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<mods:title id="2A5D1CE405B61834A46D7A3C8D2C4924">Ant fauna (Hymenoptera: Formicidae) of the Socotra Archipelago (Yemen): zoogeography, distribution and description of a new species</mods:title>
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<emphasis id="B940DB511623FFCCB496E04469ADCE00" bold="true" box="[424,756,867,892]" italics="true" pageId="53" pageNumber="369">Hypoponera punctatissima</emphasis>
(
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)
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(
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(ac))
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<taxonomicName id="4C347CC01623FFCCB59EE09D6920CEAE" authority="Roger, 1859: 246" authorityName="Roger" authorityPageNumber="246" authorityYear="1859" box="[160,633,954,978]" class="Liliopsida" family="Orchidaceae" genus="Ponera" kingdom="Plantae" order="Asparagales" pageId="53" pageNumber="369" phylum="Tracheophyta" rank="species" species="punctatissima">
<emphasis id="B940DB511623FFCCB59EE09D6ACCCEAE" box="[160,405,954,978]" italics="true" pageId="53" pageNumber="369">Ponera punctatissima</emphasis>
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, pl. 7,
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(w. q.)
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,
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. Palaearctic.
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<emphasis id="B940DB511623FFCCB59EE7076A4DC946" bold="true" box="[160,276,1056,1082]" italics="true" pageId="53" pageNumber="369">Diagnosis</emphasis>
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<emphasis id="B940DB511623FFCCB59EE7626BB5C921" box="[160,236,1093,1117]" italics="true" pageId="53" pageNumber="369">Worker</emphasis>
. Eyes small but distinct, situated far forward on sides of head; impression or feeble longitudinal line running back along midline of head from frontal lobes to midlength of vertex; scapes, when laid straight back from their insertions, fail to reach or less commonly just touch midpoint of posterior margin of head in full-face view; metanotal groove deeply impressed; mesonotum with a well-defined posterior margin; mesopleuron smooth and shining; petiole in profile with anterior and posterior faces of node weakly convergent dorsally. Dark brown to yellowish brown, antennae and legs yellowish.
</paragraph>
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<emphasis id="B940DB511623FFCCB59EE67F6AD9C80E" bold="true" box="[160,384,1368,1394]" italics="true" pageId="53" pageNumber="369">Material examined</emphasis>
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21 April
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, 196 m, (M.
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leg.),
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(21 w,
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,
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, W.
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,
<date id="FF8A21831623FFCCB716E520699FCB63" box="[552,710,1543,1567]" pageId="53" pageNumber="369" value="2014-04-20">
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20 April
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, 214 m, (M.
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.
<collectorName id="26C162951623FFCCB650E52068EDCB63" box="[878,948,1543,1567]" pageId="53" pageNumber="369">Sharaf</collectorName>
leg.),
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,
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(3 w,
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);
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<materialsCitation id="3B5C0D1E1623FFCCB486E50D6853CB19" ID-GBIF-Occurrence="3762058394" collectingDate="2014-04-24" collectingDateMax="2014-04-27" collectingDateMin="2014-04-25" collectionCode="R, KSMA" collectorName="W. Zereg &amp; Sharaf" country="Yemen" elevation="2014214" latitude="12.54038" location="Dixam" longLatPrecision="1" longitude="54.17186" pageId="53" pageNumber="369" specimenCount="1">
<collectingCountry id="F32347D31623FFCCB486E50D6950CB3E" box="[440,521,1578,1602]" name="Yemen" pageId="53" pageNumber="369">Yemen</collectingCountry>
,
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,
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(
<collectorName id="26C162951623FFCCB628E50D68D9CB3E" box="[790,896,1578,1602]" pageId="53" pageNumber="369">W. Zereg</collectorName>
),
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<collectingDate id="EFCED86B1623FFCCB6ACE50D6F72CB3E" box="[914,1067,1578,1602]" pageId="53" pageNumber="369" value="2014-04-24">
24 April
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</date>
, 279 m, (M.
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.
<collectorName id="26C162951623FFCCB5E1E56A6A7CCB19" box="[223,293,1613,1637]" pageId="53" pageNumber="369">Sharaf</collectorName>
leg.),
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,
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(1 w,
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)
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.
</paragraph>
</subSubSection>
<caption id="DF4B57CB1620FFCFB59EE58A69F7CB9D" ID-DOI="http://doi.org/10.5281/zenodo.5184351" ID-Zenodo-Dep="5184351" httpUri="https://zenodo.org/record/5184351/files/figure.png" pageId="54" pageNumber="370" startId="54.[160,225,1709,1732]" targetBox="[253,1061,142,1667]" targetPageId="54">
<paragraph id="8B8B07431620FFCFB59EE58A69F7CB9D" blockId="54.[160,1157,1709,1761]" pageId="54" pageNumber="370">
<emphasis id="B940DB511620FFCFB59EE58A6A5ECBB8" bold="true" box="[160,263,1709,1732]" pageId="54" pageNumber="370">Figure 28.</emphasis>
<taxonomicName id="4C347CC01620FFCFB42EE58A6949CBB8" baseAuthorityName="Roger" baseAuthorityYear="1859" box="[272,528,1709,1732]" class="Insecta" family="Formicidae" genus="Hypoponera" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="370" phylum="Arthropoda" rank="species" species="punctatissima">
<emphasis id="B940DB511620FFCFB42EE58A6949CBB8" box="[272,528,1709,1732]" italics="true" pageId="54" pageNumber="370">Hypoponera punctatissima</emphasis>
</taxonomicName>
. (a) Body in profile. (b) Head in full-face view, CASENT0746620, AntWeb.org, (Zach Lieberman). (c) Distribution map.
</paragraph>
</caption>
<subSubSection id="C32E54C81621FFCEB59EE3AA6BA2CC0B" pageId="55" pageNumber="371" type="distribution">
<paragraph id="8B8B07431621FFCEB59EE3AA6A2ECDDB" blockId="55.[160,1156,141,375]" box="[160,375,141,167]" pageId="55" pageNumber="371">
<emphasis id="B940DB511621FFCEB59EE3AA6A2ECDDB" bold="true" box="[160,375,141,167]" italics="true" pageId="55" pageNumber="371">Geographic range</emphasis>
</paragraph>
<paragraph id="8B8B07431621FFCEB59EE3956BA2CC0B" blockId="55.[160,1156,141,375]" pageId="55" pageNumber="371">
This species is broadly distributed throughout the Afrotropical region (
<bibRefCitation id="EFA57AB21621FFCEB682E3956B8ECD90" author="Bolton B &amp; Fisher BL" pageId="55" pageNumber="371" pagination="1 - 118" refId="ref17955" refString="Bolton B, Fisher BL. 2011. Taxonomy of Afrotropical and West Palaearctic ants of the ponerine genus Hypoponera Santschi (Hymenoptera: Formicidae). Zootaxa. 2843: 1 - 118." type="journal article" year="2011">Bolton and Fisher 2011</bibRefCitation>
) and Europe (
<collectingCountry id="F32347D31621FFCEB441E3F36A86CD90" box="[383,479,212,236]" name="Belgium" pageId="55" pageNumber="371">Belgium</collectingCountry>
, Britain,
<collectingCountry id="F32347D31621FFCEB778E3F369F6CD90" box="[582,687,212,236]" name="Denmark" pageId="55" pageNumber="371">Denmark</collectingCountry>
,
<collectingCountry id="F32347D31621FFCEB782E3F3684BCD90" box="[700,786,212,236]" name="Finland" pageId="55" pageNumber="371">Finland</collectingCountry>
,
<collectingCountry id="F32347D31621FFCEB61EE3F36834CD90" box="[800,877,212,236]" name="France" pageId="55" pageNumber="371">France</collectingCountry>
,
<collectingCountry id="F32347D31621FFCEB645E3F368BDCD90" box="[891,996,212,236]" name="Germany" pageId="55" pageNumber="371">Germany</collectingCountry>
,
<collectingCountry id="F32347D31621FFCEB6CFE3F36F18CD90" box="[1009,1089,212,236]" name="Ireland" pageId="55" pageNumber="371">Ireland</collectingCountry>
,
<collectingCountry id="F32347D31621FFCEB171E3F36FD9CD90" box="[1103,1152,212,236]" name="Italy" pageId="55" pageNumber="371">Italy</collectingCountry>
,
<collectingCountry id="F32347D31621FFCEB59EE3D06A6BCC73" box="[160,306,247,271]" name="Luxembourg" pageId="55" pageNumber="371">Luxembourg</collectingCountry>
,
<collectingCountry id="F32347D31621FFCEB47EE3D06AC0CC73" box="[320,409,247,271]" name="Norway" pageId="55" pageNumber="371">Norway</collectingCountry>
,
<collectingCountry id="F32347D31621FFCEB498E3D06AAFCC73" box="[422,502,247,271]" name="Poland" pageId="55" pageNumber="371">Poland</collectingCountry>
,
<collectingCountry id="F32347D31621FFCEB73AE3D0691CCC73" box="[516,581,247,271]" name="Spain" pageId="55" pageNumber="371">Spain</collectingCountry>
,
<collectingCountry id="F32347D31621FFCEB76DE3D0698CCC73" box="[595,725,247,271]" name="Switzerland" pageId="55" pageNumber="371">Switzerland</collectingCountry>
and the Balkans) (
<bibRefCitation id="EFA57AB21621FFCEB68DE3D06B8ECC4E" author="Delabie JHC &amp; Blard F" pageId="55" pageNumber="371" pagination="149 - 151" refId="ref18406" refString="Delabie JHC, Blard F. 2002. The tramp ant Hypoponera punctatissima (Roger) (Hymenoptera: Formicidae: Ponerinae): new records from the southern hemisphere. Neotrop Entomol. 31: 149 - 151." type="journal article" year="2002">Delabie and Blard 2002</bibRefCitation>
). For the Arabian Peninsula, it has been recorded from
<collectingCountry id="F32347D31621FFCEB656E23D68CDCC4E" box="[872,916,282,306]" name="Saudi Arabia" pageId="55" pageNumber="371">KSA</collectingCountry>
(
<bibRefCitation id="EFA57AB21621FFCEB69BE23D6F2FCC4E" author="Collingwood CA" box="[933,1142,282,306]" pageId="55" pageNumber="371" pagination="230 - 302" refId="ref18149" refString="Collingwood CA. 1985. Hymenoptera: fam. Formicidae of Saudi Arabia. Fauna Saudi Arabia. 7: 230 - 302." type="journal article" year="1985">Collingwood 1985</bibRefCitation>
),
<collectingCountry id="F32347D31621FFCEB59EE21B6BBDCC28" box="[160,228,316,340]" name="Oman" pageId="55" pageNumber="371">Oman</collectingCountry>
and
<collectingCountry id="F32347D31621FFCEB417E21B6A2ECC28" box="[297,375,316,340]" name="Yemen" pageId="55" pageNumber="371">Yemen</collectingCountry>
(
<bibRefCitation id="EFA57AB21621FFCEB4B4E21B69B0CC28" author="Collingwood CA &amp; Agosti D" box="[394,745,316,340]" pageId="55" pageNumber="371" pagination="300 - 385" refId="ref18173" refString="Collingwood CA, Agosti D. 1996. Formicidae (Insecta: Hymenoptera) of Saudi Arabia (part 2). Fauna of Saudi Arabia. 15: 300 - 385." type="journal article" year="1996">Collingwood and Agosti 1996</bibRefCitation>
). This species is a new record for
<collectingCountry id="F32347D31621FFCEB59EE2786BA1CC0B" box="[160,248,351,375]" name="Yemen" pageId="55" pageNumber="371">Socotra</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C32E54C81621FFCEB59EE284696ACFD3" pageId="55" pageNumber="371" type="biology_ecology">
<paragraph id="8B8B07431621FFCEB59EE284694CCCC1" blockId="55.[160,1155,419,687]" box="[160,533,419,445]" pageId="55" pageNumber="371">
<emphasis id="B940DB511621FFCEB59EE284694CCCC1" bold="true" box="[160,533,419,445]" italics="true" pageId="55" pageNumber="371">Ecological and biological notes</emphasis>
</paragraph>
<paragraph id="8B8B07431621FFCEB59EE2E0696ACFD3" blockId="55.[160,1155,419,687]" pageId="55" pageNumber="371">
This genus is recorded for the first time from
<collectingCountry id="F32347D31621FFCEB7FBE2E06845CCA3" box="[709,796,455,479]" name="Yemen" pageId="55" pageNumber="371">Socotra</collectingCountry>
; based on its distribution, this species has probably been introduced. This species was collected from moist soil and leaf litter under mango and date palm trees where the soil was rich in decaying organic material. It was also observed nesting under a stone next to a dragon blood tree. This species was also nesting directly in moist soil under a dead palm tree where organic materials are found. Another nest was found among old moist leaf sheaths surrounding the trunk base of a date palm tree.
</paragraph>
</subSubSection>
</treatment>
</document>