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<mods:title id="7158EE1B910847CF47F9149CA1C984C9">New World Orphanostigma Guenée, 1854, including a new species from Costa Rica (Lepidoptera: Crambidae: Spilomelinae)</mods:title>
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<mods:affiliation id="C1DA425742476B86733A5477E83E9923">Systematic Entomology Laboratory, Beltsville Agriculture Research Center, Agricultural Research Service, U. S. Department of Agriculture, c / o National Museum Natural History, MRC 168, Smithsonian Institution, P. O. Box 37012, Washington, District of Columbia, United States of America, 20013 - 7012.</mods:affiliation>
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<mods:namePart id="A0AD74D6B0AC7B70E27BDA4DD2D51CC5">Cock, Matthew J. W.</mods:namePart>
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<taxonomicName id="4C384D7BFFAAFFF34EFDF8C5FDD8F887" authority="Solis" authorityName="Solis &amp; Cock &amp; Janzen &amp; Hallwachs" authorityYear="2024" box="[151,622,1810,1837]" class="Insecta" family="Crambidae" genus="Orphanostigma" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="385" phylum="Arthropoda" rank="species" species="eugeniephillipsia" status="sp. nov.">
<emphasis id="B94CEAEAFFAAFFF34EFDF8C5FD9DF886" bold="true" box="[151,555,1810,1837]" italics="true" pageId="2" pageNumber="385">Orphanostigma eugeniephillipsia</emphasis>
Solis
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,
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<paragraph id="8B8736F8FFAAFFF34EFDF88CFE26F8DE" blockId="2.[151,1198,1810,1910]" box="[151,400,1883,1910]" pageId="2" pageNumber="385">
(
<figureCitation id="13032A7DFFAAFFF34EF5F88CFEBDF8DE" box="[159,267,1883,1909]" captionStart="FIGURES 17" captionStartId="2.[152,270,1560,1584]" captionTargetBox="[153,1419,306,1511]" captionTargetId="figure-76@2.[153,1434,306,1534]" captionTargetPageId="2" captionText="FIGURES 17. 1, Orphanostigma futilalis, male lectotype, with abdomen before dissection and anal segments pattern visible, San Benito, Texas [USNMENT01433287]. 2, O. futilalis, female paralectotype, with wings spread, Brownsville, Texas [USNMENT01521297]. 3, O. futilalis lectotype labels. 4, O. haemorrhoidalis, male lectotype, Brazil [USNMENT01894048]. 5, O. haemorrhoidalis lectotype labels. 6, Orphanostigma eugeniephillipsia, sp. nov., female holotype, ACG, Costa Rica [USNMENT01894128], forewing length = 5 mm. 7, O. eugeniephillipsia, sp. nov., adult holotype labels." figureDoi="http://doi.org/10.5281/zenodo.12190260" httpUri="https://zenodo.org/record/12190260/files/figure.png" pageId="2" pageNumber="385">Figs. 67</figureCitation>
,
<figureCitation id="13032A7DFFAAFFF34F7DF88CFEE9F8DD" box="[279,351,1883,1910]" captionStart="FIGURES 815" captionStartId="4.[152,270,1773,1797]" captionTargetBox="[237,1359,181,1734]" captionTargetId="figure-20@4.[218,1370,181,1749]" captionTargetPageId="4" captionText="FIGURES 815. 8, Orphanostigma futilalis lectotype male genitalia, San Benito, Texas, USNM slide #116366. 9, O. futilalis phallus, USNM slide #116366. 10, O. haemorrhoidalis male genitalia, ACG, Costa Rica, USNM slide #116402 [USNMENT01894170]. 11, O. haemorrhoidalis phallus, USNM slide #116363. 12, O. abruptalis male genitalia, Sri Lanka, USNM slide #116379 [USNMENT01894037]. 13, O. abruptalis phallus, USNM slide #116379. 14, O. eugeniephillipsia, sp. nov., paratype male genitalia, ACG, Costa Rica, USNM slide #116382 [USNMENT01894117]. 15, O. eugeniephillipsia, sp. nov., paratype phallus, USNM slide #116382. [black arrows indicate the saccular margin and species-specific ornamentation on the valva]" figureDoi="http://doi.org/10.5281/zenodo.12190264" httpUri="https://zenodo.org/record/12190264/files/figure.png" pageId="2" pageNumber="385">14, 15</figureCitation>
,
<figureCitation id="13032A7DFFAAFFF34F01F88CFE3EF8DE" box="[363,392,1883,1909]" captionStart="FIGURES 1619" captionStartId="5.[152,270,1848,1872]" captionTargetBox="[351,1239,195,1813]" captionTargetId="figure-18@5.[331,1257,181,1824]" captionTargetPageId="5" captionText="FIGURES 1619. 16, Orphanostigma futilalis paralectotype female genitalia, Brownsville, Texas, USNM slide #116,365 [USNMENT01521289]. 17, O. eugeniephillipsia, sp. nov., paratype, ACG, Costa Rica, USNM slide #116383 [USNMENT01894118]. 18, O. haemorrhoidalis, ACG, Costa Rica, USNM slide #116403 [USNMENT01894170]. 19, O. abruptalis, Sri Lanka, USNM slide #116380 [USNMENT01894185]. [black arrows indicate projections from anterior section of antrum]" figureDoi="http://doi.org/10.5281/zenodo.12190266" httpUri="https://zenodo.org/record/12190266/files/figure.png" pageId="2" pageNumber="385">17</figureCitation>
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<collectingDate id="EFC2E9D0FFABFFF24D3CFF6CFC7AFF7E" box="[854,972,187,213]" pageId="3" pageNumber="386" value="2011-02-30">30.II.2011</collectingDate>
</date>
, @
<collectingMethod id="52794EEFFFABFFF24D90FF6CFBD5FF7E" box="[1018,1123,187,213]" pageId="3" pageNumber="386">light trap</collectingMethod>
,
<collectorName id="26CD532EFFABFFF24A05FF6CFAA8FF7E" box="[1135,1310,187,213]" pageId="3" pageNumber="386">H. Cambronero</collectorName>
&amp;
<collectorName id="26CD532EFFABFFF24B2EFF6CFA2EFF7E" box="[1348,1432,187,213]" pageId="3" pageNumber="386">S. Rios</collectorName>
, 11-SRNP-104819, USNM slide #116383 [USNMENT01894118]
</materialsCitation>
;
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,
<collectingCountry id="F32F7668FFABFFF24DB5FF08FB3FFF52" box="[991,1161,223,249]" name="Costa Rica" pageId="3" pageNumber="386">COSTA RICA</collectingCountry>
,
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,
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,
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,
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, el.
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,
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<collectingDate id="EFC2E9D0FFABFFF24A1AFED4FB45FEB6" box="[1136,1267,259,285]" pageId="3" pageNumber="386" value="2017-01-29">01/29/2017</collectingDate>
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,
<collectingMethod id="52794EEFFFABFFF24B69FED4FAC7FEB6" box="[1283,1393,259,285]" pageId="3" pageNumber="386">light trap</collectingMethod>
,
<collectorName id="26CD532EFFABFFF24BEBFED4FEA6FEEA" pageId="3" pageNumber="386">A. Guadamuz</collectorName>
&amp;
<collectorName id="26CD532EFFABFFF24F5AFEF0FE17FEEA" box="[304,417,295,321]" pageId="3" pageNumber="386">G. Pereira</collectorName>
, 17-SRNP-100299, USNM slide #116382 [USNMENT01894117]
</materialsCitation>
;
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,
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,
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,
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,
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,
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, el.
<quantity id="4CC09B1DFFABFFF24AD6FE9DFABEFECE" box="[1212,1288,330,357]" metricMagnitude="2" metricUnit="m" metricValue="9.45" pageId="3" pageNumber="386" unit="m" value="945.0">
<elevation id="0015D1CBFFABFFF24AD6FE9DFABEFECE" box="[1212,1288,330,357]" metricMagnitude="2" metricUnit="m" metricValue="9.45" pageId="3" pageNumber="386" unit="m" value="945.0">945 m</elevation>
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,
<date id="FF861038FFABFFF24B7CFE9CFA2EFECE" box="[1302,1432,331,357]" pageId="3" pageNumber="386" value="2017-01-29">
<collectingDate id="EFC2E9D0FFABFFF24B7CFE9CFA2EFECE" box="[1302,1432,331,357]" pageId="3" pageNumber="386" value="2017-01-29">01/29/2017</collectingDate>
</date>
,
<collectingMethod id="52794EEFFFABFFF24EFDFEB8FEB3FE22" box="[151,261,367,393]" pageId="3" pageNumber="386">light trap</collectingMethod>
,
<collectorName id="26CD532EFFABFFF24F7EFEB8FE02FE22" box="[276,436,367,393]" pageId="3" pageNumber="386">A. Guadamuz</collectorName>
&amp;
<collectorName id="26CD532EFFABFFF24F88FEB8FDEAFE22" box="[482,604,367,393]" pageId="3" pageNumber="386">G. Pereira</collectorName>
, 17-SRNP-100298, [USNMENT01894091]. The Barcode Identification Number (BIN) for the holotype and paratypes is BOLD:AAA0296
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3226573FFABFFF74EADFE61FDE0FF52" lastPageId="6" lastPageNumber="389" pageId="3" pageNumber="386" type="diagnosis">
<paragraph id="8B8736F8FFABFFF24EADFE61FC2DFD2E" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="386">
<emphasis id="B94CEAEAFFABFFF24EADFE61FE8CFE7B" bold="true" box="[199,314,438,464]" pageId="3" pageNumber="386">Diagnosis</emphasis>
. Forewing length:
<quantity id="4CC09B1DFFABFFF24C77FE61FDC9FE7A" box="[541,639,438,465]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="3" pageNumber="386" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">56 mm</quantity>
(n=4). Thick band of black and white scales distally on abdominal segments 5-6 and 8
<superScript id="7C4D9BB0FFABFFF24F1EFE0DFE37FE43" attach="left" box="[372,385,474,488]" fontSize="6" pageId="3" pageNumber="386">th</superScript>
segment with lateral dots of black scales (
<figureCitation id="13032A7DFFABFFF24D0FFE0CFC1CFE5E" box="[869,938,475,501]" captionStart="FIGURES 17" captionStartId="2.[152,270,1560,1584]" captionTargetBox="[153,1419,306,1511]" captionTargetId="figure-76@2.[153,1434,306,1534]" captionTargetPageId="2" captionText="FIGURES 17. 1, Orphanostigma futilalis, male lectotype, with abdomen before dissection and anal segments pattern visible, San Benito, Texas [USNMENT01433287]. 2, O. futilalis, female paralectotype, with wings spread, Brownsville, Texas [USNMENT01521297]. 3, O. futilalis lectotype labels. 4, O. haemorrhoidalis, male lectotype, Brazil [USNMENT01894048]. 5, O. haemorrhoidalis lectotype labels. 6, Orphanostigma eugeniephillipsia, sp. nov., female holotype, ACG, Costa Rica [USNMENT01894128], forewing length = 5 mm. 7, O. eugeniephillipsia, sp. nov., adult holotype labels." figureDoi="http://doi.org/10.5281/zenodo.12190260" httpUri="https://zenodo.org/record/12190260/files/figure.png" pageId="3" pageNumber="386">Fig. 6</figureCitation>
). Male genitalia valva saccular margin with a medially, dorsally directed thumb-like protrusion, and another more distal dorsally directed protrusion that is broadly curved and distally toothed and sclerotized, both as ½ as long as valva width (
<figureCitation id="13032A7DFFABFFF24A26FDF4FB28FD96" box="[1100,1182,547,573]" captionStart="FIGURES 815" captionStartId="4.[152,270,1773,1797]" captionTargetBox="[237,1359,181,1734]" captionTargetId="figure-20@4.[218,1370,181,1749]" captionTargetPageId="4" captionText="FIGURES 815. 8, Orphanostigma futilalis lectotype male genitalia, San Benito, Texas, USNM slide #116366. 9, O. futilalis phallus, USNM slide #116366. 10, O. haemorrhoidalis male genitalia, ACG, Costa Rica, USNM slide #116402 [USNMENT01894170]. 11, O. haemorrhoidalis phallus, USNM slide #116363. 12, O. abruptalis male genitalia, Sri Lanka, USNM slide #116379 [USNMENT01894037]. 13, O. abruptalis phallus, USNM slide #116379. 14, O. eugeniephillipsia, sp. nov., paratype male genitalia, ACG, Costa Rica, USNM slide #116382 [USNMENT01894117]. 15, O. eugeniephillipsia, sp. nov., paratype phallus, USNM slide #116382. [black arrows indicate the saccular margin and species-specific ornamentation on the valva]" figureDoi="http://doi.org/10.5281/zenodo.12190264" httpUri="https://zenodo.org/record/12190264/files/figure.png" pageId="3" pageNumber="386">Fig. 14</figureCitation>
); phallus with two flat, sclerotized plates, posterior plate with a sclerotized spine (
<figureCitation id="13032A7DFFABFFF24D47FD90FC37FDCA" box="[813,897,582,609]" captionStart="FIGURES 815" captionStartId="4.[152,270,1773,1797]" captionTargetBox="[237,1359,181,1734]" captionTargetId="figure-20@4.[218,1370,181,1749]" captionTargetPageId="4" captionText="FIGURES 815. 8, Orphanostigma futilalis lectotype male genitalia, San Benito, Texas, USNM slide #116366. 9, O. futilalis phallus, USNM slide #116366. 10, O. haemorrhoidalis male genitalia, ACG, Costa Rica, USNM slide #116402 [USNMENT01894170]. 11, O. haemorrhoidalis phallus, USNM slide #116363. 12, O. abruptalis male genitalia, Sri Lanka, USNM slide #116379 [USNMENT01894037]. 13, O. abruptalis phallus, USNM slide #116379. 14, O. eugeniephillipsia, sp. nov., paratype male genitalia, ACG, Costa Rica, USNM slide #116382 [USNMENT01894117]. 15, O. eugeniephillipsia, sp. nov., paratype phallus, USNM slide #116382. [black arrows indicate the saccular margin and species-specific ornamentation on the valva]" figureDoi="http://doi.org/10.5281/zenodo.12190264" httpUri="https://zenodo.org/record/12190264/files/figure.png" pageId="3" pageNumber="386">Fig. 15</figureCitation>
). Female genitalia with antrum sclerotized with two distal arm-like, lateral projections, bending posteriorly (
<figureCitation id="13032A7DFFABFFF24D50FDBCFC3BFD2E" box="[826,909,619,645]" captionStart="FIGURES 1619" captionStartId="5.[152,270,1848,1872]" captionTargetBox="[351,1239,195,1813]" captionTargetId="figure-18@5.[331,1257,181,1824]" captionTargetPageId="5" captionText="FIGURES 1619. 16, Orphanostigma futilalis paralectotype female genitalia, Brownsville, Texas, USNM slide #116,365 [USNMENT01521289]. 17, O. eugeniephillipsia, sp. nov., paratype, ACG, Costa Rica, USNM slide #116383 [USNMENT01894118]. 18, O. haemorrhoidalis, ACG, Costa Rica, USNM slide #116403 [USNMENT01894170]. 19, O. abruptalis, Sri Lanka, USNM slide #116380 [USNMENT01894185]. [black arrows indicate projections from anterior section of antrum]" figureDoi="http://doi.org/10.5281/zenodo.12190266" httpUri="https://zenodo.org/record/12190266/files/figure.png" pageId="3" pageNumber="386">Fig. 17</figureCitation>
).
</paragraph>
<paragraph id="8B8736F8FFABFFF24EADFD59FAB7FB6E" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="386">
<emphasis id="B94CEAEAFFABFFF24EADFD59FDA4FD03" bold="true" box="[199,530,654,680]" pageId="3" pageNumber="386">Morphological comparison</emphasis>
. Although the best way to confirm identification of these species is with internal genitalia characters, we provide a few external characters of the head and abdomen. The antennae of
<taxonomicName id="4C384D7BFFABFFF24BEBFD64FEE7FD5A" class="Insecta" family="Crambidae" genus="Orphanostigma" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="386" phylum="Arthropoda" rank="species" species="haemorrhoidalis">
<emphasis id="B94CEAEAFFABFFF24BEBFD64FEE7FD5A" italics="true" pageId="3" pageNumber="386">O. haemorrhoidalis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C384D7BFFABFFF24FE4FD00FDC2FD5A" authorityName="Solis &amp; Cock &amp; Janzen &amp; Hallwachs" authorityYear="2024" box="[398,628,727,753]" class="Insecta" family="Crambidae" genus="Orphanostigma" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="386" phylum="Arthropoda" rank="species" species="eugeniephillipsia">
<emphasis id="B94CEAEAFFABFFF24FE4FD00FDC2FD5A" box="[398,628,727,753]" italics="true" pageId="3" pageNumber="386">O. eugeniephillipsia</emphasis>
</taxonomicName>
have an anterior brown line extending onto the shaft, and
<taxonomicName id="4C384D7BFFABFFF24B4BFD00FA2BFD5A" authorityName="Solis &amp; Cock &amp; Janzen &amp; Hallwachs" authorityYear="2024" baseAuthorityName="Barnes &amp; McDunnough" baseAuthorityYear="1914" box="[1313,1437,727,753]" class="Insecta" family="Crambidae" genus="Orphanostigma" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="386" phylum="Arthropoda" rank="species" species="futilalis">
<emphasis id="B94CEAEAFFABFFF24B4BFD00FA2BFD5A" box="[1313,1437,727,753]" italics="true" pageId="3" pageNumber="386">O. futilalis</emphasis>
</taxonomicName>
lacks these lines.
<taxonomicName id="4C384D7BFFABFFF24F33FD2CFD66FCBE" authorityName="Solis &amp; Cock &amp; Janzen &amp; Hallwachs" authorityYear="2024" box="[345,720,763,789]" class="Insecta" family="Crambidae" genus="Orphanostigma" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="386" phylum="Arthropoda" rank="species" species="eugeniephillipsia">
<emphasis id="B94CEAEAFFABFFF24F33FD2CFD66FCBE" box="[345,720,763,789]" italics="true" pageId="3" pageNumber="386">Orphanostigma eugeniephillipsia</emphasis>
</taxonomicName>
has a thick band of black and white scales distally on abdominal segments 56 and lateral dots with black scales,
<taxonomicName id="4C384D7BFFABFFF24CDFFCC8FC27FC92" box="[693,913,799,825]" class="Insecta" family="Crambidae" genus="Orphanostigma" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="386" phylum="Arthropoda" rank="species" species="haemorrhoidalis">
<emphasis id="B94CEAEAFFABFFF24CDFFCC8FC27FC92" box="[693,913,799,825]" italics="true" pageId="3" pageNumber="386">O. haemorrhoidalis</emphasis>
</taxonomicName>
has a few black scales on abdominal segments 56 and sometimes with a few black scales laterally, and
<taxonomicName id="4C384D7BFFABFFF24D70FC94FC24FCF6" authorityName="Solis &amp; Cock &amp; Janzen &amp; Hallwachs" authorityYear="2024" baseAuthorityName="Barnes &amp; McDunnough" baseAuthorityYear="1914" box="[794,914,835,861]" class="Insecta" family="Crambidae" genus="Orphanostigma" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="386" phylum="Arthropoda" rank="species" species="futilalis">
<emphasis id="B94CEAEAFFABFFF24D70FC94FC24FCF6" box="[794,914,835,861]" italics="true" pageId="3" pageNumber="386">O. futilalis</emphasis>
</taxonomicName>
has no black scales on these segments. On the dorsum of the 8
<superScript id="7C4D9BB0FFABFFF24F26FCB1FEEFFCDF" attach="left" box="[332,345,870,884]" fontSize="6" pageId="3" pageNumber="386">th</superScript>
abdominal segment of
<emphasis id="B94CEAEAFFABFFF24C0FFCB0FC7AFC2A" box="[613,972,871,897]" italics="true" pageId="3" pageNumber="386">
<taxonomicName id="4C384D7BFFABFFF24C0FFCB0FCF0FC2A" box="[613,838,871,897]" class="Insecta" family="Crambidae" genus="Orphanostigma" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="386" phylum="Arthropoda" rank="species" species="haemorrhoidalis">O. haemorrhoidalis</taxonomicName>
,
<taxonomicName id="4C384D7BFFABFFF24D39FCB0FC7AFC2A" authorityName="Solis &amp; Cock &amp; Janzen &amp; Hallwachs" authorityYear="2024" baseAuthorityName="Barnes &amp; McDunnough" baseAuthorityYear="1914" box="[851,972,871,897]" class="Insecta" family="Crambidae" genus="Orphanostigma" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="386" phylum="Arthropoda" rank="species" species="futilalis">O. futilalis</taxonomicName>
</emphasis>
, and
<taxonomicName id="4C384D7BFFABFFF24A66FCB0FB47FC2A" authorityName="Solis &amp; Cock &amp; Janzen &amp; Hallwachs" authorityYear="2024" box="[1036,1265,871,897]" class="Insecta" family="Crambidae" genus="Orphanostigma" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="386" phylum="Arthropoda" rank="species" species="eugeniephillipsia">
<emphasis id="B94CEAEAFFABFFF24A66FCB0FB47FC2A" box="[1036,1265,871,897]" italics="true" pageId="3" pageNumber="386">O. eugeniephillipsia</emphasis>
</taxonomicName>
the segment is mainly orange yellow in color, but distally it has a white u-shaped line and within it are dark brown scales that can be seen prominently in the
<typeStatus id="5483885AFFABFFF24FA1FC78FD84FC62" box="[459,562,943,969]" pageId="3" pageNumber="386" type="lectotype">lectotype</typeStatus>
(
<figureCitation id="13032A7DFFABFFF24C28FC78FD31FC62" box="[578,647,943,969]" captionStart="FIGURES 17" captionStartId="2.[152,270,1560,1584]" captionTargetBox="[153,1419,306,1511]" captionTargetId="figure-76@2.[153,1434,306,1534]" captionTargetPageId="2" captionText="FIGURES 17. 1, Orphanostigma futilalis, male lectotype, with abdomen before dissection and anal segments pattern visible, San Benito, Texas [USNMENT01433287]. 2, O. futilalis, female paralectotype, with wings spread, Brownsville, Texas [USNMENT01521297]. 3, O. futilalis lectotype labels. 4, O. haemorrhoidalis, male lectotype, Brazil [USNMENT01894048]. 5, O. haemorrhoidalis lectotype labels. 6, Orphanostigma eugeniephillipsia, sp. nov., female holotype, ACG, Costa Rica [USNMENT01894128], forewing length = 5 mm. 7, O. eugeniephillipsia, sp. nov., adult holotype labels." figureDoi="http://doi.org/10.5281/zenodo.12190260" httpUri="https://zenodo.org/record/12190260/files/figure.png" pageId="3" pageNumber="386">Fig. 4</figureCitation>
), but this brown may appear very dark or just as a smudge depending on the condition of the specimen. Wing coloration is not included in the diagnostic characters because, although the specimens of
<taxonomicName id="4C384D7BFFABFFF24F45FC20FDA7FBBA" authorityName="Solis &amp; Cock &amp; Janzen &amp; Hallwachs" authorityYear="2024" box="[303,529,1015,1041]" class="Insecta" family="Crambidae" genus="Orphanostigma" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="386" phylum="Arthropoda" rank="species" species="eugeniephillipsia">
<emphasis id="B94CEAEAFFABFFF24F45FC20FDA7FBBA" box="[303,529,1015,1041]" italics="true" pageId="3" pageNumber="386">O. eugeniephillipsia</emphasis>
</taxonomicName>
are dark yellow orange beyond the postmedial line in both wings and only yellow in
<taxonomicName id="4C384D7BFFABFFF24ED2FBCCFE82FB9E" authorityName="Solis &amp; Cock &amp; Janzen &amp; Hallwachs" authorityYear="2024" baseAuthorityName="Barnes &amp; McDunnough" baseAuthorityYear="1914" box="[184,308,1051,1077]" class="Insecta" family="Crambidae" genus="Orphanostigma" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="386" phylum="Arthropoda" rank="species" species="futilalis">
<emphasis id="B94CEAEAFFABFFF24ED2FBCCFE82FB9E" box="[184,308,1051,1077]" italics="true" pageId="3" pageNumber="386">O. futilalis</emphasis>
</taxonomicName>
, there is wide variation in
<taxonomicName id="4C384D7BFFABFFF24C1CFBCCFCE0FB9E" box="[630,854,1051,1077]" class="Insecta" family="Crambidae" genus="Orphanostigma" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="386" phylum="Arthropoda" rank="species" species="haemorrhoidalis">
<emphasis id="B94CEAEAFFABFFF24C1CFBCCFCE0FB9E" box="[630,854,1051,1077]" italics="true" pageId="3" pageNumber="386">O. haemorrhoidalis</emphasis>
</taxonomicName>
(that
<taxonomicName id="4C384D7BFFABFFF24DF5FBCCFB30FB9E" authorityName="Solis &amp; Cock &amp; Janzen &amp; Hallwachs" authorityYear="2024" box="[927,1158,1051,1077]" class="Insecta" family="Crambidae" genus="Orphanostigma" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="386" phylum="Arthropoda" rank="species" species="eugeniephillipsia">
<emphasis id="B94CEAEAFFABFFF24DF5FBCCFB30FB9E" box="[927,1158,1051,1077]" italics="true" pageId="3" pageNumber="386">O. eugeniephillipsia</emphasis>
</taxonomicName>
most closely resembles externally) from plain yellow to dark or “deep orange yellow” in this area of the forewing (as in
<taxonomicName id="4C384D7BFFABFFF24ABAFBE8FAD0FBF2" baseAuthorityName="Walker" baseAuthorityYear="1859" box="[1232,1382,1087,1113]" class="Insecta" family="Crambidae" genus="Orphanostigma" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="386" phylum="Arthropoda" rank="species" species="abruptalis">
<emphasis id="B94CEAEAFFABFFF24ABAFBE8FAD0FBF2" box="[1232,1382,1087,1113]" italics="true" pageId="3" pageNumber="386">O. abruptalis</emphasis>
</taxonomicName>
, and as described in
<bibRefCitation id="EFA94B09FFABFFF24F2FFBB4FDEEFBD6" author="Shaffer, J. C. &amp; Munroe, E. G." box="[325,600,1123,1149]" pageId="3" pageNumber="386" pagination="1 - 110" refId="ref4975" refString="Shaffer, J. C. &amp; Munroe, E. G. (2007) Crambidae of Aldabra Atoll (Lepidoptera: Pyraloidea). Tropical Lepidoptera, 14 (1 - 2), 1 - 110." type="journal article" year="2007">Shaffer &amp; Munroe 2007</bibRefCitation>
). There are only
<specimenCount id="9D3EFD71FFABFFF24D7FFBB4FC29FBD6" box="[789,927,1123,1149]" count="4" pageId="3" pageNumber="386" type="generic">4 specimens</specimenCount>
in the type series of
<taxonomicName id="4C384D7BFFABFFF24AE3FBB4FADBFBD6" authorityName="Solis &amp; Cock &amp; Janzen &amp; Hallwachs" authorityYear="2024" box="[1161,1389,1123,1149]" class="Insecta" family="Crambidae" genus="Orphanostigma" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="386" phylum="Arthropoda" rank="species" species="eugeniephillipsia">
<emphasis id="B94CEAEAFFABFFF24AE3FBB4FADBFBD6" box="[1161,1389,1123,1149]" italics="true" pageId="3" pageNumber="386">O. eugeniephillipsia</emphasis>
</taxonomicName>
and only
<specimenCount id="9D3EFD71FFABFFF24EBAFB50FE32FB0A" box="[208,388,1159,1185]" count="3" pageId="3" pageNumber="386" type="generic">three specimens</specimenCount>
were found of the type series of
<taxonomicName id="4C384D7BFFABFFF24C9CFB50FCD8FB0A" authorityName="Solis &amp; Cock &amp; Janzen &amp; Hallwachs" authorityYear="2024" baseAuthorityName="Barnes &amp; McDunnough" baseAuthorityYear="1914" box="[758,878,1159,1185]" class="Insecta" family="Crambidae" genus="Orphanostigma" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="386" phylum="Arthropoda" rank="species" species="futilalis">
<emphasis id="B94CEAEAFFABFFF24C9CFB50FCD8FB0A" box="[758,878,1159,1185]" italics="true" pageId="3" pageNumber="386">O. futilalis</emphasis>
</taxonomicName>
(
<specimenCount id="9D3EFD71FFABFFF24D17FB50FBBEFB0A" box="[893,1032,1159,1185]" count="2" pageId="3" pageNumber="386" type="generic">2 specimens</specimenCount>
were not found), so we recommend caution on identifications based only on wing coloration and recommend dissections of the genitalia.
</paragraph>
<paragraph id="8B8736F8FFABFFF24EADFB18FC65F916" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="386">
Although the male genitalia of all three New World species have an elongated, sclerotized line from a broad transtilla just below the length of the costal or dorsal margin of the valvae, the armature of the saccular margin of the valva is distinctive in each of the three species (
<figureCitation id="13032A7DFFABFFF24CBCFAC0FCCFFA9A" box="[726,889,1303,1329]" captionStart="FIGURES 815" captionStartId="4.[152,270,1773,1797]" captionTargetBox="[237,1359,181,1734]" captionTargetId="figure-20@4.[218,1370,181,1749]" captionTargetPageId="4" captionText="FIGURES 815. 8, Orphanostigma futilalis lectotype male genitalia, San Benito, Texas, USNM slide #116366. 9, O. futilalis phallus, USNM slide #116366. 10, O. haemorrhoidalis male genitalia, ACG, Costa Rica, USNM slide #116402 [USNMENT01894170]. 11, O. haemorrhoidalis phallus, USNM slide #116363. 12, O. abruptalis male genitalia, Sri Lanka, USNM slide #116379 [USNMENT01894037]. 13, O. abruptalis phallus, USNM slide #116379. 14, O. eugeniephillipsia, sp. nov., paratype male genitalia, ACG, Costa Rica, USNM slide #116382 [USNMENT01894117]. 15, O. eugeniephillipsia, sp. nov., paratype phallus, USNM slide #116382. [black arrows indicate the saccular margin and species-specific ornamentation on the valva]" figureDoi="http://doi.org/10.5281/zenodo.12190264" httpUri="https://zenodo.org/record/12190264/files/figure.png" pageId="3" pageNumber="386">Figs. 8, 10, 14</figureCitation>
). In
<taxonomicName id="4C384D7BFFABFFF24DC6FAC0FB39FA9A" authorityName="Solis &amp; Cock &amp; Janzen &amp; Hallwachs" authorityYear="2024" box="[940,1167,1303,1329]" class="Insecta" family="Crambidae" genus="Orphanostigma" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="386" phylum="Arthropoda" rank="species" species="eugeniephillipsia">
<emphasis id="B94CEAEAFFABFFF24DC6FAC0FB39FA9A" box="[940,1167,1303,1329]" italics="true" pageId="3" pageNumber="386">O. eugeniephillipsia</emphasis>
</taxonomicName>
the saccular margin has two dorsally directed protrusions, medially a thumb-like protrusion and distally a broadly curved, distally toothed and sclerotized protrusion (
<figureCitation id="13032A7DFFABFFF24FA3FA88FDAAFAD2" box="[457,540,1375,1401]" captionStart="FIGURES 815" captionStartId="4.[152,270,1773,1797]" captionTargetBox="[237,1359,181,1734]" captionTargetId="figure-20@4.[218,1370,181,1749]" captionTargetPageId="4" captionText="FIGURES 815. 8, Orphanostigma futilalis lectotype male genitalia, San Benito, Texas, USNM slide #116366. 9, O. futilalis phallus, USNM slide #116366. 10, O. haemorrhoidalis male genitalia, ACG, Costa Rica, USNM slide #116402 [USNMENT01894170]. 11, O. haemorrhoidalis phallus, USNM slide #116363. 12, O. abruptalis male genitalia, Sri Lanka, USNM slide #116379 [USNMENT01894037]. 13, O. abruptalis phallus, USNM slide #116379. 14, O. eugeniephillipsia, sp. nov., paratype male genitalia, ACG, Costa Rica, USNM slide #116382 [USNMENT01894117]. 15, O. eugeniephillipsia, sp. nov., paratype phallus, USNM slide #116382. [black arrows indicate the saccular margin and species-specific ornamentation on the valva]" figureDoi="http://doi.org/10.5281/zenodo.12190264" httpUri="https://zenodo.org/record/12190264/files/figure.png" pageId="3" pageNumber="386">Fig. 14</figureCitation>
), in
<taxonomicName id="4C384D7BFFABFFF24C24FA88FC9CFAD2" box="[590,810,1375,1401]" class="Insecta" family="Crambidae" genus="Orphanostigma" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="386" phylum="Arthropoda" rank="species" species="haemorrhoidalis">
<emphasis id="B94CEAEAFFABFFF24C24FA88FC9CFAD2" box="[590,810,1375,1401]" italics="true" pageId="3" pageNumber="386">O. haemorrhoidalis</emphasis>
</taxonomicName>
the saccular margin medially has a knife-like elongated ridge, basally with a variously-shaped dorsally directed triangle, i.e., pointed, 1/3 the width of the valva (the shape of the triangle sides can vary and can be difficult to see with some slide preparations) (
<figureCitation id="13032A7DFFABFFF24A07FA70FB76FA6A" box="[1133,1216,1447,1473]" captionStart="FIGURES 815" captionStartId="4.[152,270,1773,1797]" captionTargetBox="[237,1359,181,1734]" captionTargetId="figure-20@4.[218,1370,181,1749]" captionTargetPageId="4" captionText="FIGURES 815. 8, Orphanostigma futilalis lectotype male genitalia, San Benito, Texas, USNM slide #116366. 9, O. futilalis phallus, USNM slide #116366. 10, O. haemorrhoidalis male genitalia, ACG, Costa Rica, USNM slide #116402 [USNMENT01894170]. 11, O. haemorrhoidalis phallus, USNM slide #116363. 12, O. abruptalis male genitalia, Sri Lanka, USNM slide #116379 [USNMENT01894037]. 13, O. abruptalis phallus, USNM slide #116379. 14, O. eugeniephillipsia, sp. nov., paratype male genitalia, ACG, Costa Rica, USNM slide #116382 [USNMENT01894117]. 15, O. eugeniephillipsia, sp. nov., paratype phallus, USNM slide #116382. [black arrows indicate the saccular margin and species-specific ornamentation on the valva]" figureDoi="http://doi.org/10.5281/zenodo.12190264" httpUri="https://zenodo.org/record/12190264/files/figure.png" pageId="3" pageNumber="386">Fig. 10</figureCitation>
), and in
<taxonomicName id="4C384D7BFFABFFF24B4EFA70FA2BFA6A" authorityName="Solis &amp; Cock &amp; Janzen &amp; Hallwachs" authorityYear="2024" baseAuthorityName="Barnes &amp; McDunnough" baseAuthorityYear="1914" box="[1316,1437,1447,1473]" class="Insecta" family="Crambidae" genus="Orphanostigma" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="386" phylum="Arthropoda" rank="species" species="futilalis">
<emphasis id="B94CEAEAFFABFFF24B4EFA70FA2BFA6A" box="[1316,1437,1447,1473]" italics="true" pageId="3" pageNumber="386">O. futilalis</emphasis>
</taxonomicName>
there is a basal, sclerotized ridge near the saccular margin with a ventrally directed, elongate protrusion (almost as long as the dorsal sclerotized ridge), with a square-shaped apex (
<figureCitation id="13032A7DFFABFFF24DF2FA38FC68F9A2" box="[920,990,1519,1545]" captionStart="FIGURES 815" captionStartId="4.[152,270,1773,1797]" captionTargetBox="[237,1359,181,1734]" captionTargetId="figure-20@4.[218,1370,181,1749]" captionTargetPageId="4" captionText="FIGURES 815. 8, Orphanostigma futilalis lectotype male genitalia, San Benito, Texas, USNM slide #116366. 9, O. futilalis phallus, USNM slide #116366. 10, O. haemorrhoidalis male genitalia, ACG, Costa Rica, USNM slide #116402 [USNMENT01894170]. 11, O. haemorrhoidalis phallus, USNM slide #116363. 12, O. abruptalis male genitalia, Sri Lanka, USNM slide #116379 [USNMENT01894037]. 13, O. abruptalis phallus, USNM slide #116379. 14, O. eugeniephillipsia, sp. nov., paratype male genitalia, ACG, Costa Rica, USNM slide #116382 [USNMENT01894117]. 15, O. eugeniephillipsia, sp. nov., paratype phallus, USNM slide #116382. [black arrows indicate the saccular margin and species-specific ornamentation on the valva]" figureDoi="http://doi.org/10.5281/zenodo.12190264" httpUri="https://zenodo.org/record/12190264/files/figure.png" pageId="3" pageNumber="386">Fig. 8</figureCitation>
). The saccular margin of all three New World species varies from that of
<taxonomicName id="4C384D7BFFABFFF24C7FF9C4FD1DF986" baseAuthorityName="Walker" baseAuthorityYear="1859" box="[533,683,1555,1581]" class="Insecta" family="Crambidae" genus="Orphanostigma" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="386" phylum="Arthropoda" rank="species" species="abruptalis">
<emphasis id="B94CEAEAFFABFFF24C7FF9C4FD1DF986" box="[533,683,1555,1581]" italics="true" pageId="3" pageNumber="386">O. abruptalis</emphasis>
</taxonomicName>
as shown in
<figureCitation id="13032A7DFFABFFF24D55F9C4FC24F986" box="[831,914,1555,1581]" captionStart="FIGURES 815" captionStartId="4.[152,270,1773,1797]" captionTargetBox="[237,1359,181,1734]" captionTargetId="figure-20@4.[218,1370,181,1749]" captionTargetPageId="4" captionText="FIGURES 815. 8, Orphanostigma futilalis lectotype male genitalia, San Benito, Texas, USNM slide #116366. 9, O. futilalis phallus, USNM slide #116366. 10, O. haemorrhoidalis male genitalia, ACG, Costa Rica, USNM slide #116402 [USNMENT01894170]. 11, O. haemorrhoidalis phallus, USNM slide #116363. 12, O. abruptalis male genitalia, Sri Lanka, USNM slide #116379 [USNMENT01894037]. 13, O. abruptalis phallus, USNM slide #116379. 14, O. eugeniephillipsia, sp. nov., paratype male genitalia, ACG, Costa Rica, USNM slide #116382 [USNMENT01894117]. 15, O. eugeniephillipsia, sp. nov., paratype phallus, USNM slide #116382. [black arrows indicate the saccular margin and species-specific ornamentation on the valva]" figureDoi="http://doi.org/10.5281/zenodo.12190264" httpUri="https://zenodo.org/record/12190264/files/figure.png" pageId="3" pageNumber="386">Fig. 12</figureCitation>
. The cornutus of the phallus is also distinctive between the three species:
<taxonomicName id="4C384D7BFFABFFF24FA8F9E0FD13F9FA" authorityName="Solis &amp; Cock &amp; Janzen &amp; Hallwachs" authorityYear="2024" box="[450,677,1591,1617]" class="Insecta" family="Crambidae" genus="Orphanostigma" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="386" phylum="Arthropoda" rank="species" species="eugeniephillipsia">
<emphasis id="B94CEAEAFFABFFF24FA8F9E0FD13F9FA" box="[450,677,1591,1617]" italics="true" pageId="3" pageNumber="386">O. eugeniephillipsia</emphasis>
</taxonomicName>
has a posterior sclerotized spine on a flat plate (
<figureCitation id="13032A7DFFABFFF24AD4F9E0FAA6F9FA" box="[1214,1296,1590,1617]" captionStart="FIGURES 815" captionStartId="4.[152,270,1773,1797]" captionTargetBox="[237,1359,181,1734]" captionTargetId="figure-20@4.[218,1370,181,1749]" captionTargetPageId="4" captionText="FIGURES 815. 8, Orphanostigma futilalis lectotype male genitalia, San Benito, Texas, USNM slide #116366. 9, O. futilalis phallus, USNM slide #116366. 10, O. haemorrhoidalis male genitalia, ACG, Costa Rica, USNM slide #116402 [USNMENT01894170]. 11, O. haemorrhoidalis phallus, USNM slide #116363. 12, O. abruptalis male genitalia, Sri Lanka, USNM slide #116379 [USNMENT01894037]. 13, O. abruptalis phallus, USNM slide #116379. 14, O. eugeniephillipsia, sp. nov., paratype male genitalia, ACG, Costa Rica, USNM slide #116382 [USNMENT01894117]. 15, O. eugeniephillipsia, sp. nov., paratype phallus, USNM slide #116382. [black arrows indicate the saccular margin and species-specific ornamentation on the valva]" figureDoi="http://doi.org/10.5281/zenodo.12190264" httpUri="https://zenodo.org/record/12190264/files/figure.png" pageId="3" pageNumber="386">Fig. 15</figureCitation>
),
<taxonomicName id="4C384D7BFFABFFF24B4FF9E0FA2BF9FA" authorityName="Solis &amp; Cock &amp; Janzen &amp; Hallwachs" authorityYear="2024" baseAuthorityName="Barnes &amp; McDunnough" baseAuthorityYear="1914" box="[1317,1437,1591,1617]" class="Insecta" family="Crambidae" genus="Orphanostigma" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="386" phylum="Arthropoda" rank="species" species="futilalis">
<emphasis id="B94CEAEAFFABFFF24B4FF9E0FA2BF9FA" box="[1317,1437,1591,1617]" italics="true" pageId="3" pageNumber="386">O. futilalis</emphasis>
</taxonomicName>
has a posterior broad, toothed, and spoon-like shaped sclerotization (
<figureCitation id="13032A7DFFABFFF24DE4F98CFC64F9DE" box="[910,978,1627,1653]" captionStart="FIGURES 815" captionStartId="4.[152,270,1773,1797]" captionTargetBox="[237,1359,181,1734]" captionTargetId="figure-20@4.[218,1370,181,1749]" captionTargetPageId="4" captionText="FIGURES 815. 8, Orphanostigma futilalis lectotype male genitalia, San Benito, Texas, USNM slide #116366. 9, O. futilalis phallus, USNM slide #116366. 10, O. haemorrhoidalis male genitalia, ACG, Costa Rica, USNM slide #116402 [USNMENT01894170]. 11, O. haemorrhoidalis phallus, USNM slide #116363. 12, O. abruptalis male genitalia, Sri Lanka, USNM slide #116379 [USNMENT01894037]. 13, O. abruptalis phallus, USNM slide #116379. 14, O. eugeniephillipsia, sp. nov., paratype male genitalia, ACG, Costa Rica, USNM slide #116382 [USNMENT01894117]. 15, O. eugeniephillipsia, sp. nov., paratype phallus, USNM slide #116382. [black arrows indicate the saccular margin and species-specific ornamentation on the valva]" figureDoi="http://doi.org/10.5281/zenodo.12190264" httpUri="https://zenodo.org/record/12190264/files/figure.png" pageId="3" pageNumber="386">Fig. 9</figureCitation>
), and
<taxonomicName id="4C384D7BFFABFFF24A7EF98CFB59F9DE" box="[1044,1263,1627,1653]" class="Insecta" family="Crambidae" genus="Orphanostigma" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="386" phylum="Arthropoda" rank="species" species="haemorrhoidalis">
<emphasis id="B94CEAEAFFABFFF24A7EF98CFB59F9DE" box="[1044,1263,1627,1653]" italics="true" pageId="3" pageNumber="386">O. haemorrhoidalis</emphasis>
</taxonomicName>
has a posterior, sclerotized broad, curved, toothed band (
<figureCitation id="13032A7DFFABFFF24C3CF9A8FD13F932" box="[598,677,1663,1689]" captionStart="FIGURES 815" captionStartId="4.[152,270,1773,1797]" captionTargetBox="[237,1359,181,1734]" captionTargetId="figure-20@4.[218,1370,181,1749]" captionTargetPageId="4" captionText="FIGURES 815. 8, Orphanostigma futilalis lectotype male genitalia, San Benito, Texas, USNM slide #116366. 9, O. futilalis phallus, USNM slide #116366. 10, O. haemorrhoidalis male genitalia, ACG, Costa Rica, USNM slide #116402 [USNMENT01894170]. 11, O. haemorrhoidalis phallus, USNM slide #116363. 12, O. abruptalis male genitalia, Sri Lanka, USNM slide #116379 [USNMENT01894037]. 13, O. abruptalis phallus, USNM slide #116379. 14, O. eugeniephillipsia, sp. nov., paratype male genitalia, ACG, Costa Rica, USNM slide #116382 [USNMENT01894117]. 15, O. eugeniephillipsia, sp. nov., paratype phallus, USNM slide #116382. [black arrows indicate the saccular margin and species-specific ornamentation on the valva]" figureDoi="http://doi.org/10.5281/zenodo.12190264" httpUri="https://zenodo.org/record/12190264/files/figure.png" pageId="3" pageNumber="386">Fig. 11</figureCitation>
). The phallus of New World species differs from that of
<taxonomicName id="4C384D7BFFABFFF24B63F9A8FA2AF932" baseAuthorityName="Walker" baseAuthorityYear="1859" box="[1289,1436,1663,1689]" class="Insecta" family="Crambidae" genus="Orphanostigma" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="386" phylum="Arthropoda" rank="species" species="abruptalis">
<emphasis id="B94CEAEAFFABFFF24B63F9A8FA2AF932" box="[1289,1436,1663,1689]" italics="true" pageId="3" pageNumber="386">O. abruptalis</emphasis>
</taxonomicName>
that has “numerous minute denticles” (
<figureCitation id="13032A7DFFABFFF24C20F974FD2DF916" box="[586,667,1699,1725]" captionStart="FIGURES 815" captionStartId="4.[152,270,1773,1797]" captionTargetBox="[237,1359,181,1734]" captionTargetId="figure-20@4.[218,1370,181,1749]" captionTargetPageId="4" captionText="FIGURES 815. 8, Orphanostigma futilalis lectotype male genitalia, San Benito, Texas, USNM slide #116366. 9, O. futilalis phallus, USNM slide #116366. 10, O. haemorrhoidalis male genitalia, ACG, Costa Rica, USNM slide #116402 [USNMENT01894170]. 11, O. haemorrhoidalis phallus, USNM slide #116363. 12, O. abruptalis male genitalia, Sri Lanka, USNM slide #116379 [USNMENT01894037]. 13, O. abruptalis phallus, USNM slide #116379. 14, O. eugeniephillipsia, sp. nov., paratype male genitalia, ACG, Costa Rica, USNM slide #116382 [USNMENT01894117]. 15, O. eugeniephillipsia, sp. nov., paratype phallus, USNM slide #116382. [black arrows indicate the saccular margin and species-specific ornamentation on the valva]" figureDoi="http://doi.org/10.5281/zenodo.12190264" httpUri="https://zenodo.org/record/12190264/files/figure.png" pageId="3" pageNumber="386">Fig. 13</figureCitation>
) (
<bibRefCitation id="EFA94B09FFABFFF24CD8F974FC71F916" author="Shaffer, J. C. &amp; Munroe, E. G." box="[690,967,1699,1725]" pageId="3" pageNumber="386" pagination="1 - 110" refId="ref4975" refString="Shaffer, J. C. &amp; Munroe, E. G. (2007) Crambidae of Aldabra Atoll (Lepidoptera: Pyraloidea). Tropical Lepidoptera, 14 (1 - 2), 1 - 110." type="journal article" year="2007">Shaffer &amp; Munroe 2007</bibRefCitation>
).
</paragraph>
<paragraph id="8B8736F8FFABFFF74EADF910FDE0FF52" blockId="3.[151,1437,151,2013]" lastBlockId="6.[151,1437,151,609]" lastPageId="6" lastPageNumber="389" pageId="3" pageNumber="386">
In the female genitalia, all three species have a broad ostium bursae, almost the width of the segment, the antrum is sclerotized and variously shaped, half as long as the corpus bursae, corpus bursae with very short spines, the corpus bursae posterior is 1/3 less broad than the anterior 2/3
<superScript id="7C4D9BB0FFABFFF24DE7F8D9FC2DF8B7" attach="left" box="[909,923,1806,1820]" fontSize="6" pageId="3" pageNumber="386">rd</superScript>
that is broadly rounded, with a small round indentation from the corpus bursae wall or signum, and a ductus seminalis from the flat, square-like posterior edge of corpus bursae. The antrum consists of anterior and posterior sections, the shape is distinctive in the three species:
<taxonomicName id="4C384D7BFFABFFF24E9DF8ACFE6CF83E" authorityName="Solis &amp; Cock &amp; Janzen &amp; Hallwachs" authorityYear="2024" box="[247,474,1915,1941]" class="Insecta" family="Crambidae" genus="Orphanostigma" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="386" phylum="Arthropoda" rank="species" species="eugeniephillipsia">
<emphasis id="B94CEAEAFFABFFF24E9DF8ACFE6CF83E" box="[247,474,1915,1941]" italics="true" pageId="3" pageNumber="386">O. eugeniephillipsia</emphasis>
</taxonomicName>
has two arm-like, lateral projections bending posteriorly originating from the anterior section (
<figureCitation id="13032A7DFFABFFF24E93F848FEF8F812" box="[249,334,1951,1977]" captionStart="FIGURES 1619" captionStartId="5.[152,270,1848,1872]" captionTargetBox="[351,1239,195,1813]" captionTargetId="figure-18@5.[331,1257,181,1824]" captionTargetPageId="5" captionText="FIGURES 1619. 16, Orphanostigma futilalis paralectotype female genitalia, Brownsville, Texas, USNM slide #116,365 [USNMENT01521289]. 17, O. eugeniephillipsia, sp. nov., paratype, ACG, Costa Rica, USNM slide #116383 [USNMENT01894118]. 18, O. haemorrhoidalis, ACG, Costa Rica, USNM slide #116403 [USNMENT01894170]. 19, O. abruptalis, Sri Lanka, USNM slide #116380 [USNMENT01894185]. [black arrows indicate projections from anterior section of antrum]" figureDoi="http://doi.org/10.5281/zenodo.12190266" httpUri="https://zenodo.org/record/12190266/files/figure.png" pageId="3" pageNumber="386">Fig. 17</figureCitation>
), but in
<taxonomicName id="4C384D7BFFABFFF24FD9F848FD24F812" box="[435,658,1951,1977]" class="Insecta" family="Crambidae" genus="Orphanostigma" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="386" phylum="Arthropoda" rank="species" species="haemorrhoidalis">
<emphasis id="B94CEAEAFFABFFF24FD9F848FD24F812" box="[435,658,1951,1977]" italics="true" pageId="3" pageNumber="386">O. haemorrhoidalis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C384D7BFFABFFF24CA4F848FCFFF812" authorityName="Solis &amp; Cock &amp; Janzen &amp; Hallwachs" authorityYear="2024" baseAuthorityName="Barnes &amp; McDunnough" baseAuthorityYear="1914" box="[718,841,1951,1977]" class="Insecta" family="Crambidae" genus="Orphanostigma" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="386" phylum="Arthropoda" rank="species" species="futilalis">
<emphasis id="B94CEAEAFFABFFF24CA4F848FCFFF812" box="[718,841,1951,1977]" italics="true" pageId="3" pageNumber="386">O. futilalis</emphasis>
</taxonomicName>
the lateral projections from the anterior section are short and stubby (
<figureCitation id="13032A7DFFABFFF24F35F814FE57F876" box="[351,481,1987,2013]" captionStart="FIGURES 1619" captionStartId="5.[152,270,1848,1872]" captionTargetBox="[351,1239,195,1813]" captionTargetId="figure-18@5.[331,1257,181,1824]" captionTargetPageId="5" captionText="FIGURES 1619. 16, Orphanostigma futilalis paralectotype female genitalia, Brownsville, Texas, USNM slide #116,365 [USNMENT01521289]. 17, O. eugeniephillipsia, sp. nov., paratype, ACG, Costa Rica, USNM slide #116383 [USNMENT01894118]. 18, O. haemorrhoidalis, ACG, Costa Rica, USNM slide #116403 [USNMENT01894170]. 19, O. abruptalis, Sri Lanka, USNM slide #116380 [USNMENT01894185]. [black arrows indicate projections from anterior section of antrum]" figureDoi="http://doi.org/10.5281/zenodo.12190266" httpUri="https://zenodo.org/record/12190266/files/figure.png" pageId="3" pageNumber="386">Figs 16, 18</figureCitation>
). The lateral projections originate dorsolaterally in
<taxonomicName id="4C384D7BFFABFFF24A4BF814FB2FF876" authorityName="Solis &amp; Cock &amp; Janzen &amp; Hallwachs" authorityYear="2024" baseAuthorityName="Barnes &amp; McDunnough" baseAuthorityYear="1914" box="[1057,1177,1987,2013]" class="Insecta" family="Crambidae" genus="Orphanostigma" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="386" phylum="Arthropoda" rank="species" species="futilalis">
<emphasis id="B94CEAEAFFABFFF24A4BF814FB2FF876" box="[1057,1177,1987,2013]" italics="true" pageId="3" pageNumber="386">O. futilalis</emphasis>
</taxonomicName>
(
<figureCitation id="13032A7DFFABFFF24AC2F814FB4DF876" box="[1192,1275,1987,2013]" captionStart="FIGURES 1619" captionStartId="5.[152,270,1848,1872]" captionTargetBox="[351,1239,195,1813]" captionTargetId="figure-18@5.[331,1257,181,1824]" captionTargetPageId="5" captionText="FIGURES 1619. 16, Orphanostigma futilalis paralectotype female genitalia, Brownsville, Texas, USNM slide #116,365 [USNMENT01521289]. 17, O. eugeniephillipsia, sp. nov., paratype, ACG, Costa Rica, USNM slide #116383 [USNMENT01894118]. 18, O. haemorrhoidalis, ACG, Costa Rica, USNM slide #116403 [USNMENT01894170]. 19, O. abruptalis, Sri Lanka, USNM slide #116380 [USNMENT01894185]. [black arrows indicate projections from anterior section of antrum]" figureDoi="http://doi.org/10.5281/zenodo.12190266" httpUri="https://zenodo.org/record/12190266/files/figure.png" pageId="3" pageNumber="386">Fig. 16</figureCitation>
, black arrow), and ventrally in
<taxonomicName id="4C384D7BFFAEFFF74F3EFF40FD85FF1A" box="[340,563,151,177]" class="Insecta" family="Crambidae" genus="Orphanostigma" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="389" phylum="Arthropoda" rank="species" species="haemorrhoidalis">
<emphasis id="B94CEAEAFFAEFFF74F3EFF40FD85FF1A" box="[340,563,151,177]" italics="true" pageId="6" pageNumber="389">O. haemorrhoidalis</emphasis>
</taxonomicName>
(
<figureCitation id="13032A7DFFAEFFF74C2FFF40FD2FFF1A" box="[581,665,151,177]" captionStart="FIGURES 1619" captionStartId="5.[152,270,1848,1872]" captionTargetBox="[351,1239,195,1813]" captionTargetId="figure-18@5.[331,1257,181,1824]" captionTargetPageId="5" captionText="FIGURES 1619. 16, Orphanostigma futilalis paralectotype female genitalia, Brownsville, Texas, USNM slide #116,365 [USNMENT01521289]. 17, O. eugeniephillipsia, sp. nov., paratype, ACG, Costa Rica, USNM slide #116383 [USNMENT01894118]. 18, O. haemorrhoidalis, ACG, Costa Rica, USNM slide #116403 [USNMENT01894170]. 19, O. abruptalis, Sri Lanka, USNM slide #116380 [USNMENT01894185]. [black arrows indicate projections from anterior section of antrum]" figureDoi="http://doi.org/10.5281/zenodo.12190266" httpUri="https://zenodo.org/record/12190266/files/figure.png" pageId="6" pageNumber="389">Fig. 18</figureCitation>
, black arrow). The antrum of
<taxonomicName id="4C384D7BFFAEFFF74D9FFF40FB3AFF1A" baseAuthorityName="Walker" baseAuthorityYear="1859" box="[1013,1164,151,177]" class="Insecta" family="Crambidae" genus="Orphanostigma" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="389" phylum="Arthropoda" rank="species" species="abruptalis">
<emphasis id="B94CEAEAFFAEFFF74D9FFF40FB3AFF1A" box="[1013,1164,151,177]" italics="true" pageId="6" pageNumber="389">O. abruptalis</emphasis>
</taxonomicName>
lacks distinctive lateral projections, and the antrum is broader than New World species with a strap-like sclerotization anteriorly that is lacking in New World species (
<figureCitation id="13032A7DFFAEFFF74F9FFF08FDFFFF52" box="[501,585,223,249]" captionStart="FIGURES 1619" captionStartId="5.[152,270,1848,1872]" captionTargetBox="[351,1239,195,1813]" captionTargetId="figure-18@5.[331,1257,181,1824]" captionTargetPageId="5" captionText="FIGURES 1619. 16, Orphanostigma futilalis paralectotype female genitalia, Brownsville, Texas, USNM slide #116,365 [USNMENT01521289]. 17, O. eugeniephillipsia, sp. nov., paratype, ACG, Costa Rica, USNM slide #116383 [USNMENT01894118]. 18, O. haemorrhoidalis, ACG, Costa Rica, USNM slide #116403 [USNMENT01894170]. 19, O. abruptalis, Sri Lanka, USNM slide #116380 [USNMENT01894185]. [black arrows indicate projections from anterior section of antrum]" figureDoi="http://doi.org/10.5281/zenodo.12190266" httpUri="https://zenodo.org/record/12190266/files/figure.png" pageId="6" pageNumber="389">Fig. 19</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF476670FFACFFF54EF2F93AFEAAF876" ID-DOI="http://doi.org/10.5281/zenodo.12190264" ID-Zenodo-Dep="12190264" httpUri="https://zenodo.org/record/12190264/files/figure.png" pageId="4" pageNumber="387" startId="4.[152,270,1773,1797]" targetBox="[237,1359,181,1734]" targetPageId="4" targetType="figure">
<paragraph id="8B8736F8FFACFFF54EF2F93AFEAAF876" blockId="4.[151,1437,1773,2013]" pageId="4" pageNumber="387">
<emphasis id="B94CEAEAFFACFFF54EF2F93AFEE5F8AE" bold="true" box="[152,339,1773,1797]" pageId="4" pageNumber="387">FIGURES 815.</emphasis>
8,
<taxonomicName id="4C384D7BFFACFFF54F15F93AFDC1F8AE" authorityName="Solis &amp; Cock &amp; Janzen &amp; Hallwachs" authorityYear="2024" baseAuthorityName="Barnes &amp; McDunnough" baseAuthorityYear="1914" box="[383,631,1773,1797]" class="Insecta" family="Crambidae" genus="Orphanostigma" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="387" phylum="Arthropoda" rank="species" species="futilalis">
<emphasis id="B94CEAEAFFACFFF54F15F93AFDC1F8AE" box="[383,631,1773,1797]" italics="true" pageId="4" pageNumber="387">Orphanostigma futilalis</emphasis>
</taxonomicName>
lectotype male genitalia, San Benito, Texas, USNM slide #116366. 9,
<taxonomicName id="4C384D7BFFACFFF54BEEF939FF52F882" authorityName="Solis &amp; Cock &amp; Janzen &amp; Hallwachs" authorityYear="2024" baseAuthorityName="Barnes &amp; McDunnough" baseAuthorityYear="1914" class="Insecta" family="Crambidae" genus="Orphanostigma" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="387" phylum="Arthropoda" rank="species" species="futilalis">
<emphasis id="B94CEAEAFFACFFF54BEEF939FF52F882" italics="true" pageId="4" pageNumber="387">O. futilalis</emphasis>
</taxonomicName>
phallus, USNM slide #116366. 10,
<taxonomicName id="4C384D7BFFACFFF54C07F8C5FC8FF882" box="[621,825,1809,1833]" class="Insecta" family="Crambidae" genus="Orphanostigma" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="387" phylum="Arthropoda" rank="species" species="haemorrhoidalis">
<emphasis id="B94CEAEAFFACFFF54C07F8C5FDC9F882" box="[621,639,1810,1833]" italics="true" pageId="4" pageNumber="387">O</emphasis>
.
<emphasis id="B94CEAEAFFACFFF54CFBF8C6FC8FF882" box="[657,825,1809,1833]" italics="true" pageId="4" pageNumber="387">haemorrhoidalis</emphasis>
</taxonomicName>
male genitalia, ACG, Costa Rica, USNM slide #116402 [USNMENT01894170]. 11,
<taxonomicName id="4C384D7BFFACFFF54FD4F8E1FD31F8E6" box="[446,647,1845,1869]" class="Insecta" family="Crambidae" genus="Orphanostigma" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="387" phylum="Arthropoda" rank="species" species="haemorrhoidalis">
<emphasis id="B94CEAEAFFACFFF54FD4F8E1FE66F8E6" box="[446,464,1846,1869]" italics="true" pageId="4" pageNumber="387">O</emphasis>
.
<emphasis id="B94CEAEAFFACFFF54FB5F8E2FD31F8E6" box="[479,647,1845,1869]" italics="true" pageId="4" pageNumber="387">haemorrhoidalis</emphasis>
</taxonomicName>
phallus, USNM slide #116363. 12,
<taxonomicName id="4C384D7BFFACFFF54A6AF8E1FB3FF8E6" baseAuthorityName="Walker" baseAuthorityYear="1859" box="[1024,1161,1845,1869]" class="Insecta" family="Crambidae" genus="Orphanostigma" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="387" phylum="Arthropoda" rank="species" species="abruptalis">
<emphasis id="B94CEAEAFFACFFF54A6AF8E1FB3FF8E6" box="[1024,1161,1845,1869]" italics="true" pageId="4" pageNumber="387">O. abruptalis</emphasis>
</taxonomicName>
male genitalia, Sri Lanka, USNM slide #116379 [USNMENT01894037]. 13,
<taxonomicName id="4C384D7BFFACFFF54CCFF88DFC9BF8DA" baseAuthorityName="Walker" baseAuthorityYear="1859" box="[677,813,1881,1905]" class="Insecta" family="Crambidae" genus="Orphanostigma" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="387" phylum="Arthropoda" rank="species" species="abruptalis">
<emphasis id="B94CEAEAFFACFFF54CCFF88DFC9BF8DA" box="[677,813,1881,1905]" italics="true" pageId="4" pageNumber="387">O. abruptalis</emphasis>
</taxonomicName>
phallus, USNM slide #116379. 14,
<emphasis id="B94CEAEAFFACFFF54ACBF88DFAC0F8DA" box="[1185,1398,1881,1905]" italics="true" pageId="4" pageNumber="387">
<taxonomicName id="4C384D7BFFACFFF54ACBF88DFAC4F8DA" authority="Solis &amp; Cock &amp; Janzen &amp; Hallwachs, 2024" authorityName="Solis &amp; Cock &amp; Janzen &amp; Hallwachs" authorityYear="2024" box="[1185,1394,1881,1905]" class="Insecta" family="Crambidae" genus="Orphanostigma" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="387" phylum="Arthropoda" rank="species" species="eugeniephillipsia" status="sp. nov.">O. eugeniephillipsia</taxonomicName>
,
</emphasis>
<emphasis id="B94CEAEAFFACFFF54B14F88EFF75F83E" bold="true" pageId="4" pageNumber="387">
<taxonomicNameLabel id="A27F5791FFACFFF54B14F88EFF75F83E" pageId="4" pageNumber="387" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype male genitalia, ACG, Costa Rica, USNM slide #116382 [USNMENT01894117]. 15,
<emphasis id="B94CEAEAFFACFFF54ACAF8A9FAC0F83E" box="[1184,1398,1917,1941]" italics="true" pageId="4" pageNumber="387">
<taxonomicName id="4C384D7BFFACFFF54ACAF8A9FAC4F83E" authority="Solis &amp; Cock &amp; Janzen &amp; Hallwachs, 2024" authorityName="Solis &amp; Cock &amp; Janzen &amp; Hallwachs" authorityYear="2024" box="[1184,1394,1917,1941]" class="Insecta" family="Crambidae" genus="Orphanostigma" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="387" phylum="Arthropoda" rank="species" species="eugeniephillipsia" status="sp. nov.">O. eugeniephillipsia</taxonomicName>
,
</emphasis>
<emphasis id="B94CEAEAFFACFFF54B14F8AAFF75F812" bold="true" pageId="4" pageNumber="387">
<taxonomicNameLabel id="A27F5791FFACFFF54B14F8AAFF75F812" pageId="4" pageNumber="387" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype phallus, USNM slide #116382. [black arrows indicate the saccular margin and species-specific ornamentation on the valva]
</paragraph>
</caption>
<caption id="DF476670FFADFFF44EF2F8EFFEB5F84B" ID-DOI="http://doi.org/10.5281/zenodo.12190266" ID-Zenodo-Dep="12190266" httpUri="https://zenodo.org/record/12190266/files/figure.png" pageId="5" pageNumber="388" startId="5.[152,270,1848,1872]" targetBox="[351,1239,195,1813]" targetPageId="5" targetType="figure">
<paragraph id="8B8736F8FFADFFF44EF2F8EFFEB5F84B" blockId="5.[151,1437,1848,2016]" pageId="5" pageNumber="388">
<emphasis id="B94CEAEAFFADFFF44EF2F8EFFED1F8FB" bold="true" box="[152,359,1848,1872]" pageId="5" pageNumber="388">FIGURES 1619.</emphasis>
16,
<taxonomicName id="4C384D7BFFADFFF44FC4F8EFFD1BF8FB" authorityName="Solis &amp; Cock &amp; Janzen &amp; Hallwachs" authorityYear="2024" baseAuthorityName="Barnes &amp; McDunnough" baseAuthorityYear="1914" box="[430,685,1848,1872]" class="Insecta" family="Crambidae" genus="Orphanostigma" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="388" phylum="Arthropoda" rank="species" species="futilalis">
<emphasis id="B94CEAEAFFADFFF44FC4F8EFFD1BF8FB" box="[430,685,1848,1872]" italics="true" pageId="5" pageNumber="388">Orphanostigma futilalis</emphasis>
</taxonomicName>
paralectotype female genitalia, Brownsville, Texas, USNM slide #116,365 [USNMENT01521289]. 17,
<emphasis id="B94CEAEAFFADFFF44C5AF88AFCBFF8DF" box="[560,777,1884,1908]" italics="true" pageId="5" pageNumber="388">
<taxonomicName id="4C384D7BFFADFFF44C5AF88AFCB3F8DF" authority="Solis &amp; Cock &amp; Janzen &amp; Hallwachs, 2024" authorityName="Solis &amp; Cock &amp; Janzen &amp; Hallwachs" authorityYear="2024" box="[560,773,1884,1908]" class="Insecta" family="Crambidae" genus="Orphanostigma" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="388" phylum="Arthropoda" rank="species" species="eugeniephillipsia" status="sp. nov.">O. eugeniephillipsia</taxonomicName>
,
</emphasis>
<emphasis id="B94CEAEAFFADFFF44D7FF88BFCDDF8DF" bold="true" box="[789,875,1884,1908]" pageId="5" pageNumber="388">
<taxonomicNameLabel id="A27F5791FFADFFF44D7FF88BFCDDF8DF" box="[789,875,1884,1908]" pageId="5" pageNumber="388" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype, ACG, Costa Rica, USNM slide #116383 [USNMENT01894118]. 18,
<taxonomicName id="4C384D7BFFADFFF44FA8F856FD63F833" authority=", ACG" authorityName="ACG" box="[450,725,1920,1944]" class="Insecta" family="Crambidae" genus="Orphanostigma" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="388" phylum="Arthropoda" rank="species" species="haemorrhoidalis">
<emphasis id="B94CEAEAFFADFFF44FA8F856FE62F833" box="[450,468,1921,1944]" italics="true" pageId="5" pageNumber="388">O</emphasis>
.
<emphasis id="B94CEAEAFFADFFF44F8FF857FD3BF833" box="[485,653,1920,1944]" italics="true" pageId="5" pageNumber="388">haemorrhoidalis</emphasis>
, ACG
</taxonomicName>
, Costa Rica, USNM slide #116403 [USNMENT01894170]. 19,
<emphasis id="B94CEAEAFFADFFF44BEEF856FEB0F817" italics="true" pageId="5" pageNumber="388">
<taxonomicName id="4C384D7BFFADFFF44BEEF856FEB4F817" baseAuthorityName="Walker" baseAuthorityYear="1859" class="Insecta" family="Crambidae" genus="Orphanostigma" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="388" phylum="Arthropoda" rank="species" species="abruptalis">O. abruptalis</taxonomicName>
,
</emphasis>
Sri Lanka, USNM slide #116380 [USNMENT01894185]. [black arrows indicate projections from anterior section of antrum]
</paragraph>
</caption>
<subSubSection id="C3226573FFAEFFF74EADFED5FBD9FEB6" box="[199,1135,258,285]" pageId="6" pageNumber="389" type="distribution">
<paragraph id="8B8736F8FFAEFFF74EADFED5FBD9FEB6" blockId="6.[151,1437,151,609]" box="[199,1135,258,285]" pageId="6" pageNumber="389">
<emphasis id="B94CEAEAFFAEFFF74EADFED5FED7FEB7" bold="true" box="[199,353,258,284]" pageId="6" pageNumber="389">Distribution.</emphasis>
Known only from the Area de Conservación
<collectingRegion id="49FCF81AFFAEFFF74D35FED4FC55FEB6" box="[863,995,259,285]" country="Costa Rica" name="Guanacaste" pageId="6" pageNumber="389">Guanacaste</collectingRegion>
,
<collectingCountry id="F32F7668FFAEFFF74D84FED4FBDDFEB6" box="[1006,1131,259,285]" name="Costa Rica" pageId="6" pageNumber="389">Costa Rica</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3226573FFAEFFF74EADFEF0FB85FE7A" pageId="6" pageNumber="389" type="discussion">
<paragraph id="8B8736F8FFAEFFF74EADFEF0FB85FE7A" blockId="6.[151,1437,151,609]" pageId="6" pageNumber="389">
<emphasis id="B94CEAEAFFAEFFF74EADFEF0FE8DFEEA" bold="true" box="[199,315,295,321]" pageId="6" pageNumber="389">Remarks.</emphasis>
The four light-caught specimens of the
<typeStatus id="5483885AFFAEFFF74C9AFEF0FC96FEEA" box="[752,800,295,321]" pageId="6" pageNumber="389">type</typeStatus>
series were not reared, although its host plant is predicted to be in the
<taxonomicName id="4C384D7BFFAEFFF74F70FE9CFE11FECE" box="[282,423,331,357]" class="Magnoliopsida" family="Verbenaceae" kingdom="Plantae" order="Lamiales" pageId="6" pageNumber="389" phylum="Tracheophyta" rank="family">Verbenaceae</taxonomicName>
because most of the
<taxonomicName id="4C384D7BFFAEFFF74CF8FE9CFCDBFECE" box="[658,877,331,357]" class="Insecta" family="Crambidae" genus="Orphanostigma" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="389" phylum="Arthropoda" rank="species" species="haemorrhoidalis">
<emphasis id="B94CEAEAFFAEFFF74CF8FE9CFCDBFECE" box="[658,877,331,357]" italics="true" pageId="6" pageNumber="389">O. haemorrhoidalis</emphasis>
</taxonomicName>
specimens from the ACG,
<collectingCountry id="F32F7668FFAEFFF74AF7FE9CFAA1FECE" box="[1181,1303,331,357]" name="Costa Rica" pageId="6" pageNumber="389">Costa Rica</collectingCountry>
were found feeding on species in this plant family (Janzen &amp; Hallwachs 2009). Most were reared on
<taxonomicName id="4C384D7BFFAEFFF74AFEFEB8FAE6FE23" box="[1172,1360,367,392]" class="Magnoliopsida" family="Verbenaceae" genus="Lantana" kingdom="Plantae" order="Lamiales" pageId="6" pageNumber="389" phylum="Tracheophyta" rank="species" species="camara">
<emphasis id="B94CEAEAFFAEFFF74AFEFEB8FAE6FE23" box="[1172,1360,367,392]" italics="true" pageId="6" pageNumber="389">Lantana camara</emphasis>
</taxonomicName>
, and a much fewer number were found feeding on
<taxonomicName id="4C384D7BFFAEFFF74C11FE44FC9AFE06" box="[635,812,403,429]" class="Magnoliopsida" family="Verbenaceae" genus="Lantana" kingdom="Plantae" order="Lamiales" pageId="6" pageNumber="389" phylum="Tracheophyta" rank="species" species="trifolia">
<emphasis id="B94CEAEAFFAEFFF74C11FE44FC9AFE06" box="[635,812,403,429]" italics="true" pageId="6" pageNumber="389">Lantana trifolia</emphasis>
</taxonomicName>
L., and additionally on
<taxonomicName id="4C384D7BFFAEFFF74A5BFE44FB74FE06" box="[1073,1218,403,429]" class="Magnoliopsida" family="Verbenaceae" kingdom="Plantae" order="Lamiales" pageId="6" pageNumber="389" phylum="Tracheophyta" rank="family">Verbenaceae</taxonomicName>
,
<specimenCount id="9D3EFD71FFAEFFF74AA1FE44FAD5FE06" box="[1227,1379,403,429]" count="1" pageId="6" pageNumber="389" type="generic">one specimen</specimenCount>
each was reared on
<taxonomicName id="4C384D7BFFAEFFF74F50FE60FDDFFE7A" authority="Schauer" authorityName="Schauer" box="[314,617,439,465]" class="Magnoliopsida" family="Verbenaceae" genus="Lippia" kingdom="Plantae" order="Lamiales" pageId="6" pageNumber="389" phylum="Tracheophyta" rank="species" species="berlandieri">
<emphasis id="B94CEAEAFFAEFFF74F50FE60FDBEFE7A" box="[314,520,439,465]" italics="true" pageId="6" pageNumber="389">Lippia berlandieri</emphasis>
Schauer
</taxonomicName>
and
<taxonomicName id="4C384D7BFFAEFFF74CF5FE60FB85FE7A" authority="(Donn. Sm.) Standl." authorityName="Standl." baseAuthorityName="Donn. Sm." box="[671,1075,439,465]" class="Magnoliopsida" family="Verbenaceae" genus="Lippia" kingdom="Plantae" order="Lamiales" pageId="6" pageNumber="389" phylum="Tracheophyta" rank="species" species="oxyphyllaria">
<emphasis id="B94CEAEAFFAEFFF74CF5FE60FCFDFE7A" box="[671,843,439,465]" italics="true" pageId="6" pageNumber="389">L. oxyphyllaria</emphasis>
(Donn. Sm.) Standl.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C3226573FFAEFFF74EADFE0CFC8EFDCA" pageId="6" pageNumber="389" type="etymology">
<paragraph id="8B8736F8FFAEFFF74EADFE0CFC8EFDCA" blockId="6.[151,1437,151,609]" pageId="6" pageNumber="389">
<emphasis id="B94CEAEAFFAEFFF74EADFE0CFEF8FE5E" bold="true" box="[199,334,475,501]" pageId="6" pageNumber="389">Etymology.</emphasis>
This species is named in honor of Dr. Eugenie Phillips of
<collectingRegion id="49FCF81AFFAEFFF74DA4FE0CFB87FE5E" box="[974,1073,475,501]" country="Costa Rica" name="San Jose" pageId="6" pageNumber="389">San Jose</collectingRegion>
,
<collectingCountry id="F32F7668FFAEFFF74A51FE0CFB02FE5E" box="[1083,1204,475,501]" name="Costa Rica" pageId="6" pageNumber="389">Costa Rica</collectingCountry>
in recognition of her decades of dedicated attention to the taxonomy of Costa Rican microlepidoptera, and especially her efforts to the integration of Costa Rican biodiversity with social processes for widespread public understanding of the importance of the survival of Costa Ricas wildland and its biodiversity.
</paragraph>
</subSubSection>
</treatment>
</document>