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<document id="A3E6444E5BC6EA6C102A957938467CE8" ID-DOI="10.5281/zenodo.274748" ID-GBIF-Dataset="e687202a-a625-4aab-94de-312277c7e3fb" ID-ISSN="1175-5326" ID-Zenodo-Dep="274748" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461047074832" checkinUser="plazi" docAuthor="Aarvik, Leif &amp; Karisch, Timm" docDate="2009" docId="039687B6FFA5FFF3FF6E5A81FBEA48B9" docLanguage="en" docName="zt02026p032.pdf" docOrigin="Zootaxa 2026" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Multiquaestia purana Aarvik &amp; Karisch, new species" docType="treatment" docVersion="8" lastPageNumber="25" masterDocId="FFAFFFCEFFA2FFF4FFF95853FFB14F5D" masterDocTitle="Revision of Multiquaestia Karisch (Lepidoptera: Tortricidae: Grapholitini)" masterLastPageNumber="32" masterPageNumber="18" pageNumber="25" updateTime="1698577463540" updateUser="plazi">
<mods:mods id="E11F0FD4F5370BEB0E7624F7950B32A3" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8C7232B38A65667AA60467CD22D4CA44">
<mods:title id="A1426512BB6EC2D4802C3700A9A2A6C5">Revision of Multiquaestia Karisch (Lepidoptera: Tortricidae: Grapholitini)</mods:title>
</mods:titleInfo>
<mods:name id="54CA3E9B3E2965C8D11BA000E73CA480" type="personal">
<mods:role id="4C5544B45EFEE4EE2F41C5303D35C8B5">
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</mods:role>
<mods:namePart id="BA55B28168247AA6920234E96B2467E9">Aarvik, Leif</mods:namePart>
</mods:name>
<mods:name id="0DAF2A682D0B44687C5D848F4926F86D" type="personal">
<mods:role id="90D8CEA9D0833D5481F5223AC44EE1DF">
<mods:roleTerm id="39079A457C4A947C4683F5B8F6172CA4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EFE26C3065624E1A1AED5BAA01A237D2">Karisch, Timm</mods:namePart>
</mods:name>
<mods:typeOfResource id="0CB814A3F8B3F15B4556B22C301D2C67">text</mods:typeOfResource>
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<mods:titleInfo id="A57AB90C2EBC9A22FEF0B64C4A61292F">
<mods:title id="7C99D42E3316B77B9009EFF20A7827A4">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="82313412D5ECDDEB4F5E0671053F9AEE">
<mods:date id="BD7D00DE8421BCAB805E822297C2CA71">2009</mods:date>
<mods:detail id="5DFC19B7DA4621E05254F093745CA9C8" type="volume">
<mods:number id="B2B9406D1828E7813AC7AEAD203284CE">2026</mods:number>
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<mods:start id="E99A4B308648D0A5CB9B5F0AA0DF8D99">18</mods:start>
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<mods:classification id="A1FBFB6D53733BC0137CD5C8E12A0499">journal article</mods:classification>
<mods:identifier id="D266331E5B37056BCD9CAF4DC7C1855F" type="DOI">10.5281/zenodo.274748</mods:identifier>
<mods:identifier id="62CA0FFE123D4AFEDCE5DE59AFA0327F" type="GBIF-Dataset">e687202a-a625-4aab-94de-312277c7e3fb</mods:identifier>
<mods:identifier id="93BD559DF35E6889A7B6DEE52D305DD0" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="FA7D1DD751BED618A8E6B5F24B2020BF" type="Zenodo-Dep">274748</mods:identifier>
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<treatment id="039687B6FFA5FFF3FF6E5A81FBEA48B9" ID-DOI="http://doi.org/10.5281/zenodo.6216217" ID-GBIF-Taxon="119611111" ID-Zenodo-Dep="6216217" LSID="urn:lsid:plazi:treatment:039687B6FFA5FFF3FF6E5A81FBEA48B9" httpUri="http://treatment.plazi.org/id/039687B6FFA5FFF3FF6E5A81FBEA48B9" lastPageNumber="25" pageId="7" pageNumber="25">
<subSubSection id="C325652BFFA5FFF3FF6E5A81FE3A4C52" pageId="7" pageNumber="25" type="nomenclature">
<paragraph id="8B8036A0FFA5FFF3FF6E5A81FC974DB1" blockId="7.[151,806,722,783]" box="[151,806,722,748]" pageId="7" pageNumber="25">
<heading id="D0C881CCFFA5FFF3FF6E5A81FC974DB1" bold="true" box="[151,806,722,748]" fontSize="11" level="1" pageId="7" pageNumber="25" reason="1">
<emphasis id="B94BEAB2FFA5FFF3FF6E5A81FC974DB1" bold="true" box="[151,806,722,748]" pageId="7" pageNumber="25">
<taxonomicName id="4C3F4D23FFA5FFF3FF6E5A81FD384DB1" ID-CoL="B3XJN" authority="Aarvik &amp; Karisch" authorityName="Aarvik &amp; Karisch" box="[151,649,722,748]" class="Insecta" family="Tortricidae" genus="Multiquaestia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="25" phylum="Arthropoda" rank="species" species="purana" status="sp. nov.">
<emphasis id="B94BEAB2FFA5FFF3FF6E5A81FE2F4DB1" bold="true" box="[151,414,722,748]" italics="true" pageId="7" pageNumber="25">Multiquaestia purana</emphasis>
Aarvik &amp; Karisch
</taxonomicName>
,
<taxonomicNameLabel id="A27857C9FFA5FFF3FD6F5A81FC974DB1" box="[662,806,722,748]" pageId="7" pageNumber="25" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8036A0FFA5FFF3FF6E5AA6FE3A4C52" blockId="7.[151,806,722,783]" box="[151,395,757,783]" pageId="7" pageNumber="25">
(
<figureCitation id="13042A25FFA5FFF3FF665AA6FF4E4C52" box="[159,255,757,783]" captionStart="FIGURES 7 – 14" captionStartId="4.[151,269,1941,1965]" captionTargetBox="[151,1436,223,1920]" captionTargetId="figure@4.[151,1436,183,1922]" captionTargetPageId="4" captionText="FIGURES 7 – 14. Adults of Multiquaestia. 7. M. skulei n. sp. Holotype. 8. M. agassizi n. sp. Holotype. 9. M. dallastai n. sp. Holotype. 10. M. kingstoni n. sp. Holotype. 11. M. purana n. sp. 12. M. iringana n. sp. 13. M. albimaculana Karisch. Holotype. 14. M. andersi n. sp. Holotype. Scale 10 mm." httpUri="https://zenodo.org/record/274750/files/figure.png" pageId="7" pageNumber="25">Figs. 11</figureCitation>
,
<figureCitation id="13042A25FFA5FFF3FEF55AA6FEE84C52" box="[268,345,757,783]" captionStart="FIGURES 23 – 28" captionStartId="9.[151,274,1557,1581]" captionTargetBox="[151,1436,194,1533]" captionTargetId="figure@9.[151,1436,194,1534]" captionTargetPageId="9" captionText="FIGURES 23 – 28. Male genitalia of Multiquaestia. 23. M. dallastai n. sp. 24. M. dallastai n. sp. aedeagus. 25. M. kingstoni n. sp. 26. M. kingstoni n. sp. aedeagus. 27. M. purana n. sp. 28. M. purana n. sp. aedeagus." httpUri="https://zenodo.org/record/274753/files/figure.png" pageId="7" pageNumber="25">27, 28</figureCitation>
,
<figureCitation id="13042A25FFA5FFF3FE9D5AA6FE334C52" box="[356,386,757,783]" captionStart="FIGURES 36 – 40" captionStartId="12.[151,271,1721,1745]" captionTargetBox="[151,1429,194,1688]" captionTargetId="figure@12.[151,1436,194,1697]" captionTargetPageId="12" captionText="FIGURES 36 – 40. Female genitalia of Multiquaestia. 36. M. purana n. sp. 37. M. iringana n. sp. 38. M. albimaculana Karisch. 39. Multiquaestia sp. 1. 40. Multiquaestia sp. 2." httpUri="https://zenodo.org/record/274755/files/figure.png" pageId="7" pageNumber="25">36</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C325652BFFA5FFF3FF6E5B11FAE34CF7" pageId="7" pageNumber="25" type="materials_examined">
<paragraph id="8B8036A0FFA5FFF3FF6E5B11FAE34CF7" blockId="7.[151,1437,834,2020]" pageId="7" pageNumber="25">
<emphasis id="B94BEAB2FFA5FFF3FF6E5B11FEEC4C01" bold="true" box="[151,349,834,860]" pageId="7" pageNumber="25">
<typeStatus id="54848802FFA5FFF3FF6E5B11FF684C01" box="[151,217,834,860]" pageId="7" pageNumber="25">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54848802FFA5FFF3FE9F5B11FE514C01" box="[358,480,834,860]" pageId="7" pageNumber="25" type="holotype">Holotype</typeStatus>
, 3,
<collectingCountry id="F3287630FFA5FFF3FDEF5B11FD094C01" box="[534,696,834,860]" name="Tanzania" pageId="7" pageNumber="25">TANZANIA</collectingCountry>
, Iringa Region, Mufindi District, Kigogo Forest
<quantity id="4CC79B45FFA5FFF3FAD55B11FA224C01" box="[1324,1427,834,860]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="7" pageNumber="25" unit="m" value="1900.0">1900 m</quantity>
.,
<date id="FF811060FFA5FFF3FF6E5B3AFEF54CDE" box="[151,324,873,899]" pageId="7" pageNumber="25" value="2005-11-23" valueMax="2005-11-25" valueMin="2005-11-23">23–25.xi.2005</date>
, leg. L. Aarvik, M. Fibiger, A. Kingston, genitalia slide 2724 L. Aarvik (
<collectionCode id="ED2EAE65FFA5FFF3FB555B3AFABC4CDE" box="[1196,1293,873,899]" httpUri="http://grbio.org/cool/2knt-7f1r" name="Natural History Museum, University of Oslo" pageId="7" pageNumber="25">NHMO</collectionCode>
).
<typeStatus id="54848802FFA5FFF3FAD85B3AFA294CDE" box="[1313,1432,873,899]" pageId="7" pageNumber="25" type="paratype">Paratypes</typeStatus>
: 33, 1Ƥ, same data as
<typeStatus id="54848802FFA5FFF3FE605BC3FDB24CF7" box="[409,515,912,938]" pageId="7" pageNumber="25" type="holotype">holotype</typeStatus>
, female with genitalia slide 2722 L. Aarvik (23 LAA; 13, 1Ƥ
<collectionCode id="ED2EAE65FFA5FFF3FB1E5BC3FAF74CF7" box="[1255,1350,912,938]" httpUri="http://grbio.org/cool/2knt-7f1r" name="Natural History Museum, University of Oslo" pageId="7" pageNumber="25">NHMO</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C325652BFFA5FFF3FF3F5BE5FCEF498B" pageId="7" pageNumber="25" type="description">
<paragraph id="8B8036A0FFA5FFF3FF3F5BE5FA8D4A9A" blockId="7.[151,1437,834,2020]" pageId="7" pageNumber="25">
<emphasis id="B94BEAB2FFA5FFF3FF3F5BE5FEEE4C8D" bold="true" box="[198,351,950,976]" pageId="7" pageNumber="25">Description.</emphasis>
Adult. Male (
<figureCitation id="13042A25FFA5FFF3FDFB5BE5FDE94C8D" box="[514,600,950,976]" captionStart="FIGURES 7 – 14" captionStartId="4.[151,269,1941,1965]" captionTargetBox="[151,1436,223,1920]" captionTargetId="figure@4.[151,1436,183,1922]" captionTargetPageId="4" captionText="FIGURES 7 – 14. Adults of Multiquaestia. 7. M. skulei n. sp. Holotype. 8. M. agassizi n. sp. Holotype. 9. M. dallastai n. sp. Holotype. 10. M. kingstoni n. sp. Holotype. 11. M. purana n. sp. 12. M. iringana n. sp. 13. M. albimaculana Karisch. Holotype. 14. M. andersi n. sp. Holotype. Scale 10 mm." httpUri="https://zenodo.org/record/274750/files/figure.png" pageId="7" pageNumber="25">Fig. 11</figureCitation>
). Wingspan 15.0–16.5 mm.
<emphasis id="B94BEAB2FFA5FFF3FC585BE4FC514C8D" box="[929,992,951,976]" italics="true" pageId="7" pageNumber="25">Head</emphasis>
: Dirty white. Antenna brownish grey, scape dirty white above. Labial palpus 1.5 times diameter of eye, externally dark grey, internally light grey; rather slender as scaling of second segment is less pronounced than in other species, third segment drooping.
<emphasis id="B94BEAB2FFA5FFF3FF6E5C79FF5D4B1E" box="[151,236,1066,1091]" italics="true" pageId="7" pageNumber="25">Thorax</emphasis>
: Anteriorly greyish white, greyish brown medially, with distinct white posterior spot; tegulae brown. Legs brownish grey, tarsi with paler rings. Forewing upperside with basal 2/3 blackish brown; terminal 1/3 ochreous; on dorsum two white marks; the inner mark reaching to about 2/5 from dorsum, the distal one sharply pointed, reaching to 1/3; a white dot present somewhat beyond distal white mark, and one between wing base and inner white mark; a few dark scales present in terminal area.
<collectingCountry id="F3287630FFA5FFF3FBAF5C96FB254B82" box="[1110,1172,1221,1247]" name="India" pageId="7" pageNumber="25">Hind</collectingCountry>
wing upperside dark brownish grey; ochreous hair-pencil from base of hind margin to scale cover on dorsum of abdomen.
<emphasis id="B94BEAB2FFA5FFF3FF6E5D40FEB74A71" box="[151,262,1299,1324]" italics="true" pageId="7" pageNumber="25">Abdomen</emphasis>
: Grey, abdominal brush dark grey above and beneath, laterally light yellowish grey. Male genitalia (
<figureCitation id="13042A25FFA5FFF3FF595D6AFF464A0E" box="[160,247,1337,1363]" captionStart="FIGURES 23 – 28" captionStartId="9.[151,274,1557,1581]" captionTargetBox="[151,1436,194,1533]" captionTargetId="figure@9.[151,1436,194,1534]" captionTargetPageId="9" captionText="FIGURES 23 – 28. Male genitalia of Multiquaestia. 23. M. dallastai n. sp. 24. M. dallastai n. sp. aedeagus. 25. M. kingstoni n. sp. 26. M. kingstoni n. sp. aedeagus. 27. M. purana n. sp. 28. M. purana n. sp. aedeagus." httpUri="https://zenodo.org/record/274753/files/figure.png" pageId="7" pageNumber="25">Fig. 27</figureCitation>
) with tegumen evenly curved in dorsal part, uncus not differentiated, socii present but weak; lower edge of sacculus angled, narrow portion of valva long and curved, cucullus large, broad, subcircular with a rounded-rectangular ventral angle; aedeagus (
<figureCitation id="13042A25FFA5FFF3FD4D5DD5FCBA4AFD" box="[692,779,1414,1440]" captionStart="FIGURES 23 – 28" captionStartId="9.[151,274,1557,1581]" captionTargetBox="[151,1436,194,1533]" captionTargetId="figure@9.[151,1436,194,1534]" captionTargetPageId="9" captionText="FIGURES 23 – 28. Male genitalia of Multiquaestia. 23. M. dallastai n. sp. 24. M. dallastai n. sp. aedeagus. 25. M. kingstoni n. sp. 26. M. kingstoni n. sp. aedeagus. 27. M. purana n. sp. 28. M. purana n. sp. aedeagus." httpUri="https://zenodo.org/record/274753/files/figure.png" pageId="7" pageNumber="25">Fig. 28</figureCitation>
) with basal 1/5 broader and straight, otherwise slender, evenly curved; with 9 cornuti of different length, from large spindle-shaped to very small; 10 carinae.
</paragraph>
<paragraph id="8B8036A0FFA5FFF3FF3F5D87FCEF498B" blockId="7.[151,1437,834,2020]" pageId="7" pageNumber="25">
Female.
<emphasis id="B94BEAB2FFA5FFF3FED75D87FE634AB0" box="[302,466,1492,1517]" italics="true" pageId="7" pageNumber="25">Head, Thorax</emphasis>
: Similar to male, but blackish suffusion on basal 2/3 of forewing upperside less heavy. Labial palpus 2.0 times diameter of eye, legs paler.
<emphasis id="B94BEAB2FFA5FFF3FC9B5DA8FC624949" box="[866,979,1531,1556]" italics="true" pageId="7" pageNumber="25">Abdomen</emphasis>
: Female genitalia (
<figureCitation id="13042A25FFA5FFF3FB465DA9FAA64949" box="[1215,1303,1530,1556]" captionStart="FIGURES 36 – 40" captionStartId="12.[151,271,1721,1745]" captionTargetBox="[151,1429,194,1688]" captionTargetId="figure@12.[151,1436,194,1697]" captionTargetPageId="12" captionText="FIGURES 36 – 40. Female genitalia of Multiquaestia. 36. M. purana n. sp. 37. M. iringana n. sp. 38. M. albimaculana Karisch. 39. Multiquaestia sp. 1. 40. Multiquaestia sp. 2." httpUri="https://zenodo.org/record/274755/files/figure.png" pageId="7" pageNumber="25">Fig. 36</figureCitation>
) with both pairs of apophyses long; posterior edge of sternite 7 with deep round excavation, anterior edge strongly sclerotised and with two pairs of humps; lamella postvaginalis long, broadened posteriorly; ostium cylindrical, ventral margin deeply concave; ductus bursae gradually widened towards corpus bursae, sclerotised in medial part; ductus seminalis inserted into ductus bursae just before corpus bursae; corpus bursae with two curved, horn-shaped signa with small bases.
</paragraph>
</subSubSection>
<subSubSection id="C325652BFFA5FFF3FF3F5EB1FB3D482D" pageId="7" pageNumber="25" type="diagnosis">
<paragraph id="8B8036A0FFA5FFF3FF3F5EB1FB3D482D" blockId="7.[151,1437,834,2020]" pageId="7" pageNumber="25">
<emphasis id="B94BEAB2FFA5FFF3FF3F5EB1FEF749A1" bold="true" box="[198,326,1762,1788]" pageId="7" pageNumber="25">Diagnosis.</emphasis>
This species is externally similar to
<taxonomicName id="4C3F4D23FFA5FFF3FD095EB0FC3049A1" box="[752,897,1763,1788]" class="Insecta" family="Tortricidae" genus="Multiquaestia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="25" phylum="Arthropoda" rank="species" species="kingstoni">
<emphasis id="B94BEAB2FFA5FFF3FD095EB0FC3049A1" box="[752,897,1763,1788]" italics="true" pageId="7" pageNumber="25">M. kingstoni</emphasis>
</taxonomicName>
from which it differs by its larger size and the presence of a distinct white spot posteriorly on the thorax. In the male genitalia, the broad, nearly subcircular cucullus is characteristic of
<taxonomicName id="4C3F4D23FFA5FFF3FE255F63FDE64814" box="[476,599,1840,1865]" class="Insecta" family="Tortricidae" genus="Multiquaestia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="25" phylum="Arthropoda" rank="species" species="purana">
<emphasis id="B94BEAB2FFA5FFF3FE255F63FDE64814" box="[476,599,1840,1865]" italics="true" pageId="7" pageNumber="25">M. purana</emphasis>
</taxonomicName>
. In the female, the deeply excavated posterior edge of sternite 7 and the long lamella postvaginalis are very different from other known
<taxonomicName id="4C3F4D23FFA5FFF3FC865F04FB93482D" box="[895,1058,1879,1904]" class="Insecta" family="Tortricidae" genus="Multiquaestia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="B94BEAB2FFA5FFF3FC865F04FB93482D" box="[895,1058,1879,1904]" italics="true" pageId="7" pageNumber="25">Multiquaestia</emphasis>
</taxonomicName>
females.
</paragraph>
</subSubSection>
<subSubSection id="C325652BFFA5FFF3FF3F5F2EFE6D48CA" box="[198,476,1917,1943]" pageId="7" pageNumber="25" type="distribution">
<paragraph id="8B8036A0FFA5FFF3FF3F5F2EFE6D48CA" blockId="7.[151,1437,834,2020]" box="[198,476,1917,1943]" pageId="7" pageNumber="25">
<emphasis id="B94BEAB2FFA5FFF3FF3F5F2EFED648CA" bold="true" box="[198,359,1917,1943]" pageId="7" pageNumber="25">Distribution.</emphasis>
<collectingCountry id="F3287630FFA5FFF3FE945F2EFE6448CA" box="[365,469,1917,1943]" name="Tanzania" pageId="7" pageNumber="25">Tanzania</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C325652BFFA5FFF3FF3F5FF0FBEA48B9" pageId="7" pageNumber="25" type="etymology">
<paragraph id="8B8036A0FFA5FFF3FF3F5FF0FBEA48B9" blockId="7.[151,1437,834,2020]" pageId="7" pageNumber="25">
<emphasis id="B94BEAB2FFA5FFF3FF3F5FF0FEE248E0" bold="true" box="[198,339,1955,1981]" pageId="7" pageNumber="25">Etymology.</emphasis>
The species name refers to the uniformly coloured markings of the forewing, in contrast to
<taxonomicName id="4C3F4D23FFA5FFF3FA855FF7FF4D48B9" class="Insecta" family="Tortricidae" genus="Multiquaestia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="25" phylum="Arthropoda" rank="species" species="iringana" status="sp. nov.">
<emphasis id="B94BEAB2FFA5FFF3FA855FF7FF4D48B9" italics="true" pageId="7" pageNumber="25">M. iringana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27857C9FFA5FFF3FEFA5F99FEF448B9" box="[259,325,1994,2020]" pageId="7" pageNumber="25" rank="species">
<emphasis id="B94BEAB2FFA5FFF3FEFA5F99FEF448B9" bold="true" box="[259,325,1994,2020]" pageId="7" pageNumber="25">n. sp.</emphasis>
</taxonomicNameLabel>
from the same locality, which has a more blurred forewing pattern.
</paragraph>
</subSubSection>
</treatment>
</document>
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