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<document id="3E73C3C8DD2C4825BE3BBB58134F4F7B" ID-DOI="10.11646/zootaxa.5583.2.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="14798833" ID-ZooBank="FC4C2C8C-B379-4E7C-9FBB-116783DDD4A0" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1738625443653" checkinUser="plazi" docAuthor="Heppner, J. B., Arita, Yutaka &amp; Bae, Yang-Seop" docDate="2025" docId="03946562FFCCFF97FF1564C06760FE50" docLanguage="en" docName="zootaxa.5583.2.3.pdf" docOrigin="Zootaxa 5583 (2)" docSource="https://doi.org/10.11646/zootaxa.5583.2.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Thaumatographa aurosa subsp. taiwana Heppner &amp; Arita 2025, new subspecies" docType="treatment" docVersion="2" lastPageNumber="286" masterDocId="FFAD1D1AFFC2FF98FF8264576327FFD9" masterDocTitle="Review of Thaumatographa tortricids in Taiwan (Lepidoptera: Tortricidae: Chlidanotinae: Hilarographini)" masterLastPageNumber="292" masterPageNumber="271" pageNumber="285" updateTime="1738628064399" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="B0500CD44A4DD0F694DA95D487502824">
<mods:title id="FF9DE04C56811C3F0551AB3D33B011D7">Review of Thaumatographa tortricids in Taiwan (Lepidoptera: Tortricidae: Chlidanotinae: Hilarographini)</mods:title>
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<mods:name id="32B11AA4F0E5978210EB3B0871D79A16" type="personal">
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<mods:namePart id="E30708502A0FB3C632CB967434B79BEB">Heppner, J. B.</mods:namePart>
<mods:affiliation id="BD9CA798D8EA999E3EFB01ADCB11C026">McGuire Center for Lepidoptera and Biodiversity, Florida Museum of Natural History, University of Florida, Gainesville, Florida, USA. &amp; Bio ˗ Resource and Environmental Center, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012, Republic of Korea.</mods:affiliation>
</mods:name>
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<mods:namePart id="06BB0C2ED34E80115B2306E79FA13734">Arita, Yutaka</mods:namePart>
<mods:affiliation id="302B892A7BFBC8C673FDD2BB613DB4D6">Zoological Laboratory, Faculty of Agriculture, Meijo University, Tempaku-ku, Nagoya 468 - 85 - 2, Japan.</mods:affiliation>
</mods:name>
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<mods:namePart id="20349DD9BDC8610498DA447B7CB054B4">Bae, Yang-Seop</mods:namePart>
<mods:nameIdentifier id="8D5C4760E9957C3CD811CA2E65F04E52" type="ORCID">0000-0001-7356-5633</mods:nameIdentifier>
<mods:affiliation id="2C833597149C9539048150B4608D4316">Convergence Research Center for Insect Vectors, Division of Life Sciences, College of Life Sciences and Bioengineering, Incheon National University, Songdo ˗ dong, Incheon, 22012, Republic of Korea. &amp; Bio ˗ Resource and Environmental Center, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012, Republic of Korea.</mods:affiliation>
<mods:nameIdentifier id="F94CAE54B1C60437CAAADFE7AF030D2C" type="email">baeys@inu.ac.kr</mods:nameIdentifier>
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<mods:typeOfResource id="6D0CCBE8EF01C6E506F1BD0D3BEBBEBC">text</mods:typeOfResource>
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<mods:title id="5C5DD50AD26C3E91BA646DC057DE6ED9">Zootaxa</mods:title>
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<mods:date id="DCF2E1EEBC91966F47D11E778AB4102C">2025</mods:date>
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<mods:number id="2137B171990B11F43280325B25D174B2">2025-02-03</mods:number>
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<treatment id="03946562FFCCFF97FF1564C06760FE50" LSID="urn:lsid:plazi:treatment:03946562FFCCFF97FF1564C06760FE50" httpUri="http://treatment.plazi.org/id/03946562FFCCFF97FF1564C06760FE50" lastPageId="15" lastPageNumber="286" pageId="14" pageNumber="285">
<subSubSection id="C32787FFFFCCFF96FF1564C0673EFF0F" pageId="14" pageNumber="285" type="nomenclature">
<paragraph id="8B82D474FFCCFF96FF1564C060F2FF68" blockId="14.[151,1049,151,250]" box="[151,981,151,178]" pageId="14" pageNumber="285">
<heading id="D0CA6318FFCCFF96FF1564C060F2FF68" bold="true" box="[151,981,151,178]" fontSize="11" level="1" pageId="14" pageNumber="285" reason="1">
<emphasis id="B9490866FFCCFF96FF1564C060F2FF68" bold="true" box="[151,981,151,178]" pageId="14" pageNumber="285">
<taxonomicName id="4C3DAFF7FFCCFF96FF1564C0602AFF68" authority="Heppner &amp; Arita" authorityName="Heppner &amp; Arita" authorityYear="2025" box="[151,781,151,178]" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="285" phylum="Arthropoda" rank="subSpecies" species="aurosa" status="subsp. nov." subSpecies="taiwana">
<emphasis id="B9490866FFCCFF96FF1564C0610AFF68" bold="true" box="[151,557,151,178]" italics="true" pageId="14" pageNumber="285">Thaumatographa aurosa taiwana</emphasis>
Heppner &amp; Arita
</taxonomicName>
,
<taxonomicNameLabel id="A27AB51DFFCCFF96FC9864C060F2FF68" box="[794,981,151,177]" pageId="14" pageNumber="285" rank="subSpecies">new subspecies</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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LSID:
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urn:lsid:zoobank.org:act:
<uuid id="FF9BEEA1FFCCFF96FE7764EC673EFF0F" box="[501,1049,187,214]" pageId="14" pageNumber="285">4A472AE9-FC2A-47BB-80C5-E5028A88169C</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C32787FFFFCCFF96FF1564886017FEE5" pageId="14" pageNumber="285" type="description">
<paragraph id="8B82D474FFCCFF96FF156488623EFF23" blockId="14.[151,1049,151,250]" box="[151,281,223,250]" pageId="14" pageNumber="285">
(
<figureCitation id="1306C8F1FFCCFF96FF1D648863CFFF23" box="[159,232,223,250]" captionStart="FIGURES 16" captionStartId="4.[151,264,1560,1584]" captionTargetBox="[151,1436,555,1536]" captionTargetId="figure-133@4.[151,1436,555,1536]" captionTargetPageId="4" captionText="FIGURES 16. Adults of Taiwan Thaumatographa species, with lateral views of heads (a) for each moth: 1) T. mesostigmatias Diakonoff &amp; Arita, male (Chiayi Co.: Fennchihwu, photo 5569) (WS = 13.0mm); 2) same data, female (photo14173) (WS = 14.8 mm) (MGCL). 3) T. fennchihwu Heppner &amp; Arita, sp. nov., male holotype (Chiayi Co.: Fennchihwu, photo 5570) (WS = 15.5 mm); 4) same data, female paratype (photo 5571) (WS = 19.0 mm) (MGCL). 5) T. aurosa taiwana Heppner &amp;Arita, subsp. nov., female paratype (Ilan Co.: Nanao, photo 16014) (WS = 13.6 mm) (MGCL). 6) T. kentingana Heppner &amp; Bae, sp. nov., male holotype (Pingtung Co.: Kenting, photo 5562) (WS = 7.5 mm) (USNM). Note: the white mark in middle of wing Fig. 2 is partially descaled, thus otherwise as small as in the male Fig. 1. Also, note the bluish iridescence of the labial palpi in Fig. 1a and 2a." figureDoi="http://doi.org/10.5281/zenodo.14798835" httpUri="https://zenodo.org/record/14798835/files/figure.png" pageId="14" pageNumber="285">Figs 5</figureCitation>
,
<figureCitation id="1306C8F1FFCCFF96FF7664886236FF23" box="[244,273,223,250]" captionStart="FIGURE 15" captionStartId="15.[151,250,1858,1882]" captionTargetBox="[151,1436,453,1835]" captionTargetId="figure-128@15.[151,1436,453,1835]" captionTargetPageId="15" captionText="FIGURE 15. Thaumatographa aurosa taiwana Heppner &amp; Arita, subsp. nov., paratype female (Taiwan: Lien-Hua-Chih For. Sta., Nantou Co.), with details of ostium, sterigma, and antrum diverticulum (a), and signum, (b) (gen. slide JBH-4644 (MGCL)." figureDoi="http://doi.org/10.5281/zenodo.14798853" httpUri="https://zenodo.org/record/14798853/files/figure.png" pageId="14" pageNumber="285">15</figureCitation>
)
</paragraph>
<paragraph id="8B82D474FFCCFF96FF1565736017FEE5" blockId="14.[151,816,292,316]" box="[151,816,292,316]" pageId="14" pageNumber="285">
Type-locality:
<collectingCountry id="F32A94E4FFCCFF96FEA96573625EFEE5" box="[299,377,292,316]" name="Taiwan" pageId="14" pageNumber="285">Taiwan</collectingCountry>
: Lien-Hua-Chih For. Sta., Puli,
<collectingRegion id="49F91A96FFCCFF96FD3965736021FEE2" box="[699,774,292,315]" country="Taiwan" name="Nantou" pageId="14" pageNumber="285">Nantou</collectingRegion>
Co.
</paragraph>
</subSubSection>
<subSubSection id="C32787FFFFCCFF96FF15653D6786FE29" pageId="14" pageNumber="285" type="materials_examined">
<paragraph id="8B82D474FFCCFF96FF15653D6786FE29" blockId="14.[151,1437,361,1972]" pageId="14" pageNumber="285">
<emphasis id="B9490866FFCCFF96FF15653D6261FE5A" bold="true" box="[151,326,361,388]" pageId="14" pageNumber="285">Type material.</emphasis>
<materialsCitation id="3B55DE29FFCCFF96FECC653D6015FE71" collectingDate="1983-09-07" collectingDateMax="1983-09-12" collectingDateMin="1983-09-07" collectionCode="MGCL" collectorName="J. B. Heppner" country="Taiwan" elevation="700" location="Nantou" pageId="14" pageNumber="285" specimenCount="1" specimenCount-female="1" stateProvince="Nantou" typeStatus="holotype">
<emphasis id="B9490866FFCCFF96FECC653D6112FE5A" bold="true" box="[334,565,361,388]" pageId="14" pageNumber="285">
<typeStatus id="54866AD6FFCCFF96FECC653D629AFE5D" box="[334,445,362,388]" pageId="14" pageNumber="285" type="holotype">Holotype</typeStatus>
. [
<collectingCountry id="F32A94E4FFCCFF96FE53653E6109FE5A" box="[465,558,361,387]" name="Taiwan" pageId="14" pageNumber="285">Taiwan</collectingCountry>
]
</emphasis>
<specimenCount id="9D3B1FFDFFCCFF96FDBF653D6175FE5D" box="[573,594,362,388]" count="1" pageId="14" pageNumber="285" type="female"></specimenCount>
Lien-Hua-Chih For. Sta. (
<quantity id="4CC57991FFCCFF96FCFE653D60EEFE5D" box="[892,969,362,388]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="14" pageNumber="285" unit="m" value="700.0">
<elevation id="00103347FFCCFF96FCFE653D60EEFE5D" box="[892,969,362,388]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="14" pageNumber="285" unit="m" value="700.0">700 m</elevation>
</quantity>
), [nr. Puli],
<collectingRegion id="49F91A96FFCCFF96FBD0653D6783FE5D" box="[1106,1188,362,388]" country="Taiwan" name="Nantou" pageId="14" pageNumber="285">Nantou</collectingRegion>
Co.,
<date id="FF83F2B4FFCCFF96FB66653D66BFFE5D" box="[1252,1432,362,388]" pageId="14" pageNumber="285" value="1983-09-07" valueMax="1983-09-12" valueMin="1983-09-07">
<collectingDate id="EFC70B5CFFCCFF96FB66653D66BFFE5D" box="[1252,1432,362,388]" pageId="14" pageNumber="285" value="1983-09-07" valueMax="1983-09-12" valueMin="1983-09-07">712 Sep. 1983</collectingDate>
</date>
,
<collectorName id="26C8B1A2FFCCFF96FF1565D96216FE71" box="[151,305,398,424]" pageId="14" pageNumber="285">J. B. Heppner</collectorName>
(adult photo 5572; gen. JBH-4643) (
<collectionCode id="ED2C4CB1FFCCFF96FD5065D9600EFE71" box="[722,809,398,424]" pageId="14" pageNumber="285">MGCL</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B55DE29FFCCFF96FCBE65D967BAFE29" collectingDate="1984-10-11" collectingDateMax="1984-10-13" collectingDateMin="1984-10-11" collectionCode="MGCL" collectorName="J. B. Heppner &amp; H. - Y. Wang" country="Taiwan" county="Ilan Co." elevation="100" location="Taiwan" municipality="Taiwan" pageId="14" pageNumber="285" specimenCount="2" specimenCount-female="2" stateProvince="Nantou" typeStatus="paratype">
<emphasis id="B9490866FFCCFF96FCBE65D96093FE71" bold="true" box="[828,948,398,424]" pageId="14" pageNumber="285">
<typeStatus id="54866AD6FFCCFF96FCBE65D96093FE71" box="[828,948,398,424]" pageId="14" pageNumber="285" type="paratype">Paratypes</typeStatus>
</emphasis>
. (n = 2).
<collectingMunicipality id="6BE64E0EFFCCFF96FB9F65D96755FE71" box="[1053,1138,398,424]" pageId="14" pageNumber="285">Taiwan</collectingMunicipality>
,
<collectingRegion id="49F91A96FFCCFF96FBFC65D967F7FE71" box="[1150,1232,398,424]" country="Taiwan" name="Nantou" pageId="14" pageNumber="285">Nantou</collectingRegion>
Co.: same data as holotype (
<specimenCount id="9D3B1FFDFFCCFF96FE8F65E5620BFE15" box="[269,300,434,460]" count="1" pageId="14" pageNumber="285" type="female">1♀</specimenCount>
) (adult photo 16013; gen. JBH-4644) (
<collectionCode id="ED2C4CB1FFCCFF96FD6C65E56062FE15" box="[750,837,434,460]" pageId="14" pageNumber="285">MGCL</collectionCode>
).
<location id="8EE282AFFFCCFF96FCDB65E56089FE15" LSID="urn:lsid:plazi:treatment:03946562FFCCFF97FF1564C06760FE50:8EE282AFFFCCFF96FCDB65E56089FE15" box="[857,942,434,460]" county="Ilan Co." municipality="Taiwan" name="Taiwan" pageId="14" pageNumber="285" stateProvince="Nantou">Taiwan</location>
,
<collectingCounty id="62E3ACF8FFCCFF96FC3965E5673CFE15" box="[955,1051,434,460]" pageId="14" pageNumber="285">Ilan Co.</collectingCounty>
:
<location id="8EE282AFFFCCFF96FBA565E56757FE15" LSID="urn:lsid:plazi:treatment:03946562FFCCFF97FF1564C06760FE50:8EE282AFFFCCFF96FBA565E56757FE15" box="[1063,1136,434,460]" county="Ilan Co." municipality="Taiwan" name="Nanao" pageId="14" pageNumber="285" stateProvince="Nantou">Nanao</location>
(
<quantity id="4CC57991FFCCFF96FB0065E56799FE15" box="[1154,1214,434,461]" metricMagnitude="3" metricUnit="m" metricValue="8.04672" pageId="14" pageNumber="285" unit="mi" value="5.0">5 mi.</quantity>
N) (
<quantity id="4CC57991FFCCFF96FB7665E56666FE15" box="[1268,1345,434,460]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="285" unit="m" value="100.0">
<elevation id="00103347FFCCFF96FB7665E56666FE15" box="[1268,1345,434,460]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="285" unit="m" value="100.0">100 m</elevation>
</quantity>
),
<date id="FF83F2B4FFCCFF96FAD565E56221FE29" pageId="14" pageNumber="285" value="1984-10-11" valueMax="1984-10-13" valueMin="1984-10-11">
<collectingDate id="EFC70B5CFFCCFF96FAD565E56221FE29" pageId="14" pageNumber="285" value="1984-10-11" valueMax="1984-10-13" valueMin="1984-10-11">1113 Oct. 1984</collectingDate>
</date>
(
<specimenCount id="9D3B1FFDFFCCFF96FE9A6581621FFE29" box="[280,312,470,496]" count="1" pageId="14" pageNumber="285" type="female">1♀</specimenCount>
),
<collectorName id="26C8B1A2FFCCFF96FED2658162CDFE29" box="[336,490,470,496]" pageId="14" pageNumber="285">J. B. Heppner</collectorName>
&amp;
<collectorName id="26C8B1A2FFCCFF96FD8C658161B5FE29" box="[526,658,470,496]" pageId="14" pageNumber="285">H.-Y. Wang</collectorName>
(adult photo 16014; gen. JBH-4642) (
<collectionCode id="ED2C4CB1FFCCFF96FBBC658167B2FE29" box="[1086,1173,470,496]" pageId="14" pageNumber="285">MGCL</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32787FFFFCCFF96FF4565AE62C0FD35" pageId="14" pageNumber="285" type="diagnosis">
<paragraph id="8B82D474FFCCFF96FF4565AE62C0FD35" blockId="14.[151,1437,361,1972]" pageId="14" pageNumber="285">
<emphasis id="B9490866FFCCFF96FF4565AE621DFDCA" bold="true" box="[199,314,505,531]" pageId="14" pageNumber="285">Diagnosis</emphasis>
. This new subspecies is very similar to
<taxonomicName id="4C3DAFF7FFCCFF96FC9265AD667BFDCD" authority="Diakonoff &amp; Arita (1976)" authorityName="Diakonoff &amp; Arita" authorityYear="1976" box="[784,1372,506,532]" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="285" phylum="Arthropoda" rank="species" species="aurosa">
<emphasis id="B9490866FFCCFF96FC9265AD670DFDCD" box="[784,1066,506,532]" italics="true" pageId="14" pageNumber="285">Thaumatographa aurosa</emphasis>
<bibRefCitation id="EFACA985FFCCFF96FBB165AD667BFDCD" author="Diakonoff, A. &amp; Arita, Y." box="[1075,1372,506,532]" pageId="14" pageNumber="285" pagination="179 - 219" refId="ref11038" refString="Diakonoff, A. &amp; Arita, Y. (1976) New and old species of Glyphipterigidae and a new species of Acrolepiidae from Japan and the Far East (Lepidoptera). Zoologische Mededelingen, 50, 179-219." type="journal article" year="1976">Diakonoff &amp; Arita (1976)</bibRefCitation>
</taxonomicName>
from southern
<collectingCountry id="F32A94E4FFCCFF96FE8066496265FDE1" box="[258,322,542,568]" name="Japan" pageId="14" pageNumber="285">Japan</collectingCountry>
(Kyushu and Yakushima), and has the following distinctions: on the forewing the basal pair of vertical orange lines are more curved distally from the dorsal margin (straight in nominate
<taxonomicName id="4C3DAFF7FFCCFF96FB09661467D4FD85" authorityName="Diakonoff &amp; Arita" authorityYear="1976" box="[1163,1267,579,604]" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="285" phylum="Arthropoda" rank="species" species="aurosa">
<emphasis id="B9490866FFCCFF96FB09661467D4FD85" box="[1163,1267,579,604]" italics="true" pageId="14" pageNumber="285">T. aurosa</emphasis>
</taxonomicName>
) and the costal angled streaks are white on the costal margin (silvery blue in nominate
<taxonomicName id="4C3DAFF7FFCCFF96FC2F66306733FD59" authorityName="Diakonoff &amp; Arita" authorityYear="1976" box="[941,1044,615,640]" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="285" phylum="Arthropoda" rank="species" species="aurosa">
<emphasis id="B9490866FFCCFF96FC2F66306733FD59" box="[941,1044,615,640]" italics="true" pageId="14" pageNumber="285">T. aurosa</emphasis>
</taxonomicName>
). The female genitalia are virtually identical and
<taxonomicName id="4C3DAFF7FFCCFF96FEAC66DC62DFFD7D" authorityName="Heppner &amp; Arita" authorityYear="2025" box="[302,504,651,676]" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="285" phylum="Arthropoda" rank="subSpecies" species="aurosa" subSpecies="taiwana">
<emphasis id="B9490866FFCCFF96FEAC66DC62DFFD7D" box="[302,504,651,676]" italics="true" pageId="14" pageNumber="285">T. aurosa taiwana</emphasis>
</taxonomicName>
has the ovipositor somewhat more slender caudally, especially very narrow papilla anales (slightly wider in the nominate subspecies from
<collectingCountry id="F32A94E4FFCCFF96FC8566F9606BFD11" box="[775,844,686,712]" name="Japan" pageId="14" pageNumber="285">Japan</collectingCountry>
). Male genitalia could not be compared, lacking any male specimens from
<collectingCountry id="F32A94E4FFCCFF96FE0F668562C4FD35" box="[397,483,722,748]" name="Taiwan" pageId="14" pageNumber="285">Taiwan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C32787FFFFCCFF96FF4566A26209F86D" pageId="14" pageNumber="285" type="description">
<paragraph id="8B82D474FFCCFF96FF4566A26785FCC9" blockId="14.[151,1437,361,1972]" box="[199,1186,757,784]" pageId="14" pageNumber="285">
<emphasis id="B9490866FFCCFF96FF4566A26276FCD6" bold="true" box="[199,337,757,783]" pageId="14" pageNumber="285">Description</emphasis>
. Wing expanse: male 10.014.0 mm (
<collectingCountry id="F32A94E4FFCCFF96FD7966A16067FCC9" box="[763,832,758,784]" name="Japan" pageId="14" pageNumber="285">Japan</collectingCountry>
), female 11.013.6 mm (n = 3).
</paragraph>
<paragraph id="8B82D474FFCCFF96FF45674D6109FCEC" blockId="14.[151,1437,361,1972]" box="[199,558,794,821]" pageId="14" pageNumber="285">
Female (
<figureCitation id="1306C8F1FFCCFF96FEAB674D624AFCEC" box="[297,365,794,821]" captionStart="FIGURES 16" captionStartId="4.[151,264,1560,1584]" captionTargetBox="[151,1436,555,1536]" captionTargetId="figure-133@4.[151,1436,555,1536]" captionTargetPageId="4" captionText="FIGURES 16. Adults of Taiwan Thaumatographa species, with lateral views of heads (a) for each moth: 1) T. mesostigmatias Diakonoff &amp; Arita, male (Chiayi Co.: Fennchihwu, photo 5569) (WS = 13.0mm); 2) same data, female (photo14173) (WS = 14.8 mm) (MGCL). 3) T. fennchihwu Heppner &amp; Arita, sp. nov., male holotype (Chiayi Co.: Fennchihwu, photo 5570) (WS = 15.5 mm); 4) same data, female paratype (photo 5571) (WS = 19.0 mm) (MGCL). 5) T. aurosa taiwana Heppner &amp;Arita, subsp. nov., female paratype (Ilan Co.: Nanao, photo 16014) (WS = 13.6 mm) (MGCL). 6) T. kentingana Heppner &amp; Bae, sp. nov., male holotype (Pingtung Co.: Kenting, photo 5562) (WS = 7.5 mm) (USNM). Note: the white mark in middle of wing Fig. 2 is partially descaled, thus otherwise as small as in the male Fig. 1. Also, note the bluish iridescence of the labial palpi in Fig. 1a and 2a." figureDoi="http://doi.org/10.5281/zenodo.14798835" httpUri="https://zenodo.org/record/14798835/files/figure.png" pageId="14" pageNumber="285">Fig. 5</figureCitation>
).
<emphasis id="B9490866FFCCFF96FE01674D62E7FCED" box="[387,448,794,820]" italics="true" pageId="14" pageNumber="285">Head</emphasis>
(
<figureCitation id="1306C8F1FFCCFF96FE4D674D6106FCEC" box="[463,545,794,821]" captionStart="FIGURES 16" captionStartId="4.[151,264,1560,1584]" captionTargetBox="[151,1436,555,1536]" captionTargetId="figure-133@4.[151,1436,555,1536]" captionTargetPageId="4" captionText="FIGURES 16. Adults of Taiwan Thaumatographa species, with lateral views of heads (a) for each moth: 1) T. mesostigmatias Diakonoff &amp; Arita, male (Chiayi Co.: Fennchihwu, photo 5569) (WS = 13.0mm); 2) same data, female (photo14173) (WS = 14.8 mm) (MGCL). 3) T. fennchihwu Heppner &amp; Arita, sp. nov., male holotype (Chiayi Co.: Fennchihwu, photo 5570) (WS = 15.5 mm); 4) same data, female paratype (photo 5571) (WS = 19.0 mm) (MGCL). 5) T. aurosa taiwana Heppner &amp;Arita, subsp. nov., female paratype (Ilan Co.: Nanao, photo 16014) (WS = 13.6 mm) (MGCL). 6) T. kentingana Heppner &amp; Bae, sp. nov., male holotype (Pingtung Co.: Kenting, photo 5562) (WS = 7.5 mm) (USNM). Note: the white mark in middle of wing Fig. 2 is partially descaled, thus otherwise as small as in the male Fig. 1. Also, note the bluish iridescence of the labial palpi in Fig. 1a and 2a." figureDoi="http://doi.org/10.5281/zenodo.14798835" httpUri="https://zenodo.org/record/14798835/files/figure.png" pageId="14" pageNumber="285">Fig. 5a</figureCitation>
).
</paragraph>
<paragraph id="8B82D474FFCCFF96FF45676860E1FB45" blockId="14.[151,1437,361,1972]" pageId="14" pageNumber="285">
<emphasis id="B9490866FFCCFF96FF4567686214FC81" box="[199,307,831,856]" italics="true" pageId="14" pageNumber="285">Forewing</emphasis>
(
<figureCitation id="1306C8F1FFCCFF96FEC6676962ADFC80" box="[324,394,830,857]" captionStart="FIGURES 16" captionStartId="4.[151,264,1560,1584]" captionTargetBox="[151,1436,555,1536]" captionTargetId="figure-133@4.[151,1436,555,1536]" captionTargetPageId="4" captionText="FIGURES 16. Adults of Taiwan Thaumatographa species, with lateral views of heads (a) for each moth: 1) T. mesostigmatias Diakonoff &amp; Arita, male (Chiayi Co.: Fennchihwu, photo 5569) (WS = 13.0mm); 2) same data, female (photo14173) (WS = 14.8 mm) (MGCL). 3) T. fennchihwu Heppner &amp; Arita, sp. nov., male holotype (Chiayi Co.: Fennchihwu, photo 5570) (WS = 15.5 mm); 4) same data, female paratype (photo 5571) (WS = 19.0 mm) (MGCL). 5) T. aurosa taiwana Heppner &amp;Arita, subsp. nov., female paratype (Ilan Co.: Nanao, photo 16014) (WS = 13.6 mm) (MGCL). 6) T. kentingana Heppner &amp; Bae, sp. nov., male holotype (Pingtung Co.: Kenting, photo 5562) (WS = 7.5 mm) (USNM). Note: the white mark in middle of wing Fig. 2 is partially descaled, thus otherwise as small as in the male Fig. 1. Also, note the bluish iridescence of the labial palpi in Fig. 1a and 2a." figureDoi="http://doi.org/10.5281/zenodo.14798835" httpUri="https://zenodo.org/record/14798835/files/figure.png" pageId="14" pageNumber="285">Fig. 5</figureCitation>
): overall black-brown, with usual geneedric striae pattern, with five thin costal striae white and slightly silvery from margin (more silvery in nominate subspecies), with basal two strongly bent to termen on radius; basal streaks straight and long, with dorsal on orange-yellow and middle streak yellow-white, and at end of forewing cell is a prominent tan-white spot; three curved light yellow-white lines from dorsal margin at middle of forewing, with a 4
<superScript id="7C48793CFFCCFF96FE07679A62B5FC02" attach="left" box="[389,402,973,987]" fontSize="6" pageId="14" pageNumber="285">th</superScript>
slightly further distad; tornal quarter with four series of interrupted striae curved slightly to termen; forewing apex with a white subapical line convergent and recurved as a loop to similar preapical line, both margined with orange; termen orange, with mid-termen white margin on termen and toward tornus 3 black spots; fringe white; venter dark brown with light yellow in cell and dorsal markings faintly repeated except for distinct white marks on costa, and dorsal margin white; venter brown with dorsal lines repeated as dull light tan lines, with white liens of apex repeated, and likewise the tornal markings but duller.
</paragraph>
<paragraph id="8B82D474FFCCFF96FF4560F163E5FAD1" blockId="14.[151,1437,361,1972]" pageId="14" pageNumber="285">
<emphasis id="B9490866FFCCFF96FF4560F16212FB19" box="[199,309,1190,1216]" italics="true" pageId="14" pageNumber="285">Hindwing</emphasis>
(
<figureCitation id="1306C8F1FFCCFF96FEC660F162A0FB18" box="[324,391,1190,1217]" captionStart="FIGURES 16" captionStartId="4.[151,264,1560,1584]" captionTargetBox="[151,1436,555,1536]" captionTargetId="figure-133@4.[151,1436,555,1536]" captionTargetPageId="4" captionText="FIGURES 16. Adults of Taiwan Thaumatographa species, with lateral views of heads (a) for each moth: 1) T. mesostigmatias Diakonoff &amp; Arita, male (Chiayi Co.: Fennchihwu, photo 5569) (WS = 13.0mm); 2) same data, female (photo14173) (WS = 14.8 mm) (MGCL). 3) T. fennchihwu Heppner &amp; Arita, sp. nov., male holotype (Chiayi Co.: Fennchihwu, photo 5570) (WS = 15.5 mm); 4) same data, female paratype (photo 5571) (WS = 19.0 mm) (MGCL). 5) T. aurosa taiwana Heppner &amp;Arita, subsp. nov., female paratype (Ilan Co.: Nanao, photo 16014) (WS = 13.6 mm) (MGCL). 6) T. kentingana Heppner &amp; Bae, sp. nov., male holotype (Pingtung Co.: Kenting, photo 5562) (WS = 7.5 mm) (USNM). Note: the white mark in middle of wing Fig. 2 is partially descaled, thus otherwise as small as in the male Fig. 1. Also, note the bluish iridescence of the labial palpi in Fig. 1a and 2a." figureDoi="http://doi.org/10.5281/zenodo.14798835" httpUri="https://zenodo.org/record/14798835/files/figure.png" pageId="14" pageNumber="285">Fig. 5</figureCitation>
): dark brown with thin orange-yellow streak diffusely margined in cell area, and diffuse tawny orange area on anal sector to base; fringe white; venter bronze-brown, with pale white markings on costa on basal 1/3.
</paragraph>
<paragraph id="8B82D474FFCCFF96FF4561456149FAAD" blockId="14.[151,1437,361,1972]" pageId="14" pageNumber="285">
<emphasis id="B9490866FFCCFF96FF456145623FFAF5" box="[199,280,1298,1324]" italics="true" pageId="14" pageNumber="285">Thorax</emphasis>
(
<figureCitation id="1306C8F1FFCCFF96FEAB61456257FAF4" box="[297,368,1298,1325]" captionStart="FIGURES 16" captionStartId="4.[151,264,1560,1584]" captionTargetBox="[151,1436,555,1536]" captionTargetId="figure-133@4.[151,1436,555,1536]" captionTargetPageId="4" captionText="FIGURES 16. Adults of Taiwan Thaumatographa species, with lateral views of heads (a) for each moth: 1) T. mesostigmatias Diakonoff &amp; Arita, male (Chiayi Co.: Fennchihwu, photo 5569) (WS = 13.0mm); 2) same data, female (photo14173) (WS = 14.8 mm) (MGCL). 3) T. fennchihwu Heppner &amp; Arita, sp. nov., male holotype (Chiayi Co.: Fennchihwu, photo 5570) (WS = 15.5 mm); 4) same data, female paratype (photo 5571) (WS = 19.0 mm) (MGCL). 5) T. aurosa taiwana Heppner &amp;Arita, subsp. nov., female paratype (Ilan Co.: Nanao, photo 16014) (WS = 13.6 mm) (MGCL). 6) T. kentingana Heppner &amp; Bae, sp. nov., male holotype (Pingtung Co.: Kenting, photo 5562) (WS = 7.5 mm) (USNM). Note: the white mark in middle of wing Fig. 2 is partially descaled, thus otherwise as small as in the male Fig. 1. Also, note the bluish iridescence of the labial palpi in Fig. 1a and 2a." figureDoi="http://doi.org/10.5281/zenodo.14798835" httpUri="https://zenodo.org/record/14798835/files/figure.png" pageId="14" pageNumber="285">Fig. 5</figureCitation>
): like nominate subspecies, dark brown, lacking any longitudinal lines; tegula slightly lighter brown than thorax; venter white; legs white, with dark brown markings dorsally on tibiae like related species, as dorsal spots on tibiae and tarsal segments.
</paragraph>
<paragraph id="8B82D474FFCCFF96FF45612966AAFA41" blockId="14.[151,1437,361,1972]" box="[199,1421,1406,1433]" pageId="14" pageNumber="285">
<emphasis id="B9490866FFCCFF96FF4561296216FA41" box="[199,305,1406,1432]" italics="true" pageId="14" pageNumber="285">Abdomen</emphasis>
(
<figureCitation id="1306C8F1FFCCFF96FEC2612962A2FA40" box="[320,389,1406,1433]" captionStart="FIGURES 16" captionStartId="4.[151,264,1560,1584]" captionTargetBox="[151,1436,555,1536]" captionTargetId="figure-133@4.[151,1436,555,1536]" captionTargetPageId="4" captionText="FIGURES 16. Adults of Taiwan Thaumatographa species, with lateral views of heads (a) for each moth: 1) T. mesostigmatias Diakonoff &amp; Arita, male (Chiayi Co.: Fennchihwu, photo 5569) (WS = 13.0mm); 2) same data, female (photo14173) (WS = 14.8 mm) (MGCL). 3) T. fennchihwu Heppner &amp; Arita, sp. nov., male holotype (Chiayi Co.: Fennchihwu, photo 5570) (WS = 15.5 mm); 4) same data, female paratype (photo 5571) (WS = 19.0 mm) (MGCL). 5) T. aurosa taiwana Heppner &amp;Arita, subsp. nov., female paratype (Ilan Co.: Nanao, photo 16014) (WS = 13.6 mm) (MGCL). 6) T. kentingana Heppner &amp; Bae, sp. nov., male holotype (Pingtung Co.: Kenting, photo 5562) (WS = 7.5 mm) (USNM). Note: the white mark in middle of wing Fig. 2 is partially descaled, thus otherwise as small as in the male Fig. 1. Also, note the bluish iridescence of the labial palpi in Fig. 1a and 2a." figureDoi="http://doi.org/10.5281/zenodo.14798835" httpUri="https://zenodo.org/record/14798835/files/figure.png" pageId="14" pageNumber="285">Fig. 5</figureCitation>
): dark brown, with posterior margins of tergites tan; venter cream-white; genital tuft brown.
</paragraph>
<paragraph id="8B82D474FFCCFF96FF4561F560AFF961" blockId="14.[151,1437,361,1972]" pageId="14" pageNumber="285">
<emphasis id="B9490866FFCCFF96FF4561F562AEFA65" box="[199,393,1442,1468]" italics="true" pageId="14" pageNumber="285">Female genitalia</emphasis>
(
<figureCitation id="1306C8F1FFCCFF96FE1961F562D6FA64" box="[411,497,1442,1469]" captionStart="FIGURE 15" captionStartId="15.[151,250,1858,1882]" captionTargetBox="[151,1436,453,1835]" captionTargetId="figure-128@15.[151,1436,453,1835]" captionTargetPageId="15" captionText="FIGURE 15. Thaumatographa aurosa taiwana Heppner &amp; Arita, subsp. nov., paratype female (Taiwan: Lien-Hua-Chih For. Sta., Nantou Co.), with details of ostium, sterigma, and antrum diverticulum (a), and signum, (b) (gen. slide JBH-4644 (MGCL)." figureDoi="http://doi.org/10.5281/zenodo.14798853" httpUri="https://zenodo.org/record/14798853/files/figure.png" pageId="14" pageNumber="285">Fig. 15</figureCitation>
): ovipositor short (subequal to length of segment 7) (dorsal flap lacking); papilla anales narrow, with caudal ends semi-acute; apophyses with anterior pair slightly longer; ostium (
<figureCitation id="1306C8F1FFCCFF96FB5F6191661BFA38" box="[1245,1340,1478,1505]" captionStart="FIGURE 15" captionStartId="15.[151,250,1858,1882]" captionTargetBox="[151,1436,453,1835]" captionTargetId="figure-128@15.[151,1436,453,1835]" captionTargetPageId="15" captionText="FIGURE 15. Thaumatographa aurosa taiwana Heppner &amp; Arita, subsp. nov., paratype female (Taiwan: Lien-Hua-Chih For. Sta., Nantou Co.), with details of ostium, sterigma, and antrum diverticulum (a), and signum, (b) (gen. slide JBH-4644 (MGCL)." figureDoi="http://doi.org/10.5281/zenodo.14798853" httpUri="https://zenodo.org/record/14798853/files/figure.png" pageId="14" pageNumber="285">Fig. 15a</figureCitation>
), a short funnel, narrowed proximally with a sclerotized and scobinate proximal ventral wall (divided into two halves); sterigma membranous, widened from ostial funnel and slightly concave; ductus bursae evenly narrow and long (ca. 4x as long as width of bursa), with slight sclerotization on caudal 1/10, and with small ventral diverticular sac (internally partially spiculate) by antrum; corpus bursae oblong-ovate; ductus seminalis from junction of ductus bursae with bursa; accessory sac elongate-oblong from short and thin tube at ductus bursae junction; signum (
<figureCitation id="1306C8F1FFCCFF96FAED622D63E3F960" captionStart="FIGURE 15" captionStartId="15.[151,250,1858,1882]" captionTargetBox="[151,1436,453,1835]" captionTargetId="figure-128@15.[151,1436,453,1835]" captionTargetPageId="15" captionText="FIGURE 15. Thaumatographa aurosa taiwana Heppner &amp; Arita, subsp. nov., paratype female (Taiwan: Lien-Hua-Chih For. Sta., Nantou Co.), with details of ostium, sterigma, and antrum diverticulum (a), and signum, (b) (gen. slide JBH-4644 (MGCL)." figureDoi="http://doi.org/10.5281/zenodo.14798853" httpUri="https://zenodo.org/record/14798853/files/figure.png" pageId="14" pageNumber="285">Fig. 15b</figureCitation>
), a typical form for genus but in inverted V-shaped spine array.
</paragraph>
<paragraph id="8B82D474FFCCFF96FF4562956209F86D" blockId="14.[151,1437,361,1972]" pageId="14" pageNumber="285">
Male: unknown in
<collectingCountry id="F32A94E4FFCCFF96FE1F629562D4F905" box="[413,499,1730,1756]" name="Taiwan" pageId="14" pageNumber="285">Taiwan</collectingCountry>
. The male genitalia for the Japanese nominate subspecies are figured by
<bibRefCitation id="EFACA985FFCCFF96FAA8629563C1F8D9" author="Diakonoff, A." pageId="14" pageNumber="285" pagination="1 - 436" refId="ref10974" refString="Diakonoff, A. (1986) Glyphipterigidae auct. sensu lata (sensu Meyrick, 1913). In: Amsel, H. G., Reisser, H. &amp; Gregor, F. (Eds.), Microlepidoptera Palaearctic. Vol. 7. G. Braun, Karlsruhe, pp. 1-436, 175 pls." type="book chapter" year="1986">Diakonoff (1986)</bibRefCitation>
: uncus short and narrow, very evenly recurved from base to blunt apex; hamus stout and somewhat upwardly curved to acute apex, with small basal process; socius nearly subequal to hamus and relatively straight and setose; gnathos membranous; transtilla quadratic, thin; juxta a small concave plate, dorsally blunt; aedeagus moderate, thin, with short decumbent phallobase; vesica scobinate, with narrow cornutus tube; valva oblong, with costal margin somewhat convex with a slight invagination at middle, and termen very truncated; vinculum with saccus undeveloped.
</paragraph>
</subSubSection>
<subSubSection id="C32787FFFFCDFF97FF4564C060A2FF68" box="[199,901,151,177]" pageId="15" pageNumber="286" type="etymology">
<paragraph id="8B82D474FFCDFF97FF4564C060A2FF68" blockId="15.[151,1436,151,393]" box="[199,901,151,177]" pageId="15" pageNumber="286">
<emphasis id="B9490866FFCDFF97FF4564C0626FFF68" bold="true" box="[199,328,151,177]" pageId="15" pageNumber="286">Etymology</emphasis>
. The subspecies is named for the island of
<collectingCountry id="F32A94E4FFCDFF97FCA964C060A6FF68" box="[811,897,151,177]" name="Taiwan" pageId="15" pageNumber="286">Taiwan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C32787FFFFCDFF97FF4564ED67CFFF0C" box="[199,1256,186,213]" pageId="15" pageNumber="286" type="biology_ecology">
<paragraph id="8B82D474FFCDFF97FF4564ED67CFFF0C" blockId="15.[151,1436,151,393]" box="[199,1256,186,213]" pageId="15" pageNumber="286">
<emphasis id="B9490866FFCDFF97FF4564ED6206FF0D" bold="true" box="[199,289,186,212]" pageId="15" pageNumber="286">Biology</emphasis>
. Unknown. Flight period (n = 3 records): mid-September to mid-October (in
<collectingRegion id="49F91A96FFCDFF97FB0764EC67C3FF0C" box="[1157,1252,187,213]" country="China" name="Taiwan)" pageId="15" pageNumber="286">Taiwan)</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C32787FFFFCDFF97FF456489608AFF20" box="[199,941,222,249]" pageId="15" pageNumber="286" type="distribution">
<paragraph id="8B82D474FFCDFF97FF456489608AFF20" blockId="15.[151,1436,151,393]" box="[199,941,222,249]" pageId="15" pageNumber="286">
<emphasis id="B9490866FFCDFF97FF456489627EFF21" bold="true" box="[199,345,222,248]" pageId="15" pageNumber="286">Distribution</emphasis>
. The
<collectingCountry id="F32A94E4FFCDFF97FE1B648862CCFF20" box="[409,491,223,249]" name="Taiwan" pageId="15" pageNumber="286">Taiwan</collectingCountry>
subspecies is known only from
<collectingCountry id="F32A94E4FFCDFF97FCD16488608EFF20" box="[851,937,223,249]" name="Taiwan" pageId="15" pageNumber="286">Taiwan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C32787FFFFCDFF97FF4565556760FE50" pageId="15" pageNumber="286" type="discussion">
<paragraph id="8B82D474FFCDFF97FF4565556760FE50" blockId="15.[151,1436,151,393]" pageId="15" pageNumber="286">
<emphasis id="B9490866FFCDFF97FF4565556262FEC5" bold="true" box="[199,325,258,284]" pageId="15" pageNumber="286">Discussion</emphasis>
. While some maculation distinctions are evident between the two subspecies, further study is needed once males are found of
<taxonomicName id="4C3DAFF7FFCDFF97FE2E65706152FE99" authorityName="Heppner &amp; Arita" authorityYear="2025" box="[428,629,295,320]" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="286" phylum="Arthropoda" rank="subSpecies" species="aurosa" subSpecies="taiwana">
<emphasis id="B9490866FFCDFF97FE2E65706152FE99" box="[428,629,295,320]" italics="true" pageId="15" pageNumber="286">T. aurosa taiwana</emphasis>
</taxonomicName>
to compare the male genitalia of the two subspecies, or to verify if they should be considered distinct species. All available
<collectingCountry id="F32A94E4FFCDFF97FD56651C6001FEBC" box="[724,806,331,357]" name="Taiwan" pageId="15" pageNumber="286">Taiwan</collectingCountry>
specimens are females. The purpose of the small ventral flap (or pouch) external to the base of the ostial funnel near the antrum, is unknown.
</paragraph>
</subSubSection>
</treatment>
</document>