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<document id="291363CE79412BB591FC584B4708A187" ID-CLB-Dataset="22222" ID-DOI="10.5852/ejt.2012.6" ID-GBIF-Dataset="4474424d-176b-4799-b758-85f890cedb7a" ID-Zenodo-Dep="3822694" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589313615730" checkinUser="carolina" docAuthor="Johanson, Kjell A. &amp; Pham, Hong-Thai" docDate="2012" docId="03AF87F9527CFF9BB468FED6FB2DFE19" docLanguage="en" docName="ejt-2011-6.pdf.imf" docOrigin="European Journal of Taxonomy 6" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Helicopsyche meander Johanson &amp; Pham 2012, sp. nov." docType="treatment" docUuid="254700A7-8643-4D20-8474-F4AD71215009" docUuidSource="ZooBank" docVersion="6" lastPageNumber="5" masterDocId="FF96FF815278FF9EB608FFDDFFD5FFAB" masterDocTitle="Three new species of Helicopsyche (Trichoptera, Helicopsychidae) from northern Vietnam, with a key to Helicopsyche species of Vietnam" masterLastPageNumber="10" masterPageNumber="1" pageNumber="5" updateTime="1698839278927" updateUser="ExternalLinkService">
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<mods:title id="8EB125E93300C5F6A370F2B58CA69E79">Three new species of Helicopsyche (Trichoptera, Helicopsychidae) from northern Vietnam, with a key to Helicopsyche species of Vietnam</mods:title>
</mods:titleInfo>
<mods:name id="31F0FE7A0B00DDE408BD96F539DD0E74" type="personal">
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<mods:namePart id="69E61A908DFD8177AE4BE700516A6BFC">Johanson, Kjell A.</mods:namePart>
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<mods:namePart id="FBD63750567B2C363E8E1F77CE28FDB1">Pham, Hong-Thai</mods:namePart>
</mods:name>
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<mods:title id="3EB7537546543E2668ECFC77E5C244FC">European Journal of Taxonomy</mods:title>
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<mods:part id="BD02C7FFB74FD34C24748EADC1955E46">
<mods:date id="B541D7768266C2D8556A0EE98644EBB3">2012</mods:date>
<mods:detail id="D78C59F668ECEFBE74D0BBA259529588" type="pubDate">
<mods:number id="47BE20CE86C56D2137CAAF8AB351584B">2012-02-02</mods:number>
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<mods:identifier id="F01256D996B93ADC06EC42AB7D0D726B" type="DOI">10.5852/ejt.2012.6</mods:identifier>
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<treatment id="03AF87F9527CFF9BB468FED6FB2DFE19" ID-DOI="http://doi.org/10.5281/zenodo.3844363" ID-GBIF-Taxon="164126243" ID-Zenodo-Dep="3844363" LSID="urn:lsid:zoobank.org:act:254700A7-8643-4D20-8474-F4AD71215009" httpUri="http://treatment.plazi.org/id/03AF87F9527CFF9BB468FED6FB2DFE19" lastPageId="5" lastPageNumber="5" pageId="4" pageNumber="5">
<subSubSection id="C31C6564527CFF9AB468FED6FC01FE8D" box="[608,980,267,294]" pageId="4" pageNumber="5" type="nomenclature">
<paragraph id="8BB936EF527CFF9AB468FED6FC01FE8D" blockId="4.[608,980,267,329]" box="[608,980,267,294]" pageId="4" pageNumber="5">
<heading id="D0F18183527CFF9AB468FED6FC01FE8D" box="[608,980,267,294]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="5" reason="2">
<taxonomicName id="4C064D6C527CFF9AB468FED6FCA7FE8D" authority="Johanson &amp; Pham, 2012" authorityName="Johanson &amp; Pham" authorityYear="2012" box="[608,882,267,294]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="meander" status="sp. nov.">
<emphasis id="B972EAFD527CFF9AB468FED6FCA7FE8D" bold="true" box="[608,882,267,294]" italics="true" pageId="4" pageNumber="5">Helicopsyche meander</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2415786527CFF9AB572FED1FC01FE8D" box="[890,980,268,294]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31C6564527CFF9AB4FFFEF2FCE9FEE2" box="[759,828,303,329]" pageId="4" pageNumber="5" type="description">
<paragraph id="8BB936EF527CFF9AB4FFFEF2FCE9FEE2" blockId="4.[608,980,267,329]" box="[759,828,303,329]" pageId="4" pageNumber="5">
<figureCitation id="133D2A6A527CFF9AB4FFFEF2FCE9FEE2" box="[759,828,303,329]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[213,1376,611,1844]" captionTargetId="figure@5.[189,1398,598,1890]" captionTargetPageId="5" captionText="Fig. 2. Helicopsyche meander sp. nov., ♂ holotype. A. abdominal sternum VI, lateral view. B. abdominal sternal process VI, ventral view. C. genitalia, lateral view. D. genitalia, dorsal view. E. genitalia, ventral view. F. phallus, lateral view. G. phallus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3822700" httpUri="https://zenodo.org/record/3822700/files/figure.png" pageId="4" pageNumber="5">Fig. 2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31C6564527CFF9AB6B5FEACFAC3FD39" pageId="4" pageNumber="5" type="diagnosis">
<paragraph id="8BB936EF527CFF9AB6B5FEACFEE3FE20" blockId="4.[189,310,369,395]" box="[189,310,369,395]" pageId="4" pageNumber="5">
<heading id="D0F18183527CFF9AB6B5FEACFEE3FE20" bold="true" box="[189,310,369,395]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="B972EAFD527CFF9AB6B5FEACFEE3FE20" bold="true" box="[189,310,369,395]" pageId="4" pageNumber="5">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BB936EF527CFF9AB6B5FE7DFAC3FD39" blockId="4.[189,1399,416,658]" pageId="4" pageNumber="5">
<taxonomicName id="4C064D6C527CFF9AB6B5FE7DFE12FE11" authority="Johanson &amp; Pham, 2012" authorityName="Johanson &amp; Pham" authorityYear="2012" box="[189,455,416,442]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="meander" status="sp. nov.">
<emphasis id="B972EAFD527CFF9AB6B5FE7DFE12FE11" box="[189,455,416,442]" italics="true" pageId="4" pageNumber="5">Helicopsyche meander</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2415786527CFF9AB7C6FE7CFDFDFE10" box="[462,552,417,443]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
is most similar to
<taxonomicName id="4C064D6C527CFF9AB4F6FE7CFC57FE11" authority="Johanson &amp; Pham, 2012" authorityName="Johanson &amp; Pham" authorityYear="2012" box="[766,898,416,443]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="lamnata" status="sp. nov.">
<emphasis id="B972EAFD527CFF9AB4F6FE7CFCC6FE11" box="[766,787,417,442]" italics="true" pageId="4" pageNumber="5">H</emphasis>
.
<emphasis id="B972EAFD527CFF9AB52AFE7DFC57FE11" box="[802,898,416,442]" italics="true" pageId="4" pageNumber="5">lamnata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2415786527CFF9AB580FE7CFC31FE10" box="[904,996,417,443]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C064D6C527CFF9AB5F8FE7CFA9BFE10" authority="Malicky, 1995" authorityName="Malicky" authorityYear="1995" box="[1008,1358,416,443]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="anaktangga">
<emphasis id="B972EAFD527CFF9AB5F8FE7CFB48FE11" box="[1008,1181,416,442]" italics="true" pageId="4" pageNumber="5">H. anaktangga</emphasis>
Malicky, 1995
</taxonomicName>
,
<taxonomicName id="4C064D6C527CFF9AB352FE7CFE0CFE74" authority="Malicky, 1995" authorityName="Malicky" authorityYear="1995" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="anaksaku">
<emphasis id="B972EAFD527CFF9AB352FE7CFEFEFE75" italics="true" pageId="4" pageNumber="5">H. anaksaku</emphasis>
Malicky, 1995
</taxonomicName>
,
<taxonomicName id="4C064D6C527CFF9AB7EBFE18FB93FE75" authority="Malicky &amp; Nawvong (Malicky et al. 2004)" box="[483,1094,452,479]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="minyas">
<emphasis id="B972EAFD527CFF9AB7EBFE18FD82FE75" box="[483,599,453,478]" italics="true" pageId="4" pageNumber="5">H. minyas</emphasis>
Malicky &amp; Nawvong (
<bibRefCitation id="EF974B1E527CFF9AB557FE19FBEBFE75" author="Malicky H. &amp; Chantaramongkol P. &amp; Bunlue P. &amp; Changthong N. &amp; Nawvong J. &amp; Nuntakwang A. &amp; Prommi T. &amp; Thamsenanupap P. &amp; Thapanya D." box="[863,1086,452,478]" pageId="4" pageNumber="5" pagination="287 - 304" refId="ref4944" refString="Malicky H., Chantaramongkol P., Bunlue P., Changthong N., Nawvong J., Nuntakwang A., Prommi T., Thamsenanupap P. &amp; Thapanya D. 2004. 27 neue Kocherfliegen aus Thailand (Insecta, Trichoptera). Linzer Biologische Beitrage 36: 287 - 304." type="journal article" year="2004">
Malicky
<emphasis id="B972EAFD527CFF9AB5CDFE18FC28FE75" box="[965,1021,452,478]" italics="true" pageId="4" pageNumber="5">et al.</emphasis>
2004
</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C064D6C527CFF9AB273FE18FF2FFDA9" authority="Schmid, 1993" authorityName="Schmid" authorityYear="1993" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="cymodoce">
<emphasis id="B972EAFD527CFF9AB273FE18FAC4FE75" box="[1147,1297,452,478]" italics="true" pageId="4" pageNumber="5">H. cymodoce</emphasis>
<bibRefCitation id="EF974B1E527CFF9AB31EFE19FF2FFDA9" author="Schmid F." pageId="4" pageNumber="5" pagination="65 - 100" refId="ref5172" refString="Schmid F. 1993. Considerations sur les helicopsychides (Trichoptera, Integripalpia). Beaufortia 43: 65 - 100." type="journal article" year="1993">Schmid, 1993</bibRefCitation>
</taxonomicName>
, particularly due to the presence of two-segmented maxillary palps in combination with the nearly similar shape of the gonopods in lateral view. The new species is distinguished from
<taxonomicName id="4C064D6C527CFF9AB296FDD0FA99FD8D" authorityName="Malicky" authorityYear="1995" box="[1182,1356,524,550]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="anaktangga">
<emphasis id="B972EAFD527CFF9AB296FDD0FA99FD8D" box="[1182,1356,524,550]" italics="true" pageId="4" pageNumber="5">H. anaktangga</emphasis>
</taxonomicName>
,
<taxonomicName id="4C064D6C527CFF9AB352FDD0FEFEFDE1" authorityName="Malicky" authorityYear="1995" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="anaksaku">
<emphasis id="B972EAFD527CFF9AB352FDD0FEFEFDE1" italics="true" pageId="4" pageNumber="5">H. anaksaku</emphasis>
</taxonomicName>
,
<taxonomicName id="4C064D6C527CFF9AB734FDECFE60FDE1" box="[316,437,561,586]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="minyas">
<emphasis id="B972EAFD527CFF9AB734FDECFE60FDE1" box="[316,437,561,586]" italics="true" pageId="4" pageNumber="5">H. minyas</emphasis>
</taxonomicName>
and
<taxonomicName id="4C064D6C527CFF9AB7FAFDECFD58FDE1" authorityName="Schmid" authorityYear="1993" box="[498,653,560,586]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="cymodoce">
<emphasis id="B972EAFD527CFF9AB7FAFDECFD58FDE1" box="[498,653,560,586]" italics="true" pageId="4" pageNumber="5">H. cymodoce</emphasis>
</taxonomicName>
by the pointed apex of tergum X in lateral view; and from
<taxonomicName id="4C064D6C527CFF9AB352FDECFEC8FDC5" authority="Johanson &amp; Pham, 2012" authorityName="Johanson &amp; Pham" authorityYear="2012" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="lamnata" status="sp. nov.">
<emphasis id="B972EAFD527CFF9AB352FDECFEC8FDC5" italics="true" pageId="4" pageNumber="5">H. lamnata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2415786527CFF9AB72BFD89FEAAFDC5" box="[291,383,596,622]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C064D6C527CFF9AB783FD88FDC9FDC5" authorityName="Malicky" authorityYear="1995" box="[395,540,596,622]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="anaksaku">
<emphasis id="B972EAFD527CFF9AB783FD88FDC9FDC5" box="[395,540,596,622]" italics="true" pageId="4" pageNumber="5">H. anaksaku</emphasis>
</taxonomicName>
and
<taxonomicName id="4C064D6C527CFF9AB45CFD88FD1FFDC5" box="[596,714,597,622]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="minyas">
<emphasis id="B972EAFD527CFF9AB45CFD88FD1FFDC5" box="[596,714,597,622]" italics="true" pageId="4" pageNumber="5">H. minyas</emphasis>
</taxonomicName>
by the more strongly S-shaped gonopods. It is furthermore separated from
<taxonomicName id="4C064D6C527CFF9AB77AFDA4FE23FD39" authority="Johanson &amp; Pham, 2012" authorityName="Johanson &amp; Pham" authorityYear="2012" box="[370,502,632,658]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="lamnata" status="sp. nov.">
<emphasis id="B972EAFD527CFF9AB77AFDA4FE23FD39" box="[370,502,632,658]" italics="true" pageId="4" pageNumber="5">H. lamnata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2415786527CFF9AB7F5FDA5FD83FD39" box="[509,598,632,658]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
by the absence of a central posterior process of segment IX.
</paragraph>
</subSubSection>
<subSubSection id="C31C6564527CFF9AB6B5FD67FBA3FCAF" box="[189,1142,698,772]" pageId="4" pageNumber="5" type="etymology">
<paragraph id="8BB936EF527CFF9AB6B5FD67FE91FD7F" blockId="4.[189,1142,698,772]" box="[189,324,698,724]" pageId="4" pageNumber="5">
<heading id="D0F18183527CFF9AB6B5FD67FE91FD7F" bold="true" box="[189,324,698,724]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="B972EAFD527CFF9AB6B5FD67FE91FD7F" bold="true" box="[189,324,698,724]" pageId="4" pageNumber="5">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BB936EF527CFF9AB6B5FD34FBA3FCAF" blockId="4.[189,1142,698,772]" box="[189,1142,745,772]" pageId="4" pageNumber="5">
<emphasis id="B972EAFD527CFF9AB6B5FD34FEF2FCA8" box="[189,295,745,771]" italics="true" pageId="4" pageNumber="5">Meander</emphasis>
, named after the shape of the gonopods, resembling a meandering river.
</paragraph>
</subSubSection>
<subSubSection id="C31C6564527CFF9AB6B5FCF6FC3CFC4B" pageId="4" pageNumber="5" type="materials_examined">
<paragraph id="8BB936EF527CFF9AB6B5FCF6FE7EFCEE" blockId="4.[189,427,811,837]" box="[189,427,811,837]" pageId="4" pageNumber="5">
<heading id="D0F18183527CFF9AB6B5FCF6FE7EFCEE" bold="true" box="[189,427,811,837]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="B972EAFD527CFF9AB6B5FCF6FE7EFCEE" bold="true" box="[189,427,811,837]" pageId="4" pageNumber="5">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B6E3CB2527CFF9AB6E2FC86FC3CFC4B" ID-GBIF-Occurrence="2612312303" collectingDate="2011-04-04" collectingDateMax="2011-04-12" collectingDateMin="2011-04-04" collectionCode="NHRS" collectorName="K. A. Johanson &amp; P. H. Thai" country="Vietnam" latitude="21.391167" location="Me Linh District" longLatPrecision="1" longitude="105.71297" municipality="Prov" pageId="4" pageNumber="5" specimenCount="1" specimenCount-male="1" stateProvince="Vinh Phuc" typeStatus="holotype">
<paragraph id="8BB936EF527CFF9AB6E2FC86FEAFFCDE" blockId="4.[189,1399,857,993]" box="[234,378,857,885]" pageId="4" pageNumber="5">
<emphasis id="B972EAFD527CFF9AB6E2FC86FEAFFCDE" bold="true" box="[234,378,857,885]" pageId="4" pageNumber="5">
<typeStatus id="54BD884D527CFF9AB6E2FC86FE89FCDE" box="[234,348,859,885]" pageId="4" pageNumber="5" type="holotype">Holotype</typeStatus>
<specimenCount id="9D00FD66527CFF9AB76CFC84FEAFFCDE" box="[356,378,857,885]" pageId="4" pageNumber="5" type="male">♂</specimenCount>
</emphasis>
</paragraph>
<paragraph id="8BB936EF527CFF9AB6B5FCA3FC3CFC4B" blockId="4.[189,1399,857,993]" pageId="4" pageNumber="5">
<collectingCountry id="F311767F527CFF9AB6B5FCA3FE91FC33" box="[189,324,894,920]" name="Vietnam" pageId="4" pageNumber="5">VIETNAM</collectingCountry>
:
<collectingRegion id="49C2F80D527CFF9AB75EFCA3FE01FC33" box="[342,468,894,920]" country="Vietnam" name="Vinh Phuc" pageId="4" pageNumber="5">Vinh Phuc</collectingRegion>
<collectingMunicipality id="6BDDAC95527CFF9AB7D6FCA2FDC2FC32" box="[478,535,895,921]" pageId="4" pageNumber="5">Prov</collectingMunicipality>
,
<location id="8ED96034527CFF9AB42CFCA2FD27FC33" LSID="urn:lsid:plazi:treatment:03AF87F9527CFF9BB468FED6FB2DFE19:8ED96034527CFF9AB42CFCA2FD27FC33" box="[548,754,894,921]" country="Vietnam" latitude="21.391167" longLatPrecision="1" longitude="105.71297" municipality="Prov" name="Me Linh District" pageId="4" pageNumber="5" stateProvince="Vinh Phuc">Me Linh District</location>
, stream near
<location id="8ED96034527CFF9AB59DFCA2FB5FFC33" LSID="urn:lsid:plazi:treatment:03AF87F9527CFF9BB468FED6FB2DFE19:8ED96034527CFF9AB59DFCA2FB5FFC33" box="[917,1162,894,921]" country="Vietnam" latitude="21.391167" longLatPrecision="1" longitude="105.71297" municipality="Prov" name="Ngoc Thanh Village" pageId="4" pageNumber="5" stateProvince="Vinh Phuc">Ngoc Thanh Village</location>
,
<location id="8ED96034527CFF9AB290FCA3FE9AFC17" LSID="urn:lsid:plazi:treatment:03AF87F9527CFF9BB468FED6FB2DFE19:8ED96034527CFF9AB290FCA3FE9AFC17" country="Vietnam" latitude="21.391167" longLatPrecision="1" longitude="105.71297" municipality="Prov" name="Melinh Station for Biodiversity" pageId="4" pageNumber="5" stateProvince="Vinh Phuc">Melinh Station for Biodiversity</location>
,
<geoCoordinate id="EE325028527CFF9AB753FC7FFE2BFC17" box="[347,510,930,956]" degrees="21" direction="north" minutes="23" orientation="latitude" pageId="4" pageNumber="5" precision="1" seconds="28.2" value="21.391167">21°23'28.2&quot;N</geoCoordinate>
,
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,
<quantity id="4CFE9B0A527CFF9AB4CCFC7FFCC5FC16" box="[708,784,930,957]" metricMagnitude="2" metricUnit="m" metricValue="2.33" pageId="4" pageNumber="5" unit="m" value="233.0">233 m</quantity>
, Malaise trap
<date id="FFB8102F527CFF9AB5BDFC7FFBB2FC17" box="[949,1127,930,956]" pageId="4" pageNumber="5" value="2011-04-04" valueMax="2011-04-12" valueMin="2011-04-04">
<collectingDate id="EFFCE9C7527CFF9AB5BDFC7FFBB2FC17" box="[949,1127,930,956]" pageId="4" pageNumber="5" value="2011-04-04" valueMax="2011-04-12" valueMin="2011-04-04">4-12 Apr. 2011</collectingDate>
</date>
, loc#VN001, leg.
<collectorName id="26F35339527CFF9AB335FC7FFEF8FC4B" pageId="4" pageNumber="5">K.A. Johanson</collectorName>
,
<collectorName id="26F35339527CFF9AB731FC1AFE72FC4B" box="[313,423,966,993]" pageId="4" pageNumber="5">P.H. Thai</collectorName>
&amp; T.T. Du (
<collectionCode id="ED17AE2A527CFF9AB43CFC1BFD52FC4B" box="[564,647,966,992]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="4" pageNumber="5">NHRS</collectionCode>
, alcohol) [DNA voucher IP7].
</paragraph>
</materialsCitation>
</subSubSection>
<subSubSection id="C31C6564527CFF9BB6B5FBD5FB2DFE19" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="description">
<paragraph id="8BB936EF527CFF9AB6B5FBD5FE9BFB89" blockId="4.[189,334,1032,1058]" box="[189,334,1032,1058]" pageId="4" pageNumber="5">
<heading id="D0F18183527CFF9AB6B5FBD5FE9BFB89" bold="true" box="[189,334,1032,1058]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="B972EAFD527CFF9AB6B5FBD5FE9BFB89" bold="true" box="[189,334,1032,1058]" pageId="4" pageNumber="5">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BB936EF527CFF9AB6B5FBE5FACAFB31" blockId="4.[189,1398,1079,1178]" pageId="4" pageNumber="5">MAXILLARY PALPS. Two-segmented, with distal segment slightly longer than basal segment. Pair of interantennal setal warts semi-spherical, elevated. Each scapus about as long as each basal segment of maxillary palps. Spur formula 1,2,4; each foreleg spur slightly shorter than each first tarsomere.</paragraph>
<paragraph id="8BB936EF527CFF9AB6B5FB1FFCE5FB70" blockId="4.[189,816,1217,1244]" box="[189,816,1217,1244]" pageId="4" pageNumber="5">
LENTH. Forewing
<quantity id="4CFE9B0A527CFF9AB784FB1CFE3CFB77" box="[396,489,1217,1244]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="4" pageNumber="5" unit="mm" value="3.7">3.7 mm</quantity>
, hind wing
<quantity id="4CFE9B0A527CFF9AB47AFB1CFD19FB77" box="[626,716,1217,1244]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="4" pageNumber="5" unit="mm" value="2.8">2.8 mm</quantity>
(N = 1).
</paragraph>
<paragraph id="8BB936EF527CFF9BB6B5FAD9FB2DFE19" blockId="4.[189,1399,1283,2027]" lastBlockId="5.[189,1399,267,435]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
MALE ABDOMEN AND GENITALIA. (
<figureCitation id="133D2A6A527CFF9AB436FADEFD5CFAB6" box="[574,649,1283,1309]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[213,1376,611,1844]" captionTargetId="figure@5.[189,1398,598,1890]" captionTargetPageId="5" captionText="Fig. 2. Helicopsyche meander sp. nov., ♂ holotype. A. abdominal sternum VI, lateral view. B. abdominal sternal process VI, ventral view. C. genitalia, lateral view. D. genitalia, dorsal view. E. genitalia, ventral view. F. phallus, lateral view. G. phallus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3822700" httpUri="https://zenodo.org/record/3822700/files/figure.png" pageId="4" pageNumber="5">Fig. 2</figureCitation>
) Sternal process VI about one-eighth as long as sternum VI, straight, oriented posteroventrad; slightly tapering in lateral view (
<figureCitation id="133D2A6A527CFF9AB5EAFAFAFB91FAEA" box="[994,1092,1319,1345]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[213,1376,611,1844]" captionTargetId="figure@5.[189,1398,598,1890]" captionTargetPageId="5" captionText="Fig. 2. Helicopsyche meander sp. nov., ♂ holotype. A. abdominal sternum VI, lateral view. B. abdominal sternal process VI, ventral view. C. genitalia, lateral view. D. genitalia, dorsal view. E. genitalia, ventral view. F. phallus, lateral view. G. phallus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3822700" httpUri="https://zenodo.org/record/3822700/files/figure.png" pageId="4" pageNumber="5">Fig. 2A</figureCitation>
); pointed in ventral view (
<figureCitation id="133D2A6A527CFF9AB6CEFA96FEF3FACE" box="[198,294,1355,1381]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[213,1376,611,1844]" captionTargetId="figure@5.[189,1398,598,1890]" captionTargetPageId="5" captionText="Fig. 2. Helicopsyche meander sp. nov., ♂ holotype. A. abdominal sternum VI, lateral view. B. abdominal sternal process VI, ventral view. C. genitalia, lateral view. D. genitalia, dorsal view. E. genitalia, ventral view. F. phallus, lateral view. G. phallus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3822700" httpUri="https://zenodo.org/record/3822700/files/figure.png" pageId="4" pageNumber="5">Fig. 2B</figureCitation>
). Segment IX about 1.5 x taller than long in lateral view (
<figureCitation id="133D2A6A527CFF9AB5F5FA96FB8BFACE" box="[1021,1118,1355,1381]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[213,1376,611,1844]" captionTargetId="figure@5.[189,1398,598,1890]" captionTargetPageId="5" captionText="Fig. 2. Helicopsyche meander sp. nov., ♂ holotype. A. abdominal sternum VI, lateral view. B. abdominal sternal process VI, ventral view. C. genitalia, lateral view. D. genitalia, dorsal view. E. genitalia, ventral view. F. phallus, lateral view. G. phallus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3822700" httpUri="https://zenodo.org/record/3822700/files/figure.png" pageId="4" pageNumber="5">Fig. 2C</figureCitation>
); each anterior lobe of segment IX (
<figureCitation id="133D2A6A527CFF9AB750FAB2FE61FA22" box="[344,436,1391,1417]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[213,1376,611,1844]" captionTargetId="figure@5.[189,1398,598,1890]" captionTargetPageId="5" captionText="Fig. 2. Helicopsyche meander sp. nov., ♂ holotype. A. abdominal sternum VI, lateral view. B. abdominal sternal process VI, ventral view. C. genitalia, lateral view. D. genitalia, dorsal view. E. genitalia, ventral view. F. phallus, lateral view. G. phallus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3822700" httpUri="https://zenodo.org/record/3822700/files/figure.png" pageId="4" pageNumber="5">Fig. 2C</figureCitation>
) irregularly narrowing anteriorly in lateral view, asymmetrical, oriented anterad; anterodorsal margin almost straight; anteroventral margin concave (
<figureCitation id="133D2A6A527CFF9AB5F4FA4EFB8BFA06" box="[1020,1118,1427,1453]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[213,1376,611,1844]" captionTargetId="figure@5.[189,1398,598,1890]" captionTargetPageId="5" captionText="Fig. 2. Helicopsyche meander sp. nov., ♂ holotype. A. abdominal sternum VI, lateral view. B. abdominal sternal process VI, ventral view. C. genitalia, lateral view. D. genitalia, dorsal view. E. genitalia, ventral view. F. phallus, lateral view. G. phallus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3822700" httpUri="https://zenodo.org/record/3822700/files/figure.png" pageId="4" pageNumber="5">Fig. 2C</figureCitation>
); in dorsal view, inner margin forming narrowly ellipsoid cavity (
<figureCitation id="133D2A6A527CFF9AB4B2FA6AFCCFFA7A" box="[698,794,1463,1489]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[213,1376,611,1844]" captionTargetId="figure@5.[189,1398,598,1890]" captionTargetPageId="5" captionText="Fig. 2. Helicopsyche meander sp. nov., ♂ holotype. A. abdominal sternum VI, lateral view. B. abdominal sternal process VI, ventral view. C. genitalia, lateral view. D. genitalia, dorsal view. E. genitalia, ventral view. F. phallus, lateral view. G. phallus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3822700" httpUri="https://zenodo.org/record/3822700/files/figure.png" pageId="4" pageNumber="5">Fig. 2D</figureCitation>
); in ventral view, without central posterior process (
<figureCitation id="133D2A6A527CFF9AB6CEFA07FEF0FA5E" box="[198,293,1498,1525]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[213,1376,611,1844]" captionTargetId="figure@5.[189,1398,598,1890]" captionTargetPageId="5" captionText="Fig. 2. Helicopsyche meander sp. nov., ♂ holotype. A. abdominal sternum VI, lateral view. B. abdominal sternal process VI, ventral view. C. genitalia, lateral view. D. genitalia, dorsal view. E. genitalia, ventral view. F. phallus, lateral view. G. phallus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3822700" httpUri="https://zenodo.org/record/3822700/files/figure.png" pageId="4" pageNumber="5">Fig. 2E</figureCitation>
); inner margin forming wide, almost rectangular cavity; lateral apodemes nearly horizontal (
<figureCitation id="133D2A6A527CFF9AB6CEFA23FEF3F9B3" box="[198,294,1534,1560]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[213,1376,611,1844]" captionTargetId="figure@5.[189,1398,598,1890]" captionTargetPageId="5" captionText="Fig. 2. Helicopsyche meander sp. nov., ♂ holotype. A. abdominal sternum VI, lateral view. B. abdominal sternal process VI, ventral view. C. genitalia, lateral view. D. genitalia, dorsal view. E. genitalia, ventral view. F. phallus, lateral view. G. phallus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3822700" httpUri="https://zenodo.org/record/3822700/files/figure.png" pageId="4" pageNumber="5">Fig. 2C</figureCitation>
), each parallel-sided along their length, meeting anterior margin; sub-marginal line present above lateral apodemes; tergal transverse apodemes present, fading immediately above superior appendages; sternal transverse apodeme present along posterior margin of segment IX (
<figureCitation id="133D2A6A527CFF9AB30EF99BFABCF9CB" box="[1286,1385,1606,1632]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[213,1376,611,1844]" captionTargetId="figure@5.[189,1398,598,1890]" captionTargetPageId="5" captionText="Fig. 2. Helicopsyche meander sp. nov., ♂ holotype. A. abdominal sternum VI, lateral view. B. abdominal sternal process VI, ventral view. C. genitalia, lateral view. D. genitalia, dorsal view. E. genitalia, ventral view. F. phallus, lateral view. G. phallus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3822700" httpUri="https://zenodo.org/record/3822700/files/figure.png" pageId="4" pageNumber="5">Fig. 2C</figureCitation>
). Transverse plate absent. Tergum X, in lateral view, oriented posteroventrad (
<figureCitation id="133D2A6A527CFF9AB237F9B7FB49F92F" box="[1087,1180,1642,1668]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[213,1376,611,1844]" captionTargetId="figure@5.[189,1398,598,1890]" captionTargetPageId="5" captionText="Fig. 2. Helicopsyche meander sp. nov., ♂ holotype. A. abdominal sternum VI, lateral view. B. abdominal sternal process VI, ventral view. C. genitalia, lateral view. D. genitalia, dorsal view. E. genitalia, ventral view. F. phallus, lateral view. G. phallus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3822700" httpUri="https://zenodo.org/record/3822700/files/figure.png" pageId="4" pageNumber="5">Fig. 2C</figureCitation>
), slightly sigmoid, tapering towards mid-length, apex strongly club-shaped, slightly curving dorsally (
<figureCitation id="133D2A6A527CFF9AB291F953FB2DF903" box="[1177,1272,1678,1704]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[213,1376,611,1844]" captionTargetId="figure@5.[189,1398,598,1890]" captionTargetPageId="5" captionText="Fig. 2. Helicopsyche meander sp. nov., ♂ holotype. A. abdominal sternum VI, lateral view. B. abdominal sternal process VI, ventral view. C. genitalia, lateral view. D. genitalia, dorsal view. E. genitalia, ventral view. F. phallus, lateral view. G. phallus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3822700" httpUri="https://zenodo.org/record/3822700/files/figure.png" pageId="4" pageNumber="5">Fig. 2C</figureCitation>
); in dorsal view (
<figureCitation id="133D2A6A527CFF9AB70DF96FFEB4F967" box="[261,353,1714,1740]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[213,1376,611,1844]" captionTargetId="figure@5.[189,1398,598,1890]" captionTargetPageId="5" captionText="Fig. 2. Helicopsyche meander sp. nov., ♂ holotype. A. abdominal sternum VI, lateral view. B. abdominal sternal process VI, ventral view. C. genitalia, lateral view. D. genitalia, dorsal view. E. genitalia, ventral view. F. phallus, lateral view. G. phallus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3822700" httpUri="https://zenodo.org/record/3822700/files/figure.png" pageId="4" pageNumber="5">Fig. 2D</figureCitation>
), deeply and widely divided into pair of parallel-sided and parallel-running branches, with 3 pairs of equally long megasetae in apical group, starting well before apex of gonopods in lateral view (
<figureCitation id="133D2A6A527CFF9AB6CEF924FEF1F8BF" box="[198,292,1785,1812]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[213,1376,611,1844]" captionTargetId="figure@5.[189,1398,598,1890]" captionTargetPageId="5" captionText="Fig. 2. Helicopsyche meander sp. nov., ♂ holotype. A. abdominal sternum VI, lateral view. B. abdominal sternal process VI, ventral view. C. genitalia, lateral view. D. genitalia, dorsal view. E. genitalia, ventral view. F. phallus, lateral view. G. phallus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3822700" httpUri="https://zenodo.org/record/3822700/files/figure.png" pageId="4" pageNumber="5">Fig. 2C</figureCitation>
). Superior appendages club-shaped (
<figureCitation id="133D2A6A527CFF9AB4D0F924FCE3F8BF" box="[728,822,1785,1812]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[213,1376,611,1844]" captionTargetId="figure@5.[189,1398,598,1890]" captionTargetPageId="5" captionText="Fig. 2. Helicopsyche meander sp. nov., ♂ holotype. A. abdominal sternum VI, lateral view. B. abdominal sternal process VI, ventral view. C. genitalia, lateral view. D. genitalia, dorsal view. E. genitalia, ventral view. F. phallus, lateral view. G. phallus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3822700" httpUri="https://zenodo.org/record/3822700/files/figure.png" pageId="4" pageNumber="5">Fig. 2C</figureCitation>
), oriented ventrad. Primary branch of gonopods, in lateral view (
<figureCitation id="133D2A6A527CFF9AB788F8C0FE34F89C" box="[384,481,1821,1847]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[213,1376,611,1844]" captionTargetId="figure@5.[189,1398,598,1890]" captionTargetPageId="5" captionText="Fig. 2. Helicopsyche meander sp. nov., ♂ holotype. A. abdominal sternum VI, lateral view. B. abdominal sternal process VI, ventral view. C. genitalia, lateral view. D. genitalia, dorsal view. E. genitalia, ventral view. F. phallus, lateral view. G. phallus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3822700" httpUri="https://zenodo.org/record/3822700/files/figure.png" pageId="4" pageNumber="5">Fig. 2C</figureCitation>
), with basal part oriented posterad before strongly bending dorsad, angling posterad at half-length, apical part produced mesad above tergum X; almost parallel-sided along their length; anterior margin smooth, strongly concave; dorsal and posterior margins undulating. Secondary branch of gonopods absent. Basomesal lobes long, tuboid, each with pair of apical setae; sigmoid in lateral view (
<figureCitation id="133D2A6A527CFF9AB757F870FE15F86C" box="[351,448,1965,1991]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[213,1376,611,1844]" captionTargetId="figure@5.[189,1398,598,1890]" captionTargetPageId="5" captionText="Fig. 2. Helicopsyche meander sp. nov., ♂ holotype. A. abdominal sternum VI, lateral view. B. abdominal sternal process VI, ventral view. C. genitalia, lateral view. D. genitalia, dorsal view. E. genitalia, ventral view. F. phallus, lateral view. G. phallus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3822700" httpUri="https://zenodo.org/record/3822700/files/figure.png" pageId="4" pageNumber="5">Fig. 2D</figureCitation>
), parallel-sided with diverging apex in ventral view (
<figureCitation id="133D2A6A527CFF9AB247F870FB7AF86C" box="[1103,1199,1965,1991]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[213,1376,611,1844]" captionTargetId="figure@5.[189,1398,598,1890]" captionTargetPageId="5" captionText="Fig. 2. Helicopsyche meander sp. nov., ♂ holotype. A. abdominal sternum VI, lateral view. B. abdominal sternal process VI, ventral view. C. genitalia, lateral view. D. genitalia, dorsal view. E. genitalia, ventral view. F. phallus, lateral view. G. phallus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3822700" httpUri="https://zenodo.org/record/3822700/files/figure.png" pageId="4" pageNumber="5">Fig. 2E</figureCitation>
). Basal plate, in lateral view (
<figureCitation id="133D2A6A527CFF9AB757F80CFE15F840" box="[351,448,2001,2027]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[213,1376,611,1844]" captionTargetId="figure@5.[189,1398,598,1890]" captionTargetPageId="5" captionText="Fig. 2. Helicopsyche meander sp. nov., ♂ holotype. A. abdominal sternum VI, lateral view. B. abdominal sternal process VI, ventral view. C. genitalia, lateral view. D. genitalia, dorsal view. E. genitalia, ventral view. F. phallus, lateral view. G. phallus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3822700" httpUri="https://zenodo.org/record/3822700/files/figure.png" pageId="4" pageNumber="5">Fig. 2C</figureCitation>
), curving anteroventrad, narrowing along its length from posterior to anterior end, slightly produced beyond anteroventral margin of segment IX in lateral view (
<figureCitation id="133D2A6A527DFF9BB2A6FED6FADBFE8E" box="[1198,1294,267,293]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[213,1376,611,1844]" captionTargetId="figure@5.[189,1398,598,1890]" captionTargetPageId="5" captionText="Fig. 2. Helicopsyche meander sp. nov., ♂ holotype. A. abdominal sternum VI, lateral view. B. abdominal sternal process VI, ventral view. C. genitalia, lateral view. D. genitalia, dorsal view. E. genitalia, ventral view. F. phallus, lateral view. G. phallus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3822700" httpUri="https://zenodo.org/record/3822700/files/figure.png" pageId="5" pageNumber="6">Fig. 2C</figureCitation>
); widely triangular in ventral view (
<figureCitation id="133D2A6A527DFF9BB40AFEF3FDB5FEE2" box="[514,608,302,329]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[213,1376,611,1844]" captionTargetId="figure@5.[189,1398,598,1890]" captionTargetPageId="5" captionText="Fig. 2. Helicopsyche meander sp. nov., ♂ holotype. A. abdominal sternum VI, lateral view. B. abdominal sternal process VI, ventral view. C. genitalia, lateral view. D. genitalia, dorsal view. E. genitalia, ventral view. F. phallus, lateral view. G. phallus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3822700" httpUri="https://zenodo.org/record/3822700/files/figure.png" pageId="5" pageNumber="6">Fig. 2E</figureCitation>
). Phallic apparatus, lateral view, irregularly tube-shaped along its length (
<figureCitation id="133D2A6A527DFF9BB711FE8FFEA3FEC7" box="[281,374,338,364]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[213,1376,611,1844]" captionTargetId="figure@5.[189,1398,598,1890]" captionTargetPageId="5" captionText="Fig. 2. Helicopsyche meander sp. nov., ♂ holotype. A. abdominal sternum VI, lateral view. B. abdominal sternal process VI, ventral view. C. genitalia, lateral view. D. genitalia, dorsal view. E. genitalia, ventral view. F. phallus, lateral view. G. phallus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3822700" httpUri="https://zenodo.org/record/3822700/files/figure.png" pageId="5" pageNumber="6">Fig. 2F</figureCitation>
), apex about as broad as base; nearly straight after mid-length; in ventral view, with basis narrow (
<figureCitation id="133D2A6A527DFF9BB76FFEA8FE1CFE24" box="[359,457,373,399]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[213,1376,611,1844]" captionTargetId="figure@5.[189,1398,598,1890]" captionTargetPageId="5" captionText="Fig. 2. Helicopsyche meander sp. nov., ♂ holotype. A. abdominal sternum VI, lateral view. B. abdominal sternal process VI, ventral view. C. genitalia, lateral view. D. genitalia, dorsal view. E. genitalia, ventral view. F. phallus, lateral view. G. phallus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3822700" httpUri="https://zenodo.org/record/3822700/files/figure.png" pageId="5" pageNumber="6">Fig. 2G</figureCitation>
); widest after mid-length; endotheca weakly produced, posteroventral part not sclerotized, except single, weakly sclerotized triangular processes distally of phallic sclerite.
</paragraph>
</subSubSection>
</treatment>
</document>
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