235 lines
22 KiB
XML
235 lines
22 KiB
XML
<document ID-DOI="10.1080/00222933.2018.1524032" ID-GBIF-Dataset="034ebcd9-efa7-416a-822c-b6aee0e57a1c" ID-ISSN="1464-5262" ID-Zenodo-Dep="3670229" checkinTime="1581941790231" checkinUser="valdenar" docAuthor="DeVries, Thomas J." docDate="2019" docId="03BB2338E368E21AFE70A5825989FC06" docLanguage="en" docName="JNATHIST.53.25–26.1533–1584.pdf.imf" docOrigin="Journal of Natural History 53 (25)" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Nodifaunus Olssoni 1944" docType="treatment" docVersion="20" lastPageNumber="1565" masterDocId="FF825B40E348E23BFFA1A1435A0CFFCC" masterDocTitle="Early Paleogene brackish-water molluscs from the Caballas Formation of the East Pisco Basin (Southern Peru)" masterLastPageNumber="1584" masterPageNumber="1533" pageNumber="1564" updateTime="1636522412784" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Early Paleogene brackish-water molluscs from the Caballas Formation of the East Pisco Basin (Southern Peru)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>DeVries, Thomas J.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Journal of Natural History</mods:title>
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</mods:titleInfo>
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||
<mods:part>
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<mods:date>2019</mods:date>
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||
<mods:detail type="pubDate">
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<mods:number>2018-12-17</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>53</mods:number>
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</mods:detail>
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||
<mods:detail type="issue">
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<mods:number>25</mods:number>
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||
</mods:detail>
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||
<mods:extent unit="page">
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<mods:start>1533</mods:start>
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||
<mods:end>1584</mods:end>
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||
</mods:extent>
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||
</mods:part>
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||
</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1080/00222933.2018.1524032</mods:identifier>
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<mods:identifier type="GBIF-Dataset">034ebcd9-efa7-416a-822c-b6aee0e57a1c</mods:identifier>
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<mods:identifier type="ISSN">1464-5262</mods:identifier>
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<mods:identifier type="Zenodo-Dep">3670229</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3671260" ID-GBIF-Taxon="162146487" ID-Zenodo-Dep="3671260" LSID="urn:lsid:plazi:treatment:03BB2338E368E21AFE70A5825989FC06" httpUri="http://treatment.plazi.org/id/03BB2338E368E21AFE70A5825989FC06" lastPageId="33" lastPageNumber="1565" pageId="32" pageNumber="1564">
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<subSubSection box="[465,850,1217,1242]" pageId="32" pageNumber="1564" type="nomenclature">
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<paragraph blockId="32.[465,850,1217,1242]" box="[465,850,1217,1242]" pageId="32" pageNumber="1564">
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<heading box="[465,850,1217,1242]" centered="true" fontSize="11" level="2" pageId="32" pageNumber="1564" reason="2">
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Genus
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<taxonomicName authority="Olssoni 1944" authorityName="Olssoni" authorityYear="1944" box="[548,850,1217,1242]" class="Gastropoda" family="Pachychilidae" genus="Nodifaunus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="32" pageNumber="1564" phylum="Mollusca" rank="genus">
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<emphasis bold="true" box="[548,689,1217,1242]" italics="true" pageId="32" pageNumber="1564">Nodifaunus</emphasis>
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Olsson, 1944
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="32" pageNumber="1564" type="materials_examined">
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<paragraph blockId="32.[160,1155,1284,1343]" pageId="32" pageNumber="1564">
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<typeStatus box="[160,214,1284,1308]" pageId="32" pageNumber="1564">Type</typeStatus>
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species:
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<taxonomicName authorityName="Olsson" authorityYear="1944" box="[316,538,1284,1308]" class="Gastropoda" family="Pachychilidae" genus="Nodifaunus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="32" pageNumber="1564" phylum="Mollusca" rank="species" species="nodosus">
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<emphasis box="[316,538,1284,1308]" italics="true" pageId="32" pageNumber="1564">Nodifaunus nodosus</emphasis>
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</taxonomicName>
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,
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<date box="[552,612,1284,1308]" pageId="32" pageNumber="1564" value="1944-01">1944,</date>
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by original designation. Late Cretaceous, northern
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<collectingCountry box="[206,260,1319,1343]" name="Peru" pageId="32" pageNumber="1564">Peru</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="33" lastPageNumber="1565" pageId="32" pageNumber="1564" type="discussion">
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<paragraph blockId="32.[160,1156,1389,1727]" box="[160,262,1389,1415]" pageId="32" pageNumber="1564">
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<emphasis bold="true" box="[160,262,1389,1415]" italics="true" pageId="32" pageNumber="1564">Remarks</emphasis>
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</paragraph>
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<paragraph blockId="32.[160,1156,1389,1727]" pageId="32" pageNumber="1564">
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<taxonomicName authorityName="Olssoni" authorityYear="1944" box="[160,285,1426,1450]" class="Gastropoda" family="Pachychilidae" genus="Nodifaunus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="32" pageNumber="1564" phylum="Mollusca" rank="genus">
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<emphasis box="[160,285,1426,1450]" italics="true" pageId="32" pageNumber="1564">Nodifaunus</emphasis>
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</taxonomicName>
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was created by
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<bibRefCitation author="Olsson AA" box="[474,629,1426,1450]" pageId="32" pageNumber="1564" pagination="1 - 147" refId="ref25769" refString="Olsson AA. 1944. Contributions to the Tertiary paleontology of northern Peru. Part VII. The Cretaceous of the Paita region. Bulletins Am Paleontol. 19 (68): 1 - 147." type="journal article" year="1944">Olsson (1944)</bibRefCitation>
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to include high-spired cerithiform gastropods with smooth early teleoconch whorls and tuberculate or coarsely ribbed later whorls, an aperture having a broadly callused inner lip, an outer lip extended posteriorly at the suture, a short anterior canal, and a smooth base.
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<bibRefCitation author="Olsson AA" box="[749,905,1530,1554]" pageId="32" pageNumber="1564" pagination="1 - 147" refId="ref25769" refString="Olsson AA. 1944. Contributions to the Tertiary paleontology of northern Peru. Part VII. The Cretaceous of the Paita region. Bulletins Am Paleontol. 19 (68): 1 - 147." type="journal article" year="1944">Olsson (1944)</bibRefCitation>
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noted morphological gradations between
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<taxonomicName authorityName="Olsson" authorityYear="1944" box="[404,529,1565,1589]" class="Gastropoda" family="Pachychilidae" genus="Nodifaunus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="32" pageNumber="1564" phylum="Mollusca" rank="species" species="nodosus">
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<emphasis box="[404,529,1565,1589]" italics="true" pageId="32" pageNumber="1564">N. nodosus</emphasis>
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</taxonomicName>
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and
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<taxonomicName authority="Olssoni 1944" authorityName="Olssoni" authorityYear="1944" box="[598,885,1565,1589]" class="Gastropoda" family="Pachychilidae" genus="Nodifaunus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="32" pageNumber="1564" phylum="Mollusca" rank="species" species="costatus">
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<emphasis box="[598,723,1565,1589]" italics="true" pageId="32" pageNumber="1564">N. costatus</emphasis>
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Olsson, 1944
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</taxonomicName>
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; both from the Upper Cretaceous
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<collectingCountry box="[298,386,1599,1623]" name="Haiti" pageId="32" pageNumber="1564">Tortuga</collectingCountry>
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Formation of northern
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<collectingCountry box="[679,733,1599,1623]" name="Peru" pageId="32" pageNumber="1564">Peru</collectingCountry>
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. According to
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<bibRefCitation author="Olsson AA" box="[916,1076,1599,1623]" pageId="32" pageNumber="1564" pagination="1 - 147" refId="ref25769" refString="Olsson AA. 1944. Contributions to the Tertiary paleontology of northern Peru. Part VII. The Cretaceous of the Paita region. Bulletins Am Paleontol. 19 (68): 1 - 147." type="journal article" year="1944">Olsson (1944)</bibRefCitation>
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, some specimens of
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<taxonomicName authorityName="Olsson" authorityYear="1944" box="[324,448,1634,1658]" class="Gastropoda" family="Pachychilidae" genus="Nodifaunus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="32" pageNumber="1564" phylum="Mollusca" rank="species" species="nodosus">
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<emphasis box="[324,448,1634,1658]" italics="true" pageId="32" pageNumber="1564">N. nodosus</emphasis>
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</taxonomicName>
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have weak spiral cords on the base, while others have the smooth base that characterises
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<emphasis box="[537,804,1669,1693]" italics="true" pageId="32" pageNumber="1564">
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<taxonomicName authorityName="Olssoni" authorityYear="1944" box="[537,663,1669,1693]" class="Gastropoda" family="Pachychilidae" genus="Nodifaunus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="32" pageNumber="1564" phylum="Mollusca" rank="species" species="costatus">N. costatus</taxonomicName>
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.
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<taxonomicName authorityName="Olssoni" authorityYear="1944" box="[679,804,1669,1693]" class="Gastropoda" family="Pachychilidae" genus="Nodifaunus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="32" pageNumber="1564" phylum="Mollusca" rank="species" species="costatus">N. costatus</taxonomicName>
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</emphasis>
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is herein considered a junior synonym of
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<taxonomicName authorityName="Olsson" authorityYear="1944" box="[302,423,1703,1727]" class="Gastropoda" family="Pachychilidae" genus="Nodifaunus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="32" pageNumber="1564" phylum="Mollusca" rank="species" species="nodosus">
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<emphasis box="[302,423,1703,1727]" italics="true" pageId="32" pageNumber="1564">N. nodosus</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<paragraph blockId="33.[160,1156,142,755]" pageId="33" pageNumber="1565">
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<taxonomicName authority="Olssoni 1944" authorityName="Olssoni" authorityYear="1944" box="[192,556,142,166]" class="Gastropoda" family="Thiaridae" genus="Melanatria" kingdom="Animalia" pageId="33" pageNumber="1565" phylum="Mollusca" rank="species" species="medialis">
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<emphasis box="[192,407,142,166]" italics="true" pageId="33" pageNumber="1565">Melanatria medialis</emphasis>
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Olsson, 1944
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</taxonomicName>
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, also from the
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<collectingCountry box="[723,811,142,166]" name="Haiti" pageId="33" pageNumber="1565">Tortuga</collectingCountry>
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Formation of northern Peru, is figured by
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<bibRefCitation author="Olsson AA" box="[284,429,177,201]" pageId="33" pageNumber="1565" pagination="1 - 147" refId="ref25769" refString="Olsson AA. 1944. Contributions to the Tertiary paleontology of northern Peru. Part VII. The Cretaceous of the Paita region. Bulletins Am Paleontol. 19 (68): 1 - 147." type="journal article" year="1944">
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Olsson (
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<date box="[374,429,177,201]" pageId="33" pageNumber="1565" value="1944-01">1944</date>
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</bibRefCitation>
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, pl. 12, figs. 8, 11) alongside
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<taxonomicName authorityName="Olsson" authorityYear="1944" box="[755,977,177,201]" class="Gastropoda" family="Pachychilidae" genus="Nodifaunus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="33" pageNumber="1565" phylum="Mollusca" rank="species" species="nodosus">
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<emphasis box="[755,977,177,201]" italics="true" pageId="33" pageNumber="1565">Nodifaunus nodosus</emphasis>
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</taxonomicName>
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and
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<taxonomicName authorityName="Olssoni" authorityYear="1944" box="[1035,1155,177,201]" class="Gastropoda" family="Pachychilidae" genus="Nodifaunus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="33" pageNumber="1565" phylum="Mollusca" rank="species" species="costatus">
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<emphasis box="[1035,1155,177,201]" italics="true" pageId="33" pageNumber="1565">N. costatus</emphasis>
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</taxonomicName>
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(
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<bibRefCitation author="Olsson AA" box="[166,307,212,236]" pageId="33" pageNumber="1565" pagination="1 - 147" refId="ref25769" refString="Olsson AA. 1944. Contributions to the Tertiary paleontology of northern Peru. Part VII. The Cretaceous of the Paita region. Bulletins Am Paleontol. 19 (68): 1 - 147." type="journal article" year="1944">
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Olsson
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<date box="[252,307,212,236]" pageId="33" pageNumber="1565" value="1944-01">1944</date>
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</bibRefCitation>
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, pl. 12). Both in illustration and description, the specimens of
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<taxonomicName authorityName="Olssoni" authorityYear="1944" box="[1029,1155,212,236]" class="Gastropoda" family="Thiaridae" genus="Melanatria" kingdom="Animalia" pageId="33" pageNumber="1565" phylum="Mollusca" rank="species" species="medialis">
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<emphasis box="[1029,1155,212,236]" italics="true" pageId="33" pageNumber="1565">M. medialis</emphasis>
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</taxonomicName>
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differ little from specimens of
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<taxonomicName authorityName="Olssoni" authorityYear="1944" box="[511,636,246,270]" class="Gastropoda" family="Pachychilidae" genus="Nodifaunus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="33" pageNumber="1565" phylum="Mollusca" rank="genus">
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<emphasis box="[511,636,246,270]" italics="true" pageId="33" pageNumber="1565">Nodifaunus</emphasis>
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</taxonomicName>
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, except that the
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<taxonomicName authorityName="Olssoni" authorityYear="1944" box="[838,964,246,270]" class="Gastropoda" family="Thiaridae" genus="Melanatria" kingdom="Animalia" pageId="33" pageNumber="1565" phylum="Mollusca" rank="species" species="medialis">
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<emphasis box="[838,964,246,270]" italics="true" pageId="33" pageNumber="1565">M. medialis</emphasis>
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</taxonomicName>
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specimens have two spiral rows of tubercles.
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<bibRefCitation author="Olsson AA" box="[516,675,281,305]" pageId="33" pageNumber="1565" pagination="1 - 147" refId="ref25769" refString="Olsson AA. 1944. Contributions to the Tertiary paleontology of northern Peru. Part VII. The Cretaceous of the Paita region. Bulletins Am Paleontol. 19 (68): 1 - 147." type="journal article" year="1944">Olsson (1944)</bibRefCitation>
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also noted that some specimens of
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<taxonomicName authorityName="Olssoni" authorityYear="1944" class="Gastropoda" family="Thiaridae" genus="Melanatria" kingdom="Animalia" pageId="33" pageNumber="1565" phylum="Mollusca" rank="species" species="medialis">
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<emphasis italics="true" pageId="33" pageNumber="1565">M. medialis</emphasis>
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</taxonomicName>
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have fine spiral lines on the early teleoconch whorls.
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<taxonomicName authorityName="Olssoni" authorityYear="1944" box="[906,1126,316,340]" class="Gastropoda" family="Thiaridae" genus="Melanatria" kingdom="Animalia" pageId="33" pageNumber="1565" phylum="Mollusca" rank="species" species="medialis">
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<emphasis box="[906,1126,316,340]" italics="true" pageId="33" pageNumber="1565">Melanatria medialis</emphasis>
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</taxonomicName>
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is herein transferred to
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<taxonomicName authorityName="Olssoni" authorityYear="1944" box="[404,529,350,374]" class="Gastropoda" family="Pachychilidae" genus="Nodifaunus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="33" pageNumber="1565" phylum="Mollusca" rank="genus">
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<emphasis box="[404,529,350,374]" italics="true" pageId="33" pageNumber="1565">Nodifaunus</emphasis>
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</taxonomicName>
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, with the diagnosis of
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<taxonomicName authorityName="Olssoni" authorityYear="1944" box="[792,917,350,374]" class="Gastropoda" family="Pachychilidae" genus="Nodifaunus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="33" pageNumber="1565" phylum="Mollusca" rank="genus">
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<emphasis box="[792,917,350,374]" italics="true" pageId="33" pageNumber="1565">Nodifaunus</emphasis>
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</taxonomicName>
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amended to include species that have spiral cords or threads on early teleoconch whorls or on the base, but that nonetheless are largely smooth on later teleoconch whorls.
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</paragraph>
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<paragraph blockId="33.[160,1156,142,755]" pageId="33" pageNumber="1565">
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<bibRefCitation author="Olsson AA" box="[192,345,454,478]" pageId="33" pageNumber="1565" pagination="1 - 147" refId="ref25769" refString="Olsson AA. 1944. Contributions to the Tertiary paleontology of northern Peru. Part VII. The Cretaceous of the Paita region. Bulletins Am Paleontol. 19 (68): 1 - 147." type="journal article" year="1944">Olsson (1944)</bibRefCitation>
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||
compared
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<taxonomicName authorityName="Olssoni" authorityYear="1944" box="[473,689,454,478]" class="Gastropoda" family="Thiaridae" genus="Melanatria" kingdom="Animalia" pageId="33" pageNumber="1565" phylum="Mollusca" rank="species" species="medialis">
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<emphasis box="[473,689,454,478]" italics="true" pageId="33" pageNumber="1565">Melanatria medialis</emphasis>
|
||
</taxonomicName>
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||
with
|
||
<taxonomicName authority="Woodsi 1922" authorityName="Woodsi" authorityYear="1922" box="[753,1063,454,478]" class="Gastropoda" family="Thiaridae" genus="Melanatria" kingdom="Animalia" pageId="33" pageNumber="1565" phylum="Mollusca" rank="species" species="dimorphica">
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||
<emphasis box="[753,911,454,478]" italics="true" pageId="33" pageNumber="1565">M. dimorphica</emphasis>
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Woods, 1922
|
||
</taxonomicName>
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||
, a highspired species with two spiral rows of tubercles found in lower Eocene strata near Negritos, northern
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<collectingCountry box="[382,436,524,548]" name="Peru" pageId="33" pageNumber="1565">Peru</collectingCountry>
|
||
.
|
||
<taxonomicName authorityName="Woodsi" authorityYear="1922" box="[450,703,524,548]" class="Gastropoda" family="Thiaridae" genus="Melanatria" kingdom="Animalia" pageId="33" pageNumber="1565" phylum="Mollusca" rank="species" species="dimorphica">
|
||
<emphasis box="[450,703,524,548]" italics="true" pageId="33" pageNumber="1565">Melanatria dimorphica</emphasis>
|
||
</taxonomicName>
|
||
has fine spiral threads on early teleoconch whorls and spiral cords on the base (
|
||
<bibRefCitation author="Woods H" box="[657,800,558,582]" pageId="33" pageNumber="1565" pagination="51 - 113" refId="ref27434" refString="Woods H. 1922. Mollusca from Eocene and Miocene deposits of Peru. In: Bosworth TOi editor. Geology of the Tertiary and Quaternary periods in the northern part of Peru. London: Macmillan; p. 51 - 113." type="book chapter" year="1922">
|
||
Woods
|
||
<date box="[745,800,558,582]" pageId="33" pageNumber="1565" value="1922-01">1922</date>
|
||
</bibRefCitation>
|
||
, pl. 9, fig. 11a, 11b).
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||
<taxonomicName authorityName="Woodsi" authorityYear="1922" class="Gastropoda" family="Thiaridae" genus="Melanatria" kingdom="Animalia" pageId="33" pageNumber="1565" phylum="Mollusca" rank="species" species="dimorphica">
|
||
<emphasis italics="true" pageId="33" pageNumber="1565">Melanatria dimorphica</emphasis>
|
||
</taxonomicName>
|
||
is herein transferred to
|
||
<taxonomicName authorityName="Olssoni" authorityYear="1944" box="[556,681,593,617]" class="Gastropoda" family="Pachychilidae" genus="Nodifaunus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="33" pageNumber="1565" phylum="Mollusca" rank="genus">
|
||
<emphasis box="[556,681,593,617]" italics="true" pageId="33" pageNumber="1565">Nodifaunus</emphasis>
|
||
</taxonomicName>
|
||
, as are four other early Eocene cerithioids from northern
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||
<collectingCountry box="[326,380,627,651]" name="Peru" pageId="33" pageNumber="1565">Peru</collectingCountry>
|
||
:
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||
<taxonomicName authority="Woodsi 1922" authorityName="Woodsi" authorityYear="1922" box="[390,774,627,651]" class="Gastropoda" family="Thiaridae" genus="Melanatria" kingdom="Animalia" pageId="33" pageNumber="1565" phylum="Mollusca" rank="species" species="acanthica">
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||
<emphasis box="[390,622,627,651]" italics="true" pageId="33" pageNumber="1565">Melanatria acanthica</emphasis>
|
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Woods, 1922
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||
</taxonomicName>
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||
, a large form of
|
||
<emphasis box="[959,1122,627,651]" italics="true" pageId="33" pageNumber="1565">
|
||
<taxonomicName authorityName="Woodsi" authorityYear="1922" box="[959,1116,627,651]" class="Gastropoda" family="Thiaridae" genus="Melanatria" kingdom="Animalia" pageId="33" pageNumber="1565" phylum="Mollusca" rank="species" species="dimorphica">M. dimorphica</taxonomicName>
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||
;
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</emphasis>
|
||
<taxonomicName authority="Woodsi 1922" authorityName="Woodsi" authorityYear="1922" class="Gastropoda" family="Thiaridae" genus="Melanatria" kingdom="Animalia" pageId="33" pageNumber="1565" phylum="Mollusca" rank="species" species="propinqua">
|
||
<emphasis italics="true" pageId="33" pageNumber="1565">M. propinqua</emphasis>
|
||
Woods, 1922
|
||
</taxonomicName>
|
||
, a species with a single row of medially situated tubercles and thought by
|
||
<bibRefCitation author="Woods H" box="[294,450,697,721]" pageId="33" pageNumber="1565" pagination="51 - 113" refId="ref27434" refString="Woods H. 1922. Mollusca from Eocene and Miocene deposits of Peru. In: Bosworth TOi editor. Geology of the Tertiary and Quaternary periods in the northern part of Peru. London: Macmillan; p. 51 - 113." type="book chapter" year="1922">Woods (1922)</bibRefCitation>
|
||
to be a variety of
|
||
<taxonomicName authorityName="Woodsi" authorityYear="1922" box="[657,815,697,721]" class="Gastropoda" family="Thiaridae" genus="Melanatria" kingdom="Animalia" pageId="33" pageNumber="1565" phylum="Mollusca" rank="species" species="dimorphica">
|
||
<emphasis box="[657,815,697,721]" italics="true" pageId="33" pageNumber="1565">M. dimorphica</emphasis>
|
||
</taxonomicName>
|
||
; and
|
||
<taxonomicName authority="Woodsi 1922" authorityName="Woodsi" authorityYear="1922" box="[878,1150,697,721]" class="Gastropoda" family="Thiaridae" genus="Melanatria" kingdom="Animalia" pageId="33" pageNumber="1565" phylum="Mollusca" rank="species" species="venusta">
|
||
<emphasis box="[878,997,697,721]" italics="true" pageId="33" pageNumber="1565">M. venusta</emphasis>
|
||
Woods, 1922
|
||
</taxonomicName>
|
||
, a species with tuberculate axial ribs resembling those of
|
||
<taxonomicName authorityName="Olssoni" authorityYear="1944" box="[813,1036,731,755]" class="Gastropoda" family="Pachychilidae" genus="Nodifaunus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="33" pageNumber="1565" phylum="Mollusca" rank="species" species="costatus">
|
||
<emphasis box="[813,1036,731,755]" italics="true" pageId="33" pageNumber="1565">Nodifaunus costatus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="33.[160,1156,805,970]" box="[160,391,805,831]" pageId="33" pageNumber="1565">
|
||
<emphasis bold="true" box="[160,391,805,831]" italics="true" pageId="33" pageNumber="1565">Emended diagnosis</emphasis>
|
||
</paragraph>
|
||
<paragraph blockId="33.[160,1156,805,970]" pageId="33" pageNumber="1565">Whorls flat sided, sutures adpressed or canaliculate. Early whorls smooth or with spiral threads, later whorls smooth, with tubercles or tuberculate axial ribs. Aperture with broad parietal and columellar thickening; anterior canal short, wide; outer lip flaring adjacent to suture of penultimate whorl; posterior canal shallow.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |