treatments-xml/data/03/AF/87/03AF87E48E4DFFDBFF49FA98FE448287.xml
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<document id="587EFDFB29F2FF3BDDC0E36E443865E1" ID-CLB-Dataset="29479" ID-DOI="10.11646/zootaxa.4471.2.3" ID-GBIF-Dataset="694f85ad-7913-43ea-89b9-8eb4e20f0821" ID-ISSN="1175-5326" ID-Zenodo-Dep="1439674" ID-ZooBank="CE4C6E73-09CB-4E74-91C5-771B3696DC52" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538364447371" checkinUser="plazi" docAuthor="Smith, Andrew B. T. &amp; Evans, Arthur V." docDate="2018" docId="03AF87E48E4DFFDBFF49FA98FE448287" docLanguage="en" docName="zootaxa.4471.2.3.pdf" docOrigin="Zootaxa 4471 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Apteroathlia Smith &amp; Evans 2018, new genus" docType="treatment" docUuid="778D2B0C-E445-46BC-84CB-338DA4F2F7F6" docUuidSource="ZooBank" docVersion="8" lastPageNumber="282" masterDocId="FF96FF9C8E4FFFD8FFDEFFC7FFE38255" masterDocTitle="Taxonomic review of Athliini (Coleoptera: Scarabaeidae: Melolonthinae), a new tribe of scarab beetles endemic to South America" masterLastPageNumber="308" masterPageNumber="279" pageNumber="281" updateTime="1698558211898" updateUser="ExternalLinkService">
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<mods:title id="0BA6F28F3FC7B057EBA49BF29BAAEC53">Taxonomic review of Athliini (Coleoptera: Scarabaeidae: Melolonthinae), a new tribe of scarab beetles endemic to South America</mods:title>
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<mods:namePart id="16DB173D273E096C5445087168C49FE3">Smith, Andrew B. T.</mods:namePart>
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<mods:namePart id="97BB2FF346BC5DC9FBEAFD5F30D4CBEB">Evans, Arthur V.</mods:namePart>
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<mods:date id="2FD8B8913182CA4A078A4B48A3E2BD50">2018</mods:date>
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<treatment id="03AF87E48E4DFFDBFF49FA98FE448287" ID-DOI="http://doi.org/10.5281/zenodo.5958484" ID-GBIF-Taxon="148415673" ID-Zenodo-Dep="5958484" LSID="urn:lsid:zoobank.org:act:778D2B0C-E445-46BC-84CB-338DA4F2F7F6" httpUri="http://treatment.plazi.org/id/03AF87E48E4DFFDBFF49FA98FE448287" lastPageId="3" lastPageNumber="282" pageId="2" pageNumber="281">
<subSubSection id="C31C65798E4DFFDAFF49FA98FB4287CE" pageId="2" pageNumber="281" type="nomenclature">
<paragraph id="8BB936F28E4DFFDAFF49FA98FD67872F" blockId="2.[151,1185,1375,1435]" box="[151,644,1375,1402]" pageId="2" pageNumber="281">
<heading id="D0F1819E8E4DFFDAFF49FA98FD67872F" bold="true" box="[151,644,1375,1402]" fontSize="11" level="1" pageId="2" pageNumber="281" reason="1">
<emphasis id="B972EAE08E4DFFDAFF49FA98FD67872F" bold="true" box="[151,644,1375,1402]" pageId="2" pageNumber="281">
<taxonomicName id="4C064D718E4DFFDAFF49FA98FE16872F" ID-CoL="ZCG" authority="Smith &amp; Evans" authorityName="Smith &amp; Evans" authorityYear="2018" box="[151,501,1375,1402]" class="Insecta" family="Scarabaeidae" genus="Apteroathlia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="281" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B972EAE08E4DFFDAFF49FA98FECC872C" bold="true" box="[151,303,1375,1401]" italics="true" pageId="2" pageNumber="281">Apteroathlia</emphasis>
Smith &amp; Evans
</taxonomicName>
,
<taxonomicNameLabel id="A241579B8E4DFFDAFDDDFAA7FD67872F" box="[515,644,1376,1402]" pageId="2" pageNumber="281" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
</heading>
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<paragraph id="8BB936F28E4DFFDAFF49FA45FB4287CE" blockId="2.[151,1185,1375,1435]" box="[151,1185,1410,1435]" pageId="2" pageNumber="281">http://zoobank.org/urn:lsid:zoobank.org:act:778D2B0C-E445-46BC-84CB-338DA4F2F7F6</paragraph>
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<subSubSection id="C31C65798E4DFFDAFF49FA02FC1D878E" box="[151,1022,1477,1499]" pageId="2" pageNumber="281" type="materials_examined">
<paragraph id="8BB936F28E4DFFDAFF49FA02FC1D878E" blockId="2.[151,1022,1477,1499]" box="[151,1022,1477,1499]" pageId="2" pageNumber="281">
<emphasis id="B972EAE08E4DFFDAFF49FA02FEC1878E" bold="true" box="[151,290,1477,1499]" pageId="2" pageNumber="281">Type species.</emphasis>
<taxonomicName id="4C064D718E4DFFDAFEF7FA02FD26878E" authority="Smith &amp; Evans" authorityName="Smith &amp; Evans" authorityYear="2018" box="[297,709,1477,1499]" class="Insecta" family="Scarabaeidae" genus="Apteroathlia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="281" phylum="Arthropoda" rank="species" species="translucida" status="sp. nov.">
<emphasis id="B972EAE08E4DFFDAFEF7FA02FDC1878E" box="[297,546,1477,1499]" italics="true" pageId="2" pageNumber="281">Apteroathlia translucida</emphasis>
Smith &amp; Evans
</taxonomicName>
,
<taxonomicNameLabel id="A241579B8E4DFFDAFD0EFA02FCAF878E" box="[720,844,1477,1499]" pageId="2" pageNumber="281" rank="species">
<emphasis id="B972EAE08E4DFFDAFD0EFA02FCAF878E" bold="true" box="[720,844,1477,1499]" pageId="2" pageNumber="281">new species</emphasis>
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, here designated.
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<subSubSection id="C31C65798E4DFFDAFF49F9C5FA9885DF" pageId="2" pageNumber="281" type="description">
<paragraph id="8BB936F28E4DFFDAFF49F9C5FA9885DF" blockId="2.[151,1437,1538,2030]" pageId="2" pageNumber="281">
<emphasis id="B972EAE08E4DFFDAFF49F9C5FEC2844E" bold="true" box="[151,289,1538,1563]" pageId="2" pageNumber="281">Description</emphasis>
. Length
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. Ovate, widest behind middle of elytra; testaceous to black, or black with testaceous appendages and partly or entirely testaceous pronotum; surfaces moderately to strongly shiny, and mostly glabrous. Head: labrum wide, broadly emarginate, evenly and shallowly reflexed anteriorly, completely fused to and slightly below plane of clypeus exposing anterior clypeal margin; labroclypeal suture distinct and terminating in small, marginal notches. Mentum longer than wide, narrowly emarginate in front, with sides gradually converging posteriorly. Antennae with 8 antennomeres, club with 3 short, thick lamellae. Pronotum: convex, wider than long, widest medially. Elytra: convex and without costae or distinct interval, punctostriate. Metathoracic wings absent. Abdomen: with lateral carinae on ventrites. Legs: protibia weakly tridentate on outer margin, spur present and reaching to basal third or half of protarsomere 1. Claws symmetrical and toothed with a short, bisetose onychium in between. Metatibial apices with spurs set in notch, each set on either side of tarsal articulation, longest spur reaching about two-thirds to three-quarters the length of first tarsomere. Male genitalia: symmetrical parameres simple in form, not split or with strong sinuations, and subequal in length to phallobase.
</paragraph>
</subSubSection>
<subSubSection id="C31C65798E4DFFDAFF19F855FB7885B8" pageId="2" pageNumber="281" type="etymology">
<paragraph id="8BB936F28E4DFFDAFF19F855FDDC8599" blockId="2.[151,1437,1538,2030]" pageId="2" pageNumber="281">
<emphasis id="B972EAE08E4DFFDAFF19F855FEAD85FE" bold="true" box="[199,334,1938,1963]" pageId="2" pageNumber="281">Etymology.</emphasis>
<taxonomicName id="4C064D718E4DFFDAFE8AF854FE0285FE" authority="Smith &amp; Evans, 2018" authorityName="Smith &amp; Evans" authorityYear="2018" box="[340,481,1939,1963]" class="Insecta" family="Scarabaeidae" genus="Apteroathlia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="282" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B972EAE08E4DFFDAFE8AF854FE0285FE" box="[340,481,1939,1963]" italics="true" pageId="2" pageNumber="281">Apteroathlia</emphasis>
</taxonomicName>
means “wingless
<taxonomicName id="4C064D718E4DFFDAFD73F854FD1285FE" box="[685,753,1939,1963]" class="Insecta" family="Scarabaeidae" genus="Athlia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="281" phylum="Arthropoda" rank="genus">
<emphasis id="B972EAE08E4DFFDAFD73F854FD1285FE" box="[685,753,1939,1963]" italics="true" pageId="2" pageNumber="281">Athlia</emphasis>
</taxonomicName>
” to indicate the similarities with the genus
<taxonomicName id="4C064D718E4DFFDAFB04F854FAFD85FE" box="[1242,1310,1939,1963]" class="Insecta" family="Scarabaeidae" genus="Athlia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="281" phylum="Arthropoda" rank="genus">
<emphasis id="B972EAE08E4DFFDAFB04F854FAFD85FE" box="[1242,1310,1939,1963]" italics="true" pageId="2" pageNumber="281">Athlia</emphasis>
</taxonomicName>
. The name is feminine in gender to match
<taxonomicName id="4C064D718E4DFFDAFE2DF873FDD48599" box="[499,567,1972,1996]" class="Insecta" family="Scarabaeidae" genus="Athlia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="281" phylum="Arthropoda" rank="genus">
<emphasis id="B972EAE08E4DFFDAFE2DF873FDD48599" box="[499,567,1972,1996]" italics="true" pageId="2" pageNumber="281">Athlia</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BB936F28E4DFFDAFF19F812FB7885B8" blockId="2.[151,1437,1538,2030]" box="[199,1179,2005,2030]" pageId="2" pageNumber="281">
<emphasis id="B972EAE08E4DFFDAFF19F812FE8485BB" bold="true" box="[199,359,2005,2030]" pageId="2" pageNumber="281">Composition.</emphasis>
<taxonomicName id="4C064D718E4DFFDAFEB3F812FDC985B8" authority="Smith &amp; Evans, 2018" authorityName="Smith &amp; Evans" authorityYear="2018" box="[365,554,2005,2029]" class="Insecta" family="Scarabaeidae" genus="Apteroathlia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="281" phylum="Arthropoda" rank="species" species="nox" status="sp. nov.">
<emphasis id="B972EAE08E4DFFDAFEB3F812FDC985B8" box="[365,554,2005,2029]" italics="true" pageId="2" pageNumber="281">Apteroathlia nox</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A241579B8E4DFFDAFDEEF812FD5985BB" box="[560,698,2005,2030]" pageId="2" pageNumber="281" rank="species">
<emphasis id="B972EAE08E4DFFDAFDEEF812FD5985BB" bold="true" box="[560,698,2005,2030]" pageId="2" pageNumber="281">new species</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="4C064D718E4DFFDAFD2EF812FBE085B8" authority="Smith &amp; Evans, 2018" authorityName="Smith &amp; Evans" authorityYear="2018" box="[752,1027,2005,2029]" class="Insecta" family="Scarabaeidae" genus="Apteroathlia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="281" phylum="Arthropoda" rank="species" species="translucida" status="sp. nov.">
<emphasis id="B972EAE08E4DFFDAFD2EF812FBE085B8" box="[752,1027,2005,2029]" italics="true" pageId="2" pageNumber="281">Apteroathlia translucida</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A241579B8E4DFFDAFBD4F812FB7785BB" box="[1034,1172,2005,2030]" pageId="2" pageNumber="281" rank="species">
<emphasis id="B972EAE08E4DFFDAFBD4F812FB7785BB" bold="true" box="[1034,1172,2005,2030]" pageId="2" pageNumber="281">new species</emphasis>
</taxonomicNameLabel>
.
</paragraph>
</subSubSection>
<subSubSection id="C31C65798E4CFFDBFF19FF50FE448287" pageId="3" pageNumber="282" type="discussion">
<paragraph id="8BB936F28E4CFFDBFF19FF50FA7F82E5" blockId="3.[151,1436,151,210]" box="[199,1436,151,176]" pageId="3" pageNumber="282">
<emphasis id="B972EAE08E4CFFDBFF19FF50FED882E5" bold="true" box="[199,315,151,176]" pageId="3" pageNumber="282">Remarks.</emphasis>
The two known species in this genus are both endemic to the region around Río de la Plata in
</paragraph>
<paragraph id="8BB936F28E4CFFDBFF49FF7DFE448287" blockId="3.[151,1436,151,210]" box="[151,423,185,210]" pageId="3" pageNumber="282">Uruguay and Argentina.</paragraph>
</subSubSection>
</treatment>
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