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<document id="3BE5A2BD5E284440E51C6288ADF49441" ID-CLB-Dataset="8896" ID-DOI="10.11646/zootaxa.4868.1.4" ID-GBIF-Dataset="cfd5c06e-06f0-4f1f-ae4b-40b7b46f44a3" ID-ISSN="1175-5326" ID-Zenodo-Dep="4417281" ID-ZooBank="DC4B8EC5-CBDA-49D2-B370-381C96AF3A10" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609805123592" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2020" docId="03A887A1A96A1969AFE9FB0E430E6574" docLanguage="en" docName="zootaxa.4868.1.4.pdf" docOrigin="Zootaxa 4868 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hydrochus dualis Perkins 2020, new species" docType="treatment" docVersion="8" lastPageNumber="73" masterDocId="FF91FFD9A9611964AF7EFFC8470B6435" masterDocTitle="Taxonomy of Peruvian water beetles in the genus Hydrochus Leach, 1817 (Coleoptera: Hydrochidae)" masterLastPageNumber="89" masterPageNumber="61" pageNumber="72" updateTime="1698852338306" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="9FCA9C6E5B3A82D3904BE979BDD8E19B">Taxonomy of Peruvian water beetles in the genus Hydrochus Leach, 1817 (Coleoptera: Hydrochidae)</mods:title>
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<mods:namePart id="30A2BEEBB8EA5A50E6CB71B0BC02CA05">Perkins, Philip D.</mods:namePart>
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<mods:date id="3C35C9F398FF9E509512FE6E2D961638">2020</mods:date>
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<treatment id="03A887A1A96A1969AFE9FB0E430E6574" ID-DOI="http://doi.org/10.5281/zenodo.4417468" ID-GBIF-Taxon="176581837" ID-Zenodo-Dep="4417468" LSID="urn:lsid:plazi:treatment:03A887A1A96A1969AFE9FB0E430E6574" httpUri="http://treatment.plazi.org/id/03A887A1A96A1969AFE9FB0E430E6574" lastPageId="13" lastPageNumber="73" pageId="11" pageNumber="72">
<subSubSection id="C31B653CA96A196FAFE9FB0E450760D5" box="[151,524,1222,1249]" pageId="11" pageNumber="72" type="nomenclature">
<paragraph id="8BBE36B7A96A196FAFE9FB0E450760D5" blockId="11.[151,524,1222,1285]" box="[151,524,1222,1249]" pageId="11" pageNumber="72">
<heading id="D0F681DBA96A196FAFE9FB0E450760D5" bold="true" box="[151,524,1222,1249]" fontSize="11" level="1" pageId="11" pageNumber="72" reason="1">
<emphasis id="B975EAA5A96A196FAFE9FB0E450760D5" bold="true" box="[151,524,1222,1249]" pageId="11" pageNumber="72">
<taxonomicName id="4C014D34A96A196FAFE9FB0E466660D4" ID-CoL="8Y4Q3" authority="Perkins, 2020" authorityName="Perkins" authorityYear="2020" box="[151,365,1222,1249]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="72" phylum="Arthropoda" rank="species" species="dualis" status="sp. nov.">
<emphasis id="B975EAA5A96A196FAFE9FB0E466660D4" bold="true" box="[151,365,1222,1249]" italics="true" pageId="11" pageNumber="72">Hydrochus dualis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A24657DEA96A196FAE02FB0F450760D5" box="[380,524,1222,1249]" pageId="11" pageNumber="72" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C31B653CA96A196FAFE9FB2347F46130" box="[151,255,1259,1285]" pageId="11" pageNumber="72" type="description">
<paragraph id="8BBE36B7A96A196FAFE9FB2347F46130" blockId="11.[151,524,1222,1285]" box="[151,255,1259,1285]" pageId="11" pageNumber="72">
<figureCitation id="133A2A32A96A196FAFE9FB2347E86130" box="[151,227,1259,1285]" captionStart="FIGURE 1" captionStartId="2.[151,250,1971,1995]" captionTargetBox="[157,1430,309,1948]" captionTargetId="figure-37@2.[157,1430,309,1948]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal habitus of Hydrochus species." figureDoi="http://doi.org/10.5281/zenodo.4417283" httpUri="https://zenodo.org/record/4417283/files/figure.png" pageId="11" pageNumber="72">Figs. 1</figureCitation>
,
<figureCitation id="133A2A32A96A196FAF8FFB2347F46130" box="[241,255,1259,1285]" captionStart="FIGURE 8" captionStartId="12.[151,250,1861,1885]" captionTargetBox="[168,1427,534,1821]" captionTargetId="figure-166@12.[151,1435,510,1845]" captionTargetPageId="12" captionText="FIGURE 8. Hydrochus dualis n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.4417299" httpUri="https://zenodo.org/record/4417299/files/figure.png" pageId="11" pageNumber="72">8</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31B653CA96A196FAFE9FAFB45C86144" pageId="11" pageNumber="72" type="materials_examined">
<paragraph id="8BBE36B7A96A196FAFE9FAFB45C86144" blockId="11.[151,1437,1330,2041]" pageId="11" pageNumber="72">
<emphasis id="B975EAA5A96A196FAFE9FAFB464C6179" bold="true" box="[151,327,1330,1357]" pageId="11" pageNumber="72">Type Material.</emphasis>
<materialsCitation id="3B693CEAA96A196FAE33FAFB45C86144" ID-GBIF-Occurrence="3015416366" collectingDate="1937-12-11" collectingDateMax="1937-12-30" collectingDateMin="1937-12-11" collectionCode="SEMC" collectorName="F. Woytkowski No." country="Peru" elevation="800" location="Huanuco" pageId="11" pageNumber="72" specimenCount="800" stateProvince="Huanuco" typeStatus="holotype">
<emphasis id="B975EAA5A96A196FAE33FAFB451E6179" bold="true" box="[333,533,1330,1357]" pageId="11" pageNumber="72">
<typeStatus id="54BA8815A96A196FAE33FAFB46B16178" box="[333,442,1331,1357]" pageId="11" pageNumber="72" type="holotype">Holotype</typeStatus>
(male):
</emphasis>
<collectingCountry id="F3167627A96A196FAD58FAFB45546178" box="[550,607,1331,1357]" name="Peru" pageId="11" pageNumber="72">Peru</collectingCountry>
, S. A.
<date id="FFBF1077A96A196FADDBFAFB44666178" box="[677,877,1331,1357]" pageId="11" pageNumber="72" value="1937-12-11" valueMax="1937-12-30" valueMin="1937-12-11">
<collectingDate id="EFFBE99FA96A196FADDBFAFB44666178" box="[677,877,1331,1357]" pageId="11" pageNumber="72" value="1937-12-11" valueMax="1937-12-30" valueMin="1937-12-11">Dec. 1130, 1937</collectingDate>
</date>
,
<collectorName id="26F45361A96A196FAC09FAFB43466178" box="[887,1101,1331,1357]" pageId="11" pageNumber="72">F. Woytkowski No.</collectorName>
3811 // Department
<collectingRegion id="49C5F855A96A196FAA4EFAFB42936178" box="[1328,1432,1331,1357]" country="Peru" name="Huanuco" pageId="11" pageNumber="72">Huanuco</collectingRegion>
, Vic. Leonpampa, jungle,
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<quantity id="4CF99B52A96A196FAECCFA9F46F76144" box="[434,508,1367,1393]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="11" pageNumber="72" unit="m" value="800.0">
<elevation id="002CD184A96A196FAECCFA9F46F76144" box="[434,508,1367,1393]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="11" pageNumber="72" unit="m" value="800.0">800 m</elevation>
</quantity>
.
</specimenCount>
a. s. l.” (
<collectionCode id="ED10AE72A96A196FAD1AFA9F45BD6144" box="[612,694,1367,1393]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="11" pageNumber="72" type="Museum">SEMC</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31B653CA96A196FAFB9FAB243696234" pageId="11" pageNumber="72" type="diagnosis">
<paragraph id="8BBE36B7A96A196FAFB9FAB243696234" blockId="11.[151,1437,1330,2041]" pageId="11" pageNumber="72">
<emphasis id="B975EAA5A96A196FAFB9FAB246D861A1" bold="true" box="[199,467,1402,1428]" pageId="11" pageNumber="72">Differential Diagnosis.</emphasis>
Among Peruvian species, recognized by the combination of large size (ca.
<quantity id="4CF99B52A96A196FAA56FAB3429B61A0" box="[1320,1424,1403,1429]" metricMagnitude="-3" metricUnit="m" metricValue="3.92" pageId="11" pageNumber="72" unit="mm" value="3.92">3.92 mm</quantity>
), the moderately deep pronotal depressions, the roundly raised, almost impunctate pronotal reliefs, the 5
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elytral interstria with two callosities, the usual posterior and another, smaller, slightly anterior, and the male genitalia (
<figureCitation id="133A2A32A96A196FAA2DFA0B429D61E8" box="[1363,1430,1475,1501]" captionStart="FIGURE 8" captionStartId="12.[151,250,1861,1885]" captionTargetBox="[168,1427,534,1821]" captionTargetId="figure-166@12.[151,1435,510,1845]" captionTargetPageId="12" captionText="FIGURE 8. Hydrochus dualis n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.4417299" httpUri="https://zenodo.org/record/4417299/files/figure.png" pageId="11" pageNumber="72">Fig. 8</figureCitation>
; described below). Completely reliable determinations will include dissection of males.
</paragraph>
</subSubSection>
<subSubSection id="C31B653CA96A1968AFB9F9C2443E65C0" lastPageId="12" lastPageNumber="73" pageId="11" pageNumber="72" type="description">
<paragraph id="8BBE36B7A96A196FAFB9F9C244F662EC" blockId="11.[151,1437,1330,2041]" pageId="11" pageNumber="72">
<emphasis id="B975EAA5A96A196FAFB9F9C2465A6211" bold="true" box="[199,337,1546,1572]" pageId="11" pageNumber="72">Description</emphasis>
. Habitus as illustrated (
<figureCitation id="133A2A32A96A196FAD23F9C345A96210" box="[605,674,1547,1573]" captionStart="FIGURE 8" captionStartId="12.[151,250,1861,1885]" captionTargetBox="[168,1427,534,1821]" captionTargetId="figure-166@12.[151,1435,510,1845]" captionTargetPageId="12" captionText="FIGURE 8. Hydrochus dualis n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.4417299" httpUri="https://zenodo.org/record/4417299/files/figure.png" pageId="11" pageNumber="72">Fig. 8</figureCitation>
). Size:
<typeStatus id="54BA8815A96A196FAD89F9C344526210" box="[759,857,1547,1573]" pageId="11" pageNumber="72" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 3.92/1.54; head width 0.90; pronotum l/w 0.87/0.92; PA 0.82; PB 0.77; elytra 2.54/1.54. Body size large (ca.
<quantity id="4CF99B52A96A196FAA15F9E747CC6258" metricMagnitude="-3" metricUnit="m" metricValue="3.92" pageId="11" pageNumber="72" unit="mm" value="3.92">3.92 mm</quantity>
). Dorsum grey with some black on head, with slight to moderate iridescence, elytra with black spots. Legs brown, with tibiofemoral joints darker. Punctation on elytra ca. 0.51x that of pronotum. Elytra interstriae ca. 13x strial puncture diameter. Interstria 9
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more convex than others, overhanging 10
<superScript id="7C749BFFA96A196FAB6DF952432B629D" attach="left" box="[1043,1056,1690,1704]" fontSize="6" pageId="11" pageNumber="72">th</superScript>
interstria. Usual area of callus on 5
<superScript id="7C749BFFA96A196FAFDBF97647B962F9" attach="left" box="[165,178,1726,1740]" fontSize="6" pageId="11" pageNumber="72">th</superScript>
markedly raised, moderately elongate, ca. 5 punctures in row on each side.
</paragraph>
<paragraph id="8BBE36B7A96A196FAFB9F92B468B635C" blockId="11.[151,1437,1330,2041]" pageId="11" pageNumber="72">Pronotum very slightly wider than long, widest slightly behind anterior margin, narrowed at base, sides arcuate or very weakly sinuate, smooth; anterior margin slightly bisinuate; depressions moderately deep, more densely punctate than low reliefs between depressions, latter almost entirely impunctate; anteromedian depression less punctate than others.</paragraph>
<paragraph id="8BBE36B7A96A196FAFB9F8BB47FF63CC" blockId="11.[151,1437,1330,2041]" pageId="11" pageNumber="72">
Elytra with spaces between strial punctures ca. 12x their diameter; apices rather sharply conjointly rounded in dorsal view, in lateral view outer margin with moderate angulation; 3
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and 4
<superScript id="7C749BFFA96A196FAC9AF85E44FA6391" attach="left" box="[996,1009,1942,1956]" fontSize="6" pageId="11" pageNumber="72">th</superScript>
interstriae slightly raised immediately anterior to usual 5
<superScript id="7C749BFFA96A196FAE14F872467C63FD" attach="left" box="[362,375,1978,1992]" fontSize="6" pageId="11" pageNumber="72">th</superScript>
callosity; 5
<superScript id="7C749BFFA96A196FAE81F872450763FD" attach="left" box="[511,524,1978,1992]" fontSize="6" pageId="11" pageNumber="72">th</superScript>
interstria with two callosities, the usual posterior and another, smaller, slightly anterior.
</paragraph>
<paragraph id="8BBE36B7A96D1968AFB9FF5F42726484" blockId="12.[151,1437,151,503]" box="[199,1401,151,177]" pageId="12" pageNumber="73">Ventral characters: Mentum densely punctate, with large, deep median fovea. Submentum with two foveae.</paragraph>
<paragraph id="8BBE36B7A96D1968AFB9FF7343D86550" blockId="12.[151,1437,151,503]" pageId="12" pageNumber="73">
Male genitalia (
<figureCitation id="133A2A32A96D1968AE05FF7346B464E0" box="[379,447,187,213]" captionStart="FIGURE 8" captionStartId="12.[151,250,1861,1885]" captionTargetBox="[168,1427,534,1821]" captionTargetId="figure-166@12.[151,1435,510,1845]" captionTargetPageId="12" captionText="FIGURE 8. Hydrochus dualis n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.4417299" httpUri="https://zenodo.org/record/4417299/files/figure.png" pageId="12" pageNumber="73">Fig. 8</figureCitation>
) general characters: genitalia very stout, wide in lateral view; paramere tips extending slightly beyond distal end of aedeagus, lateral margin between plsa and distal end emarginate, medial margin arcuate, in lateral view extreme end of tips upturned slightly; aedeagus lanceolate, greatest width distal to midlength, with alsm; pseudoparameres large, extending to level of distal end of aedeagus, pseudoparameres wider in lateral than dorsal/ventral views; basal piece shorter than parameres, orifice damaged during dissection.
</paragraph>
<paragraph id="8BBE36B7A96D1968AFB9FEA7441B6598" blockId="12.[151,1437,151,503]" pageId="12" pageNumber="73">Dorsal surface: adtl located proximal to midlength of aedeagus; pdmm sinuate, overlapping lateral margins of aedeagus only over proximal 1/4; adbl moderately large.</paragraph>
<paragraph id="8BBE36B7A96D1968AFB9FE7F443E65C0" blockId="12.[151,1437,151,503]" pageId="12" pageNumber="73">Ventral surface: avtl U-shaped, near basal 1/5 of aedeagus; pvmm not overlapping lateral margins of aedeagus; pseudoparameres enveloping most of basal ¼ of parameres.</paragraph>
</subSubSection>
<caption id="DF7E663FA96D1968AFE9F88D44A86368" ID-DOI="http://doi.org/10.5281/zenodo.4417299" ID-Zenodo-Dep="4417299" box="[151,931,1861,1885]" httpUri="https://zenodo.org/record/4417299/files/figure.png" pageId="12" pageNumber="73" startId="12.[151,250,1861,1885]" targetBox="[168,1427,534,1821]" targetPageId="12">
<paragraph id="8BBE36B7A96D1968AFE9F88D44A86368" blockId="12.[151,931,1861,1885]" box="[151,931,1861,1885]" pageId="12" pageNumber="73">
<emphasis id="B975EAA5A96D1968AFE9F88D46186368" bold="true" box="[151,275,1861,1885]" pageId="12" pageNumber="73">FIGURE 8.</emphasis>
<taxonomicName id="4C014D34A96D1968AE67F88D46C76368" authority="Perkins, 2020" authorityName="Perkins" authorityYear="2020" box="[281,460,1861,1885]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="73" phylum="Arthropoda" rank="species" species="dualis" status="sp. nov.">
<emphasis id="B975EAA5A96D1968AE67F88D46C76368" box="[281,460,1861,1885]" italics="true" pageId="12" pageNumber="73">Hydrochus dualis</emphasis>
</taxonomicName>
<emphasis id="B975EAA5A96D1968AEACF88D45006368" bold="true" box="[466,523,1861,1885]" pageId="12" pageNumber="73">
<taxonomicNameLabel id="A24657DEA96D1968AEACF88D45006368" box="[466,523,1861,1885]" pageId="12" pageNumber="73" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, habitus and male genitalia of holotype.
</paragraph>
</caption>
<subSubSection id="C31B653CA96D1968AFB9F843430B6390" box="[199,1024,1931,1958]" pageId="12" pageNumber="73" type="etymology">
<paragraph id="8BBE36B7A96D1968AFB9F843430B6390" blockId="12.[151,1436,1931,2030]" box="[199,1024,1931,1958]" pageId="12" pageNumber="73">
<emphasis id="B975EAA5A96D1968AFB9F84346456390" bold="true" box="[199,334,1931,1957]" pageId="12" pageNumber="73">Etymology.</emphasis>
Named in reference to the two callosities on the 5
<superScript id="7C749BFFA96D1968ACFFF843448563AC" attach="left" box="[897,910,1931,1945]" fontSize="6" pageId="12" pageNumber="73">th</superScript>
interstria.
</paragraph>
</subSubSection>
<subSubSection id="C31B653CA96D1969AFB9F867430E6574" lastPageId="13" lastPageNumber="74" pageId="12" pageNumber="73" type="discussion">
<paragraph id="8BBE36B7A96D1969AFB9F867430E6574" blockId="12.[151,1436,1931,2030]" lastBlockId="13.[151,1437,151,321]" lastPageId="13" lastPageNumber="74" pageId="12" pageNumber="73">
<emphasis id="B975EAA5A96D1968AFB9F867463063FC" bold="true" box="[199,315,1967,1993]" pageId="12" pageNumber="73">Remarks.</emphasis>
The male genitalia of
<taxonomicName id="4C014D34A96D1968AD4AF878459163FC" authorityName="Perkins" authorityYear="2020" box="[564,666,1967,1993]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="73" phylum="Arthropoda" rank="species" species="dualis">
<emphasis id="B975EAA5A96D1968AD4AF878459163FC" box="[564,666,1967,1993]" italics="true" pageId="12" pageNumber="73">H. dualis</emphasis>
</taxonomicName>
are similar in some characters to that of
<taxonomicName id="4C014D34A96D1968AB22F878426F63FC" authority="Makhan" authorityName="Makhan" authorityYear="1992" box="[1116,1380,1967,1993]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="73" phylum="Arthropoda" rank="species" species="ramcharani">
<emphasis id="B975EAA5A96D1968AB22F878420963FC" box="[1116,1282,1967,1993]" italics="true" pageId="12" pageNumber="73">H. ramcharani</emphasis>
Makhan
</taxonomicName>
(figure of aedeagus given in
<bibRefCitation id="EF904B46A96D1968AECFF81B455763D8" author="Perkins, P. D." box="[433,604,2003,2029]" pageId="12" pageNumber="73" pagination="1 - 59" refId="ref12912" refString="Perkins, P. D. (2019 c) Taxonomy of Venezuelan water beetles in the genus Hydrochus Leach, 1817, and an analysis of male genitalia morphology (Coleoptera: Hydrochidae). Zootaxa, 4708 (1), 1 - 59. https: // doi. org / 10.11646 / zootaxa. 4708.1.1" type="journal article" year="2019">Perkins, 2019c</bibRefCitation>
). Differences in the two species include: 1) in
<taxonomicName id="4C014D34A96D1968AB18F81C43C763D8" authorityName="Perkins" authorityYear="2020" box="[1126,1228,2003,2029]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="73" phylum="Arthropoda" rank="species" species="dualis">
<emphasis id="B975EAA5A96D1968AB18F81C43C763D8" box="[1126,1228,2003,2029]" italics="true" pageId="12" pageNumber="73">H. dualis</emphasis>
</taxonomicName>
the paramere tips, from plsa to distal end, are shorter and the lateral margin is entirely emarginate, whereas in
<taxonomicName id="4C014D34A96C1969ABE2FF5F42486484" authorityName="Makhan" authorityYear="1992" box="[1180,1347,151,177]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="74" phylum="Arthropoda" rank="species" species="ramcharani">
<emphasis id="B975EAA5A96C1969ABE2FF5F42486484" box="[1180,1347,151,177]" italics="true" pageId="13" pageNumber="74">H. ramcharani</emphasis>
</taxonomicName>
the tips are nearly twice as long and the lateral margin is straight except emarginate subapically; 2) in
<taxonomicName id="4C014D34A96C1969ABD0FF73421F64E0" authorityName="Perkins" authorityYear="2020" box="[1198,1300,187,213]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="74" phylum="Arthropoda" rank="species" species="dualis">
<emphasis id="B975EAA5A96C1969ABD0FF73421F64E0" box="[1198,1300,187,213]" italics="true" pageId="13" pageNumber="74">H. dualis</emphasis>
</taxonomicName>
the pseudoparameres are much wider in lateral view for the entire length, whereas in
<taxonomicName id="4C014D34A96C1969ACA4FF17438B64CC" authorityName="Makhan" authorityYear="1992" box="[986,1152,223,249]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="74" phylum="Arthropoda" rank="species" species="ramcharani">
<emphasis id="B975EAA5A96C1969ACA4FF17438B64CC" box="[986,1152,223,249]" italics="true" pageId="13" pageNumber="74">H. ramcharani</emphasis>
</taxonomicName>
the pseudoparameres are cylindrical beyond ca. the basal 1/3; and 3) in
<taxonomicName id="4C014D34A96C1969ADE0FECB440F6528" authorityName="Perkins" authorityYear="2020" box="[670,772,259,285]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="74" phylum="Arthropoda" rank="species" species="dualis">
<emphasis id="B975EAA5A96C1969ADE0FECB440F6528" box="[670,772,259,285]" italics="true" pageId="13" pageNumber="74">H. dualis</emphasis>
</taxonomicName>
the adtl is located more proximally than in
<taxonomicName id="4C014D34A96C1969AB91FECB429E6528" authorityName="Makhan" authorityYear="1992" box="[1263,1429,259,285]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="74" phylum="Arthropoda" rank="species" species="ramcharani">
<emphasis id="B975EAA5A96C1969AB91FECB429E6528" box="[1263,1429,259,285]" italics="true" pageId="13" pageNumber="74">H. ramcharani</emphasis>
</taxonomicName>
, hence the membranous dorsal part of the aedeagus is much larger in
<taxonomicName id="4C014D34A96C1969ACE9FEEF44F66574" authorityName="Perkins" authorityYear="2020" box="[919,1021,295,321]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="74" phylum="Arthropoda" rank="species" species="dualis">
<emphasis id="B975EAA5A96C1969ACE9FEEF44F66574" box="[919,1021,295,321]" italics="true" pageId="13" pageNumber="74">H. dualis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>