treatments-xml/data/03/91/87/0391879409774F63FC52FA06FD71EA31.xml

<document id="3BF1B1D14092FB015DF73AC6996F0992" ID-CLB-Dataset="133823" ID-DOI="10.1093/zoolinnean/zlac002" ID-GBIF-Dataset="c3c3d111-1fdd-48e2-a738-40935b0165ca" ID-ISSN="0024-4082" ID-Zenodo-Dep="7037692" ID-ZooBank="B5B83725-6D9F-4153-8B89-4001FDCE46CD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1661767516544" checkinUser="plazi" docAuthor="Paz-Sedano, Sofía, Smirnoff, Dimitri, Candás, María, Gosliner, Terrence M. &amp; Pola, Marta" docDate="2022" docId="0391879409774F63FC52FA06FD71EA31" docLanguage="en" docName="zlac002.pdf" docOrigin="Zoological Journal of the Linnean Society 196" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Murphydoris adusta Paz-Sedano &amp; Smirnoff &amp; Candás &amp; Gosliner &amp; Pola 2022, SP. NOV." docType="treatment" docVersion="9" lastPageNumber="259" masterDocId="FFA8FFEC09734F6AFFE2FFF6FFC3EE07" masterDocTitle="Rediscovering the overlooked genus Murphydoris (Nudibranchia: Goniodorididae): the first phylogeny and addition of four new species" masterLastPageNumber="269" masterPageNumber="250" pageNumber="254" updateTime="1699374003752" updateUser="ExternalLinkService">
<mods:mods id="2084F445BFA7EE0D1DB7C0171D09D2DA" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="9452D3FD0BD0E2AB6E2BC2EA4BAF8C48">
<mods:title id="F31E9EC338C366DD0C93719145D67A49">Rediscovering the overlooked genus Murphydoris (Nudibranchia: Goniodorididae): the first phylogeny and addition of four new species</mods:title>
</mods:titleInfo>
<mods:name id="08ED69DD4C9959F1BD7AAE2252670916" type="personal">
<mods:role id="571ABFC0CF13E8E560465C9E019DD1A0">
<mods:roleTerm id="A432B0079C45FC97ED337D72030E0B74">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D65B7333754B75F1577D859200932069">Paz-Sedano, Sofía</mods:namePart>
</mods:name>
<mods:name id="8E38AD1AC9C82F7DE7F492A59406261B" type="personal">
<mods:role id="F69B1A1FFD467EFD464AAEF67A00B889">
<mods:roleTerm id="C37E1C81FBD4DD0511CC562E7810104E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="690E306649626BB19C5CD10B091B364D">Smirnoff, Dimitri</mods:namePart>
</mods:name>
<mods:name id="29BFC682D7BC50E914D0CB91C6CEE5D7" type="personal">
<mods:role id="B8B3665CD72B3BD93023BE69B3690594">
<mods:roleTerm id="D3F742A86618F8892B06616EFB3C9EB7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="684A9B3C7C3DA7C3CFCE62A7819CE145">Candás, María</mods:namePart>
</mods:name>
<mods:name id="8687617EAE320CA606BE0A1591361389" type="personal">
<mods:role id="B0E1956E6E76649D5935482453F941E4">
<mods:roleTerm id="CC79BA9A41336998BA46E9B04B01239D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5216959DF06371BFD9A451D4DB42F06A">Gosliner, Terrence M.</mods:namePart>
</mods:name>
<mods:name id="2EF21C042E161AF6400E270727D5D1C2" type="personal">
<mods:role id="24F221A83AF893F0768AD634DF263248">
<mods:roleTerm id="5FBCD7EB175F5FAD8425ABD3546D6CC7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="626DA45D7D8363972009883C3B2D43A4">Pola, Marta</mods:namePart>
</mods:name>
<mods:typeOfResource id="67FE2DAF63952B97101CE1A39E9B18B0">text</mods:typeOfResource>
<mods:relatedItem id="CE8EBF110CEE92E84E73DB7E3B3314C6" type="host">
<mods:titleInfo id="2C144B42C1816B76BA55B85934DD4D9E">
<mods:title id="393F6A0A1B3F51014B9FD03F405E7FBA">Zoological Journal of the Linnean Society</mods:title>
</mods:titleInfo>
<mods:part id="D589A3D979319071F9AB833DBAFD2168">
<mods:date id="05B240BB9059EEF9C92B54BBD79D6B26">2022</mods:date>
<mods:detail id="F43798F342BB27FDBE84799CFCAD70F3" type="volume">
<mods:number id="A97796D3DD072BE5C4A0A86CAAAC8F57">196</mods:number>
</mods:detail>
<mods:extent id="678FB2B49BCE853CF249E804C72C3E2E" unit="page">
<mods:start id="54333323E148CCD5FA786E70F492D27B">250</mods:start>
<mods:end id="60FCE496D4FCC0D1FE3A5F0AB2EED7D0">269</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="30F8CD573EE265BF69075BFC62B57117">journal article</mods:classification>
<mods:identifier id="10729B674FCD942AFAB16CB2E6449FC2" type="CLB-Dataset">133823</mods:identifier>
<mods:identifier id="F243AB84B131785FA44EAFE98B5E67AC" type="DOI">10.1093/zoolinnean/zlac002</mods:identifier>
<mods:identifier id="CC1E91A5A695D53567BEAD5D1A12853E" type="GBIF-Dataset">c3c3d111-1fdd-48e2-a738-40935b0165ca</mods:identifier>
<mods:identifier id="99A2FC8643454E11FBACE04D62D0D290" type="ISSN">0024-4082</mods:identifier>
<mods:identifier id="C438A724B7139794902F4151697E87A6" type="Zenodo-Dep">7037692</mods:identifier>
<mods:identifier id="ABF3CC377D3D3192B2722B3357DF38FF" type="ZooBank">B5B83725-6D9F-4153-8B89-4001FDCE46CD</mods:identifier>
</mods:mods>
<treatment id="0391879409774F63FC52FA06FD71EA31" ID-DOI="http://doi.org/10.5281/zenodo.7043790" ID-GBIF-Taxon="199901873" ID-Zenodo-Dep="7043790" LSID="urn:lsid:plazi:treatment:0391879409774F63FC52FA06FD71EA31" httpUri="http://treatment.plazi.org/id/0391879409774F63FC52FA06FD71EA31" lastPageId="9" lastPageNumber="259" pageId="4" pageNumber="254">
<subSubSection id="C322650909774F6EFC52FA06FAEEE80F" box="[944,1325,1520,1544]" pageId="4" pageNumber="254" type="nomenclature">
<paragraph id="8B87368209774F6EFC52FA06FAEEE80F" blockId="4.[944,1325,1520,1544]" box="[944,1325,1520,1544]" pageId="4" pageNumber="254">
<heading id="D0CF81EE09774F6EFC52FA06FAEEE80F" box="[944,1325,1520,1544]" centered="true" fontSize="9" level="2" pageId="4" pageNumber="254" reason="2">
<taxonomicName id="4C384D0109774F6EFC52FA06FB00E801" ID-CoL="8YN8R" authority="Paz-Sedano &amp; Smirnoff &amp; Candás &amp; Gosliner &amp; Pola, 2022" authorityName="Paz-Sedano &amp; Smirnoff &amp; Candás &amp; Gosliner &amp; Pola" authorityYear="2022" box="[944,1219,1520,1543]" class="Gastropoda" family="Goniodorididae" genus="Murphydoris" kingdom="Animalia" order="Nudibranchia" pageId="4" pageNumber="267" phylum="Mollusca" rank="species" species="adusta" status="sp. nov.">MURPHYDORIS ADUSTA</taxonomicName>
<emphasis id="B94CEA9009774F6EFB28FA05FAEEE80F" bold="true" box="[1226,1325,1520,1544]" pageId="4" pageNumber="254">
<taxonomicNameLabel id="A27F57EB09774F6EFB28FA05FAEEE80F" box="[1226,1325,1520,1544]" pageId="4" pageNumber="254" rank="species">SP. NOV.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C322650909774F6EFC38F9E0FAC0E828" box="[986,1283,1558,1583]" pageId="4" pageNumber="254" type="description">
<paragraph id="8B87368209774F6EFC38F9E0FAC0E828" blockId="4.[986,1283,1558,1583]" box="[986,1283,1558,1583]" pageId="4" pageNumber="254">
(
<figureCitation id="13032A0709774F6EFC01F9E0FB9DE828" box="[995,1118,1558,1583]" captionStart="Figure 2" captionStartId="6.[163,241,1491,1513]" captionTargetBox="[323,1283,196,1451]" captionTargetId="figure-159@6.[323,1283,195,1451]" captionTargetPageId="6" captionText="Figure 2. Photographs of living animals. A, B, Murphydoris adusta sp. nov., photographed by G. Cobb: A, (WAM S72660); B, (QMMO 85995). C, Murphydoris cobbi sp. nov. (CASIZ 206795), photographed by T. M. Gosliner. D, M. cobbi sp. nov. (QMMO 85979), photographed by G. Cobb. E, Murphydoris maracabranchia sp. nov. (CASIZ 186120), photographed by P. Paleracio. F, G, M. puncticulata sp. nov. photographed by G. Cobb. F, (WAM S72711), G, (WAM S72710)." figureDoi="http://doi.org/10.5281/zenodo.7037696" httpUri="https://zenodo.org/record/7037696/files/figure.png" pageId="4" pageNumber="254">FIGS 2A, B</figureCitation>
, 
<figureCitation id="13032A0709774F6EFB89F9E0FB68E828" box="[1131,1195,1558,1583]" captionStart="Figure 3" captionStartId="7.[144,221,1453,1475]" captionTargetBox="[224,1343,196,1404]" captionTargetId="figure-135@7.[223,1346,195,1413]" captionTargetPageId="7" captionText="Figure 3. A–C, drawings of Murphydoris adusta sp. nov.:A, gill branches; B, buccal bulb (MNCN 15.05/200144); C, reproductive system (MNCN 15.05/200144). D–F, drawings of Murphydoris cobbi sp. nov.: D, modification of gill branches; E, buccal bulb (QMMO 85983); F, reproductive system (QMMO 85982).G–H, drawings ofMurphydoris maracabranchiasp. nov.:G,modification of gill branches; H, reproductive system (CASIZ 224657). I–K, drawings of Murphydoris puncticulata sp. nov.: I, gill branches; J, buccal bulb (WAM S72711); K, reproductive system (WAM S72711). Abbreviations: am, ampulla; bc, bursa copulatrix; bp, buccal pump; fgm, female gland mass; hd, hermaphroditic duct; oe, oesophagus; op, oesophagal pump; p, penis; pr, prostate; ra, radular sac; rs, receptaculum seminis; sgl, salivary gland; ud, uterine duct; va, vagina; vd, vas deferens. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7037702" httpUri="https://zenodo.org/record/7037702/files/figure.png" pageId="4" pageNumber="254">3A–C</figureCitation>
, 
<figureCitation id="13032A0709774F6EFB5AF9E0FB39E828" box="[1208,1274,1558,1583]" captionStart="Figure 4" captionStartId="8.[163,241,1584,1606]" captionTargetBox="[323,1283,196,1544]" captionTargetId="figure-87@8.[323,1283,195,1544]" captionTargetPageId="8" captionText="Figure 4. Scanning electron microscope (SEM) photographs and light microscope photographs (LMP) of A–D, Murphydoris adusta sp. nov. and E–H, Murphydoris cobbi sp. nov. A, frontal view of radula (QMMO 85995) (SEM); B, lateral view of teeth (CASIZ 181290) (SEM); C, detail of masticatory margin of teeth (QMMO 85995) (SEM); D, detail of penial spines (QMMO 85986) (SEM); E, detail of labial cuticle elements (QMMO 85984) (SEM); F, frontal view of radula – detail of rachis, internal and external teeth (QMMO 85983) (SEM); G, detail of internal teeth (QMMO 85983) (SEM); H, detail of penial spines (LMP) (QMMO 85983). Scale bars: A, 50 µm; B, 40 µm; C, 10 µm; D, 10 µm; E, 5µm; F, 30 µm; G, 20 µm; H, 10 µm." figureDoi="http://doi.org/10.5281/zenodo.7037704" httpUri="https://zenodo.org/record/7037704/files/figure.png" pageId="4" pageNumber="254">4A–D</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C322650909774F6EFCD9F9C9FAE9E8D7" pageId="4" pageNumber="254" type="reference_group">
<paragraph id="8B87368209774F6EFCD9F9C9FAE9E8D7" blockId="4.[827,1414,1599,1744]" pageId="4" pageNumber="254">
<treatmentCitationGroup id="AB2811AC09774F6EFCD9F9C9FAE9E8D7" pageId="4" pageNumber="254">
‘ 
<taxonomicName id="4C384D0109774F6EFCA2F9C9FBDAE852" ID-CoL="3GWR8" authorityName="Alder &amp; Hancock" authorityYear="1864" box="[832,1049,1599,1621]" class="Gastropoda" family="Goniodorididae" genus="Goniodoris" kingdom="Animalia" order="Nudibranchia" pageId="4" pageNumber="254" phylum="Mollusca" rank="species" species="aspersa">
<emphasis id="B94CEA9009774F6EFCA2F9C9FBDAE852" box="[832,1049,1599,1621]" italics="true" pageId="4" pageNumber="254">Goniodoris aspersa</emphasis>
</taxonomicName>
’ 
<emphasis id="B94CEA9009774F6EFBC7F9B6FBF2E852" box="[1061,1073,1600,1621]" italics="true" pageId="4" pageNumber="254">–</emphasis>
<treatmentCitation id="0A99109309774F6EFBDAF9C9FA41E853" author="Debelius H &amp; Kuiter RH" box="[1080,1410,1599,1621]" page="28" pageId="4" pageNumber="254" year="2007">
<bibRefCitation id="EFA94B7309774F6EFBDAF9C9FA41E853" author="Debelius H &amp; Kuiter RH" box="[1080,1410,1599,1621]" pageId="4" pageNumber="254" refId="ref14257" refString="Debelius H, Kuiter RH. 2007. Nudibranchs of the world. Frankfurt: IKAN." type="book" year="2007">Debelius &amp; Kuiter (2007: 28)</bibRefCitation>
</treatmentCitation>
. ‘ 
<taxonomicName id="4C384D0109774F6EFCA2F9A8FBD3E874" box="[832,1040,1630,1651]" class="Gastropoda" family="Goniodorididae" genus="Goniodoridella" kingdom="Animalia" order="Nudibranchia" pageId="4" pageNumber="254" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="B94CEA9009774F6EFCA2F9A8FC29E874" box="[832,1002,1630,1651]" italics="true" pageId="4" pageNumber="254">Goniodoridella</emphasis>
sp.
</taxonomicName>
’ – 
<treatmentCitation id="0A99109309774F6EFBD2F9A8FAC8E873" author="Su Y &amp; Huang LJ &amp; Chang YW &amp; Mok HK" box="[1072,1291,1630,1652]" page="457" pageId="4" pageNumber="254" year="2009">
<bibRefCitation id="EFA94B7309774F6EFBD2F9A8FAC8E873" author="Su Y &amp; Huang LJ &amp; Chang YW &amp; Mok HK" box="[1072,1291,1630,1652]" pageId="4" pageNumber="254" pagination="448 - 459" refId="ref15961" refString="Su Y, Huang LJ, Chang YW, Mok HK. 2009. Temporal changes in nudibranch composition at a coastal site off Penghu (the Pescadores) in the Taiwan Strait. Zoological Studies 48: 448 - 459." type="journal article" year="2009">
Su 
<emphasis id="B94CEA9009774F6EFBB4F9A8FB4DE874" box="[1110,1166,1630,1651]" italics="true" pageId="4" pageNumber="254">et al.</emphasis>
(2009: 457
</bibRefCitation>
</treatmentCitation>
, fig. 11B). ‘ 
<taxonomicName id="4C384D0109774F6EFCA2F98AFBE6E895" box="[832,1061,1660,1682]" class="Gastropoda" family="Goniodorididae" genus="Goniodoridella" kingdom="Animalia" order="Nudibranchia" pageId="4" pageNumber="254" phylum="Mollusca" rank="species" species="undefined-6">
<emphasis id="B94CEA9009774F6EFCA2F98AFC29E896" box="[832,1002,1660,1681]" italics="true" pageId="4" pageNumber="254">Goniodoridella</emphasis>
sp. 6
</taxonomicName>
’ – 
<treatmentCitation id="0A99109309774F6EFBA7F98AFAAFE896" author="Gosliner TM &amp; Valdes A &amp; Behrens D" box="[1093,1388,1660,1682]" page="133" pageId="4" pageNumber="254" year="2015">
<bibRefCitation id="EFA94B7309774F6EFBA7F98AFAAFE896" author="Gosliner TM &amp; Valdes A &amp; Behrens D" box="[1093,1388,1660,1682]" pageId="4" pageNumber="254" refId="ref14453" refString="Gosliner TM, Valdes A, Behrens D. 2015. Nudibranch &amp; sea slug identification - Indo-Pacific. Jacksonville, Florida: New World Publications." type="book" year="2015">
Gosliner 
<emphasis id="B94CEA9009774F6EFB4FF98BFB26E896" box="[1197,1253,1660,1682]" italics="true" pageId="4" pageNumber="254">et al.</emphasis>
(2015: 133)
</bibRefCitation>
</treatmentCitation>
. ‘ 
<taxonomicName id="4C384D0109774F6EFCA2F96DFBE6E8B6" box="[832,1061,1691,1713]" class="Gastropoda" family="Goniodorididae" genus="Goniodoridella" kingdom="Animalia" order="Nudibranchia" pageId="4" pageNumber="254" phylum="Mollusca" rank="species" species="undefined-6">
<emphasis id="B94CEA9009774F6EFCA2F96DFC29E8B7" box="[832,1002,1691,1712]" italics="true" pageId="4" pageNumber="254">Goniodoridella</emphasis>
sp. 6
</taxonomicName>
’ – 
<treatmentCitation id="0A99109309774F6EFBA7F96DFA9DE8B6" author="Gosliner TM &amp; Valdes A &amp; Behrens D" box="[1093,1374,1691,1713]" page="51" pageId="4" pageNumber="254" year="2018">
<bibRefCitation id="EFA94B7309774F6EFBA7F96DFA9DE8B6" author="Gosliner TM &amp; Valdes A &amp; Behrens D" box="[1093,1374,1691,1713]" pageId="4" pageNumber="254" refId="ref14480" refString="Gosliner TM, Valdes A, Behrens D. 2018. Nudibranch &amp; sea slug identification - Indo-Pacific. Jacksonville, Florida: New World Publications." type="book" year="2018">
Gosliner 
<emphasis id="B94CEA9009774F6EFB4FF96AFB26E8B7" box="[1197,1253,1691,1713]" italics="true" pageId="4" pageNumber="254">et al.</emphasis>
(2018: 51)
</bibRefCitation>
</treatmentCitation>
. ‘ 
<taxonomicName id="4C384D0109774F6EFCA2F94CFBE6E8C8" box="[832,1061,1722,1743]" class="Gastropoda" family="Goniodorididae" genus="Goniodoridella" kingdom="Animalia" order="Nudibranchia" pageId="4" pageNumber="254" phylum="Mollusca" rank="species" species="undefined-3">
<emphasis id="B94CEA9009774F6EFCA2F94CFC29E8C8" box="[832,1002,1722,1743]" italics="true" pageId="4" pageNumber="254">Goniodoridella</emphasis>
sp. 3
</taxonomicName>
’ – 
<treatmentCitation id="0A99109309774F6EFBA7F94CFAE5E8D7" author="Nakano R" box="[1093,1318,1722,1744]" page="166" pageId="4" pageNumber="254" year="2018">
<bibRefCitation id="EFA94B7309774F6EFBA7F94CFAE5E8D7" author="Nakano R" box="[1093,1318,1722,1744]" pageId="4" pageNumber="254" refId="ref15081" refString="Nakano R. 2018. Field guide to sea slugs and nudibranchs of Japan. Tokyo: Bun-ichi Co., Ltd." type="book" year="2018">Nakano (2018: 166)</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C322650909774F6EFCD9F91FFA63E91A" pageId="4" pageNumber="254" type="description">
<paragraph id="8B87368209774F6EFCD9F91FFA63E91A" blockId="4.[827,1444,1768,1821]" pageId="4" pageNumber="254">
<emphasis id="B94CEA9009774F6EFCD9F91FFB4BE8F9" box="[827,1160,1768,1790]" italics="true" pageId="4" pageNumber="254">Z o o b a n k r e g i s t r a t i o n:</emphasis>
u r n: l s i d: z o o b a n k. org:act: 
<uuid id="FF9E0C5709774F6EFC6BF8F1FA58E91A" box="[905,1435,1799,1821]" pageId="4" pageNumber="254">A6372BF6-E947-4470-9B1D-D22A9AB5FE58</uuid>
.
</paragraph>
</subSubSection>
<subSubSection id="C322650909774F6FFCD9F8CDFF36E8FC" lastPageId="5" lastPageNumber="255" pageId="4" pageNumber="254" type="materials_examined">
<paragraph id="8B87368209774F6FFCD9F8CDFF36E8FC" blockId="4.[827,1442,1851,1904]" lastBlockId="5.[144,762,629,1787]" lastPageId="5" lastPageNumber="255" pageId="4" pageNumber="254">
<emphasis id="B94CEA9009774F6EFCD9F8CDFBF9E957" box="[827,1082,1851,1872]" italics="true" pageId="4" pageNumber="254">Material examined:</emphasis>
<materialsCitation id="3B503CDF09774F6EFBB4F8CDFA61E977" ID-GBIF-Occurrence="3897190345" collectionCode="QMMO" collectorName="La Balsa Park" country="Australia" location="Queensland" pageId="4" pageNumber="254" specimenCode="QMMO 85985" specimenCount="1" stateProvince="Queensland" typeStatus="holotype">
<emphasis id="B94CEA9009774F6EFBB4F8CDFB22E957" bold="true" box="[1110,1249,1851,1872]" pageId="4" pageNumber="254">
<typeStatus id="5483882009774F6EFBB4F8CDFB18E957" box="[1110,1243,1851,1872]" pageId="4" pageNumber="254" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="DB9E9EF909774F6EFB0CF8CDFA62E956" box="[1262,1441,1851,1873]" collectionCode="QMMO" pageId="4" pageNumber="254">QMMO 85985</specimenCode>
(
<collectingCountry id="F32F761209774F6EFCA1F8ACFC7BE977" box="[835,952,1882,1904]" name="Australia" pageId="4" pageNumber="254">Australia</collectingCountry>
, 
<collectingRegion id="49FCF86009774F6EFC25F8ACFB9AE977" box="[967,1113,1882,1904]" country="Australia" name="Queensland" pageId="4" pageNumber="254">Queensland</collectingRegion>
, 
<collectorName id="26CD535409774F6EFB85F8ACFADAE977" box="[1127,1305,1882,1904]" pageId="4" pageNumber="254">La Balsa Park</collectorName>
, Mooloolah
</materialsCitation>
River, 
<geoCoordinate id="EE0C504509764F6FFF3FFD81FEF4EC8B" box="[221,311,629,653]" degrees="26" direction="south" minutes="41" orientation="latitude" pageId="5" pageNumber="255" precision="925" value="-26.683332">26°41′S</geoCoordinate>
, 
<geoCoordinate id="EE0C504509764F6FFEA6FD80FE73EC8B" box="[324,432,629,652]" degrees="153" direction="east" minutes="08" orientation="longitude" pageId="5" pageNumber="255" precision="925" value="153.13333">153°08′E</geoCoordinate>
, 
<quantity id="4CC09B6709764F6FFE5FFD81FDC6EC8B" box="[445,517,631,653]" metricMagnitude="0" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="5" pageNumber="255" unit="m" value="4.5" valueMax="6.0" valueMin="3.0">3–6 m</quantity>
depth; coll. G. Cobb, 
<date id="FF86104209764F6FFF70FD63FEBAECAD" box="[146,377,661,683]" pageId="5" pageNumber="255" value="2019-09-29">29 September 2019</date>
), one preserved specimen, 
<quantity id="4CC09B6709764F6FFD55FD63FD39ECAC" box="[695,762,661,683]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="255" unit="mm" value="3.0">3 mm</quantity>
long, dissected. 
<materialsCitation id="3B503CDF09764F6FFEB1FD42FD29ECCD" ID-GBIF-Occurrence="3897190385" box="[339,746,692,714]" collectionCode="MNCN" pageId="5" pageNumber="255" specimenCode="MNCN 15.05" specimenCount="1" typeStatus="paratype">
<emphasis id="B94CEA9009764F6FFEB1FD42FE20ECCE" bold="true" box="[339,483,692,713]" pageId="5" pageNumber="255">
<typeStatus id="5483882009764F6FFEB1FD42FE1EECCE" box="[339,477,692,713]" pageId="5" pageNumber="255" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCode id="DB9E9EF909764F6FFE0EFD42FD4AECCD" box="[492,649,692,714]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="5" pageNumber="255" type="Museum">MNCN 15.05</specimenCode>
/200143
</materialsCitation>
– 
<materialsCitation id="3B503CDF09764F6FFF73FD25FE49ECEF" ID-GBIF-Occurrence="3897190305" box="[145,394,722,744]" collectionCode="MNCN" pageId="5" pageNumber="255" specimenCode="MNCN 15.05" specimenCount="1" typeStatus="paratype">
<specimenCode id="DB9E9EF909764F6FFF73FD25FEE9ECEF" box="[145,298,722,744]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="5" pageNumber="255" type="Museum">MNCN 15.05</specimenCode>
/200148
</materialsCitation>
, 
<materialsCitation id="3B503CDF09764F6FFE74FD25FDFFECEE" ID-GBIF-Occurrence="3897190351" accessionNumber="S72660" box="[406,572,723,745]" collectionCode="WAM" pageId="5" pageNumber="255" specimenCount="1" typeStatus="paratype">
<collectionCode id="ED29AE4709764F6FFE74FD25FE14ECEF" box="[406,471,723,744]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="5" pageNumber="255" type="Museum">WAM</collectionCode>
S72660,
</materialsCitation>
<materialsCitation id="3B503CDF09764F6FFDA7FD25FD29ECEF" ID-GBIF-Occurrence="3897190347" box="[581,746,722,744]" collectionCode="QMMO" pageId="5" pageNumber="255" specimenCode="QMMO 85986" specimenCount="1" typeStatus="paratype">
<specimenCode id="DB9E9EF909764F6FFDA7FD25FD29ECEF" box="[581,746,722,744]" collectionCode="QMMO" pageId="5" pageNumber="255">QMMO 85986</specimenCode>
</materialsCitation>
– 
<materialsCitation id="3B503CDF09764F6FFF73FD07FEF7ED00" ID-GBIF-Occurrence="3897190309" box="[145,308,753,775]" collectionCode="QMMO" pageId="5" pageNumber="255" specimenCode="QMMO 85988" specimenCount="1" typeStatus="paratype">
<specimenCode id="DB9E9EF909764F6FFF73FD07FEF7ED00" box="[145,308,753,775]" collectionCode="QMMO" pageId="5" pageNumber="255">QMMO 85988</specimenCode>
</materialsCitation>
; 
<materialsCitation id="3B503CDF09764F6FFEDDFD07FE2AED42" ID-GBIF-Occurrence="3897190393" collectionCode="QMMO" collectorName="G. Cobb" country="Australia" county="La Balsa Park" elevation="5" latitude="-26.683332" location="Mooloolah River" longLatPrecision="1240" longitude="153.13333" municipality="La Balsa Park" pageId="5" pageNumber="255" specimenCode="QMMO 85995" specimenCount="1" stateProvince="Queensland" typeStatus="paratype">
<specimenCode id="DB9E9EF909764F6FFEDDFD07FE22ED00" box="[319,481,753,775]" collectionCode="QMMO" pageId="5" pageNumber="255">QMMO 85995</specimenCode>
(
<collectingCountry id="F32F761209764F6FFE12FD07FD9CED00" box="[496,607,753,775]" name="Australia" pageId="5" pageNumber="255">Australia</collectingCountry>
, 
<collectingRegion id="49FCF86009764F6FFD88FD07FD36ED00" box="[618,757,753,775]" country="Australia" name="Queensland" pageId="5" pageNumber="255">Queensland</collectingRegion>
, 
<collectingMunicipality id="6BE3ACF809764F6FFF73FCE6FE83ED21" box="[145,320,784,806]" pageId="5" pageNumber="255">
<collectingCounty id="62E64E0E09764F6FFF73FCE6FE83ED21" box="[145,320,784,806]" pageId="5" pageNumber="255">La Balsa Park</collectingCounty>
</collectingMunicipality>
, 
<location id="8EE7605909764F6FFEAFFCE6FDD2ED21" LSID="urn:lsid:plazi:treatment:0391879409774F63FC52FA06FD71EA31:8EE7605909764F6FFEAFFCE6FDD2ED21" box="[333,529,784,806]" country="Australia" county="La Balsa Park" latitude="-26.683332" longLatPrecision="1240" longitude="153.13333" municipality="La Balsa Park" name="Mooloolah River" pageId="5" pageNumber="255" stateProvince="Queensland">Mooloolah River</location>
, 
<geoCoordinate id="EE0C504509764F6FFDFCFCE6FDBAED22" box="[542,633,783,806]" degrees="26" direction="south" minutes="41" orientation="latitude" pageId="5" pageNumber="255" precision="925" value="-26.683332">26°41′S</geoCoordinate>
, 
<geoCoordinate id="EE0C504509764F6FFD65FCE6FD37ED22" box="[647,756,783,806]" degrees="153" direction="east" minutes="08" orientation="longitude" pageId="5" pageNumber="255" precision="925" value="153.13333">153°08′E</geoCoordinate>
, 
<elevation id="0015D1B109764F6FFF73FCD9FEF3ED42" box="[145,304,815,837]" metricMagnitude="0" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="5" pageNumber="255" unit="m" value="4.5" valueMax="6.0" valueMin="3.0">
<quantity id="4CC09B6709764F6FFF73FCD9FF1EED43" box="[145,221,815,837]" metricMagnitude="0" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="5" pageNumber="255" unit="m" value="4.5" valueMax="6.0" valueMin="3.0">3–6 m</quantity>
depth
</elevation>
; coll. 
<collectorName id="26CD535409764F6FFE99FCD9FE22ED42" box="[379,481,815,837]" pageId="5" pageNumber="255">G. Cobb</collectorName>
)
</materialsCitation>
. 
<materialsCitation id="3B503CDF09764F6FFE15FCD9FE46ED65" ID-GBIF-Occurrence="3897190340" collectingDate="2019-09-29" collectionCode="MNCN" country="Australia" pageId="5" pageNumber="255" specimenCode="MNCN 15.05" specimenCount="1" stateProvince="Queensland" typeStatus="paratype">
<specimenCode id="DB9E9EF909764F6FFE15FCD9FD55ED43" box="[503,662,814,836]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="5" pageNumber="255" type="Museum">MNCN 15.05</specimenCode>
/200143 (
<date id="FF86104209764F6FFF7BFCBBFEBDED65" box="[153,382,845,867]" pageId="5" pageNumber="255" value="2019-09-29">
<collectingDate id="EFC2E9AA09764F6FFF7BFCBBFEBDED65" box="[153,382,845,867]" pageId="5" pageNumber="255" value="2019-09-29">29 September 2019</collectingDate>
</date>
)
</materialsCitation>
, 
<materialsCitation id="3B503CDF09764F6FFE73FCB8FE34ED85" ID-GBIF-Occurrence="3897190353" collectionCode="SEM" collectorName="Radula" country="Australia" pageId="5" pageNumber="255" specimenCount="1" stateProvince="Queensland" typeStatus="paratype">
one preserved specimen, 
<quantity id="4CC09B6709764F6FFD55FCBBFD3AED64" box="[695,761,845,867]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="255" unit="mm" value="2.0">2 mm</quantity>
long, dissected (
<collectionCode id="ED29AE4709764F6FFEACFC9AFE48ED86" box="[334,395,876,897]" pageId="5" pageNumber="255">SEM</collectionCode>
: 
<collectorName id="26CD535409764F6FFE75FC9AFE2CED85" box="[407,495,876,898]" pageId="5" pageNumber="255">Radula</collectorName>
)
</materialsCitation>
. 
<materialsCitation id="3B503CDF09764F6FFDE1FC9AFDFBEDA7" ID-GBIF-Occurrence="3897190325" collectingDate="2019-09-29" collectionCode="MNCN" country="Australia" pageId="5" pageNumber="255" specimenCode="MNCN 15.05" specimenCount="1" stateProvince="Queensland" typeStatus="paratype">
<specimenCode id="DB9E9EF909764F6FFDE1FC9AFD58ED85" box="[515,667,876,898]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="5" pageNumber="255" type="Museum">MNCN 15.05</specimenCode>
/200144 (
<figureCitation id="13032A0709764F6FFF78FC7DFEDAEDA7" box="[154,281,907,929]" captionStart="Figure 3" captionStartId="7.[144,221,1453,1475]" captionTargetBox="[224,1343,196,1404]" captionTargetId="figure-135@7.[223,1346,195,1413]" captionTargetPageId="7" captionText="Figure 3. A–C, drawings of Murphydoris adusta sp. nov.:A, gill branches; B, buccal bulb (MNCN 15.05/200144); C, reproductive system (MNCN 15.05/200144). D–F, drawings of Murphydoris cobbi sp. nov.: D, modification of gill branches; E, buccal bulb (QMMO 85983); F, reproductive system (QMMO 85982).G–H, drawings ofMurphydoris maracabranchiasp. nov.:G,modification of gill branches; H, reproductive system (CASIZ 224657). I–K, drawings of Murphydoris puncticulata sp. nov.: I, gill branches; J, buccal bulb (WAM S72711); K, reproductive system (WAM S72711). Abbreviations: am, ampulla; bc, bursa copulatrix; bp, buccal pump; fgm, female gland mass; hd, hermaphroditic duct; oe, oesophagus; op, oesophagal pump; p, penis; pr, prostate; ra, radular sac; rs, receptaculum seminis; sgl, salivary gland; ud, uterine duct; va, vagina; vd, vas deferens. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7037702" httpUri="https://zenodo.org/record/7037702/files/figure.png" pageId="5" pageNumber="255">Fig. 3B, C</figureCitation>
) (
<date id="FF86104209764F6FFED4FC7DFDF3EDA7" box="[310,560,907,929]" pageId="5" pageNumber="255" value="2019-09-29">
<collectingDate id="EFC2E9AA09764F6FFED4FC7DFDF3EDA7" box="[310,560,907,929]" pageId="5" pageNumber="255" value="2019-09-29">29 September 2019</collectingDate>
</date>
)
</materialsCitation>
, 
<materialsCitation id="3B503CDF09764F6FFDAAFC7DFD5EEDB8" ID-GBIF-Occurrence="3897190372" collectionCode="SEM" collectorName="Radula" country="Australia" pageId="5" pageNumber="255" specimenCount="1" stateProvince="Queensland" typeStatus="paratype">
one preserved specimen 
<quantity id="4CC09B6709764F6FFEE3FC5FFE82EDB8" box="[257,321,937,959]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="255" unit="mm" value="2.0">2 mm</quantity>
long, dissected (
<collectionCode id="ED29AE4709764F6FFE19FC5FFDF4EDB9" box="[507,567,937,958]" pageId="5" pageNumber="255">SEM</collectionCode>
: 
<collectorName id="26CD535409764F6FFDA2FC5FFD56EDB8" box="[576,661,937,959]" pageId="5" pageNumber="255">Radula</collectorName>
)
</materialsCitation>
. 
<materialsCitation id="3B503CDF09764F6FFD45FC5FFDFDEDDA" ID-GBIF-Occurrence="3897190373" collectingDate="2019-09-29" collectionCode="MNCN" country="Australia" pageId="5" pageNumber="255" specimenCode="MNCN 15.05" specimenCount="1" stateProvince="Queensland" typeStatus="paratype">
<specimenCode id="DB9E9EF909764F6FFD45FC5FFF0CEDD9" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="5" pageNumber="255" type="Museum">MNCN 15.05</specimenCode>
/200145 (
<date id="FF86104209764F6FFEA6FC3EFDF5EDDA" box="[324,566,968,990]" pageId="5" pageNumber="255" value="2019-09-29">
<collectingDate id="EFC2E9AA09764F6FFEA6FC3EFDF5EDDA" box="[324,566,968,990]" pageId="5" pageNumber="255" value="2019-09-29">29 September 2019</collectingDate>
</date>
)
</materialsCitation>
, 
<materialsCitation id="3B503CDF09764F6FFDAFFC3EFD5DEDFA" ID-GBIF-Occurrence="3897190363" collectionCode="SEM" collectorName="Radula" country="Australia" pageId="5" pageNumber="255" specimenCount="1" stateProvince="Queensland" typeStatus="paratype">
one preserved specimen, 
<quantity id="4CC09B6709764F6FFEE4FC11FE86EDFB" box="[262,325,999,1020]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="255" unit="mm" value="3.0">3 mm</quantity>
long, dissected (
<collectionCode id="ED29AE4709764F6FFE1EFC11FDFBEDFB" box="[508,568,999,1020]" pageId="5" pageNumber="255">SEM</collectionCode>
: 
<collectorName id="26CD535409764F6FFDA3FC11FD55EDFA" box="[577,662,999,1021]" pageId="5" pageNumber="255">Radula</collectorName>
)
</materialsCitation>
. 
<materialsCitation id="3B503CDF09764F6FFD45FC11FDF8EA1D" ID-GBIF-Occurrence="3897190301" collectingDate="2019-11-28" collectionCode="MNCN" country="Australia" pageId="5" pageNumber="255" specimenCode="MNCN 15.05" specimenCount="1" stateProvince="Queensland" typeStatus="paratype">
<specimenCode id="DB9E9EF909764F6FFD45FC11FF13EA1C" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="5" pageNumber="255" type="Museum">MNCN 15.05</specimenCode>
/200146 (
<date id="FF86104209764F6FFEA4FBF3FDF0EA1D" box="[326,563,1029,1051]" pageId="5" pageNumber="255" value="2019-11-28">
<collectingDate id="EFC2E9AA09764F6FFEA4FBF3FDF0EA1D" box="[326,563,1029,1051]" pageId="5" pageNumber="255" value="2019-11-28">28 November 2019</collectingDate>
</date>
)
</materialsCitation>
, one preserved specimen, 
<quantity id="4CC09B6709764F6FFEF9FBD2FEA0EA3E" box="[283,355,1060,1082]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="255" unit="mm" value="5.0">5 mm</quantity>
long. 
<materialsCitation id="3B503CDF09764F6FFE5BFBD2FE8CEA5F" ID-GBIF-Occurrence="3897190352" collectingDate="2019-11-28" collectionCode="MNCN" country="Australia" pageId="5" pageNumber="255" specimenCode="MNCN 15.05" specimenCount="1" stateProvince="Queensland" typeStatus="paratype">
<specimenCode id="DB9E9EF909764F6FFE5BFBD2FD9DEA3D" box="[441,606,1060,1082]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="5" pageNumber="255" type="Museum">MNCN 15.05</specimenCode>
/200147 (
<date id="FF86104209764F6FFD38FBD2FE8BEA5F" pageId="5" pageNumber="255" value="2019-11-28">
<collectingDate id="EFC2E9AA09764F6FFD38FBD2FE8BEA5F" pageId="5" pageNumber="255" value="2019-11-28">28 November 2019</collectingDate>
</date>
)
</materialsCitation>
, 
<materialsCitation id="3B503CDF09764F6FFEB9FBB5FE38EA70" ID-GBIF-Occurrence="3897190349" collectionCode="SEM" collectorName="Radula" country="Australia" pageId="5" pageNumber="255" specimenCount="1" stateProvince="Queensland" typeStatus="paratype">
one preserved specimen, 
<quantity id="4CC09B6709764F6FFD98FBB4FD7FEA5F" box="[634,700,1090,1112]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="255" unit="mm" value="5.0">5 mm</quantity>
long, dissected (
<collectionCode id="ED29AE4709764F6FFEE8FB97FE85EA71" box="[266,326,1121,1142]" pageId="5" pageNumber="255">SEM</collectionCode>
: 
<collectorName id="26CD535409764F6FFEB0FB97FE64EA70" box="[338,423,1121,1143]" pageId="5" pageNumber="255">Radula</collectorName>
, penis)
</materialsCitation>
. 
<materialsCitation id="3B503CDF09764F6FFDE4FB97FEB1EA92" ID-GBIF-Occurrence="3897190324" collectingDate="2019-12-05" collectionCode="MNCN" country="Australia" pageId="5" pageNumber="255" specimenCode="MNCN 15.05" specimenCount="1" stateProvince="Queensland" typeStatus="paratype">
<specimenCode id="DB9E9EF909764F6FFDE4FB97FD5FEA70" box="[518,668,1121,1143]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="5" pageNumber="255" type="Museum">MNCN 15.05</specimenCode>
/200148 (
<date id="FF86104209764F6FFF78FB76FEA9EA92" box="[154,362,1152,1174]" pageId="5" pageNumber="255" value="2019-12-05">
<collectingDate id="EFC2E9AA09764F6FFF78FB76FEA9EA92" box="[154,362,1152,1174]" pageId="5" pageNumber="255" value="2019-12-05">5 December 2019</collectingDate>
</date>
)
</materialsCitation>
<materialsCitation id="3B503CDF09764F6FFE99FB76FDC7EAB2" ID-GBIF-Occurrence="3897190394" collectionCode="SEM" collectorName="Radula" country="Australia" pageId="5" pageNumber="255" specimenCount="1" stateProvince="Queensland" typeStatus="paratype">
one preserved specimen, 
<quantity id="4CC09B6709764F6FFD45FB76FD3BEA92" box="[679,760,1152,1173]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="255" unit="mm" value="10.0">10 mm</quantity>
long, dissected (
<collectionCode id="ED29AE4709764F6FFEB4FB69FE57EAB3" box="[342,404,1183,1204]" pageId="5" pageNumber="255">SEM</collectionCode>
: 
<collectorName id="26CD535409764F6FFE40FB69FE3FEAB2" box="[418,508,1183,1205]" pageId="5" pageNumber="255">Radula</collectorName>
)
</materialsCitation>
. 
<materialsCitation id="3B503CDF09764F6FFDF3FB69FEBDEAD5" ID-GBIF-Occurrence="3897190377" accessionNumber="S72660" collectingDate="2018-05-02" collectionCode="WAM" country="Australia" pageId="5" pageNumber="255" specimenCount="1" stateProvince="Queensland" typeStatus="paratype">
<collectionCode id="ED29AE4709764F6FFDF3FB69FD90EAB3" box="[529,595,1183,1204]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="5" pageNumber="255" type="Museum">WAM</collectionCode>
S72660 (
<figureCitation id="13032A0709764F6FFD24FB69FF5CEAD5" captionStart="Figure 1" captionStartId="4.[163,244,1066,1088]" captionTargetBox="[164,1443,197,1025]" captionTargetId="figure-402@4.[163,1446,195,1026]" captionTargetPageId="4" captionText="Figure 1. Phylogenetic relationships (BI/ML) based on the concatenated mitochondrial (COI and 16S) and nuclear (H3) genes. Purple branches indicate the node supporting the Goniodorididae family. Blue shading represents the genus Murphydoris. Murphydoris species newly sequenced in bold. Colours highlighted in ABGD and bPTP species delimitation analyses represent potential different taxa. Photographs of species from top to bottom: Murphydoris maracabranchia (CASIZ 186120), photographed by P.Palereacio;Murphydoris adusta (WAM S72660), photographed by G.Cobb;Murphydoris puncticulata (WAM S72711), photographed by G. Cobb; Murphydoris cobbi (CASIZ 206795), photographed by T.M. Gosliner." figureDoi="http://doi.org/10.5281/zenodo.7037694" httpUri="https://zenodo.org/record/7037694/files/figure.png" pageId="5" pageNumber="255">Figs 1</figureCitation>
, 
<figureCitation id="13032A0709764F6FFF4CFB4BFF0EEAD5" box="[174,205,1213,1234]" captionStart="Figure 2" captionStartId="6.[163,241,1491,1513]" captionTargetBox="[323,1283,196,1451]" captionTargetId="figure-159@6.[323,1283,195,1451]" captionTargetPageId="6" captionText="Figure 2. Photographs of living animals. A, B, Murphydoris adusta sp. nov., photographed by G. Cobb: A, (WAM S72660); B, (QMMO 85995). C, Murphydoris cobbi sp. nov. (CASIZ 206795), photographed by T. M. Gosliner. D, M. cobbi sp. nov. (QMMO 85979), photographed by G. Cobb. E, Murphydoris maracabranchia sp. nov. (CASIZ 186120), photographed by P. Paleracio. F, G, M. puncticulata sp. nov. photographed by G. Cobb. F, (WAM S72711), G, (WAM S72710)." figureDoi="http://doi.org/10.5281/zenodo.7037696" httpUri="https://zenodo.org/record/7037696/files/figure.png" pageId="5" pageNumber="255">2A</figureCitation>
) (
<date id="FF86104209764F6FFF05FB4BFEB6EAD5" box="[231,373,1213,1235]" pageId="5" pageNumber="255" value="2018-05-02">
<collectingDate id="EFC2E9AA09764F6FFF05FB4BFEB6EAD5" box="[231,373,1213,1235]" pageId="5" pageNumber="255" value="2018-05-02">2 May 2018</collectingDate>
</date>
)
</materialsCitation>
, one preserved specimen, 
<quantity id="4CC09B6709764F6FFD54FB4BFD39EAD4" box="[694,762,1213,1235]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="255" unit="mm" value="5.0">5 mm</quantity>
long. 
<materialsCitation id="3B503CDF09764F6FFF36FB2AFEF9EB28" ID-GBIF-Occurrence="3897190338" collectionCode="SEM" collectorName="Radula" country="Australia" pageId="5" pageNumber="255" specimenCode="QMMO 85986" specimenCount="1" stateProvince="Queensland" typeStatus="paratype">
<specimenCode id="DB9E9EF909764F6FFF36FB2AFEB8EAF5" box="[212,379,1244,1266]" collectionCode="QMMO" pageId="5" pageNumber="255">QMMO 85986</specimenCode>
(
<figureCitation id="13032A0709764F6FFE6EFB2AFE27EAF6" box="[396,484,1244,1266]" captionStart="Figure 4" captionStartId="8.[163,241,1584,1606]" captionTargetBox="[323,1283,196,1544]" captionTargetId="figure-87@8.[323,1283,195,1544]" captionTargetPageId="8" captionText="Figure 4. Scanning electron microscope (SEM) photographs and light microscope photographs (LMP) of A–D, Murphydoris adusta sp. nov. and E–H, Murphydoris cobbi sp. nov. A, frontal view of radula (QMMO 85995) (SEM); B, lateral view of teeth (CASIZ 181290) (SEM); C, detail of masticatory margin of teeth (QMMO 85995) (SEM); D, detail of penial spines (QMMO 85986) (SEM); E, detail of labial cuticle elements (QMMO 85984) (SEM); F, frontal view of radula – detail of rachis, internal and external teeth (QMMO 85983) (SEM); G, detail of internal teeth (QMMO 85983) (SEM); H, detail of penial spines (LMP) (QMMO 85983). Scale bars: A, 50 µm; B, 40 µm; C, 10 µm; D, 10 µm; E, 5µm; F, 30 µm; G, 20 µm; H, 10 µm." figureDoi="http://doi.org/10.5281/zenodo.7037704" httpUri="https://zenodo.org/record/7037704/files/figure.png" pageId="5" pageNumber="255">Fig. 4D</figureCitation>
) (2019), one preserved specimen, 
<quantity id="4CC09B6709764F6FFEEFFB0DFE8EEB17" box="[269,333,1275,1296]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="255" unit="mm" value="4.0">4 mm</quantity>
long, dissected (
<collectionCode id="ED29AE4709764F6FFDECFB0DFD88EB17" box="[526,587,1275,1296]" pageId="5" pageNumber="255">SEM</collectionCode>
: 
<collectorName id="26CD535409764F6FFDB4FB0DFD6FEB16" box="[598,684,1275,1297]" pageId="5" pageNumber="255">Radula</collectorName>
, labial cuticle, penis)
</materialsCitation>
. 
<materialsCitation id="3B503CDF09764F6FFEA5FAEFFD37EB49" ID-GBIF-Occurrence="3897190381" collectionCode="SEM" collectorName="Radula" country="Australia" pageId="5" pageNumber="255" specimenCode="QMMO 85987" specimenCount="1" stateProvince="Queensland" typeStatus="paratype">
<specimenCode id="DB9E9EF909764F6FFEA5FAEFFE2DEB28" box="[327,494,1305,1327]" collectionCode="QMMO" pageId="5" pageNumber="255">QMMO 85987</specimenCode>
(2019), one preserved specimen, 
<quantity id="4CC09B6709764F6FFEF9FACEFEA3EB4A" box="[283,352,1336,1357]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="255" unit="mm" value="3.0">3 mm</quantity>
long, dissected (
<collectionCode id="ED29AE4709764F6FFDDFFACEFDBDEB4A" box="[573,638,1336,1357]" pageId="5" pageNumber="255">SEM</collectionCode>
: 
<collectorName id="26CD535409764F6FFD6CFACEFD28EB49" box="[654,747,1336,1358]" pageId="5" pageNumber="255">Radula</collectorName>
)
</materialsCitation>
. 
<materialsCitation id="3B503CDF09764F6FFF73FAA1FD36EB6A" ID-GBIF-Occurrence="3897190364" box="[145,757,1366,1389]" collectingDate="2019-09-29" collectionCode="QMMO" country="Australia" pageId="5" pageNumber="255" specimenCode="QMMO 85988" specimenCount="1" stateProvince="Queensland" typeStatus="paratype">
<specimenCode id="DB9E9EF909764F6FFF73FAA1FEF4EB6B" box="[145,311,1366,1388]" collectionCode="QMMO" pageId="5" pageNumber="255">QMMO 85988</specimenCode>
(
<date id="FF86104209764F6FFEAAFAA1FDF2EB6B" box="[328,561,1367,1389]" pageId="5" pageNumber="255" value="2019-09-29">
<collectingDate id="EFC2E9AA09764F6FFEAAFAA1FDF2EB6B" box="[328,561,1367,1389]" pageId="5" pageNumber="255" value="2019-09-29">29 September 2019</collectingDate>
</date>
), 
<specimenCount id="9D3EFD0B09764F6FFDA4FAA1FD36EB6A" box="[582,757,1367,1389]" pageId="5" pageNumber="255" type="generic">two specimens</specimenCount>
</materialsCitation>
. 
<materialsCitation id="3B503CDF09764F6FFF73FA83FD37EB8D" ID-GBIF-Occurrence="3897190318" box="[145,756,1397,1419]" collectingDate="2019-12-11" collectionCode="QMMO" country="Australia" pageId="5" pageNumber="255" specimenCode="QMMO 85995" specimenCount="1" stateProvince="Queensland" typeStatus="paratype">
<specimenCode id="DB9E9EF909764F6FFF73FA83FEF4EB8C" box="[145,311,1397,1419]" collectionCode="QMMO" pageId="5" pageNumber="255">QMMO 85995</specimenCode>
(
<figureCitation id="13032A0709764F6FFEABFA83FE67EB8D" box="[329,420,1397,1419]" captionStart="Figure 2" captionStartId="6.[163,241,1491,1513]" captionTargetBox="[323,1283,196,1451]" captionTargetId="figure-159@6.[323,1283,195,1451]" captionTargetPageId="6" captionText="Figure 2. Photographs of living animals. A, B, Murphydoris adusta sp. nov., photographed by G. Cobb: A, (WAM S72660); B, (QMMO 85995). C, Murphydoris cobbi sp. nov. (CASIZ 206795), photographed by T. M. Gosliner. D, M. cobbi sp. nov. (QMMO 85979), photographed by G. Cobb. E, Murphydoris maracabranchia sp. nov. (CASIZ 186120), photographed by P. Paleracio. F, G, M. puncticulata sp. nov. photographed by G. Cobb. F, (WAM S72711), G, (WAM S72710)." figureDoi="http://doi.org/10.5281/zenodo.7037696" httpUri="https://zenodo.org/record/7037696/files/figure.png" pageId="5" pageNumber="255">Figs 2B</figureCitation>
, 
<figureCitation id="13032A0709764F6FFE50FA83FE37EB8D" box="[434,500,1397,1419]" captionStart="Figure 4" captionStartId="8.[163,241,1584,1606]" captionTargetBox="[323,1283,196,1544]" captionTargetId="figure-87@8.[323,1283,195,1544]" captionTargetPageId="8" captionText="Figure 4. Scanning electron microscope (SEM) photographs and light microscope photographs (LMP) of A–D, Murphydoris adusta sp. nov. and E–H, Murphydoris cobbi sp. nov. A, frontal view of radula (QMMO 85995) (SEM); B, lateral view of teeth (CASIZ 181290) (SEM); C, detail of masticatory margin of teeth (QMMO 85995) (SEM); D, detail of penial spines (QMMO 85986) (SEM); E, detail of labial cuticle elements (QMMO 85984) (SEM); F, frontal view of radula – detail of rachis, internal and external teeth (QMMO 85983) (SEM); G, detail of internal teeth (QMMO 85983) (SEM); H, detail of penial spines (LMP) (QMMO 85983). Scale bars: A, 50 µm; B, 40 µm; C, 10 µm; D, 10 µm; E, 5µm; F, 30 µm; G, 20 µm; H, 10 µm." figureDoi="http://doi.org/10.5281/zenodo.7037704" httpUri="https://zenodo.org/record/7037704/files/figure.png" pageId="5" pageNumber="255">4A, C</figureCitation>
) (
<date id="FF86104209764F6FFDECFA83FD2FEB8D" box="[526,748,1397,1419]" pageId="5" pageNumber="255" value="2019-12-11">
<collectingDate id="EFC2E9AA09764F6FFDECFA83FD2FEB8D" box="[526,748,1397,1419]" pageId="5" pageNumber="255" value="2019-12-11">11 December 2019</collectingDate>
</date>
)
</materialsCitation>
, 
<materialsCitation id="3B503CDF09764F6FFF70FA62FE5DEBCE" ID-GBIF-Occurrence="3897190322" collectionCode="SEM" collectorName="Radula" country="Australia" pageId="5" pageNumber="255" specimenCount="1" stateProvince="Queensland" typeStatus="paratype">
one preserved specimen, 
<quantity id="4CC09B6709764F6FFE55FA62FE3BEBAE" box="[439,504,1428,1449]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="255" unit="mm" value="4.0">4 mm</quantity>
long, dissected (
<collectionCode id="ED29AE4709764F6FFD5BFA62FD36EBAE" box="[697,757,1428,1449]" pageId="5" pageNumber="255">SEM</collectionCode>
: 
<collectorName id="26CD535409764F6FFF70FA45FF29EBCE" box="[146,234,1459,1481]" pageId="5" pageNumber="255">Radula</collectorName>
, labial cuticle)
</materialsCitation>
. 
<materialsCitation id="3B503CDF09764F6FFE49FA45FE6AE823" ID-GBIF-Occurrence="3897190332" collectingDate="2008-04-23" collectionCode="T" collectorName="M. Gosliner" country="Philippines" county="Tingloy" latitude="13.761595" location="Aphol's Rock" longLatPrecision="1" longitude="121.383484" municipality="Maricaban Island" pageId="5" pageNumber="255" specimenCode="CASIZ 177775" specimenCount="1" stateProvince="Batangas" typeStatus="paratype">
<specimenCode id="DB9E9EF909764F6FFE49FA45FD9AEBCF" box="[427,601,1458,1480]" collectionCode="CASIZ" pageId="5" pageNumber="255">CASIZ 177775</specimenCode>
(
<collectingCountry id="F32F761209764F6FFD88FA45FD37EBCE" box="[618,756,1459,1481]" name="Philippines" pageId="5" pageNumber="255">Philippines</collectingCountry>
, 
<collectingRegion id="49FCF86009764F6FFF73FA24FE4BEBE0" box="[145,392,1489,1511]" country="Philippines" name="Batangas" pageId="5" pageNumber="255">Batangas Province</collectingRegion>
, 
<collectingCounty id="62E64E0E09764F6FFE7BFA27FE3AEBE0" box="[409,505,1489,1511]" pageId="5" pageNumber="255">Tingloy</collectingCounty>
, 
<collectingMunicipality id="6BE3ACF809764F6FFDE9FA27FD31EBE0" box="[523,754,1489,1511]" pageId="5" pageNumber="255">Maricaban Island</collectingMunicipality>
, 
<location id="8EE7605909764F6FFF73FA06FEE7E801" LSID="urn:lsid:plazi:treatment:0391879409774F63FC52FA06FD71EA31:8EE7605909764F6FFF73FA06FEE7E801" box="[145,292,1520,1542]" country="Philippines" county="Tingloy" latitude="13.761595" longLatPrecision="1" longitude="121.383484" municipality="Maricaban Island" name="Aphol's Rock" pageId="5" pageNumber="255" stateProvince="Batangas">Aphol’s Rock</location>
, 
<geoCoordinate id="EE0C504509764F6FFECCFA06FE16E802" box="[302,469,1519,1542]" degrees="13" direction="north" minutes="45" orientation="latitude" pageId="5" pageNumber="255" precision="1" seconds="41.74" value="13.761595">13°45′41.74′′N</geoCoordinate>
, 
<geoCoordinate id="EE0C504509764F6FFE3DFA06FD47E802" box="[479,644,1519,1542]" degrees="121" direction="east" minutes="23" orientation="longitude" pageId="5" pageNumber="255" precision="1" seconds="0.56" value="121.383484">121°23′0.56′′E</geoCoordinate>
; coll. 
<collectionCode id="ED29AE4709764F6FFD20FA06FD10E802" box="[706,723,1520,1541]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="255">T</collectionCode>
. 
<collectorName id="26CD535409764F6FFD3EFA06FF30E822" pageId="5" pageNumber="255">M. Gosliner</collectorName>
, 
<date id="FF86104209764F6FFF1DF9F9FE62E823" box="[255,417,1551,1573]" pageId="5" pageNumber="255" value="2008-04-23">
<collectingDate id="EFC2E9AA09764F6FFF1DF9F9FE62E823" box="[255,417,1551,1573]" pageId="5" pageNumber="255" value="2008-04-23">23 April 2008</collectingDate>
</date>
)
</materialsCitation>
. 
<materialsCitation id="3B503CDF09764F6FFE57F9F9FDC4E887" ID-GBIF-Occurrence="3897190392" collectingDate="2011-05-04" collectionCode="CASIZ" collectorName="D. Raymundo" country="Philippines" county="Calumpan Peninsula" latitude="13.69902" location="Murals" longLatPrecision="1" longitude="120.8827" municipality="Mabini" pageId="5" pageNumber="255" specimenCode="CASIZ 186117" specimenCount="1" stateProvince="Batangas" typeStatus="paratype">
<specimenCode id="DB9E9EF909764F6FFE57F9F9FD9DE822" box="[437,606,1551,1573]" collectionCode="CASIZ" pageId="5" pageNumber="255">CASIZ 186117</specimenCode>
(
<collectingCountry id="F32F761209764F6FFD8FF9F9FD37E822" box="[621,756,1551,1573]" name="Philippines" pageId="5" pageNumber="255">Philippines</collectingCountry>
, 
<collectingRegion id="49FCF86009764F6FFF73F9D8FEB9E844" box="[145,378,1581,1603]" country="Philippines" name="Batangas" pageId="5" pageNumber="255">Batangas Province</collectingRegion>
, 
<collectingCounty id="62E64E0E09764F6FFE6BF9DBFD4EE844" box="[393,653,1581,1603]" pageId="5" pageNumber="255">Calumpan Peninsula</collectingCounty>
, 
<collectingMunicipality id="6BE3ACF809764F6FFD7EF9DBFD37E844" box="[668,756,1581,1603]" pageId="5" pageNumber="255">Mabini</collectingMunicipality>
, 
<location id="8EE7605909764F6FFF73F9BAFF25E865" LSID="urn:lsid:plazi:treatment:0391879409774F63FC52FA06FD71EA31:8EE7605909764F6FFF73F9BAFF25E865" box="[145,230,1612,1634]" country="Philippines" county="Calumpan Peninsula" latitude="13.69902" longLatPrecision="1" longitude="120.8827" municipality="Mabini" name="Murals" pageId="5" pageNumber="255" stateProvince="Batangas">Murals</location>
dive site, 
<geoCoordinate id="EE0C504509764F6FFE80F9BAFDDDE866" box="[354,542,1611,1634]" degrees="13" direction="north" minutes="41" orientation="latitude" pageId="5" pageNumber="255" precision="1" seconds="56.472" value="13.69902">13°41′56.472′′N</geoCoordinate>
, 
<geoCoordinate id="EE0C504509764F6FFDC9F9BAFD37E866" box="[555,756,1611,1634]" degrees="120" direction="east" minutes="52" orientation="longitude" pageId="5" pageNumber="255" precision="1" seconds="57.719" value="120.8827">120°52′57.719′′E</geoCoordinate>
; coll. 
<collectorName id="26CD535409764F6FFF25F99DFEA4E886" box="[199,359,1643,1665]" pageId="5" pageNumber="255">D. Raymundo</collectorName>
, 
<date id="FF86104209764F6FFE91F99DFE3CE887" box="[371,511,1643,1664]" pageId="5" pageNumber="255" value="2011-05-04">
<collectingDate id="EFC2E9AA09764F6FFE91F99DFE3CE887" box="[371,511,1643,1664]" pageId="5" pageNumber="255" value="2011-05-04">4 May 2011</collectingDate>
</date>
)
</materialsCitation>
. 
<materialsCitation id="3B503CDF09764F6FFDF1F99DFDEAE8DB" ID-GBIF-Occurrence="3897190366" collectingDate="2009-05-21" collectionCode="T" collectorName="M. Gosliner" country="Philippines" county="Luzon" latitude="13.685917" location="Mainit Bubbles" longLatPrecision="1" longitude="120.89522" municipality="Mabini" pageId="5" pageNumber="255" specimenCode="CASIZ 181290" specimenCount="1" stateProvince="Batangas" typeStatus="paratype">
<specimenCode id="DB9E9EF909764F6FFDF1F99DFD7FE887" box="[531,700,1643,1664]" collectionCode="CASIZ" pageId="5" pageNumber="255">CASIZ 181290</specimenCode>
(
<figureCitation id="13032A0709764F6FFD2FF99DFF72E899" captionStart="Figure 4" captionStartId="8.[163,241,1584,1606]" captionTargetBox="[323,1283,196,1544]" captionTargetId="figure-87@8.[323,1283,195,1544]" captionTargetPageId="8" captionText="Figure 4. Scanning electron microscope (SEM) photographs and light microscope photographs (LMP) of A–D, Murphydoris adusta sp. nov. and E–H, Murphydoris cobbi sp. nov. A, frontal view of radula (QMMO 85995) (SEM); B, lateral view of teeth (CASIZ 181290) (SEM); C, detail of masticatory margin of teeth (QMMO 85995) (SEM); D, detail of penial spines (QMMO 85986) (SEM); E, detail of labial cuticle elements (QMMO 85984) (SEM); F, frontal view of radula – detail of rachis, internal and external teeth (QMMO 85983) (SEM); G, detail of internal teeth (QMMO 85983) (SEM); H, detail of penial spines (LMP) (QMMO 85983). Scale bars: A, 50 µm; B, 40 µm; C, 10 µm; D, 10 µm; E, 5µm; F, 30 µm; G, 20 µm; H, 10 µm." figureDoi="http://doi.org/10.5281/zenodo.7037704" httpUri="https://zenodo.org/record/7037704/files/figure.png" pageId="5" pageNumber="255">Fig. 4B</figureCitation>
) (
<collectingCountry id="F32F761209764F6FFF2BF97FFE92E898" box="[201,337,1673,1695]" name="Philippines" pageId="5" pageNumber="255">Philippines</collectingCountry>
, 
<collectingCounty id="62E64E0E09764F6FFEBFF97CFE64E898" box="[349,423,1674,1695]" pageId="5" pageNumber="255">Luzon</collectingCounty>
, 
<collectingRegion id="49FCF86009764F6FFE56F97CFD51E898" box="[436,658,1673,1695]" country="Philippines" name="Batangas" pageId="5" pageNumber="255">Batangas Province</collectingRegion>
, 
<collectingMunicipality id="6BE3ACF809764F6FFD7DF97FFD37E898" box="[671,756,1673,1695]" pageId="5" pageNumber="255">Mabini</collectingMunicipality>
, 
<location id="8EE7605909764F6FFF73F95EFE86E8B9" LSID="urn:lsid:plazi:treatment:0391879409774F63FC52FA06FD71EA31:8EE7605909764F6FFF73F95EFE86E8B9" box="[145,325,1704,1726]" country="Philippines" county="Luzon" latitude="13.685917" longLatPrecision="1" longitude="120.89522" municipality="Mabini" name="Mainit Bubbles" pageId="5" pageNumber="255" stateProvince="Batangas">Mainit Bubbles</location>
dive site, 
<geoCoordinate id="EE0C504509764F6FFE5FF95EFD8EE8BA" box="[445,589,1703,1726]" degrees="13" direction="north" minutes="41.155" orientation="latitude" pageId="5" pageNumber="255" precision="1" value="13.685917">13°41.155′N</geoCoordinate>
, 
<geoCoordinate id="EE0C504509764F6FFDBBF95EFD36E8BA" box="[601,757,1703,1726]" degrees="120" direction="east" minutes="53.713" orientation="longitude" pageId="5" pageNumber="255" precision="1" value="120.89522">120°53.713′E</geoCoordinate>
; coll. 
<collectionCode id="ED29AE4709764F6FFF2BF931FF18E8DB" box="[201,219,1735,1756]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="255">T</collectionCode>
. 
<collectorName id="26CD535409764F6FFF05F931FEB7E8DA" box="[231,372,1735,1757]" pageId="5" pageNumber="255">M. Gosliner</collectorName>
, 
<date id="FF86104209764F6FFE60F931FDE2E8DB" box="[386,545,1735,1756]" pageId="5" pageNumber="255" value="2009-05-21">
<collectingDate id="EFC2E9AA09764F6FFE60F931FDE2E8DB" box="[386,545,1735,1756]" pageId="5" pageNumber="255" value="2009-05-21">21 May 2009</collectingDate>
</date>
)
</materialsCitation>
, 
<materialsCitation id="3B503CDF09764F6FFDD5F931FF32E8FC" ID-GBIF-Occurrence="3897190310" collectionCode="SEM" collectorName="Radula" country="Philippines" pageId="5" pageNumber="255" specimenCount="1" stateProvince="Batangas" typeStatus="paratype">
dissected (
<collectionCode id="ED29AE4709764F6FFD55F931FD36E8DB" box="[695,757,1735,1756]" pageId="5" pageNumber="255">SEM</collectionCode>
: 
<collectorName id="26CD535409764F6FFF71F913FF2AE8FC" box="[147,233,1765,1787]" pageId="5" pageNumber="255">Radula</collectorName>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF47660A09764F6FFF73FF32FB28EEDD" ID-Table-UUID="DF47660A09764F6FFF73FF32FB28EEDD" box="[145,1259,196,218]" httpUri="http://table.plazi.org/id/DF47660A09764F6FFF73FF32FB28EEDD" pageId="5" pageNumber="255" startId="5.[145,209,196,217]" targetBox="[145,1423,254,571]" targetIsTable="true" targetPageId="5">
<paragraph id="8B87368209764F6FFF73FF32FB28EEDD" blockId="5.[145,1259,196,218]" box="[145,1259,196,218]" pageId="5" pageNumber="255">
<emphasis id="B94CEA9009764F6FFF73FF32FF28EEDE" bold="true" box="[145,235,196,217]" pageId="5" pageNumber="255">Table 2.</emphasis>
<emphasis id="B94CEA9009764F6FFF14FF32FEE2EEDD" box="[246,289,196,218]" italics="true" pageId="5" pageNumber="255">COI</emphasis>
and 16S gene pairwise uncorrected 
<emphasis id="B94CEA9009764F6FFD7EFF32FD6AEEDE" box="[668,681,196,217]" italics="true" pageId="5" pageNumber="255">p</emphasis>
-distances (%) amongst the new species of 
<taxonomicName id="4C384D0109764F6FFB81FF32FB28EEDD" authorityName="Sigurdson" authorityYear="1991" box="[1123,1259,196,218]" class="Gastropoda" family="Goniodorididae" genus="Murphydoris" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="255" phylum="Mollusca" rank="genus">Murphydoris</taxonomicName>
</paragraph>
</caption>
<paragraph id="8B87368209764F6FFE48FF08FA86EC3C" pageId="5" pageNumber="255">
<table id="F938C4220976B095FF73FF08FA4CEC3C" box="[145,1423,254,571]" gridcols="6" gridrows="10" pageId="5" pageNumber="255">
<tr id="350834C00976B095FF73FF08FA4CEF13" box="[145,1423,254,276]" gridrow="0" pageId="5" pageNumber="255" rowspan-0="1">
<th id="76D95DBC0976B095FE48FF08FDBBEF13" box="[426,632,254,276]" gridcol="1" gridrow="0" pageId="5" pageNumber="255">Species</th>
<th id="76D95DBC0976B095FD49FF08FCF0EF13" box="[683,819,254,276]" gridcol="2" gridrow="0" pageId="5" pageNumber="255">
<taxonomicName id="4C384D0109764F6FFD49FF08FCF0EF13" authorityName="Sigurdson" authorityYear="1991" box="[683,819,254,276]" class="Gastropoda" family="Goniodorididae" genus="Murphydoris" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="255" phylum="Mollusca" rank="genus">
<emphasis id="B94CEA9009764F6FFD49FF08FCF0EF13" box="[683,819,254,276]" italics="true" pageId="5" pageNumber="255">Murphydoris</emphasis>
</taxonomicName>
</th>
<th id="76D95DBC0976B095FC84FF08FC2DEF13" box="[870,1006,254,276]" gridcol="3" gridrow="0" pageId="5" pageNumber="255">
<taxonomicName id="4C384D0109764F6FFC84FF08FC2DEF13" authorityName="Sigurdson" authorityYear="1991" box="[870,1006,254,276]" class="Gastropoda" family="Goniodorididae" genus="Murphydoris" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="255" phylum="Mollusca" rank="genus">
<emphasis id="B94CEA9009764F6FFC84FF08FC2DEF13" box="[870,1006,254,276]" italics="true" pageId="5" pageNumber="255">Murphydoris</emphasis>
</taxonomicName>
</th>
<th id="76D95DBC0976B095FBC3FF08FB0DEF13" box="[1057,1230,254,276]" gridcol="4" gridrow="0" pageId="5" pageNumber="255">
<taxonomicName id="4C384D0109764F6FFBC3FF08FB6AEF13" authorityName="Sigurdson" authorityYear="1991" box="[1057,1193,254,276]" class="Gastropoda" family="Goniodorididae" genus="Murphydoris" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="255" phylum="Mollusca" rank="genus">
<emphasis id="B94CEA9009764F6FFBC3FF08FB6AEF13" box="[1057,1193,254,276]" italics="true" pageId="5" pageNumber="255">Murphydoris</emphasis>
</taxonomicName>
</th>
<th id="76D95DBC0976B095FAE5FF08FA4CEF13" box="[1287,1423,254,276]" gridcol="5" gridrow="0" pageId="5" pageNumber="255">
<emphasis id="B94CEA9009764F6FFAE5FF08FA4CEF13" box="[1287,1423,254,276]" italics="true" pageId="5" pageNumber="255">Murphydoris</emphasis>
</th>
</tr>
<tr id="350834C00976B095FF73FEEDFA4CEF36" box="[145,1423,283,305]" gridrow="1" pageId="5" pageNumber="255" rowspan-0="1" rowspan-1="1">
<td id="76D95DBC0976B095FD49FEEDFCF0EF36" box="[683,819,283,305]" gridcol="2" gridrow="1" pageId="5" pageNumber="255">
<emphasis id="B94CEA9009764F6FFD49FEEDFD30EF36" box="[683,755,283,305]" italics="true" pageId="5" pageNumber="255">adusta</emphasis>
</td>
<td id="76D95DBC0976B095FC84FEEDFC2DEF36" box="[870,1006,283,305]" gridcol="3" gridrow="1" pageId="5" pageNumber="255">
<taxonomicName id="4C384D0109764F6FFC84FEEDFC5FEF36" box="[870,924,283,305]" class="Gastropoda" family="Goniodorididae" genus="Murphydoris" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="255" phylum="Mollusca" rank="species" species="cobbi">
<emphasis id="B94CEA9009764F6FFC84FEEDFC5FEF36" box="[870,924,283,305]" italics="true" pageId="5" pageNumber="255">cobbi</emphasis>
</taxonomicName>
</td>
<td id="76D95DBC0976B095FBC3FEEDFB0DEF36" box="[1057,1230,283,305]" gridcol="4" gridrow="1" pageId="5" pageNumber="255">
<taxonomicName id="4C384D0109764F6FFBC3FEEDFB0DEF36" box="[1057,1230,283,305]" class="Gastropoda" family="Goniodorididae" genus="Murphydoris" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="255" phylum="Mollusca" rank="species" species="maracabranchia">
<emphasis id="B94CEA9009764F6FFBC3FEEDFB0DEF36" box="[1057,1230,283,305]" italics="true" pageId="5" pageNumber="255">maracabranchia</emphasis>
</taxonomicName>
</td>
<td id="76D95DBC0976B095FAE5FEEDFA4CEF36" box="[1287,1423,283,305]" gridcol="5" gridrow="1" pageId="5" pageNumber="255">
<taxonomicName id="4C384D0109764F6FFAE5FEEDFA48EF36" box="[1287,1419,283,305]" class="Gastropoda" family="Goniodorididae" genus="Murphydoris" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="255" phylum="Mollusca" rank="species" species="puncticulata">
<emphasis id="B94CEA9009764F6FFAE5FEEDFA48EF36" box="[1287,1419,283,305]" italics="true" pageId="5" pageNumber="255">puncticulata</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="350834C00976B095FF73FEA4FA4CEF6F" box="[145,1423,338,360]" gridrow="2" pageId="5" pageNumber="255">
<th id="76D95DBC0976B095FF73FEA4FEB4EF6F" box="[145,375,338,360]" gridcol="0" gridrow="2" pageId="5" pageNumber="255">
<emphasis id="B94CEA9009764F6FFF73FEA4FEB4EF6F" bold="true" box="[145,375,338,360]" pageId="5" pageNumber="255">
<emphasis id="B94CEA9009764F6FFF73FEA4FF15EF6F" box="[145,214,338,360]" italics="true" pageId="5" pageNumber="255">COI P</emphasis>
-distances (%)
</emphasis>
</th>
<td id="76D95DBC0976B095FE48FEA4FDBBEF6F" box="[426,632,338,360]" gridcol="1" gridrow="2" pageId="5" pageNumber="255">
<taxonomicName id="4C384D0109764F6FFE48FEA5FDD0EF6F" authorityName="Paz-Sedano &amp; Smirnoff &amp; Candás &amp; Gosliner &amp; Pola" authorityYear="2022" box="[426,531,338,360]" class="Gastropoda" family="Goniodorididae" genus="Murphydoris" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="255" phylum="Mollusca" rank="species" species="adusta">
<emphasis id="B94CEA9009764F6FFE48FEA5FDD0EF6F" box="[426,531,338,360]" italics="true" pageId="5" pageNumber="255">M. adusta</emphasis>
</taxonomicName>
</td>
<td id="76D95DBC0976B095FD49FEA4FCF0EF6F" box="[683,819,338,360]" gridcol="2" gridrow="2" pageId="5" pageNumber="255">-</td>
<td id="76D95DBC0976B095FC84FEA4FC2DEF6F" box="[870,1006,338,360]" gridcol="3" gridrow="2" pageId="5" pageNumber="255">-</td>
<td id="76D95DBC0976B095FBC3FEA4FB0DEF6F" box="[1057,1230,338,360]" gridcol="4" gridrow="2" pageId="5" pageNumber="255">-</td>
<td id="76D95DBC0976B095FAE5FEA4FA4CEF6F" box="[1287,1423,338,360]" gridcol="5" gridrow="2" pageId="5" pageNumber="255">-</td>
</tr>
<tr id="350834C00976B095FF73FE86FA4CEF81" box="[145,1423,368,390]" gridrow="3" pageId="5" pageNumber="255" rowspan-0="1">
<td id="76D95DBC0976B095FE48FE86FDBBEF81" box="[426,632,368,390]" gridcol="1" gridrow="3" pageId="5" pageNumber="255">
<taxonomicName id="4C384D0109764F6FFE48FE87FDC2EF81" box="[426,513,368,390]" class="Gastropoda" family="Goniodorididae" genus="Murphydoris" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="255" phylum="Mollusca" rank="species" species="cobbi">
<emphasis id="B94CEA9009764F6FFE48FE87FDC2EF81" box="[426,513,368,390]" italics="true" pageId="5" pageNumber="255">M. cobbi</emphasis>
</taxonomicName>
</td>
<td id="76D95DBC0976B095FD49FE86FCF0EF81" box="[683,819,368,390]" gridcol="2" gridrow="3" pageId="5" pageNumber="255">-</td>
<td id="76D95DBC0976B095FC84FE86FC2DEF81" box="[870,1006,368,390]" gridcol="3" gridrow="3" pageId="5" pageNumber="255">0–0.8</td>
<td id="76D95DBC0976B095FBC3FE86FB0DEF81" box="[1057,1230,368,390]" gridcol="4" gridrow="3" pageId="5" pageNumber="255">21.4–21.9</td>
<td id="76D95DBC0976B095FAE5FE86FA4CEF81" box="[1287,1423,368,390]" gridcol="5" gridrow="3" pageId="5" pageNumber="255">10.7–11.2</td>
</tr>
<tr id="350834C00976B095FF73FE78FA4CEFA3" box="[145,1423,398,420]" gridrow="4" pageId="5" pageNumber="255" rowspan-0="1">
<td id="76D95DBC0976B095FE48FE78FDBBEFA3" box="[426,632,398,420]" gridcol="1" gridrow="4" pageId="5" pageNumber="255">
<taxonomicName id="4C384D0109764F6FFE48FE79FDBBEFA3" box="[426,632,398,420]" class="Gastropoda" family="Goniodorididae" genus="Murphydoris" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="255" phylum="Mollusca" rank="species" species="maracabranchia">
<emphasis id="B94CEA9009764F6FFE48FE79FDBBEFA3" box="[426,632,398,420]" italics="true" pageId="5" pageNumber="255">M. maracabranchia</emphasis>
</taxonomicName>
</td>
<td id="76D95DBC0976B095FD49FE78FCF0EFA3" box="[683,819,398,420]" gridcol="2" gridrow="4" pageId="5" pageNumber="255">-</td>
<td id="76D95DBC0976B095FC84FE78FC2DEFA3" box="[870,1006,398,420]" gridcol="3" gridrow="4" pageId="5" pageNumber="255">21.4–21.9</td>
<td id="76D95DBC0976B095FBC3FE78FB0DEFA3" box="[1057,1230,398,420]" gridcol="4" gridrow="4" pageId="5" pageNumber="255">0</td>
<td id="76D95DBC0976B095FAE5FE78FA4CEFA3" box="[1287,1423,398,420]" gridcol="5" gridrow="4" pageId="5" pageNumber="255">19.4</td>
</tr>
<tr id="350834C00976B095FF73FE5BFA4CEFC4" box="[145,1423,429,451]" gridrow="5" pageId="5" pageNumber="255" rowspan-0="1">
<td id="76D95DBC0976B095FE48FE5BFDBBEFC4" box="[426,632,429,451]" gridcol="1" gridrow="5" pageId="5" pageNumber="255">
<taxonomicName id="4C384D0109764F6FFE48FE5BFD8CEFC4" box="[426,591,429,451]" class="Gastropoda" family="Goniodorididae" genus="Murphydoris" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="255" phylum="Mollusca" rank="species" species="puncticulata">
<emphasis id="B94CEA9009764F6FFE48FE5BFD8CEFC4" box="[426,591,429,451]" italics="true" pageId="5" pageNumber="255">M. puncticulata</emphasis>
</taxonomicName>
</td>
<td id="76D95DBC0976B095FD49FE5BFCF0EFC4" box="[683,819,429,451]" gridcol="2" gridrow="5" pageId="5" pageNumber="255">-</td>
<td id="76D95DBC0976B095FC84FE5BFC2DEFC4" box="[870,1006,429,451]" gridcol="3" gridrow="5" pageId="5" pageNumber="255">10.7–11.2</td>
<td id="76D95DBC0976B095FBC3FE5BFB0DEFC4" box="[1057,1230,429,451]" gridcol="4" gridrow="5" pageId="5" pageNumber="255">19.4</td>
<td id="76D95DBC0976B095FAE5FE5BFA4CEFC4" box="[1287,1423,429,451]" gridcol="5" gridrow="5" pageId="5" pageNumber="255">0–0.5</td>
</tr>
<tr id="350834C00976B095FF73FE3CFA4CEFE6" box="[145,1423,458,481]" gridrow="6" pageId="5" pageNumber="255">
<th id="76D95DBC0976B095FF73FE3CFEB4EFE6" box="[145,375,458,481]" gridcol="0" gridrow="6" pageId="5" pageNumber="255">
<emphasis id="B94CEA9009764F6FFF73FE3CFEB1EFE7" bold="true" box="[145,370,458,480]" pageId="5" pageNumber="255">
16S 
<emphasis id="B94CEA9009764F6FFF22FE3DFF12EFE7" bold="true" box="[192,209,459,480]" italics="true" pageId="5" pageNumber="255">P</emphasis>
-distances (%)
</emphasis>
</th>
<td id="76D95DBC0976B095FE48FE3CFDBBEFE6" box="[426,632,458,481]" gridcol="1" gridrow="6" pageId="5" pageNumber="255">
<taxonomicName id="4C384D0109764F6FFE48FE3DFDD0EFE6" authorityName="Paz-Sedano &amp; Smirnoff &amp; Candás &amp; Gosliner &amp; Pola" authorityYear="2022" box="[426,531,459,481]" class="Gastropoda" family="Goniodorididae" genus="Murphydoris" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="255" phylum="Mollusca" rank="species" species="adusta">
<emphasis id="B94CEA9009764F6FFE48FE3DFDD0EFE6" box="[426,531,459,481]" italics="true" pageId="5" pageNumber="255">M. adusta</emphasis>
</taxonomicName>
</td>
<td id="76D95DBC0976B095FD49FE3CFCF0EFE6" box="[683,819,458,481]" gridcol="2" gridrow="6" pageId="5" pageNumber="255">0.6–0.9</td>
<td id="76D95DBC0976B095FC84FE3CFC2DEFE6" box="[870,1006,458,481]" gridcol="3" gridrow="6" pageId="5" pageNumber="255">14.5–15.1</td>
<td id="76D95DBC0976B095FBC3FE3CFB0DEFE6" box="[1057,1230,458,481]" gridcol="4" gridrow="6" pageId="5" pageNumber="255">18–18.6</td>
<td id="76D95DBC0976B095FAE5FE3CFA4CEFE6" box="[1287,1423,458,481]" gridcol="5" gridrow="6" pageId="5" pageNumber="255">13.2–14.5</td>
</tr>
<tr id="350834C00976B095FF73FE1FFA4CEFF8" box="[145,1423,489,511]" gridrow="7" pageId="5" pageNumber="255" rowspan-0="1">
<td id="76D95DBC0976B095FE48FE1FFDBBEFF8" box="[426,632,489,511]" gridcol="1" gridrow="7" pageId="5" pageNumber="255">
<taxonomicName id="4C384D0109764F6FFE48FE1FFDC2EFF8" box="[426,513,489,511]" class="Gastropoda" family="Goniodorididae" genus="Murphydoris" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="255" phylum="Mollusca" rank="species" species="cobbi">
<emphasis id="B94CEA9009764F6FFE48FE1FFDC2EFF8" box="[426,513,489,511]" italics="true" pageId="5" pageNumber="255">M. cobbi</emphasis>
</taxonomicName>
</td>
<td id="76D95DBC0976B095FD49FE1FFCF0EFF8" box="[683,819,489,511]" gridcol="2" gridrow="7" pageId="5" pageNumber="255">14.5–15.1</td>
<td id="76D95DBC0976B095FC84FE1FFC2DEFF8" box="[870,1006,489,511]" gridcol="3" gridrow="7" pageId="5" pageNumber="255">0–0.6</td>
<td id="76D95DBC0976B095FBC3FE1FFB0DEFF8" box="[1057,1230,489,511]" gridcol="4" gridrow="7" pageId="5" pageNumber="255">17.7–18.6</td>
<td id="76D95DBC0976B095FAE5FE1FFA4CEFF8" box="[1287,1423,489,511]" gridcol="5" gridrow="7" pageId="5" pageNumber="255">7.9–8.2</td>
</tr>
<tr id="350834C00976B095FF73FDF1FA4CEC1A" box="[145,1423,519,541]" gridrow="8" pageId="5" pageNumber="255" rowspan-0="1">
<td id="76D95DBC0976B095FE48FDF1FDBBEC1A" box="[426,632,519,541]" gridcol="1" gridrow="8" pageId="5" pageNumber="255">
<taxonomicName id="4C384D0109764F6FFE48FDF1FDBBEC1A" box="[426,632,519,541]" class="Gastropoda" family="Goniodorididae" genus="Murphydoris" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="255" phylum="Mollusca" rank="species" species="maracabranchia">
<emphasis id="B94CEA9009764F6FFE48FDF1FDBBEC1A" box="[426,632,519,541]" italics="true" pageId="5" pageNumber="255">M. maracabranchia</emphasis>
</taxonomicName>
</td>
<td id="76D95DBC0976B095FD49FDF1FCF0EC1A" box="[683,819,519,541]" gridcol="2" gridrow="8" pageId="5" pageNumber="255">18–18.6</td>
<td id="76D95DBC0976B095FC84FDF1FC2DEC1A" box="[870,1006,519,541]" gridcol="3" gridrow="8" pageId="5" pageNumber="255">17.7–18.6</td>
<td id="76D95DBC0976B095FBC3FDF1FB0DEC1A" box="[1057,1230,519,541]" gridcol="4" gridrow="8" pageId="5" pageNumber="255">0–0.6</td>
<td id="76D95DBC0976B095FAE5FDF1FA4CEC1A" box="[1287,1423,519,541]" gridcol="5" gridrow="8" pageId="5" pageNumber="255">18.9–19.2</td>
</tr>
<tr id="350834C00976B095FF73FDD3FA4CEC3C" box="[145,1423,549,571]" gridrow="9" pageId="5" pageNumber="255" rowspan-0="1">
<td id="76D95DBC0976B095FE48FDD3FDBBEC3C" box="[426,632,549,571]" gridcol="1" gridrow="9" pageId="5" pageNumber="255">
<taxonomicName id="4C384D0109764F6FFE48FDD3FD8CEC3C" box="[426,591,549,571]" class="Gastropoda" family="Goniodorididae" genus="Murphydoris" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="255" phylum="Mollusca" rank="species" species="puncticulata">
<emphasis id="B94CEA9009764F6FFE48FDD3FD8CEC3C" box="[426,591,549,571]" italics="true" pageId="5" pageNumber="255">M. puncticulata</emphasis>
</taxonomicName>
</td>
<td id="76D95DBC0976B095FD49FDD3FCF0EC3C" box="[683,819,549,571]" gridcol="2" gridrow="9" pageId="5" pageNumber="255">13.2–14.5</td>
<td id="76D95DBC0976B095FC84FDD3FC2DEC3C" box="[870,1006,549,571]" gridcol="3" gridrow="9" pageId="5" pageNumber="255">7.9–8.2</td>
<td id="76D95DBC0976B095FBC3FDD3FB0DEC3C" box="[1057,1230,549,571]" gridcol="4" gridrow="9" pageId="5" pageNumber="255">18.9–19.2</td>
<td id="76D95DBC0976B095FAE5FDD3FA4CEC3C" box="[1287,1423,549,571]" gridcol="5" gridrow="9" pageId="5" pageNumber="255">0–0.9</td>
</tr>
</table>
</paragraph>
<subSubSection id="C322650909764F6FFF73F8C7FBA5ED43" pageId="5" pageNumber="255" type="distribution">
<paragraph id="8B87368209764F6FFF73F8C7FBA5ED43" blockId="5.[145,761,1841,1894]" lastBlockId="5.[809,1426,630,836]" pageId="5" pageNumber="255">
<emphasis id="B94CEA9009764F6FFF73F8C7FEEEE941" box="[145,301,1841,1862]" italics="true" pageId="5" pageNumber="255">Distribution:</emphasis>
Indian Ocean to 
<collectingCountry id="F32F761209764F6FFDF0F8C7FD4FE940" box="[530,652,1841,1863]" name="Indonesia" pageId="5" pageNumber="255">Indonesia</collectingCountry>
, western Pacific Ocean; 
<collectingCountry id="F32F761209764F6FFED1F8A6FE7DE961" box="[307,446,1872,1894]" name="Madagascar" pageId="5" pageNumber="255">Madagascar</collectingCountry>
(
<bibRefCitation id="EFA94B7309764F6FFE29F8A6FD6AE961" author="Gosliner TM &amp; Valdes A &amp; Behrens D" box="[459,681,1872,1894]" pageId="5" pageNumber="255" refId="ref14453" refString="Gosliner TM, Valdes A, Behrens D. 2015. Nudibranch &amp; sea slug identification - Indo-Pacific. Jacksonville, Florida: New World Publications." type="book" year="2015">
Gosliner 
<emphasis id="B94CEA9009764F6FFDD3F8A7FD9DE962" box="[561,606,1872,1894]" italics="true" pageId="5" pageNumber="255">et al</emphasis>
., 2015
</bibRefCitation>
, 
<bibRefCitation id="EFA94B7309764F6FFD56F8A6FD28E962" author="Gosliner TM &amp; Valdes A &amp; Behrens D" box="[692,747,1872,1893]" pageId="5" pageNumber="255" refId="ref14480" refString="Gosliner TM, Valdes A, Behrens D. 2018. Nudibranch &amp; sea slug identification - Indo-Pacific. Jacksonville, Florida: New World Publications." type="book" year="2018">2018</bibRefCitation>
); 
<collectingCountry id="F32F761209764F6FFCCBFD80FC5FEC8B" box="[809,924,630,652]" name="Indonesia" pageId="5" pageNumber="255">Indonesia</collectingCountry>
: 
<collectingCountry id="F32F761209764F6FFC44FD80FC2BEC8B" box="[934,1000,630,652]" name="Palau" pageId="5" pageNumber="255">Palau</collectingCountry>
Sangeang (
<bibRefCitation id="EFA94B7309764F6FFB8DFD80FAE3EC8C" author="Anderson J" box="[1135,1312,630,652]" pageId="5" pageNumber="255" pagination="6" refId="ref13836" refString="Anderson J. 2021. Goniodoridella sp 6. In: Indonesia nudibranchs &amp; sea slugs. Available at: http: // www. nudibranch. org / Indonesia % 20 Sea % 20 Slugs / html / nudibranchs / goniodoridella-sp 6 - 01. html" type="book chapter" year="2021">Anderson, 2021</bibRefCitation>
) and Bali (
<bibRefCitation id="EFA94B7309764F6FFCD0FD62FBA1ECAE" author="Debelius H &amp; Kuiter RH" box="[818,1122,660,682]" pageId="5" pageNumber="255" refId="ref14257" refString="Debelius H, Kuiter RH. 2007. Nudibranchs of the world. Frankfurt: IKAN." type="book" year="2007">Debelius &amp; Kuiter, 2007</bibRefCitation>
); 
<collectingCountry id="F32F761209764F6FFB9FFD62FACDECAD" box="[1149,1294,660,682]" name="Philippines" pageId="5" pageNumber="255">Philippines</collectingCountry>
: 
<collectingRegion id="49FCF86009764F6FFAFFFD63FA53ECAD" box="[1309,1424,661,682]" country="Philippines" name="Batangas" pageId="5" pageNumber="255">Batangas</collectingRegion>
(
<bibRefCitation id="EFA94B7309764F6FFCD0FD45FBE4ECCF" author="Femia Marzo G" box="[818,1063,691,713]" pageId="5" pageNumber="255" pagination="1933" refId="ref14274" refString="Femia Marzo G. 2014. Goniodoridella sp. 01 Genus: PruvotFol, 1933. In: Sea slugs of the Mediterranean Sea and elsewhere. Available at: http: // www. medslugs. de / E / Pac-W / Goniodoridella _ sp _ 01 / Goniodoridella _ sp _ 01 _ 02. htm" type="book chapter" year="2014">Femia Marzo, 2014</bibRefCitation>
; present study) and 
<collectingRegion id="49FCF86009764F6FFADBFD45FC4CECEF" country="Philippines" name="Negros Oriental" pageId="5" pageNumber="255">Negros Oriental</collectingRegion>
(
<bibRefCitation id="EFA94B7309764F6FFC40FD24FBF0ECE0" author="Tillen G" box="[930,1075,721,744]" pageId="5" pageNumber="255" pagination="1933" refId="ref16041" refString="Tillen G. 2015. Goniodoridella sp. 03 Genus: Pruvot-Fol, 1933. In: Sea-slugs from the Philippine Islands. Available at: https: // www. philippine-sea-slugs. com / Nudibranchia / Doridina / Goniodoridella _ sp _ 03 _ 10. htm" type="book chapter" year="2015">Tillen, 2015</bibRefCitation>
); 
<collectingCountry id="F32F761209764F6FFBAEFD24FB6BECEF" box="[1100,1192,722,744]" name="Taiwan" pageId="5" pageNumber="255">Taiwan</collectingCountry>
: 
<collectingRegion id="49FCF86009764F6FFB54FD24FAD2ECEF" box="[1206,1297,722,744]" country="Taiwan" name="Penghu" pageId="5" pageNumber="255">Penghu</collectingRegion>
(
<bibRefCitation id="EFA94B7309764F6FFAC1FD24FCA3ED02" author="Su Y &amp; Huang LJ &amp; Chang YW &amp; Mok HK" pageId="5" pageNumber="255" pagination="448 - 459" refId="ref15961" refString="Su Y, Huang LJ, Chang YW, Mok HK. 2009. Temporal changes in nudibranch composition at a coastal site off Penghu (the Pescadores) in the Taiwan Strait. Zoological Studies 48: 448 - 459." type="journal article" year="2009">
Su 
<emphasis id="B94CEA9009764F6FFAAFFD24FA49ECE0" box="[1357,1418,722,743]" italics="true" pageId="5" pageNumber="255">et al.</emphasis>
, 2009
</bibRefCitation>
); 
<collectingCountry id="F32F761209764F6FFC96FD07FC7FED01" box="[884,956,753,774]" name="Japan" pageId="5" pageNumber="255">Japan</collectingCountry>
: Izu Peninsula (
<bibRefCitation id="EFA94B7309764F6FFB92FD06FACFED02" author="Nakano R" box="[1136,1292,752,774]" pageId="5" pageNumber="255" refId="ref15081" refString="Nakano R. 2018. Field guide to sea slugs and nudibranchs of Japan. Tokyo: Bun-ichi Co., Ltd." type="book" year="2018">Nakano, 2018</bibRefCitation>
); 
<collectingCountry id="F32F761209764F6FFAFDFD06FA4EED01" box="[1311,1421,752,774]" name="Australia" pageId="5" pageNumber="255">Australia</collectingCountry>
: 
<collectingRegion id="49FCF86009764F6FFCCBFCF9FBC7ED22" box="[809,1028,783,805]" country="Australia" name="New South Wales" pageId="5" pageNumber="255">New South Wales</collectingRegion>
(
<bibRefCitation id="EFA94B7309764F6FFBFAFCF9FB71ED23" author="Mayes B" box="[1048,1202,783,804]" pageId="5" pageNumber="255" refId="ref14809" refString="Mayes B. 2020. Nudibranch (Goniodoridella sp). In: Brian Mayes (Flickr photostream). Available at: https: // www. flickr. com / photos / brianmayes / 50884781246 /" type="url" year="2020">Mayes, 2020</bibRefCitation>
) and 
<collectingRegion id="49FCF86009764F6FFB1CFCF9FA4CED22" box="[1278,1423,783,805]" country="Australia" name="Queensland" pageId="5" pageNumber="255">Queensland</collectingRegion>
(
<bibRefCitation id="EFA94B7309764F6FFCD3FCD8FC6DED44" author="Cobb G" box="[817,942,814,836]" pageId="5" pageNumber="255" pagination="7" refId="ref13956" refString="Cobb G. 2017. Goniodoridella sp 7. In: Nudibranchs of the Sunshine Coast, Queensland, Australia, Indo-Pacific. Available at: http: // nudibranch. com. au / pages / Cap-Goni-sp 7 - 009 w. htm" type="book chapter" year="2017">Cobb, 2017</bibRefCitation>
; present study).
</paragraph>
</subSubSection>
<subSubSection id="C322650909764F6FFCCBFC9AFBA4EA1C" pageId="5" pageNumber="255" type="biology_ecology">
<paragraph id="8B87368209764F6FFCCBFC9AFBA4EA1C" blockId="5.[809,1426,876,1052]" pageId="5" pageNumber="255">
<emphasis id="B94CEA9009764F6FFCCBFC9AFC4EED86" box="[809,909,876,897]" italics="true" pageId="5" pageNumber="255">Ecology:</emphasis>
The species is found from 
<quantity id="4CC09B6709764F6FFB0BFC9AFAD4ED85" box="[1257,1303,876,898]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="255" unit="m" value="1.0">1 m</quantity>
(
<bibRefCitation id="EFA94B7309764F6FFAC8FC9AFC6BEDA6" author="Gosliner TM &amp; Valdes A &amp; Behrens D" pageId="5" pageNumber="255" refId="ref14453" refString="Gosliner TM, Valdes A, Behrens D. 2015. Nudibranch &amp; sea slug identification - Indo-Pacific. Jacksonville, Florida: New World Publications." type="book" year="2015">
Gosliner 
<emphasis id="B94CEA9009764F6FFCCBFC7AFC98EDA7" box="[809,859,907,929]" italics="true" pageId="5" pageNumber="255">et al</emphasis>
., 2015
</bibRefCitation>
, 
<bibRefCitation id="EFA94B7309764F6FFC54FC7DFC2DEDA7" author="Gosliner TM &amp; Valdes A &amp; Behrens D" box="[950,1006,907,928]" pageId="5" pageNumber="255" refId="ref14480" refString="Gosliner TM, Valdes A, Behrens D. 2018. Nudibranch &amp; sea slug identification - Indo-Pacific. Jacksonville, Florida: New World Publications." type="book" year="2018">2018</bibRefCitation>
) to up to 
<quantity id="4CC09B6709764F6FFBBDFC7DFB5BEDA7" box="[1119,1176,907,928]" metricMagnitude="1" metricUnit="m" metricValue="2.1" pageId="5" pageNumber="255" unit="m" value="21.0">21 m</quantity>
depth (
<bibRefCitation id="EFA94B7309764F6FFB10FC7DFCA2EDB8" author="Femia Marzo G" pageId="5" pageNumber="255" pagination="1933" refId="ref14274" refString="Femia Marzo G. 2014. Goniodoridella sp. 01 Genus: PruvotFol, 1933. In: Sea slugs of the Mediterranean Sea and elsewhere. Available at: http: // www. medslugs. de / E / Pac-W / Goniodoridella _ sp _ 01 / Goniodoridella _ sp _ 01 _ 02. htm" type="book chapter" year="2014">Femia Marzo, 2014</bibRefCitation>
). It has been found on red algae (
<bibRefCitation id="EFA94B7309764F6FFB14FC5CFA4BEDB8" author="Mayes B" box="[1270,1416,938,959]" pageId="5" pageNumber="255" refId="ref14809" refString="Mayes B. 2020. Nudibranch (Goniodoridella sp). In: Brian Mayes (Flickr photostream). Available at: https: // www. flickr. com / photos / brianmayes / 50884781246 /" type="url" year="2020">Mayes, 2020</bibRefCitation>
) and under coral rubble (
<bibRefCitation id="EFA94B7309764F6FFBA9FC3EFAF9EDD9" author="Gosliner TM &amp; Valdes A &amp; Behrens D" box="[1099,1338,968,990]" pageId="5" pageNumber="255" refId="ref14453" refString="Gosliner TM, Valdes A, Behrens D. 2015. Nudibranch &amp; sea slug identification - Indo-Pacific. Jacksonville, Florida: New World Publications." type="book" year="2015">
Gosliner 
<emphasis id="B94CEA9009764F6FFB5AFC3FFB2FEDDA" box="[1208,1260,968,990]" italics="true" pageId="5" pageNumber="255">et al</emphasis>
., 2015
</bibRefCitation>
, 
<bibRefCitation id="EFA94B7309764F6FFAABFC3EFA42EDDA" author="Gosliner TM &amp; Valdes A &amp; Behrens D" box="[1353,1409,968,989]" pageId="5" pageNumber="255" refId="ref14480" refString="Gosliner TM, Valdes A, Behrens D. 2018. Nudibranch &amp; sea slug identification - Indo-Pacific. Jacksonville, Florida: New World Publications." type="book" year="2018">2018</bibRefCitation>
). 
<taxonomicName id="4C384D0109764F6FFCCBFC11FBE0EDFB" authorityName="Paz-Sedano &amp; Smirnoff &amp; Candás &amp; Gosliner &amp; Pola" authorityYear="2022" box="[809,1059,999,1020]" class="Gastropoda" family="Goniodorididae" genus="Murphydoris" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="255" phylum="Mollusca" rank="species" species="adusta">
<emphasis id="B94CEA9009764F6FFCCBFC11FBE0EDFB" box="[809,1059,999,1020]" italics="true" pageId="5" pageNumber="255">Murphydoris adusta</emphasis>
</taxonomicName>
feeds on compound tunicates (
<bibRefCitation id="EFA94B7309764F6FFCD3FBF0FBD7EA1C" author="Gosliner TM &amp; Valdes A &amp; Behrens D" box="[817,1044,1029,1052]" pageId="5" pageNumber="255" refId="ref14453" refString="Gosliner TM, Valdes A, Behrens D. 2015. Nudibranch &amp; sea slug identification - Indo-Pacific. Jacksonville, Florida: New World Publications." type="book" year="2015">
Gosliner 
<emphasis id="B94CEA9009764F6FFC7BFBF0FC0AEA1C" box="[921,969,1030,1051]" italics="true" pageId="5" pageNumber="255">et al</emphasis>
., 2015
</bibRefCitation>
, 
<bibRefCitation id="EFA94B7309764F6FFBC3FBF0FB9BEA1C" author="Gosliner TM &amp; Valdes A &amp; Behrens D" box="[1057,1112,1030,1051]" pageId="5" pageNumber="255" refId="ref14480" refString="Gosliner TM, Valdes A, Behrens D. 2018. Nudibranch &amp; sea slug identification - Indo-Pacific. Jacksonville, Florida: New World Publications." type="book" year="2018">2018</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C322650909764F6FFCCBFBB2FB11E8C6" pageId="5" pageNumber="255" type="etymology">
<paragraph id="8B87368209764F6FFCCBFBB2FB6EEA9F" blockId="5.[809,1425,1092,1176]" pageId="5" pageNumber="255">
<emphasis id="B94CEA9009764F6FFCCBFBB2FC6FEA5E" box="[809,940,1092,1113]" italics="true" pageId="5" pageNumber="255">Etymology:</emphasis>
From Latin 
<emphasis id="B94CEA9009764F6FFBB5FBB2FB77EA5E" box="[1111,1204,1092,1113]" italics="true" pageId="5" pageNumber="255">adustus</emphasis>
, perfect participle passive of 
<emphasis id="B94CEA9009764F6FFC7CFB95FC21EA7F" box="[926,994,1123,1144]" italics="true" pageId="5" pageNumber="255">aduro</emphasis>
, brown. 
<taxonomicName id="4C384D0109764F6FFBA0FB95FAE8EA7F" authorityName="Paz-Sedano &amp; Smirnoff &amp; Candás &amp; Gosliner &amp; Pola" authorityYear="2022" box="[1090,1323,1123,1144]" class="Gastropoda" family="Goniodorididae" genus="Murphydoris" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="255" phylum="Mollusca" rank="species" species="adusta">
<emphasis id="B94CEA9009764F6FFBA0FB95FAE8EA7F" box="[1090,1323,1123,1144]" italics="true" pageId="5" pageNumber="255">Murphydoris adusta</emphasis>
</taxonomicName>
refers to the dark brown colour of the body.
</paragraph>
<paragraph id="8B87368209764F6FFCCBFB36FB11E8C6" blockId="5.[807,1425,1216,1729]" pageId="5" pageNumber="255">
<emphasis id="B94CEA9009764F6FFCCBFB36FB3CEAD1" box="[809,1279,1216,1238]" italics="true" pageId="5" pageNumber="255">
External morphology (
<figureCitation id="13032A0709764F6FFBDFFB37FB7DEAD1" box="[1085,1214,1216,1238]" captionStart="Figure 2" captionStartId="6.[163,241,1491,1513]" captionTargetBox="[323,1283,196,1451]" captionTargetId="figure-159@6.[323,1283,195,1451]" captionTargetPageId="6" captionText="Figure 2. Photographs of living animals. A, B, Murphydoris adusta sp. nov., photographed by G. Cobb: A, (WAM S72660); B, (QMMO 85995). C, Murphydoris cobbi sp. nov. (CASIZ 206795), photographed by T. M. Gosliner. D, M. cobbi sp. nov. (QMMO 85979), photographed by G. Cobb. E, Murphydoris maracabranchia sp. nov. (CASIZ 186120), photographed by P. Paleracio. F, G, M. puncticulata sp. nov. photographed by G. Cobb. F, (WAM S72711), G, (WAM S72710)." figureDoi="http://doi.org/10.5281/zenodo.7037696" httpUri="https://zenodo.org/record/7037696/files/figure.png" pageId="5" pageNumber="255">Figs 2A, B</figureCitation>
, 
<figureCitation id="13032A0709764F6FFB2CFB36FB2CEAD2" box="[1230,1263,1216,1237]" captionStart="Figure 3" captionStartId="7.[144,221,1453,1475]" captionTargetBox="[224,1343,196,1404]" captionTargetId="figure-135@7.[223,1346,195,1413]" captionTargetPageId="7" captionText="Figure 3. A–C, drawings of Murphydoris adusta sp. nov.:A, gill branches; B, buccal bulb (MNCN 15.05/200144); C, reproductive system (MNCN 15.05/200144). D–F, drawings of Murphydoris cobbi sp. nov.: D, modification of gill branches; E, buccal bulb (QMMO 85983); F, reproductive system (QMMO 85982).G–H, drawings ofMurphydoris maracabranchiasp. nov.:G,modification of gill branches; H, reproductive system (CASIZ 224657). I–K, drawings of Murphydoris puncticulata sp. nov.: I, gill branches; J, buccal bulb (WAM S72711); K, reproductive system (WAM S72711). Abbreviations: am, ampulla; bc, bursa copulatrix; bp, buccal pump; fgm, female gland mass; hd, hermaphroditic duct; oe, oesophagus; op, oesophagal pump; p, penis; pr, prostate; ra, radular sac; rs, receptaculum seminis; sgl, salivary gland; ud, uterine duct; va, vagina; vd, vas deferens. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7037702" httpUri="https://zenodo.org/record/7037702/files/figure.png" pageId="5" pageNumber="255">3A</figureCitation>
):
</emphasis>
Preserved specimens between 2 and 
<quantity id="4CC09B6709764F6FFBA9FB29FB4BEAF3" box="[1099,1160,1247,1268]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="255" unit="mm" value="3.0">3 mm</quantity>
length. Body elongated and narrow. Foot tapered at posterior end, with slightly rounded edge at anterior part. Foot narrow, does not protrude from sides of notum. Notal border well developed, with thickened edge at anterior part around head, continuing with slightly serrated margin, supported by internal spicules. Gill located at posterior two-thirds of body. Three bipinnate gill branches arise from common base. Middle branch larger with wider and rounded tip (
<figureCitation id="13032A0709764F6FFC1AFA05FB8FE80F" box="[1016,1100,1523,1545]" captionStart="Figure 3" captionStartId="7.[144,221,1453,1475]" captionTargetBox="[224,1343,196,1404]" captionTargetId="figure-135@7.[223,1346,195,1413]" captionTargetPageId="7" captionText="Figure 3. A–C, drawings of Murphydoris adusta sp. nov.:A, gill branches; B, buccal bulb (MNCN 15.05/200144); C, reproductive system (MNCN 15.05/200144). D–F, drawings of Murphydoris cobbi sp. nov.: D, modification of gill branches; E, buccal bulb (QMMO 85983); F, reproductive system (QMMO 85982).G–H, drawings ofMurphydoris maracabranchiasp. nov.:G,modification of gill branches; H, reproductive system (CASIZ 224657). I–K, drawings of Murphydoris puncticulata sp. nov.: I, gill branches; J, buccal bulb (WAM S72711); K, reproductive system (WAM S72711). Abbreviations: am, ampulla; bc, bursa copulatrix; bp, buccal pump; fgm, female gland mass; hd, hermaphroditic duct; oe, oesophagus; op, oesophagal pump; p, penis; pr, prostate; ra, radular sac; rs, receptaculum seminis; sgl, salivary gland; ud, uterine duct; va, vagina; vd, vas deferens. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7037702" httpUri="https://zenodo.org/record/7037702/files/figure.png" pageId="5" pageNumber="255">Fig. 3A</figureCitation>
). Dorsal or lateral papillae absent. Rhinophores non-retractile and smooth, located at anterior part of body. Rhinophoral sheath absent. Mouth displaced towards ventral zone, lacking oral tentacles. Reproductive opening located halfway up right side. Whole body, including lateral processes, covered by dense network of spicules.
</paragraph>
</subSubSection>
<subSubSection id="C322650909764F63FCCBF902FE49EF73" lastPageId="9" lastPageNumber="259" pageId="5" pageNumber="255" type="description">
<paragraph id="8B87368209764F6CFCCBF902FC09E89C" blockId="5.[809,1425,1780,1894]" lastBlockId="6.[827,1443,1639,1692]" lastPageId="6" lastPageNumber="256" pageId="5" pageNumber="255">
<emphasis id="B94CEA9009764F6FFCCBF902FBB7E90E" box="[809,1140,1780,1802]" italics="true" pageId="5" pageNumber="255">
Colour pattern (
<figureCitation id="13032A0709764F6FFC09F903FBA0E90D" box="[1003,1123,1780,1802]" captionStart="Figure 2" captionStartId="6.[163,241,1491,1513]" captionTargetBox="[323,1283,196,1451]" captionTargetId="figure-159@6.[323,1283,195,1451]" captionTargetPageId="6" captionText="Figure 2. Photographs of living animals. A, B, Murphydoris adusta sp. nov., photographed by G. Cobb: A, (WAM S72660); B, (QMMO 85995). C, Murphydoris cobbi sp. nov. (CASIZ 206795), photographed by T. M. Gosliner. D, M. cobbi sp. nov. (QMMO 85979), photographed by G. Cobb. E, Murphydoris maracabranchia sp. nov. (CASIZ 186120), photographed by P. Paleracio. F, G, M. puncticulata sp. nov. photographed by G. Cobb. F, (WAM S72711), G, (WAM S72710)." figureDoi="http://doi.org/10.5281/zenodo.7037696" httpUri="https://zenodo.org/record/7037696/files/figure.png" pageId="5" pageNumber="255">Fig. 2A, B</figureCitation>
):
</emphasis>
Body chocolate brown with patches between pale blue and iridescent white. Edge of notum same colour as patches with discontinuous pale-yellow coloration at border. Middle dorsal part of the body with transversal iridescent white line and yellow patch on each side. Anteriormost part of head iridescent white with chocolate brown band in middle. Rhinophores with same coloration as body. Gill branches translucent white with yellow tips, sometimes more striking on larger middle branch. Thin line of iridescent white arises from each gill base that join at beginning of tail, continuing with wide iridescent white dorsal band until end of tail. Yellowish pigmentation may be present. Foot blueish with small brown spots.
</paragraph>
<caption id="DF47660A09754F6CFF41FA25FBAAE846" ID-DOI="http://doi.org/10.5281/zenodo.7037696" ID-Zenodo-Dep="7037696" httpUri="https://zenodo.org/record/7037696/files/figure.png" pageId="6" pageNumber="256" startId="6.[163,241,1491,1513]" targetBox="[323,1283,196,1451]" targetPageId="6">
<paragraph id="8B87368209754F6CFF41FA25FBAAE846" blockId="6.[162,1444,1491,1601]" pageId="6" pageNumber="256">
<emphasis id="B94CEA9009754F6CFF41FA25FEC8EBEF" bold="true" box="[163,267,1491,1513]" pageId="6" pageNumber="256">Figure 2.</emphasis>
Photographs of living animals. A, B, 
<taxonomicName id="4C384D0109754F6CFD6EFA25FCA2EBEE" authority="Paz-Sedano &amp; Smirnoff &amp; Candás &amp; Gosliner &amp; Pola, 2022" authorityName="Paz-Sedano &amp; Smirnoff &amp; Candás &amp; Gosliner &amp; Pola" authorityYear="2022" box="[652,865,1491,1513]" class="Gastropoda" family="Goniodorididae" genus="Murphydoris" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="256" phylum="Mollusca" rank="species" species="adusta" status="sp. nov.">
<emphasis id="B94CEA9009754F6CFD6EFA25FCA2EBEE" box="[652,865,1491,1513]" italics="true" pageId="6" pageNumber="256">Murphydoris adusta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27F57EB09754F6CFC84FA25FC70EBEE" box="[870,947,1491,1513]" pageId="6" pageNumber="256" rank="species">sp. nov.</taxonomicNameLabel>
, photographed by G. Cobb: A, (WAM S72660); B, (QMMO 85995). C, 
<taxonomicName id="4C384D0109754F6CFE89FA06FDECE801" authority="Paz-Sedano &amp; Smirnoff &amp; Candás &amp; Gosliner &amp; Pola, 2022" authorityName="Paz-Sedano &amp; Smirnoff &amp; Candás &amp; Gosliner &amp; Pola" authorityYear="2022" box="[363,559,1520,1542]" class="Gastropoda" family="Goniodorididae" genus="Murphydoris" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="256" phylum="Mollusca" rank="species" species="cobbi" status="sp. nov.">
<emphasis id="B94CEA9009754F6CFE89FA06FDECE801" box="[363,559,1520,1542]" italics="true" pageId="6" pageNumber="256">Murphydoris cobbi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27F57EB09754F6CFDD7FA06FD43E801" box="[565,640,1520,1542]" pageId="6" pageNumber="256" rank="species">sp. nov.</taxonomicNameLabel>
(CASIZ 206795), photographed by T. M. Gosliner. D, 
<taxonomicName id="4C384D0109754F6CFB42FA07FB34E801" authority="Paz-Sedano &amp; Smirnoff &amp; Candás &amp; Gosliner &amp; Pola, 2022" authorityName="Paz-Sedano &amp; Smirnoff &amp; Candás &amp; Gosliner &amp; Pola" authorityYear="2022" box="[1184,1271,1520,1542]" class="Gastropoda" family="Goniodorididae" genus="Murphydoris" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="256" phylum="Mollusca" rank="species" species="cobbi" status="sp. nov.">
<emphasis id="B94CEA9009754F6CFB42FA07FB34E801" box="[1184,1271,1520,1542]" italics="true" pageId="6" pageNumber="256">M. cobbi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27F57EB09754F6CFB1EFA06FA8BE801" box="[1276,1352,1520,1542]" pageId="6" pageNumber="256" rank="species">sp. nov.</taxonomicNameLabel>
(QMMO 85979), photographed by G. Cobb. E, 
<taxonomicName id="4C384D0109754F6CFDC4F9F8FCA1E823" authority="Paz-Sedano &amp; Smirnoff &amp; Candás &amp; Gosliner &amp; Pola, 2022" authorityName="Paz-Sedano &amp; Smirnoff &amp; Candás &amp; Gosliner &amp; Pola" authorityYear="2022" box="[550,866,1550,1572]" class="Gastropoda" family="Goniodorididae" genus="Murphydoris" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="256" phylum="Mollusca" rank="species" species="maracabranchia" status="sp. nov.">
<emphasis id="B94CEA9009754F6CFDC4F9F8FCA1E823" box="[550,866,1550,1572]" italics="true" pageId="6" pageNumber="256">Murphydoris maracabranchia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27F57EB09754F6CFC8BF9F8FC76E823" box="[873,949,1550,1572]" pageId="6" pageNumber="256" rank="species">sp. nov.</taxonomicNameLabel>
(CASIZ 186120), photographed by P. Paleracio. F, G, 
<taxonomicName id="4C384D0109754F6CFF3AF9DAFEBEE846" authority="Paz-Sedano &amp; Smirnoff &amp; Candás &amp; Gosliner &amp; Pola, 2022" authorityName="Paz-Sedano &amp; Smirnoff &amp; Candás &amp; Gosliner &amp; Pola" authorityYear="2022" box="[216,381,1579,1601]" class="Gastropoda" family="Goniodorididae" genus="Murphydoris" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="256" phylum="Mollusca" rank="species" species="puncticulata" status="sp. nov.">
<emphasis id="B94CEA9009754F6CFF3AF9DAFEBEE846" box="[216,381,1579,1601]" italics="true" pageId="6" pageNumber="256">M. puncticulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27F57EB09754F6CFE60F9DDFE0DE846" box="[386,462,1579,1601]" pageId="6" pageNumber="256" rank="species">sp. nov.</taxonomicNameLabel>
photographed by G. Cobb. F, (WAM 
<accessionNumber id="946BAB6109754F6CFCA5F9DDFC54E846" box="[839,919,1579,1601]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S72711" pageId="6" pageNumber="256" type="EnaNcbi">S72711</accessionNumber>
), G, (WAM 
<accessionNumber id="946BAB6109754F6CFBEFF9DDFB9EE846" box="[1037,1117,1579,1601]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S72710" pageId="6" pageNumber="256" type="EnaNcbi">S72710</accessionNumber>
).
</paragraph>
</caption>
<paragraph id="8B87368209754F6DFCD9F933FCBFE8EA" blockId="6.[827,1443,1733,1908]" lastBlockId="7.[809,1424,1689,1773]" lastPageId="7" lastPageNumber="257" pageId="6" pageNumber="256">
<emphasis id="B94CEA9009754F6CFCD9F933FB32E8DD" box="[827,1265,1733,1754]" italics="true" pageId="6" pageNumber="256">
Foregut anatomy (
<figureCitation id="13032A0709754F6CFBCAF933FB48E8DD" box="[1064,1163,1733,1754]" captionStart="Figure 3" captionStartId="7.[144,221,1453,1475]" captionTargetBox="[224,1343,196,1404]" captionTargetId="figure-135@7.[223,1346,195,1413]" captionTargetPageId="7" captionText="Figure 3. A–C, drawings of Murphydoris adusta sp. nov.:A, gill branches; B, buccal bulb (MNCN 15.05/200144); C, reproductive system (MNCN 15.05/200144). D–F, drawings of Murphydoris cobbi sp. nov.: D, modification of gill branches; E, buccal bulb (QMMO 85983); F, reproductive system (QMMO 85982).G–H, drawings ofMurphydoris maracabranchiasp. nov.:G,modification of gill branches; H, reproductive system (CASIZ 224657). I–K, drawings of Murphydoris puncticulata sp. nov.: I, gill branches; J, buccal bulb (WAM S72711); K, reproductive system (WAM S72711). Abbreviations: am, ampulla; bc, bursa copulatrix; bp, buccal pump; fgm, female gland mass; hd, hermaphroditic duct; oe, oesophagus; op, oesophagal pump; p, penis; pr, prostate; ra, radular sac; rs, receptaculum seminis; sgl, salivary gland; ud, uterine duct; va, vagina; vd, vas deferens. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7037702" httpUri="https://zenodo.org/record/7037702/files/figure.png" pageId="6" pageNumber="256">Figs 3B</figureCitation>
, 
<figureCitation id="13032A0709754F6CFB7CF933FB1CE8DD" box="[1182,1247,1733,1754]" captionStart="Figure 4" captionStartId="8.[163,241,1584,1606]" captionTargetBox="[323,1283,196,1544]" captionTargetId="figure-87@8.[323,1283,195,1544]" captionTargetPageId="8" captionText="Figure 4. Scanning electron microscope (SEM) photographs and light microscope photographs (LMP) of A–D, Murphydoris adusta sp. nov. and E–H, Murphydoris cobbi sp. nov. A, frontal view of radula (QMMO 85995) (SEM); B, lateral view of teeth (CASIZ 181290) (SEM); C, detail of masticatory margin of teeth (QMMO 85995) (SEM); D, detail of penial spines (QMMO 85986) (SEM); E, detail of labial cuticle elements (QMMO 85984) (SEM); F, frontal view of radula – detail of rachis, internal and external teeth (QMMO 85983) (SEM); G, detail of internal teeth (QMMO 85983) (SEM); H, detail of penial spines (LMP) (QMMO 85983). Scale bars: A, 50 µm; B, 40 µm; C, 10 µm; D, 10 µm; E, 5µm; F, 30 µm; G, 20 µm; H, 10 µm." figureDoi="http://doi.org/10.5281/zenodo.7037704" httpUri="https://zenodo.org/record/7037704/files/figure.png" pageId="6" pageNumber="256">4A–C</figureCitation>
):
</emphasis>
Buccal bulb muscular. Dorsal buccal pump elongated posteriorly (
<figureCitation id="13032A0709754F6CFCA1F8F4FC56E910" box="[835,917,1794,1816]" captionStart="Figure 3" captionStartId="7.[144,221,1453,1475]" captionTargetBox="[224,1343,196,1404]" captionTargetId="figure-135@7.[223,1346,195,1413]" captionTargetPageId="7" captionText="Figure 3. A–C, drawings of Murphydoris adusta sp. nov.:A, gill branches; B, buccal bulb (MNCN 15.05/200144); C, reproductive system (MNCN 15.05/200144). D–F, drawings of Murphydoris cobbi sp. nov.: D, modification of gill branches; E, buccal bulb (QMMO 85983); F, reproductive system (QMMO 85982).G–H, drawings ofMurphydoris maracabranchiasp. nov.:G,modification of gill branches; H, reproductive system (CASIZ 224657). I–K, drawings of Murphydoris puncticulata sp. nov.: I, gill branches; J, buccal bulb (WAM S72711); K, reproductive system (WAM S72711). Abbreviations: am, ampulla; bc, bursa copulatrix; bp, buccal pump; fgm, female gland mass; hd, hermaphroditic duct; oe, oesophagus; op, oesophagal pump; p, penis; pr, prostate; ra, radular sac; rs, receptaculum seminis; sgl, salivary gland; ud, uterine duct; va, vagina; vd, vas deferens. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7037702" httpUri="https://zenodo.org/record/7037702/files/figure.png" pageId="6" pageNumber="256">Fig. 3B</figureCitation>
). Radular sac short, descending ventrally and expanding posteriorly. Thin oesophagus originates at buccal bulb behind buccal pump. Nervous system surrounds this area. Oesophagus becomes wider and continues with oesophageal pump. One small and rounded salivary gland on each side, at the junction of oesophagus and buccal pump (
<figureCitation id="13032A0709744F6DFE11F921FD86E8EB" box="[499,581,1751,1773]" captionStart="Figure 3" captionStartId="7.[144,221,1453,1475]" captionTargetBox="[224,1343,196,1404]" captionTargetId="figure-135@7.[223,1346,195,1413]" captionTargetPageId="7" captionText="Figure 3. A–C, drawings of Murphydoris adusta sp. nov.:A, gill branches; B, buccal bulb (MNCN 15.05/200144); C, reproductive system (MNCN 15.05/200144). D–F, drawings of Murphydoris cobbi sp. nov.: D, modification of gill branches; E, buccal bulb (QMMO 85983); F, reproductive system (QMMO 85982).G–H, drawings ofMurphydoris maracabranchiasp. nov.:G,modification of gill branches; H, reproductive system (CASIZ 224657). I–K, drawings of Murphydoris puncticulata sp. nov.: I, gill branches; J, buccal bulb (WAM S72711); K, reproductive system (WAM S72711). Abbreviations: am, ampulla; bc, bursa copulatrix; bp, buccal pump; fgm, female gland mass; hd, hermaphroditic duct; oe, oesophagus; op, oesophagal pump; p, penis; pr, prostate; ra, radular sac; rs, receptaculum seminis; sgl, salivary gland; ud, uterine duct; va, vagina; vd, vas deferens. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7037702" httpUri="https://zenodo.org/record/7037702/files/figure.png" pageId="7" pageNumber="257">Fig. 3B</figureCitation>
). Labial cuticle surrounds lips and expands within buccal pump. Radular formula 11–13 × 1.0.1. Lateral tooth large and robust, with two upper rounded cusps (
<figureCitation id="13032A0709744F6DFD9EF8C5FD28E94F" box="[636,747,1843,1865]" captionStart="Figure 4" captionStartId="8.[163,241,1584,1606]" captionTargetBox="[323,1283,196,1544]" captionTargetId="figure-87@8.[323,1283,195,1544]" captionTargetPageId="8" captionText="Figure 4. Scanning electron microscope (SEM) photographs and light microscope photographs (LMP) of A–D, Murphydoris adusta sp. nov. and E–H, Murphydoris cobbi sp. nov. A, frontal view of radula (QMMO 85995) (SEM); B, lateral view of teeth (CASIZ 181290) (SEM); C, detail of masticatory margin of teeth (QMMO 85995) (SEM); D, detail of penial spines (QMMO 85986) (SEM); E, detail of labial cuticle elements (QMMO 85984) (SEM); F, frontal view of radula – detail of rachis, internal and external teeth (QMMO 85983) (SEM); G, detail of internal teeth (QMMO 85983) (SEM); H, detail of penial spines (LMP) (QMMO 85983). Scale bars: A, 50 µm; B, 40 µm; C, 10 µm; D, 10 µm; E, 5µm; F, 30 µm; G, 20 µm; H, 10 µm." figureDoi="http://doi.org/10.5281/zenodo.7037704" httpUri="https://zenodo.org/record/7037704/files/figure.png" pageId="7" pageNumber="257">Fig. 4A, B</figureCitation>
). Thick masticatory margin. Masticatory margin with few, small, thin denticles (
<figureCitation id="13032A0709744F6DFBBFF96FFB72E8A9" box="[1117,1201,1689,1711]" captionStart="Figure 4" captionStartId="8.[163,241,1584,1606]" captionTargetBox="[323,1283,196,1544]" captionTargetId="figure-87@8.[323,1283,195,1544]" captionTargetPageId="8" captionText="Figure 4. Scanning electron microscope (SEM) photographs and light microscope photographs (LMP) of A–D, Murphydoris adusta sp. nov. and E–H, Murphydoris cobbi sp. nov. A, frontal view of radula (QMMO 85995) (SEM); B, lateral view of teeth (CASIZ 181290) (SEM); C, detail of masticatory margin of teeth (QMMO 85995) (SEM); D, detail of penial spines (QMMO 85986) (SEM); E, detail of labial cuticle elements (QMMO 85984) (SEM); F, frontal view of radula – detail of rachis, internal and external teeth (QMMO 85983) (SEM); G, detail of internal teeth (QMMO 85983) (SEM); H, detail of penial spines (LMP) (QMMO 85983). Scale bars: A, 50 µm; B, 40 µm; C, 10 µm; D, 10 µm; E, 5µm; F, 30 µm; G, 20 µm; H, 10 µm." figureDoi="http://doi.org/10.5281/zenodo.7037704" httpUri="https://zenodo.org/record/7037704/files/figure.png" pageId="7" pageNumber="257">Fig. 4C</figureCitation>
). Base of the teeth wide and straight (
<figureCitation id="13032A0709744F6DFBF7F94EFB4DE8CA" box="[1045,1166,1720,1742]" captionStart="Figure 4" captionStartId="8.[163,241,1584,1606]" captionTargetBox="[323,1283,196,1544]" captionTargetId="figure-87@8.[323,1283,195,1544]" captionTargetPageId="8" captionText="Figure 4. Scanning electron microscope (SEM) photographs and light microscope photographs (LMP) of A–D, Murphydoris adusta sp. nov. and E–H, Murphydoris cobbi sp. nov. A, frontal view of radula (QMMO 85995) (SEM); B, lateral view of teeth (CASIZ 181290) (SEM); C, detail of masticatory margin of teeth (QMMO 85995) (SEM); D, detail of penial spines (QMMO 85986) (SEM); E, detail of labial cuticle elements (QMMO 85984) (SEM); F, frontal view of radula – detail of rachis, internal and external teeth (QMMO 85983) (SEM); G, detail of internal teeth (QMMO 85983) (SEM); H, detail of penial spines (LMP) (QMMO 85983). Scale bars: A, 50 µm; B, 40 µm; C, 10 µm; D, 10 µm; E, 5µm; F, 30 µm; G, 20 µm; H, 10 µm." figureDoi="http://doi.org/10.5281/zenodo.7037704" httpUri="https://zenodo.org/record/7037704/files/figure.png" pageId="7" pageNumber="257">Fig. 4A, B</figureCitation>
). Outer lateral teeth absent.
</paragraph>
<caption id="DF47660A09744F6DFF72FA5BFAC4E874" ID-DOI="http://doi.org/10.5281/zenodo.7037702" ID-Zenodo-Dep="7037702" httpUri="https://zenodo.org/record/7037702/files/figure.png" pageId="7" pageNumber="257" startId="7.[144,221,1453,1475]" targetBox="[224,1343,196,1404]" targetPageId="7">
<paragraph id="8B87368209744F6DFF72FA5BFAC4E874" blockId="7.[143,1426,1453,1651]" pageId="7" pageNumber="257">
<emphasis id="B94CEA9009744F6DFF72FA5BFF36EBC5" bold="true" box="[144,245,1453,1475]" pageId="7" pageNumber="257">Figure 3.</emphasis>
A–C, drawings of 
<taxonomicName id="4C384D0109744F6DFE4CFA5BFDBEEBC4" authority="Paz-Sedano &amp; Smirnoff &amp; Candás &amp; Gosliner &amp; Pola, 2022" authorityName="Paz-Sedano &amp; Smirnoff &amp; Candás &amp; Gosliner &amp; Pola" authorityYear="2022" box="[430,637,1453,1475]" class="Gastropoda" family="Goniodorididae" genus="Murphydoris" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="257" phylum="Mollusca" rank="species" species="adusta" status="sp. nov.">
<emphasis id="B94CEA9009744F6DFE4CFA5BFDBEEBC4" box="[430,637,1453,1475]" italics="true" pageId="7" pageNumber="257">Murphydoris adusta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27F57EB09744F6DFD60FA5BFD0EEBC4" box="[642,717,1453,1475]" pageId="7" pageNumber="257" rank="species">sp. nov.</taxonomicNameLabel>
: A, gill branches; B, buccal bulb (MNCN 15.05/200144); C, reproductive system (MNCN 15.05/200144). D–F, drawings of 
<taxonomicName id="4C384D0109744F6DFD6EFA3CFC93EBE7" authority="Paz-Sedano &amp; Smirnoff &amp; Candás &amp; Gosliner &amp; Pola, 2022" authorityName="Paz-Sedano &amp; Smirnoff &amp; Candás &amp; Gosliner &amp; Pola" authorityYear="2022" box="[652,848,1482,1504]" class="Gastropoda" family="Goniodorididae" genus="Murphydoris" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="257" phylum="Mollusca" rank="species" species="cobbi" status="sp. nov.">
<emphasis id="B94CEA9009744F6DFD6EFA3CFC93EBE7" box="[652,848,1482,1504]" italics="true" pageId="7" pageNumber="257">Murphydoris cobbi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27F57EB09744F6DFCB5FA3CFC66EBE7" box="[855,933,1482,1504]" pageId="7" pageNumber="257" rank="species">sp. nov.</taxonomicNameLabel>
: D, modification of gill branches; E, buccal bulb (QMMO 85983); F, reproductive system (QMMO 85982).G–H, drawings of 
<taxonomicName id="4C384D0109744F6DFC91FA1EFB67EBF9" authority="Paz-Sedano &amp; Smirnoff &amp; Candás &amp; Gosliner &amp; Pola, 2022" authorityName="Paz-Sedano &amp; Smirnoff &amp; Candás &amp; Gosliner &amp; Pola" authorityYear="2022" box="[883,1188,1512,1534]" class="Gastropoda" family="Goniodorididae" genus="Murphydoris" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="257" phylum="Mollusca" rank="species" species="maracabranchia" status="sp. nov.">
<emphasis id="B94CEA9009744F6DFC91FA1EFB67EBF9" box="[883,1188,1512,1534]" italics="true" pageId="7" pageNumber="257">Murphydoris maracabranchia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27F57EB09744F6DFB44FA1EFB30EBF9" box="[1190,1267,1512,1534]" pageId="7" pageNumber="257" rank="species">sp. nov.</taxonomicNameLabel>
: G, modification of gill branches; H, reproductive system (CASIZ 224657). I–K, drawings of 
<taxonomicName id="4C384D0109744F6DFC9CF9F3FB4BE81C" authority="Paz-Sedano &amp; Smirnoff &amp; Candás &amp; Gosliner &amp; Pola, 2022" authorityName="Paz-Sedano &amp; Smirnoff &amp; Candás &amp; Gosliner &amp; Pola" authorityYear="2022" box="[894,1160,1541,1563]" class="Gastropoda" family="Goniodorididae" genus="Murphydoris" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="257" phylum="Mollusca" rank="species" species="puncticulata" status="sp. nov.">
<emphasis id="B94CEA9009744F6DFC9CF9F3FB4BE81C" box="[894,1160,1541,1563]" italics="true" pageId="7" pageNumber="257">Murphydoris puncticulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27F57EB09744F6DFB6FF9F3FB1AE81C" box="[1165,1241,1541,1563]" pageId="7" pageNumber="257" rank="species">sp. nov.</taxonomicNameLabel>
: I, gill branches; J, buccal bulb (WAM 
<accessionNumber id="946BAB6109744F6DFEADF9D4FE5FE83F" box="[335,412,1570,1592]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S72711" pageId="7" pageNumber="257" type="EnaNcbi">S72711</accessionNumber>
); K, reproductive system (WAM 
<accessionNumber id="946BAB6109744F6DFD02F9D4FCEEE83F" box="[736,813,1570,1592]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S72711" pageId="7" pageNumber="257" type="EnaNcbi">S72711</accessionNumber>
). Abbreviations: am, ampulla; bc, bursa copulatrix; bp, buccal pump; fgm, female gland mass; hd, hermaphroditic duct; oe, oesophagus; op, oesophagal pump; p, penis; pr, prostate; ra, radular sac; rs, receptaculum seminis; sgl, salivary gland; ud, uterine duct; va, vagina; vd, vas deferens. Scale bars: 1 mm.
</paragraph>
</caption>
<paragraph id="8B87368209744F63FCCBF8E3FE49EF73" blockId="7.[809,1425,1813,1897]" lastBlockId="9.[145,762,197,372]" lastPageId="9" lastPageNumber="259" pageId="7" pageNumber="257">
<emphasis id="B94CEA9009744F6DFCCBF8E3FB1BE92C" box="[809,1240,1813,1835]" italics="true" pageId="7" pageNumber="257">
Reproductive system (
<figureCitation id="13032A0709744F6DFBD4F8E0FB56E92D" box="[1078,1173,1813,1835]" captionStart="Figure 3" captionStartId="7.[144,221,1453,1475]" captionTargetBox="[224,1343,196,1404]" captionTargetId="figure-135@7.[223,1346,195,1413]" captionTargetPageId="7" captionText="Figure 3. A–C, drawings of Murphydoris adusta sp. nov.:A, gill branches; B, buccal bulb (MNCN 15.05/200144); C, reproductive system (MNCN 15.05/200144). D–F, drawings of Murphydoris cobbi sp. nov.: D, modification of gill branches; E, buccal bulb (QMMO 85983); F, reproductive system (QMMO 85982).G–H, drawings ofMurphydoris maracabranchiasp. nov.:G,modification of gill branches; H, reproductive system (CASIZ 224657). I–K, drawings of Murphydoris puncticulata sp. nov.: I, gill branches; J, buccal bulb (WAM S72711); K, reproductive system (WAM S72711). Abbreviations: am, ampulla; bc, bursa copulatrix; bp, buccal pump; fgm, female gland mass; hd, hermaphroditic duct; oe, oesophagus; op, oesophagal pump; p, penis; pr, prostate; ra, radular sac; rs, receptaculum seminis; sgl, salivary gland; ud, uterine duct; va, vagina; vd, vas deferens. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7037702" httpUri="https://zenodo.org/record/7037702/files/figure.png" pageId="7" pageNumber="257">Figs 3C</figureCitation>
, 
<figureCitation id="13032A0709744F6DFB47F8E3FB05E92D" box="[1189,1222,1813,1834]" captionStart="Figure 4" captionStartId="8.[163,241,1584,1606]" captionTargetBox="[323,1283,196,1544]" captionTargetId="figure-87@8.[323,1283,195,1544]" captionTargetPageId="8" captionText="Figure 4. Scanning electron microscope (SEM) photographs and light microscope photographs (LMP) of A–D, Murphydoris adusta sp. nov. and E–H, Murphydoris cobbi sp. nov. A, frontal view of radula (QMMO 85995) (SEM); B, lateral view of teeth (CASIZ 181290) (SEM); C, detail of masticatory margin of teeth (QMMO 85995) (SEM); D, detail of penial spines (QMMO 85986) (SEM); E, detail of labial cuticle elements (QMMO 85984) (SEM); F, frontal view of radula – detail of rachis, internal and external teeth (QMMO 85983) (SEM); G, detail of internal teeth (QMMO 85983) (SEM); H, detail of penial spines (LMP) (QMMO 85983). Scale bars: A, 50 µm; B, 40 µm; C, 10 µm; D, 10 µm; E, 5µm; F, 30 µm; G, 20 µm; H, 10 µm." figureDoi="http://doi.org/10.5281/zenodo.7037704" httpUri="https://zenodo.org/record/7037704/files/figure.png" pageId="7" pageNumber="257">4D</figureCitation>
):
</emphasis>
Reproductive system located at anterior-third of body. Thin, elongate hermaphroditic duct begins at ovotestis, located inside digestive-hermaphroditic gland. Hermaphroditic duct expands into large, bean-shaped ampulla. Thin post-ampullary duct enters in female gland mass and divides into the prostatic portion of vas deferens and oviduct. Prostate large, wide and elongated, narrowing and continuing as long vas deferens, ending in penial sac at most distal part. Penial sac short and spherical. Penis armed. Penial spines hooked at base, long and thin at most distal part (
<figureCitation id="13032A07097A4F63FDD4FF33FD4DEEDD" box="[566,654,197,219]" captionStart="Figure 4" captionStartId="8.[163,241,1584,1606]" captionTargetBox="[323,1283,196,1544]" captionTargetId="figure-87@8.[323,1283,195,1544]" captionTargetPageId="8" captionText="Figure 4. Scanning electron microscope (SEM) photographs and light microscope photographs (LMP) of A–D, Murphydoris adusta sp. nov. and E–H, Murphydoris cobbi sp. nov. A, frontal view of radula (QMMO 85995) (SEM); B, lateral view of teeth (CASIZ 181290) (SEM); C, detail of masticatory margin of teeth (QMMO 85995) (SEM); D, detail of penial spines (QMMO 85986) (SEM); E, detail of labial cuticle elements (QMMO 85984) (SEM); F, frontal view of radula – detail of rachis, internal and external teeth (QMMO 85983) (SEM); G, detail of internal teeth (QMMO 85983) (SEM); H, detail of penial spines (LMP) (QMMO 85983). Scale bars: A, 50 µm; B, 40 µm; C, 10 µm; D, 10 µm; E, 5µm; F, 30 µm; G, 20 µm; H, 10 µm." figureDoi="http://doi.org/10.5281/zenodo.7037704" httpUri="https://zenodo.org/record/7037704/files/figure.png" pageId="9" pageNumber="259">Fig. 4D</figureCitation>
). Vagina wide and short, followed by rounded bursa copulatrix. From base of bursa arises a thin duct that connects with elongated receptaculum seminis. Thin and long uterine duct arises at distal part of vagina and enters in female gland mass.
</paragraph>
</subSubSection>
<caption id="DF47660A097B4F62FF41F9C6FB3BE8DE" ID-DOI="http://doi.org/10.5281/zenodo.7037704" ID-Zenodo-Dep="7037704" httpUri="https://zenodo.org/record/7037704/files/figure.png" pageId="8" pageNumber="258" startId="8.[163,241,1584,1606]" targetBox="[323,1283,196,1544]" targetPageId="8">
<paragraph id="8B873682097B4F62FF41F9C6FB3BE8DE" blockId="8.[162,1443,1584,1753]" pageId="8" pageNumber="258">
<emphasis id="B94CEA90097B4F62FF41F9C6FEC9E842" bold="true" box="[163,266,1584,1606]" pageId="8" pageNumber="258">Figure 4.</emphasis>
Scanning electron microscope (SEM) photographs and light microscope photographs (LMP) of A–D, 
<taxonomicName id="4C384D01097B4F62FAF8F9C6FF28E863" authority="Paz-Sedano &amp; Smirnoff &amp; Candás &amp; Gosliner &amp; Pola, 2022" authorityName="Paz-Sedano &amp; Smirnoff &amp; Candás &amp; Gosliner &amp; Pola" authorityYear="2022" class="Gastropoda" family="Goniodorididae" genus="Murphydoris" kingdom="Animalia" order="Nudibranchia" pageId="8" pageNumber="258" phylum="Mollusca" rank="species" species="adusta" status="sp. nov.">
<emphasis id="B94CEA90097B4F62FAF8F9C6FF28E863" italics="true" pageId="8" pageNumber="258">Murphydoris adusta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27F57EB097B4F62FF12F9B8FEF8E863" box="[240,315,1614,1636]" pageId="8" pageNumber="258" rank="species">sp. nov.</taxonomicNameLabel>
and E–H, 
<taxonomicName id="4C384D01097B4F62FE4AF9B8FDAFE863" authority="Paz-Sedano &amp; Smirnoff &amp; Candás &amp; Gosliner &amp; Pola, 2022" authorityName="Paz-Sedano &amp; Smirnoff &amp; Candás &amp; Gosliner &amp; Pola" authorityYear="2022" box="[424,620,1614,1636]" class="Gastropoda" family="Goniodorididae" genus="Murphydoris" kingdom="Animalia" order="Nudibranchia" pageId="8" pageNumber="258" phylum="Mollusca" rank="species" species="cobbi" status="sp. nov.">
<emphasis id="B94CEA90097B4F62FE4AF9B8FDAFE863" box="[424,620,1614,1636]" italics="true" pageId="8" pageNumber="258">Murphydoris cobbi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27F57EB097B4F62FD90F9B8FD7EE863" box="[626,701,1614,1636]" pageId="8" pageNumber="258" rank="species">sp. nov.</taxonomicNameLabel>
A, frontal view of radula (QMMO 85995) (SEM); B, lateral view of teeth (CASIZ 181290) (SEM); C, detail of masticatory margin of teeth (QMMO 85995) (SEM); D, detail of penial spines (QMMO 85986) (SEM); E, detail of labial cuticle elements (QMMO 85984) (SEM); F, frontal view of radula – detail of rachis, internal and external teeth (QMMO 85983) (SEM); G, detail of internal teeth (QMMO 85983) (SEM); H, detail of penial spines (LMP) (QMMO 85983). Scale bars: A, 50 µm; B, 40 µm; C, 10 µm; D, 10 µm; E, 5µm; F, 30 µm; G, 20 µm; H, 10 µm.
</paragraph>
</caption>
<subSubSection id="C3226509097A4F63FF73FE6BFD71EA31" pageId="9" pageNumber="259" type="discussion">
<paragraph id="8B873682097A4F63FF73FE6BFD71EA31" blockId="9.[145,762,413,1079]" pageId="9" pageNumber="259">
<emphasis id="B94CEA90097A4F63FF73FE6BFE3EEFB5" box="[145,509,413,434]" italics="true" pageId="9" pageNumber="259">
Remarks: 
<taxonomicName id="4C384D01097A4F63FEF3FE6BFE3EEFB5" authorityName="Paz-Sedano &amp; Smirnoff &amp; Candás &amp; Gosliner &amp; Pola" authorityYear="2022" box="[273,509,413,434]" class="Gastropoda" family="Goniodorididae" genus="Murphydoris" kingdom="Animalia" order="Nudibranchia" pageId="9" pageNumber="259" phylum="Mollusca" rank="species" species="adusta">Murphydoris adusta</taxonomicName>
</emphasis>
and 
<taxonomicName id="4C384D01097A4F63FDDAFE68FD3BEFB4" authorityName="Sigurdson" authorityYear="1991" box="[568,760,414,435]" class="Gastropoda" family="Goniodorididae" genus="Murphydoris" kingdom="Animalia" order="Nudibranchia" pageId="9" pageNumber="259" phylum="Mollusca" rank="species" species="singaporensis">
<emphasis id="B94CEA90097A4F63FDDAFE68FD3BEFB4" box="[568,760,414,435]" italics="true" pageId="9" pageNumber="259">M. singaporensis</emphasis>
</taxonomicName>
share brownish coloration (
<bibRefCitation id="EFA94B73097A4F63FE26FE4AFD47EFD6" author="Sigurdsson JB" box="[452,644,444,466]" pageId="9" pageNumber="259" pagination="259 - 263" refId="ref15924" refString="Sigurdsson JB. 1991. A nudibranch, Murphydoris singaporensis, new genus and species, from Singapore mangroves (Mollusca: Opisthobranchia: Goniodorididae). Raffles Bulletin of Zoology 39: 259 - 263." type="journal article" year="1991">Sigurdsson, 1991</bibRefCitation>
; 
<bibRefCitation id="EFA94B73097A4F63FD73FE4AFE80EFF7" author="Swennen C &amp; Buatip S" pageId="9" pageNumber="259" pagination="9 - 14" refId="ref16003" refString="Swennen C, Buatip S. 2012. Murphydoris singaporensis (Gastropoda: Nudibranchia: Goniodorididae) found in Thailand; with external and first internal description of the specimens. Strombus 19: 9 - 14." type="journal article" year="2012">Swennen &amp; Buatip, 2012</bibRefCitation>
), but 
<taxonomicName id="4C384D01097A4F63FE68FE2DFD8DEFF7" authorityName="Sigurdson" authorityYear="1991" box="[394,590,475,496]" class="Gastropoda" family="Goniodorididae" genus="Murphydoris" kingdom="Animalia" order="Nudibranchia" pageId="9" pageNumber="259" phylum="Mollusca" rank="species" species="singaporensis">
<emphasis id="B94CEA90097A4F63FE68FE2DFD8DEFF7" box="[394,590,475,496]" italics="true" pageId="9" pageNumber="259">M. singaporensis</emphasis>
</taxonomicName>
is translucent with reddish-brown blotches (
<bibRefCitation id="EFA94B73097A4F63FDF4FE0FFD32EC09" author="Sigurdsson JB" box="[534,753,505,527]" pageId="9" pageNumber="259" pagination="259 - 263" refId="ref15924" refString="Sigurdsson JB. 1991. A nudibranch, Murphydoris singaporensis, new genus and species, from Singapore mangroves (Mollusca: Opisthobranchia: Goniodorididae). Raffles Bulletin of Zoology 39: 259 - 263." type="journal article" year="1991">Sigurdsson, 1991</bibRefCitation>
; 
<bibRefCitation id="EFA94B73097A4F63FF73FDEEFE6DEC2A" author="Swennen C &amp; Buatip S" box="[145,430,536,558]" pageId="9" pageNumber="259" pagination="9 - 14" refId="ref16003" refString="Swennen C, Buatip S. 2012. Murphydoris singaporensis (Gastropoda: Nudibranchia: Goniodorididae) found in Thailand; with external and first internal description of the specimens. Strombus 19: 9 - 14." type="journal article" year="2012">Swennen &amp; Buatip, 2012</bibRefCitation>
), while 
<taxonomicName id="4C384D01097A4F63FDEBFDEFFDB8EC2A" authorityName="Paz-Sedano &amp; Smirnoff &amp; Candás &amp; Gosliner &amp; Pola" authorityYear="2022" box="[521,635,536,558]" class="Gastropoda" family="Goniodorididae" genus="Murphydoris" kingdom="Animalia" order="Nudibranchia" pageId="9" pageNumber="259" phylum="Mollusca" rank="species" species="adusta">
<emphasis id="B94CEA90097A4F63FDEBFDEFFDB8EC2A" box="[521,635,536,558]" italics="true" pageId="9" pageNumber="259">M. adusta</emphasis>
</taxonomicName>
has brown colour with pale blue, iridescent white and yellow in different parts of the body. Moreover, 
<taxonomicName id="4C384D01097A4F63FDA3FDA0FD70EC6D" authorityName="Paz-Sedano &amp; Smirnoff &amp; Candás &amp; Gosliner &amp; Pola" authorityYear="2022" box="[577,691,597,619]" class="Gastropoda" family="Goniodorididae" genus="Murphydoris" kingdom="Animalia" order="Nudibranchia" pageId="9" pageNumber="259" phylum="Mollusca" rank="species" species="adusta">
<emphasis id="B94CEA90097A4F63FDA3FDA0FD70EC6D" box="[577,691,597,619]" italics="true" pageId="9" pageNumber="259">M. adusta</emphasis>
</taxonomicName>
has a prolonged edge at the anterior part of the notal border, whereas 
<taxonomicName id="4C384D01097A4F63FF17FD65FE71ECAF" authorityName="Sigurdson" authorityYear="1991" box="[245,434,659,680]" class="Gastropoda" family="Goniodorididae" genus="Murphydoris" kingdom="Animalia" order="Nudibranchia" pageId="9" pageNumber="259" phylum="Mollusca" rank="species" species="singaporensis">
<emphasis id="B94CEA90097A4F63FF17FD65FE71ECAF" box="[245,434,659,680]" italics="true" pageId="9" pageNumber="259">M. singaporensis</emphasis>
</taxonomicName>
lacks this (
<bibRefCitation id="EFA94B73097A4F63FDD1FD64FD30ECAF" author="Sigurdsson JB" box="[563,755,658,680]" pageId="9" pageNumber="259" pagination="259 - 263" refId="ref15924" refString="Sigurdsson JB. 1991. A nudibranch, Murphydoris singaporensis, new genus and species, from Singapore mangroves (Mollusca: Opisthobranchia: Goniodorididae). Raffles Bulletin of Zoology 39: 259 - 263." type="journal article" year="1991">Sigurdsson, 1991</bibRefCitation>
; 
<bibRefCitation id="EFA94B73097A4F63FF73FD47FE74ECC1" author="Swennen C &amp; Buatip S" box="[145,439,689,711]" pageId="9" pageNumber="259" pagination="9 - 14" refId="ref16003" refString="Swennen C, Buatip S. 2012. Murphydoris singaporensis (Gastropoda: Nudibranchia: Goniodorididae) found in Thailand; with external and first internal description of the specimens. Strombus 19: 9 - 14." type="journal article" year="2012">Swennen &amp; Buatip, 2012</bibRefCitation>
). Internally, 
<taxonomicName id="4C384D01097A4F63FDB1FD44FD0BECC1" authorityName="Paz-Sedano &amp; Smirnoff &amp; Candás &amp; Gosliner &amp; Pola" authorityYear="2022" box="[595,712,689,711]" class="Gastropoda" family="Goniodorididae" genus="Murphydoris" kingdom="Animalia" order="Nudibranchia" pageId="9" pageNumber="259" phylum="Mollusca" rank="species" species="adusta">
<emphasis id="B94CEA90097A4F63FDB1FD44FD0BECC1" box="[595,712,689,711]" italics="true" pageId="9" pageNumber="259">M. adusta</emphasis>
</taxonomicName>
has only one row of teeth per side, which have two cusps, while 
<taxonomicName id="4C384D01097A4F63FF3BFD19FE62ED03" authorityName="Sigurdson" authorityYear="1991" box="[217,417,751,772]" class="Gastropoda" family="Goniodorididae" genus="Murphydoris" kingdom="Animalia" order="Nudibranchia" pageId="9" pageNumber="259" phylum="Mollusca" rank="species" species="singaporensis">
<emphasis id="B94CEA90097A4F63FF3BFD19FE62ED03" box="[217,417,751,772]" italics="true" pageId="9" pageNumber="259">M. singaporensis</emphasis>
</taxonomicName>
has one inner and one outer lateral tooth and the inner teeth have only a single cusp (
<bibRefCitation id="EFA94B73097A4F63FF7BFCDAFE7DED46" author="Swennen C &amp; Buatip S" box="[153,446,812,834]" pageId="9" pageNumber="259" pagination="9 - 14" refId="ref16003" refString="Swennen C, Buatip S. 2012. Murphydoris singaporensis (Gastropoda: Nudibranchia: Goniodorididae) found in Thailand; with external and first internal description of the specimens. Strombus 19: 9 - 14." type="journal article" year="2012">Swennen &amp; Buatip, 2012</bibRefCitation>
; 
<bibRefCitation id="EFA94B73097A4F63FE2EFCDAFD29ED45" author="Paz-Sedano S &amp; Diaz-Agras G &amp; Gosliner TM &amp; Pola M" box="[460,746,812,834]" pageId="9" pageNumber="259" refId="ref15263" refString="Paz-Sedano S, Diaz-Agras G, Gosliner TM, Pola M. 2021 b. Revealing morphological characteristics of Goniodorididae genera (Mollusca: Nudibranchia). Organisms Biodiversity and Evolutionary Research. Available at: https: // link. springer. com / article / 10.1007 / s 13127 - 021 - 00508 - w # citeas." type="book" year="2021">
Paz-Sedano 
<emphasis id="B94CEA90097A4F63FDBEFCDBFD55ED46" box="[604,662,812,834]" italics="true" pageId="9" pageNumber="259">et al.</emphasis>
, 2021b
</bibRefCitation>
). Moreover, the denticles of the masticatory margin are bigger and more numerous in 
<taxonomicName id="4C384D01097A4F63FE11FC9CFEB9ED9A" authority="(PazSedano et al., 2021 b)" baseAuthorityName="PazSedano" baseAuthorityYear="2021" class="Gastropoda" family="Goniodorididae" genus="Murphydoris" kingdom="Animalia" order="Nudibranchia" pageId="9" pageNumber="259" phylum="Mollusca" rank="species" species="singaporensis">
<emphasis id="B94CEA90097A4F63FE11FC9CFD75ED78" box="[499,694,874,895]" italics="true" pageId="9" pageNumber="259">M. singaporensis</emphasis>
(PazSedano 
<emphasis id="B94CEA90097A4F63FF08FC7FFEE2ED99" box="[234,289,904,926]" italics="true" pageId="9" pageNumber="259">et al.</emphasis>
, 2021b)
</taxonomicName>
than in 
<taxonomicName id="4C384D01097A4F63FE38FC7FFD88ED9A" authorityName="Paz-Sedano &amp; Smirnoff &amp; Candás &amp; Gosliner &amp; Pola" authorityYear="2022" box="[474,587,904,926]" class="Gastropoda" family="Goniodorididae" genus="Murphydoris" kingdom="Animalia" order="Nudibranchia" pageId="9" pageNumber="259" phylum="Mollusca" rank="species" species="adusta">
<emphasis id="B94CEA90097A4F63FE38FC7FFD88ED9A" box="[474,587,904,926]" italics="true" pageId="9" pageNumber="259">M. adusta</emphasis>
</taxonomicName>
. Regarding the reproductive system, in 
<taxonomicName id="4C384D01097A4F63FE7BFC51FD96EDBB" box="[409,597,935,956]" class="Gastropoda" family="Goniodorididae" genus="Murphydoris" kingdom="Animalia" order="Nudibranchia" pageId="9" pageNumber="259" phylum="Mollusca" rank="species" species="singaporensis">
<emphasis id="B94CEA90097A4F63FE7BFC51FD96EDBB" box="[409,597,935,956]" italics="true" pageId="9" pageNumber="259">M. singaporensis</emphasis>
</taxonomicName>
there is a duct arising from the base of the bursa copulatrix connecting with the uterine duct (
<bibRefCitation id="EFA94B73097A4F63FE7AFC12FD7AEDFE" author="Swennen C &amp; Buatip S" box="[408,697,996,1018]" pageId="9" pageNumber="259" pagination="9 - 14" refId="ref16003" refString="Swennen C, Buatip S. 2012. Murphydoris singaporensis (Gastropoda: Nudibranchia: Goniodorididae) found in Thailand; with external and first internal description of the specimens. Strombus 19: 9 - 14." type="journal article" year="2012">Swennen &amp; Buatip, 2012</bibRefCitation>
; PazSedano 
<emphasis id="B94CEA90097A4F63FF0BFBF5FEE3EA10" box="[233,288,1026,1048]" italics="true" pageId="9" pageNumber="259">et al.</emphasis>
, 2021b), whereas in 
<taxonomicName id="4C384D01097A4F63FDE7FBF5FDB5EA10" authorityName="Paz-Sedano &amp; Smirnoff &amp; Candás &amp; Gosliner &amp; Pola" authorityYear="2022" box="[517,630,1026,1048]" class="Gastropoda" family="Goniodorididae" genus="Murphydoris" kingdom="Animalia" order="Nudibranchia" pageId="9" pageNumber="259" phylum="Mollusca" rank="species" species="adusta">
<emphasis id="B94CEA90097A4F63FDE7FBF5FDB5EA10" box="[517,630,1026,1048]" italics="true" pageId="9" pageNumber="259">M. adusta</emphasis>
</taxonomicName>
the uterine duct begins at the middle of the vagina (
<tableCitation id="C6BA0339097A4F63FDB0FBD7FD60EA31" box="[594,675,1057,1079]" captionStart="Table 3" captionStartId="14.[143,207,514,536]" captionTargetPageId="14" captionText="Table 3. Comparative morphology among new species of Murphydoris" httpUri="http://table.plazi.org/id/DF47660A097D4F64FF6DFDF4FCB2EC1F" pageId="9" pageNumber="259" tableUuid="DF47660A097D4F64FF6DFDF4FCB2EC1F">Table 3</tableCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>