313 lines
60 KiB
XML
313 lines
60 KiB
XML
<document id="758C04DB54A10C1D3E9BC09DE440F8EE" ID-DOI="10.11646/zootaxa.5529.3.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="14022748" ID-ZooBank="716EDF67-C933-484D-911B-4585B11187A6" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1730453336952" checkinUser="plazi" docAuthor="Wesener, Thomas" docDate="2024" docId="0394B500FFDAFFADFF322FFDD595FC8C" docLanguage="en" docName="zootaxa.5529.3.3.pdf" docOrigin="Zootaxa 5529 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5529.3.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Eucarlia tampolo Wesener, 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="477" masterDocId="FFADCD78FFD4FFBDFFA52C0BD158FF99" masterDocTitle="Five new species and numerous locality records of Spirobolida millipedes from Madagascar (Diplopoda, Spirobolida, Pachybolidae)" masterLastPageNumber="486" masterPageNumber="461" pageNumber="475" updateTime="1730454045455" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="8DD8345FB35A734ACA021B77CB060232">
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<mods:title id="012533BF02FC645C9FC2613550C1E4A1">Five new species and numerous locality records of Spirobolida millipedes from Madagascar (Diplopoda, Spirobolida, Pachybolidae)</mods:title>
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<mods:namePart id="FB15785E2B5FD472CC040D8D04ABE6FF">Wesener, Thomas</mods:namePart>
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<mods:title id="55594DA8342558CE9B89E2B424BF66AD">Zootaxa</mods:title>
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<mods:part id="BC88D04E9C3177D6D173C862CC12A9A2">
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<mods:date id="CA5A3C705015AC6F91F2679A6D537010">2024</mods:date>
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<mods:number id="91FA9B479D38852C25DB835E98404EE4">2024-10-29</mods:number>
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<mods:number id="3C15EECD50DC8F4CC3703E77E3078372">5529</mods:number>
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<mods:start id="56B6BE96DD88FA763BE38A9274D36C89">461</mods:start>
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<mods:url id="04960AE5C2B1B118618C7F1C798A84E2">http://dx.doi.org/10.11646/zootaxa.5529.3.3</mods:url>
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<mods:classification id="CC25CC78A749BF09BC2BB2D4951D5B8B">journal article</mods:classification>
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<mods:identifier id="A4C10E90B98185B2C16EB3DC8845C4FF" type="DOI">10.11646/zootaxa.5529.3.3</mods:identifier>
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<mods:identifier id="201280608CB0E3519BC9DA21615E6FCF" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="13A9A1E513231EA52B0D3916DF38F305" type="Zenodo-Dep">14022748</mods:identifier>
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<mods:identifier id="5F949D2FCA8DE730FE056768DD73B1D3" type="ZooBank">716EDF67-C933-484D-911B-4585B11187A6</mods:identifier>
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</mods:mods>
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<treatment id="0394B500FFDAFFADFF322FFDD595FC8C" LSID="urn:lsid:plazi:treatment:0394B500FFDAFFADFF322FFDD595FC8C" httpUri="http://treatment.plazi.org/id/0394B500FFDAFFADFF322FFDD595FC8C" lastPageId="16" lastPageNumber="477" pageId="14" pageNumber="475">
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<subSubSection id="C327579DFFDAFFB3FF322FFDD086FB88" box="[151,478,1014,1041]" pageId="14" pageNumber="475" type="nomenclature">
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<paragraph id="8B820416FFDAFFB3FF322FFDD086FB88" blockId="14.[151,478,1014,1077]" box="[151,478,1014,1041]" pageId="14" pageNumber="475">
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<heading id="D0CAB37AFFDAFFB3FF322FFDD086FB88" bold="true" box="[151,478,1014,1041]" fontSize="11" level="1" pageId="14" pageNumber="475" reason="1">
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<emphasis id="B949D804FFDAFFB3FF322FFDD086FB88" bold="true" box="[151,478,1014,1041]" pageId="14" pageNumber="475">
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||
<emphasis id="B949D804FFDAFFB3FF322FFDD021FB88" bold="true" box="[151,377,1014,1041]" italics="true" pageId="14" pageNumber="475">
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||
?
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<taxonomicName id="4C3D7F95FFDAFFB3FF062FFDD021FB88" authorityName="Wesener" authorityYear="2024" box="[163,377,1014,1041]" class="Diplopoda" family="Pachybolidae" genus="Eucarlia" kingdom="Animalia" order="Spirobolida" pageId="14" pageNumber="475" phylum="Arthropoda" rank="species" species="tampolo" status="sp. nov.">Eucarlia tampolo</taxonomicName>
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</emphasis>
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<taxonomicNameLabel id="A27A657FFFDAFFB3FE252FFCD086FB88" box="[384,478,1015,1041]" pageId="14" pageNumber="475" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C327579DFFDAFFB3FF322810D045FBAC" box="[151,285,1051,1077]" pageId="14" pageNumber="475" type="description">
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||
<paragraph id="8B820416FFDAFFB3FF322810D045FBAC" blockId="14.[151,478,1014,1077]" box="[151,285,1051,1077]" pageId="14" pageNumber="475">
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<figureCitation id="13061893FFDAFFB3FF322810D045FBAC" box="[151,285,1051,1077]" captionStart="FIGURE 10" captionStartId="15.[151,250,1765,1789]" captionTargetBox="[166,1423,184,1741]" captionTargetId="figure-15@15.[164,1423,181,1741]" captionTargetPageId="15" captionText="FIGURE 10.?Eucarliatampolo, sp. nov., holotype (FMMC 7796), drawings.A. Gnathochilarium, ventral view; B. Coxosternite of female leg 1, posterior view; C. Coxosternite of female leg 2, posterior view; D. Male midbody leg, posterior view; E. Posterior body end with telson, lateral view; F. Anterior gonopod, anterior view; G. Anterior gonopod, posterior view; H. Right posterior gonopod, tip, anterior view; I. Right posterior gonopod, posterior view. Abbreviations: av = paraprocts; Cx = coxa; Cx-St = coxosternite; cx = coxite; Fe = femur; L = ledge on mentum of gnathochilarium; LL = lamella linguales; main = main branch of posterior telopod; Me = mentum; mes = mesal branch of posterior gonopod; pre = epiproct; Pre = prefemur; Po = postfemur; st = sternite; St = stipites; sub = hypoproct; t = telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022776" httpUri="https://zenodo.org/record/14022776/files/figure.png" pageId="14" pageNumber="475">Fig. 10A–H</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C327579DFFDAFFB3FF322869D2A4FB5C" pageId="14" pageNumber="475" type="materials_examined">
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||
<paragraph id="8B820416FFDAFFB3FF322869D279FB38" blockId="14.[151,1437,1122,2013]" pageId="14" pageNumber="475">
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||
<emphasis id="B949D804FFDAFFB3FF322869D021FBE5" bold="true" box="[151,377,1122,1148]" pageId="14" pageNumber="475">Material examined</emphasis>
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.
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<materialsCitation id="3B550E4BFFDAFFB3FE222868D279FB38" collectingDate="1997-04-03" collectingDateMax="1997-04-16" collectingDateMin="1997-04-03" collectorName="S. M. Goodman" elevation="10" latitude="-17.286667" location="Station Forestiere de Tampolo" longLatPrecision="127" longitude="49.408333" pageId="14" pageNumber="475" specimenCode="FMNH-INS 7796" specimenCount="1" stateProvince="Toamasina" typeStatus="holotype">
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<typeStatus id="5486BAB4FFDAFFB3FE222868D0ACFBE4" box="[391,500,1123,1149]" pageId="14" pageNumber="475" type="holotype">Holotype</typeStatus>
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||
: 1 M,
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||
<emphasis id="B949D804FFDAFFB3FD9C2869D25DFBE5" bold="true" box="[569,773,1122,1148]" pageId="14" pageNumber="475">
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||
<specimenCode id="DB9BAC6DFFDAFFB3FD9C2869D25DFBE5" box="[569,773,1122,1148]" collectionCode="FMNH-INS" pageId="14" pageNumber="475">FMNH-INS 7796</specimenCode>
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||
</emphasis>
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,
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<collectingRegion id="49F9CAF4FFDAFFB3FCB72868D2D6FBE4" box="[786,910,1123,1149]" country="Madagascar" name="Toamasina" pageId="14" pageNumber="475">Toamasina</collectingRegion>
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||
,
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||
<location id="8EE252CDFFDAFFB3FC3C2868D5B4FBE4" LSID="urn:lsid:plazi:treatment:0394B500FFDAFFADFF322FFDD595FC8C:8EE252CDFFDAFFB3FC3C2868D5B4FBE4" box="[921,1260,1123,1149]" latitude="-17.286667" longLatPrecision="127" longitude="49.408333" name="Station Forestiere de Tampolo" pageId="14" pageNumber="475" stateProvince="Toamasina">Station Forestiere de Tampolo</location>
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,
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||
<quantity id="4CC5A9F3FFDAFFB3FB522868D469FBE4" box="[1271,1329,1123,1149]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="475" unit="m" value="10.0">
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||
<elevation id="0010E325FFDAFFB3FB522868D469FBE4" box="[1271,1329,1123,1149]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="475" unit="m" value="10.0">10 m</elevation>
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</quantity>
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,
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<geoCoordinate id="EE0962D1FFDAFFB3FA982868D1F0FB38" degrees="17" direction="south" minutes="17.2" orientation="latitude" pageId="14" pageNumber="475" precision="92" value="-17.286667">17°17.2’ S</geoCoordinate>
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,
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<geoCoordinate id="EE0962D1FFDAFFB3FF10288DD076FB38" box="[181,302,1158,1185]" degrees="49" direction="east" minutes="24.5" orientation="longitude" pageId="14" pageNumber="475" precision="92" value="49.408333">49°24.5’ E</geoCoordinate>
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, coll.
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<collectorName id="26C861C0FFDAFFB3FED7288CD379FB38" box="[370,545,1159,1185]" pageId="14" pageNumber="475">S.M. Goodman</collectorName>
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,
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<collectingMethod id="527C7C01FFDAFFB3FD89288CD325FB38" box="[556,637,1159,1185]" pageId="14" pageNumber="475">pitfalls</collectingMethod>
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,
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<date id="FF8322D6FFDAFFB3FD2D288CD246FB38" box="[648,798,1159,1185]" pageId="14" pageNumber="475" value="1997-04-03" valueMax="1997-04-16" valueMin="1997-04-03">
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||
<collectingDate id="EFC7DB3EFFDAFFB3FD2D288CD246FB38" box="[648,798,1159,1185]" pageId="14" pageNumber="475" value="1997-04-03" valueMax="1997-04-16" valueMin="1997-04-03">3–16.iv.1997</collectingDate>
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</date>
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||
.
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||
</materialsCitation>
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||
</paragraph>
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<paragraph id="8B820416FFDAFFB3FF6228A0D2A4FB5C" blockId="14.[151,1437,1122,2013]" box="[199,1020,1194,1221]" pageId="14" pageNumber="475">
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||
<materialsCitation id="3B550E4BFFDAFFB3FF6228A0D2A0FB5C" box="[199,1016,1194,1221]" collectingDate="1997-04-03" collectingDateMax="1997-04-16" collectingDateMin="1997-04-03" collectorName="S. M. Goodman" elevation="10" latitude="-17.286667" location="Station Forestiere de Tampolo" longLatPrecision="127" longitude="49.408333" pageId="14" pageNumber="475" specimenCode="FMNH-INS 7796" specimenCount="1" stateProvince="Toamasina" typeStatus="paratype">
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||
<typeStatus id="5486BAB4FFDAFFB3FF6228A0D060FB5C" box="[199,312,1195,1221]" pageId="14" pageNumber="475" type="paratype">Paratypes</typeStatus>
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||
: 4 M (mature?), 5 F,
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||
<emphasis id="B949D804FFDAFFB3FD8628A1D3B7FB5D" bold="true" box="[547,751,1194,1220]" pageId="14" pageNumber="475">
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||
<specimenCode id="DB9BAC6DFFDAFFB3FD8628A1D3B7FB5D" box="[547,751,1194,1220]" collectionCode="FMNH-INS" pageId="14" pageNumber="475">FMNH-INS 7796</specimenCode>
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</emphasis>
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, same data as holotype
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C327579DFFDAFFB3FF6228C5D0CAFA94" pageId="14" pageNumber="475" type="etymology">
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<paragraph id="8B820416FFDAFFB3FF6228C5D0CAFA94" blockId="14.[151,1437,1122,2013]" pageId="14" pageNumber="475">
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<emphasis id="B949D804FFDAFFB3FF6228C5D0C6FB71" bold="true" box="[199,414,1230,1256]" pageId="14" pageNumber="475">Derivatio nominis</emphasis>
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. tampolo, noun in apposition, after the littoral forest of Tampolo, the
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<typeStatus id="5486BAB4FFDAFFB3FB0A28C4D587FB70" box="[1199,1247,1231,1257]" pageId="14" pageNumber="475">type</typeStatus>
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and only known locality of the species.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C327579DFFDAFFB3FF62291DD2CFF9B4" pageId="14" pageNumber="475" type="diagnosis">
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||
<paragraph id="8B820416FFDAFFB3FF62291DD2CFF9B4" blockId="14.[151,1437,1122,2013]" pageId="14" pageNumber="475">
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||
<emphasis id="B949D804FFDAFFB3FF62291DD062FAA9" bold="true" box="[199,314,1302,1328]" pageId="14" pageNumber="475">Diagnosis</emphasis>
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.?
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<taxonomicName id="4C3D7F95FFDAFFB3FEFD291CD378FAA8" authorityName="Wesener" authorityYear="2024" box="[344,544,1303,1329]" class="Diplopoda" family="Pachybolidae" genus="Eucarlia" kingdom="Animalia" order="Spirobolida" pageId="14" pageNumber="475" phylum="Arthropoda" rank="species" species="tampolo" status="sp. nov.">
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<emphasis id="B949D804FFDAFFB3FEFD291CD378FAA8" box="[344,544,1303,1329]" italics="true" pageId="14" pageNumber="475">Eucarlia tampolo</emphasis>
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</taxonomicName>
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<emphasis id="B949D804FFDAFFB3FD8C291CD3DEFAA8" bold="true" box="[553,646,1303,1329]" pageId="14" pageNumber="475">
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||
<taxonomicNameLabel id="A27A657FFFDAFFB3FD8C291CD3DEFAA8" box="[553,646,1303,1329]" pageId="14" pageNumber="475" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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shares the presence of a strong transverse ridge across the mentum, separating a basal, transversely striate part and an apical, smooth part, which is recessed when the gnathochilarium is viewed from below (
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<figureCitation id="13061893FFDAFFB3FE382954D35CFAE0" box="[413,516,1375,1401]" captionStart="FIGURE 10" captionStartId="15.[151,250,1765,1789]" captionTargetBox="[166,1423,184,1741]" captionTargetId="figure-15@15.[164,1423,181,1741]" captionTargetPageId="15" captionText="FIGURE 10.?Eucarliatampolo, sp. nov., holotype (FMMC 7796), drawings.A. Gnathochilarium, ventral view; B. Coxosternite of female leg 1, posterior view; C. Coxosternite of female leg 2, posterior view; D. Male midbody leg, posterior view; E. Posterior body end with telson, lateral view; F. Anterior gonopod, anterior view; G. Anterior gonopod, posterior view; H. Right posterior gonopod, tip, anterior view; I. Right posterior gonopod, posterior view. Abbreviations: av = paraprocts; Cx = coxa; Cx-St = coxosternite; cx = coxite; Fe = femur; L = ledge on mentum of gnathochilarium; LL = lamella linguales; main = main branch of posterior telopod; Me = mentum; mes = mesal branch of posterior gonopod; pre = epiproct; Pre = prefemur; Po = postfemur; st = sternite; St = stipites; sub = hypoproct; t = telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022776" httpUri="https://zenodo.org/record/14022776/files/figure.png" pageId="14" pageNumber="475">Fig. 10A</figureCitation>
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||
) with species of the genera
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||
<taxonomicName id="4C3D7F95FFDAFFB3FCE42954D5D4FAE0" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[833,1164,1375,1401]" class="Diplopoda" family="Pachybolidae" genus="Flagellobolus" kingdom="Animalia" order="Spirobolida" pageId="14" pageNumber="475" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B949D804FFDAFFB3FCE42954D285FAE0" box="[833,989,1375,1401]" italics="true" pageId="14" pageNumber="475">Flagellobolus</emphasis>
|
||
Wesener, 2009
|
||
</taxonomicName>
|
||
,
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||
<taxonomicName id="4C3D7F95FFDAFFB3FB3D2954D18AFA04" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="14" pageNumber="475" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B949D804FFDAFFB3FB3D2954D477FAE0" box="[1176,1327,1375,1401]" italics="true" pageId="14" pageNumber="475">Riotintobolus</emphasis>
|
||
Wesener, 2009
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3D7F95FFDAFFB3FF7A2988D330FA04" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[223,616,1411,1437]" class="Diplopoda" family="Pachybolidae" genus="Pseudocentrobolus" kingdom="Animalia" order="Spirobolida" pageId="14" pageNumber="475" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B949D804FFDAFFB3FF7A2988D0EDFA04" box="[223,437,1411,1437]" italics="true" pageId="14" pageNumber="475">Pseudocentrobolus</emphasis>
|
||
Wesener, 2009
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3D7F95FFDAFFB3FDD02988D2E6FA04" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[629,958,1411,1437]" class="Diplopoda" family="Pachybolidae" genus="Granitobolus" kingdom="Animalia" order="Spirobolida" pageId="14" pageNumber="475" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B949D804FFDAFFB3FDD02988D252FA04" box="[629,778,1411,1437]" italics="true" pageId="14" pageNumber="475">Granitobolus</emphasis>
|
||
Wesener, 2009
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3D7F95FFDAFFB3FC6E2988D459FA04" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[971,1281,1411,1437]" class="Diplopoda" family="Pachybolidae" genus="Caprobolus" kingdom="Animalia" order="Spirobolida" pageId="14" pageNumber="475" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B949D804FFDAFFB3FC6E2988D516FA04" box="[971,1102,1411,1437]" italics="true" pageId="14" pageNumber="475">Caprobolus</emphasis>
|
||
Wesener, 2009
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||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3D7F95FFDAFFB3FAAA2988D018FA58" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" class="Diplopoda" family="Pachybolidae" genus="Alluviobolus" kingdom="Animalia" order="Spirobolida" pageId="14" pageNumber="475" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B949D804FFDAFFB3FAAA2988D4C5FA04" box="[1295,1437,1411,1437]" italics="true" pageId="14" pageNumber="475">Alluviobolus</emphasis>
|
||
Wesener, 2009
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3D7F95FFDAFFB3FEE829ACD3DAFA58" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[333,642,1447,1473]" class="Diplopoda" family="Pachybolidae" genus="Ostinobolus" kingdom="Animalia" order="Spirobolida" pageId="14" pageNumber="475" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B949D804FFDAFFB3FEE829ACD08CFA58" box="[333,468,1447,1473]" italics="true" pageId="14" pageNumber="475">Ostinobolus</emphasis>
|
||
Wesener, 2009
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B949D804FFDAFFB3FD1929ACD205FA58" box="[700,861,1447,1473]" italics="true" pageId="14" pageNumber="475">Zehnterobolus</emphasis>
|
||
Wesener, 2009. Sperm canal of posterior gonopod apically free (
|
||
<figureCitation id="13061893FFDAFFB3FE9129C0D0ECFA7C" box="[308,436,1483,1509]" captionStart="FIGURE 10" captionStartId="15.[151,250,1765,1789]" captionTargetBox="[166,1423,184,1741]" captionTargetId="figure-15@15.[164,1423,181,1741]" captionTargetPageId="15" captionText="FIGURE 10.?Eucarliatampolo, sp. nov., holotype (FMMC 7796), drawings.A. Gnathochilarium, ventral view; B. Coxosternite of female leg 1, posterior view; C. Coxosternite of female leg 2, posterior view; D. Male midbody leg, posterior view; E. Posterior body end with telson, lateral view; F. Anterior gonopod, anterior view; G. Anterior gonopod, posterior view; H. Right posterior gonopod, tip, anterior view; I. Right posterior gonopod, posterior view. Abbreviations: av = paraprocts; Cx = coxa; Cx-St = coxosternite; cx = coxite; Fe = femur; L = ledge on mentum of gnathochilarium; LL = lamella linguales; main = main branch of posterior telopod; Me = mentum; mes = mesal branch of posterior gonopod; pre = epiproct; Pre = prefemur; Po = postfemur; st = sternite; St = stipites; sub = hypoproct; t = telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022776" httpUri="https://zenodo.org/record/14022776/files/figure.png" pageId="14" pageNumber="475">Fig. 10H, I</figureCitation>
|
||
), like in species of
|
||
<taxonomicName id="4C3D7F95FFDAFFB3FD3129C0D268FA7C" authorityName="Wesener" authorityYear="2009" box="[660,816,1483,1509]" class="Diplopoda" family="Pachybolidae" genus="Flagellobolus" kingdom="Animalia" order="Spirobolida" pageId="14" pageNumber="475" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B949D804FFDAFFB3FD3129C0D268FA7C" box="[660,816,1483,1509]" italics="true" pageId="14" pageNumber="475">Flagellobolus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3D7F95FFDAFFB3FCCC29C0D2B4FA7C" authorityName="Wesener" authorityYear="2009" box="[873,1004,1483,1509]" class="Diplopoda" family="Pachybolidae" genus="Caprobolus" kingdom="Animalia" order="Spirobolida" pageId="14" pageNumber="475" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B949D804FFDAFFB3FCCC29C0D2B4FA7C" box="[873,1004,1483,1509]" italics="true" pageId="14" pageNumber="475">Caprobolus</emphasis>
|
||
</taxonomicName>
|
||
, but differs from all known Malagasy
|
||
<taxonomicName id="4C3D7F95FFDAFFB3FF3229E4D042F990" authorityName="Bollman" authorityYear="1893" box="[151,282,1519,1545]" class="Diplopoda" kingdom="Animalia" order="Spirobolida" pageId="14" pageNumber="475" phylum="Arthropoda" rank="order">Spirobolida</taxonomicName>
|
||
species in the combination of a simple wide anterior gonopod lacking projections or processes (
|
||
<figureCitation id="13061893FFDAFFB3FACA29E4D1BCF9B4" captionStart="FIGURE 10" captionStartId="15.[151,250,1765,1789]" captionTargetBox="[166,1423,184,1741]" captionTargetId="figure-15@15.[164,1423,181,1741]" captionTargetPageId="15" captionText="FIGURE 10.?Eucarliatampolo, sp. nov., holotype (FMMC 7796), drawings.A. Gnathochilarium, ventral view; B. Coxosternite of female leg 1, posterior view; C. Coxosternite of female leg 2, posterior view; D. Male midbody leg, posterior view; E. Posterior body end with telson, lateral view; F. Anterior gonopod, anterior view; G. Anterior gonopod, posterior view; H. Right posterior gonopod, tip, anterior view; I. Right posterior gonopod, posterior view. Abbreviations: av = paraprocts; Cx = coxa; Cx-St = coxosternite; cx = coxite; Fe = femur; L = ledge on mentum of gnathochilarium; LL = lamella linguales; main = main branch of posterior telopod; Me = mentum; mes = mesal branch of posterior gonopod; pre = epiproct; Pre = prefemur; Po = postfemur; st = sternite; St = stipites; sub = hypoproct; t = telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022776" httpUri="https://zenodo.org/record/14022776/files/figure.png" pageId="14" pageNumber="475">Fig. 10F, G</figureCitation>
|
||
)) with a special shape of the posterior gonopod (
|
||
<figureCitation id="13061893FFDAFFB3FCA22A18D2D1F9B4" box="[775,905,1555,1581]" captionStart="FIGURE 10" captionStartId="15.[151,250,1765,1789]" captionTargetBox="[166,1423,184,1741]" captionTargetId="figure-15@15.[164,1423,181,1741]" captionTargetPageId="15" captionText="FIGURE 10.?Eucarliatampolo, sp. nov., holotype (FMMC 7796), drawings.A. Gnathochilarium, ventral view; B. Coxosternite of female leg 1, posterior view; C. Coxosternite of female leg 2, posterior view; D. Male midbody leg, posterior view; E. Posterior body end with telson, lateral view; F. Anterior gonopod, anterior view; G. Anterior gonopod, posterior view; H. Right posterior gonopod, tip, anterior view; I. Right posterior gonopod, posterior view. Abbreviations: av = paraprocts; Cx = coxa; Cx-St = coxosternite; cx = coxite; Fe = femur; L = ledge on mentum of gnathochilarium; LL = lamella linguales; main = main branch of posterior telopod; Me = mentum; mes = mesal branch of posterior gonopod; pre = epiproct; Pre = prefemur; Po = postfemur; st = sternite; St = stipites; sub = hypoproct; t = telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022776" httpUri="https://zenodo.org/record/14022776/files/figure.png" pageId="14" pageNumber="475">Figs 10H, I</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C327579DFFDAFFADFF622A3DD23CFD54" lastPageId="16" lastPageNumber="477" pageId="14" pageNumber="475" type="description">
|
||
<paragraph id="8B820416FFDAFFB3FF622A3DD009F9C9" blockId="14.[151,1437,1122,2013]" box="[199,337,1590,1616]" pageId="14" pageNumber="475">
|
||
<emphasis id="B949D804FFDAFFB3FF622A3DD009F9C9" bold="true" box="[199,337,1590,1616]" pageId="14" pageNumber="475">Description</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B820416FFDAFFB3FF622A50D510F900" blockId="14.[151,1437,1122,2013]" pageId="14" pageNumber="475">
|
||
Measurements: All 43+0 rings. Male
|
||
<typeStatus id="5486BAB4FFDAFFB3FDC02A50D39FF9EC" box="[613,711,1627,1653]" pageId="14" pageNumber="475" type="holotype">holotype</typeStatus>
|
||
<quantity id="4CC5A9F3FFDAFFB3FD682A50D243F9EC" box="[717,795,1627,1653]" metricMagnitude="-2" metricUnit="m" metricValue="3.8" pageId="14" pageNumber="475" unit="mm" value="38.0">38 mm</quantity>
|
||
long, 3.2 wide. Largest female circa
|
||
<quantity id="4CC5A9F3FFDAFFB3FB102A50D45BF9EC" box="[1205,1283,1627,1653]" metricMagnitude="-2" metricUnit="m" metricValue="4.6" pageId="14" pageNumber="475" unit="mm" value="46.0">46 mm</quantity>
|
||
long,
|
||
<quantity id="4CC5A9F3FFDAFFB3FAED2A50D4C5F9EC" box="[1352,1437,1627,1653]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="14" pageNumber="475" unit="mm" value="4.8">4.8 mm</quantity>
|
||
wide. Habitus small and slender, body rings like
|
||
<taxonomicName id="4C3D7F95FFDAFFB3FD1F2A74D2A5F900" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[698,1021,1663,1689]" class="Diplopoda" family="Pachybolidae" genus="Granitobolus" kingdom="Animalia" order="Spirobolida" pageId="14" pageNumber="475" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B949D804FFDAFFB3FD1F2A74D217F900" box="[698,847,1663,1689]" italics="true" pageId="14" pageNumber="475">Granitobolus</emphasis>
|
||
Wesener, 2009
|
||
</taxonomicName>
|
||
, thick.
|
||
</paragraph>
|
||
<paragraph id="8B820416FFDAFFB3FF622AA8D563F978" blockId="14.[151,1437,1122,2013]" pageId="14" pageNumber="475">Colour strongly faded, mesozonites dark, metazonite white. Collum dark except for posterior margin. Anal scale and part of anal valves white, rest of telson black. Legs and antennae whitish.</paragraph>
|
||
<paragraph id="8B820416FFDAFFB3FF622AE0D04DF844" blockId="14.[151,1437,1122,2013]" pageId="14" pageNumber="475">
|
||
Head: Eyes very large, male eyes with 35 ommatidia, arranged in 7 rows, female similar. Antennae short, male antennae reaching back to third, female to second ring. Mandible basal joint without projection. Gnathochilarium with ridge, stipites each with three marginal setae, lamellae linguales each with two setae located behind one another (
|
||
<figureCitation id="13061893FFDAFFB3FF3A2B5CD052F8E8" box="[159,266,1879,1905]" captionStart="FIGURE 10" captionStartId="15.[151,250,1765,1789]" captionTargetBox="[166,1423,184,1741]" captionTargetId="figure-15@15.[164,1423,181,1741]" captionTargetPageId="15" captionText="FIGURE 10.?Eucarliatampolo, sp. nov., holotype (FMMC 7796), drawings.A. Gnathochilarium, ventral view; B. Coxosternite of female leg 1, posterior view; C. Coxosternite of female leg 2, posterior view; D. Male midbody leg, posterior view; E. Posterior body end with telson, lateral view; F. Anterior gonopod, anterior view; G. Anterior gonopod, posterior view; H. Right posterior gonopod, tip, anterior view; I. Right posterior gonopod, posterior view. Abbreviations: av = paraprocts; Cx = coxa; Cx-St = coxosternite; cx = coxite; Fe = femur; L = ledge on mentum of gnathochilarium; LL = lamella linguales; main = main branch of posterior telopod; Me = mentum; mes = mesal branch of posterior gonopod; pre = epiproct; Pre = prefemur; Po = postfemur; st = sternite; St = stipites; sub = hypoproct; t = telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022776" httpUri="https://zenodo.org/record/14022776/files/figure.png" pageId="14" pageNumber="475">Fig. 10A</figureCitation>
|
||
). Female leg 1 with fused coxo-sternite (
|
||
<figureCitation id="13061893FFDAFFB3FD442B5CD211F8E8" box="[737,841,1879,1905]" captionStart="FIGURE 10" captionStartId="15.[151,250,1765,1789]" captionTargetBox="[166,1423,184,1741]" captionTargetId="figure-15@15.[164,1423,181,1741]" captionTargetPageId="15" captionText="FIGURE 10.?Eucarliatampolo, sp. nov., holotype (FMMC 7796), drawings.A. Gnathochilarium, ventral view; B. Coxosternite of female leg 1, posterior view; C. Coxosternite of female leg 2, posterior view; D. Male midbody leg, posterior view; E. Posterior body end with telson, lateral view; F. Anterior gonopod, anterior view; G. Anterior gonopod, posterior view; H. Right posterior gonopod, tip, anterior view; I. Right posterior gonopod, posterior view. Abbreviations: av = paraprocts; Cx = coxa; Cx-St = coxosternite; cx = coxite; Fe = femur; L = ledge on mentum of gnathochilarium; LL = lamella linguales; main = main branch of posterior telopod; Me = mentum; mes = mesal branch of posterior gonopod; pre = epiproct; Pre = prefemur; Po = postfemur; st = sternite; St = stipites; sub = hypoproct; t = telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022776" httpUri="https://zenodo.org/record/14022776/files/figure.png" pageId="14" pageNumber="475">Fig. 10B</figureCitation>
|
||
), leg 2 also with coxo-sternite, coxal part elongated (
|
||
<figureCitation id="13061893FFDAFFB3FF3A2B70D050F80C" box="[159,264,1915,1941]" captionStart="FIGURE 10" captionStartId="15.[151,250,1765,1789]" captionTargetBox="[166,1423,184,1741]" captionTargetId="figure-15@15.[164,1423,181,1741]" captionTargetPageId="15" captionText="FIGURE 10.?Eucarliatampolo, sp. nov., holotype (FMMC 7796), drawings.A. Gnathochilarium, ventral view; B. Coxosternite of female leg 1, posterior view; C. Coxosternite of female leg 2, posterior view; D. Male midbody leg, posterior view; E. Posterior body end with telson, lateral view; F. Anterior gonopod, anterior view; G. Anterior gonopod, posterior view; H. Right posterior gonopod, tip, anterior view; I. Right posterior gonopod, posterior view. Abbreviations: av = paraprocts; Cx = coxa; Cx-St = coxosternite; cx = coxite; Fe = femur; L = ledge on mentum of gnathochilarium; LL = lamella linguales; main = main branch of posterior telopod; Me = mentum; mes = mesal branch of posterior gonopod; pre = epiproct; Pre = prefemur; Po = postfemur; st = sternite; St = stipites; sub = hypoproct; t = telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022776" httpUri="https://zenodo.org/record/14022776/files/figure.png" pageId="14" pageNumber="475">Fig. 10C</figureCitation>
|
||
). Legs 8+ in both sexes at each podomere, except tarsus, with single ventral spine. Tarsus in male (
|
||
<figureCitation id="13061893FFDAFFB3FACA2B70D191F820" captionStart="FIGURE 10" captionStartId="15.[151,250,1765,1789]" captionTargetBox="[166,1423,184,1741]" captionTargetId="figure-15@15.[164,1423,181,1741]" captionTargetPageId="15" captionText="FIGURE 10.?Eucarliatampolo, sp. nov., holotype (FMMC 7796), drawings.A. Gnathochilarium, ventral view; B. Coxosternite of female leg 1, posterior view; C. Coxosternite of female leg 2, posterior view; D. Male midbody leg, posterior view; E. Posterior body end with telson, lateral view; F. Anterior gonopod, anterior view; G. Anterior gonopod, posterior view; H. Right posterior gonopod, tip, anterior view; I. Right posterior gonopod, posterior view. Abbreviations: av = paraprocts; Cx = coxa; Cx-St = coxosternite; cx = coxite; Fe = femur; L = ledge on mentum of gnathochilarium; LL = lamella linguales; main = main branch of posterior telopod; Me = mentum; mes = mesal branch of posterior gonopod; pre = epiproct; Pre = prefemur; Po = postfemur; st = sternite; St = stipites; sub = hypoproct; t = telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022776" httpUri="https://zenodo.org/record/14022776/files/figure.png" pageId="14" pageNumber="475">Fig. 10D</figureCitation>
|
||
) with tarsal pad (except for posterior legs) and a pair of ventral spines. Claw large, apical spine quite small (
|
||
<figureCitation id="13061893FFDAFFB3FF3A2BC8D050F844" box="[159,264,1987,2013]" captionStart="FIGURE 10" captionStartId="15.[151,250,1765,1789]" captionTargetBox="[166,1423,184,1741]" captionTargetId="figure-15@15.[164,1423,181,1741]" captionTargetPageId="15" captionText="FIGURE 10.?Eucarliatampolo, sp. nov., holotype (FMMC 7796), drawings.A. Gnathochilarium, ventral view; B. Coxosternite of female leg 1, posterior view; C. Coxosternite of female leg 2, posterior view; D. Male midbody leg, posterior view; E. Posterior body end with telson, lateral view; F. Anterior gonopod, anterior view; G. Anterior gonopod, posterior view; H. Right posterior gonopod, tip, anterior view; I. Right posterior gonopod, posterior view. Abbreviations: av = paraprocts; Cx = coxa; Cx-St = coxosternite; cx = coxite; Fe = femur; L = ledge on mentum of gnathochilarium; LL = lamella linguales; main = main branch of posterior telopod; Me = mentum; mes = mesal branch of posterior gonopod; pre = epiproct; Pre = prefemur; Po = postfemur; st = sternite; St = stipites; sub = hypoproct; t = telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022776" httpUri="https://zenodo.org/record/14022776/files/figure.png" pageId="14" pageNumber="475">Fig. 10D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF42549EFFDBFFB2FF322AEED07CF861" ID-DOI="http://doi.org/10.5281/zenodo.14022776" ID-Zenodo-Dep="14022776" httpUri="https://zenodo.org/record/14022776/files/figure.png" pageId="15" pageNumber="476" startId="15.[151,250,1765,1789]" targetBox="[166,1423,184,1741]" targetPageId="15" targetType="figure">
|
||
<paragraph id="8B820416FFDBFFB2FF322AEED07CF861" blockId="15.[151,1436,1765,2041]" pageId="15" pageNumber="476">
|
||
<emphasis id="B949D804FFDBFFB2FF322AEED045F964" bold="true" box="[151,285,1765,1789]" pageId="15" pageNumber="476">FIGURE 10.</emphasis>
|
||
?
|
||
<taxonomicName id="4C3D7F95FFDBFFB2FE882AEED084F964" authorityName="Wesener" authorityYear="2024" box="[301,476,1765,1789]" class="Diplopoda" family="Pachybolidae" genus="Eucarlia" kingdom="Animalia" order="Spirobolida" pageId="15" pageNumber="476" phylum="Arthropoda" rank="species" species="tampolo" status="sp. nov.">
|
||
<emphasis id="B949D804FFDBFFB2FE882AEED084F964" box="[301,476,1765,1789]" italics="true" pageId="15" pageNumber="476">Eucarlia tampolo</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B949D804FFDBFFB2FE432AEED36DF964" bold="true" box="[486,565,1765,1789]" pageId="15" pageNumber="476">
|
||
<taxonomicNameLabel id="A27A657FFFDBFFB2FE432AEED36DF964" box="[486,565,1765,1789]" pageId="15" pageNumber="476" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype (
|
||
<emphasis id="B949D804FFDBFFB2FD012AEED275F964" bold="true" box="[676,813,1765,1789]" pageId="15" pageNumber="476">FMMC 7796</emphasis>
|
||
), drawings.
|
||
<emphasis id="B949D804FFDBFFB2FC022AEED2E1F964" bold="true" box="[935,953,1765,1789]" pageId="15" pageNumber="476">A</emphasis>
|
||
. Gnathochilarium, ventral view;
|
||
<emphasis id="B949D804FFDBFFB2FAA52AEED449F964" bold="true" box="[1280,1297,1765,1789]" pageId="15" pageNumber="476">B</emphasis>
|
||
. Coxosternite of female leg 1, posterior view;
|
||
<emphasis id="B949D804FFDBFFB2FE482B02D0A7F8B8" bold="true" box="[493,511,1801,1825]" pageId="15" pageNumber="476">C</emphasis>
|
||
. Coxosternite of female leg 2, posterior view;
|
||
<emphasis id="B949D804FFDBFFB2FC4A2B02D550F8B8" bold="true" box="[1007,1032,1801,1825]" pageId="15" pageNumber="476">D.</emphasis>
|
||
Male midbody leg, posterior view;
|
||
<emphasis id="B949D804FFDBFFB2FA202B02D4CEF8B8" bold="true" box="[1413,1430,1801,1825]" pageId="15" pageNumber="476">E</emphasis>
|
||
. Posterior body end with telson, lateral view;
|
||
<emphasis id="B949D804FFDBFFB2FDFF2B26D331F8DC" bold="true" box="[602,617,1837,1861]" pageId="15" pageNumber="476">F</emphasis>
|
||
. Anterior gonopod, anterior view;
|
||
<emphasis id="B949D804FFDBFFB2FC622B26D283F8DC" bold="true" box="[967,987,1837,1861]" pageId="15" pageNumber="476">G</emphasis>
|
||
. Anterior gonopod, posterior view;
|
||
<emphasis id="B949D804FFDBFFB2FAE12B26D400F8DC" bold="true" box="[1348,1368,1837,1861]" pageId="15" pageNumber="476">H</emphasis>
|
||
. Right posterior gonopod, tip, anterior view;
|
||
<emphasis id="B949D804FFDBFFB2FDB92B5AD37EF8F0" bold="true" box="[540,550,1873,1897]" pageId="15" pageNumber="476">I</emphasis>
|
||
. Right posterior gonopod, posterior view.
|
||
<emphasis id="B949D804FFDBFFB2FC7D2B5AD529F8F0" bold="true" box="[984,1137,1873,1897]" pageId="15" pageNumber="476">Abbreviations</emphasis>
|
||
: av = paraprocts; Cx = coxa; Cx-St = coxosternite; cx = coxite; Fe = femur; L = ledge on mentum of gnathochilarium; LL = lamella linguales; main = main branch of posterior telopod; Me = mentum; mes = mesal branch of posterior gonopod; pre = epiproct; Pre = prefemur; Po = postfemur; st = sternite; St = stipites; sub = hypoproct; t = telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; Vu = vulva.
|
||
<emphasis id="B949D804FFDBFFB2FAC12BB6D19FF860" bold="true" pageId="15" pageNumber="476">Scale bars</emphasis>
|
||
= 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B820416FFC4FFADFF622C9CD57CFF28" blockId="16.[151,1437,151,789]" box="[199,1060,151,177]" pageId="16" pageNumber="477">
|
||
Ozopores starting at ring 6, located on or even posterior of suture (
|
||
<figureCitation id="13061893FFC4FFADFC142C9CD54FFF28" box="[945,1047,151,177]" captionStart="FIGURE 10" captionStartId="15.[151,250,1765,1789]" captionTargetBox="[166,1423,184,1741]" captionTargetId="figure-15@15.[164,1423,181,1741]" captionTargetPageId="15" captionText="FIGURE 10.?Eucarliatampolo, sp. nov., holotype (FMMC 7796), drawings.A. Gnathochilarium, ventral view; B. Coxosternite of female leg 1, posterior view; C. Coxosternite of female leg 2, posterior view; D. Male midbody leg, posterior view; E. Posterior body end with telson, lateral view; F. Anterior gonopod, anterior view; G. Anterior gonopod, posterior view; H. Right posterior gonopod, tip, anterior view; I. Right posterior gonopod, posterior view. Abbreviations: av = paraprocts; Cx = coxa; Cx-St = coxosternite; cx = coxite; Fe = femur; L = ledge on mentum of gnathochilarium; LL = lamella linguales; main = main branch of posterior telopod; Me = mentum; mes = mesal branch of posterior gonopod; pre = epiproct; Pre = prefemur; Po = postfemur; st = sternite; St = stipites; sub = hypoproct; t = telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022776" httpUri="https://zenodo.org/record/14022776/files/figure.png" pageId="16" pageNumber="477">Fig. 10E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B820416FFC4FFADFF622CB0D4D8FF4C" blockId="16.[151,1437,151,789]" box="[199,1408,187,213]" pageId="16" pageNumber="477">
|
||
Telson without projecting preanal scale, inconspicuous subanal scale, anal valves with weak lips (
|
||
<figureCitation id="13061893FFC4FFADFAA82CB0D42BFF4C" box="[1293,1395,187,213]" captionStart="FIGURE 10" captionStartId="15.[151,250,1765,1789]" captionTargetBox="[166,1423,184,1741]" captionTargetId="figure-15@15.[164,1423,181,1741]" captionTargetPageId="15" captionText="FIGURE 10.?Eucarliatampolo, sp. nov., holotype (FMMC 7796), drawings.A. Gnathochilarium, ventral view; B. Coxosternite of female leg 1, posterior view; C. Coxosternite of female leg 2, posterior view; D. Male midbody leg, posterior view; E. Posterior body end with telson, lateral view; F. Anterior gonopod, anterior view; G. Anterior gonopod, posterior view; H. Right posterior gonopod, tip, anterior view; I. Right posterior gonopod, posterior view. Abbreviations: av = paraprocts; Cx = coxa; Cx-St = coxosternite; cx = coxite; Fe = femur; L = ledge on mentum of gnathochilarium; LL = lamella linguales; main = main branch of posterior telopod; Me = mentum; mes = mesal branch of posterior gonopod; pre = epiproct; Pre = prefemur; Po = postfemur; st = sternite; St = stipites; sub = hypoproct; t = telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022776" httpUri="https://zenodo.org/record/14022776/files/figure.png" pageId="16" pageNumber="477">Fig. 10E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B820416FFC4FFADFF622CD4D2B1FF60" blockId="16.[151,1437,151,789]" box="[199,1001,223,249]" pageId="16" pageNumber="477">Vulvae bivalve-like, posterior valve slightly overlapping anterior valve.</paragraph>
|
||
<paragraph id="8B820416FFC4FFADFF622D08D2AEFE84" blockId="16.[151,1437,151,789]" box="[199,1014,259,285]" pageId="16" pageNumber="477">Male coxae 3–7 unmodified, but tarsi 3 - midbody legs with a tarsal pad.</paragraph>
|
||
<paragraph id="8B820416FFC4FFADFF622D2CD514FE34" blockId="16.[151,1437,151,789]" pageId="16" pageNumber="477">
|
||
Anterior gonopods sternite well-rounded, not protruding as high as coxite process (
|
||
<figureCitation id="13061893FFC4FFADFBC42D2CD59CFED8" box="[1121,1220,295,321]" captionStart="FIGURE 10" captionStartId="15.[151,250,1765,1789]" captionTargetBox="[166,1423,184,1741]" captionTargetId="figure-15@15.[164,1423,181,1741]" captionTargetPageId="15" captionText="FIGURE 10.?Eucarliatampolo, sp. nov., holotype (FMMC 7796), drawings.A. Gnathochilarium, ventral view; B. Coxosternite of female leg 1, posterior view; C. Coxosternite of female leg 2, posterior view; D. Male midbody leg, posterior view; E. Posterior body end with telson, lateral view; F. Anterior gonopod, anterior view; G. Anterior gonopod, posterior view; H. Right posterior gonopod, tip, anterior view; I. Right posterior gonopod, posterior view. Abbreviations: av = paraprocts; Cx = coxa; Cx-St = coxosternite; cx = coxite; Fe = femur; L = ledge on mentum of gnathochilarium; LL = lamella linguales; main = main branch of posterior telopod; Me = mentum; mes = mesal branch of posterior gonopod; pre = epiproct; Pre = prefemur; Po = postfemur; st = sternite; St = stipites; sub = hypoproct; t = telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022776" httpUri="https://zenodo.org/record/14022776/files/figure.png" pageId="16" pageNumber="477">Fig. 10F</figureCitation>
|
||
). Sternal apodemes (
|
||
<figureCitation id="13061893FFC4FFADFF3A2D40D05AFEFC" box="[159,258,331,357]" captionStart="FIGURE 10" captionStartId="15.[151,250,1765,1789]" captionTargetBox="[166,1423,184,1741]" captionTargetId="figure-15@15.[164,1423,181,1741]" captionTargetPageId="15" captionText="FIGURE 10.?Eucarliatampolo, sp. nov., holotype (FMMC 7796), drawings.A. Gnathochilarium, ventral view; B. Coxosternite of female leg 1, posterior view; C. Coxosternite of female leg 2, posterior view; D. Male midbody leg, posterior view; E. Posterior body end with telson, lateral view; F. Anterior gonopod, anterior view; G. Anterior gonopod, posterior view; H. Right posterior gonopod, tip, anterior view; I. Right posterior gonopod, posterior view. Abbreviations: av = paraprocts; Cx = coxa; Cx-St = coxosternite; cx = coxite; Fe = femur; L = ledge on mentum of gnathochilarium; LL = lamella linguales; main = main branch of posterior telopod; Me = mentum; mes = mesal branch of posterior gonopod; pre = epiproct; Pre = prefemur; Po = postfemur; st = sternite; St = stipites; sub = hypoproct; t = telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022776" httpUri="https://zenodo.org/record/14022776/files/figure.png" pageId="16" pageNumber="477">Fig. 10F</figureCitation>
|
||
) short. Coxite wide, projecting mesally into a wide, broadly-rounded process (
|
||
<figureCitation id="13061893FFC4FFADFB222D40D5B4FEFC" box="[1159,1260,331,357]" captionStart="FIGURE 10" captionStartId="15.[151,250,1765,1789]" captionTargetBox="[166,1423,184,1741]" captionTargetId="figure-15@15.[164,1423,181,1741]" captionTargetPageId="15" captionText="FIGURE 10.?Eucarliatampolo, sp. nov., holotype (FMMC 7796), drawings.A. Gnathochilarium, ventral view; B. Coxosternite of female leg 1, posterior view; C. Coxosternite of female leg 2, posterior view; D. Male midbody leg, posterior view; E. Posterior body end with telson, lateral view; F. Anterior gonopod, anterior view; G. Anterior gonopod, posterior view; H. Right posterior gonopod, tip, anterior view; I. Right posterior gonopod, posterior view. Abbreviations: av = paraprocts; Cx = coxa; Cx-St = coxosternite; cx = coxite; Fe = femur; L = ledge on mentum of gnathochilarium; LL = lamella linguales; main = main branch of posterior telopod; Me = mentum; mes = mesal branch of posterior gonopod; pre = epiproct; Pre = prefemur; Po = postfemur; st = sternite; St = stipites; sub = hypoproct; t = telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022776" httpUri="https://zenodo.org/record/14022776/files/figure.png" pageId="16" pageNumber="477">Fig. 10F</figureCitation>
|
||
), tip of process slightly overreaching telopodite process mesally but not apically (
|
||
<figureCitation id="13061893FFC4FFADFC272D64D2B3FE10" box="[898,1003,367,393]" captionStart="FIGURE 10" captionStartId="15.[151,250,1765,1789]" captionTargetBox="[166,1423,184,1741]" captionTargetId="figure-15@15.[164,1423,181,1741]" captionTargetPageId="15" captionText="FIGURE 10.?Eucarliatampolo, sp. nov., holotype (FMMC 7796), drawings.A. Gnathochilarium, ventral view; B. Coxosternite of female leg 1, posterior view; C. Coxosternite of female leg 2, posterior view; D. Male midbody leg, posterior view; E. Posterior body end with telson, lateral view; F. Anterior gonopod, anterior view; G. Anterior gonopod, posterior view; H. Right posterior gonopod, tip, anterior view; I. Right posterior gonopod, posterior view. Abbreviations: av = paraprocts; Cx = coxa; Cx-St = coxosternite; cx = coxite; Fe = femur; L = ledge on mentum of gnathochilarium; LL = lamella linguales; main = main branch of posterior telopod; Me = mentum; mes = mesal branch of posterior gonopod; pre = epiproct; Pre = prefemur; Po = postfemur; st = sternite; St = stipites; sub = hypoproct; t = telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022776" httpUri="https://zenodo.org/record/14022776/files/figure.png" pageId="16" pageNumber="477">Fig. 10G</figureCitation>
|
||
). Telopodite apically expanding into a wide, well-rounded process, process almost as wide as base of telopodite (
|
||
<figureCitation id="13061893FFC4FFADFC722D98D518FE34" box="[983,1088,403,429]" captionStart="FIGURE 10" captionStartId="15.[151,250,1765,1789]" captionTargetBox="[166,1423,184,1741]" captionTargetId="figure-15@15.[164,1423,181,1741]" captionTargetPageId="15" captionText="FIGURE 10.?Eucarliatampolo, sp. nov., holotype (FMMC 7796), drawings.A. Gnathochilarium, ventral view; B. Coxosternite of female leg 1, posterior view; C. Coxosternite of female leg 2, posterior view; D. Male midbody leg, posterior view; E. Posterior body end with telson, lateral view; F. Anterior gonopod, anterior view; G. Anterior gonopod, posterior view; H. Right posterior gonopod, tip, anterior view; I. Right posterior gonopod, posterior view. Abbreviations: av = paraprocts; Cx = coxa; Cx-St = coxosternite; cx = coxite; Fe = femur; L = ledge on mentum of gnathochilarium; LL = lamella linguales; main = main branch of posterior telopod; Me = mentum; mes = mesal branch of posterior gonopod; pre = epiproct; Pre = prefemur; Po = postfemur; st = sternite; St = stipites; sub = hypoproct; t = telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022776" httpUri="https://zenodo.org/record/14022776/files/figure.png" pageId="16" pageNumber="477">Fig. 10G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B820416FFC4FFADFF622DBCD23CFD54" blockId="16.[151,1437,151,789]" pageId="16" pageNumber="477">
|
||
Posterior gonopod clearly separated into coxite and telopodite, with coxite greatly extended and longer than telopodite (
|
||
<figureCitation id="13061893FFC4FFADFEBF2DD0D023FE6C" box="[282,379,475,501]" captionStart="FIGURE 10" captionStartId="15.[151,250,1765,1789]" captionTargetBox="[166,1423,184,1741]" captionTargetId="figure-15@15.[164,1423,181,1741]" captionTargetPageId="15" captionText="FIGURE 10.?Eucarliatampolo, sp. nov., holotype (FMMC 7796), drawings.A. Gnathochilarium, ventral view; B. Coxosternite of female leg 1, posterior view; C. Coxosternite of female leg 2, posterior view; D. Male midbody leg, posterior view; E. Posterior body end with telson, lateral view; F. Anterior gonopod, anterior view; G. Anterior gonopod, posterior view; H. Right posterior gonopod, tip, anterior view; I. Right posterior gonopod, posterior view. Abbreviations: av = paraprocts; Cx = coxa; Cx-St = coxosternite; cx = coxite; Fe = femur; L = ledge on mentum of gnathochilarium; LL = lamella linguales; main = main branch of posterior telopod; Me = mentum; mes = mesal branch of posterior gonopod; pre = epiproct; Pre = prefemur; Po = postfemur; st = sternite; St = stipites; sub = hypoproct; t = telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022776" httpUri="https://zenodo.org/record/14022776/files/figure.png" pageId="16" pageNumber="477">Fig. 10I</figureCitation>
|
||
). Sternite small but sclerotized, triangular (
|
||
<figureCitation id="13061893FFC4FFADFCD32DD0D28EFE6C" box="[886,982,475,501]" captionStart="FIGURE 10" captionStartId="15.[151,250,1765,1789]" captionTargetBox="[166,1423,184,1741]" captionTargetId="figure-15@15.[164,1423,181,1741]" captionTargetPageId="15" captionText="FIGURE 10.?Eucarliatampolo, sp. nov., holotype (FMMC 7796), drawings.A. Gnathochilarium, ventral view; B. Coxosternite of female leg 1, posterior view; C. Coxosternite of female leg 2, posterior view; D. Male midbody leg, posterior view; E. Posterior body end with telson, lateral view; F. Anterior gonopod, anterior view; G. Anterior gonopod, posterior view; H. Right posterior gonopod, tip, anterior view; I. Right posterior gonopod, posterior view. Abbreviations: av = paraprocts; Cx = coxa; Cx-St = coxosternite; cx = coxite; Fe = femur; L = ledge on mentum of gnathochilarium; LL = lamella linguales; main = main branch of posterior telopod; Me = mentum; mes = mesal branch of posterior gonopod; pre = epiproct; Pre = prefemur; Po = postfemur; st = sternite; St = stipites; sub = hypoproct; t = telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022776" httpUri="https://zenodo.org/record/14022776/files/figure.png" pageId="16" pageNumber="477">Fig. 10I</figureCitation>
|
||
). Coxite mesally with a single groove, elongated into a long stem (
|
||
<figureCitation id="13061893FFC4FFADFE772DF4D368FD80" box="[466,560,511,537]" captionStart="FIGURE 10" captionStartId="15.[151,250,1765,1789]" captionTargetBox="[166,1423,184,1741]" captionTargetId="figure-15@15.[164,1423,181,1741]" captionTargetPageId="15" captionText="FIGURE 10.?Eucarliatampolo, sp. nov., holotype (FMMC 7796), drawings.A. Gnathochilarium, ventral view; B. Coxosternite of female leg 1, posterior view; C. Coxosternite of female leg 2, posterior view; D. Male midbody leg, posterior view; E. Posterior body end with telson, lateral view; F. Anterior gonopod, anterior view; G. Anterior gonopod, posterior view; H. Right posterior gonopod, tip, anterior view; I. Right posterior gonopod, posterior view. Abbreviations: av = paraprocts; Cx = coxa; Cx-St = coxosternite; cx = coxite; Fe = femur; L = ledge on mentum of gnathochilarium; LL = lamella linguales; main = main branch of posterior telopod; Me = mentum; mes = mesal branch of posterior gonopod; pre = epiproct; Pre = prefemur; Po = postfemur; st = sternite; St = stipites; sub = hypoproct; t = telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022776" httpUri="https://zenodo.org/record/14022776/files/figure.png" pageId="16" pageNumber="477">Fig. 10I</figureCitation>
|
||
). Telopodite long, but shorter than elongated coxite (
|
||
<figureCitation id="13061893FFC4FFADFB2D2DF4D5BDFD80" box="[1160,1253,511,537]" captionStart="FIGURE 10" captionStartId="15.[151,250,1765,1789]" captionTargetBox="[166,1423,184,1741]" captionTargetId="figure-15@15.[164,1423,181,1741]" captionTargetPageId="15" captionText="FIGURE 10.?Eucarliatampolo, sp. nov., holotype (FMMC 7796), drawings.A. Gnathochilarium, ventral view; B. Coxosternite of female leg 1, posterior view; C. Coxosternite of female leg 2, posterior view; D. Male midbody leg, posterior view; E. Posterior body end with telson, lateral view; F. Anterior gonopod, anterior view; G. Anterior gonopod, posterior view; H. Right posterior gonopod, tip, anterior view; I. Right posterior gonopod, posterior view. Abbreviations: av = paraprocts; Cx = coxa; Cx-St = coxosternite; cx = coxite; Fe = femur; L = ledge on mentum of gnathochilarium; LL = lamella linguales; main = main branch of posterior telopod; Me = mentum; mes = mesal branch of posterior gonopod; pre = epiproct; Pre = prefemur; Po = postfemur; st = sternite; St = stipites; sub = hypoproct; t = telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022776" httpUri="https://zenodo.org/record/14022776/files/figure.png" pageId="16" pageNumber="477">Fig. 10I</figureCitation>
|
||
), basally deeply divided into two branches, main branch and mesal branch, both curved mesally (
|
||
<figureCitation id="13061893FFC4FFADFB9E2E28D5C2FDA4" box="[1083,1178,547,573]" captionStart="FIGURE 10" captionStartId="15.[151,250,1765,1789]" captionTargetBox="[166,1423,184,1741]" captionTargetId="figure-15@15.[164,1423,181,1741]" captionTargetPageId="15" captionText="FIGURE 10.?Eucarliatampolo, sp. nov., holotype (FMMC 7796), drawings.A. Gnathochilarium, ventral view; B. Coxosternite of female leg 1, posterior view; C. Coxosternite of female leg 2, posterior view; D. Male midbody leg, posterior view; E. Posterior body end with telson, lateral view; F. Anterior gonopod, anterior view; G. Anterior gonopod, posterior view; H. Right posterior gonopod, tip, anterior view; I. Right posterior gonopod, posterior view. Abbreviations: av = paraprocts; Cx = coxa; Cx-St = coxosternite; cx = coxite; Fe = femur; L = ledge on mentum of gnathochilarium; LL = lamella linguales; main = main branch of posterior telopod; Me = mentum; mes = mesal branch of posterior gonopod; pre = epiproct; Pre = prefemur; Po = postfemur; st = sternite; St = stipites; sub = hypoproct; t = telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022776" httpUri="https://zenodo.org/record/14022776/files/figure.png" pageId="16" pageNumber="477">Fig. 10I</figureCitation>
|
||
). Main branch slender on whole length, apical part slightly slenderer than basal 3/4, with well-rounded tip weakly curved towards mesal branch, sclerotized. Mesal branch with basal half sclerotized, apical half twice as wide as basal half and membranous (
|
||
<figureCitation id="13061893FFC4FFADFF3A2E84D05FFD30" box="[159,263,655,681]" captionStart="FIGURE 10" captionStartId="15.[151,250,1765,1789]" captionTargetBox="[166,1423,184,1741]" captionTargetId="figure-15@15.[164,1423,181,1741]" captionTargetPageId="15" captionText="FIGURE 10.?Eucarliatampolo, sp. nov., holotype (FMMC 7796), drawings.A. Gnathochilarium, ventral view; B. Coxosternite of female leg 1, posterior view; C. Coxosternite of female leg 2, posterior view; D. Male midbody leg, posterior view; E. Posterior body end with telson, lateral view; F. Anterior gonopod, anterior view; G. Anterior gonopod, posterior view; H. Right posterior gonopod, tip, anterior view; I. Right posterior gonopod, posterior view. Abbreviations: av = paraprocts; Cx = coxa; Cx-St = coxosternite; cx = coxite; Fe = femur; L = ledge on mentum of gnathochilarium; LL = lamella linguales; main = main branch of posterior telopod; Me = mentum; mes = mesal branch of posterior gonopod; pre = epiproct; Pre = prefemur; Po = postfemur; st = sternite; St = stipites; sub = hypoproct; t = telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022776" httpUri="https://zenodo.org/record/14022776/files/figure.png" pageId="16" pageNumber="477">Fig. 10H</figureCitation>
|
||
). Tip of mesal branch with two openings of the sperm canal (
|
||
<figureCitation id="13061893FFC4FFADFC0E2E84D55EFD30" box="[939,1030,655,681]" captionStart="FIGURE 10" captionStartId="15.[151,250,1765,1789]" captionTargetBox="[166,1423,184,1741]" captionTargetId="figure-15@15.[164,1423,181,1741]" captionTargetPageId="15" captionText="FIGURE 10.?Eucarliatampolo, sp. nov., holotype (FMMC 7796), drawings.A. Gnathochilarium, ventral view; B. Coxosternite of female leg 1, posterior view; C. Coxosternite of female leg 2, posterior view; D. Male midbody leg, posterior view; E. Posterior body end with telson, lateral view; F. Anterior gonopod, anterior view; G. Anterior gonopod, posterior view; H. Right posterior gonopod, tip, anterior view; I. Right posterior gonopod, posterior view. Abbreviations: av = paraprocts; Cx = coxa; Cx-St = coxosternite; cx = coxite; Fe = femur; L = ledge on mentum of gnathochilarium; LL = lamella linguales; main = main branch of posterior telopod; Me = mentum; mes = mesal branch of posterior gonopod; pre = epiproct; Pre = prefemur; Po = postfemur; st = sternite; St = stipites; sub = hypoproct; t = telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022776" httpUri="https://zenodo.org/record/14022776/files/figure.png" pageId="16" pageNumber="477">Fig. 10I</figureCitation>
|
||
), the upper canal strongly and freely projecting mesally from membranous part of branch (
|
||
<figureCitation id="13061893FFC4FFADFD4A2EB8D200FD54" box="[751,856,691,717]" captionStart="FIGURE 10" captionStartId="15.[151,250,1765,1789]" captionTargetBox="[166,1423,184,1741]" captionTargetId="figure-15@15.[164,1423,181,1741]" captionTargetPageId="15" captionText="FIGURE 10.?Eucarliatampolo, sp. nov., holotype (FMMC 7796), drawings.A. Gnathochilarium, ventral view; B. Coxosternite of female leg 1, posterior view; C. Coxosternite of female leg 2, posterior view; D. Male midbody leg, posterior view; E. Posterior body end with telson, lateral view; F. Anterior gonopod, anterior view; G. Anterior gonopod, posterior view; H. Right posterior gonopod, tip, anterior view; I. Right posterior gonopod, posterior view. Abbreviations: av = paraprocts; Cx = coxa; Cx-St = coxosternite; cx = coxite; Fe = femur; L = ledge on mentum of gnathochilarium; LL = lamella linguales; main = main branch of posterior telopod; Me = mentum; mes = mesal branch of posterior gonopod; pre = epiproct; Pre = prefemur; Po = postfemur; st = sternite; St = stipites; sub = hypoproct; t = telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022776" httpUri="https://zenodo.org/record/14022776/files/figure.png" pageId="16" pageNumber="477">Fig. 10H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C327579DFFC4FFADFF622EDCD595FC8C" pageId="16" pageNumber="477" type="discussion">
|
||
<paragraph id="8B820416FFC4FFADFF622EDCD595FC8C" blockId="16.[151,1437,151,789]" pageId="16" pageNumber="477">
|
||
<emphasis id="B949D804FFC4FFADFF622EDCD06CFD68" bold="true" box="[199,308,727,753]" pageId="16" pageNumber="477">Remarks</emphasis>
|
||
. The littoral forest of Tampolo is currently also the only known locality of the critically endangered Tampolo giant pill-millipede
|
||
<taxonomicName id="4C3D7F95FFC4FFADFE452EF0D2D7FC8C" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[480,911,763,789]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="16" pageNumber="477" phylum="Arthropoda" rank="species" species="tampolo">
|
||
<emphasis id="B949D804FFC4FFADFE452EF0D3BDFC8C" box="[480,741,763,789]" italics="true" pageId="16" pageNumber="477">Zoosphaerium tampolo</emphasis>
|
||
Wesener, 2009
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFAC79E7FFC4FFADFC3B2EF0D598FC8C" author="Rudolf, E. & Wesener, T." box="[926,1216,763,789]" pageId="16" pageNumber="477" refId="ref14546" refString="Rudolf, E. & Wesener, T. (2017 c) Zoosphaerium tampolo. The IUCN Red List of Threatened Species, 2017, e. T 65526057 A 65528020. https: // doi. org / 10.2305 / IUCN. UK. 2017 - 1. RLTS. T 65526057 A 65528020. en" type="journal volume" year="2017">Rudolf & Wesener 2017c</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |