber="19">
<collectionCode box="[524,555,454,482]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung M<>nchen" pageId="18" pageNumber="19">M</collectionCode>
. bernoullii
</emphasis>
</taxonomicName>
, and 4.0% from
<taxonomicName authorityName="Jiří Šmíd &amp; Jiří Moravec &amp; Václav Gvoždík &amp; Jan Štundl &amp; Daniel Frynta &amp; Petros Lymberakis &amp; Paschalia Kapli &amp; Thomas Wilms &amp; Andreas Schmitz &amp; Mohammed Shobrak &amp; Saeed Hosseinian Yousefkhani &amp; Eskandar Rastegar-Pouyani &amp; Aurora M. Castilla &amp; Johannes Els &amp; Werner Mayer" authorityYear="2017" baseAuthorityName="Angel" baseAuthorityYear="1936" box="[952,1138,454,482]" class="Reptilia" family="Lacertidae" genus="Mesalina" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="microlepis">
<emphasis box="[952,1138,454,482]" italics="true" pageId="18" pageNumber="19">M. microlepis</emphasis>
</taxonomicName>
for the
<emphasis box="[1265,1313,454,482]" italics="true" pageId="18" pageNumber="19">16S</emphasis>
(after Gblocks); 9.5% from
<taxonomicName authorityName="Jiří Šmíd &amp; Jiří Moravec &amp; Václav Gvoždík &amp; Jan Štundl &amp; Daniel Frynta &amp; Petros Lymberakis &amp; Paschalia Kapli &amp; Thomas Wilms &amp; Andreas Schmitz &amp; Mohammed Shobrak &amp; Saeed Hosseinian Yousefkhani &amp; Eskandar Rastegar-Pouyani &amp; Aurora M. Castilla &amp; Johannes Els &amp; Werner Mayer" authorityYear="2017" box="[468,659,490,518]" class="Reptilia" family="Lacertidae" genus="Mesalina" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="brevirostris">
<emphasis box="[468,659,490,518]" italics="true" pageId="18" pageNumber="19">M. brevirostris</emphasis>
</taxonomicName>
, 7.5% from
<taxonomicName authorityName="Schenkel" authorityYear="1901" box="[821,990,490,518]" class="Reptilia" family="Lacertidae" genus="Eremias" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="bernoullii">
<emphasis box="[821,990,490,518]" italics="true" pageId="18" pageNumber="19">M. bernoullii</emphasis>
</taxonomicName>
, and 10.8% from
<taxonomicName authorityName="Jiří Šmíd &amp; Jiří Moravec &amp; Václav Gvoždík &amp; Jan Štundl &amp; Daniel Frynta &amp; Petros Lymberakis &amp; Paschalia Kapli &amp; Thomas Wilms &amp; Andreas Schmitz &amp; Mohammed Shobrak &amp; Saeed Hosseinian Yousefkhani &amp; Eskandar Rastegar-Pouyani &amp; Aurora M. Castilla &amp; Johannes Els &amp; Werner Mayer" authorityYear="2017" baseAuthorityName="Angel" baseAuthorityYear="1936" box="[1221,1399,490,518]" class="Reptilia" family="Lacertidae" genus="Mesalina" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="microlepis">
<emphasis box="[1221,1399,490,518]" italics="true" pageId="18" pageNumber="19">M. microlepis</emphasis>
</taxonomicName>
for the
<emphasis box="[285,338,527,555]" italics="true" pageId="18" pageNumber="19">cytb</emphasis>
; (2) low number of dorsal scales (4142); (3) low number of collar plates (68); (4) low number of preanal scales (23); (5) low number of femoral pores in males (1213); (6) having 12 large semitransparent scales in the lower eyelid window; (7) in life, dorsum light cinnamon brown with a pattern of small whitish and larger dark cinnamon spots arranged in more or less regular longitudinal rows. Most of the whitish spots are not edged with dark brown color. The dark cinnamon brown spots predominate on flanks where they form a characteristic longitudinal lateral row that continues onto the tail. Ventral side bright white, sharply contrasting with the coloration of the dorsum.
</paragraph>
<paragraph blockId="18.[188,1408,858,1438]" pageId="18" pageNumber="19">
Comparisons.
<taxonomicName authority="Jiří Šmíd &amp; Jiří Moravec &amp; Václav Gvoždík &amp; Jan Štundl &amp; Daniel Frynta &amp; Petros Lymberakis &amp; Paschalia Kapli &amp; Thomas Wilms &amp; Andreas Schmitz &amp; Mohammed Shobrak &amp; Saeed Hosseinian Yousefkhani &amp; Eskandar Rastegar-Pouyani &amp; Aurora M. Castilla &amp; Johannes Els &amp; Werner Mayer, 2017" authorityName="Jiří Šmíd &amp; Jiří Moravec &amp; Václav Gvoždík &amp; Jan Štundl &amp; Daniel Frynta &amp; Petros Lymberakis &amp; Paschalia Kapli &amp; Thomas Wilms &amp; Andreas Schmitz &amp; Mohammed Shobrak &amp; Saeed Hosseinian Yousefkhani &amp; Eskandar Rastegar-Pouyani &amp; Aurora M. Castilla &amp; Johannes Els &amp; Werner Mayer" authorityYear="2017" box="[374,672,858,886]" class="Reptilia" family="Lacertidae" genus="Mesalina" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="saudiarabica" status="sp. n.">
<emphasis box="[374,672,858,886]" italics="true" pageId="18" pageNumber="19">
<taxonomicName box="[374,493,858,886]" class="Reptilia" family="Lacertidae" genus="Mesalina" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">Mesalina</taxonomicName>
saudiarabica
</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[681,751,858,886]" pageId="18" pageNumber="19" rank="species">sp. n.</taxonomicNameLabel>
can primarily be distinguished from other species of the complex by its genetic differentiation at both mtDNA and nDNA level. Genetic distances in the mtDNA genes are given above in the diagnosis and in
<tableCitation box="[1217,1325,932,960]" captionStart="Table S" captionStartId="23.[188,266,554,583]" captionText="Table S 1. Material used for the genetic analyses. Sample column refers to codes shown in Fig. S 1; Analyses indicates what phylogenetic analyses each sample was used for." pageId="18" pageNumber="19">Table 1</tableCitation>
. The differentiation in the nDNA is obvious from the allele networks (
<figureCitation box="[1059,1140,969,997]" captionStart="Figure S" captionStartId="22.[188,278,1539,1568]" captionTargetBox="[210,1439,213,1504]" captionTargetId="figure@22.[188,1453,188,1532]" captionTargetPageId="22" captionText="Figure S 2. Environmental space of the background used for developing the SDMs (grey box) and its parts occupied by Mesalina bernoullii, M. brevirostris, M. saudiarabica sp. n., M. microlepis. Values for the nine non­correlated environmental variables are shown (for abbreviations explanation see Material and methods). The species environmental space is based on their modelled distribution. Mean values (central line), standard deviation (box) and minimum and maximum values (whiskers) are shown." httpUri="https://zenodo.org/record/1037930/files/figure.png" pageId="18" pageNumber="19">Fig. 2</figureCitation>
) that show that the species does not share alleles of any gene with any other species.
<taxonomicName authority="Jiří Šmíd &amp; Jiří Moravec &amp; Václav Gvoždík &amp; Jan Štundl &amp; Daniel Frynta &amp; Petros Lymberakis &amp; Paschalia Kapli &amp; Thomas Wilms &amp; Andreas Schmitz &amp; Mohammed Shobrak &amp; Saeed Hosseinian Yousefkhani &amp; Eskandar Rastegar-Pouyani &amp; Aurora M. Castilla &amp; Johannes Els &amp; Werner Mayer, 2017" authorityName="Jiří Šmíd &amp; Jiří Moravec &amp; Václav Gvoždík &amp; Jan Štundl &amp; Daniel Frynta &amp; Petros Lymberakis &amp; Paschalia Kapli &amp; Thomas Wilms &amp; Andreas Schmitz &amp; Mohammed Shobrak &amp; Saeed Hosseinian Yousefkhani &amp; Eskandar Rastegar-Pouyani &amp; Aurora M. Castilla &amp; Johannes Els &amp; Werner Mayer" authorityYear="2017" box="[1054,1353,1006,1034]" class="Reptilia" family="Lacertidae" genus="Mesalina" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="saudiarabica" status="sp. n.">
<emphasis box="[1054,1353,1006,1034]" italics="true" pageId="18" pageNumber="19">
<taxonomicName box="[1054,1173,1006,1034]" class="Reptilia" family="Lacertidae" genus="Mesalina" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">Mesalina</taxonomicName>
saudiarabica
</emphasis>
</taxonomicName>
<taxonomicNameLabel pageId="18" pageNumber="19" rank="species">sp. n.</taxonomicNameLabel>
is also geographically isolated from the rest of the complex. The nearest localities of
<taxonomicName authorityName="Schenkel" authorityYear="1901" class="Reptilia" family="Lacertidae" genus="Eremias" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="bernoullii">
<emphasis italics="true" pageId="18" pageNumber="19">M. bernoullii</emphasis>
</taxonomicName>
lie
<quantity box="[377,482,1079,1107]" metricMagnitude="5" metricUnit="m" metricValue="6.8" pageId="18" pageNumber="19" unit="km" value="680.0">680 km</quantity>
to the east or 860 to the north. Moreover it can be distinguished morphologically from
<taxonomicName authorityName="Jiří Šmíd &amp; Jiří Moravec &amp; Václav Gvoždík &amp; Jan Štundl &amp; Daniel Frynta &amp; Petros Lymberakis &amp; Paschalia Kapli &amp; Thomas Wilms &amp; Andreas Schmitz &amp; Mohammed Shobrak &amp; Saeed Hosseinian Yousefkhani &amp; Eskandar Rastegar-Pouyani &amp; Aurora M. Castilla &amp; Johannes Els &amp; Werner Mayer" authorityYear="2017" baseAuthorityName="Angel" baseAuthorityYear="1936" box="[482,662,1116,1144]" class="Reptilia" family="Lacertidae" genus="Mesalina" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="microlepis">
<emphasis box="[482,662,1116,1144]" italics="true" pageId="18" pageNumber="19">M. microlepis</emphasis>
</taxonomicName>
by having 12 large semitransparent scales in the lower eyelid (several roughly equal semitransparent scales in the latter), lower number of collar plates (68 vs. 1013;
<emphasis box="[479,488,1190,1218]" italics="true" pageId="18" pageNumber="19">t</emphasis>
­test t = 5.01,
<emphasis box="[667,683,1190,1218]" italics="true" pageId="18" pageNumber="19">p</emphasis>
&lt;0.001), lower number of dorsal scales (4142 vs. 48 61;
<emphasis box="[239,248,1226,1254]" italics="true" pageId="18" pageNumber="19">t</emphasis>
­test t = 3.78,
<emphasis box="[435,451,1226,1254]" italics="true" pageId="18" pageNumber="19">p</emphasis>
&lt;0.005), lower number of preanal scales (23 vs. 4;
<emphasis box="[1159,1168,1226,1254]" italics="true" pageId="18" pageNumber="19">t</emphasis>
­test t = 8.14,
<emphasis box="[1355,1371,1226,1254]" italics="true" pageId="18" pageNumber="19">p</emphasis>
&lt;0.001), and lower number of femoral pores in males (1213 vs. 1520;
<emphasis box="[1151,1160,1263,1291]" italics="true" pageId="18" pageNumber="19">t</emphasis>
­test t = 5.12,
<emphasis box="[1353,1369,1263,1291]" italics="true" pageId="18" pageNumber="19">p</emphasis>
&lt;0.001). The lower number of collar plates differentiates the new species also from
<taxonomicName authorityName="Schenkel" authorityYear="1901" class="Reptilia" family="Lacertidae" genus="Eremias" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="bernoullii">
<emphasis italics="true" pageId="18" pageNumber="19">
<collectionCodeEnum box="[1364,1402,1300,1328]" pageId="18" pageNumber="19">M</collectionCodeEnum>
. bernoullii
</emphasis>
</taxonomicName>
(68 vs. 813;
<emphasis box="[532,541,1337,1365]" italics="true" pageId="18" pageNumber="19">t</emphasis>
­test t = 3.14,
<emphasis box="[737,753,1337,1365]" italics="true" pageId="18" pageNumber="19">p</emphasis>
&lt;0.005) and
<taxonomicName authority="Jiří Šmíd &amp; Jiří Moravec &amp; Václav Gvoždík &amp; Jan Štundl &amp; Daniel Frynta &amp; Petros Lymberakis &amp; Paschalia Kapli &amp; Thomas Wilms &amp; Andreas Schmitz &amp; Mohammed Shobrak &amp; Saeed Hosseinian Yousefkhani &amp; Eskandar Rastegar-Pouyani &amp; Aurora M. Castilla &amp; Johannes Els &amp; Werner Mayer, 2017" authorityName="Jiří Šmíd &amp; Jiří Moravec &amp; Václav Gvoždík &amp; Jan Štundl &amp; Daniel Frynta &amp; Petros Lymberakis &amp; Paschalia Kapli &amp; Thomas Wilms &amp; Andreas Schmitz &amp; Mohammed Shobrak &amp; Saeed Hosseinian Yousefkhani &amp; Eskandar Rastegar-Pouyani &amp; Aurora M. Castilla &amp; Johannes Els &amp; Werner Mayer" authorityYear="2017" box="[950,1155,1337,1365]" class="Reptilia" family="Lacertidae" genus="Mesalina" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="species" sensu="stricto" species="brevirostris" status="s. s.">
<emphasis box="[950,1155,1337,1365]" italics="true" pageId="18" pageNumber="19">
<collectionCodeEnum box="[950,981,1337,1365]" pageId="18" pageNumber="19">M</collectionCodeEnum>
. brevirostris
</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1160,1214,1337,1365]" pageId="18" pageNumber="19" sensu="stricto">s. s.</taxonomicNameLabel>
, although not significantly after Bonferroni correction (68 vs. 810;
<emphasis box="[915,924,1374,1402]" italics="true" pageId="18" pageNumber="19">t</emphasis>
­test t = ­2.67,
<emphasis box="[1120,1136,1374,1402]" italics="true" pageId="18" pageNumber="19">p</emphasis>
&lt;0.05) (
<tableCitation box="[1260,1401,1373,1401]" captionStart="Table S" captionStartId="27.[188,266,612,641]" captionText="Table S 4. Material examined for morphological comparisons and original measurements for each specimen. For details on the morphological character acronyms see Material and methods." pageId="18" pageNumber="19">
Tables
<collectionCodeEnum box="[1357,1379,1373,1401]" pageId="18" pageNumber="19">S</collectionCodeEnum>
4
</tableCitation>
,
<collectionCodeEnum box="[188,207,1410,1438]" pageId="18" pageNumber="19">S</collectionCodeEnum>
<tableCitation box="[207,224,1410,1438]" captionStart="Table S" captionStartId="29.[102,180,623,652]" captionTargetBox="[210,1856,745,1708]" captionTargetId="graphics@29.[201,1871,726,1707]" captionTargetPageId="29" captionText="Table S 5. Morphological comparison of Mesalina saudiarabica sp. n. with the other species formerly included under M. brevirostris. Values marked by asterisk indicate subadult specimens." httpUri="http://table.plazi.org/id/DF130BBDAE4FFFBB5418FDB4A2C46AF9" pageId="18" pageNumber="19" tableUuid="DF130BBDAE4FFFBB5418FDB4A2C46AF9">5</tableCitation>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19" type="description">
<paragraph blockId="18.[188,1408,1484,1990]" pageId="18" pageNumber="19">
Description of the
<typeStatus box="[436,552,1484,1512]" pageId="18" pageNumber="19">holotype</typeStatus>
. Subadult male (Fig. 4). Body slender, slightly depressed; snout short with prominent elevated nostrils, occipital shield absent, two large semitransparent scales in the lower eyelid; snout­vent length 31.0 mm, tail length 56.0 mm, head length 6.9/8.0 mm (to the anterior/posterior edge of the ear), head width
<quantity box="[1031,1132,1594,1622]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="18" pageNumber="19" unit="mm" value="5.1">5.1 mm</quantity>
, head depth
<quantity box="[1293,1403,1594,1622]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="18" pageNumber="19" unit="mm" value="3.5">3.5 mm</quantity>
. Upper labials (left/right) anterior the centre of eye 5/5 (smaller fifth upper labial separating the subocular from the mouth included), gulars 25, plates in collar 8, dorsals across midbody 41, ventrals across belly 12, transverse rows of ventrals 32, preanals in straight median series 3, subdigital lamellae 23/24, femoral pores 13/13. In alcohol, dorsum light brown with a pattern of small whitish and larger dark brown spots. Most of the whitish spots are not edged with dark brown color. The spots are arranged in more or less regular longitudinal rows. The dark brown spots predominate on flanks where they form a longitudinal lateral row that continues onto the tail. The lateral row of large brown spots is bordered by a dorsolateral row of small whitish spots and a narrow inconspicuous whitish ventrolateral line. Ventral side white.
</paragraph>
<paragraph blockId="19.[188,1408,196,334]" pageId="19" pageNumber="20">
Variation. The
<typeStatus box="[388,498,196,224]" pageId="19" pageNumber="20">paratype</typeStatus>
generally corresponds in morphology with the
<typeStatus box="[1123,1239,196,224]" pageId="19" pageNumber="20">holotype</typeStatus>
. Apart from several minute differences in scalation described in the paragraph Diagnosis it differs also in the presence of a distinct occipital shield and the presence of only one large semitransparent scale in the lower eyelid. The
<typeStatus box="[573,683,306,334]" pageId="19" pageNumber="20">paratype</typeStatus>
lacks the left hind leg and the tail.
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="20" type="distribution">
<paragraph blockId="19.[188,1408,380,923]" pageId="19" pageNumber="20">
Distribution and ecology. All eight so far known localities of
<taxonomicName authority="Jiří Šmíd &amp; Jiří Moravec &amp; Václav Gvoždík &amp; Jan Štundl &amp; Daniel Frynta &amp; Petros Lymberakis &amp; Paschalia Kapli &amp; Thomas Wilms &amp; Andreas Schmitz &amp; Mohammed Shobrak &amp; Saeed Hosseinian Yousefkhani &amp; Eskandar Rastegar-Pouyani &amp; Aurora M. Castilla &amp; Johannes Els &amp; Werner Mayer, 2017" authorityName="Jiří Šmíd &amp; Jiří Moravec &amp; Václav Gvoždík &amp; Jan Štundl &amp; Daniel Frynta &amp; Petros Lymberakis &amp; Paschalia Kapli &amp; Thomas Wilms &amp; Andreas Schmitz &amp; Mohammed Shobrak &amp; Saeed Hosseinian Yousefkhani &amp; Eskandar Rastegar-Pouyani &amp; Aurora M. Castilla &amp; Johannes Els &amp; Werner Mayer" authorityYear="2017" box="[988,1285,380,408]" class="Reptilia" family="Lacertidae" genus="Mesalina" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="saudiarabica" status="sp. n.">
<emphasis box="[988,1285,380,408]" italics="true" pageId="19" pageNumber="20">
<taxonomicName box="[988,1107,380,408]" class="Reptilia" family="Lacertidae" genus="Mesalina" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="20" phylum="Chordata" rank="genus">Mesalina</taxonomicName>
<taxonomicName box="[1116,1285,380,408]" pageId="19" pageNumber="20" rank="species" species="saudiarabica" status="sp. n.">saudiarabica</taxonomicName>
</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1294,1364,380,408]" pageId="19" pageNumber="20" rank="species">sp. n.</taxonomicNameLabel>
as well as its range of suitable conditions are located in central­western
<collectingCountry box="[1129,1302,417,445]" name="Saudi Arabia" pageId="19" pageNumber="20">Saudi Arabia</collectingCountry>
on the central plateau of the Arabian Peninsula at elevations of
<quantity box="[964,1131,453,481]" metricMagnitude="2" metricUnit="m" metricValue="9.75" metricValueMax="10.5" metricValueMin="9.0" pageId="19" pageNumber="20" unit="m" value="975.0" valueMax="1050.0" valueMin="900.0">9001050 m</quantity>
a.s.l. The region is characterized by hot and semi­arid to arid climate with mean summer temperatures up to 30°C and mean annual precipitation of
<quantity box="[638,796,527,555]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="19" pageNumber="20" unit="mm" value="75.0" valueMax="100.0" valueMin="50.0">50100 mm</quantity>
with rain typically occurring between March and May (Edgell 2006; Shobrak 2011). The terrain consists mostly of flat gravel plains known as regs, occasionally intersected by dry sandy wadis and dominated by sparse vegetation of perennial grasses including
<taxonomicName authorityName="C.G.D. Nees" authorityYear="1832" box="[558,711,638,666]" class="Liliopsida" family="Poaceae" genus="Stipagrostis" kingdom="Plantae" order="Poales" pageId="19" pageNumber="20" phylum="Tracheophyta" rank="genus">
<emphasis box="[558,711,638,666]" italics="true" pageId="19" pageNumber="20">Stipagrostis</emphasis>
</taxonomicName>
sp.,
<taxonomicName authorityName="Forssk" authorityYear="1775" box="[782,1025,638,666]" class="Liliopsida" family="Poaceae" genus="Panicum" kingdom="Plantae" order="Poales" pageId="19" pageNumber="20" phylum="Tracheophyta" rank="species" species="turgidum">
<emphasis box="[782,1025,638,666]" italics="true" pageId="19" pageNumber="20">
<taxonomicName box="[782,895,638,666]" class="Liliopsida" family="Poaceae" genus="Panicum" higherTaxonomySource="CoL" kingdom="Plantae" order="Poales" pageId="19" pageNumber="20" phylum="Tracheophyta" rank="genus">Panicum</taxonomicName>
<taxonomicName box="[908,1025,638,666]" pageId="19" pageNumber="20" rank="species" species="turgidum">turgidum</taxonomicName>
</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Henrard" authorityYear="1941" box="[1097,1341,638,666]" class="Mammalia" family="Vespertilionidae" genus="Lasiurus" kingdom="Animalia" order="Chiroptera" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="scindicus">
<emphasis box="[1097,1341,638,666]" italics="true" pageId="19" pageNumber="20">
<taxonomicName box="[1097,1209,638,666]" class="Mammalia" family="Vespertilionidae" genus="Lasiurus" higherTaxonomySource="CoL" kingdom="Animalia" order="Chiroptera" pageId="19" pageNumber="20" phylum="Chordata" rank="genus">Lasiurus</taxonomicName>
<taxonomicName box="[1222,1341,638,666]" pageId="19" pageNumber="20" rank="species" species="scindicus">scindicus</taxonomicName>
</emphasis>
</taxonomicName>
and small trees, mainly
<taxonomicName box="[438,527,674,702]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="19" pageNumber="20" phylum="Tracheophyta" rank="genus">
<emphasis box="[438,527,674,702]" italics="true" pageId="19" pageNumber="20">Acacia</emphasis>
</taxonomicName>
sp. (Mandaville 1990).
</paragraph>
<paragraph blockId="19.[188,1408,380,923]" pageId="19" pageNumber="20">
The
<typeStatus box="[345,400,711,739]" pageId="19" pageNumber="20">type</typeStatus>
locality is located in the Mahazat as­Sayd Nature Reserve, approximately
<quantity box="[188,287,748,776]" metricMagnitude="5" metricUnit="m" metricValue="1.7" pageId="19" pageNumber="20" unit="km" value="170.0">170 km</quantity>
E­NE of Taif. The reserve is Saudi Arabias only completely fenced wildlife reserve and is a reintroduction site for MacQueens bustard (
<taxonomicName baseAuthorityName="J. E. Gray" baseAuthorityYear="1832" box="[880,1199,785,813]" class="Aves" family="Otididae" genus="Chlamydotis" kingdom="Animalia" order="Gruiformes" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="macqueenii">
<emphasis box="[880,1199,785,813]" italics="true" pageId="19" pageNumber="20">
<taxonomicName box="[880,1042,785,813]" class="Aves" family="Otididae" genus="Chlamydotis" higherTaxonomySource="CoL" kingdom="Animalia" order="Otidiformes" pageId="19" pageNumber="20" phylum="Chordata" rank="genus">Chlamydotis</taxonomicName>
<taxonomicName box="[1052,1199,785,813]" pageId="19" pageNumber="20" rank="species" species="macqueenii">macqueenii</taxonomicName>
</emphasis>
</taxonomicName>
), Arabian oryx (
<emphasis box="[199,384,822,850]" italics="true" pageId="19" pageNumber="20">Oryx leucoryx</emphasis>
) and Sand gazelles (
<taxonomicName baseAuthorityName="Güldenstaedt" baseAuthorityYear="1780" box="[665,944,822,850]" class="Mammalia" family="Bovidae" genus="Gazella" kingdom="Animalia" order="Artiodactyla" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="subgutturosa">
<emphasis box="[665,944,822,850]" italics="true" pageId="19" pageNumber="20">
<taxonomicName box="[665,765,822,850]" class="Mammalia" family="Bovidae" genus="Gazella" higherTaxonomySource="CoL" kingdom="Animalia" order="Artiodactyla" pageId="19" pageNumber="20" phylum="Chordata" rank="genus">Gazella</taxonomicName>
<taxonomicName box="[777,944,822,850]" pageId="19" pageNumber="20" rank="species" species="subgutturosa">subgutturosa</taxonomicName>
</emphasis>
</taxonomicName>
). At the reserve,
<taxonomicName box="[1180,1367,822,850]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="19" pageNumber="20" phylum="Tracheophyta" rank="species" species="tortilis">
<emphasis box="[1180,1367,822,850]" italics="true" pageId="19" pageNumber="20">
<taxonomicName box="[1180,1269,822,850]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" higherTaxonomySource="CoL" kingdom="Plantae" order="Fabales" pageId="19" pageNumber="20" phylum="Tracheophyta" rank="genus">Acacia</taxonomicName>
<taxonomicName box="[1281,1367,822,850]" pageId="19" pageNumber="20" rank="species" species="tortilis">tortilis</taxonomicName>
</emphasis>
</taxonomicName>
is the most common tree species,
<taxonomicName authorityName="Burm fil" authorityYear="1768" box="[599,798,858,886]" class="Magnoliopsida" family="Zygophyllaceae" genus="Fagonia" kingdom="Plantae" order="Zygophyllales" pageId="19" pageNumber="20" phylum="Tracheophyta" rank="species" species="indica">
<emphasis box="[599,798,858,886]" italics="true" pageId="19" pageNumber="20">
<taxonomicName box="[599,707,858,886]" class="Magnoliopsida" family="Zygophyllaceae" genus="Fagonia" higherTaxonomySource="CoL" kingdom="Plantae" order="Zygophyllales" pageId="19" pageNumber="20" phylum="Tracheophyta" rank="genus">Fagonia</taxonomicName>
<taxonomicName box="[718,798,858,886]" pageId="19" pageNumber="20" rank="species" species="indica">indica</taxonomicName>
</emphasis>
</taxonomicName>
and
<taxonomicName box="[864,1107,858,886]" class="Magnoliopsida" family="Fabaceae" genus="Indigofera" kingdom="Plantae" order="Fabales" pageId="19" pageNumber="20" phylum="Tracheophyta" rank="species" species="spinosa">
<emphasis box="[864,1107,858,886]" italics="true" pageId="19" pageNumber="20">
<taxonomicName box="[864,999,858,886]" class="Magnoliopsida" family="Fabaceae" genus="Indigofera" higherTaxonomySource="CoL" kingdom="Plantae" order="Fabales" pageId="19" pageNumber="20" phylum="Tracheophyta" rank="genus">Indigofera</taxonomicName>
<taxonomicName box="[1009,1107,858,886]" pageId="19" pageNumber="20" rank="species" species="spinosa">spinosa</taxonomicName>
</emphasis>
</taxonomicName>
are the most common herbs, and
<taxonomicName authorityName="Forssk" authorityYear="1775" box="[327,566,895,923]" class="Liliopsida" family="Poaceae" genus="Panicum" kingdom="Plantae" order="Poales" pageId="19" pageNumber="20" phylum="Tracheophyta" rank="species" species="turgidum">
<emphasis box="[327,566,895,923]" italics="true" pageId="19" pageNumber="20">
<taxonomicName box="[327,441,895,923]" class="Liliopsida" family="Poaceae" genus="Panicum" higherTaxonomySource="CoL" kingdom="Plantae" order="Poales" pageId="19" pageNumber="20" phylum="Tracheophyta" rank="genus">Panicum</taxonomicName>
<taxonomicName box="[449,566,895,923]" pageId="19" pageNumber="20" rank="species" species="turgidum">turgidum</taxonomicName>
</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="C.G.D. Nees" authorityYear="1832" box="[629,790,895,923]" class="Liliopsida" family="Poaceae" genus="Stipagrostis" kingdom="Plantae" order="Poales" pageId="19" pageNumber="20" phylum="Tracheophyta" rank="genus">
<emphasis box="[629,790,895,923]" italics="true" pageId="19" pageNumber="20">Stipagrostis</emphasis>
</taxonomicName>
spp. are the prevailing grasses.
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="20" type="etymology">
<paragraph blockId="19.[188,1407,969,1033]" pageId="19" pageNumber="20">Etymology. The specific epithet is derived from the name of the country where the species occurs. Proposed English name Arabian short­nosed desert lizard.</paragraph>
</subSubSection>
</treatment>
</document>