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<document id="DC22057EE379BB142AA4F5B04F153388" ID-DOI="10.11646/zootaxa.5567.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="14703735" ID-ZooBank="AF6254F6-CE26-41FE-BB85-A3FFD16FAA39" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737371447294" checkinUser="plazi" docAuthor="VIRAKTAMATH, C. A., WEBB, M. D. &amp; YESHWANTH, H. M." docDate="2025" docId="03B75F43F05DFF88FF48767034D32536" docLanguage="en" docName="zootaxa.5567.1.1.pdf" docOrigin="Zootaxa 5567 (1)" docSource="https://doi.org/10.11646/zootaxa.5567.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Madhyabus reticulatus VIRAKTAMATH, WEBB &amp; YESHWANTH, 2025, sp. nov." docType="treatment" docVersion="2" lastPageNumber="21" masterDocId="FF8E273BF049FF9CFFDF7756364D2343" masterDocTitle="Review of the Monobazus-group leafhopper genera (Hemiptera: Cicadellidae: Deltocephalinae) from the Indian subcontinent with description of five new genera and twenty-eight new species" masterLastPageNumber="105" masterPageNumber="1" pageNumber="21" updateTime="1737389185710" updateUser="ExternalLinkService">
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<mods:title id="97651C883CB61DCD9A7E9C171654BEFA">Review of the Monobazus-group leafhopper genera (Hemiptera: Cicadellidae: Deltocephalinae) from the Indian subcontinent with description of five new genera and twenty-eight new species</mods:title>
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<mods:namePart id="BFDDE5292529005D0EDB5746B98F5B55">VIRAKTAMATH, C. A.</mods:namePart>
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<mods:namePart id="A2310F8A331A76EDB83C62625D333281">WEBB, M. D.</mods:namePart>
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<mods:date id="2C4C899C8A5CA3231AD5EBE4DFDD8903">2025</mods:date>
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<treatment id="03B75F43F05DFF88FF48767034D32536" LSID="urn:lsid:plazi:treatment:03B75F43F05DFF88FF48767034D32536" httpUri="http://treatment.plazi.org/id/03B75F43F05DFF88FF48767034D32536" lastPageNumber="21" pageId="20" pageNumber="21">
<subSubSection id="C304BDDEF05DFF88FF487670345F2202" box="[151,530,294,321]" pageId="20" pageNumber="21" type="nomenclature">
<paragraph id="8BA1EE55F05DFF88FF487670345F2202" blockId="20.[151,530,294,357]" box="[151,530,294,321]" pageId="20" pageNumber="21">
<heading id="D0E95939F05DFF88FF487670345F2202" bold="true" box="[151,530,294,321]" fontSize="11" level="2" pageId="20" pageNumber="21" reason="4">
<emphasis id="B96A3247F05DFF88FF487670345F2202" bold="true" box="[151,530,294,321]" pageId="20" pageNumber="21">
<taxonomicName id="4C1E95D6F05DFF88FF48767037E02202" authorityName="VIRAKTAMATH &amp; WEBB &amp; YESHWANTH" authorityYear="2025" box="[151,429,294,321]" class="Insecta" family="Cicadellidae" genus="Madhyabus" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="reticulatus" status="sp. nov.">
<emphasis id="B96A3247F05DFF88FF48767037E02202" bold="true" box="[151,429,294,321]" italics="true" pageId="20" pageNumber="21">Madhyabus reticulatus</emphasis>
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<taxonomicNameLabel id="A2598F3CF05DFF88FE6B7671345F2202" box="[436,530,295,321]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C304BDDEF05DFF88FF48761D3449256E" pageId="20" pageNumber="21" type="description">
<paragraph id="8BA1EE55F05DFF88FF48761D37CC2226" blockId="20.[151,530,294,357]" box="[151,385,331,357]" pageId="20" pageNumber="21">
<figureCitation id="1325F2D0F05DFF88FF48761D36912226" box="[151,220,331,357]" captionStart="FIGURES 2" captionStartId="43.[151,264,1897,1921]" captionTargetBox="[163,1426,181,1869]" captionTargetId="figure-17@43.[158,1430,181,1874]" captionTargetPageId="43" captionText="FIGURES 2AR. Monobazus group of genera, habitus, dorsal and lateral views. AD, Lankabanus attenuatus gen. et sp. nov., holotype male (AB) and paratype female (CD); EH, Madhyabus biligiris gen. et sp. nov., holotype male (EF), paratype female (GH); IJ, M. reticulatus sp. nov., holotype male; KN, Mavromoustaca brevispina sp. nov., holotype male (KL), paratype female (MN); OR, M. channabasavanna sp. nov., holotype male (OP), paratype female (QR)." figureDoi="http://doi.org/10.5281/zenodo.14703743" httpUri="https://zenodo.org/record/14703743/files/figure.png" pageId="20" pageNumber="21">Figs 2</figureCitation>
IJ, 8N, 24AG
</paragraph>
<paragraph id="8BA1EE55F05DFF88FF4876C5322D21C6" blockId="20.[151,1437,403,1653]" pageId="20" pageNumber="21">
<emphasis id="B96A3247F05DFF88FF4876C5374422EF" box="[151,265,403,428]" italics="true" pageId="20" pageNumber="21">Diagnosis</emphasis>
. Ochraceous with dark brown markings on the head, thorax and forewings. Forewing venation reticulate. Male pygofer about 1.2× as long as broad in lateral view, ventral process blade-like extending along ventral margin. Valve with median short angulate projection on the posterior margin. Aedeagus compressed, with well-developed dorsal apodeme, with gonopore at midlength on the ventral surface, atrium not elongate. Externally the species resembles Palaearctic
<taxonomicName id="4C1E95D6F05DFF88FE4775753544217E" authority="Emeljanov" authorityName="Emeljanov" authorityYear="1964" box="[408,777,547,573]" class="Insecta" family="Cicadellidae" genus="Allotapes" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="nigrifacies">
<emphasis id="B96A3247F05DFF88FE47757534C8217E" box="[408,645,547,573]" italics="true" pageId="20" pageNumber="21">Allotapes nigrifacies</emphasis>
Emeljanov
</taxonomicName>
(see,
<bibRefCitation id="EF8F93A4F05DFF88FC8C75753204217E" author="Dai, W. &amp; Dietrich, C. H." box="[851,1097,547,573]" pageId="20" pageNumber="21" pagination="3 - 4" refId="ref49796" refString="Dai, W. &amp; Dietrich, C. H. (2011) Review of the Old World leafhopper genus Scaphoidella Vilbaste (Hemiptera: Cicadellidae: Deltocephalinae) with description of ten new species from Thailand and Vietnam. Annales de la Societe entomologique de France (N. S.), 47 (3 - 4), 457 - 473. https: // doi. org / 10.1080 / 00379271.2011.10697737" type="journal article" year="2011">Dai &amp; Dietrich 2011</bibRefCitation>
: figs 7077) but has entirely different male genitalia: male pygofer with a ventral process (absent in
<taxonomicName id="4C1E95D6F05DFF88FC18751132122122" authorityName="Emeljanov" authorityYear="1964" box="[967,1119,583,609]" class="Insecta" family="Cicadellidae" genus="Allotapes" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="nigrifacies">
<emphasis id="B96A3247F05DFF88FC18751132122122" box="[967,1119,583,609]" italics="true" pageId="20" pageNumber="21">A. nigrifacies</emphasis>
</taxonomicName>
), aedeagus compressed and without basal processes (aedeagus cylindrical with ventral processes in
<taxonomicName id="4C1E95D6F05DFF88FC66753D321D21C6" authorityName="Emeljanov" authorityYear="1964" box="[953,1104,619,645]" class="Insecta" family="Cicadellidae" genus="Allotapes" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="nigrifacies">
<emphasis id="B96A3247F05DFF88FC66753D321D21C6" box="[953,1104,619,645]" italics="true" pageId="20" pageNumber="21">A. nigrifacies</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BA1EE55F05DFF88FF1875D9330120AE" blockId="20.[151,1437,403,1653]" pageId="20" pageNumber="21">
<emphasis id="B96A3247F05DFF88FF1875D9370621EB" box="[199,331,655,680]" italics="true" pageId="20" pageNumber="21">Description</emphasis>
. Ochraceous with reddish brown to dark brown markings on head, pronotum, exposed mesonotum and forewings (
<figureCitation id="1325F2D0F05DFF88FE9675E537DF218E" box="[329,402,691,717]" captionStart="FIGURES 2" captionStartId="43.[151,264,1897,1921]" captionTargetBox="[163,1426,181,1869]" captionTargetId="figure-17@43.[158,1430,181,1874]" captionTargetPageId="43" captionText="FIGURES 2AR. Monobazus group of genera, habitus, dorsal and lateral views. AD, Lankabanus attenuatus gen. et sp. nov., holotype male (AB) and paratype female (CD); EH, Madhyabus biligiris gen. et sp. nov., holotype male (EF), paratype female (GH); IJ, M. reticulatus sp. nov., holotype male; KN, Mavromoustaca brevispina sp. nov., holotype male (KL), paratype female (MN); OR, M. channabasavanna sp. nov., holotype male (OP), paratype female (QR)." figureDoi="http://doi.org/10.5281/zenodo.14703743" httpUri="https://zenodo.org/record/14703743/files/figure.png" pageId="20" pageNumber="21">Figs 2</figureCitation>
IJ). Crown with one large irregular spot on either side of median line with lateral marginal dark brown border; series of small spots on anterior submargin dark brown. Frontoclypeus dark fuscous laterally and ochraceous medially; large spot surrounding base of antenna, upper half of lateral margins of lorum, one stripe on gena, lateral margins of clypoclypellar suture, dark brown. Pronotum with anterior submargin yellow with transverse dark brown spots laterally; disc piceous brown. Exposed mesonotum ochraceous, basal triangles and two small round spots between them, scutoscutellar suture, one transverse line across mesoscutellum at midlength, dark brown to black. Forewing cross veins marked with dark brown, one spot on cross vein connective claval veins, apex of clavus, one basal spot on corium, one spot on median area, three spots on costal margin (one slightly distad of median length, and other two on apices of veins reaching costal margin) dark brown. Some veins brownish.
</paragraph>
<paragraph id="8BA1EE55F05DFF88FF1874A13721271A" blockId="20.[151,1437,403,1653]" pageId="20" pageNumber="21">Head slightly wider than pronotum. Crown 0.7× as long as median length, anteriorly subconically rounded. Eyes large more or less transverse. Pronotum 2.1× as wide as median length. Forewing richly reticulate both on corium and clavus.</paragraph>
<paragraph id="8BA1EE55F05DFF88FF187335355527AA" blockId="20.[151,1437,403,1653]" pageId="20" pageNumber="21">
<emphasis id="B96A3247F05DFF88FF1873353723273E" box="[199,366,1123,1149]" italics="true" pageId="20" pageNumber="21">Male genitalia</emphasis>
. Male pygofer about 1.2× as long as broad in lateral view, ventral process blade-like running along ventral margin. Valve with median short angulate projection on posterior margin. Aedeagus strongly curved and compressed, almost U-shaped, with well-developed dorsal apodeme, atrium short, shaft apex obliquely truncate in lateral view, gonopore at midlength on ventral surface.
</paragraph>
<paragraph id="8BA1EE55F05DFF88FF1873A5373C2616" blockId="20.[151,1437,403,1653]" pageId="20" pageNumber="21">
<emphasis id="B96A3247F05DFF88FF1873A53724264F" box="[199,361,1267,1292]" italics="true" pageId="20" pageNumber="21">Measurements</emphasis>
. Male
<quantity id="4CE643B0F05DFF88FE6473A53409264E" box="[443,580,1267,1293]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" metricValueMax="4.3" metricValueMin="3.9" pageId="20" pageNumber="21" unit="mm" value="4.1" valueMax="4.3" valueMin="3.9">3.94.3 mm</quantity>
long, 1.0
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wide across eyes and 1.0 mm wide across posterolateral angles of pronotum. Female
<quantity id="4CE643B0F05DFF88FE3B724034292672" box="[484,612,1302,1329]" metricMagnitude="-3" metricUnit="m" metricValue="4.85" metricValueMax="5.1" metricValueMin="4.6" pageId="20" pageNumber="21" unit="mm" value="4.85" valueMax="5.1" valueMin="4.6">4.65.1mm</quantity>
long,
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wide across eyes and
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wide across posterolateral angles of pronotum
</paragraph>
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<emphasis id="B96A3247F05DFF88FF18720937D5263A" box="[199,408,1375,1401]" italics="true" pageId="20" pageNumber="21">Material examined</emphasis>
.
<materialsCitation id="3B76E408F05DFF88FE7C720936B526DE" collectingDate="1996-10-26" collectionCode="UASB" country="India" county="Nehru Nagar" location="Kalaichelvan" municipality="Durg Dist." pageId="20" pageNumber="21" specimenCount="1" specimenCount-male="1" stateProvince="Madhya Pradesh" typeStatus="holotype">
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,
<collectingCountry id="F309AEC5F05DFF88FD85720934E3263A" box="[602,686,1375,1401]" name="India" pageId="20" pageNumber="21">INDIA</collectingCountry>
:
<collectingRegion id="49DA20B7F05DFF88FD6A7209353F263A" box="[693,882,1375,1401]" country="India" name="Madhya Pradesh" pageId="20" pageNumber="21">Madhya Pradesh</collectingRegion>
:
<collectingCounty id="62C096D9F05DFF88FCA672093245263A" box="[889,1032,1375,1401]" pageId="20" pageNumber="21">Nehru Nagar</collectingCounty>
,
<collectingMunicipality id="6BC5742FF05DFF88FBCE720932CB263A" box="[1041,1158,1375,1401]" pageId="20" pageNumber="21">Durg Dist.</collectingMunicipality>
,
<date id="FFA0C895F05DFF88FB517209334D263A" box="[1166,1280,1375,1401]" pageId="20" pageNumber="21" value="1996-10-26">
<collectingDate id="EFE4317DF05DFF88FB517209334D263A" box="[1166,1280,1375,1401]" pageId="20" pageNumber="21" value="1996-10-26">26.x.1996</collectingDate>
</date>
,
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(
<collectionCode id="ED0F7690F05DFF88FF4072D536BD26DE" box="[159,240,1411,1437]" country="India" name="University of Agricultural Sciences" pageId="20" pageNumber="21">UASB</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B76E408F05DFF88FEDD72D537BA2682" collectingDate="2010-11-02" collectorName="Yeshwanth, H. M" country="India" elevation="277" location="Tharapuram" pageId="20" pageNumber="21" specimenCount="6" specimenCount-female="4" specimenCount-male="2" stateProvince="Tamil Nadu" typeStatus="paratype">
<typeStatus id="54A550F7F05DFF88FEDD72D537ED26DE" box="[258,416,1411,1437]" pageId="20" pageNumber="21" type="paratype">PARATYPES</typeStatus>
.
<collectingCountry id="F309AEC5F05DFF88FE7572D537B326DE" box="[426,510,1411,1437]" name="India" pageId="20" pageNumber="21">INDIA</collectingCountry>
:
<collectingRegion id="49DA20B7F05DFF88FDD972D534C226DE" box="[518,655,1411,1437]" country="India" name="Tamil Nadu" pageId="20" pageNumber="21">Tamil Nadu</collectingRegion>
:
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,
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,
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,
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89.3” N,
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810” E,
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<collectingDate id="EFE4317DF05DFF88FAF472D533D526DE" box="[1323,1432,1411,1437]" pageId="20" pageNumber="21" value="2010-11-02">2.xi.2010</collectingDate>
</date>
,
<collectingMethod id="525F9642F05DFF88FF4872F137692682" box="[151,292,1447,1473]" pageId="20" pageNumber="21">Suction trap</collectingMethod>
,
<collectorName id="26EB8B83F05DFF88FEEE72F137BA2682" box="[305,503,1447,1473]" pageId="20" pageNumber="21">Yeshwanth, H.M</collectorName>
</materialsCitation>
.;
<materialsCitation id="3B76E408F05DFF88FDD672F137D226A6" collectingDate="2010-11-11" collectionCode="NBAIR, UASB" collectorName="Yeshwanth, H. M." country="India" elevation="1311" location="Thandigudi" pageId="20" pageNumber="21" specimenCount="1" specimenCount-female="1" stateProvince="Tamil Nadu" typeStatus="paratype">
<specimenCount id="9D1825DCF05DFF88FDD672F134632682" box="[521,558,1447,1473]" count="1" pageId="20" pageNumber="21" type="female">1♀</specimenCount>
:
<location id="8EC1B88EF05DFF88FDE272F134F32682" LSID="urn:lsid:plazi:treatment:03B75F43F05DFF88FF48767034D32536:8EC1B88EF05DFF88FDE272F134F32682" box="[573,702,1447,1473]" country="India" name="Thandigudi" pageId="20" pageNumber="21" stateProvince="Tamil Nadu">Thandigudi</location>
<quantity id="4CE643B0F05DFF88FD1772F1355A2682" box="[712,791,1447,1473]" metricMagnitude="3" metricUnit="m" metricValue="1.311" pageId="20" pageNumber="21" unit="m" value="1311.0">
<elevation id="00330966F05DFF88FD1772F1355A2682" box="[712,791,1447,1473]" metricMagnitude="3" metricUnit="m" metricValue="1.311" pageId="20" pageNumber="21" unit="m" value="1311.0">1311m</elevation>
</quantity>
,
<superScript id="7C6B431DF05DFF88FCFA72F1352A2682" attach="none" box="[805,871,1446,1473]" fontSize="6" pageId="20" pageNumber="21">10018</superScript>
N,
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E,
<date id="FFA0C895F05DFF88FBD272F132CB2682" box="[1037,1158,1447,1473]" pageId="20" pageNumber="21" value="2010-11-11">
<collectingDate id="EFE4317DF05DFF88FBD272F132CB2682" box="[1037,1158,1447,1473]" pageId="20" pageNumber="21" value="2010-11-11">11.xi.2010</collectingDate>
</date>
,
<collectingMethod id="525F9642F05DFF88FB4B72F133402682" box="[1172,1293,1447,1473]" pageId="20" pageNumber="21">Sweep net</collectingMethod>
,
<collectorName id="26EB8B83F05DFF88FAC572F1369E26A6" pageId="20" pageNumber="21">Yeshwanth, H.M.</collectorName>
(
<collectionCode id="ED0F7690F05DFF88FF3E729D377026A6" box="[225,317,1483,1509]" pageId="20" pageNumber="21">NBAIR</collectionCode>
,
<collectionCode id="ED0F7690F05DFF88FE97729D37D526A6" box="[328,408,1483,1509]" country="India" name="University of Agricultural Sciences" pageId="20" pageNumber="21">UASB</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BA1EE55F05DFF88FF1872B93449256E" blockId="20.[151,1437,403,1653]" pageId="20" pageNumber="21">
<emphasis id="B96A3247F05DFF88FF1872B93772254A" box="[199,319,1519,1545]" italics="true" pageId="20" pageNumber="21">Etymology</emphasis>
. The specific epithet is the Latin adjective reticulatus, -a, -um (= net-like, reticulated), referring to the reticulate forewing venation.
</paragraph>
</subSubSection>
<subSubSection id="C304BDDEF05DFF88FF18716134D32536" pageId="20" pageNumber="21" type="discussion">
<paragraph id="8BA1EE55F05DFF88FF18716134D32536" blockId="20.[151,1437,403,1653]" pageId="20" pageNumber="21">
<emphasis id="B96A3247F05DFF88FF18716137792512" bold="true" box="[199,308,1591,1617]" pageId="20" pageNumber="21">Remarks</emphasis>
. The aedeagus in specimens from Tarapuram (
<collectingRegion id="49DA20B7F05DFF88FC9D716135872512" box="[834,970,1591,1617]" country="India" name="Tamil Nadu" pageId="20" pageNumber="21">Tamil Nadu</collectingRegion>
) have the ventral surface of shaft apex in lateral view slightly more angularly produced.
</paragraph>
</subSubSection>
</treatment>
</document>