treatments-xml/data/03/94/B5/0394B500FFD1FFBBFF3229C1D04AF86F.xml

277 lines
44 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="2EBFA3D11B133E1EA090F71080F5A4EA" ID-DOI="10.11646/zootaxa.5529.3.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="14022748" ID-ZooBank="716EDF67-C933-484D-911B-4585B11187A6" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1730453336952" checkinUser="plazi" docAuthor="Wesener, Thomas" docDate="2024" docId="0394B500FFD1FFBBFF3229C1D04AF86F" docLanguage="en" docName="zootaxa.5529.3.3.pdf" docOrigin="Zootaxa 5529 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5529.3.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Aphistogoniulus amberivery Wesener, 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="467" masterDocId="FFADCD78FFD4FFBDFFA52C0BD158FF99" masterDocTitle="Five new species and numerous locality records of Spirobolida millipedes from Madagascar (Diplopoda, Spirobolida, Pachybolidae)" masterLastPageNumber="486" masterPageNumber="461" pageNumber="466" updateTime="1730454045455" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="BF7EE29630F5D739CA4EA5F1EAB7B619" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="CAA40E50530C08F10B9BE3FF194A06ED">
<mods:title id="6E0B8FAB8241F179E408C88EA6F3329B">Five new species and numerous locality records of Spirobolida millipedes from Madagascar (Diplopoda, Spirobolida, Pachybolidae)</mods:title>
</mods:titleInfo>
<mods:name id="69858DDCDA4A52FD09ED4CB794BE1CAD" type="personal">
<mods:role id="919D96A310E8A195493A0D74DD25A31A">
<mods:roleTerm id="D78666F00E420BF627B986524ADB1A4B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="ED876AB67FC24E3840A70171F0C8C56B">Wesener, Thomas</mods:namePart>
</mods:name>
<mods:typeOfResource id="6E6B18AB5811F649DCCEABA6855099C5">text</mods:typeOfResource>
<mods:relatedItem id="F5C264F3280061D0078A75E43086001D" type="host">
<mods:titleInfo id="C5F524001D7F25A24468AF4B11E7F330">
<mods:title id="70855D203EFEBF1214408D8DDF9DCED7">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="73EF3A852ACC67B1ABF82B7686985F79">
<mods:date id="EA0E7ABE3B54A44B9A63DFC69FD345BB">2024</mods:date>
<mods:detail id="30F705EC7251B03D9DD630C5E4A78EAD" type="pubDate">
<mods:number id="EFA38F32F64B366A9566131A890474D1">2024-10-29</mods:number>
</mods:detail>
<mods:detail id="F6F2F91C3F03E16E4CF894E987DE9AC4" type="volume">
<mods:number id="46C28A0E469E2CD3DE93903F7B42B46D">5529</mods:number>
</mods:detail>
<mods:detail id="EE3CDBED36B48966868C82378ED07FE8" type="issue">
<mods:number id="9588D887118F56F6138E2E7E788923A2">3</mods:number>
</mods:detail>
<mods:extent id="3E85028DC93E6E168E904915293D834E" unit="page">
<mods:start id="3149C1030F55F4AB523EFA711305E1B6">461</mods:start>
<mods:end id="18696186E97100608C5CB45979833186">486</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="08650BD4BFD91A4E0F3578303CF0CB97">
<mods:url id="8CB5EF662CDE0F6F6C1C617EF620DD58">http://dx.doi.org/10.11646/zootaxa.5529.3.3</mods:url>
</mods:location>
<mods:classification id="DC438A144CC999A38B8501DF0701C65A">journal article</mods:classification>
<mods:identifier id="0663050DAF7655665A48DDC5784FC821" type="DOI">10.11646/zootaxa.5529.3.3</mods:identifier>
<mods:identifier id="40894BA355FD287EAF0D239A1A6FDFB7" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="7C4EADA357F6B63F3E01893A64B65CB2" type="Zenodo-Dep">14022748</mods:identifier>
<mods:identifier id="148B954FE537BE294C3615D1FB8BCD90" type="ZooBank">716EDF67-C933-484D-911B-4585B11187A6</mods:identifier>
</mods:mods>
<treatment id="0394B500FFD1FFBBFF3229C1D04AF86F" LSID="urn:lsid:plazi:treatment:0394B500FFD1FFBBFF3229C1D04AF86F" httpUri="http://treatment.plazi.org/id/0394B500FFD1FFBBFF3229C1D04AF86F" lastPageId="6" lastPageNumber="467" pageId="5" pageNumber="466">
<subSubSection id="C327579DFFD1FFB8FF3229C1D30AFA7C" box="[151,594,1482,1509]" pageId="5" pageNumber="466" type="nomenclature">
<paragraph id="8B820416FFD1FFB8FF3229C1D30AFA7C" blockId="5.[151,594,1482,1545]" box="[151,594,1482,1509]" pageId="5" pageNumber="466">
<heading id="D0CAB37AFFD1FFB8FF3229C1D30AFA7C" bold="true" box="[151,594,1482,1509]" fontSize="11" level="1" pageId="5" pageNumber="466" reason="1">
<emphasis id="B949D804FFD1FFB8FF3229C1D30AFA7C" bold="true" box="[151,594,1482,1509]" pageId="5" pageNumber="466">
<taxonomicName id="4C3D7F95FFD1FFB8FF3229C1D0B4FA7C" authorityName="Wesener" authorityYear="2024" box="[151,492,1482,1509]" class="Diplopoda" family="Pachybolidae" genus="Aphistogoniulus" kingdom="Animalia" order="Spirobolida" pageId="5" pageNumber="466" phylum="Arthropoda" rank="species" species="amberivery" status="sp. nov.">
<emphasis id="B949D804FFD1FFB8FF3229C1D0B4FA7C" bold="true" box="[151,492,1482,1509]" italics="true" pageId="5" pageNumber="466">Aphistogoniulus amberivery</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A657FFFD1FFB8FE5129C0D30AFA7C" box="[500,594,1483,1509]" pageId="5" pageNumber="466" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C327579DFFD1FFB8FF3229E4D057F990" box="[151,271,1519,1545]" pageId="5" pageNumber="466" type="description">
<paragraph id="8B820416FFD1FFB8FF3229E4D057F990" blockId="5.[151,594,1482,1545]" box="[151,271,1519,1545]" pageId="5" pageNumber="466">
<figureCitation id="13061893FFD1FFB8FF3229E4D057F990" box="[151,271,1519,1545]" captionStart="FIGURE 2" captionStartId="6.[151,250,1012,1036]" captionTargetBox="[416,1172,237,989]" captionTargetId="figure-56@6.[416,1172,237,989]" captionTargetPageId="6" captionText="FIGURE 2. Aphistogoniulus amberivery, sp. nov., holotype (FMNH-INS 3196506). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 17, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Anterior gonopod, anterior view; E. Anterior gonopod, posterior view; F. Right posterior gonopod, anterior view; G. Right posterior gonopod, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; mb = main branch of posterior gonopod; Md = basal joints of mandible; oz = ozopore; Pre = epiproct; rp = retrorse process; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022754" httpUri="https://zenodo.org/record/14022754/files/figure.png" pageId="5" pageNumber="466">Fig. 2AG</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C327579DFFD1FFB8FF322A3DD0B7F924" pageId="5" pageNumber="466" type="materials_examined">
<paragraph id="8B820416FFD1FFB8FF322A3DD0F8F900" blockId="5.[151,1437,1590,2013]" pageId="5" pageNumber="466">
<emphasis id="B949D804FFD1FFB8FF322A3DD025F9C9" bold="true" box="[151,381,1590,1616]" pageId="5" pageNumber="466">Material examined</emphasis>
.
<materialsCitation id="3B550E4BFFD1FFB8FE2B2A3CD0C4F900" collectingDate="2013-10" collectionCode="FMNH-INS" collectorName="S. M. Goodman &amp; Toky Randriamora" county="Sofia" elevation="1570" latitude="-14.346472" location="Amberivery Forest" longLatPrecision="1" longitude="48.579803" municipality="Bemanevika" pageId="5" pageNumber="466" specimenCode="FMNH-INS 3196506" specimenCount="1" stateProvince="Mahajanga" typeStatus="holotype">
1 M
<typeStatus id="5486BAB4FFD1FFB8FE6C2A3CD376F9C8" box="[457,558,1591,1617]" pageId="5" pageNumber="466" type="holotype">holotype</typeStatus>
,
<emphasis id="B949D804FFD1FFB8FD982A3DD26FF9C9" bold="true" box="[573,823,1590,1616]" pageId="5" pageNumber="466">
<specimenCode id="DB9BAC6DFFD1FFB8FD982A3DD26FF9C9" box="[573,823,1590,1616]" collectionCode="FMNH-INS" pageId="5" pageNumber="466">FMNH-INS 3196506</specimenCode>
</emphasis>
,
<collectingRegion id="49F9CAF4FFD1FFB8FCED2A3CD29FF9C8" box="[840,967,1591,1617]" country="Madagascar" name="Mahajanga" pageId="5" pageNumber="466">Mahajanga</collectingRegion>
,
<collectingCounty id="62E37C9AFFD1FFB8FC732A3CD54CF9C8" box="[982,1044,1591,1617]" pageId="5" pageNumber="466">Sofia</collectingCounty>
,
<collectingMunicipality id="6BE69E6CFFD1FFB8FB872A3CD5E9F9C8" box="[1058,1201,1591,1617]" pageId="5" pageNumber="466">Bemanevika</collectingMunicipality>
,
<location id="8EE252CDFFD1FFB8FB1B2A3CD4C0F9C8" LSID="urn:lsid:plazi:treatment:0394B500FFD1FFBBFF3229C1D04AF86F:8EE252CDFFD1FFB8FB1B2A3CD4C0F9C8" box="[1214,1432,1591,1617]" county="Sofia" latitude="-14.346472" longLatPrecision="1" longitude="48.579803" municipality="Bemanevika" name="Amberivery Forest" pageId="5" pageNumber="466" stateProvince="Mahajanga">Amberivery Forest</location>
,
<quantity id="4CC5A9F3FFD1FFB8FF322A51D1A8F9EC" box="[151,240,1626,1653]" metricMagnitude="3" metricUnit="m" metricValue="1.5699999999999998" pageId="5" pageNumber="466" unit="m" value="1570.0">
<elevation id="0010E325FFD1FFB8FF322A51D1A8F9EC" box="[151,240,1626,1653]" metricMagnitude="3" metricUnit="m" metricValue="1.5699999999999998" pageId="5" pageNumber="466" unit="m" value="1570.0">1570 m</elevation>
</quantity>
, slightly disturbed montane forest,
<geoCoordinate id="EE0962D1FFD1FFB8FD222A50D275F9EC" box="[647,813,1627,1653]" degrees="14" direction="south" minutes="20" orientation="latitude" pageId="5" pageNumber="466" precision="1" seconds="47.3" value="-14.346472">14°2047.3 S</geoCoordinate>
,
<geoCoordinate id="EE0962D1FFD1FFB8FC992A50D2BCF9EC" box="[828,996,1627,1653]" degrees="48" direction="east" minutes="34" orientation="longitude" pageId="5" pageNumber="466" precision="1" seconds="47.3" value="48.579803">48°3447.3 E</geoCoordinate>
, coll.
<collectingMethod id="527C7C01FFD1FFB8FB882A50D526F9EC" box="[1069,1150,1627,1653]" pageId="5" pageNumber="466">pitfalls</collectingMethod>
,
<date id="FF8322D6FFD1FFB8FB2E2A50D5B8F9EC" box="[1163,1248,1627,1653]" pageId="5" pageNumber="466" value="2013-10">
<collectingDate id="EFC7DB3EFFD1FFB8FB2E2A50D5B8F9EC" box="[1163,1248,1627,1653]" pageId="5" pageNumber="466" value="2013-10">X.2013</collectingDate>
</date>
,
<collectorName id="26C861C0FFD1FFB8FB4B2A50D4C4F9EC" box="[1262,1436,1627,1653]" pageId="5" pageNumber="466">S.M. Goodman</collectorName>
and
<collectorName id="26C861C0FFD1FFB8FF632A74D0C4F900" box="[198,412,1663,1689]" pageId="5" pageNumber="466">Toky Randriamora</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph id="8B820416FFD1FFB8FF622AA9D0B7F924" blockId="5.[151,1437,1590,2013]" box="[199,495,1698,1725]" pageId="5" pageNumber="466">
<emphasis id="B949D804FFD1FFB8FF622AA9D031F924" bold="true" box="[199,361,1698,1725]" pageId="5" pageNumber="466">DNA barcode</emphasis>
: PQ307145
</paragraph>
</subSubSection>
<subSubSection id="C327579DFFD1FFB8FF622ACDD43EF978" box="[199,1382,1734,1761]" pageId="5" pageNumber="466" type="etymology">
<paragraph id="8B820416FFD1FFB8FF622ACDD43EF978" blockId="5.[151,1437,1590,2013]" box="[199,1382,1734,1761]" pageId="5" pageNumber="466">
<emphasis id="B949D804FFD1FFB8FF622ACDD0FCF979" bold="true" box="[199,420,1734,1760]" pageId="5" pageNumber="466">Derivatio nominis.</emphasis>
amberivery, noun in apposition, after the
<typeStatus id="5486BAB4FFD1FFB8FCDC2ACCD2F1F978" box="[889,937,1735,1761]" pageId="5" pageNumber="466">type</typeStatus>
and only known locality of the species.
</paragraph>
</subSubSection>
<subSubSection id="C327579DFFD1FFBBFF622AE1D45BFF4C" lastPageId="6" lastPageNumber="467" pageId="5" pageNumber="466" type="diagnosis">
<paragraph id="8B820416FFD1FFBBFF622AE1D45BFF4C" blockId="5.[151,1437,1590,2013]" lastBlockId="6.[151,1437,151,213]" lastPageId="6" lastPageNumber="467" pageId="5" pageNumber="466">
<emphasis id="B949D804FFD1FFB8FF622AE1D01AF89D" bold="true" box="[199,322,1770,1796]" pageId="5" pageNumber="466">Diagnosis.</emphasis>
Characteristic colour with laterally entirely black body rings, which are above the ozopores dorsally bright red with a wide black posterior margin. Collum black, anterior and posterior margin with a red stripe. Head, antenna and legs black. The posterior gonopod with the shorter basal branch and the main branch with the numerous membranous fringes resembles only those of
<taxonomicName id="4C3D7F95FFD1FFB8FD352B5CD3BCF8E8" baseAuthorityName="de Saussure &amp; Zehntner" baseAuthorityYear="1897" box="[656,740,1879,1905]" class="Diplopoda" family="Pachybolidae" genus="Aphistogoniulus" kingdom="Animalia" order="Spirobolida" pageId="5" pageNumber="466" phylum="Arthropoda" rank="species" species="hova">
<emphasis id="B949D804FFD1FFB8FD352B5CD3BCF8E8" box="[656,740,1879,1905]" italics="true" pageId="5" pageNumber="466">A. hova</emphasis>
</taxonomicName>
(deSaussure &amp; Zehntner, 1897).
<taxonomicName id="4C3D7F95FFD1FFB8FBF12B5CD5ABF8E8" authorityName="Wesener" authorityYear="2024" box="[1108,1267,1879,1905]" class="Diplopoda" family="Pachybolidae" genus="Aphistogoniulus" kingdom="Animalia" order="Spirobolida" pageId="5" pageNumber="466" phylum="Arthropoda" rank="species" species="amberivery">
<emphasis id="B949D804FFD1FFB8FBF12B5CD5ABF8E8" box="[1108,1267,1879,1905]" italics="true" pageId="5" pageNumber="466">A. amberivery</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C3D7F95FFD1FFB8FA212B5CD195F80C" baseAuthorityName="de Saussure &amp; Zehntner" baseAuthorityYear="1897" class="Diplopoda" family="Pachybolidae" genus="Aphistogoniulus" kingdom="Animalia" order="Spirobolida" pageId="5" pageNumber="466" phylum="Arthropoda" rank="species" species="hova">
<emphasis id="B949D804FFD1FFB8FA212B5CD195F80C" italics="true" pageId="5" pageNumber="466">A. hova</emphasis>
</taxonomicName>
in fewer body rings (51 versus
<quantity id="4CC5A9F3FFD1FFB8FD962B71D3CEF80C" box="[563,662,1914,1941]" metricMagnitude="0" metricUnit="m" metricValue="1.4097" metricValueMax="1.4478" metricValueMin="1.3716" pageId="5" pageNumber="466" unit="in" value="55.5" valueMax="57.0" valueMin="54.0">5457 in</quantity>
<taxonomicName id="4C3D7F95FFD1FFB8FD382B70D3AAF80C" baseAuthorityName="de Saussure &amp; Zehntner" baseAuthorityYear="1897" box="[669,754,1915,1941]" class="Diplopoda" family="Pachybolidae" genus="Aphistogoniulus" kingdom="Animalia" order="Spirobolida" pageId="5" pageNumber="466" phylum="Arthropoda" rank="species" species="hova">
<emphasis id="B949D804FFD1FFB8FD382B70D3AAF80C" box="[669,754,1915,1941]" italics="true" pageId="5" pageNumber="466">A. hova</emphasis>
</taxonomicName>
), as well as the anterior gonopod, where the retrorse process is located apically and not medially. Compared to the
<typeStatus id="5486BAB4FFD1FFB8FD532B94D205F820" box="[758,861,1951,1977]" pageId="5" pageNumber="466" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="4C3D7F95FFD1FFB8FC272B94D280F820" baseAuthorityName="de Saussure &amp; Zehntner" baseAuthorityYear="1897" box="[898,984,1951,1977]" class="Diplopoda" family="Pachybolidae" genus="Aphistogoniulus" kingdom="Animalia" order="Spirobolida" pageId="5" pageNumber="466" phylum="Arthropoda" rank="species" species="hova">
<emphasis id="B949D804FFD1FFB8FC272B94D280F820" box="[898,984,1951,1977]" italics="true" pageId="5" pageNumber="466">A. hova</emphasis>
</taxonomicName>
, the sternite of the anterior gonopods of
<taxonomicName id="4C3D7F95FFD1FFB8FF322BC8D060F844" authorityName="Wesener" authorityYear="2024" box="[151,312,1987,2013]" class="Diplopoda" family="Pachybolidae" genus="Aphistogoniulus" kingdom="Animalia" order="Spirobolida" pageId="5" pageNumber="466" phylum="Arthropoda" rank="species" species="amberivery" status="sp. nov.">
<emphasis id="B949D804FFD1FFB8FF322BC8D060F844" box="[151,312,1987,2013]" italics="true" pageId="5" pageNumber="466">A. amberivery</emphasis>
</taxonomicName>
<emphasis id="B949D804FFD1FFB8FE9A2BC8D0C2F844" bold="true" box="[319,410,1987,2013]" pageId="5" pageNumber="466">
<taxonomicNameLabel id="A27A657FFFD1FFB8FE9A2BC8D0C2F844" box="[319,410,1987,2013]" pageId="5" pageNumber="466" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is single-tipped (double-tipped in
<taxonomicName id="4C3D7F95FFD1FFB8FC862BC8D221F844" baseAuthorityName="de Saussure &amp; Zehntner" baseAuthorityYear="1897" box="[803,889,1987,2013]" class="Diplopoda" family="Pachybolidae" genus="Aphistogoniulus" kingdom="Animalia" order="Spirobolida" pageId="5" pageNumber="466" phylum="Arthropoda" rank="species" species="hova">
<emphasis id="B949D804FFD1FFB8FC862BC8D221F844" box="[803,889,1987,2013]" italics="true" pageId="5" pageNumber="466">A. hova</emphasis>
</taxonomicName>
), and the retrorse process is strongly protruding above the lateral margin of the telopodite (only very slightly protruding in
<taxonomicName id="4C3D7F95FFD2FFBBFC7A2C9CD56CFF28" baseAuthorityName="de Saussure &amp; Zehntner" baseAuthorityYear="1897" box="[991,1076,151,177]" class="Diplopoda" family="Pachybolidae" genus="Aphistogoniulus" kingdom="Animalia" order="Spirobolida" pageId="6" pageNumber="467" phylum="Arthropoda" rank="species" species="hova">
<emphasis id="B949D804FFD2FFBBFC7A2C9CD56CFF28" box="[991,1076,151,177]" italics="true" pageId="6" pageNumber="467">A. hova</emphasis>
</taxonomicName>
).
<taxonomicName id="4C3D7F95FFD2FFBBFBE92C9CD5B4FF28" authorityName="Wesener" authorityYear="2024" box="[1100,1260,151,177]" class="Diplopoda" family="Pachybolidae" genus="Aphistogoniulus" kingdom="Animalia" order="Spirobolida" pageId="6" pageNumber="467" phylum="Arthropoda" rank="species" species="amberivery" status="sp. nov.">
<emphasis id="B949D804FFD2FFBBFBE92C9CD5B4FF28" box="[1100,1260,151,177]" italics="true" pageId="6" pageNumber="467">A. amberivery</emphasis>
</taxonomicName>
<emphasis id="B949D804FFD2FFBBFB562C9CD414FF28" bold="true" box="[1267,1356,151,177]" pageId="6" pageNumber="467">
<taxonomicNameLabel id="A27A657FFFD2FFBBFB562C9CD414FF28" box="[1267,1356,151,177]" pageId="6" pageNumber="467" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from all sampled 6 populations of
<taxonomicName id="4C3D7F95FFD2FFBBFDBC2CB0D336FF4C" baseAuthorityName="de Saussure &amp; Zehntner" baseAuthorityYear="1897" box="[537,622,187,213]" class="Diplopoda" family="Pachybolidae" genus="Aphistogoniulus" kingdom="Animalia" order="Spirobolida" pageId="6" pageNumber="467" phylum="Arthropoda" rank="species" species="hova">
<emphasis id="B949D804FFD2FFBBFDBC2CB0D336FF4C" box="[537,622,187,213]" italics="true" pageId="6" pageNumber="467">A. hova</emphasis>
</taxonomicName>
in a p-distance of the COI barcoding fragment of 1112%.
</paragraph>
</subSubSection>
<caption id="DF42549EFFD2FFBBFF322FFFD438FB7D" ID-DOI="http://doi.org/10.5281/zenodo.14022754" ID-Zenodo-Dep="14022754" httpUri="https://zenodo.org/record/14022754/files/figure.png" pageId="6" pageNumber="467" startId="6.[151,250,1012,1036]" targetBox="[416,1172,237,989]" targetPageId="6" targetType="figure">
<paragraph id="8B820416FFD2FFBBFF322FFFD438FB7D" blockId="6.[151,1437,1012,1253]" pageId="6" pageNumber="467">
<emphasis id="B949D804FFD2FFBBFF322FFFD04BFB95" bold="true" box="[151,275,1012,1036]" pageId="6" pageNumber="467">FIGURE 2.</emphasis>
<taxonomicName id="4C3D7F95FFD2FFBBFEBC2FFFD36FFB95" authorityName="Wesener" authorityYear="2024" box="[281,567,1012,1036]" class="Diplopoda" family="Pachybolidae" genus="Aphistogoniulus" kingdom="Animalia" order="Spirobolida" pageId="6" pageNumber="467" phylum="Arthropoda" rank="species" species="amberivery" status="sp. nov.">
<emphasis id="B949D804FFD2FFBBFEBC2FFFD36FFB95" box="[281,567,1012,1036]" italics="true" pageId="6" pageNumber="467">Aphistogoniulus amberivery</emphasis>
</taxonomicName>
,
<emphasis id="B949D804FFD2FFBBFDE12FFED3CDFB94" bold="true" box="[580,661,1013,1037]" pageId="6" pageNumber="467">
<taxonomicNameLabel id="A27A657FFFD2FFBBFDE12FFED3CDFB94" box="[580,661,1013,1037]" pageId="6" pageNumber="467" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype (
<emphasis id="B949D804FFD2FFBBFCAC2FFFD2B1FB95" bold="true" box="[777,1001,1012,1036]" pageId="6" pageNumber="467">FMNH-INS 3196506</emphasis>
).
<emphasis id="B949D804FFD2FFBBFC582FFFD557FB95" bold="true" box="[1021,1039,1012,1036]" pageId="6" pageNumber="467">A</emphasis>
. Multi-layer photograph, male anterior body, lateral view;
<emphasis id="B949D804FFD2FFBBFEC72812D02BFBA8" bold="true" box="[354,371,1049,1073]" pageId="6" pageNumber="467">B</emphasis>
. Multi-layer photograph, male leg pairs 17, ventral view;
<emphasis id="B949D804FFD2FFBBFC432813D2A0FBA9" bold="true" box="[998,1016,1048,1072]" pageId="6" pageNumber="467">C</emphasis>
. Multi-layer photograph, male posterior body end with telson;
<emphasis id="B949D804FFD2FFBBFEDA2836D0C9FBCC" bold="true" box="[383,401,1085,1109]" pageId="6" pageNumber="467">D</emphasis>
. Anterior gonopod, anterior view;
<emphasis id="B949D804FFD2FFBBFD5C2836D252FBCC" bold="true" box="[761,778,1085,1109]" pageId="6" pageNumber="467">E</emphasis>
. Anterior gonopod, posterior view;
<emphasis id="B949D804FFD2FFBBFBD82836D5D4FBCC" bold="true" box="[1149,1164,1085,1109]" pageId="6" pageNumber="467">F</emphasis>
. Right posterior gonopod, anterior view;
<emphasis id="B949D804FFD2FFBBFE8E286BD067FBE1" bold="true" box="[299,319,1120,1144]" pageId="6" pageNumber="467">G</emphasis>
. Right posterior gonopod, posterior view.
<emphasis id="B949D804FFD2FFBBFD49286BD2DDFBE1" bold="true" box="[748,901,1120,1144]" pageId="6" pageNumber="467">Abbreviations</emphasis>
: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; mb = main branch of posterior gonopod; Md = basal joints of mandible; oz = ozopore; Pre = epiproct; rp = retrorse process; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; numbers refer to leg pair number.
<emphasis id="B949D804FFD2FFBBFB3128C7D45BFB7C" bold="true" box="[1172,1283,1228,1253]" pageId="6" pageNumber="467">Scale bars</emphasis>
= 1 mm.
</paragraph>
</caption>
<subSubSection id="C327579DFFD2FFBBFF32293DD444F813" pageId="6" pageNumber="467" type="description">
<paragraph id="8B820416FFD2FFBBFF32293DD070FAC9" blockId="6.[151,296,1334,1360]" box="[151,296,1334,1360]" pageId="6" pageNumber="467">
<emphasis id="B949D804FFD2FFBBFF32293DD070FAC9" bold="true" box="[151,296,1334,1360]" pageId="6" pageNumber="467">Description</emphasis>
</paragraph>
<paragraph id="8B820416FFD2FFBBFF322972D54FFA0B" blockId="6.[151,1437,1400,2038]" box="[151,1047,1400,1427]" pageId="6" pageNumber="467">
Measurements: 51+0 body rings plus telson, circa
<quantity id="4CC5A9F3FFD2FFBBFD692973D270FA0A" box="[716,808,1400,1427]" metricMagnitude="-1" metricUnit="m" metricValue="1.1" pageId="6" pageNumber="467" unit="mm" value="110.0">110 mm</quantity>
long,
<quantity id="4CC5A9F3FFD2FFBBFCCB2973D28AFA0A" box="[878,978,1400,1427]" metricMagnitude="-2" metricUnit="m" metricValue="1.02" pageId="6" pageNumber="467" unit="mm" value="10.2">10.2 mm</quantity>
wide.
</paragraph>
<paragraph id="8B820416FFD2FFBBFF622997D59AFA67" blockId="6.[151,1437,1400,2038]" pageId="6" pageNumber="467">
Colour faded in ethanol. Body rings laterally entirely black, above ozopores dorsally bright red with a wide black posterior margin. Collum black, anterior and posterior margin with a red stripe (
<figureCitation id="13061893FFD2FFBBFBFB29CBD5E0FA43" box="[1118,1208,1472,1498]" captionStart="FIGURE 2" captionStartId="6.[151,250,1012,1036]" captionTargetBox="[416,1172,237,989]" captionTargetId="figure-56@6.[416,1172,237,989]" captionTargetPageId="6" captionText="FIGURE 2. Aphistogoniulus amberivery, sp. nov., holotype (FMNH-INS 3196506). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 17, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Anterior gonopod, anterior view; E. Anterior gonopod, posterior view; F. Right posterior gonopod, anterior view; G. Right posterior gonopod, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; mb = main branch of posterior gonopod; Md = basal joints of mandible; oz = ozopore; Pre = epiproct; rp = retrorse process; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022754" httpUri="https://zenodo.org/record/14022754/files/figure.png" pageId="6" pageNumber="467">Fig. 2A</figureCitation>
). Head, antenna and legs black (
<figureCitation id="13061893FFD2FFBBFEB229EFD037FA67" box="[279,367,1508,1534]" captionStart="FIGURE 2" captionStartId="6.[151,250,1012,1036]" captionTargetBox="[416,1172,237,989]" captionTargetId="figure-56@6.[416,1172,237,989]" captionTargetPageId="6" captionText="FIGURE 2. Aphistogoniulus amberivery, sp. nov., holotype (FMNH-INS 3196506). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 17, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Anterior gonopod, anterior view; E. Anterior gonopod, posterior view; F. Right posterior gonopod, anterior view; G. Right posterior gonopod, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; mb = main branch of posterior gonopod; Md = basal joints of mandible; oz = ozopore; Pre = epiproct; rp = retrorse process; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022754" httpUri="https://zenodo.org/record/14022754/files/figure.png" pageId="6" pageNumber="467">Fig. 2B</figureCitation>
). Telson black, except for anal valves which have a large red spot (
<figureCitation id="13061893FFD2FFBBFBF829EFD5EEFA67" box="[1117,1206,1508,1534]" captionStart="FIGURE 2" captionStartId="6.[151,250,1012,1036]" captionTargetBox="[416,1172,237,989]" captionTargetId="figure-56@6.[416,1172,237,989]" captionTargetPageId="6" captionText="FIGURE 2. Aphistogoniulus amberivery, sp. nov., holotype (FMNH-INS 3196506). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 17, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Anterior gonopod, anterior view; E. Anterior gonopod, posterior view; F. Right posterior gonopod, anterior view; G. Right posterior gonopod, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; mb = main branch of posterior gonopod; Md = basal joints of mandible; oz = ozopore; Pre = epiproct; rp = retrorse process; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022754" httpUri="https://zenodo.org/record/14022754/files/figure.png" pageId="6" pageNumber="467">Fig. 2C</figureCitation>
).
</paragraph>
<paragraph id="8B820416FFD2FFBBFF622A03D5E4F9BB" blockId="6.[151,1437,1400,2038]" box="[199,1212,1544,1571]" pageId="6" pageNumber="467">
Head typical for the genus (
<figureCitation id="13061893FFD2FFBBFE5B2A03D320F9BB" box="[510,632,1544,1570]" captionStart="FIGURE 2" captionStartId="6.[151,250,1012,1036]" captionTargetBox="[416,1172,237,989]" captionTargetId="figure-56@6.[416,1172,237,989]" captionTargetPageId="6" captionText="FIGURE 2. Aphistogoniulus amberivery, sp. nov., holotype (FMNH-INS 3196506). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 17, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Anterior gonopod, anterior view; E. Anterior gonopod, posterior view; F. Right posterior gonopod, anterior view; G. Right posterior gonopod, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; mb = main branch of posterior gonopod; Md = basal joints of mandible; oz = ozopore; Pre = epiproct; rp = retrorse process; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022754" httpUri="https://zenodo.org/record/14022754/files/figure.png" pageId="6" pageNumber="467">Fig. 2A, B</figureCitation>
). Antenna short, extending back to ring 4 (
<figureCitation id="13061893FFD2FFBBFBF32A03D5F6F9BB" box="[1110,1198,1544,1570]" captionStart="FIGURE 2" captionStartId="6.[151,250,1012,1036]" captionTargetBox="[416,1172,237,989]" captionTargetId="figure-56@6.[416,1172,237,989]" captionTargetPageId="6" captionText="FIGURE 2. Aphistogoniulus amberivery, sp. nov., holotype (FMNH-INS 3196506). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 17, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Anterior gonopod, anterior view; E. Anterior gonopod, posterior view; F. Right posterior gonopod, anterior view; G. Right posterior gonopod, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; mb = main branch of posterior gonopod; Md = basal joints of mandible; oz = ozopore; Pre = epiproct; rp = retrorse process; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022754" httpUri="https://zenodo.org/record/14022754/files/figure.png" pageId="6" pageNumber="467">Fig. 2B</figureCitation>
).
</paragraph>
<paragraph id="8B820416FFD2FFBBFF622A27D545F9DF" blockId="6.[151,1437,1400,2038]" box="[199,1053,1580,1607]" pageId="6" pageNumber="467">Ozopore starting at ring 6, located on suture between meso- and metazonite.</paragraph>
<paragraph id="8B820416FFD2FFBBFF622A5BD2B2F9F3" blockId="6.[151,1437,1400,2038]" box="[199,1002,1616,1643]" pageId="6" pageNumber="467">
Male legs 37 with coxae and prefemora slightly invaginated (
<figureCitation id="13061893FFD2FFBBFC212A5BD284F9F3" box="[900,988,1616,1642]" captionStart="FIGURE 2" captionStartId="6.[151,250,1012,1036]" captionTargetBox="[416,1172,237,989]" captionTargetId="figure-56@6.[416,1172,237,989]" captionTargetPageId="6" captionText="FIGURE 2. Aphistogoniulus amberivery, sp. nov., holotype (FMNH-INS 3196506). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 17, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Anterior gonopod, anterior view; E. Anterior gonopod, posterior view; F. Right posterior gonopod, anterior view; G. Right posterior gonopod, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; mb = main branch of posterior gonopod; Md = basal joints of mandible; oz = ozopore; Pre = epiproct; rp = retrorse process; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022754" httpUri="https://zenodo.org/record/14022754/files/figure.png" pageId="6" pageNumber="467">Fig. 2B</figureCitation>
).
</paragraph>
<paragraph id="8B820416FFD2FFBBFF622A7FD190F92B" blockId="6.[151,1437,1400,2038]" pageId="6" pageNumber="467">
Telson without projecting preanal scale, anal valves without developed lips, subanal scale inconspicuous (
<figureCitation id="13061893FFD2FFBBFACA2A7FD1E4F92B" captionStart="FIGURE 2" captionStartId="6.[151,250,1012,1036]" captionTargetBox="[416,1172,237,989]" captionTargetId="figure-56@6.[416,1172,237,989]" captionTargetPageId="6" captionText="FIGURE 2. Aphistogoniulus amberivery, sp. nov., holotype (FMNH-INS 3196506). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 17, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Anterior gonopod, anterior view; E. Anterior gonopod, posterior view; F. Right posterior gonopod, anterior view; G. Right posterior gonopod, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; mb = main branch of posterior gonopod; Md = basal joints of mandible; oz = ozopore; Pre = epiproct; rp = retrorse process; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022754" httpUri="https://zenodo.org/record/14022754/files/figure.png" pageId="6" pageNumber="467">Fig. 2C</figureCitation>
).
</paragraph>
<paragraph id="8B820416FFD2FFBBFF622AB7D368F887" blockId="6.[151,1437,1400,2038]" pageId="6" pageNumber="467">
Anterior gonopod sternite triangular, no shoulders, rounded tip (
<figureCitation id="13061893FFD2FFBBFC022AB7D55CF94F" box="[935,1028,1724,1750]" captionStart="FIGURE 2" captionStartId="6.[151,250,1012,1036]" captionTargetBox="[416,1172,237,989]" captionTargetId="figure-56@6.[416,1172,237,989]" captionTargetPageId="6" captionText="FIGURE 2. Aphistogoniulus amberivery, sp. nov., holotype (FMNH-INS 3196506). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 17, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Anterior gonopod, anterior view; E. Anterior gonopod, posterior view; F. Right posterior gonopod, anterior view; G. Right posterior gonopod, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; mb = main branch of posterior gonopod; Md = basal joints of mandible; oz = ozopore; Pre = epiproct; rp = retrorse process; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022754" httpUri="https://zenodo.org/record/14022754/files/figure.png" pageId="6" pageNumber="467">Fig. 2D</figureCitation>
). Coxite process elongated, slightly shorter than telopodite (
<figureCitation id="13061893FFD2FFBBFE152AEBD36AF963" box="[432,562,1760,1786]" captionStart="FIGURE 2" captionStartId="6.[151,250,1012,1036]" captionTargetBox="[416,1172,237,989]" captionTargetId="figure-56@6.[416,1172,237,989]" captionTargetPageId="6" captionText="FIGURE 2. Aphistogoniulus amberivery, sp. nov., holotype (FMNH-INS 3196506). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 17, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Anterior gonopod, anterior view; E. Anterior gonopod, posterior view; F. Right posterior gonopod, anterior view; G. Right posterior gonopod, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; mb = main branch of posterior gonopod; Md = basal joints of mandible; oz = ozopore; Pre = epiproct; rp = retrorse process; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022754" httpUri="https://zenodo.org/record/14022754/files/figure.png" pageId="6" pageNumber="467">Fig. 2D, E</figureCitation>
). Telopodite appendage only weakly swollen, sharp-edged retrorse process starting to project apically (
<figureCitation id="13061893FFD2FFBBFE692B0FD37BF887" box="[460,547,1796,1822]" captionStart="FIGURE 2" captionStartId="6.[151,250,1012,1036]" captionTargetBox="[416,1172,237,989]" captionTargetId="figure-56@6.[416,1172,237,989]" captionTargetPageId="6" captionText="FIGURE 2. Aphistogoniulus amberivery, sp. nov., holotype (FMNH-INS 3196506). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 17, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Anterior gonopod, anterior view; E. Anterior gonopod, posterior view; F. Right posterior gonopod, anterior view; G. Right posterior gonopod, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; mb = main branch of posterior gonopod; Md = basal joints of mandible; oz = ozopore; Pre = epiproct; rp = retrorse process; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022754" httpUri="https://zenodo.org/record/14022754/files/figure.png" pageId="6" pageNumber="467">Fig. 2E</figureCitation>
).
</paragraph>
<paragraph id="8B820416FFD2FFBBFF622B23D444F813" blockId="6.[151,1437,1400,2038]" pageId="6" pageNumber="467">
Posterior gonopod telopodite branches forming a C (
<figureCitation id="13061893FFD2FFBBFC872B23D2CFF8DB" box="[802,919,1832,1858]" captionStart="FIGURE 2" captionStartId="6.[151,250,1012,1036]" captionTargetBox="[416,1172,237,989]" captionTargetId="figure-56@6.[416,1172,237,989]" captionTargetPageId="6" captionText="FIGURE 2. Aphistogoniulus amberivery, sp. nov., holotype (FMNH-INS 3196506). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 17, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Anterior gonopod, anterior view; E. Anterior gonopod, posterior view; F. Right posterior gonopod, anterior view; G. Right posterior gonopod, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; mb = main branch of posterior gonopod; Md = basal joints of mandible; oz = ozopore; Pre = epiproct; rp = retrorse process; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022754" httpUri="https://zenodo.org/record/14022754/files/figure.png" pageId="6" pageNumber="467">Fig. 2F, G</figureCitation>
). Basal branch stout, shorter than main branch (
<figureCitation id="13061893FFD2FFBBFF3A2B47D1ADF8FF" box="[159,245,1868,1894]" captionStart="FIGURE 2" captionStartId="6.[151,250,1012,1036]" captionTargetBox="[416,1172,237,989]" captionTargetId="figure-56@6.[416,1172,237,989]" captionTargetPageId="6" captionText="FIGURE 2. Aphistogoniulus amberivery, sp. nov., holotype (FMNH-INS 3196506). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 17, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Anterior gonopod, anterior view; E. Anterior gonopod, posterior view; F. Right posterior gonopod, anterior view; G. Right posterior gonopod, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; mb = main branch of posterior gonopod; Md = basal joints of mandible; oz = ozopore; Pre = epiproct; rp = retrorse process; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022754" httpUri="https://zenodo.org/record/14022754/files/figure.png" pageId="6" pageNumber="467">Fig. 2F</figureCitation>
) Tips of main and basal branch distant from one another. Main branch at apically with membranous the numerous membranous fringes (
<figureCitation id="13061893FFD2FFBBFDA42B7BD323F813" box="[513,635,1904,1930]" captionStart="FIGURE 2" captionStartId="6.[151,250,1012,1036]" captionTargetBox="[416,1172,237,989]" captionTargetId="figure-56@6.[416,1172,237,989]" captionTargetPageId="6" captionText="FIGURE 2. Aphistogoniulus amberivery, sp. nov., holotype (FMNH-INS 3196506). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 17, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Anterior gonopod, anterior view; E. Anterior gonopod, posterior view; F. Right posterior gonopod, anterior view; G. Right posterior gonopod, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; mb = main branch of posterior gonopod; Md = basal joints of mandible; oz = ozopore; Pre = epiproct; rp = retrorse process; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022754" httpUri="https://zenodo.org/record/14022754/files/figure.png" pageId="6" pageNumber="467">Figs 2F, G</figureCitation>
). Main branch slenderer but much longer than basal branch.
</paragraph>
</subSubSection>
<subSubSection id="C327579DFFD2FFBBFF622B9FD04AF86F" pageId="6" pageNumber="467" type="biology_ecology">
<paragraph id="8B820416FFD2FFBBFF622B9FD04AF86F" blockId="6.[151,1437,1400,2038]" pageId="6" pageNumber="467">
<emphasis id="B949D804FFD2FFBBFF622B9FD07EF837" bold="true" box="[199,294,1940,1966]" pageId="6" pageNumber="467">Ecology</emphasis>
.
<taxonomicName id="4C3D7F95FFD2FFBBFE932B9FD080F837" authorityName="Wesener" authorityYear="2024" box="[310,472,1940,1966]" class="Diplopoda" family="Pachybolidae" genus="Aphistogoniulus" kingdom="Animalia" order="Spirobolida" pageId="6" pageNumber="467" phylum="Arthropoda" rank="species" species="amberivery">
<emphasis id="B949D804FFD2FFBBFE932B9FD080F837" box="[310,472,1940,1966]" italics="true" pageId="6" pageNumber="467">A. amberivery</emphasis>
</taxonomicName>
occurs in sympatry with two microendemic species of
<taxonomicName id="4C3D7F95FFD2FFBBFBF32B9FD4C4F836" authority="Pocock, 1895" authorityName="Pocock" authorityYear="1895" box="[1110,1436,1940,1967]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="6" pageNumber="467" phylum="Arthropoda" rank="genus">
<emphasis id="B949D804FFD2FFBBFBF32B9FD5A0F837" box="[1110,1272,1940,1966]" italics="true" pageId="6" pageNumber="467">Zoosphaerium</emphasis>
<bibRefCitation id="EFAC79E7FFD2FFBBFAA42B9FD4C4F836" author="Pocock, R. I." box="[1281,1436,1940,1967]" pageId="6" pageNumber="467" pagination="409 - 415" refId="ref14376" refString="Pocock, R. I. (1895) Descriptions of new genera of Zephroniidae, with brief preliminary Diagnoses of some new Species. Annals and Magazine of Natural History, Series 6, 16 (95), 409 - 415. https: // doi. org / 10.1080 / 00222939508680293" type="journal article" year="1895">Pocock, 1895</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="EFAC79E7FFD2FFBBFF3A2BB3D090F84B" author="Sagorny, C. &amp; Wesener, T." box="[159,456,1976,2003]" pageId="6" pageNumber="467" pagination="273 - 294" refId="ref14604" refString="Sagorny, C. &amp; Wesener, T. (2017) Two new giant pill-millipede species of the genus Zoosphaerium endemic to the Bemanevika area in northern Madagascar (Diplopoda, Sphaerotheriida, Arthrosphaeridae). Zootaxa, 4263 (2), 273 - 294. https: // doi. org / 10.11646 / zootaxa. 4263.2.4" type="journal article" year="2017">Sagorny &amp; Wesener 2017</bibRefCitation>
), highlighting the only recently protected area of Bemanevika as a center of millipede endemism.
</paragraph>
</subSubSection>
</treatment>
</document>