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<document ID-DOI="10.1080/00222933.2018.1524032" ID-GBIF-Dataset="034ebcd9-efa7-416a-822c-b6aee0e57a1c" ID-ISSN="1464-5262" ID-Zenodo-Dep="3670229" checkinTime="1581941790231" checkinUser="valdenar" docAuthor="DeVries, Thomas J." docDate="2019" docId="03BB2338E351E221FE52A0445E88FED8" docLanguage="en" docName="JNATHIST.53.25–26.1533–1584.pdf.imf" docOrigin="Journal of Natural History 53 (25)" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Potamides henryi DeVries 2019, sp. nov." docType="treatment" docVersion="20" lastPageNumber="1558" masterDocId="FF825B40E348E23BFFA1A1435A0CFFCC" masterDocTitle="Early Paleogene brackish-water molluscs from the Caballas Formation of the East Pisco Basin (Southern Peru)" masterLastPageNumber="1584" masterPageNumber="1533" pageNumber="1557" updateTime="1636522412784" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Early Paleogene brackish-water molluscs from the Caballas Formation of the East Pisco Basin (Southern Peru)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>DeVries, Thomas J.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="pubDate">
<mods:number>2018-12-17</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>53</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>25</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1533</mods:start>
<mods:end>1584</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1080/00222933.2018.1524032</mods:identifier>
<mods:identifier type="GBIF-Dataset">034ebcd9-efa7-416a-822c-b6aee0e57a1c</mods:identifier>
<mods:identifier type="ISSN">1464-5262</mods:identifier>
<mods:identifier type="Zenodo-Dep">3670229</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3671198" ID-GBIF-Taxon="162146512" ID-Zenodo-Dep="3671198" LSID="urn:lsid:plazi:treatment:03BB2338E351E221FE52A0445E88FED8" httpUri="http://treatment.plazi.org/id/03BB2338E351E221FE52A0445E88FED8" lastPageId="26" lastPageNumber="1558" pageId="25" pageNumber="1557">
<subSubSection box="[499,817,263,288]" pageId="25" pageNumber="1557" type="nomenclature">
<paragraph blockId="25.[499,817,263,322]" box="[499,817,263,288]" pageId="25" pageNumber="1557">
<heading box="[499,817,263,288]" centered="true" fontSize="11" level="2" pageId="25" pageNumber="1557" reason="2">
<taxonomicName authority="DeVries, 2019" authorityName="DeVries" authorityYear="2019" box="[499,714,263,288]" class="Gastropoda" family="Potamididae" genus="Potamides" kingdom="Animalia" pageId="25" pageNumber="1557" phylum="Mollusca" rank="species" species="henryi" status="sp. nov.">
<emphasis bold="true" box="[499,714,263,288]" italics="true" pageId="25" pageNumber="1557">Potamides henryi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[724,817,263,288]" pageId="25" pageNumber="1557" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[566,749,297,322]" pageId="25" pageNumber="1557" type="description">
<paragraph blockId="25.[499,817,263,322]" box="[566,749,297,322]" pageId="25" pageNumber="1557">
<heading box="[566,749,297,322]" centered="true" fontSize="11" level="2" pageId="25" pageNumber="1557" reason="5">
(
<figureCitation box="[574,674,297,322]" captionStart="Figure 4" captionStartId="13.[160,225,936,959]" captionTargetId="figure@14.[162,1153,142,1037]" captionTargetPageId="14" captionText="Figure 4. Gastropods from the Caballas Formation. (a–e) Nerita squatina sp. nov., B8771. (a) UWBM 107577, holotype, W 5.7 mm, spire. (b) UWBM 107648, paratype, W 8.5 mm, shoulder of last whorl. (c) UWBM 107649,paratype, W 10.1 mm, anterior of outer lip.(d) UWBM 107647,paratype,W 7.8 mm,inner surface of outer lip. (e) UWBM 107646, paratype, W 6.2 mm, columnar teeth. (f) Calyptraea cf. C. aperta Solander, 1766, UWBM 107578, B8772,W (diameter) 9.9 mm, external mould. (g) Ampullina ortoni Gabb, 1870, UWBM 107,627, B8772, L 36.8 mm, abapertural view, last whorl mostly missing; left margin annotated. (h–l) Melanella indicaformis sp. nov. (h) UWBM 107620, paratype, B8772, L 8.2 mm, apertural view. (i) UWBM 107619, paratype, B8769, L 9.4 mm, apertural/lateral view. (j) UWBM 107623, paratype, B8769,L 8.2 mm, apertural view. (k) UWBM 107618, paratype, B8772, L 10.6, abapertural view. (l) UWBM 107617, holotype, B8769, L 11.1 mm, abapertural view. (m–q) Potamides henryi sp. nov. (m) UWBM 107582, holotype, B8772, L 17.8 mm, lateral view. (n) UWBM 107582, W 6.9 mm, basal view. (o) UWBM 107586, paratype, B8771, L 11.2 mm, lateral view. (p) UWBM 107587, paratype, B8769, L 9.5 mm, abapertural view. (q) UWBM 107650, paratype, B8770, L 15.4 mm, lateral view, shell compressed. (r–t) Potamides janeae sp. nov., B8771.(r) UWBM 107579, holotype, L 9.0 mm, lateral view. (s) MUSM INV 249, L 8.8 mm, lateral and columellar view. (t) UWBM 107580, L 9.5 mm, layeral and columellar view. (u, v, x) Papposilenus utriculus gen.nov., sp. nov. (u) UWBM 107592,paratype, B8772, L 18.7 mm, oblique view of base. (v) UWBM 107589, holotype, B8772, L 25.5 mm, lateral view. (x) UWBM 107590, B8770, L 17.6 mm, lateral view.(w, y,z,jj) Rhinotamides everriculum sp.nov. (w) UWBM 107598,paratype, B8772,L 23.5 mm, abapertural view, juvenile. (y) UWBM 107593, holotype, B8769, L 21.1 mm, abapertural view. (z) UWBM 107594, paratype, B8772, L 19.4 mm, apertural view, posterior and anterior missing. (jj) UWBM 107593, apertural view.(aa,bb) Terebralia pauli gen. nov.sp.nov., UWBM 107599,holotype,B8771, L 9.2 mm.(aa) Abapertural view. (bb) Apertural view, outer lip missing. (cc) Terebralia marki gen. nov. sp. nov., UWBM 107600, holotype, B8771, L 9.8, lateral view. (dd–ii). Nodifaunus gainesi sp. nov. (dd) UWBM 107609, holotype, B8769, L 42.4 mm, apertural view. (ee) MUSM INV 264, paratype, B8772, L 31.0 mm, apertural view, anterior missing. (ff) UWBM 107609, abapertural view. (gg) UWBM 107612, paratype, B8770, L 14.2 mm, abapertural view, juvenile. (hh) UWBM 107612, apertural view. (ii) UWBM 107610, paratype, B8769, L 25.9 mm, apertural view, anterior missing." figureDoi="http://doi.org/10.5281/zenodo.3686966" httpUri="https://zenodo.org/record/3686966/files/figure.png" pageId="25" pageNumber="1557">Figure 4</figureCitation>
(m
<emphasis box="[703,717,297,322]" italics="true" pageId="25" pageNumber="1557">–</emphasis>
q))
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="25" pageNumber="1557" type="diagnosis">
<paragraph blockId="25.[160,1156,369,499]" box="[160,276,369,395]" pageId="25" pageNumber="1557">
<emphasis bold="true" box="[160,276,369,395]" italics="true" pageId="25" pageNumber="1557">Diagnosis</emphasis>
</paragraph>
<paragraph blockId="25.[160,1156,369,499]" pageId="25" pageNumber="1557">Shell with narrow spire. Whorls with four beaded primary spiral cords, posteriormost strongest; later whorls often with additional one to three beaded secondary spiral cords. Varices and ventrolateral varix absent. Columella with single fold.</paragraph>
</subSubSection>
<subSubSection pageId="25" pageNumber="1557" type="description">
<paragraph blockId="25.[160,1156,546,1127]" box="[160,294,546,572]" pageId="25" pageNumber="1557">
<emphasis bold="true" box="[160,294,546,572]" italics="true" pageId="25" pageNumber="1557">Description</emphasis>
</paragraph>
<paragraph blockId="25.[160,1156,546,1127]" pageId="25" pageNumber="1557">
Estimated shell length
<quantity box="[418,541,583,607]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="25" pageNumber="1557" unit="mm" value="17.5" valueMax="20.0" valueMin="15.0">
15
<emphasis box="[445,458,583,607]" italics="true" pageId="25" pageNumber="1557">–</emphasis>
20 mm
</quantity>
; elongate, spire slightly convex, spire angle about 16°. Protoconch poorly preserved. Spire with about eight whorls. Whorls flat-sided, sutures thinly incised, recessed. Earliest teleoconch whorls with three beaded spiral cords. Later whorls with four beaded spiral cords: posteriormost widest, adjacent to posterior suture, with axially elongate beads; anteriormost cord, nearly adjacent to suture, second widest, with spherical or spirally elongate oval beads; second cord from the anterior suture third widest, with small, partly separated spherical beads; and second cord from the posterior margin most narrow, with separated spirally elongate oval beads; lattermost spiral cord strengthens on last whorls to match strength of second most anterior cord. Secondary beaded spiral cord usually present between two posteriormost primary spiral cords; rarely with intercalation of other beaded secondary spiral threads. Anterior suture bordered with barely visible beaded spiral cord marking edge of base; latter spiral cord bordered adaxially by rounded groove with medial thread, strong spiral cord with radially elongate beads, at least one thin spiral cord near axis, and convex axial threads. Varices and ventrolateral varix absent. Columella with single fold posterior to midpoint. Inner lip with erect flange arising from parietal area; anterior end not preserved. Outer lip flaring with terminal varix but poorly preserved.
</paragraph>
</subSubSection>
<subSubSection pageId="25" pageNumber="1557" type="discussion">
<paragraph blockId="25.[160,1156,1174,1373]" box="[160,262,1174,1200]" pageId="25" pageNumber="1557">
<emphasis bold="true" box="[160,262,1174,1200]" italics="true" pageId="25" pageNumber="1557">Remarks</emphasis>
</paragraph>
<paragraph blockId="25.[160,1156,1174,1373]" pageId="25" pageNumber="1557">
Specimens of
<taxonomicName authority="DeVries, 2019" authorityName="DeVries" authorityYear="2019" box="[328,521,1211,1235]" class="Gastropoda" family="Potamididae" genus="Potamides" kingdom="Animalia" pageId="25" pageNumber="1557" phylum="Mollusca" rank="species" species="henryi" status="sp. nov.">
<emphasis box="[328,521,1211,1235]" italics="true" pageId="25" pageNumber="1557">Potamides henryi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[534,629,1211,1235]" pageId="25" pageNumber="1557" rank="species">sp. nov.</taxonomicNameLabel>
, which occur together with specimens of
<taxonomicName authority="DeVries, 2019" authorityName="DeVries" authorityYear="2019" class="Gastropoda" family="Potamididae" genus="Potamides" kingdom="Animalia" pageId="25" pageNumber="1557" phylum="Mollusca" rank="species" species="janeae" status="sp. nov.">
<emphasis italics="true" pageId="25" pageNumber="1557">P. janeae</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[241,332,1245,1269]" pageId="25" pageNumber="1557" rank="species">sp. nov.</taxonomicNameLabel>
, are generally more elongate and the spires more convexly conical than those of
<taxonomicName authorityName="DeVries" authorityYear="2019" box="[268,371,1280,1304]" class="Gastropoda" family="Potamididae" genus="Potamides" kingdom="Animalia" pageId="25" pageNumber="1557" phylum="Mollusca" rank="species" species="janeae">
<emphasis box="[268,371,1280,1304]" italics="true" pageId="25" pageNumber="1557">P. janeae</emphasis>
</taxonomicName>
. Specimens of
<taxonomicName authorityName="DeVries" authorityYear="2019" box="[555,653,1280,1304]" class="Gastropoda" family="Potamididae" genus="Potamides" kingdom="Animalia" pageId="25" pageNumber="1557" phylum="Mollusca" rank="species" species="henryi">
<emphasis box="[555,653,1280,1304]" italics="true" pageId="25" pageNumber="1557">P. henryi</emphasis>
</taxonomicName>
have four or more beaded primary cords, rather than the three beaded spiral cords on specimens of
<taxonomicName authorityName="DeVries" authorityYear="2019" box="[841,941,1315,1339]" class="Gastropoda" family="Potamididae" genus="Potamides" kingdom="Animalia" pageId="25" pageNumber="1557" phylum="Mollusca" rank="species" species="janeae">
<emphasis box="[841,941,1315,1339]" italics="true" pageId="25" pageNumber="1557">P. janeae</emphasis>
</taxonomicName>
, and have beaded spiral cords on the base.
</paragraph>
</subSubSection>
<subSubSection pageId="25" pageNumber="1557" type="materials_examined">
<paragraph blockId="25.[160,1157,1420,1654]" box="[160,259,1420,1446]" pageId="25" pageNumber="1557">
<emphasis bold="true" box="[160,259,1420,1446]" italics="true" pageId="25" pageNumber="1557">Material</emphasis>
</paragraph>
<paragraph blockId="25.[160,1157,1420,1654]" pageId="25" pageNumber="1557">
<materialsCitation ID-GBIF-Occurrence="2572159726" box="[160,864,1457,1481]" collectionCode="UWBM" pageId="25" pageNumber="1557" specimenCode="UWBM 107582" specimenCount="1" typeStatus="holotype">
<specimenCode box="[160,328,1457,1481]" pageId="25" pageNumber="1557">UWBM 107582</specimenCode>
,
<typeStatus box="[337,440,1457,1481]" pageId="25" pageNumber="1557">holotype</typeStatus>
, B8772 (type locality), L (17.8), W (6.9)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2572159719" collectionCode="UWBM" pageId="25" pageNumber="1557" specimenCode="UWBM 107583" specimenCount="1" typeStatus="paratype">
remainder are
<typeStatus box="[1040,1153,1457,1481]" pageId="25" pageNumber="1557">paratypes</typeStatus>
:
<specimenCode box="[160,330,1492,1516]" pageId="25" pageNumber="1557">UWBM 107583</specimenCode>
, B8772, L (14.9), W (6.3)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2572159725" box="[620,1066,1492,1516]" collectionCode="UWBM" pageId="25" pageNumber="1557" specimenCode="UWBM 107584" specimenCount="1">
<specimenCode box="[620,790,1492,1516]" pageId="25" pageNumber="1557">UWBM 107584</specimenCode>
, B8772, L (15.2), W (6.6)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2572159717" collectionCode="UWBM" pageId="25" pageNumber="1557" specimenCode="UWBM 107585" specimenCount="1">
<specimenCode pageId="25" pageNumber="1557">UWBM 107585</specimenCode>
, B8772, L (12.6), W (6.0)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2572159723" box="[530,971,1526,1550]" collectionCode="UWBM" pageId="25" pageNumber="1557" specimenCode="UWBM 107586" specimenCount="1">
<specimenCode box="[530,699,1526,1550]" pageId="25" pageNumber="1557">UWBM 107586</specimenCode>
, B8771, L (11.2), W (6.0)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2572159720" collectionCode="UWBM" longitude="-6.9" pageId="25" pageNumber="1557" specimenCode="UWBM 107587" specimenCount="1">
<specimenCode box="[983,1153,1526,1550]" pageId="25" pageNumber="1557">UWBM 107587</specimenCode>
, B8769, L (9.5), W 6.9
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2572159728" box="[411,861,1561,1585]" collectionCode="UWBM" pageId="25" pageNumber="1557" specimenCode="UWBM 107588" specimenCount="1">
<specimenCode box="[411,582,1561,1585]" pageId="25" pageNumber="1557">UWBM 107588</specimenCode>
, B8769, L (11.7), W (5.7)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2572159718" collectionCode="UWBM" pageId="25" pageNumber="1557" specimenCode="UWBM 107650" specimenCount="1">
<specimenCode box="[875,1045,1561,1585]" pageId="25" pageNumber="1557">UWBM 107650</specimenCode>
, B8770, L (15.4), W (7.4)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2572159721" box="[326,764,1596,1620]" collectionCode="MUSM" pageId="25" pageNumber="1557" specimenCode="INV 250" specimenCount="1">
<collectionCode box="[326,399,1596,1620]" pageId="25" pageNumber="1557">MUSM</collectionCode>
<specimenCode box="[407,497,1596,1620]" pageId="25" pageNumber="1557">INV 250</specimenCode>
, B8772, L (14.0), W (7.9)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2572159724" collectionCode="MUSM" pageId="25" pageNumber="1557" specimenCode="INV 251" specimenCount="1">
<collectionCode box="[775,848,1596,1620]" pageId="25" pageNumber="1557">MUSM</collectionCode>
<specimenCode box="[855,946,1596,1620]" pageId="25" pageNumber="1557">INV 251</specimenCode>
, B8772, L (15.8), W (7.7)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2572159733" box="[222,664,1630,1654]" collectionCode="MUSM" pageId="25" pageNumber="1557" specimenCode="INV 252" specimenCount="1">
<collectionCode box="[222,295,1630,1654]" pageId="25" pageNumber="1557">MUSM</collectionCode>
<specimenCode box="[304,396,1630,1654]" pageId="25" pageNumber="1557">INV 252</specimenCode>
, B8772, L (10.0), W (5.6
</materialsCitation>
</paragraph>
<paragraph blockId="25.[160,1157,1420,1654]" box="[664,674,1630,1654]" pageId="25" pageNumber="1557">).</paragraph>
</subSubSection>
<subSection lastPageId="26" lastPageNumber="1558" pageId="25" pageNumber="1557" type="multiple">
<subSubSection box="[160,839,142,1727]" lastPageId="26" lastPageNumber="1558" pageId="25" pageNumber="1557" type="etymology">
<paragraph blockId="25.[160,285,1701,1727]" box="[160,285,1701,1727]" pageId="25" pageNumber="1557">
<emphasis bold="true" box="[160,285,1701,1727]" italics="true" pageId="25" pageNumber="1557">Etymology</emphasis>
</paragraph>
<paragraph blockId="26.[160,839,142,166]" box="[160,839,142,166]" pageId="26" pageNumber="1558">Named in honour of Henry R. DeVries, father of the author.</paragraph>
</subSubSection>
<subSubSection box="[160,1156,215,276]" pageId="26" pageNumber="1558" type="distribution">
<paragraph blockId="26.[160,1156,215,276]" box="[160,292,215,241]" pageId="26" pageNumber="1558">
<emphasis bold="true" box="[160,292,215,241]" italics="true" pageId="26" pageNumber="1558">Occurrence</emphasis>
</paragraph>
<paragraph blockId="26.[160,1156,215,276]" box="[160,1156,252,276]" pageId="26" pageNumber="1558">
Lower Paleogene, Cuenca Member, Caballas Formation, East Pisco Basin, southern
<collectingCountry box="[1099,1153,252,276]" name="Peru" pageId="26" pageNumber="1558">Peru</collectingCountry>
.
</paragraph>
</subSubSection>
</subSection>
</treatment>
</document>
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