641 lines
121 KiB
XML
641 lines
121 KiB
XML
<document id="BF6DA844A9DF76177036A74643B88B1F" ID-DOI="10.11646/zootaxa.5529.3.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="14022748" ID-ZooBank="716EDF67-C933-484D-911B-4585B11187A6" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1730453336952" checkinUser="plazi" docAuthor="Wesener, Thomas" docDate="2024" docId="0394B500FFDDFFB3FF322CD4D249FD80" docLanguage="en" docName="zootaxa.5529.3.3.pdf" docOrigin="Zootaxa 5529 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5529.3.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Alluviobolus omega Wesener, 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="475" masterDocId="FFADCD78FFD4FFBDFFA52C0BD158FF99" masterDocTitle="Five new species and numerous locality records of Spirobolida millipedes from Madagascar (Diplopoda, Spirobolida, Pachybolidae)" masterLastPageNumber="486" masterPageNumber="461" pageNumber="470" updateTime="1730454045455" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="73AF4533E23F9EA884EAFEB046D8EAB6">Five new species and numerous locality records of Spirobolida millipedes from Madagascar (Diplopoda, Spirobolida, Pachybolidae)</mods:title>
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<mods:roleTerm id="119904ADDD797C19EA3C3119C3AF75D8">Author</mods:roleTerm>
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<mods:namePart id="36F13842A7916D0FDEE1AECC6DAC74E4">Wesener, Thomas</mods:namePart>
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<mods:typeOfResource id="5AFFB0217934016FCCFE4460FB7BF114">text</mods:typeOfResource>
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<mods:title id="E8892735D8FCCF7E88CE98D0EB190B43">Zootaxa</mods:title>
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<mods:part id="38AA7941322D39A58351D1DC5D18ABBA">
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<mods:date id="CFD32508AFA78EAEA96874826FDA4A14">2024</mods:date>
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<mods:number id="9EDE0E7C2FCCB1E89669EC847E9A61A1">2024-10-29</mods:number>
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<mods:number id="DFCFFEFEAAE2728FA8DB476411AA0831">5529</mods:number>
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<mods:url id="FC66AC871D4D0CDB37D4E16390D761E7">http://dx.doi.org/10.11646/zootaxa.5529.3.3</mods:url>
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<mods:classification id="DA1FD7A5A8415F1CC1E0B94234E143D4">journal article</mods:classification>
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<mods:identifier id="C38D8EA238A4626B588FB077036278D4" type="DOI">10.11646/zootaxa.5529.3.3</mods:identifier>
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<mods:identifier id="14BD014055567A9D5BAFFB79F960FC40" type="ISSN">1175-5326</mods:identifier>
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<treatment id="0394B500FFDDFFB3FF322CD4D249FD80" LSID="urn:lsid:plazi:treatment:0394B500FFDDFFB3FF322CD4D249FD80" httpUri="http://treatment.plazi.org/id/0394B500FFDDFFB3FF322CD4D249FD80" lastPageId="14" lastPageNumber="475" pageId="9" pageNumber="470">
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<subSubSection id="C327579DFFDDFFB4FF322CD4D0B4FF60" box="[151,492,223,250]" pageId="9" pageNumber="470" type="nomenclature">
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<paragraph id="8B820416FFDDFFB4FF322CD4D0B4FF60" blockId="9.[151,492,223,286]" box="[151,492,223,250]" pageId="9" pageNumber="470">
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<heading id="D0CAB37AFFDDFFB4FF322CD4D0B4FF60" bold="true" box="[151,492,223,250]" fontSize="11" level="1" pageId="9" pageNumber="470" reason="1">
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<emphasis id="B949D804FFDDFFB4FF322CD4D0B4FF60" bold="true" box="[151,492,223,250]" pageId="9" pageNumber="470">
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<taxonomicName id="4C3D7F95FFDDFFB4FF322CD4D0D0FF60" authorityName="Wesener" authorityYear="2024" box="[151,392,223,250]" class="Diplopoda" family="Pachybolidae" genus="Alluviobolus" kingdom="Animalia" order="Spirobolida" pageId="9" pageNumber="470" phylum="Arthropoda" rank="species" species="omega" status="sp. nov.">
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<emphasis id="B949D804FFDDFFB4FF322CD4D0D0FF60" bold="true" box="[151,392,223,250]" italics="true" pageId="9" pageNumber="470">Alluviobolus omega</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27A657FFFDDFFB4FE2A2CD4D0B4FF60" box="[399,492,223,249]" pageId="9" pageNumber="470" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C327579DFFDDFFB4FF322D08D1A1FE87" box="[151,249,259,286]" pageId="9" pageNumber="470" type="description">
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<paragraph id="8B820416FFDDFFB4FF322D08D1A1FE87" blockId="9.[151,492,223,286]" box="[151,249,259,286]" pageId="9" pageNumber="470">
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<figureCitation id="13061893FFDDFFB4FF322D08D1A1FE87" box="[151,249,259,286]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStart-2="FIGURE 7" captionStart-3="FIGURE 8" captionStart-4="FIGURE 9" captionStartId-0="9.[151,250,1692,1716]" captionStartId-1="10.[151,250,1533,1557]" captionStartId-2="11.[151,250,1380,1404]" captionStartId-3="12.[151,250,1388,1412]" captionStartId-4="13.[151,250,1222,1246]" captionTargetBox-0="[416,1172,849,1669]" captionTargetBox-1="[151,1435,188,1510]" captionTargetBox-2="[151,1435,273,1357]" captionTargetBox-3="[151,1437,237,1365]" captionTargetBox-4="[510,1077,182,1198]" captionTargetId-0="figure-246@9.[416,1172,849,1669]" captionTargetId-1="figure-18@10.[151,1435,181,1510]" captionTargetId-2="figure-54@11.[151,1435,273,1357]" captionTargetId-3="figure-47@12.[151,1437,237,1365]" captionTargetId-4="figure-15@13.[510,1077,181,1198]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="11" captionTargetPageId-3="12" captionTargetPageId-4="13" captionText-0="FIGURE 5. Alluviobolus omega, sp. nov., paratype what (FMMC 5403), multi-layer photographs. A. Male anterior body, lateral view; B. Male leg pairs 1–7, ventral view; C. Male posterior body end with telson; D. Anterior gonopod, anterior view; E. Anterior gonopod, posterior view; F. Posterior gonopod, anterior view; G. Posterior gonopod, posterior view. Abbreviations: a = spinose membranous area; Apo = apodeme; av = paraprocts; b= lateral opening of the sperm canal; bb = basal branch of posterior gonopod; c = membranous fringe below the apex; Co = collum (ring 1); Cx = coxite; d = large membranous area located mesally; Gn = gnathochilarium; h = ‘hole’-like structure on the posterior gonopod; Md = basal joints of mandible; oz = ozopore Pre = epiproct; sf = setiferous foveolae; sg = spermatic groove; St = sternite; sub = hypoproct; t = telopodite; numbers refer to leg pair number. Scale bars = 1 mm." captionText-1="FIGURE 6. Alluviobolus omega, sp. nov., holotype (FMMC 5458), drawings. A. Anterior gonopod, anterior view; B. Anterior gonopod, posterior view; C. Posterior gonopod, tip, anterior view; D. Posterior gonopod, tip, posterior view; E. Posterior gonopod, base, posterior view; F. Male midbody leg, posterior view; G. Female midbody leg, posterior view. Abbreviations: a = spinose membranous area; b= lateral opening of the sperm canal; bb = basal branch of posterior gonopod; c = membranous fringe below the apex; Cx = coxite; d = large membranous area located mesally; Fe = femur; fs-p = finger-shaped process; Gr = groove; h = ‘hole’-like structure on the posterior gonopod; Pre = prefemur; Po = postfemur; sg = spermatic groove; St = sternite; t = telopodite; Ta = tarsus; Ti = tibia. Scale bars = 1 mm." captionText-2="FIGURE 7. Alluviobolus omega, sp. nov., paratype (FMMC 5462), SEM. A. Left antenna, overview; B. Left antenna, antennomeres 5–7, apical view; C. Fine structures between antennomere 6 and 7. Abbreviations: A1–A7 = antennomeres 1–7; ac = apical cones; d = antennal disc; sb = sensilla basiconica." captionText-3="FIGURE 8. Alluviobolus omega, sp. nov., paratype (FMMC 5462), SEM. A. Gnathochilarium underside; B. Gnathochilarium, detail of central pads; C. Right mandible; D. Female midbody leg, overview; E. Female midbody leg, detail of tarsus; F. Right female vulva, mesal view. Abbreviations: 3iT = 3-combed inner tooth; as = apical spine; av = anterior orientated plate; cl = claw; cp = central pads; Cx = coxa; Endo = endochilarium; eP = lateral palpi; eT = external tooth; Fe = femur; iP = inner palpi; LT = lateral tooth; mp = molar plate; O = operculum; pL = pectinate lamellae; Po = postfemur; Pre = prefemur; pv = posterior orientated plate; vs = ventral spine; Ta = tarsus; Ti = tibia." captionText-4="FIGURE 9. Alluviobolus omega, sp. nov., population from Vohimena (FMMC 5396), multi-layer photographs.A. Male anterior body, lateral view; B. Midbody rings, lateral view; C. Male leg pairs 1–7, ventral view; D. Male posterior body end with telson; E. Anterior gonopod, anterior view; F. Anterior gonopod, posterior view; G. Right posterior gonopod, anterior view; H. Right posterior gonopod, posterior view. Abbreviations: a = spinose membranous area; av = paraprocts; b= lateral opening of the sperm canal; bb = basal branch of posterior gonopod; c = membranous fringe below the apex; Co = collum (ring 1); Cx = coxite; d = large membranous area located mesally; Gn = gnathochilarium; h = ‘hole’-like structure on the posterior gonopod; Md = basal joints of mandible; Me = mesozonite; Mt = metazonite; oz = ozopore; pe = pseudopenis; Pre = epiproct; sf = setiferous foveolae; sg = spermatic groove; St = sternite; sub = hypoproct; t = telopodite; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.14022762" figureDoi-1="http://doi.org/10.5281/zenodo.14022764" figureDoi-2="http://doi.org/10.5281/zenodo.14022768" figureDoi-3="http://doi.org/10.5281/zenodo.14022770" figureDoi-4="http://doi.org/10.5281/zenodo.14022774" httpUri-0="https://zenodo.org/record/14022762/files/figure.png" httpUri-1="https://zenodo.org/record/14022764/files/figure.png" httpUri-2="https://zenodo.org/record/14022768/files/figure.png" httpUri-3="https://zenodo.org/record/14022770/files/figure.png" httpUri-4="https://zenodo.org/record/14022774/files/figure.png" pageId="9" pageNumber="470">Figs 5–9</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C327579DFFDDFFB4FF322D40D3AEFD68" pageId="9" pageNumber="470" type="materials_examined">
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<paragraph id="8B820416FFDDFFB4FF322D40D44BFE10" blockId="9.[151,1437,331,826]" pageId="9" pageNumber="470">
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<emphasis id="B949D804FFDDFFB4FF322D40D022FEFC" bold="true" box="[151,378,331,357]" pageId="9" pageNumber="470">Material examined</emphasis>
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||
.
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<materialsCitation id="3B550E4BFFDDFFB4FE2F2D40D44BFE10" collectingDate="1996-01-10" collectingDateMax="1996-01-16" collectingDateMin="1996-01-10" collectorName="de Vohibasia & S. Goodman" country="Madagascar" elevation="780" latitude="-22.458334" location="Foret de Vohibasia" longLatPrecision="125" longitude="44.841667" municipality="Province" pageId="9" pageNumber="470" specimenCode="FMNH-INS 5458" specimenCount="1" stateProvince="Toliara" typeStatus="holotype">
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||
<typeStatus id="5486BAB4FFDDFFB4FE2F2D40D0AFFEFC" box="[394,503,331,357]" pageId="9" pageNumber="470" type="holotype">Holotype</typeStatus>
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||
: 1 M,
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||
<emphasis id="B949D804FFDDFFB4FD9A2D40D255FEFC" bold="true" box="[575,781,331,357]" pageId="9" pageNumber="470">
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||
<specimenCode id="DB9BAC6DFFDDFFB4FD9A2D40D255FEFC" box="[575,781,331,357]" collectionCode="FMNH-INS" pageId="9" pageNumber="470">FMNH-INS 5458</specimenCode>
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||
</emphasis>
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||
,
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||
<collectingCountry id="F32A4486FFDDFFB4FCB92D40D2FDFEFC" box="[796,933,331,357]" name="Madagascar" pageId="9" pageNumber="470">Madagascar</collectingCountry>
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,
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||
<collectingMunicipality id="6BE69E6CFFDDFFB4FC142D40D54DFEFC" box="[945,1045,331,357]" pageId="9" pageNumber="470">Province</collectingMunicipality>
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<collectingRegion id="49F9CAF4FFDDFFB4FBB92D40D536FEFC" box="[1052,1134,331,357]" country="Madagascar" name="Toliara" pageId="9" pageNumber="470">Toliara</collectingRegion>
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||
,
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||
<location id="8EE252CDFFDDFFB4FBDF2D40D415FEFC" LSID="urn:lsid:plazi:treatment:0394B500FFDDFFB3FF322CD4D249FD80:8EE252CDFFDDFFB4FBDF2D40D415FEFC" box="[1146,1357,331,357]" country="Madagascar" latitude="-22.458334" longLatPrecision="125" longitude="44.841667" municipality="Province" name="Foret de Vohibasia" pageId="9" pageNumber="470" stateProvince="Toliara">
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||
Forêt
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<collectorName id="26C861C0FFDDFFB4FB182D40D415FEFC" box="[1213,1357,331,357]" pageId="9" pageNumber="470">de Vohibasia</collectorName>
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</location>
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#3,
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<locationDeviation id="9A301D34FFDDFFB4FA252D40D000FE10" location="Sakaraha" pageId="9" pageNumber="470">
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<quantity id="4CC5A9F3FFDDFFB4FA252D40D1E3FE10" metricMagnitude="4" metricUnit="m" metricValue="5.9" pageId="9" pageNumber="470" unit="km" value="59.0">59 km</quantity>
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NE Sakaraha
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</locationDeviation>
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, tropical dry forest,
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<quantity id="4CC5A9F3FFDDFFB4FD9F2D64D3DBFE13" box="[570,643,367,394]" metricMagnitude="2" metricUnit="m" metricValue="7.8" pageId="9" pageNumber="470" unit="m" value="780.0">
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<elevation id="0010E325FFDDFFB4FD9F2D64D3DBFE13" box="[570,643,367,394]" metricMagnitude="2" metricUnit="m" metricValue="7.8" pageId="9" pageNumber="470" unit="m" value="780.0">780 m</elevation>
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</quantity>
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,
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<geoCoordinate id="EE0962D1FFDDFFB4FD2B2D64D25EFE10" box="[654,774,367,394]" degrees="22" direction="south" minutes="27.5" orientation="latitude" pageId="9" pageNumber="470" precision="92" value="-22.458334">22°27.5’ S</geoCoordinate>
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,
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<geoCoordinate id="EE0962D1FFDDFFB4FCB72D64D2C2FE13" box="[786,922,367,394]" degrees="044" direction="east" minutes="50.5" orientation="longitude" pageId="9" pageNumber="470" precision="92" value="44.841667">044°50.5’ E</geoCoordinate>
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, coll.
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<collectorName id="26C861C0FFDDFFB4FC7B2D64D534FE10" box="[990,1132,367,393]" pageId="9" pageNumber="470">S. Goodman</collectorName>
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,
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<date id="FF8322D6FFDDFFB4FBDD2D64D457FE10" box="[1144,1295,367,393]" pageId="9" pageNumber="470" value="1996-01-10" valueMax="1996-01-16" valueMin="1996-01-10">
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<collectingDate id="EFC7DB3EFFDDFFB4FBDD2D64D457FE10" box="[1144,1295,367,393]" pageId="9" pageNumber="470" value="1996-01-10" valueMax="1996-01-16" valueMin="1996-01-10">10–16.i.1996</collectingDate>
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||
</date>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8B820416FFDDFFB4FF622D9FD0A2FD80" blockId="9.[151,1437,331,826]" pageId="9" pageNumber="470">
|
||
<materialsCitation id="3B550E4BFFDDFFB4FF622D9FD293FE37" box="[199,971,403,430]" collectingDate="1996-01-10" collectingDateMax="1996-01-16" collectingDateMin="1996-01-10" collectorName="de Vohibasia & S. Goodman" country="Madagascar" elevation="780" latitude="-22.458334" location="Foret de Vohibasia" longLatPrecision="125" longitude="44.841667" municipality="Province" pageId="9" pageNumber="470" specimenCode="FMNH-INS 5458" specimenCount="1" stateProvince="Toliara" typeStatus="paratype">
|
||
<typeStatus id="5486BAB4FFDDFFB4FF622D9FD060FE37" box="[199,312,404,430]" pageId="9" pageNumber="470" type="paratype">Paratypes</typeStatus>
|
||
: 1 F,
|
||
<emphasis id="B949D804FFDDFFB4FEDC2D98D31FFE34" bold="true" box="[377,583,403,429]" pageId="9" pageNumber="470">
|
||
<specimenCode id="DB9BAC6DFFDDFFB4FEDC2D98D31FFE34" box="[377,583,403,429]" collectionCode="FMNH-INS" pageId="9" pageNumber="470">FMNH-INS 5458</specimenCode>
|
||
</emphasis>
|
||
, same data as holotype; 3 M, 1 F
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||
</materialsCitation>
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||
,
|
||
<materialsCitation id="3B550E4BFFDDFFB4FC7C2D98D40EFE48" collectingDate="1996-01-10" collectingDateMax="1996-01-16" collectingDateMin="1996-01-10" collectorName="S. Goodman" country="Madagascar" elevation="780" latitude="-22.458334" location="Foret de Vohibasia" longLatPrecision="125" longitude="44.841667" pageId="9" pageNumber="470" specimenCode="FMNH-INS 5462" specimenCount="1" stateProvince="Toliara" typeStatus="paratype">
|
||
<emphasis id="B949D804FFDDFFB4FC7C2D98D5FFFE34" bold="true" box="[985,1191,403,429]" pageId="9" pageNumber="470">
|
||
<specimenCode id="DB9BAC6DFFDDFFB4FC7C2D98D5FFFE34" box="[985,1191,403,429]" collectionCode="FMNH-INS" pageId="9" pageNumber="470">FMNH-INS 5462</specimenCode>
|
||
</emphasis>
|
||
,
|
||
<collectingRegion id="49F9CAF4FFDDFFB4FB122D98D4C0FE34" box="[1207,1432,403,429]" country="Madagascar" name="Toliara" pageId="9" pageNumber="470">Province de Toliara</collectingRegion>
|
||
,
|
||
<location id="8EE252CDFFDDFFB4FF322DBCD029FE48" LSID="urn:lsid:plazi:treatment:0394B500FFDDFFB3FF322CD4D249FD80:8EE252CDFFDDFFB4FF322DBCD029FE48" box="[151,369,439,465]" country="Madagascar" latitude="-22.458334" longLatPrecision="125" longitude="44.841667" name="Foret de Vohibasia" pageId="9" pageNumber="470" stateProvince="Toliara">Forêt de Vohibasia</location>
|
||
,
|
||
<locationDeviation id="9A301D34FFDDFFB4FEDA2DBCD335FE48" box="[383,621,439,466]" location="Sakaraha" pageId="9" pageNumber="470">
|
||
<quantity id="4CC5A9F3FFDDFFB4FEDA2DBCD091FE48" box="[383,457,439,466]" metricMagnitude="4" metricUnit="m" metricValue="5.9" pageId="9" pageNumber="470" unit="km" value="59.0">59 km</quantity>
|
||
NE Sakaraha
|
||
</locationDeviation>
|
||
,
|
||
<quantity id="4CC5A9F3FFDDFFB4FDDF2DBCD39EFE4B" box="[634,710,439,466]" metricMagnitude="2" metricUnit="m" metricValue="7.8" pageId="9" pageNumber="470" unit="m" value="780.0">
|
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<elevation id="0010E325FFDDFFB4FDDF2DBCD39EFE4B" box="[634,710,439,466]" metricMagnitude="2" metricUnit="m" metricValue="7.8" pageId="9" pageNumber="470" unit="m" value="780.0">780 m</elevation>
|
||
</quantity>
|
||
,
|
||
<geoCoordinate id="EE0962D1FFDDFFB4FD702DBCD208FE48" box="[725,848,439,466]" degrees="22" direction="south" minutes="27.5" orientation="latitude" pageId="9" pageNumber="470" precision="92" value="-22.458334">22°27.5’ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE0962D1FFDDFFB4FCC52DBCD284FE4B" box="[864,988,439,466]" degrees="44" direction="east" minutes="50.5" orientation="longitude" pageId="9" pageNumber="470" precision="92" value="44.841667">44°50.5’ E</geoCoordinate>
|
||
, leg.
|
||
<collectorName id="26C861C0FFDDFFB4FBBB2DBCD5E8FE48" box="[1054,1200,439,465]" pageId="9" pageNumber="470">S. Goodman</collectorName>
|
||
,
|
||
<date id="FF8322D6FFDDFFB4FB1A2DBCD40EFE48" box="[1215,1366,439,465]" pageId="9" pageNumber="470" value="1996-01-10" valueMax="1996-01-16" valueMin="1996-01-10">
|
||
<collectingDate id="EFC7DB3EFFDDFFB4FB1A2DBCD40EFE48" box="[1215,1366,439,465]" pageId="9" pageNumber="470" value="1996-01-10" valueMax="1996-01-16" valueMin="1996-01-10">10–16.i.1996</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
; 1 M,
|
||
<materialsCitation id="3B550E4BFFDDFFB4FF322DD0D0AEFD80" collectingDate="1996-01-10" collectorName="S. M. Goodman" country="Madagascar" elevation="780" latitude="-22.458334" location="Foret de Vohibasia" longLatPrecision="125" longitude="44.841667" pageId="9" pageNumber="470" specimenCode="FMNH-INS 5403" specimenCount="1" stateProvince="Toliara" typeStatus="paratype">
|
||
<emphasis id="B949D804FFDDFFB4FF322DD0D03DFE6C" bold="true" box="[151,357,475,501]" pageId="9" pageNumber="470">
|
||
<specimenCode id="DB9BAC6DFFDDFFB4FF322DD0D03DFE6C" box="[151,357,475,501]" collectionCode="FMNH-INS" pageId="9" pageNumber="470">FMNH-INS 5403</specimenCode>
|
||
</emphasis>
|
||
,
|
||
<collectingRegion id="49F9CAF4FFDDFFB4FED02DD0D30EFE6C" box="[373,598,475,501]" country="Madagascar" name="Toliara" pageId="9" pageNumber="470">Province de Toliara</collectingRegion>
|
||
,
|
||
<location id="8EE252CDFFDDFFB4FDC62DD0D262FE6C" LSID="urn:lsid:plazi:treatment:0394B500FFDDFFB3FF322CD4D249FD80:8EE252CDFFDDFFB4FDC62DD0D262FE6C" box="[611,826,475,501]" country="Madagascar" latitude="-22.458334" longLatPrecision="125" longitude="44.841667" name="Foret de Vohibasia" pageId="9" pageNumber="470" stateProvince="Toliara">Forêt de Vohibasia</location>
|
||
,
|
||
<locationDeviation id="9A301D34FFDDFFB4FCE22DD0D568FE6C" box="[839,1072,475,502]" location="Sakaraha" pageId="9" pageNumber="470">
|
||
<quantity id="4CC5A9F3FFDDFFB4FCE22DD0D2D7FE6C" box="[839,911,475,502]" metricMagnitude="4" metricUnit="m" metricValue="5.9" pageId="9" pageNumber="470" unit="km" value="59.0">59 km</quantity>
|
||
NE Sakaraha
|
||
</locationDeviation>
|
||
,
|
||
<quantity id="4CC5A9F3FFDDFFB4FB992DD0D5DFFE6F" box="[1084,1159,475,502]" metricMagnitude="2" metricUnit="m" metricValue="7.8" pageId="9" pageNumber="470" unit="m" value="780.0">
|
||
<elevation id="0010E325FFDDFFB4FB992DD0D5DFFE6F" box="[1084,1159,475,502]" metricMagnitude="2" metricUnit="m" metricValue="7.8" pageId="9" pageNumber="470" unit="m" value="780.0">780 m</elevation>
|
||
</quantity>
|
||
,
|
||
<geoCoordinate id="EE0962D1FFDDFFB4FB312DD0D456FE6C" box="[1172,1294,475,502]" degrees="22" direction="south" minutes="27.5" orientation="latitude" pageId="9" pageNumber="470" precision="92" value="-22.458334">22°27.5’ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE0962D1FFDDFFB4FAB92DD0D4CFFE6F" box="[1308,1431,475,502]" degrees="44" direction="east" minutes="50.5" orientation="longitude" pageId="9" pageNumber="470" precision="92" value="44.841667">44°50.5’ E</geoCoordinate>
|
||
, coll.
|
||
<collectorName id="26C861C0FFDDFFB4FF6A2DF4D026FD80" box="[207,382,511,537]" pageId="9" pageNumber="470">S.M. Goodman</collectorName>
|
||
,
|
||
<date id="FF8322D6FFDDFFB4FE2C2DF4D0AEFD80" box="[393,502,511,537]" pageId="9" pageNumber="470" value="1996-01-10">
|
||
<collectingDate id="EFC7DB3EFFDDFFB4FE2C2DF4D0AEFD80" box="[393,502,511,537]" pageId="9" pageNumber="470" value="1996-01-10">10.i.1996</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B820416FFDDFFB4FF622E28D3AEFD68" blockId="9.[151,1437,331,826]" pageId="9" pageNumber="470">
|
||
Other material: 1 M,
|
||
<materialsCitation id="3B550E4BFFDDFFB4FE652E28D2E1FDF8" collectingDate="1996-01-17" collectingDateMax="1996-01-24" collectingDateMin="1996-01-17" collectorName="S. M. Goodman" elevation="780" latitude="-22.683332" location="Foret de Vohimena" longLatPrecision="125" longitude="44.83" pageId="9" pageNumber="470" specimenCode="FMNH-INS 5396" specimenCount="1" stateProvince="Toliara">
|
||
<emphasis id="B949D804FFDDFFB4FE652E28D3C8FDA4" bold="true" box="[448,656,547,573]" pageId="9" pageNumber="470">
|
||
<specimenCode id="DB9BAC6DFFDDFFB4FE652E28D3C8FDA4" box="[448,656,547,573]" collectionCode="FMNH-INS" pageId="9" pageNumber="470">FMNH-INS 5396</specimenCode>
|
||
</emphasis>
|
||
,
|
||
<collectingRegion id="49F9CAF4FFDDFFB4FD052E28D2DCFDA4" box="[672,900,547,573]" country="Madagascar" name="Toliara" pageId="9" pageNumber="470">Province de Toliara</collectingRegion>
|
||
,
|
||
<location id="8EE252CDFFDDFFB4FC372E28D536FDA4" LSID="urn:lsid:plazi:treatment:0394B500FFDDFFB3FF322CD4D249FD80:8EE252CDFFDDFFB4FC372E28D536FDA4" box="[914,1134,547,573]" latitude="-22.683332" longLatPrecision="125" longitude="44.83" name="Foret de Vohimena" pageId="9" pageNumber="470" stateProvince="Toliara">Forêt de Vohimena</location>
|
||
,
|
||
<locationDeviation id="9A301D34FFDDFFB4FBD92E28D43DFDA4" box="[1148,1381,547,574]" location="Sakahara" pageId="9" pageNumber="470">
|
||
<quantity id="4CC5A9F3FFDDFFB4FBD92E28D59EFDA4" box="[1148,1222,547,574]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="9" pageNumber="470" unit="km" value="35.0">35 km</quantity>
|
||
SE Sakahara
|
||
</locationDeviation>
|
||
,
|
||
<quantity id="4CC5A9F3FFDDFFB4FAD72E28D1F7FDFB" metricMagnitude="2" metricUnit="m" metricValue="7.8" pageId="9" pageNumber="470" unit="m" value="780.0">
|
||
<elevation id="0010E325FFDDFFB4FAD72E28D1F7FDFB" metricMagnitude="2" metricUnit="m" metricValue="7.8" pageId="9" pageNumber="470" unit="m" value="780.0">780 m</elevation>
|
||
</quantity>
|
||
,
|
||
<geoCoordinate id="EE0962D1FFDDFFB4FF182E4CD06EFDF8" box="[189,310,583,609]" degrees="22" direction="south" minutes="41.0" orientation="latitude" pageId="9" pageNumber="470" precision="92" value="-22.683332">22°41.0’ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE0962D1FFDDFFB4FEE12E4CD098FDFB" box="[324,448,583,610]" degrees="44" direction="east" minutes="49.8" orientation="longitude" pageId="9" pageNumber="470" precision="92" value="44.83">44°49.8’ E</geoCoordinate>
|
||
, coll.
|
||
<collectorName id="26C861C0FFDDFFB4FDA22E4CD3E0FDF8" box="[519,696,583,609]" pageId="9" pageNumber="470">S.M. Goodman</collectorName>
|
||
,
|
||
<collectingMethod id="527C7C01FFDDFFB4FD602E4CD24EFDF8" box="[709,790,583,609]" pageId="9" pageNumber="470">pitfalls</collectingMethod>
|
||
,
|
||
<date id="FF8322D6FFDDFFB4FC872E4CD2E1FDF8" box="[802,953,583,609]" pageId="9" pageNumber="470" value="1996-01-17" valueMax="1996-01-24" valueMin="1996-01-17">
|
||
<collectingDate id="EFC7DB3EFFDDFFB4FC872E4CD2E1FDF8" box="[802,953,583,609]" pageId="9" pageNumber="470" value="1996-01-17" valueMax="1996-01-24" valueMin="1996-01-17">17–24.i.1996</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
; 10 M, 2 F,
|
||
<materialsCitation id="3B550E4BFFDDFFB4FBED2E4CD2CAFD33" collectingDate="2003-02-01" collectingDateMax="2003-02-05" collectingDateMin="2003-02-01" collectionCode="CAS" collectorName="Fisher & Griswold" country="Madagascar" elevation="700" latitude="-22.591667" location="Foret d'Analalava" longLatPrecision="21" longitude="45.12833" municipality="Province" pageId="9" pageNumber="470" specimenCode="BLF 7381" specimenCount="1" stateProvince="Fianarantsoa">
|
||
<emphasis id="B949D804FFDDFFB4FBED2E4CD5A7FDF8" bold="true" box="[1096,1279,583,609]" pageId="9" pageNumber="470">
|
||
<collectionCode id="ED2C9CD3FFDDFFB4FBED2E4CD5D8FDF8" box="[1096,1152,583,609]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="9" pageNumber="470" type="Museum">CAS</collectionCode>
|
||
<specimenCode id="DB9BAC6DFFDDFFB4FB2C2E4CD5A7FDF8" box="[1161,1279,583,609]" collectionCode="BLF" pageId="9" pageNumber="470">BLF 7381</specimenCode>
|
||
</emphasis>
|
||
,
|
||
<collectingCountry id="F32A4486FFDDFFB4FAAA2E4CD4C0FDF8" box="[1295,1432,583,609]" name="Madagascar" pageId="9" pageNumber="470">Madagascar</collectingCountry>
|
||
,
|
||
<collectingMunicipality id="6BE69E6CFFDDFFB4FF322E60D1A3FD1C" box="[151,251,619,645]" pageId="9" pageNumber="470">Province</collectingMunicipality>
|
||
<collectingRegion id="49F9CAF4FFDDFFB4FEA02E60D0CFFD1C" box="[261,407,619,645]" country="Madagascar" name="Fianarantsoa" pageId="9" pageNumber="470">Fianarantsoa</collectingRegion>
|
||
,
|
||
<location id="8EE252CDFFDDFFB4FE032E60D32FFD1C" LSID="urn:lsid:plazi:treatment:0394B500FFDDFFB3FF322CD4D249FD80:8EE252CDFFDDFFB4FE032E60D32FFD1C" box="[422,631,619,645]" country="Madagascar" latitude="-22.591667" longLatPrecision="21" longitude="45.12833" municipality="Province" name="Foret d'Analalava" pageId="9" pageNumber="470" stateProvince="Fianarantsoa">Forêt d’Analalava</location>
|
||
,
|
||
<locationDeviation id="9A301D34FFDDFFB4FD202E60D2E4FD1C" box="[645,956,619,646]" location="Ranohira" pageId="9" pageNumber="470">
|
||
<quantity id="4CC5A9F3FFDDFFB4FD202E60D3BDFD1C" box="[645,741,619,645]" metricMagnitude="4" metricUnit="m" metricValue="2.96" pageId="9" pageNumber="470" unit="km" value="29.6">29.6 km</quantity>
|
||
280° W Ranohira
|
||
</locationDeviation>
|
||
, tropical dry forest,
|
||
<quantity id="4CC5A9F3FFDDFFB4FB0E2E60D5AFFD1F" box="[1195,1271,619,646]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="9" pageNumber="470" unit="m" value="700.0">
|
||
<elevation id="0010E325FFDDFFB4FB0E2E60D5AFFD1F" box="[1195,1271,619,646]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="9" pageNumber="470" unit="m" value="700.0">700 m</elevation>
|
||
</quantity>
|
||
,
|
||
<geoCoordinate id="EE0962D1FFDDFFB4FAA32E60D4CFFD1C" box="[1286,1431,619,646]" degrees="22" direction="south" minutes="35" orientation="latitude" pageId="9" pageNumber="470" precision="15" seconds="30" value="-22.591667">22°35’30” S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE0962D1FFDDFFB4FF322E84D06DFD33" box="[151,309,655,682]" degrees="045" direction="east" minutes="07" orientation="longitude" pageId="9" pageNumber="470" precision="15" seconds="42" value="45.12833">045°07’42” E</geoCoordinate>
|
||
, coll.
|
||
<collectorName id="26C861C0FFDDFFB4FED92E84D09CFD30" box="[380,452,655,681]" pageId="9" pageNumber="470">Fisher</collectorName>
|
||
,
|
||
<collectorName id="26C861C0FFDDFFB4FE752E84D360FD30" box="[464,568,655,681]" pageId="9" pageNumber="470">Griswold</collectorName>
|
||
<emphasis id="B949D804FFDDFFB4FDE42E9BD323FD30" box="[577,635,655,681]" italics="true" pageId="9" pageNumber="470">et al.</emphasis>
|
||
,
|
||
<collectingMethod id="527C7C01FFDDFFB4FD2F2E84D25BFD33" box="[650,771,655,682]" pageId="9" pageNumber="470">pitfall trap</collectingMethod>
|
||
,
|
||
<date id="FF8322D6FFDDFFB4FCB52E84D2CAFD33" box="[784,914,655,682]" pageId="9" pageNumber="470" value="2003-02-01" valueMax="2003-02-05" valueMin="2003-02-01">
|
||
<collectingDate id="EFC7DB3EFFDDFFB4FCB52E84D2CAFD33" box="[784,914,655,682]" pageId="9" pageNumber="470" value="2003-02-01" valueMax="2003-02-05" valueMin="2003-02-01">1–5.ii.2003</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
; 1 M,
|
||
<materialsCitation id="3B550E4BFFDDFFB4FC7B2E84D3AAFD68" collectingDate="2003-01-27" collectingDateMax="2003-01-31" collectingDateMin="2003-01-27" collectionCode="CAS" collectorName="Fisher & Griswold" country="Madagascar" elevation="725" latitude="-22.481667" location="Parc National d'Isalo" longLatPrecision="21" longitude="45.461666" municipality="Province" pageId="9" pageNumber="470" specimenCode="BLF 7306" specimenCount="1" stateProvince="Fianarantsoa">
|
||
<emphasis id="B949D804FFDDFFB4FC7B2E84D5CCFD30" bold="true" box="[990,1172,655,681]" pageId="9" pageNumber="470">
|
||
<collectionCode id="ED2C9CD3FFDDFFB4FC7B2E84D54EFD30" box="[990,1046,655,681]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="9" pageNumber="470" type="Museum">CAS</collectionCode>
|
||
<specimenCode id="DB9BAC6DFFDDFFB4FBBB2E84D5CCFD30" box="[1054,1172,655,681]" collectionCode="BLF" pageId="9" pageNumber="470">BLF 7306</specimenCode>
|
||
</emphasis>
|
||
,
|
||
<collectingCountry id="F32A4486FFDDFFB4FB062E84D474FD30" box="[1187,1324,655,681]" name="Madagascar" pageId="9" pageNumber="470">Madagascar</collectingCountry>
|
||
,
|
||
<collectingMunicipality id="6BE69E6CFFDDFFB4FA9C2E84D4C5FD30" box="[1337,1437,655,681]" pageId="9" pageNumber="470">Province</collectingMunicipality>
|
||
<collectingRegion id="49F9CAF4FFDDFFB4FF322EB8D071FD54" box="[151,297,691,717]" country="Madagascar" name="Fianarantsoa" pageId="9" pageNumber="470">Fianarantsoa</collectingRegion>
|
||
,
|
||
<location id="8EE252CDFFDDFFB4FE932EBFD373FD54" LSID="urn:lsid:plazi:treatment:0394B500FFDDFFB3FF322CD4D249FD80:8EE252CDFFDDFFB4FE932EBFD373FD54" box="[310,555,691,718]" country="Madagascar" latitude="-22.481667" longLatPrecision="21" longitude="45.461666" municipality="Province" name="Parc National d'Isalo" pageId="9" pageNumber="470" stateProvince="Fianarantsoa">Parc National d’Isalo</location>
|
||
,
|
||
<locationDeviation id="9A301D34FFDDFFB4FD9C2EB8D216FD54" box="[569,846,691,718]" location="Ranohira" pageId="9" pageNumber="470">
|
||
<quantity id="4CC5A9F3FFDDFFB4FD9C2EB8D3D1FD54" box="[569,649,691,717]" metricMagnitude="3" metricUnit="m" metricValue="9.1" pageId="9" pageNumber="470" unit="km" value="9.1">9.1 km</quantity>
|
||
354°N Ranohira
|
||
</locationDeviation>
|
||
, gallery forest,
|
||
<quantity id="4CC5A9F3FFDDFFB4FBA42EB8D515FD57" box="[1025,1101,691,718]" metricMagnitude="2" metricUnit="m" metricValue="7.25" pageId="9" pageNumber="470" unit="m" value="725.0">
|
||
<elevation id="0010E325FFDDFFB4FBA42EB8D515FD57" box="[1025,1101,691,718]" metricMagnitude="2" metricUnit="m" metricValue="7.25" pageId="9" pageNumber="470" unit="m" value="725.0">725 m</elevation>
|
||
</quantity>
|
||
,
|
||
<geoCoordinate id="EE0962D1FFDDFFB4FBFF2EB8D5B1FD54" box="[1114,1257,691,718]" degrees="22" direction="south" minutes="28" orientation="latitude" pageId="9" pageNumber="470" precision="15" seconds="54" value="-22.481667">22°28’54” S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE0962D1FFDDFFB4FB5D2EB8D4CFFD57" box="[1272,1431,691,718]" degrees="045" direction="east" minutes="27" orientation="longitude" pageId="9" pageNumber="470" precision="15" seconds="42" value="45.461666">045°27’42” E</geoCoordinate>
|
||
, coll.
|
||
<collectorName id="26C861C0FFDDFFB4FF6A2EDCD04FFD68" box="[207,279,727,753]" pageId="9" pageNumber="470">Fisher</collectorName>
|
||
,
|
||
<collectorName id="26C861C0FFDDFFB4FE872EDCD0D2FD68" box="[290,394,727,753]" pageId="9" pageNumber="470">Griswold</collectorName>
|
||
<emphasis id="B949D804FFDDFFB4FE342ED3D092FD68" box="[401,458,727,753]" italics="true" pageId="9" pageNumber="470">et al.</emphasis>
|
||
,
|
||
<collectingMethod id="527C7C01FFDDFFB4FE722EDCD317FD6B" box="[471,591,727,754]" pageId="9" pageNumber="470">pitfall trap</collectingMethod>
|
||
,
|
||
<date id="FF8322D6FFDDFFB4FDFE2EDCD3AAFD68" box="[603,754,727,753]" pageId="9" pageNumber="470" value="2003-01-27" valueMax="2003-01-31" valueMin="2003-01-27">
|
||
<collectingDate id="EFC7DB3EFFDDFFB4FDFE2EDCD3AAFD68" box="[603,754,727,753]" pageId="9" pageNumber="470" value="2003-01-27" valueMax="2003-01-31" valueMin="2003-01-27">27–31.i.2003</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C327579DFFDDFFB4FF622EF0D27FFCA0" pageId="9" pageNumber="470" type="etymology">
|
||
<paragraph id="8B820416FFDDFFB4FF622EF0D27FFCA0" blockId="9.[151,1437,331,826]" pageId="9" pageNumber="470">
|
||
<emphasis id="B949D804FFDDFFB4FF622EF0D0C3FC8C" bold="true" box="[199,411,763,789]" pageId="9" pageNumber="470">Derivatio nominis</emphasis>
|
||
. omega, noun in apposition, after the last letter of the Greek alphabet.The posterior gonopods resemble the letter “Z”, the last letter in our Lain alphabet.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF42549EFFDDFFB4FF322A97D306F829" ID-DOI="http://doi.org/10.5281/zenodo.14022762" ID-Zenodo-Dep="14022762" httpUri="https://zenodo.org/record/14022762/files/figure.png" pageId="9" pageNumber="470" startId="9.[151,250,1692,1716]" targetBox="[416,1172,849,1669]" targetPageId="9" targetType="figure">
|
||
<paragraph id="8B820416FFDDFFB4FF322A97D306F829" blockId="9.[151,1437,1692,1969]" pageId="9" pageNumber="470">
|
||
<emphasis id="B949D804FFDDFFB4FF322A97D04EF92C" bold="true" box="[151,278,1692,1717]" pageId="9" pageNumber="470">FIGURE 5.</emphasis>
|
||
<taxonomicName id="4C3D7F95FFDDFFB4FEBA2A96D0B4F92D" authorityName="Wesener" authorityYear="2024" box="[287,492,1693,1717]" class="Diplopoda" family="Pachybolidae" genus="Alluviobolus" kingdom="Animalia" order="Spirobolida" pageId="9" pageNumber="470" phylum="Arthropoda" rank="species" species="omega" status="sp. nov.">
|
||
<emphasis id="B949D804FFDDFFB4FEBA2A96D0B4F92D" box="[287,492,1693,1717]" italics="true" pageId="9" pageNumber="470">Alluviobolus omega</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B949D804FFDDFFB4FE5E2A96D317F92C" bold="true" box="[507,591,1693,1717]" pageId="9" pageNumber="470">
|
||
<taxonomicNameLabel id="A27A657FFFDDFFB4FE5E2A96D317F92C" box="[507,591,1693,1717]" pageId="9" pageNumber="470" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, paratype what (
|
||
<emphasis id="B949D804FFDDFFB4FCA72A97D2C8F92C" bold="true" box="[770,912,1692,1717]" pageId="9" pageNumber="470">FMMC 5403</emphasis>
|
||
), multi-layer photographs.
|
||
<emphasis id="B949D804FFDDFFB4FB0B2A97D598F92D" bold="true" box="[1198,1216,1692,1716]" pageId="9" pageNumber="470">A</emphasis>
|
||
. Male anterior body, lateral view;
|
||
<emphasis id="B949D804FFDDFFB4FEBC2ACAD072F940" bold="true" box="[281,298,1729,1753]" pageId="9" pageNumber="470">B</emphasis>
|
||
. Male leg pairs 1–7, ventral view;
|
||
<emphasis id="B949D804FFDDFFB4FD212ACBD3CEF941" bold="true" box="[644,662,1728,1752]" pageId="9" pageNumber="470">C</emphasis>
|
||
. Male posterior body end with telson;
|
||
<emphasis id="B949D804FFDDFFB4FBBD2ACAD572F940" bold="true" box="[1048,1066,1729,1753]" pageId="9" pageNumber="470">D</emphasis>
|
||
. Anterior gonopod, anterior view;
|
||
<emphasis id="B949D804FFDDFFB4FA202ACAD4CEF940" bold="true" box="[1413,1430,1729,1753]" pageId="9" pageNumber="470">E</emphasis>
|
||
. Anterior gonopod, posterior view;
|
||
<emphasis id="B949D804FFDDFFB4FE5F2AEED351F964" bold="true" box="[506,521,1765,1789]" pageId="9" pageNumber="470">F</emphasis>
|
||
. Posterior gonopod, anterior view;
|
||
<emphasis id="B949D804FFDDFFB4FCD02AEFD2D1F965" bold="true" box="[885,905,1764,1788]" pageId="9" pageNumber="470">G</emphasis>
|
||
. Posterior gonopod, posterior view.
|
||
<emphasis id="B949D804FFDDFFB4FB592AEFD4CDF965" bold="true" box="[1276,1429,1764,1788]" pageId="9" pageNumber="470">Abbreviations</emphasis>
|
||
: a = spinose membranous area; Apo = apodeme; av = paraprocts; b= lateral opening of the sperm canal; bb = basal branch of posterior gonopod; c = membranous fringe below the apex; Co = collum (ring 1); Cx = coxite; d = large membranous area located mesally; Gn = gnathochilarium; h = ‘hole’-like structure on the posterior gonopod; Md = basal joints of mandible; oz = ozopore Pre = epiproct; sf = setiferous foveolae; sg = spermatic groove; St = sternite; sub = hypoproct; t = telopodite; numbers refer to leg pair number.
|
||
<emphasis id="B949D804FFDDFFB4FE362B93D35AF828" bold="true" box="[403,514,1944,1969]" pageId="9" pageNumber="470">Scale bars</emphasis>
|
||
= 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF42549EFFDEFFB7FF3229F6D385F974" ID-DOI="http://doi.org/10.5281/zenodo.14022764" ID-Zenodo-Dep="14022764" httpUri="https://zenodo.org/record/14022764/files/figure.png" pageId="10" pageNumber="471" startId="10.[151,250,1533,1557]" targetBox="[151,1435,188,1510]" targetPageId="10" targetType="figure">
|
||
<paragraph id="8B820416FFDEFFB7FF3229F6D385F974" blockId="10.[151,1436,1533,1774]" pageId="10" pageNumber="471">
|
||
<emphasis id="B949D804FFDEFFB7FF3229F6D04AF98C" bold="true" box="[151,274,1533,1557]" pageId="10" pageNumber="471">FIGURE 6.</emphasis>
|
||
<taxonomicName id="4C3D7F95FFDEFFB7FEBC29F5D0BAF98C" authorityName="Wesener" authorityYear="2024" box="[281,482,1534,1558]" class="Diplopoda" family="Pachybolidae" genus="Alluviobolus" kingdom="Animalia" order="Spirobolida" pageId="10" pageNumber="471" phylum="Arthropoda" rank="species" species="omega" status="sp. nov.">
|
||
<emphasis id="B949D804FFDEFFB7FEBC29F5D0BAF98C" box="[281,482,1534,1558]" italics="true" pageId="10" pageNumber="471">Alluviobolus omega</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B949D804FFDEFFB7FE4B29F5D366F98F" bold="true" box="[494,574,1534,1558]" pageId="10" pageNumber="471">
|
||
<taxonomicNameLabel id="A27A657FFFDEFFB7FE4B29F5D366F98F" box="[494,574,1534,1558]" pageId="10" pageNumber="471" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype (
|
||
<emphasis id="B949D804FFDEFFB7FD1729F6D265F98C" bold="true" box="[690,829,1533,1557]" pageId="10" pageNumber="471">FMMC 5458</emphasis>
|
||
), drawings.
|
||
<emphasis id="B949D804FFDEFFB7FC1F29F6D294F98C" bold="true" box="[954,972,1533,1557]" pageId="10" pageNumber="471">A</emphasis>
|
||
. Anterior gonopod, anterior view;
|
||
<emphasis id="B949D804FFDEFFB7FA8E29F5D464F98F" bold="true" box="[1323,1340,1534,1558]" pageId="10" pageNumber="471">B</emphasis>
|
||
. Anterior gonopod, posterior view;
|
||
<emphasis id="B949D804FFDEFFB7FE032A2AD0E0F9A0" bold="true" box="[422,440,1569,1593]" pageId="10" pageNumber="471">C</emphasis>
|
||
. Posterior gonopod, tip, anterior view;
|
||
<emphasis id="B949D804FFDEFFB7FCF82A29D237F9A3" bold="true" box="[861,879,1570,1594]" pageId="10" pageNumber="471">D</emphasis>
|
||
. Posterior gonopod, tip, posterior view;
|
||
<emphasis id="B949D804FFDEFFB7FA852A29D469F9A3" bold="true" box="[1312,1329,1570,1594]" pageId="10" pageNumber="471">E</emphasis>
|
||
. Posterior gonopod, base, posterior view;
|
||
<emphasis id="B949D804FFDEFFB7FE7C2A4DD0B4F9C7" bold="true" box="[473,492,1606,1630]" pageId="10" pageNumber="471">F.</emphasis>
|
||
Male midbody leg, posterior view;
|
||
<emphasis id="B949D804FFDEFFB7FCF82A4ED229F9C4" bold="true" box="[861,881,1605,1629]" pageId="10" pageNumber="471">G</emphasis>
|
||
. Female midbody leg, posterior view.
|
||
<emphasis id="B949D804FFDEFFB7FB592A4ED4CDF9C4" bold="true" box="[1276,1429,1605,1629]" pageId="10" pageNumber="471">Abbreviations</emphasis>
|
||
: a = spinose membranous area; b= lateral opening of the sperm canal; bb = basal branch of posterior gonopod; c = membranous fringe below the apex; Cx = coxite; d = large membranous area located mesally; Fe = femur; fs-p = finger-shaped process; Gr = groove; h = ‘hole’-like structure on the posterior gonopod; Pre = prefemur; Po = postfemur; sg = spermatic groove; St = sternite; t = telopodite; Ta = tarsus; Ti = tibia.
|
||
<emphasis id="B949D804FFDEFFB7FDB72ADED3D8F977" bold="true" box="[530,640,1749,1774]" pageId="10" pageNumber="471">Scale bars</emphasis>
|
||
= 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C327579DFFDEFFB6FF622B4BD349FF60" lastPageId="11" lastPageNumber="472" pageId="10" pageNumber="471" type="diagnosis">
|
||
<paragraph id="8B820416FFDEFFB6FF622B4BD349FF60" blockId="10.[151,1437,1856,2026]" lastBlockId="11.[151,1436,151,249]" lastPageId="11" lastPageNumber="472" pageId="10" pageNumber="471">
|
||
<emphasis id="B949D804FFDEFFB7FF622B4BD062F8C3" bold="true" box="[199,314,1856,1882]" pageId="10" pageNumber="471">Diagnosis</emphasis>
|
||
. The gnathochilarium with the division of the mentum and the sclerotized ledges on the stipites and mentum identify this species a member of a group uniting several genera of small-bodied Malagasy
|
||
<taxonomicName id="4C3D7F95FFDEFFB7FAA02B6FD4C4F8E7" authorityName="Cook, 1897 sensu Hoffman" authorityYear="1980" box="[1285,1436,1892,1918]" class="Diplopoda" family="Pachybolidae" kingdom="Animalia" order="Spirobolida" pageId="10" pageNumber="471" phylum="Arthropoda" rank="family">Pachybolidae</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EFAC79E7FFDEFFB7FF6B2B83D090F83B" author="Wesener T. & Enghoff H. & Sierwald P." box="[206,456,1928,1954]" pageId="10" pageNumber="471" pagination="1 - 128" refId="ref15399" refString="Wesener T., Enghoff H., Sierwald P. (2009 a) Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda). ZooKeys, 19, 1 - 128. https: // doi. org / 10.3897 / zookeys. 19.221" type="journal article" year="2009">
|
||
Wesener
|
||
<emphasis id="B949D804FFDEFFB7FE9D2B82D02CF83B" box="[312,372,1928,1954]" italics="true" pageId="10" pageNumber="471">et al.</emphasis>
|
||
2009a
|
||
</bibRefCitation>
|
||
). This species is placed in the genus
|
||
<taxonomicName id="4C3D7F95FFDEFFB7FCDA2B83D555F83B" authorityName="Wesener" authorityYear="2009" box="[895,1037,1928,1954]" class="Diplopoda" family="Pachybolidae" genus="Alluviobolus" kingdom="Animalia" order="Spirobolida" pageId="10" pageNumber="471" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B949D804FFDEFFB7FCDA2B83D555F83B" box="[895,1037,1928,1954]" italics="true" pageId="10" pageNumber="471">Alluviobolus</emphasis>
|
||
</taxonomicName>
|
||
because of the posterior telopods, in which the coxite and telopodite are clearly separated, the telopodite with a torsion so that the opening of the spermatic groove (or sperm channel) discharges apico-laterally.
|
||
<taxonomicName id="4C3D7F95FFDEFFB7FCCB2BDBD281F873" authorityName="Wesener" authorityYear="2024" box="[878,985,2000,2026]" class="Diplopoda" family="Pachybolidae" genus="Alluviobolus" kingdom="Animalia" order="Spirobolida" pageId="10" pageNumber="471" phylum="Arthropoda" rank="species" species="omega" status="sp. nov.">
|
||
<emphasis id="B949D804FFDEFFB7FCCB2BDBD281F873" box="[878,985,2000,2026]" italics="true" pageId="10" pageNumber="471">A. omega</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B949D804FFDEFFB7FC442BDBD564F873" bold="true" box="[993,1084,2000,2026]" pageId="10" pageNumber="471">
|
||
<taxonomicNameLabel id="A27A657FFFDEFFB7FC442BDBD564F873" box="[993,1084,2000,2026]" pageId="10" pageNumber="471" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs strongly from the other three
|
||
<taxonomicName id="4C3D7F95FFDFFFB6FF702C9CD0E1FF28" box="[213,441,151,177]" pageId="11" pageNumber="472">
|
||
<emphasis id="B949D804FFDFFFB6FF702C9CD03BFF28" box="[213,355,151,177]" italics="true" pageId="11" pageNumber="472">Alluviobolus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
distributed in the SE of
|
||
<collectingCountry id="F32A4486FFDFFFB6FD612C9CD208FF28" box="[708,848,151,177]" name="Madagascar" pageId="11" pageNumber="472">Madagascar</collectingCountry>
|
||
: the anterior telopod has a weaker telopodite process, the posterior telopod is unique, almost resembling the letter ‘Z’ with a hole in anterior view and numerous large membranous folds apico-mesally.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF42549EFFDFFFB6FF32296FD25BFA5D" ID-DOI="http://doi.org/10.5281/zenodo.14022768" ID-Zenodo-Dep="14022768" httpUri="https://zenodo.org/record/14022768/files/figure.png" pageId="11" pageNumber="472" startId="11.[151,250,1380,1404]" targetBox="[151,1435,273,1357]" targetPageId="11" targetType="figure">
|
||
<paragraph id="8B820416FFDFFFB6FF32296FD25BFA5D" blockId="11.[151,1437,1380,1476]" pageId="11" pageNumber="472">
|
||
<emphasis id="B949D804FFDFFFB6FF32296FD040FAE5" bold="true" box="[151,280,1380,1404]" pageId="11" pageNumber="472">FIGURE 7.</emphasis>
|
||
<taxonomicName id="4C3D7F95FFDFFFB6FE86296FD0AAFAE5" authorityName="Wesener" authorityYear="2024" box="[291,498,1380,1404]" class="Diplopoda" family="Pachybolidae" genus="Alluviobolus" kingdom="Animalia" order="Spirobolida" pageId="11" pageNumber="472" phylum="Arthropoda" rank="species" species="omega" status="sp. nov.">
|
||
<emphasis id="B949D804FFDFFFB6FE86296FD0AAFAE5" box="[291,498,1380,1404]" italics="true" pageId="11" pageNumber="472">Alluviobolus omega</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B949D804FFDFFFB6FDA1296FD301FAE5" bold="true" box="[516,601,1380,1404]" pageId="11" pageNumber="472">
|
||
<taxonomicNameLabel id="A27A657FFFDFFFB6FDA1296FD301FAE5" box="[516,601,1380,1404]" pageId="11" pageNumber="472" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, paratype (
|
||
<emphasis id="B949D804FFDFFFB6FD73296FD23EFAE5" bold="true" box="[726,870,1380,1404]" pageId="11" pageNumber="472">FMMC 5462</emphasis>
|
||
), SEM.
|
||
<emphasis id="B949D804FFDFFFB6FC60296FD28FFAE5" bold="true" box="[965,983,1380,1404]" pageId="11" pageNumber="472">A</emphasis>
|
||
. Left antenna, overview;
|
||
<emphasis id="B949D804FFDFFFB6FB4A296FD458FAE5" bold="true" box="[1263,1280,1380,1404]" pageId="11" pageNumber="472">B</emphasis>
|
||
. Left antenna, antennomeres 5–7, apical view;
|
||
<emphasis id="B949D804FFDFFFB6FE7B2983D0A8FA39" bold="true" box="[478,496,1416,1440]" pageId="11" pageNumber="472">C</emphasis>
|
||
. Fine structures between antennomere 6 and 7.
|
||
<emphasis id="B949D804FFDFFFB6FC772983D533FA39" bold="true" box="[978,1131,1416,1440]" pageId="11" pageNumber="472">Abbreviations</emphasis>
|
||
: A1–A7 = antennomeres 1–7; ac = apical cones; d = antennal disc; sb = sensilla basiconica.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C327579DFFDFFFB3FF322A1DD4DCFF60" lastPageId="14" lastPageNumber="475" pageId="11" pageNumber="472" type="description">
|
||
<paragraph id="8B820416FFDFFFB6FF322A1DD070F9A9" blockId="11.[151,296,1558,1584]" box="[151,296,1558,1584]" pageId="11" pageNumber="472">
|
||
<emphasis id="B949D804FFDFFFB6FF322A1DD070F9A9" bold="true" box="[151,296,1558,1584]" pageId="11" pageNumber="472">Description</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B820416FFDFFFB6FF322A53D520F923" blockId="11.[151,1437,1624,2010]" pageId="11" pageNumber="472">
|
||
Measurements:
|
||
<typeStatus id="5486BAB4FFDFFFB6FEEE2A53D0F5F9EB" box="[331,429,1624,1650]" pageId="11" pageNumber="472" type="holotype">holotype</typeStatus>
|
||
male with 45+0 body rings, ca.
|
||
<quantity id="4CC5A9F3FFDFFFB6FC812A53D22DF9EB" box="[804,885,1624,1650]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" pageId="11" pageNumber="472" unit="mm" value="36.0">36 mm</quantity>
|
||
long, midbody rings
|
||
<quantity id="4CC5A9F3FFDFFFB6FBC92A53D59CF9EB" box="[1132,1220,1624,1650]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="11" pageNumber="472" unit="mm" value="3.1">3.1 mm</quantity>
|
||
wide. Other males with 45 or 46+0 body rings, length up to
|
||
<quantity id="4CC5A9F3FFDFFFB6FDC02A77D3EEF90F" box="[613,694,1660,1686]" metricMagnitude="-2" metricUnit="m" metricValue="3.9" pageId="11" pageNumber="472" unit="mm" value="39.0">39 mm</quantity>
|
||
, midbody ring width up to
|
||
<quantity id="4CC5A9F3FFDFFFB6FC422A77D518F90F" box="[999,1088,1660,1686]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="11" pageNumber="472" unit="mm" value="3.4">3.4 mm</quantity>
|
||
. Females larger, large
|
||
<typeStatus id="5486BAB4FFDFFFB6FA992A77D4C4F90F" box="[1340,1436,1660,1686]" pageId="11" pageNumber="472" type="paratype">paratype</typeStatus>
|
||
female with eggs ca.
|
||
<quantity id="4CC5A9F3FFDFFFB6FE212AABD08BF923" box="[388,467,1696,1723]" metricMagnitude="-2" metricUnit="m" metricValue="5.6" pageId="11" pageNumber="472" unit="mm" value="56.0">56 mm</quantity>
|
||
long, at midbody rings
|
||
<quantity id="4CC5A9F3FFDFFFB6FD7B2AABD26CF923" box="[734,820,1696,1723]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="11" pageNumber="472" unit="mm" value="5.1">5.1 mm</quantity>
|
||
wide, with 46+0 body rings.
|
||
</paragraph>
|
||
<paragraph id="8B820416FFDFFFB6FF622ACFD20CF89B" blockId="11.[151,1437,1624,2010]" pageId="11" pageNumber="472">
|
||
Colour: Head and collum black, only on margins red (
|
||
<figureCitation id="13061893FFDFFFB6FC8E2ACFD2DDF946" box="[811,901,1732,1759]" captionStart="FIGURE 5" captionStartId="9.[151,250,1692,1716]" captionTargetBox="[416,1172,849,1669]" captionTargetId="figure-246@9.[416,1172,849,1669]" captionTargetPageId="9" captionText="FIGURE 5. Alluviobolus omega, sp. nov., paratype what (FMMC 5403), multi-layer photographs. A. Male anterior body, lateral view; B. Male leg pairs 1–7, ventral view; C. Male posterior body end with telson; D. Anterior gonopod, anterior view; E. Anterior gonopod, posterior view; F. Posterior gonopod, anterior view; G. Posterior gonopod, posterior view. Abbreviations: a = spinose membranous area; Apo = apodeme; av = paraprocts; b= lateral opening of the sperm canal; bb = basal branch of posterior gonopod; c = membranous fringe below the apex; Co = collum (ring 1); Cx = coxite; d = large membranous area located mesally; Gn = gnathochilarium; h = ‘hole’-like structure on the posterior gonopod; Md = basal joints of mandible; oz = ozopore Pre = epiproct; sf = setiferous foveolae; sg = spermatic groove; St = sternite; sub = hypoproct; t = telopodite; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022762" httpUri="https://zenodo.org/record/14022762/files/figure.png" pageId="11" pageNumber="472">Fig. 5A</figureCitation>
|
||
), legs, antennae (
|
||
<figureCitation id="13061893FFDFFFB6FBEF2ACFD5F9F946" box="[1098,1185,1732,1759]" captionStart="FIGURE 5" captionStartId="9.[151,250,1692,1716]" captionTargetBox="[416,1172,849,1669]" captionTargetId="figure-246@9.[416,1172,849,1669]" captionTargetPageId="9" captionText="FIGURE 5. Alluviobolus omega, sp. nov., paratype what (FMMC 5403), multi-layer photographs. A. Male anterior body, lateral view; B. Male leg pairs 1–7, ventral view; C. Male posterior body end with telson; D. Anterior gonopod, anterior view; E. Anterior gonopod, posterior view; F. Posterior gonopod, anterior view; G. Posterior gonopod, posterior view. Abbreviations: a = spinose membranous area; Apo = apodeme; av = paraprocts; b= lateral opening of the sperm canal; bb = basal branch of posterior gonopod; c = membranous fringe below the apex; Co = collum (ring 1); Cx = coxite; d = large membranous area located mesally; Gn = gnathochilarium; h = ‘hole’-like structure on the posterior gonopod; Md = basal joints of mandible; oz = ozopore Pre = epiproct; sf = setiferous foveolae; sg = spermatic groove; St = sternite; sub = hypoproct; t = telopodite; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022762" httpUri="https://zenodo.org/record/14022762/files/figure.png" pageId="11" pageNumber="472">Fig. 5B</figureCitation>
|
||
) and anal valves (
|
||
<figureCitation id="13061893FFDFFFB6FACA2ACFD1E1F89A" captionStart="FIGURE 5" captionStartId="9.[151,250,1692,1716]" captionTargetBox="[416,1172,849,1669]" captionTargetId="figure-246@9.[416,1172,849,1669]" captionTargetPageId="9" captionText="FIGURE 5. Alluviobolus omega, sp. nov., paratype what (FMMC 5403), multi-layer photographs. A. Male anterior body, lateral view; B. Male leg pairs 1–7, ventral view; C. Male posterior body end with telson; D. Anterior gonopod, anterior view; E. Anterior gonopod, posterior view; F. Posterior gonopod, anterior view; G. Posterior gonopod, posterior view. Abbreviations: a = spinose membranous area; Apo = apodeme; av = paraprocts; b= lateral opening of the sperm canal; bb = basal branch of posterior gonopod; c = membranous fringe below the apex; Co = collum (ring 1); Cx = coxite; d = large membranous area located mesally; Gn = gnathochilarium; h = ‘hole’-like structure on the posterior gonopod; Md = basal joints of mandible; oz = ozopore Pre = epiproct; sf = setiferous foveolae; sg = spermatic groove; St = sternite; sub = hypoproct; t = telopodite; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022762" httpUri="https://zenodo.org/record/14022762/files/figure.png" pageId="11" pageNumber="472">Fig. 5C</figureCitation>
|
||
) red. Telson and body black, dorsally with a thin red stripe.
|
||
</paragraph>
|
||
<paragraph id="8B820416FFDFFFB1FF622B07D20DFF4C" blockId="11.[151,1437,1624,2010]" lastBlockId="12.[151,1436,151,213]" lastPageId="12" lastPageNumber="473" pageId="11" pageNumber="472">
|
||
Head:Eyes with 30 ommatidia arranged in 4–5 rows (
|
||
<figureCitation id="13061893FFDFFFB6FCAC2B07D238F8BE" box="[777,864,1804,1831]" captionStart="FIGURE 5" captionStartId="9.[151,250,1692,1716]" captionTargetBox="[416,1172,849,1669]" captionTargetId="figure-246@9.[416,1172,849,1669]" captionTargetPageId="9" captionText="FIGURE 5. Alluviobolus omega, sp. nov., paratype what (FMMC 5403), multi-layer photographs. A. Male anterior body, lateral view; B. Male leg pairs 1–7, ventral view; C. Male posterior body end with telson; D. Anterior gonopod, anterior view; E. Anterior gonopod, posterior view; F. Posterior gonopod, anterior view; G. Posterior gonopod, posterior view. Abbreviations: a = spinose membranous area; Apo = apodeme; av = paraprocts; b= lateral opening of the sperm canal; bb = basal branch of posterior gonopod; c = membranous fringe below the apex; Co = collum (ring 1); Cx = coxite; d = large membranous area located mesally; Gn = gnathochilarium; h = ‘hole’-like structure on the posterior gonopod; Md = basal joints of mandible; oz = ozopore Pre = epiproct; sf = setiferous foveolae; sg = spermatic groove; St = sternite; sub = hypoproct; t = telopodite; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022762" httpUri="https://zenodo.org/record/14022762/files/figure.png" pageId="11" pageNumber="472">Fig.5A</figureCitation>
|
||
).Antennae, length of antennomeres1<<2>3>4>5=6 (
|
||
<figureCitation id="13061893FFDFFFB6FF3A2B3BD1AAF8D3" box="[159,242,1840,1866]" captionStart="FIGURE 7" captionStartId="11.[151,250,1380,1404]" captionTargetBox="[151,1435,273,1357]" captionTargetId="figure-54@11.[151,1435,273,1357]" captionTargetPageId="11" captionText="FIGURE 7. Alluviobolus omega, sp. nov., paratype (FMMC 5462), SEM. A. Left antenna, overview; B. Left antenna, antennomeres 5–7, apical view; C. Fine structures between antennomere 6 and 7. Abbreviations: A1–A7 = antennomeres 1–7; ac = apical cones; d = antennal disc; sb = sensilla basiconica." figureDoi="http://doi.org/10.5281/zenodo.14022768" httpUri="https://zenodo.org/record/14022768/files/figure.png" pageId="11" pageNumber="472">Fig 7A</figureCitation>
|
||
), with fields on sensilla basiconica on 5th and 6th (
|
||
<figureCitation id="13061893FFDFFFB6FC8B2B3BD2F2F8D3" box="[814,938,1840,1866]" captionStart="FIGURE 7" captionStartId="11.[151,250,1380,1404]" captionTargetBox="[151,1435,273,1357]" captionTargetId="figure-54@11.[151,1435,273,1357]" captionTargetPageId="11" captionText="FIGURE 7. Alluviobolus omega, sp. nov., paratype (FMMC 5462), SEM. A. Left antenna, overview; B. Left antenna, antennomeres 5–7, apical view; C. Fine structures between antennomere 6 and 7. Abbreviations: A1–A7 = antennomeres 1–7; ac = apical cones; d = antennal disc; sb = sensilla basiconica." figureDoi="http://doi.org/10.5281/zenodo.14022768" httpUri="https://zenodo.org/record/14022768/files/figure.png" pageId="11" pageNumber="472">Figs 7B, C</figureCitation>
|
||
), 4 apical cones (
|
||
<figureCitation id="13061893FFDFFFB6FBC92B3BD59DF8D3" box="[1132,1221,1840,1866]" captionStart="FIGURE 7" captionStartId="11.[151,250,1380,1404]" captionTargetBox="[151,1435,273,1357]" captionTargetId="figure-54@11.[151,1435,273,1357]" captionTargetPageId="11" captionText="FIGURE 7. Alluviobolus omega, sp. nov., paratype (FMMC 5462), SEM. A. Left antenna, overview; B. Left antenna, antennomeres 5–7, apical view; C. Fine structures between antennomere 6 and 7. Abbreviations: A1–A7 = antennomeres 1–7; ac = apical cones; d = antennal disc; sb = sensilla basiconica." figureDoi="http://doi.org/10.5281/zenodo.14022768" httpUri="https://zenodo.org/record/14022768/files/figure.png" pageId="11" pageNumber="472">Fig. 7B</figureCitation>
|
||
). Antennae in male long, reaching back to 5th body ring (
|
||
<figureCitation id="13061893FFDFFFB6FDF02B5FD3EAF8F6" box="[597,690,1876,1903]" captionStart="FIGURE 5" captionStartId="9.[151,250,1692,1716]" captionTargetBox="[416,1172,849,1669]" captionTargetId="figure-246@9.[416,1172,849,1669]" captionTargetPageId="9" captionText="FIGURE 5. Alluviobolus omega, sp. nov., paratype what (FMMC 5403), multi-layer photographs. A. Male anterior body, lateral view; B. Male leg pairs 1–7, ventral view; C. Male posterior body end with telson; D. Anterior gonopod, anterior view; E. Anterior gonopod, posterior view; F. Posterior gonopod, anterior view; G. Posterior gonopod, posterior view. Abbreviations: a = spinose membranous area; Apo = apodeme; av = paraprocts; b= lateral opening of the sperm canal; bb = basal branch of posterior gonopod; c = membranous fringe below the apex; Co = collum (ring 1); Cx = coxite; d = large membranous area located mesally; Gn = gnathochilarium; h = ‘hole’-like structure on the posterior gonopod; Md = basal joints of mandible; oz = ozopore Pre = epiproct; sf = setiferous foveolae; sg = spermatic groove; St = sternite; sub = hypoproct; t = telopodite; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022762" httpUri="https://zenodo.org/record/14022762/files/figure.png" pageId="11" pageNumber="472">Fig. 5A</figureCitation>
|
||
), in female much shorter, protruding back to body ring 2. Head with 2/2 labral grooves and 2/2 setae on lamellae linguales of gnathochilarium (
|
||
<figureCitation id="13061893FFDFFFB6FB852B73D524F80A" box="[1056,1148,1912,1939]" captionStart="FIGURE 5" captionStartId="9.[151,250,1692,1716]" captionTargetBox="[416,1172,849,1669]" captionTargetId="figure-246@9.[416,1172,849,1669]" captionTargetPageId="9" captionText="FIGURE 5. Alluviobolus omega, sp. nov., paratype what (FMMC 5403), multi-layer photographs. A. Male anterior body, lateral view; B. Male leg pairs 1–7, ventral view; C. Male posterior body end with telson; D. Anterior gonopod, anterior view; E. Anterior gonopod, posterior view; F. Posterior gonopod, anterior view; G. Posterior gonopod, posterior view. Abbreviations: a = spinose membranous area; Apo = apodeme; av = paraprocts; b= lateral opening of the sperm canal; bb = basal branch of posterior gonopod; c = membranous fringe below the apex; Co = collum (ring 1); Cx = coxite; d = large membranous area located mesally; Gn = gnathochilarium; h = ‘hole’-like structure on the posterior gonopod; Md = basal joints of mandible; oz = ozopore Pre = epiproct; sf = setiferous foveolae; sg = spermatic groove; St = sternite; sub = hypoproct; t = telopodite; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022762" httpUri="https://zenodo.org/record/14022762/files/figure.png" pageId="11" pageNumber="472">Fig. 5A</figureCitation>
|
||
). Gnathochilarium with a mentum divided by a suture, stipites each with three apical setae (
|
||
<figureCitation id="13061893FFDFFFB6FC202B97D2B8F82F" box="[901,992,1948,1974]" captionStart="FIGURE 8" captionStartId="12.[151,250,1388,1412]" captionTargetBox="[151,1437,237,1365]" captionTargetId="figure-47@12.[151,1437,237,1365]" captionTargetPageId="12" captionText="FIGURE 8. Alluviobolus omega, sp. nov., paratype (FMMC 5462), SEM. A. Gnathochilarium underside; B. Gnathochilarium, detail of central pads; C. Right mandible; D. Female midbody leg, overview; E. Female midbody leg, detail of tarsus; F. Right female vulva, mesal view. Abbreviations: 3iT = 3-combed inner tooth; as = apical spine; av = anterior orientated plate; cl = claw; cp = central pads; Cx = coxa; Endo = endochilarium; eP = lateral palpi; eT = external tooth; Fe = femur; iP = inner palpi; LT = lateral tooth; mp = molar plate; O = operculum; pL = pectinate lamellae; Po = postfemur; Pre = prefemur; pv = posterior orientated plate; vs = ventral spine; Ta = tarsus; Ti = tibia." figureDoi="http://doi.org/10.5281/zenodo.14022770" httpUri="https://zenodo.org/record/14022770/files/figure.png" pageId="11" pageNumber="472">Fig. 8A</figureCitation>
|
||
). Central pads of gnathochilarium with two sets of sensory cones, one rounded spot with 6 or 7 located more apically and an oval field with cones in 3–4 rows closer to the endochilarium (
|
||
<figureCitation id="13061893FFD8FFB1FDB02C9CD335FF28" box="[533,621,151,177]" captionStart="FIGURE 8" captionStartId="12.[151,250,1388,1412]" captionTargetBox="[151,1437,237,1365]" captionTargetId="figure-47@12.[151,1437,237,1365]" captionTargetPageId="12" captionText="FIGURE 8. Alluviobolus omega, sp. nov., paratype (FMMC 5462), SEM. A. Gnathochilarium underside; B. Gnathochilarium, detail of central pads; C. Right mandible; D. Female midbody leg, overview; E. Female midbody leg, detail of tarsus; F. Right female vulva, mesal view. Abbreviations: 3iT = 3-combed inner tooth; as = apical spine; av = anterior orientated plate; cl = claw; cp = central pads; Cx = coxa; Endo = endochilarium; eP = lateral palpi; eT = external tooth; Fe = femur; iP = inner palpi; LT = lateral tooth; mp = molar plate; O = operculum; pL = pectinate lamellae; Po = postfemur; Pre = prefemur; pv = posterior orientated plate; vs = ventral spine; Ta = tarsus; Ti = tibia." figureDoi="http://doi.org/10.5281/zenodo.14022770" httpUri="https://zenodo.org/record/14022770/files/figure.png" pageId="12" pageNumber="473">Fig. 8B</figureCitation>
|
||
). Mandible with a 3-combed inner tooth, pectinate lamellae with teeth in 5–6 rows, molar plate relatively small, with 6 ridges (
|
||
<figureCitation id="13061893FFD8FFB1FD4A2CB0D210FF4C" box="[751,840,187,213]" captionStart="FIGURE 8" captionStartId="12.[151,250,1388,1412]" captionTargetBox="[151,1437,237,1365]" captionTargetId="figure-47@12.[151,1437,237,1365]" captionTargetPageId="12" captionText="FIGURE 8. Alluviobolus omega, sp. nov., paratype (FMMC 5462), SEM. A. Gnathochilarium underside; B. Gnathochilarium, detail of central pads; C. Right mandible; D. Female midbody leg, overview; E. Female midbody leg, detail of tarsus; F. Right female vulva, mesal view. Abbreviations: 3iT = 3-combed inner tooth; as = apical spine; av = anterior orientated plate; cl = claw; cp = central pads; Cx = coxa; Endo = endochilarium; eP = lateral palpi; eT = external tooth; Fe = femur; iP = inner palpi; LT = lateral tooth; mp = molar plate; O = operculum; pL = pectinate lamellae; Po = postfemur; Pre = prefemur; pv = posterior orientated plate; vs = ventral spine; Ta = tarsus; Ti = tibia." figureDoi="http://doi.org/10.5281/zenodo.14022770" httpUri="https://zenodo.org/record/14022770/files/figure.png" pageId="12" pageNumber="473">Fig. 8C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF42549EFFD8FFB1FF322967D3BBF9A1" ID-DOI="http://doi.org/10.5281/zenodo.14022770" ID-Zenodo-Dep="14022770" httpUri="https://zenodo.org/record/14022770/files/figure.png" pageId="12" pageNumber="473" startId="12.[151,250,1388,1412]" targetBox="[151,1437,237,1365]" targetPageId="12" targetType="figure">
|
||
<paragraph id="8B820416FFD8FFB1FF322967D3BBF9A1" blockId="12.[151,1437,1388,1592]" pageId="12" pageNumber="473">
|
||
<emphasis id="B949D804FFD8FFB1FF322967D04AFA1D" bold="true" box="[151,274,1388,1412]" pageId="12" pageNumber="473">FIGURE 8.</emphasis>
|
||
<taxonomicName id="4C3D7F95FFD8FFB1FEBD2967D0BAFA1D" authorityName="Wesener" authorityYear="2024" box="[280,482,1388,1412]" class="Diplopoda" family="Pachybolidae" genus="Alluviobolus" kingdom="Animalia" order="Spirobolida" pageId="12" pageNumber="473" phylum="Arthropoda" rank="species" species="omega" status="sp. nov.">
|
||
<emphasis id="B949D804FFD8FFB1FEBD2967D0BAFA1D" box="[280,482,1388,1412]" italics="true" pageId="12" pageNumber="473">Alluviobolus omega</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B949D804FFD8FFB1FE4B2966D366FA1C" bold="true" box="[494,574,1389,1413]" pageId="12" pageNumber="473">
|
||
<taxonomicNameLabel id="A27A657FFFD8FFB1FE4B2966D366FA1C" box="[494,574,1389,1413]" pageId="12" pageNumber="473" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, paratype (
|
||
<emphasis id="B949D804FFD8FFB1FD152967D262FA1D" bold="true" box="[688,826,1388,1412]" pageId="12" pageNumber="473">FMMC 5462</emphasis>
|
||
), SEM.
|
||
<emphasis id="B949D804FFD8FFB1FC2B2967D2F8FA1D" bold="true" box="[910,928,1388,1412]" pageId="12" pageNumber="473">A</emphasis>
|
||
. Gnathochilarium underside;
|
||
<emphasis id="B949D804FFD8FFB1FB682966D586FA1C" bold="true" box="[1229,1246,1389,1413]" pageId="12" pageNumber="473">B</emphasis>
|
||
. Gnathochilarium, detail of central pads;
|
||
<emphasis id="B949D804FFD8FFB1FEDC299BD0D3FA31" bold="true" box="[377,395,1424,1448]" pageId="12" pageNumber="473">C</emphasis>
|
||
. Right mandible;
|
||
<emphasis id="B949D804FFD8FFB1FDE6299AD30DFA30" bold="true" box="[579,597,1425,1449]" pageId="12" pageNumber="473">D</emphasis>
|
||
. Female midbody leg, overview;
|
||
<emphasis id="B949D804FFD8FFB1FC0E299AD2E4FA30" bold="true" box="[939,956,1425,1449]" pageId="12" pageNumber="473">E</emphasis>
|
||
. Female midbody leg, detail of tarsus;
|
||
<emphasis id="B949D804FFD8FFB1FAEF299AD405FA30" bold="true" box="[1354,1373,1425,1449]" pageId="12" pageNumber="473">F.</emphasis>
|
||
Right female vulva, mesal view.
|
||
<emphasis id="B949D804FFD8FFB1FE0E29BFD31CFA55" bold="true" box="[427,580,1460,1484]" pageId="12" pageNumber="473">Abbreviations</emphasis>
|
||
: 3iT = 3-combed inner tooth; as = apical spine; av = anterior orientated plate; cl = claw; cp = central pads; Cx = coxa; Endo = endochilarium; eP = lateral palpi; eT = external tooth; Fe = femur; iP = inner palpi; LT = lateral tooth; mp = molar plate; O = operculum; pL = pectinate lamellae; Po = postfemur; Pre = prefemur; pv = posterior orientated plate; vs = ventral spine; Ta = tarsus; Ti = tibia.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B820416FFD8FFB1FF622A80D41AF93C" blockId="12.[151,1437,1675,2026]" box="[199,1346,1675,1702]" pageId="12" pageNumber="473">
|
||
Ozopore starting at 6th ring (
|
||
<figureCitation id="13061893FFD8FFB1FDA92A80D33DF93F" box="[524,613,1675,1702]" captionStart="FIGURE 5" captionStartId="9.[151,250,1692,1716]" captionTargetBox="[416,1172,849,1669]" captionTargetId="figure-246@9.[416,1172,849,1669]" captionTargetPageId="9" captionText="FIGURE 5. Alluviobolus omega, sp. nov., paratype what (FMMC 5403), multi-layer photographs. A. Male anterior body, lateral view; B. Male leg pairs 1–7, ventral view; C. Male posterior body end with telson; D. Anterior gonopod, anterior view; E. Anterior gonopod, posterior view; F. Posterior gonopod, anterior view; G. Posterior gonopod, posterior view. Abbreviations: a = spinose membranous area; Apo = apodeme; av = paraprocts; b= lateral opening of the sperm canal; bb = basal branch of posterior gonopod; c = membranous fringe below the apex; Co = collum (ring 1); Cx = coxite; d = large membranous area located mesally; Gn = gnathochilarium; h = ‘hole’-like structure on the posterior gonopod; Md = basal joints of mandible; oz = ozopore Pre = epiproct; sf = setiferous foveolae; sg = spermatic groove; St = sternite; sub = hypoproct; t = telopodite; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022762" httpUri="https://zenodo.org/record/14022762/files/figure.png" pageId="12" pageNumber="473">Fig. 5A</figureCitation>
|
||
), located on suture between meso- and metazonite (
|
||
<figureCitation id="13061893FFD8FFB1FB0C2A80D45DF93F" box="[1193,1285,1675,1702]" captionStart="FIGURE 5" captionStartId="9.[151,250,1692,1716]" captionTargetBox="[416,1172,849,1669]" captionTargetId="figure-246@9.[416,1172,849,1669]" captionTargetPageId="9" captionText="FIGURE 5. Alluviobolus omega, sp. nov., paratype what (FMMC 5403), multi-layer photographs. A. Male anterior body, lateral view; B. Male leg pairs 1–7, ventral view; C. Male posterior body end with telson; D. Anterior gonopod, anterior view; E. Anterior gonopod, posterior view; F. Posterior gonopod, anterior view; G. Posterior gonopod, posterior view. Abbreviations: a = spinose membranous area; Apo = apodeme; av = paraprocts; b= lateral opening of the sperm canal; bb = basal branch of posterior gonopod; c = membranous fringe below the apex; Co = collum (ring 1); Cx = coxite; d = large membranous area located mesally; Gn = gnathochilarium; h = ‘hole’-like structure on the posterior gonopod; Md = basal joints of mandible; oz = ozopore Pre = epiproct; sf = setiferous foveolae; sg = spermatic groove; St = sternite; sub = hypoproct; t = telopodite; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022762" httpUri="https://zenodo.org/record/14022762/files/figure.png" pageId="12" pageNumber="473">Figs 5C</figureCitation>
|
||
,
|
||
<figureCitation id="13061893FFD8FFB1FAB52A80D46DF93C" box="[1296,1333,1675,1701]" captionStart="FIGURE 9" captionStartId="13.[151,250,1222,1246]" captionTargetBox="[510,1077,182,1198]" captionTargetId="figure-15@13.[510,1077,181,1198]" captionTargetPageId="13" captionText="FIGURE 9. Alluviobolus omega, sp. nov., population from Vohimena (FMMC 5396), multi-layer photographs.A. Male anterior body, lateral view; B. Midbody rings, lateral view; C. Male leg pairs 1–7, ventral view; D. Male posterior body end with telson; E. Anterior gonopod, anterior view; F. Anterior gonopod, posterior view; G. Right posterior gonopod, anterior view; H. Right posterior gonopod, posterior view. Abbreviations: a = spinose membranous area; av = paraprocts; b= lateral opening of the sperm canal; bb = basal branch of posterior gonopod; c = membranous fringe below the apex; Co = collum (ring 1); Cx = coxite; d = large membranous area located mesally; Gn = gnathochilarium; h = ‘hole’-like structure on the posterior gonopod; Md = basal joints of mandible; Me = mesozonite; Mt = metazonite; oz = ozopore; pe = pseudopenis; Pre = epiproct; sf = setiferous foveolae; sg = spermatic groove; St = sternite; sub = hypoproct; t = telopodite; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022774" httpUri="https://zenodo.org/record/14022774/files/figure.png" pageId="12" pageNumber="473">9B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B820416FFD8FFB1FF622AA4D34FF85C" blockId="12.[151,1437,1675,2026]" pageId="12" pageNumber="473">
|
||
Legs in both sexes long, longer than width of body rings (
|
||
<figureCitation id="13061893FFD8FFB1FCD42AA4D2B5F950" box="[881,1005,1711,1737]" captionStart="FIGURE 6" captionStartId="10.[151,250,1533,1557]" captionTargetBox="[151,1435,188,1510]" captionTargetId="figure-18@10.[151,1435,181,1510]" captionTargetPageId="10" captionText="FIGURE 6. Alluviobolus omega, sp. nov., holotype (FMMC 5458), drawings. A. Anterior gonopod, anterior view; B. Anterior gonopod, posterior view; C. Posterior gonopod, tip, anterior view; D. Posterior gonopod, tip, posterior view; E. Posterior gonopod, base, posterior view; F. Male midbody leg, posterior view; G. Female midbody leg, posterior view. Abbreviations: a = spinose membranous area; b= lateral opening of the sperm canal; bb = basal branch of posterior gonopod; c = membranous fringe below the apex; Cx = coxite; d = large membranous area located mesally; Fe = femur; fs-p = finger-shaped process; Gr = groove; h = ‘hole’-like structure on the posterior gonopod; Pre = prefemur; Po = postfemur; sg = spermatic groove; St = sternite; t = telopodite; Ta = tarsus; Ti = tibia. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022764" httpUri="https://zenodo.org/record/14022764/files/figure.png" pageId="12" pageNumber="473">Fig. 6F, G</figureCitation>
|
||
). Male legs without tarsal pads (
|
||
<figureCitation id="13061893FFD8FFB1FACE2AA4D1EEF974" captionStart="FIGURE 6" captionStartId="10.[151,250,1533,1557]" captionTargetBox="[151,1435,188,1510]" captionTargetId="figure-18@10.[151,1435,181,1510]" captionTargetPageId="10" captionText="FIGURE 6. Alluviobolus omega, sp. nov., holotype (FMMC 5458), drawings. A. Anterior gonopod, anterior view; B. Anterior gonopod, posterior view; C. Posterior gonopod, tip, anterior view; D. Posterior gonopod, tip, posterior view; E. Posterior gonopod, base, posterior view; F. Male midbody leg, posterior view; G. Female midbody leg, posterior view. Abbreviations: a = spinose membranous area; b= lateral opening of the sperm canal; bb = basal branch of posterior gonopod; c = membranous fringe below the apex; Cx = coxite; d = large membranous area located mesally; Fe = femur; fs-p = finger-shaped process; Gr = groove; h = ‘hole’-like structure on the posterior gonopod; Pre = prefemur; Po = postfemur; sg = spermatic groove; St = sternite; t = telopodite; Ta = tarsus; Ti = tibia. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022764" httpUri="https://zenodo.org/record/14022764/files/figure.png" pageId="12" pageNumber="473">Figs 6F</figureCitation>
|
||
,
|
||
<figureCitation id="13061893FFD8FFB1FF612AD8D054F974" box="[196,268,1747,1773]" captionStart="FIGURE 8" captionStartId="12.[151,250,1388,1412]" captionTargetBox="[151,1437,237,1365]" captionTargetId="figure-47@12.[151,1437,237,1365]" captionTargetPageId="12" captionText="FIGURE 8. Alluviobolus omega, sp. nov., paratype (FMMC 5462), SEM. A. Gnathochilarium underside; B. Gnathochilarium, detail of central pads; C. Right mandible; D. Female midbody leg, overview; E. Female midbody leg, detail of tarsus; F. Right female vulva, mesal view. Abbreviations: 3iT = 3-combed inner tooth; as = apical spine; av = anterior orientated plate; cl = claw; cp = central pads; Cx = coxa; Endo = endochilarium; eP = lateral palpi; eT = external tooth; Fe = femur; iP = inner palpi; LT = lateral tooth; mp = molar plate; O = operculum; pL = pectinate lamellae; Po = postfemur; Pre = prefemur; pv = posterior orientated plate; vs = ventral spine; Ta = tarsus; Ti = tibia." figureDoi="http://doi.org/10.5281/zenodo.14022770" httpUri="https://zenodo.org/record/14022770/files/figure.png" pageId="12" pageNumber="473">8D, E</figureCitation>
|
||
). Male legs 3–5 with cylindrical coxa processes, tip broadly rounded. Process of coxae 3 and 4 bent posteriorly, 3rd tip protruding to 4th, 4th tip protruding to 5th coxa. 5th coxa process not bent, short. 6th and 7th coxae flat & broad (
|
||
<figureCitation id="13061893FFD8FFB1FED92B10D08EF8AF" box="[380,470,1819,1846]" captionStart="FIGURE 5" captionStartId="9.[151,250,1692,1716]" captionTargetBox="[416,1172,849,1669]" captionTargetId="figure-246@9.[416,1172,849,1669]" captionTargetPageId="9" captionText="FIGURE 5. Alluviobolus omega, sp. nov., paratype what (FMMC 5403), multi-layer photographs. A. Male anterior body, lateral view; B. Male leg pairs 1–7, ventral view; C. Male posterior body end with telson; D. Anterior gonopod, anterior view; E. Anterior gonopod, posterior view; F. Posterior gonopod, anterior view; G. Posterior gonopod, posterior view. Abbreviations: a = spinose membranous area; Apo = apodeme; av = paraprocts; b= lateral opening of the sperm canal; bb = basal branch of posterior gonopod; c = membranous fringe below the apex; Co = collum (ring 1); Cx = coxite; d = large membranous area located mesally; Gn = gnathochilarium; h = ‘hole’-like structure on the posterior gonopod; Md = basal joints of mandible; oz = ozopore Pre = epiproct; sf = setiferous foveolae; sg = spermatic groove; St = sternite; sub = hypoproct; t = telopodite; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022762" httpUri="https://zenodo.org/record/14022762/files/figure.png" pageId="12" pageNumber="473">Fig. 5B</figureCitation>
|
||
). Legs 8+ in male with a cylindrical, basally tapering coxa, femur longest podomere, length of podomeres: coxa <prefemur<femur>postfemur>tibia<tarsus, tarsus as long as prefemur. Each podomere in both sexes with an apical spine (
|
||
<figureCitation id="13061893FFD8FFB1FDB92B68D3C9F8E4" box="[540,657,1891,1917]" captionStart="FIGURE 6" captionStartId="10.[151,250,1533,1557]" captionTargetBox="[151,1435,188,1510]" captionTargetId="figure-18@10.[151,1435,181,1510]" captionTargetPageId="10" captionText="FIGURE 6. Alluviobolus omega, sp. nov., holotype (FMMC 5458), drawings. A. Anterior gonopod, anterior view; B. Anterior gonopod, posterior view; C. Posterior gonopod, tip, anterior view; D. Posterior gonopod, tip, posterior view; E. Posterior gonopod, base, posterior view; F. Male midbody leg, posterior view; G. Female midbody leg, posterior view. Abbreviations: a = spinose membranous area; b= lateral opening of the sperm canal; bb = basal branch of posterior gonopod; c = membranous fringe below the apex; Cx = coxite; d = large membranous area located mesally; Fe = femur; fs-p = finger-shaped process; Gr = groove; h = ‘hole’-like structure on the posterior gonopod; Pre = prefemur; Po = postfemur; sg = spermatic groove; St = sternite; t = telopodite; Ta = tarsus; Ti = tibia. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022764" httpUri="https://zenodo.org/record/14022764/files/figure.png" pageId="12" pageNumber="473">Fig. 6F, G</figureCitation>
|
||
). Male tarsus with short sharp claw with small paronychium (
|
||
<figureCitation id="13061893FFD8FFB1FA9A2B68D1F3F838" captionStart="FIGURE 8" captionStartId="12.[151,250,1388,1412]" captionTargetBox="[151,1437,237,1365]" captionTargetId="figure-47@12.[151,1437,237,1365]" captionTargetPageId="12" captionText="FIGURE 8. Alluviobolus omega, sp. nov., paratype (FMMC 5462), SEM. A. Gnathochilarium underside; B. Gnathochilarium, detail of central pads; C. Right mandible; D. Female midbody leg, overview; E. Female midbody leg, detail of tarsus; F. Right female vulva, mesal view. Abbreviations: 3iT = 3-combed inner tooth; as = apical spine; av = anterior orientated plate; cl = claw; cp = central pads; Cx = coxa; Endo = endochilarium; eP = lateral palpi; eT = external tooth; Fe = femur; iP = inner palpi; LT = lateral tooth; mp = molar plate; O = operculum; pL = pectinate lamellae; Po = postfemur; Pre = prefemur; pv = posterior orientated plate; vs = ventral spine; Ta = tarsus; Ti = tibia." figureDoi="http://doi.org/10.5281/zenodo.14022770" httpUri="https://zenodo.org/record/14022770/files/figure.png" pageId="12" pageNumber="473">Fig. 8D, E</figureCitation>
|
||
), single short apical spine above claw (
|
||
<figureCitation id="13061893FFD8FFB1FDF92B8CD3EBF838" box="[604,691,1927,1953]" captionStart="FIGURE 6" captionStartId="10.[151,250,1533,1557]" captionTargetBox="[151,1435,188,1510]" captionTargetId="figure-18@10.[151,1435,181,1510]" captionTargetPageId="10" captionText="FIGURE 6. Alluviobolus omega, sp. nov., holotype (FMMC 5458), drawings. A. Anterior gonopod, anterior view; B. Anterior gonopod, posterior view; C. Posterior gonopod, tip, anterior view; D. Posterior gonopod, tip, posterior view; E. Posterior gonopod, base, posterior view; F. Male midbody leg, posterior view; G. Female midbody leg, posterior view. Abbreviations: a = spinose membranous area; b= lateral opening of the sperm canal; bb = basal branch of posterior gonopod; c = membranous fringe below the apex; Cx = coxite; d = large membranous area located mesally; Fe = femur; fs-p = finger-shaped process; Gr = groove; h = ‘hole’-like structure on the posterior gonopod; Pre = prefemur; Po = postfemur; sg = spermatic groove; St = sternite; t = telopodite; Ta = tarsus; Ti = tibia. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022764" httpUri="https://zenodo.org/record/14022764/files/figure.png" pageId="12" pageNumber="473">Figs 6F</figureCitation>
|
||
,
|
||
<figureCitation id="13061893FFD8FFB1FD182B8CD385F838" box="[701,733,1927,1953]" captionStart="FIGURE 8" captionStartId="12.[151,250,1388,1412]" captionTargetBox="[151,1437,237,1365]" captionTargetId="figure-47@12.[151,1437,237,1365]" captionTargetPageId="12" captionText="FIGURE 8. Alluviobolus omega, sp. nov., paratype (FMMC 5462), SEM. A. Gnathochilarium underside; B. Gnathochilarium, detail of central pads; C. Right mandible; D. Female midbody leg, overview; E. Female midbody leg, detail of tarsus; F. Right female vulva, mesal view. Abbreviations: 3iT = 3-combed inner tooth; as = apical spine; av = anterior orientated plate; cl = claw; cp = central pads; Cx = coxa; Endo = endochilarium; eP = lateral palpi; eT = external tooth; Fe = femur; iP = inner palpi; LT = lateral tooth; mp = molar plate; O = operculum; pL = pectinate lamellae; Po = postfemur; Pre = prefemur; pv = posterior orientated plate; vs = ventral spine; Ta = tarsus; Ti = tibia." figureDoi="http://doi.org/10.5281/zenodo.14022770" httpUri="https://zenodo.org/record/14022770/files/figure.png" pageId="12" pageNumber="473">8E</figureCitation>
|
||
) and 5 or 6 short ventral spines. Female leg similar to male leg, with four ventral spines (
|
||
<figureCitation id="13061893FFD8FFB1FE152BA0D352F85C" box="[432,522,1963,1989]" captionStart="FIGURE 6" captionStartId="10.[151,250,1533,1557]" captionTargetBox="[151,1435,188,1510]" captionTargetId="figure-18@10.[151,1435,181,1510]" captionTargetPageId="10" captionText="FIGURE 6. Alluviobolus omega, sp. nov., holotype (FMMC 5458), drawings. A. Anterior gonopod, anterior view; B. Anterior gonopod, posterior view; C. Posterior gonopod, tip, anterior view; D. Posterior gonopod, tip, posterior view; E. Posterior gonopod, base, posterior view; F. Male midbody leg, posterior view; G. Female midbody leg, posterior view. Abbreviations: a = spinose membranous area; b= lateral opening of the sperm canal; bb = basal branch of posterior gonopod; c = membranous fringe below the apex; Cx = coxite; d = large membranous area located mesally; Fe = femur; fs-p = finger-shaped process; Gr = groove; h = ‘hole’-like structure on the posterior gonopod; Pre = prefemur; Po = postfemur; sg = spermatic groove; St = sternite; t = telopodite; Ta = tarsus; Ti = tibia. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022764" httpUri="https://zenodo.org/record/14022764/files/figure.png" pageId="12" pageNumber="473">Fig. 6G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B820416FFD8FFB1FF622BC4D223F873" blockId="12.[151,1437,1675,2026]" box="[199,891,1999,2026]" pageId="12" pageNumber="473">
|
||
Anal valves not invaginated. No pre-anal processes (
|
||
<figureCitation id="13061893FFD8FFB1FCB02BC4D236F873" box="[789,878,1999,2026]" captionStart="FIGURE 5" captionStartId="9.[151,250,1692,1716]" captionTargetBox="[416,1172,849,1669]" captionTargetId="figure-246@9.[416,1172,849,1669]" captionTargetPageId="9" captionText="FIGURE 5. Alluviobolus omega, sp. nov., paratype what (FMMC 5403), multi-layer photographs. A. Male anterior body, lateral view; B. Male leg pairs 1–7, ventral view; C. Male posterior body end with telson; D. Anterior gonopod, anterior view; E. Anterior gonopod, posterior view; F. Posterior gonopod, anterior view; G. Posterior gonopod, posterior view. Abbreviations: a = spinose membranous area; Apo = apodeme; av = paraprocts; b= lateral opening of the sperm canal; bb = basal branch of posterior gonopod; c = membranous fringe below the apex; Co = collum (ring 1); Cx = coxite; d = large membranous area located mesally; Gn = gnathochilarium; h = ‘hole’-like structure on the posterior gonopod; Md = basal joints of mandible; oz = ozopore Pre = epiproct; sf = setiferous foveolae; sg = spermatic groove; St = sternite; sub = hypoproct; t = telopodite; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022762" httpUri="https://zenodo.org/record/14022762/files/figure.png" pageId="12" pageNumber="473">Fig. 5C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF42549EFFD9FFB0FF3228CDD181FA67" ID-DOI="http://doi.org/10.5281/zenodo.14022774" ID-Zenodo-Dep="14022774" httpUri="https://zenodo.org/record/14022774/files/figure.png" pageId="13" pageNumber="474" startId="13.[151,250,1222,1246]" targetBox="[510,1077,182,1198]" targetPageId="13" targetType="figure">
|
||
<paragraph id="8B820416FFD9FFB0FF3228CDD181FA67" blockId="13.[151,1436,1222,1535]" pageId="13" pageNumber="474">
|
||
<emphasis id="B949D804FFD9FFB0FF3228CDD049FB47" bold="true" box="[151,273,1222,1246]" pageId="13" pageNumber="474">FIGURE 9.</emphasis>
|
||
<taxonomicName id="4C3D7F95FFD9FFB0FEB328CCD087FB47" authorityName="Wesener" authorityYear="2024" box="[278,479,1223,1247]" class="Diplopoda" family="Pachybolidae" genus="Alluviobolus" kingdom="Animalia" order="Spirobolida" pageId="13" pageNumber="474" phylum="Arthropoda" rank="species" species="omega" status="sp. nov.">
|
||
<emphasis id="B949D804FFD9FFB0FEB328CCD087FB47" box="[278,479,1223,1247]" italics="true" pageId="13" pageNumber="474">Alluviobolus omega</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B949D804FFD9FFB0FE4C28CCD360FB46" bold="true" box="[489,568,1223,1247]" pageId="13" pageNumber="474">
|
||
<taxonomicNameLabel id="A27A657FFFD9FFB0FE4C28CCD360FB46" box="[489,568,1223,1247]" pageId="13" pageNumber="474" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, population from Vohimena (
|
||
<emphasis id="B949D804FFD9FFB0FCFB28CDD2BFFB47" bold="true" box="[862,999,1222,1246]" pageId="13" pageNumber="474">FMMC 5396</emphasis>
|
||
), multi-layer photographs.
|
||
<emphasis id="B949D804FFD9FFB0FB5228CDD451FB47" bold="true" box="[1271,1289,1222,1246]" pageId="13" pageNumber="474">A</emphasis>
|
||
. Male anterior body, lateral view;
|
||
<emphasis id="B949D804FFD9FFB0FEFD28E0D031FA9A" bold="true" box="[344,361,1259,1283]" pageId="13" pageNumber="474">B</emphasis>
|
||
. Midbody rings, lateral view;
|
||
<emphasis id="B949D804FFD9FFB0FD3F28E1D3F4FA9B" bold="true" box="[666,684,1258,1282]" pageId="13" pageNumber="474">C</emphasis>
|
||
. Male leg pairs 1–7, ventral view;
|
||
<emphasis id="B949D804FFD9FFB0FBAE28E0D545FA9A" bold="true" box="[1035,1053,1259,1283]" pageId="13" pageNumber="474">D</emphasis>
|
||
. Male posterior body end with telson;
|
||
<emphasis id="B949D804FFD9FFB0FF322904D1F0FABE" bold="true" box="[151,168,1295,1319]" pageId="13" pageNumber="474">E</emphasis>
|
||
. Anterior gonopod, anterior view;
|
||
<emphasis id="B949D804FFD9FFB0FDAE2904D342FABE" bold="true" box="[523,538,1295,1319]" pageId="13" pageNumber="474">F</emphasis>
|
||
. Anterior gonopod, posterior view;
|
||
<emphasis id="B949D804FFD9FFB0FC2D2905D2C4FABF" bold="true" box="[904,924,1294,1318]" pageId="13" pageNumber="474">G</emphasis>
|
||
. Right posterior gonopod, anterior view;
|
||
<emphasis id="B949D804FFD9FFB0FAE62904D40FFABE" bold="true" box="[1347,1367,1295,1319]" pageId="13" pageNumber="474">H</emphasis>
|
||
. Right posterior gonopod, posterior view.
|
||
<emphasis id="B949D804FFD9FFB0FDA42939D3C2FAD3" bold="true" box="[513,666,1330,1354]" pageId="13" pageNumber="474">Abbreviations</emphasis>
|
||
: a = spinose membranous area; av = paraprocts; b= lateral opening of the sperm canal; bb = basal branch of posterior gonopod; c = membranous fringe below the apex; Co = collum (ring 1); Cx = coxite; d = large membranous area located mesally; Gn = gnathochilarium; h = ‘hole’-like structure on the posterior gonopod; Md = basal joints of mandible; Me = mesozonite; Mt = metazonite; oz = ozopore; pe = pseudopenis; Pre = epiproct; sf = setiferous foveolae; sg = spermatic groove; St = sternite; sub = hypoproct; t = telopodite; numbers refer to leg pair number.
|
||
<emphasis id="B949D804FFD9FFB0FABD29C9D4DFFA42" bold="true" box="[1304,1415,1474,1499]" pageId="13" pageNumber="474">Scale bars</emphasis>
|
||
= 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B820416FFD9FFB0FF622A5AD4D9F916" blockId="13.[151,1437,1617,2040]" pageId="13" pageNumber="474">
|
||
Female vulva simple, bivalve-like, with a small, poorly sclerotized operculum at base (
|
||
<figureCitation id="13061893FFD9FFB0FB292A5AD5B9F9F2" box="[1164,1249,1617,1643]" captionStart="FIGURE 8" captionStartId="12.[151,250,1388,1412]" captionTargetBox="[151,1437,237,1365]" captionTargetId="figure-47@12.[151,1437,237,1365]" captionTargetPageId="12" captionText="FIGURE 8. Alluviobolus omega, sp. nov., paratype (FMMC 5462), SEM. A. Gnathochilarium underside; B. Gnathochilarium, detail of central pads; C. Right mandible; D. Female midbody leg, overview; E. Female midbody leg, detail of tarsus; F. Right female vulva, mesal view. Abbreviations: 3iT = 3-combed inner tooth; as = apical spine; av = anterior orientated plate; cl = claw; cp = central pads; Cx = coxa; Endo = endochilarium; eP = lateral palpi; eT = external tooth; Fe = femur; iP = inner palpi; LT = lateral tooth; mp = molar plate; O = operculum; pL = pectinate lamellae; Po = postfemur; Pre = prefemur; pv = posterior orientated plate; vs = ventral spine; Ta = tarsus; Ti = tibia." figureDoi="http://doi.org/10.5281/zenodo.14022770" httpUri="https://zenodo.org/record/14022770/files/figure.png" pageId="13" pageNumber="474">Fig. 8F</figureCitation>
|
||
). Posterior valve apically overlapping part of anterior one. Basally on each valve towards opening with two or three rows of setae
|
||
</paragraph>
|
||
<paragraph id="8B820416FFD9FFB0FF622A92D3B2F8DA" blockId="13.[151,1437,1617,2040]" pageId="13" pageNumber="474">
|
||
Anterior gonopods massive and wide (
|
||
<figureCitation id="13061893FFD9FFB0FDDE2A92D3A0F92A" box="[635,760,1689,1716]" captionStart="FIGURE 5" captionStartId="9.[151,250,1692,1716]" captionTargetBox="[416,1172,849,1669]" captionTargetId="figure-246@9.[416,1172,849,1669]" captionTargetPageId="9" captionText="FIGURE 5. Alluviobolus omega, sp. nov., paratype what (FMMC 5403), multi-layer photographs. A. Male anterior body, lateral view; B. Male leg pairs 1–7, ventral view; C. Male posterior body end with telson; D. Anterior gonopod, anterior view; E. Anterior gonopod, posterior view; F. Posterior gonopod, anterior view; G. Posterior gonopod, posterior view. Abbreviations: a = spinose membranous area; Apo = apodeme; av = paraprocts; b= lateral opening of the sperm canal; bb = basal branch of posterior gonopod; c = membranous fringe below the apex; Co = collum (ring 1); Cx = coxite; d = large membranous area located mesally; Gn = gnathochilarium; h = ‘hole’-like structure on the posterior gonopod; Md = basal joints of mandible; oz = ozopore Pre = epiproct; sf = setiferous foveolae; sg = spermatic groove; St = sternite; sub = hypoproct; t = telopodite; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022762" httpUri="https://zenodo.org/record/14022762/files/figure.png" pageId="13" pageNumber="474">Figs 5D, E</figureCitation>
|
||
,
|
||
<figureCitation id="13061893FFD9FFB0FCA02A92D213F92A" box="[773,843,1689,1715]" captionStart="FIGURE 6" captionStartId="10.[151,250,1533,1557]" captionTargetBox="[151,1435,188,1510]" captionTargetId="figure-18@10.[151,1435,181,1510]" captionTargetPageId="10" captionText="FIGURE 6. Alluviobolus omega, sp. nov., holotype (FMMC 5458), drawings. A. Anterior gonopod, anterior view; B. Anterior gonopod, posterior view; C. Posterior gonopod, tip, anterior view; D. Posterior gonopod, tip, posterior view; E. Posterior gonopod, base, posterior view; F. Male midbody leg, posterior view; G. Female midbody leg, posterior view. Abbreviations: a = spinose membranous area; b= lateral opening of the sperm canal; bb = basal branch of posterior gonopod; c = membranous fringe below the apex; Cx = coxite; d = large membranous area located mesally; Fe = femur; fs-p = finger-shaped process; Gr = groove; h = ‘hole’-like structure on the posterior gonopod; Pre = prefemur; Po = postfemur; sg = spermatic groove; St = sternite; t = telopodite; Ta = tarsus; Ti = tibia. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022764" httpUri="https://zenodo.org/record/14022764/files/figure.png" pageId="13" pageNumber="474">6A, B</figureCitation>
|
||
). Median sternal projection triangular and elongated with a well-rounded, rectangular tip (
|
||
<figureCitation id="13061893FFD9FFB0FD8A2AB6D3FFF94E" box="[559,679,1725,1752]" captionStart="FIGURE 5" captionStartId="9.[151,250,1692,1716]" captionTargetBox="[416,1172,849,1669]" captionTargetId="figure-246@9.[416,1172,849,1669]" captionTargetPageId="9" captionText="FIGURE 5. Alluviobolus omega, sp. nov., paratype what (FMMC 5403), multi-layer photographs. A. Male anterior body, lateral view; B. Male leg pairs 1–7, ventral view; C. Male posterior body end with telson; D. Anterior gonopod, anterior view; E. Anterior gonopod, posterior view; F. Posterior gonopod, anterior view; G. Posterior gonopod, posterior view. Abbreviations: a = spinose membranous area; Apo = apodeme; av = paraprocts; b= lateral opening of the sperm canal; bb = basal branch of posterior gonopod; c = membranous fringe below the apex; Co = collum (ring 1); Cx = coxite; d = large membranous area located mesally; Gn = gnathochilarium; h = ‘hole’-like structure on the posterior gonopod; Md = basal joints of mandible; oz = ozopore Pre = epiproct; sf = setiferous foveolae; sg = spermatic groove; St = sternite; sub = hypoproct; t = telopodite; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022762" httpUri="https://zenodo.org/record/14022762/files/figure.png" pageId="13" pageNumber="474">Figs 5D, E</figureCitation>
|
||
,
|
||
<figureCitation id="13061893FFD9FFB0FD142AB6D3ACF94E" box="[689,756,1725,1751]" captionStart="FIGURE 6" captionStartId="10.[151,250,1533,1557]" captionTargetBox="[151,1435,188,1510]" captionTargetId="figure-18@10.[151,1435,181,1510]" captionTargetPageId="10" captionText="FIGURE 6. Alluviobolus omega, sp. nov., holotype (FMMC 5458), drawings. A. Anterior gonopod, anterior view; B. Anterior gonopod, posterior view; C. Posterior gonopod, tip, anterior view; D. Posterior gonopod, tip, posterior view; E. Posterior gonopod, base, posterior view; F. Male midbody leg, posterior view; G. Female midbody leg, posterior view. Abbreviations: a = spinose membranous area; b= lateral opening of the sperm canal; bb = basal branch of posterior gonopod; c = membranous fringe below the apex; Cx = coxite; d = large membranous area located mesally; Fe = femur; fs-p = finger-shaped process; Gr = groove; h = ‘hole’-like structure on the posterior gonopod; Pre = prefemur; Po = postfemur; sg = spermatic groove; St = sternite; t = telopodite; Ta = tarsus; Ti = tibia. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022764" httpUri="https://zenodo.org/record/14022764/files/figure.png" pageId="13" pageNumber="474">6A, B</figureCitation>
|
||
). Sternal process always far longer than coxite and telopodite, except for telopodite process. Coxite process weakly developed, broadly rounded mesally and barely visible behind massive sternite (
|
||
<figureCitation id="13061893FFD9FFB0FEFF2B0ED08CF886" box="[346,468,1797,1823]" captionStart="FIGURE 6" captionStartId="10.[151,250,1533,1557]" captionTargetBox="[151,1435,188,1510]" captionTargetId="figure-18@10.[151,1435,181,1510]" captionTargetPageId="10" captionText="FIGURE 6. Alluviobolus omega, sp. nov., holotype (FMMC 5458), drawings. A. Anterior gonopod, anterior view; B. Anterior gonopod, posterior view; C. Posterior gonopod, tip, anterior view; D. Posterior gonopod, tip, posterior view; E. Posterior gonopod, base, posterior view; F. Male midbody leg, posterior view; G. Female midbody leg, posterior view. Abbreviations: a = spinose membranous area; b= lateral opening of the sperm canal; bb = basal branch of posterior gonopod; c = membranous fringe below the apex; Cx = coxite; d = large membranous area located mesally; Fe = femur; fs-p = finger-shaped process; Gr = groove; h = ‘hole’-like structure on the posterior gonopod; Pre = prefemur; Po = postfemur; sg = spermatic groove; St = sternite; t = telopodite; Ta = tarsus; Ti = tibia. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022764" httpUri="https://zenodo.org/record/14022764/files/figure.png" pageId="13" pageNumber="474">Fig. 6A, B</figureCitation>
|
||
). Telopodite mesally with a slender, apically tapering process (
|
||
<figureCitation id="13061893FFD9FFB0FB312B0ED448F886" box="[1172,1296,1797,1824]" captionStart="FIGURE 5" captionStartId="9.[151,250,1692,1716]" captionTargetBox="[416,1172,849,1669]" captionTargetId="figure-246@9.[416,1172,849,1669]" captionTargetPageId="9" captionText="FIGURE 5. Alluviobolus omega, sp. nov., paratype what (FMMC 5403), multi-layer photographs. A. Male anterior body, lateral view; B. Male leg pairs 1–7, ventral view; C. Male posterior body end with telson; D. Anterior gonopod, anterior view; E. Anterior gonopod, posterior view; F. Posterior gonopod, anterior view; G. Posterior gonopod, posterior view. Abbreviations: a = spinose membranous area; Apo = apodeme; av = paraprocts; b= lateral opening of the sperm canal; bb = basal branch of posterior gonopod; c = membranous fringe below the apex; Co = collum (ring 1); Cx = coxite; d = large membranous area located mesally; Gn = gnathochilarium; h = ‘hole’-like structure on the posterior gonopod; Md = basal joints of mandible; oz = ozopore Pre = epiproct; sf = setiferous foveolae; sg = spermatic groove; St = sternite; sub = hypoproct; t = telopodite; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022762" httpUri="https://zenodo.org/record/14022762/files/figure.png" pageId="13" pageNumber="474">Figs 5D, E</figureCitation>
|
||
,
|
||
<figureCitation id="13061893FFD9FFB0FAB82B0ED43AF886" box="[1309,1378,1797,1823]" captionStart="FIGURE 6" captionStartId="10.[151,250,1533,1557]" captionTargetBox="[151,1435,188,1510]" captionTargetId="figure-18@10.[151,1435,181,1510]" captionTargetPageId="10" captionText="FIGURE 6. Alluviobolus omega, sp. nov., holotype (FMMC 5458), drawings. A. Anterior gonopod, anterior view; B. Anterior gonopod, posterior view; C. Posterior gonopod, tip, anterior view; D. Posterior gonopod, tip, posterior view; E. Posterior gonopod, base, posterior view; F. Male midbody leg, posterior view; G. Female midbody leg, posterior view. Abbreviations: a = spinose membranous area; b= lateral opening of the sperm canal; bb = basal branch of posterior gonopod; c = membranous fringe below the apex; Cx = coxite; d = large membranous area located mesally; Fe = femur; fs-p = finger-shaped process; Gr = groove; h = ‘hole’-like structure on the posterior gonopod; Pre = prefemur; Po = postfemur; sg = spermatic groove; St = sternite; t = telopodite; Ta = tarsus; Ti = tibia. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022764" httpUri="https://zenodo.org/record/14022764/files/figure.png" pageId="13" pageNumber="474">6A, B</figureCitation>
|
||
). Tip with a small, sharp-edged, retrorse process (
|
||
<figureCitation id="13061893FFD9FFB0FD202B22D386F8DA" box="[645,734,1833,1859]" captionStart="FIGURE 6" captionStartId="10.[151,250,1533,1557]" captionTargetBox="[151,1435,188,1510]" captionTargetId="figure-18@10.[151,1435,181,1510]" captionTargetPageId="10" captionText="FIGURE 6. Alluviobolus omega, sp. nov., holotype (FMMC 5458), drawings. A. Anterior gonopod, anterior view; B. Anterior gonopod, posterior view; C. Posterior gonopod, tip, anterior view; D. Posterior gonopod, tip, posterior view; E. Posterior gonopod, base, posterior view; F. Male midbody leg, posterior view; G. Female midbody leg, posterior view. Abbreviations: a = spinose membranous area; b= lateral opening of the sperm canal; bb = basal branch of posterior gonopod; c = membranous fringe below the apex; Cx = coxite; d = large membranous area located mesally; Fe = femur; fs-p = finger-shaped process; Gr = groove; h = ‘hole’-like structure on the posterior gonopod; Pre = prefemur; Po = postfemur; sg = spermatic groove; St = sternite; t = telopodite; Ta = tarsus; Ti = tibia. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022764" httpUri="https://zenodo.org/record/14022764/files/figure.png" pageId="13" pageNumber="474">Fig. 6B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B820416FFD9FFB0FF622B46D4C4F861" blockId="13.[151,1437,1617,2040]" pageId="13" pageNumber="474">
|
||
Posterior gonopods
|
||
<emphasis id="B949D804FFD9FFB0FE032B45D0B3F8FE" box="[422,491,1870,1895]" italics="true" pageId="13" pageNumber="474">in situ</emphasis>
|
||
almost completely covered by coxite and telopodite of anterior gonopod. Coxite and telopodite of posterior gonopod clearly separated, with telopodite located at different level and angle than coxite (
|
||
<figureCitation id="13061893FFD9FFB0FF3A2B9ED04FF836" box="[159,279,1941,1968]" captionStart="FIGURE 5" captionStartId="9.[151,250,1692,1716]" captionTargetBox="[416,1172,849,1669]" captionTargetId="figure-246@9.[416,1172,849,1669]" captionTargetPageId="9" captionText="FIGURE 5. Alluviobolus omega, sp. nov., paratype what (FMMC 5403), multi-layer photographs. A. Male anterior body, lateral view; B. Male leg pairs 1–7, ventral view; C. Male posterior body end with telson; D. Anterior gonopod, anterior view; E. Anterior gonopod, posterior view; F. Posterior gonopod, anterior view; G. Posterior gonopod, posterior view. Abbreviations: a = spinose membranous area; Apo = apodeme; av = paraprocts; b= lateral opening of the sperm canal; bb = basal branch of posterior gonopod; c = membranous fringe below the apex; Co = collum (ring 1); Cx = coxite; d = large membranous area located mesally; Gn = gnathochilarium; h = ‘hole’-like structure on the posterior gonopod; Md = basal joints of mandible; oz = ozopore Pre = epiproct; sf = setiferous foveolae; sg = spermatic groove; St = sternite; sub = hypoproct; t = telopodite; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022762" httpUri="https://zenodo.org/record/14022762/files/figure.png" pageId="13" pageNumber="474">Figs 5F, G</figureCitation>
|
||
). Sternite sclerotized and well-visible (
|
||
<figureCitation id="13061893FFD9FFB0FD602B9ED279F829" box="[709,801,1941,1968]" captionStart="FIGURE 5" captionStartId="9.[151,250,1692,1716]" captionTargetBox="[416,1172,849,1669]" captionTargetId="figure-246@9.[416,1172,849,1669]" captionTargetPageId="9" captionText="FIGURE 5. Alluviobolus omega, sp. nov., paratype what (FMMC 5403), multi-layer photographs. A. Male anterior body, lateral view; B. Male leg pairs 1–7, ventral view; C. Male posterior body end with telson; D. Anterior gonopod, anterior view; E. Anterior gonopod, posterior view; F. Posterior gonopod, anterior view; G. Posterior gonopod, posterior view. Abbreviations: a = spinose membranous area; Apo = apodeme; av = paraprocts; b= lateral opening of the sperm canal; bb = basal branch of posterior gonopod; c = membranous fringe below the apex; Co = collum (ring 1); Cx = coxite; d = large membranous area located mesally; Gn = gnathochilarium; h = ‘hole’-like structure on the posterior gonopod; Md = basal joints of mandible; oz = ozopore Pre = epiproct; sf = setiferous foveolae; sg = spermatic groove; St = sternite; sub = hypoproct; t = telopodite; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022762" httpUri="https://zenodo.org/record/14022762/files/figure.png" pageId="13" pageNumber="474">Figs 5G</figureCitation>
|
||
,
|
||
<figureCitation id="13061893FFD9FFB0FC8E2B9ED216F836" box="[811,846,1941,1967]" captionStart="FIGURE 6" captionStartId="10.[151,250,1533,1557]" captionTargetBox="[151,1435,188,1510]" captionTargetId="figure-18@10.[151,1435,181,1510]" captionTargetPageId="10" captionText="FIGURE 6. Alluviobolus omega, sp. nov., holotype (FMMC 5458), drawings. A. Anterior gonopod, anterior view; B. Anterior gonopod, posterior view; C. Posterior gonopod, tip, anterior view; D. Posterior gonopod, tip, posterior view; E. Posterior gonopod, base, posterior view; F. Male midbody leg, posterior view; G. Female midbody leg, posterior view. Abbreviations: a = spinose membranous area; b= lateral opening of the sperm canal; bb = basal branch of posterior gonopod; c = membranous fringe below the apex; Cx = coxite; d = large membranous area located mesally; Fe = femur; fs-p = finger-shaped process; Gr = groove; h = ‘hole’-like structure on the posterior gonopod; Pre = prefemur; Po = postfemur; sg = spermatic groove; St = sternite; t = telopodite; Ta = tarsus; Ti = tibia. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022764" httpUri="https://zenodo.org/record/14022764/files/figure.png" pageId="13" pageNumber="474">6E</figureCitation>
|
||
). Coxite mesally with a single groove,protruding into a short stem towards telopodite. Telopodite longer than coxite, curved posteriorly (
|
||
<figureCitation id="13061893FFD9FFB0FB922BB2D5E8F84A" box="[1079,1200,1977,2004]" captionStart="FIGURE 5" captionStartId="9.[151,250,1692,1716]" captionTargetBox="[416,1172,849,1669]" captionTargetId="figure-246@9.[416,1172,849,1669]" captionTargetPageId="9" captionText="FIGURE 5. Alluviobolus omega, sp. nov., paratype what (FMMC 5403), multi-layer photographs. A. Male anterior body, lateral view; B. Male leg pairs 1–7, ventral view; C. Male posterior body end with telson; D. Anterior gonopod, anterior view; E. Anterior gonopod, posterior view; F. Posterior gonopod, anterior view; G. Posterior gonopod, posterior view. Abbreviations: a = spinose membranous area; Apo = apodeme; av = paraprocts; b= lateral opening of the sperm canal; bb = basal branch of posterior gonopod; c = membranous fringe below the apex; Co = collum (ring 1); Cx = coxite; d = large membranous area located mesally; Gn = gnathochilarium; h = ‘hole’-like structure on the posterior gonopod; Md = basal joints of mandible; oz = ozopore Pre = epiproct; sf = setiferous foveolae; sg = spermatic groove; St = sternite; sub = hypoproct; t = telopodite; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022762" httpUri="https://zenodo.org/record/14022762/files/figure.png" pageId="13" pageNumber="474">Figs 5F, G</figureCitation>
|
||
,
|
||
<figureCitation id="13061893FFD9FFB0FB192BB2D450F84A" box="[1212,1288,1977,2003]" captionStart="FIGURE 6" captionStartId="10.[151,250,1533,1557]" captionTargetBox="[151,1435,188,1510]" captionTargetId="figure-18@10.[151,1435,181,1510]" captionTargetPageId="10" captionText="FIGURE 6. Alluviobolus omega, sp. nov., holotype (FMMC 5458), drawings. A. Anterior gonopod, anterior view; B. Anterior gonopod, posterior view; C. Posterior gonopod, tip, anterior view; D. Posterior gonopod, tip, posterior view; E. Posterior gonopod, base, posterior view; F. Male midbody leg, posterior view; G. Female midbody leg, posterior view. Abbreviations: a = spinose membranous area; b= lateral opening of the sperm canal; bb = basal branch of posterior gonopod; c = membranous fringe below the apex; Cx = coxite; d = large membranous area located mesally; Fe = femur; fs-p = finger-shaped process; Gr = groove; h = ‘hole’-like structure on the posterior gonopod; Pre = prefemur; Po = postfemur; sg = spermatic groove; St = sternite; t = telopodite; Ta = tarsus; Ti = tibia. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022764" httpUri="https://zenodo.org/record/14022764/files/figure.png" pageId="13" pageNumber="474">6 C–E</figureCitation>
|
||
). Telopodites arranged parallel to one another. A torsion present in telopodites, sperm canal discharging apico-laterally (
|
||
<figureCitation id="13061893FFD9FFB0FAE02BD6D4C4F861" box="[1349,1436,2013,2040]" captionStart="FIGURE 5" captionStartId="9.[151,250,1692,1716]" captionTargetBox="[416,1172,849,1669]" captionTargetId="figure-246@9.[416,1172,849,1669]" captionTargetPageId="9" captionText="FIGURE 5. Alluviobolus omega, sp. nov., paratype what (FMMC 5403), multi-layer photographs. A. Male anterior body, lateral view; B. Male leg pairs 1–7, ventral view; C. Male posterior body end with telson; D. Anterior gonopod, anterior view; E. Anterior gonopod, posterior view; F. Posterior gonopod, anterior view; G. Posterior gonopod, posterior view. Abbreviations: a = spinose membranous area; Apo = apodeme; av = paraprocts; b= lateral opening of the sperm canal; bb = basal branch of posterior gonopod; c = membranous fringe below the apex; Co = collum (ring 1); Cx = coxite; d = large membranous area located mesally; Gn = gnathochilarium; h = ‘hole’-like structure on the posterior gonopod; Md = basal joints of mandible; oz = ozopore Pre = epiproct; sf = setiferous foveolae; sg = spermatic groove; St = sternite; sub = hypoproct; t = telopodite; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022762" httpUri="https://zenodo.org/record/14022762/files/figure.png" pageId="13" pageNumber="474">Fig. 5F,</figureCitation>
|
||
</paragraph>
|
||
<paragraph id="8B820416FFDAFFB3FF322C9CD4DCFF60" blockId="14.[151,1437,151,537]" pageId="14" pageNumber="475">
|
||
G), telopodites resembling the letter ‘Z’. Telopodite in anterior view centrally with a ‘hole-like’ structure (
|
||
<figureCitation id="13061893FFDAFFB3FA922C9CD4D7FF28" box="[1335,1423,151,177]" captionStart="FIGURE 6" captionStartId="10.[151,250,1533,1557]" captionTargetBox="[151,1435,188,1510]" captionTargetId="figure-18@10.[151,1435,181,1510]" captionTargetPageId="10" captionText="FIGURE 6. Alluviobolus omega, sp. nov., holotype (FMMC 5458), drawings. A. Anterior gonopod, anterior view; B. Anterior gonopod, posterior view; C. Posterior gonopod, tip, anterior view; D. Posterior gonopod, tip, posterior view; E. Posterior gonopod, base, posterior view; F. Male midbody leg, posterior view; G. Female midbody leg, posterior view. Abbreviations: a = spinose membranous area; b= lateral opening of the sperm canal; bb = basal branch of posterior gonopod; c = membranous fringe below the apex; Cx = coxite; d = large membranous area located mesally; Fe = femur; fs-p = finger-shaped process; Gr = groove; h = ‘hole’-like structure on the posterior gonopod; Pre = prefemur; Po = postfemur; sg = spermatic groove; St = sternite; t = telopodite; Ta = tarsus; Ti = tibia. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022764" httpUri="https://zenodo.org/record/14022764/files/figure.png" pageId="14" pageNumber="475">Fig. 6C</figureCitation>
|
||
), apex with four specific areas labelled a–d: (a) is a spinose membranous area, (b) the lateral opening of the sperm channel, (c) a membranous fringe below the apex and (d) a large membranous area located mesally (
|
||
<figureCitation id="13061893FFDAFFB3FB592CD4D42FFF60" box="[1276,1399,223,249]" captionStart="FIGURE 6" captionStartId="10.[151,250,1533,1557]" captionTargetBox="[151,1435,188,1510]" captionTargetId="figure-18@10.[151,1435,181,1510]" captionTargetPageId="10" captionText="FIGURE 6. Alluviobolus omega, sp. nov., holotype (FMMC 5458), drawings. A. Anterior gonopod, anterior view; B. Anterior gonopod, posterior view; C. Posterior gonopod, tip, anterior view; D. Posterior gonopod, tip, posterior view; E. Posterior gonopod, base, posterior view; F. Male midbody leg, posterior view; G. Female midbody leg, posterior view. Abbreviations: a = spinose membranous area; b= lateral opening of the sperm canal; bb = basal branch of posterior gonopod; c = membranous fringe below the apex; Cx = coxite; d = large membranous area located mesally; Fe = femur; fs-p = finger-shaped process; Gr = groove; h = ‘hole’-like structure on the posterior gonopod; Pre = prefemur; Po = postfemur; sg = spermatic groove; St = sternite; t = telopodite; Ta = tarsus; Ti = tibia. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022764" httpUri="https://zenodo.org/record/14022764/files/figure.png" pageId="14" pageNumber="475">Fig. 6C, D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C327579DFFDAFFB3FF622D09D249FD80" pageId="14" pageNumber="475" type="distribution">
|
||
<paragraph id="8B820416FFDAFFB3FF622D09D47FFE84" blockId="14.[151,1437,151,537]" box="[199,1319,258,285]" pageId="14" pageNumber="475">
|
||
<emphasis id="B949D804FFDAFFB3FF622D09D001FE85" bold="true" box="[199,345,258,284]" pageId="14" pageNumber="475">Distribution</emphasis>
|
||
. Intermediate rain-dry forests (
|
||
<bibRefCitation id="EFAC79E7FFDAFFB3FD132D08D2C3FE84" author="Moat, J. & Smith, P." box="[694,923,259,285]" pageId="14" pageNumber="475" refId="ref13788" refString="Moat, J. & Smith, P. (2007) Atlas of the Vegetation of Madagascar. Kew Publishing, London, 124 pp." type="book" year="2007">Moat & Smith 2007</bibRefCitation>
|
||
) in Vohibasia, Isalo and Vohimena.
|
||
</paragraph>
|
||
<paragraph id="8B820416FFDAFFB3FF622D2DD249FD80" blockId="14.[151,1437,151,537]" pageId="14" pageNumber="475">
|
||
<emphasis id="B949D804FFDAFFB3FF622D2DD088FED9" bold="true" box="[199,464,294,320]" pageId="14" pageNumber="475">Intraspecific variation</emphasis>
|
||
. The three populations at Vohibasia, Isalo and Vohimena differ from one another slightly in their size, ring number and gonopods. The population from Vohimena has 47 body rings, differs slightly in the colour pattern (
|
||
<figureCitation id="13061893FFDAFFB3FEE42D64D0E1FE10" box="[321,441,367,393]" captionStart="FIGURE 9" captionStartId="13.[151,250,1222,1246]" captionTargetBox="[510,1077,182,1198]" captionTargetId="figure-15@13.[510,1077,181,1198]" captionTargetPageId="13" captionText="FIGURE 9. Alluviobolus omega, sp. nov., population from Vohimena (FMMC 5396), multi-layer photographs.A. Male anterior body, lateral view; B. Midbody rings, lateral view; C. Male leg pairs 1–7, ventral view; D. Male posterior body end with telson; E. Anterior gonopod, anterior view; F. Anterior gonopod, posterior view; G. Right posterior gonopod, anterior view; H. Right posterior gonopod, posterior view. Abbreviations: a = spinose membranous area; av = paraprocts; b= lateral opening of the sperm canal; bb = basal branch of posterior gonopod; c = membranous fringe below the apex; Co = collum (ring 1); Cx = coxite; d = large membranous area located mesally; Gn = gnathochilarium; h = ‘hole’-like structure on the posterior gonopod; Md = basal joints of mandible; Me = mesozonite; Mt = metazonite; oz = ozopore; pe = pseudopenis; Pre = epiproct; sf = setiferous foveolae; sg = spermatic groove; St = sternite; sub = hypoproct; t = telopodite; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022774" httpUri="https://zenodo.org/record/14022774/files/figure.png" pageId="14" pageNumber="475">Fig. 9A, B</figureCitation>
|
||
), missing a coxal processes on male coxa 5 (
|
||
<figureCitation id="13061893FFDAFFB3FC0C2D64D558FE10" box="[937,1024,367,393]" captionStart="FIGURE 9" captionStartId="13.[151,250,1222,1246]" captionTargetBox="[510,1077,182,1198]" captionTargetId="figure-15@13.[510,1077,181,1198]" captionTargetPageId="13" captionText="FIGURE 9. Alluviobolus omega, sp. nov., population from Vohimena (FMMC 5396), multi-layer photographs.A. Male anterior body, lateral view; B. Midbody rings, lateral view; C. Male leg pairs 1–7, ventral view; D. Male posterior body end with telson; E. Anterior gonopod, anterior view; F. Anterior gonopod, posterior view; G. Right posterior gonopod, anterior view; H. Right posterior gonopod, posterior view. Abbreviations: a = spinose membranous area; av = paraprocts; b= lateral opening of the sperm canal; bb = basal branch of posterior gonopod; c = membranous fringe below the apex; Co = collum (ring 1); Cx = coxite; d = large membranous area located mesally; Gn = gnathochilarium; h = ‘hole’-like structure on the posterior gonopod; Md = basal joints of mandible; Me = mesozonite; Mt = metazonite; oz = ozopore; pe = pseudopenis; Pre = epiproct; sf = setiferous foveolae; sg = spermatic groove; St = sternite; sub = hypoproct; t = telopodite; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022774" httpUri="https://zenodo.org/record/14022774/files/figure.png" pageId="14" pageNumber="475">Fig. 9C</figureCitation>
|
||
), and in a completely red telson (
|
||
<figureCitation id="13061893FFDAFFB3FACA2D64D1E5FE34" captionStart="FIGURE 9" captionStartId="13.[151,250,1222,1246]" captionTargetBox="[510,1077,182,1198]" captionTargetId="figure-15@13.[510,1077,181,1198]" captionTargetPageId="13" captionText="FIGURE 9. Alluviobolus omega, sp. nov., population from Vohimena (FMMC 5396), multi-layer photographs.A. Male anterior body, lateral view; B. Midbody rings, lateral view; C. Male leg pairs 1–7, ventral view; D. Male posterior body end with telson; E. Anterior gonopod, anterior view; F. Anterior gonopod, posterior view; G. Right posterior gonopod, anterior view; H. Right posterior gonopod, posterior view. Abbreviations: a = spinose membranous area; av = paraprocts; b= lateral opening of the sperm canal; bb = basal branch of posterior gonopod; c = membranous fringe below the apex; Co = collum (ring 1); Cx = coxite; d = large membranous area located mesally; Gn = gnathochilarium; h = ‘hole’-like structure on the posterior gonopod; Md = basal joints of mandible; Me = mesozonite; Mt = metazonite; oz = ozopore; pe = pseudopenis; Pre = epiproct; sf = setiferous foveolae; sg = spermatic groove; St = sternite; sub = hypoproct; t = telopodite; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022774" httpUri="https://zenodo.org/record/14022774/files/figure.png" pageId="14" pageNumber="475">Fig. 9D</figureCitation>
|
||
). The anterior (
|
||
<figureCitation id="13061893FFDAFFB3FEC92D98D0B9FE34" box="[364,481,403,429]" captionStart="FIGURE 9" captionStartId="13.[151,250,1222,1246]" captionTargetBox="[510,1077,182,1198]" captionTargetId="figure-15@13.[510,1077,181,1198]" captionTargetPageId="13" captionText="FIGURE 9. Alluviobolus omega, sp. nov., population from Vohimena (FMMC 5396), multi-layer photographs.A. Male anterior body, lateral view; B. Midbody rings, lateral view; C. Male leg pairs 1–7, ventral view; D. Male posterior body end with telson; E. Anterior gonopod, anterior view; F. Anterior gonopod, posterior view; G. Right posterior gonopod, anterior view; H. Right posterior gonopod, posterior view. Abbreviations: a = spinose membranous area; av = paraprocts; b= lateral opening of the sperm canal; bb = basal branch of posterior gonopod; c = membranous fringe below the apex; Co = collum (ring 1); Cx = coxite; d = large membranous area located mesally; Gn = gnathochilarium; h = ‘hole’-like structure on the posterior gonopod; Md = basal joints of mandible; Me = mesozonite; Mt = metazonite; oz = ozopore; pe = pseudopenis; Pre = epiproct; sf = setiferous foveolae; sg = spermatic groove; St = sternite; sub = hypoproct; t = telopodite; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022774" httpUri="https://zenodo.org/record/14022774/files/figure.png" pageId="14" pageNumber="475">Fig. 9E, F</figureCitation>
|
||
) and posterior gonopods (
|
||
<figureCitation id="13061893FFDAFFB3FCAE2D98D2DEFE34" box="[779,902,403,429]" captionStart="FIGURE 9" captionStartId="13.[151,250,1222,1246]" captionTargetBox="[510,1077,182,1198]" captionTargetId="figure-15@13.[510,1077,181,1198]" captionTargetPageId="13" captionText="FIGURE 9. Alluviobolus omega, sp. nov., population from Vohimena (FMMC 5396), multi-layer photographs.A. Male anterior body, lateral view; B. Midbody rings, lateral view; C. Male leg pairs 1–7, ventral view; D. Male posterior body end with telson; E. Anterior gonopod, anterior view; F. Anterior gonopod, posterior view; G. Right posterior gonopod, anterior view; H. Right posterior gonopod, posterior view. Abbreviations: a = spinose membranous area; av = paraprocts; b= lateral opening of the sperm canal; bb = basal branch of posterior gonopod; c = membranous fringe below the apex; Co = collum (ring 1); Cx = coxite; d = large membranous area located mesally; Gn = gnathochilarium; h = ‘hole’-like structure on the posterior gonopod; Md = basal joints of mandible; Me = mesozonite; Mt = metazonite; oz = ozopore; pe = pseudopenis; Pre = epiproct; sf = setiferous foveolae; sg = spermatic groove; St = sternite; sub = hypoproct; t = telopodite; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022774" httpUri="https://zenodo.org/record/14022774/files/figure.png" pageId="14" pageNumber="475">Fig. 9G, H</figureCitation>
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) appear very similar to those of the
|
||
<typeStatus id="5486BAB4FFDAFFB3FA812D98D40CFE34" box="[1316,1364,403,429]" pageId="14" pageNumber="475">type</typeStatus>
|
||
series from Vohibasia. Future studies should elucidate if these are three different species, like what was observed in the giant pill-millipede genus
|
||
<taxonomicName id="4C3D7F95FFDAFFB3FE612DD0D33EFE6C" authorityName="Pocock" authorityYear="1895" box="[452,614,475,501]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="14" pageNumber="475" phylum="Arthropoda" rank="genus">
|
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<emphasis id="B949D804FFDAFFB3FE612DD0D33EFE6C" box="[452,614,475,501]" italics="true" pageId="14" pageNumber="475">Zoosphaerium</emphasis>
|
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</taxonomicName>
|
||
, where similar but distinct species can be found in Vohibasia, Isalo and Vohimena (Wesener 2009;
|
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<bibRefCitation id="EFAC79E7FFDAFFB3FE602DF4D25DFD80" author="Wesener, T. & Anilkumar, P. A." box="[453,773,511,537]" pageId="14" pageNumber="475" pagination="3 - 35" refId="ref15013" refString="Wesener, T. & Anilkumar, P. A. (2020) The millipedes collected by the Museum " La Specola " on Madagascar 1989 / 1991, with the description of three new species of giant pill-millipedes (Diplopoda, Sphaerotheriida, Arthrosphaeridae). ZooKeys, 930, 3 - 35. https: // doi. org / 10.3897 / zookeys. 930.47620" type="journal article" year="2020">Wesener & Anilkumar 2020</bibRefCitation>
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||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |