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<document id="8FE86637680DC99A4B9822CCE5AE28C8" ID-DOI="10.1206/0003-0082(2000)287&lt;0001:ROTNTT&gt;2.0.CO;2" ID-ISSN="0003-0082" ID-Zenodo-Dep="10110744" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630265040447" checkinUser="carolina" docAuthor="SMETANA, ALEŠ &amp; DAVIES, ANTHONY" docDate="2000" docId="0384878FD834FB38FC496BE50D6FF917" docLanguage="en" docName="N3287.pdf" docOrigin="American Museum Novitates 3287" docSource="http://www.bioone.org/doi/abs/10.1206/0003-0082%282000%29287%3C0001%3AROTNTT%3E2.0.CO%3B2" docStyle="DocumentStyle:059C9B7006157ECE636E30DEF6793B3A.3:AmMusNovit.2000-2010.journal_article.type2" docStyleId="059C9B7006157ECE636E30DEF6793B3A" docStyleName="AmMusNovit.2000-2010.journal_article.type2" docStyleVersion="3" docTitle="Protogoerius Coiffait 1956" docType="treatment" docVersion="3" masterDocId="FFBDFFF7D829FB25FFD86B530951FFD4" masterDocTitle="Reclassification of the North Temperate Taxa Associated with Staphylinus Sensu Lato, Including Comments on Relevant Subtribes of Staphylinini (Coleoptera: Staphylinidae)" masterLastPageNumber="1" masterPageNumber="1" pageNumber="1" updateTime="1699682540511" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="51EFB2A9F5DED4BF24A942774003FB0C">Reclassification of the North Temperate Taxa Associated with Staphylinus Sensu Lato, Including Comments on Relevant Subtribes of Staphylinini (Coleoptera: Staphylinidae)</mods:title>
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<mods:namePart id="FAB73E2BE6371C37BE9340B743574C26">SMETANA, ALEŠ</mods:namePart>
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<mods:namePart id="F4A336A6C157D395562E907E6DE15390">DAVIES, ANTHONY</mods:namePart>
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<mods:typeOfResource id="A4E9D46D7C22BA1BE9660752CAFB4C8B">text</mods:typeOfResource>
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<mods:title id="F6EB79261C8E77444A060F5F9AA5757B">American Museum Novitates</mods:title>
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<mods:date id="B1564354543BDA46B11F99BC39EB8BD6">2000</mods:date>
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<mods:number id="47F5F760B12117CFCF862894D3F62E9A">2000-02-02</mods:number>
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<treatment id="0384878FD834FB38FC496BE50D6FF917" LSID="urn:lsid:plazi:treatment:0384878FD834FB38FC496BE50D6FF917" httpUri="http://treatment.plazi.org/id/0384878FD834FB38FC496BE50D6FF917" pageId="29">
<subSubSection id="C3376512D834FB38FC496BE50D8BFF19" box="[913,1242,182,206]" pageId="29" type="nomenclature">
<paragraph id="8B923699D834FB38FC496BE50D8BFF19" blockId="29.[913,1242,182,206]" box="[913,1242,182,206]" pageId="29">
<heading id="D0DA81F5D834FB38FC496BE50D8BFF19" box="[913,1242,182,206]" centered="true" fontSize="36" level="2" pageId="29" reason="8">
<taxonomicName id="4C2D4D1AD834FB38FC496BE50D8BFF19" ID-CoL="8XP6S" authority="Coiffait, 1956" authorityName="Coiffait" authorityYear="1956" box="[913,1242,182,206]" class="Insecta" family="Staphylinidae" genus="Protogoerius" kingdom="Animalia" order="Coleoptera" pageId="29" phylum="Arthropoda" rank="genus">
<emphasis id="B959EA8BD834FB38FC496BE50D7DFF1A" bold="true" box="[913,1068,182,206]" italics="true" pageId="29">Protogoerius</emphasis>
Coiffait, 1956
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3376512D834FB38FC936BA10DC0FE92" pageId="29" type="materials_examined">
<paragraph id="8B923699D834FB38FC936BA10DC0FE92" blockId="29.[814,1343,242,1731]" pageId="29">
<collectionCode id="ED3CAE5CD834FB38FC936BA10ABEFEDC" box="[843,1007,242,265]" pageId="29">
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SPECIES
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:
<taxonomicName id="4C2D4D1AD834FB38FBD36BA10C6FFEDD" authorityName="Marsham" authorityYear="1802" box="[1035,1342,242,265]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="29" phylum="Arthropoda" rank="species" species="brachypterus">
<emphasis id="B959EA8BD834FB38FBD36BA10C6FFEDD" box="[1035,1342,242,265]" italics="true" pageId="29">Staphylinus brachypterus</emphasis>
</taxonomicName>
Brulle´, 1839 (nec Geoffroy, 1785) (=
<taxonomicName id="4C2D4D1AD834FB38FAD16A420AA2FE92" authorityName="SMETANA &amp; DAVIES" authorityYear="2000" class="Insecta" family="Staphylinidae" genus="Protogoerius" kingdom="Animalia" order="Coleoptera" pageId="29" phylum="Arthropoda" rank="species" species="brullei" status="nom. nov.">
<emphasis id="B959EA8BD834FB38FAD16A420AA2FE92" italics="true" pageId="29">Protogoerius brullei</emphasis>
</taxonomicName>
,
<emphasis id="B959EA8BD834FB38FBDB6A7C0DD1FE93" bold="true" box="[1027,1152,303,327]" pageId="29">
<taxonomicNameLabel id="A26A57F0D834FB38FBDB6A7C0DD1FE93" box="[1027,1152,303,327]" pageId="29">nom. nov.</taxonomicNameLabel>
</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="C3376512D834FB38FC936A1E0D6FF917" pageId="29" type="discussion">
<paragraph id="8B923699D834FB38FC936A1E0D7FFDCF" blockId="29.[814,1343,242,1731]" pageId="29">
COMMENTS: The single species of this genus has the habitus of an
<taxonomicName id="4C2D4D1AD834FB38FBB96A3F0DE9FE57" authorityName="SMETANA &amp; DAVIES" authorityYear="2000" box="[1121,1208,364,387]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="29" phylum="Arthropoda" rank="genus">
<emphasis id="B959EA8BD834FB38FBB96A3F0DE9FE57" box="[1121,1208,364,387]" italics="true" pageId="29">Ocypus</emphasis>
</taxonomicName>
; it has two main identifying character states: (
<bibRefCitation id="EFBC4B68D834FB38FB1D6AD90D80FE75" author="Natural History Museum, London" box="[1221,1233,394,417]" pageId="29" pagination="204" refId="ref25242" refString="1. Ocypus almorensis (Cameron, 1932). Cameron (1932: 204) described the species (in Staphylinus) from specimens from '' West Almora: Kali Valley, alt. 9000 feet' ' and '' Punjab: Triun, alt. 9300 feet. '' We were able to study one of the male syntypes in the Natural History Museum, London, England, labelled as follows: '' Triun, 9,300 ft Dharm- sala. Punjab. H. G. C. / M. Cameron. Bequest. B. M. 1955 - 147. / Staphylinus (Ocypus) almorensis. '' The specimen is in very good shape. It was dissected, and the aedeagus and the genital segment were glued to the plate with the beetle. It is hereby designated as the lectotype of Staphylinus almorensis; the label '' LECTOTYPE Staphylinus almorensis Cameron A. Smetana des. 1999 '' was attached to it. The specimen cannot be distinguished from those of Ocypus helleni J. Muller that occur in northern Pakistan and in Kashmir. The name O. almorensis becomes a junior synonym of O. helleni (syn. nov.). The corresponding determination label was attached to the specimen." type="journal article" year="1955">1</bibRefCitation>
) gula tumid almost to fine basal carina, transverse impression very faint (
<figureCitation id="13162A1CD834FB38FB906A940DFCFE0A" box="[1096,1197,455,478]" captionStart="Figs" captionStartId="85.[264,312,1477,1498]" captionTargetBox="[256,1323,176,1456]" captionTargetId="figure-34@85.[256,1323,176,1456]" captionTargetPageId="85" captionText="Figs. 146151. 146. Protogoerius brullei, underside of head, showing gula. 147. Protogoerius brullei, left mandible, dorsal view. 148. Protogoerius brullei, maxillary (left) and labial palpus (right). 149. Ocypus tenebricosus, maxilla, showing setae on palpifer. 150. Ocypus tenebricosus, maxillary palpus. 151. Ocypus tenebricosus, labial palpus." figureDoi="http://doi.org/10.5281/zenodo.10110800" httpUri="https://zenodo.org/record/10110800/files/figure.png" pageId="29">fig. 146</figureCitation>
); (
<bibRefCitation id="EFBC4B68D834FB38FB096A940DB0FE0A" author="Ocypus" box="[1233,1249,455,478]" pageId="29" pagination="170" refId="ref25465" refString="2. Ocypus auricomus (Lindberg, 1953: 4), described in Staphylinus, is preoccupied by Staphylinus auricomus Brulle', 1842: pl. 5, fig. 6, now a synonym of Glenus chrysis (Gravenhorst, 1806), and by Staphylinus auricomus Cameron, 1929: 65. The name Ocypus mateui (Coiffait, 1954: 170), listed at present as a junior synonym of O. auricomus, becomes the valid name of this species (stat. nov.)." type="journal article" year="1842">2</bibRefCitation>
) dorsal portion of median lobe of aedeagus almost entirely membranous.
</paragraph>
<paragraph id="8B923699D834FB38FC9369710DEDFA7B" blockId="29.[814,1343,242,1731]" pageId="29">
Further character states include: mandibles each with one strong, broadly triangular tooth on medial edge in front of a deep, angular indentation, with dorsolateral ridge not distinctly removed from lateral margin (
<figureCitation id="13162A1CD834FB38FACF69CF0A0CFD05" captionStart="Figs" captionStartId="85.[264,312,1477,1498]" captionTargetBox="[256,1323,176,1456]" captionTargetId="figure-34@85.[256,1323,176,1456]" captionTargetPageId="85" captionText="Figs. 146151. 146. Protogoerius brullei, underside of head, showing gula. 147. Protogoerius brullei, left mandible, dorsal view. 148. Protogoerius brullei, maxillary (left) and labial palpus (right). 149. Ocypus tenebricosus, maxilla, showing setae on palpifer. 150. Ocypus tenebricosus, maxillary palpus. 151. Ocypus tenebricosus, labial palpus." figureDoi="http://doi.org/10.5281/zenodo.10110800" httpUri="https://zenodo.org/record/10110800/files/figure.png" pageId="29">fig. 147</figureCitation>
); palpifer with one subapical and three shorter apical setae; apical segment of maxillary and labial palpi moderately elongate, distinctly fusiform, only that of labial palpus setose (
<figureCitation id="13162A1CD834FB38FC5768670AA9FC9F" box="[911,1016,820,843]" captionStart="Figs" captionStartId="85.[264,312,1477,1498]" captionTargetBox="[256,1323,176,1456]" captionTargetId="figure-34@85.[256,1323,176,1456]" captionTargetPageId="85" captionText="Figs. 146151. 146. Protogoerius brullei, underside of head, showing gula. 147. Protogoerius brullei, left mandible, dorsal view. 148. Protogoerius brullei, maxillary (left) and labial palpus (right). 149. Ocypus tenebricosus, maxilla, showing setae on palpifer. 150. Ocypus tenebricosus, maxillary palpus. 151. Ocypus tenebricosus, labial palpus." figureDoi="http://doi.org/10.5281/zenodo.10110800" httpUri="https://zenodo.org/record/10110800/files/figure.png" pageId="29">fig. 148</figureCitation>
); prostheca parallel­sided, elongate with long ciliae only at apex; nuchal ridge present only dorsally, becoming obsolete well before base of infraorbital ridge laterally; dorsal basal ridge very short, ending at very base of infraorbital ridge; superior and inferior lines of hypomeron run very closely parallel for some distance anteriorly, gradually merging along prosternal margin; prothoracic epimeron absent; prosternum with short transverse ridge on keel behind two macrosetae; mesocoxal acetabulum deeply depressed, with straight carina sloping obliquely behind intercoxal process; margins of anterior metasternal projection converging at a highly obtuse angle, projection very short; metasternum very short, at an oblique angle to plane of body, with no visible medial line; dorsal apicolateral lobe of hind coxa with only one or two very fine spines; empodial setae very long.
</paragraph>
<paragraph id="8B923699D834FB38FC936EE50D6FF917" blockId="29.[814,1343,242,1731]" pageId="29">
Brullés (1839: 59) name
<taxonomicName id="4C2D4D1AD834FB38FBA46EE40ACEFA38" authorityName="Marsham" authorityYear="1802" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="29" phylum="Arthropoda" rank="species" species="brachypterus">
<emphasis id="B959EA8BD834FB38FBA46EE40ACEFA38" italics="true" pageId="29">Staphylinus brachypterus</emphasis>
</taxonomicName>
is preoccupied by
<taxonomicName id="4C2D4D1AD834FB38FBA56E860DF5F9DF" authority="Geoffroy, 1785: 176" authorityName="Geoffroy" authorityPageNumber="176" authorityYear="1785" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="29" phylum="Arthropoda" rank="species" species="brachypterus">
<emphasis id="B959EA8BD834FB38FBA56E860ACEF9DF" italics="true" pageId="29">Staphylinus brachypterus</emphasis>
Geoffroy, 1785: 176
</taxonomicName>
, now a synonym of
<taxonomicName id="4C2D4D1AD834FB38FC7B6D400C68F9FD" authority="(Goeze, 1777)" baseAuthorityName="Goeze" baseAuthorityYear="1777" box="[931,1337,1554,1578]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="29" phylum="Arthropoda" rank="species" species="curtula">
<emphasis id="B959EA8BD834FB38FC7B6D400D2CF9FE" box="[931,1149,1555,1578]" italics="true" pageId="29">Aleochara curtula</emphasis>
(Goeze, 1777)
</taxonomicName>
, and by
<taxonomicName id="4C2D4D1AD834FB38FC576D620AFFF9B3" authority="Marsham, 1802: 510" authorityName="Marsham" authorityPageNumber="510" authorityYear="1802" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="29" phylum="Arthropoda" rank="species" species="brachypterus">
<emphasis id="B959EA8BD834FB38FC576D620DEDF99C" box="[911,1212,1585,1608]" italics="true" pageId="29">Staphylinus brachypterus</emphasis>
Marsham, 1802: 510
</taxonomicName>
, now a synonym of
<taxonomicName id="4C2D4D1AD834FB38FB1F6D030D23F951" authority="Gravenhorst, 1802" authorityName="Gravenhorst" authorityYear="1802" class="Insecta" family="Staphylinidae" genus="Aploderus" kingdom="Animalia" order="Coleoptera" pageId="29" phylum="Arthropoda" rank="species" species="caelatus">
<emphasis id="B959EA8BD834FB38FB1F6D030ADEF952" italics="true" pageId="29">Aploderus caelatus</emphasis>
Gravenhorst, 1802
</taxonomicName>
. The replacement name
<taxonomicName id="4C2D4D1AD834FB38FCA66DDE0D27F970" authorityName="SMETANA &amp; DAVIES" authorityYear="2000" box="[894,1142,1677,1700]" class="Insecta" family="Staphylinidae" genus="Protogoerius" kingdom="Animalia" order="Coleoptera" pageId="29" phylum="Arthropoda" rank="species" species="brullei" status="nom. nov.">
<emphasis id="B959EA8BD834FB38FCA66DDE0D27F970" box="[894,1142,1677,1700]" italics="true" pageId="29">Protogoerius brullei</emphasis>
</taxonomicName>
,
<emphasis id="B959EA8BD834FB38FB566DDE0C43F971" bold="true" box="[1166,1298,1677,1701]" pageId="29">
<taxonomicNameLabel id="A26A57F0D834FB38FB566DDE0C43F971" box="[1166,1298,1677,1701]" pageId="29">nom. nov.</taxonomicNameLabel>
</emphasis>
, is hereby proposed for it.
</paragraph>
</subSubSection>
</treatment>
</document>