treatments-xml/data/03/81/87/038187D2FFE33652FC7372F616FB6DB6.xml

336 lines
44 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="4D136A3CF9DE90356DE480B5633780F2" ID-CLB-Dataset="296796" ID-DOI="10.1093/zoolinnean/zlad080" ID-GBIF-Dataset="c4c0762a-b832-4c15-b02e-3e6ff1e48ef4" ID-ISSN="0024-4082" ID-Zenodo-Dep="11240237" ID-ZooBank="E024A072-8083-4D91-BC40-A420D1280BEDC" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="juliana" checkinTime="1716361565008" checkinUser="plazi" docAuthor="Shofner, Ryan &amp; Cassis, Gerasimos" docDate="2024" docId="038187D2FFE33652FC7372F616FB6DB6" docLanguage="en" docName="zlad080.pdf" docOrigin="Zoological Journal of the Linnean Society 200 (2)" docSource="http://dx.doi.org/10.1093/zoolinnean/zlad080" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.6:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="6" docTitle="Tingidae , Drake and Ruhoff 1965" docType="key" docVersion="1" lastPageNumber="376" masterDocId="FFB8FFAAFFEC3642FFD4702E13626C67" masterDocTitle="Phylogenetics and taxonomy of the Indo-Australian genus Ulonemia sensu Drake (Hemiptera: Tingidae), with the recognition of new genera and species collected from Proteaceae in Australia" masterLastPageNumber="425" masterPageNumber="360" pageNumber="375" updateTime="1728431177839" updateUser="juliana">
<mods:mods id="FBE32619C21D80B8C95ECA17680185AB" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E6963A858BCC81C95FC4F539BC26FE2A">
<mods:title id="6F9D1225DE80E5EF7CF80B76FC1EB042">Phylogenetics and taxonomy of the Indo-Australian genus Ulonemia sensu Drake (Hemiptera: Tingidae), with the recognition of new genera and species collected from Proteaceae in Australia</mods:title>
</mods:titleInfo>
<mods:name id="FA36D6A39C785E7CC43375CB659C38BC" type="personal">
<mods:role id="F6634162E4B816C2C169186772054827">
<mods:roleTerm id="BA95AE8218FABD3B4B9AAC90C7B40443">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7D26F6900B5994B7CA696B049302FE9F">Shofner, Ryan</mods:namePart>
<mods:affiliation id="21FBE8C423F71ED0B7628223A87EDC67">Evolution and Ecology Research Centre, School of Biological, Earth, and Environmental Sciences, University of New South Wales, UNSW Sydney, NSW 2052, Australia</mods:affiliation>
<mods:nameIdentifier id="FB09FCA96307103481C32485E9D59E80" type="email">r.shofner@unsw.edu.au</mods:nameIdentifier>
</mods:name>
<mods:name id="0C80F7757FEC069351C135D6474B745F" type="personal">
<mods:role id="DFDACE880A65E72A0691BD0A5FC4274E">
<mods:roleTerm id="731C05D25EF8E45130AB4C4C31C776F0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5BF526127401E776B820803D87F33355">Cassis, Gerasimos</mods:namePart>
<mods:affiliation id="C89BAC2A0130846854E5915FCBD27BDB">Evolution and Ecology Research Centre, School of Biological, Earth, and Environmental Sciences, University of New South Wales, UNSW Sydney, NSW 2052, Australia</mods:affiliation>
</mods:name>
<mods:typeOfResource id="A2870006474AA9F3FD78B3C5B2034DAF">text</mods:typeOfResource>
<mods:relatedItem id="B97396CECA77FB11F63FAA62C32277BE" type="host">
<mods:titleInfo id="87396157A5851B463A8284911CF01D44">
<mods:title id="CF7E0D18B8D13FE68DEBF5B2E79F2B71">Zoological Journal of the Linnean Society</mods:title>
</mods:titleInfo>
<mods:part id="183B6B18DBCB0758A3D917F4E33889B9">
<mods:date id="BA9A85856447EE987005B4A1218F57CB">2024</mods:date>
<mods:detail id="15B78A05CB08BC88E23B5D96793F3F53" type="pubDate">
<mods:number id="3F2FB3E0F144E1D7052FBCAB7E81BAE7">2023-08-28</mods:number>
</mods:detail>
<mods:detail id="67A5CE978EFE1B8CB6A07F39D70236D9" type="volume">
<mods:number id="0B91BE899E7F9D73F5602D088DF61BD2">200</mods:number>
</mods:detail>
<mods:detail id="F305BAB32D4DA90C0FF20B4FA6032657" type="issue">
<mods:number id="F54D87C4D760BCADD4CAE58F86D9E9B5">2</mods:number>
</mods:detail>
<mods:extent id="6F7C11E8557D75F0436C376D0C80FFE6" unit="page">
<mods:start id="5F5B3829F3F0D9BC23EEBBEAEEB53372">360</mods:start>
<mods:end id="1E5279407B11F35C589ED785EEE8E35F">425</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="CA2C77200F4BA38AB3D0AC27BBAAFCB0">
<mods:url id="83DCF3AFD29707659A11E6FEC09C2805">http://dx.doi.org/10.1093/zoolinnean/zlad080</mods:url>
</mods:location>
<mods:classification id="CEA4FE709D9516A2BF4B818F2A353560">journal article</mods:classification>
<mods:identifier id="84418808FAC1F77FDB91C0A03CEE6320" type="CLB-Dataset">296796</mods:identifier>
<mods:identifier id="04F775E8AA710E4F4A95B4DEBFAAD8B4" type="DOI">10.1093/zoolinnean/zlad080</mods:identifier>
<mods:identifier id="84AB9D8F5A83FD91271B6249A1639094" type="GBIF-Dataset">c4c0762a-b832-4c15-b02e-3e6ff1e48ef4</mods:identifier>
<mods:identifier id="9863C419E904A2F699D47CC8327DF88B" type="ISSN">0024-4082</mods:identifier>
<mods:identifier id="71D3DC1F7D2C9CCCC1715984423FC25E" type="Zenodo-Dep">11240237</mods:identifier>
<mods:identifier id="88D64120FE700AA0CFED4DD63152BA4F" type="ZooBank">E024A072-8083-4D91-BC40-A420D1280BEDC</mods:identifier>
</mods:mods>
<treatment id="038187D2FFE33652FC7372F616FB6DB6" ID-DOI="http://doi.org/10.5281/zenodo.11277945" ID-GBIF-Taxon="230977586" ID-Zenodo-Dep="11277945" LSID="urn:lsid:plazi:treatment:038187D2FFE33652FC7372F616FB6DB6" httpUri="http://treatment.plazi.org/id/038187D2FFE33652FC7372F616FB6DB6" lastPageId="16" lastPageNumber="376" pageId="15" pageNumber="375">
<subSubSection id="C332654FFFE3364DFC7372F616366E95" box="[935,1364,728,754]" pageId="15" pageNumber="375" type="nomenclature">
<paragraph id="8B9736C4FFE3364DFC7372F616366E95" blockId="15.[935,1364,728,754]" box="[935,1364,728,754]" pageId="15" pageNumber="375">
<emphasis id="B95CEAD6FFE3364DFC7372F616366E95" box="[935,1364,728,754]" italics="true" pageId="15" pageNumber="375">
<taxonomicName id="4C284D47FFE3364DFC7372F610986E95" ID-CoL="H9W" ID-ENA="236450" authorityName=", Drake and Ruhoff" authorityYear="1965" box="[935,1018,728,754]" class="Insecta" family="Tingidae" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="375" phylum="Arthropoda" rank="family">Tingidae</taxonomicName>
genera associated with the
<taxonomicName id="4C284D47FFE3364DFB2572F716366E95" box="[1265,1364,729,754]" class="Magnoliopsida" family="Proteaceae" kingdom="Plantae" order="Proteales" pageId="15" pageNumber="375" phylum="Tracheophyta" rank="family">Proteaceae</taxonomicName>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C332654FFFE3364DFCED72D110CC68E9" pageId="15" pageNumber="375" type="description">
<paragraph id="8B9736C4FFE3364DFCED72D110CC68E9" blockId="15.[825,1474,767,1166]" pageId="15" pageNumber="375">
Tingid clades are often associated with a narrow range of plant genera (
<bibRefCitation id="EFB94B35FFE3364DFC44733017C66F51" author="Cassis G &amp; Symonds CL" box="[912,1188,798,822]" pageId="15" pageNumber="375" pagination="1 - 63" refId="ref48140" refString="Cassis G, Symonds CL. Systematics, biogeography and host plant associations of the lace bug genus Lasiacantha Stal in Australia (Insecta: Hemiptera: Heteroptera: Tingidae). Zootaxa 2011; 2818: 1 - 63. https: // doi. org / 10.11646 / zootaxa. 2818.1.1" type="journal article" year="2011">Cassis and Symonds 2011</bibRefCitation>
,
<bibRefCitation id="EFB94B35FFE3364DFB67733016086F51" author="Cassis G &amp; Koenig P &amp; Symonds CL" box="[1203,1386,798,822]" pageId="15" pageNumber="375" pagination="1 - 95" refId="ref48246" refString="Cassis G, Koenig P, Symonds CL et al. Systematics and host plant associations of the Australian lace bug genus Nethersia (Insecta: Heteroptera: Tingidae), including the description of eighteen new species. Insect Systematics and Evolution 2017; 48: 1 - 95. https: // doi. org / 10.1163 / 1876312 X- 47052151" type="journal article" year="2017">
Cassis
<emphasis id="B95CEAD6FFE3364DFB2F733116456F51" box="[1275,1319,798,822]" italics="true" pageId="15" pageNumber="375">et al</emphasis>
. 2017
</bibRefCitation>
), which are often closely related. Based on recent surveys (
<bibRefCitation id="EFB94B35FFE3364DFA9E7313100F6F12" author="Cassis G &amp; Wall M &amp; Schuh R" pageId="15" pageNumber="375" pagination="193" refId="ref48193" refString="Cassis G, Wall M, Schuh R. Insect biodiversity and industrialising the taxonomic process: the plant bug case study (Insecta: Heteroptera: Miridae). Systematics Association Special Volume 2007; 72: 193. https: // doi. org / 10.1201 / 9781420009538. ch 13" type="journal article" year="2007">
Cassis
<emphasis id="B95CEAD6FFE3364DFA45731016DE6F32" box="[1425,1468,829,853]" italics="true" pageId="15" pageNumber="375">et al</emphasis>
. 2007
</bibRefCitation>
) numerous tingid taxa are associated with the plant family
<taxonomicName id="4C284D47FFE3364DFCED735210C86FF4" box="[825,938,892,915]" class="Magnoliopsida" family="Proteaceae" kingdom="Plantae" order="Proteales" pageId="15" pageNumber="375" phylum="Tracheophyta" rank="family">Proteaceae</taxonomicName>
. Host records from the PBI database were harvested for a list of tingid taxa associated with
<taxonomicName id="4C284D47FFE3364DFB1A73B2165F6FD4" box="[1230,1341,924,947]" class="Magnoliopsida" family="Proteaceae" kingdom="Plantae" order="Proteales" pageId="15" pageNumber="375" phylum="Tracheophyta" rank="family">Proteaceae</taxonomicName>
(Supporting Information, Table S4). Three genus-group taxa were recovered from our molecular analysis that are specialized on the
<taxonomicName id="4C284D47FFE3364DFCED73D410C86876" box="[825,938,1018,1041]" class="Magnoliopsida" family="Proteaceae" kingdom="Plantae" order="Proteales" pageId="15" pageNumber="375" phylum="Tracheophyta" rank="family">Proteaceae</taxonomicName>
. In addition, there are several genera that have one to several species associated with the
<taxonomicName id="4C284D47FFE3364DFB6C743716456857" box="[1208,1319,1049,1072]" class="Magnoliopsida" family="Proteaceae" kingdom="Plantae" order="Proteales" pageId="15" pageNumber="375" phylum="Tracheophyta" rank="family">Proteaceae</taxonomicName>
but are not exclusively associated with the family. The following key accounts for all genera that have at least one species associated with the
<taxonomicName id="4C284D47FFE3364DFCED745910C868E9" box="[825,938,1143,1166]" class="Magnoliopsida" family="Proteaceae" kingdom="Plantae" order="Proteales" pageId="15" pageNumber="375" phylum="Tracheophyta" rank="family">Proteaceae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C332654FFFE33652FDEF74E716FB6DB6" lastPageId="16" lastPageNumber="376" pageId="15" pageNumber="375" type="key">
<paragraph id="8B9736C4FFE3364DFDEF74E7176A6884" blockId="15.[155,1448,1224,1971]" box="[571,1032,1224,1251]" pageId="15" pageNumber="375">
Key to
<taxonomicName id="4C284D47FFE3364DFD5374E611846885" ID-CoL="H9W" ID-ENA="236450" authority="Drake and Ruhoff 1965" authorityName="Drake and Ruhoff" authorityYear="1965" box="[647,742,1224,1250]" class="Insecta" family="Tingidae" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="375" phylum="Arthropoda" rank="family">Tingidae</taxonomicName>
associated with
<taxonomicName id="4C284D47FFE3364DFC4074E7176A6884" box="[916,1032,1225,1251]" class="Magnoliopsida" family="Proteaceae" kingdom="Plantae" order="Proteales" pageId="15" pageNumber="375" phylum="Tracheophyta" rank="family">Proteaceae</taxonomicName>
</paragraph>
<key id="FFB961F5FFE33652FF4F752716FB6DB6" lastPageId="16" lastPageNumber="376" pageId="15" pageNumber="375">
<paragraph id="8B9736C4FFE3364DFF4F752716C56926" blockId="15.[155,1448,1224,1971]" pageId="15" pageNumber="375">
<keyLead id="30D995F1FFE3364DFF4F752716C56926" pageId="15" pageNumber="375">
<keyStep id="30DC2E61FFE3364DFF4F752716C56926" pageId="15" pageNumber="375">
1 Paranota highly modified, extending dorsally onto pronotal disc and exceeding lateral carinae, inflated to produce paired bulbous structures (
<figureCitation id="13132A41FFE3364DFE4D7507129E6927" box="[409,508,1321,1345]" captionStart="Figure 4" captionStartId="14.[113,178,1600,1624]" captionTargetBox="[151,1421,144,1571]" captionTargetId="figure-276@14.[786,1422,619,862]" captionTargetPageId="14" captionText="Figure 4. Scanning electron micrographs of Cysteochila sp. (A, B), Engynoma spinicollis (Horváth 1925) (C), Epimixia sp. (D), and Eritingis trivirgata (Horváth 1925) (EG.). Abbreviations: fmr, femur; frn, frons; mdsp, medial spine; para, paranotum; pdsp, paired spines; pmg, peritreme of the metathoracic gland, tro, trochanter." figureDoi="http://doi.org/10.5281/zenodo.11240247" httpUri="https://zenodo.org/record/11240247/files/figure.png" pageId="15" pageNumber="375">Fig. 4A, B</figureCitation>
,
<emphasis id="B95CEAD6FFE3364DFDD3750711566927" box="[519,564,1321,1344]" italics="true" pageId="15" pageNumber="375">para</emphasis>
) .........................................................................................................................................................2
</keyStep>
</keyLead>
</paragraph>
<paragraph id="8B9736C4FFE3364DFF4F756616C56907" blockId="15.[155,1448,1224,1971]" box="[155,1447,1352,1376]" pageId="15" pageNumber="375">
<keyLead id="30D995F1FFE3364DFF4F756616C56907" box="[155,1447,1352,1376]" pageId="15" pageNumber="375">
<keyStep id="30DC2E61FFE3364DFF4F756616C56907" box="[155,1447,1352,1376]" pageId="15" pageNumber="375">2 Paranota not as above ....................................................................................................................................................................................3</keyStep>
</keyLead>
</paragraph>
<paragraph id="8B9736C4FFE3364DFF4F754916C56918" blockId="15.[155,1448,1224,1971]" box="[155,1447,1383,1407]" pageId="15" pageNumber="375">
<keyLead id="30D995F1FFE3364DFF4F754916C56918" box="[155,1447,1383,1407]" pageId="15" pageNumber="375">
<keyStep id="30DC2E61FFE3364DFF4F754916C56918" box="[155,1447,1383,1407]" pageId="15" pageNumber="375">
3 Frons porrect, extending beyond antenniferous tubercles (
<figureCitation id="13132A41FFE3364DFCC8754910E26918" box="[796,896,1383,1407]" captionStart="Figure 4" captionStartId="14.[113,178,1600,1624]" captionTargetBox="[151,1421,144,1571]" captionTargetId="figure-276@14.[786,1422,619,862]" captionTargetPageId="14" captionText="Figure 4. Scanning electron micrographs of Cysteochila sp. (A, B), Engynoma spinicollis (Horváth 1925) (C), Epimixia sp. (D), and Eritingis trivirgata (Horváth 1925) (EG.). Abbreviations: fmr, femur; frn, frons; mdsp, medial spine; para, paranotum; pdsp, paired spines; pmg, peritreme of the metathoracic gland, tro, trochanter." figureDoi="http://doi.org/10.5281/zenodo.11240247" httpUri="https://zenodo.org/record/11240247/files/figure.png" pageId="15" pageNumber="375">Fig. 4A, B</figureCitation>
,
<emphasis id="B95CEAD6FFE3364DFC5E754910C46918" box="[906,934,1383,1407]" italics="true" pageId="15" pageNumber="375">frn</emphasis>
) ...................................................
<taxonomicName id="4C284D47FFE3364DFB01754916C56918" authority="Stal 1873" authorityName=", Stal" authorityYear="1873" box="[1237,1447,1383,1407]" class="Insecta" family="Tingidae" genus="Cysteochila" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="375" phylum="Arthropoda" rank="genus">
<emphasis id="B95CEAD6FFE3364DFB01754916206918" box="[1237,1346,1383,1407]" italics="true" pageId="15" pageNumber="375">Cysteochila</emphasis>
<bibRefCitation id="EFB94B35FFE3364DFA9C754916C56918" author="Stal C" box="[1352,1447,1383,1407]" pageId="15" pageNumber="375" pagination="1 - 163" refId="ref50672" refString="Stal C. Enumeratio hemipterorum. Bidrag till en forteckning ofver aller hittills kanda Hemiptera, jemte systematiska meddelanden. 3. Kongliga Svenska Vetenskaps-Academiens Nya Handlingar, Stockholm 1873; 11: 1 - 163." type="journal article" year="1873">Stål 1873</bibRefCitation>
</taxonomicName>
</keyStep>
</keyLead>
</paragraph>
<paragraph id="8B9736C4FFE3364DFF4F75A916C569F8" blockId="15.[155,1448,1224,1971]" box="[155,1447,1414,1439]" pageId="15" pageNumber="375">
<keyLead id="30D995F1FFE3364DFF4F75A916C569F8" box="[155,1447,1414,1439]" pageId="15" pageNumber="375">
<keyStep id="30DC2E61FFE3364DFF4F75A916C569F8" box="[155,1447,1414,1439]" pageId="15" pageNumber="375">
4 Frons flattened to slightly protruding, but never exceeding antenniferous tubercles (
<figureCitation id="13132A41FFE3364DFBC875A9170569F8" box="[1052,1127,1415,1439]" captionStart="Figure 4" captionStartId="14.[113,178,1600,1624]" captionTargetBox="[151,1421,144,1571]" captionTargetId="figure-276@14.[786,1422,619,862]" captionTargetPageId="14" captionText="Figure 4. Scanning electron micrographs of Cysteochila sp. (A, B), Engynoma spinicollis (Horváth 1925) (C), Epimixia sp. (D), and Eritingis trivirgata (Horváth 1925) (EG.). Abbreviations: fmr, femur; frn, frons; mdsp, medial spine; para, paranotum; pdsp, paired spines; pmg, peritreme of the metathoracic gland, tro, trochanter." figureDoi="http://doi.org/10.5281/zenodo.11240247" httpUri="https://zenodo.org/record/11240247/files/figure.png" pageId="15" pageNumber="375">Fig. 4G</figureCitation>
,
<emphasis id="B95CEAD6FFE3364DFBA675A917EC69F8" box="[1138,1166,1415,1439]" italics="true" pageId="15" pageNumber="375">frn</emphasis>
) ..........
<taxonomicName id="4C284D47FFE3364DFB0375A916C569F8" authority="Stal 1873" authorityName="Stal" authorityYear="1873" box="[1239,1447,1414,1439]" class="Insecta" family="Tingidae" genus="Oncophysa" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="375" phylum="Arthropoda" rank="genus">
<emphasis id="B95CEAD6FFE3364DFB0375A9162169F8" box="[1239,1347,1415,1439]" italics="true" pageId="15" pageNumber="375">Oncophysa</emphasis>
<bibRefCitation id="EFB94B35FFE3364DFA9C75A816C569F8" author="Stal C" box="[1352,1447,1414,1439]" pageId="15" pageNumber="375" pagination="1 - 163" refId="ref50672" refString="Stal C. Enumeratio hemipterorum. Bidrag till en forteckning ofver aller hittills kanda Hemiptera, jemte systematiska meddelanden. 3. Kongliga Svenska Vetenskaps-Academiens Nya Handlingar, Stockholm 1873; 11: 1 - 163." type="journal article" year="1873">Stål 1873</bibRefCitation>
</taxonomicName>
</keyStep>
</keyLead>
</paragraph>
<paragraph id="8B9736C4FFE3364DFF4F758816C569D9" blockId="15.[155,1448,1224,1971]" box="[155,1447,1446,1470]" pageId="15" pageNumber="375">
<keyLead id="30D995F1FFE3364DFF4F758816C569D9" box="[155,1447,1446,1470]" pageId="15" pageNumber="375">
<keyStep id="30DC2E61FFE3364DFF4F758816C569D9" box="[155,1447,1446,1470]" pageId="15" pageNumber="375">
5 Pair of spines present on collum, their bases proximal to medial carina (
<figureCitation id="13132A41FFE3364DFC4B7588108B69D9" box="[927,1001,1446,1470]" captionStart="Figure 4" captionStartId="14.[113,178,1600,1624]" captionTargetBox="[151,1421,144,1571]" captionTargetId="figure-276@14.[786,1422,619,862]" captionTargetPageId="14" captionText="Figure 4. Scanning electron micrographs of Cysteochila sp. (A, B), Engynoma spinicollis (Horváth 1925) (C), Epimixia sp. (D), and Eritingis trivirgata (Horváth 1925) (EG.). Abbreviations: fmr, femur; frn, frons; mdsp, medial spine; para, paranotum; pdsp, paired spines; pmg, peritreme of the metathoracic gland, tro, trochanter." figureDoi="http://doi.org/10.5281/zenodo.11240247" httpUri="https://zenodo.org/record/11240247/files/figure.png" pageId="15" pageNumber="375">Fig. 4C</figureCitation>
,
<emphasis id="B95CEAD6FFE3364DFC207588174069D9" box="[1012,1058,1446,1470]" italics="true" pageId="15" pageNumber="375">pdsp</emphasis>
) .......................
<taxonomicName id="4C284D47FFE3364DFB60758916C569D9" authority="Drake 1942 b" authorityName="Drake" authorityYear="1942" box="[1204,1447,1446,1470]" class="Insecta" family="Tingidae" genus="Engynoma" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="375" phylum="Arthropoda" rank="genus">
<emphasis id="B95CEAD6FFE3364DFB607589167F69D9" box="[1204,1309,1447,1470]" italics="true" pageId="15" pageNumber="375">Engynoma</emphasis>
<bibRefCitation id="EFB94B35FFE3364DFAF6758816C569D9" author="Drake CJ" box="[1314,1447,1446,1470]" pageId="15" pageNumber="375" pagination="359 - 64" refId="ref48826" refString="Drake CJ. New Australian Tingitidae (Hemiptera). Journal of the Washington Academy of Sciences 1942 b; 32: 359 - 64." type="journal article" year="1942">Drake 1942b</bibRefCitation>
</taxonomicName>
</keyStep>
</keyLead>
</paragraph>
<paragraph id="8B9736C4FFE3364DFF4F75EB16C569BA" blockId="15.[155,1448,1224,1971]" box="[155,1447,1477,1501]" pageId="15" pageNumber="375">
<keyLead id="30D995F1FFE3364DFF4F75EB16C569BA" box="[155,1447,1477,1501]" pageId="15" pageNumber="375">
<keyStep id="30DC2E61FFE3364DFF4F75EB16C569BA" box="[155,1447,1477,1501]" pageId="15" pageNumber="375">6 Collum without spines ..................................................................................................................................................................................4</keyStep>
</keyLead>
</paragraph>
<paragraph id="8B9736C4FFE3364DFF4F75CB16C5699B" blockId="15.[155,1448,1224,1971]" box="[155,1447,1508,1533]" pageId="15" pageNumber="375">
<keyLead id="30D995F1FFE3364DFF4F75CB16C5699B" box="[155,1447,1508,1533]" pageId="15" pageNumber="375">
<keyStep id="30DC2E61FFE3364DFF4F75CB16C5699B" box="[155,1447,1508,1533]" pageId="15" pageNumber="375">7 Frons porrect, extending beyond antenniferous tubercles.....................................................................................................................5</keyStep>
</keyLead>
</paragraph>
<paragraph id="8B9736C4FFE3364DFF4F762A16C56A7B" blockId="15.[155,1448,1224,1971]" box="[155,1447,1540,1564]" pageId="15" pageNumber="375">
<keyLead id="30D995F1FFE3364DFF4F762A16C56A7B" box="[155,1447,1540,1564]" pageId="15" pageNumber="375">
<keyStep id="30DC2E61FFE3364DFF4F762A16C56A7B" box="[155,1447,1540,1564]" pageId="15" pageNumber="375">8 Frons flattened to slightly protruding, but never exceeding antenniferous tubercles.......................................................................6</keyStep>
</keyLead>
</paragraph>
<paragraph id="8B9736C4FFE3364DFF4F760D16C56A5C" blockId="15.[155,1448,1224,1971]" box="[155,1447,1571,1595]" pageId="15" pageNumber="375">
<keyLead id="30D995F1FFE3364DFF4F760D16C56A5C" box="[155,1447,1571,1595]" pageId="15" pageNumber="375">
<keyStep id="30DC2E61FFE3364DFF4F760D16C56A5C" box="[155,1447,1571,1595]" pageId="15" pageNumber="375">
9 Subcostal area four areolae wide; dorsal surface of pronotum and hemelytra glabrous .........................
<taxonomicName id="4C284D47FFE3364DFB6E760D16C56A5C" authority="Drake 1961" authorityName="Drake" authorityYear="1961" box="[1210,1447,1571,1595]" class="Insecta" family="Tingidae" genus="Chorotingis" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="375" phylum="Arthropoda" rank="genus">
<emphasis id="B95CEAD6FFE3364DFB6E760D16486A5C" box="[1210,1322,1571,1595]" italics="true" pageId="15" pageNumber="375">Chorotingis</emphasis>
Drake 1961
</taxonomicName>
</keyStep>
</keyLead>
</paragraph>
<paragraph id="8B9736C4FFE3364DFF4F766D16C56A1D" blockId="15.[155,1448,1224,1971]" pageId="15" pageNumber="375">
<keyLead id="30D995F1FFE3364DFF4F766D16C56A1D" pageId="15" pageNumber="375">
<keyStep id="30DC2E61FFE3364DFF4F766D16C56A1D" pageId="15" pageNumber="375">
10 Subcostal area three or four areolae wide, if four then dorsal surface of pronotum and hemelytra with short hairs................... .......................................................................................................................................................................................
<taxonomicName id="4C284D47FFE3364DFB18764C16C56A1D" authority="Drake 1945" authorityName="Drake" authorityYear="1945" box="[1228,1447,1634,1658]" class="Insecta" family="Tingidae" genus="Euaulana" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="375" phylum="Arthropoda" rank="genus">
<emphasis id="B95CEAD6FFE3364DFB18764C16496A1D" box="[1228,1323,1634,1658]" italics="true" pageId="15" pageNumber="375">Euaulana</emphasis>
Drake 1945
</taxonomicName>
</keyStep>
</keyLead>
</paragraph>
<paragraph id="8B9736C4FFE3364DFF4F76AF16C56AFE" blockId="15.[155,1448,1224,1971]" box="[155,1447,1665,1689]" pageId="15" pageNumber="375">
<keyLead id="30D995F1FFE3364DFF4F76AF16C56AFE" box="[155,1447,1665,1689]" pageId="15" pageNumber="375">
<keyStep id="30DC2E61FFE3364DFF4F76AF16C56AFE" box="[155,1447,1665,1689]" pageId="15" pageNumber="375">
11 Trochanters fused to femurs (
<figureCitation id="13132A41FFE3364DFE2D76AF11266AFE" box="[505,580,1665,1689]" captionStart="Figure 4" captionStartId="14.[113,178,1600,1624]" captionTargetBox="[151,1421,144,1571]" captionTargetId="figure-276@14.[786,1422,619,862]" captionTargetPageId="14" captionText="Figure 4. Scanning electron micrographs of Cysteochila sp. (A, B), Engynoma spinicollis (Horváth 1925) (C), Epimixia sp. (D), and Eritingis trivirgata (Horváth 1925) (EG.). Abbreviations: fmr, femur; frn, frons; mdsp, medial spine; para, paranotum; pdsp, paired spines; pmg, peritreme of the metathoracic gland, tro, trochanter." figureDoi="http://doi.org/10.5281/zenodo.11240247" httpUri="https://zenodo.org/record/11240247/files/figure.png" pageId="15" pageNumber="375">Fig. 4D</figureCitation>
)..............................................................................................................
<taxonomicName id="4C284D47FFE3364DFB6376AC16C56AFE" authority="Kirkaldy 1908" authorityName="Kirkaldy" authorityYear="1908" box="[1207,1447,1665,1689]" class="Insecta" family="Tingidae" genus="Epimixia" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="375" phylum="Arthropoda" rank="genus">
<emphasis id="B95CEAD6FFE3364DFB6376AC16736AFE" box="[1207,1297,1666,1689]" italics="true" pageId="15" pageNumber="375">Epimixia</emphasis>
<bibRefCitation id="EFB94B35FFE3364DFAC276AF16C56AFE" author="Kirkaldy GW" box="[1302,1447,1665,1689]" pageId="15" pageNumber="375" pagination="768 - 788" refId="ref49893" refString="Kirkaldy GW. Memoir on a few heteropterous Hemiptera from eastern Australia. Proceedings of the Linnean Society of New South Wales 1908; 32: 768 - 788." type="journal article" year="1908">Kirkaldy 1908</bibRefCitation>
</taxonomicName>
</keyStep>
</keyLead>
</paragraph>
<paragraph id="8B9736C4FFE3364DFF4F768F16C56ADF" blockId="15.[155,1448,1224,1971]" box="[155,1447,1696,1721]" pageId="15" pageNumber="375">
<keyLead id="30D995F1FFE3364DFF4F768F16C56ADF" box="[155,1447,1696,1721]" pageId="15" pageNumber="375">
<keyStep id="30DC2E61FFE3364DFF4F768F16C56ADF" box="[155,1447,1696,1721]" pageId="15" pageNumber="375">
12 Trochanters differentiated from femurs (
<figureCitation id="13132A41FFE3364DFDB3768E11CF6ADE" box="[615,685,1696,1721]" captionStart="Figure 4" captionStartId="14.[113,178,1600,1624]" captionTargetBox="[151,1421,144,1571]" captionTargetId="figure-276@14.[786,1422,619,862]" captionTargetPageId="14" captionText="Figure 4. Scanning electron micrographs of Cysteochila sp. (A, B), Engynoma spinicollis (Horváth 1925) (C), Epimixia sp. (D), and Eritingis trivirgata (Horváth 1925) (EG.). Abbreviations: fmr, femur; frn, frons; mdsp, medial spine; para, paranotum; pdsp, paired spines; pmg, peritreme of the metathoracic gland, tro, trochanter." figureDoi="http://doi.org/10.5281/zenodo.11240247" httpUri="https://zenodo.org/record/11240247/files/figure.png" pageId="15" pageNumber="375">Fig. 4E</figureCitation>
)....................................................................................................................................7
</keyStep>
</keyLead>
</paragraph>
<paragraph id="8B9736C4FFE3364DFF4F76EE16C56A90" blockId="15.[155,1448,1224,1971]" pageId="15" pageNumber="375">
<keyLead id="30D995F1FFE3364DFF4F76EE16C56A90" pageId="15" pageNumber="375">
<keyStep id="30DC2E61FFE3364DFF4F76EE16C56A90" pageId="15" pageNumber="375">
13 Paranota absent or reduced to carina-like structure on pronotal disc; if present anteriorly, upturned and adpressed to collum (
<figureCitation id="13132A41FFE3364DFF0276F1125A6A90" box="[214,312,1759,1783]" captionStart="Figure 4" captionStartId="14.[113,178,1600,1624]" captionTargetBox="[151,1421,144,1571]" captionTargetId="figure-276@14.[786,1422,619,862]" captionTargetPageId="14" captionText="Figure 4. Scanning electron micrographs of Cysteochila sp. (A, B), Engynoma spinicollis (Horváth 1925) (C), Epimixia sp. (D), and Eritingis trivirgata (Horváth 1925) (EG.). Abbreviations: fmr, femur; frn, frons; mdsp, medial spine; para, paranotum; pdsp, paired spines; pmg, peritreme of the metathoracic gland, tro, trochanter." figureDoi="http://doi.org/10.5281/zenodo.11240247" httpUri="https://zenodo.org/record/11240247/files/figure.png" pageId="15" pageNumber="375">Fig. 4F, G</figureCitation>
,
<emphasis id="B95CEAD6FFE3364DFE9776CE12126A90" box="[323,368,1760,1783]" italics="true" pageId="15" pageNumber="375">para</emphasis>
).............................................................................................................................................................................................8
</keyStep>
</keyLead>
</paragraph>
<paragraph id="8B9736C4FFE3364DFF4F76D116C56B51" blockId="15.[155,1448,1224,1971]" pageId="15" pageNumber="375">
<keyLead id="30D995F1FFE3364DFF4F76D116C56B51" pageId="15" pageNumber="375">
<keyStep id="30DC2E61FFE3364DFF4F76D116C56B51" pageId="15" pageNumber="375">
14 Paranota present, areolate over entire length; if reduced posteriorly on pronotal disc, paranota never adpressed to collum (
<figureCitation id="13132A41FFE3364DFF02773012096B52" box="[214,363,1822,1846]" captionStart="Figure 5" captionStartId="17.[129,194,1596,1620]" captionTargetBox="[166,1436,143,1568]" captionTargetId="figure-188@17.[801,1437,618,1095]" captionTargetPageId="17" captionText="Figure 5. Scanning electron micrographs of Proteatingis xouthos sp. nov. (A, B), Malandiola semota Drake 1942b (C, D), and Cercotingis tasmaniensis sp. nov. (E, F). Abbreviations:hd, hood; mdsp, medial spine; para, paranotum; pmg, peritreme of the metathoracic gland." figureDoi="http://doi.org/10.5281/zenodo.11240249" httpUri="https://zenodo.org/record/11240249/files/figure.png" pageId="15" pageNumber="375">Fig. 5A, B, E, F</figureCitation>
,
<emphasis id="B95CEAD6FFE3364DFEA2773112C16B51" box="[374,419,1823,1846]" italics="true" pageId="15" pageNumber="375">para</emphasis>
)................................................................................................................................................................................. 10
</keyStep>
</keyLead>
</paragraph>
<paragraph id="8B9736C4FFE3364DFF4F771316C56B13" blockId="15.[155,1448,1224,1971]" pageId="15" pageNumber="375">
<keyLead id="30D995F1FFE3364DFF4F771316C56B13" pageId="15" pageNumber="375">
<keyStep id="30DC2E61FFE3364DFF4F771316C56B13" pageId="15" pageNumber="375">
15 Peritreme of metathoracic gland present (
<figureCitation id="13132A41FFE3364DFDA7771311DC6B32" box="[627,702,1853,1877]" captionStart="Figure 4" captionStartId="14.[113,178,1600,1624]" captionTargetBox="[151,1421,144,1571]" captionTargetId="figure-276@14.[786,1422,619,862]" captionTargetPageId="14" captionText="Figure 4. Scanning electron micrographs of Cysteochila sp. (A, B), Engynoma spinicollis (Horváth 1925) (C), Epimixia sp. (D), and Eritingis trivirgata (Horváth 1925) (EG.). Abbreviations: fmr, femur; frn, frons; mdsp, medial spine; para, paranotum; pdsp, paired spines; pmg, peritreme of the metathoracic gland, tro, trochanter." figureDoi="http://doi.org/10.5281/zenodo.11240247" httpUri="https://zenodo.org/record/11240247/files/figure.png" pageId="15" pageNumber="375">Fig. 4G</figureCitation>
); medial spine present (
<figureCitation id="13132A41FFE3364DFC637713177B6B32" box="[951,1049,1853,1877]" captionStart="Figure 4" captionStartId="14.[113,178,1600,1624]" captionTargetBox="[151,1421,144,1571]" captionTargetId="figure-276@14.[786,1422,619,862]" captionTargetPageId="14" captionText="Figure 4. Scanning electron micrographs of Cysteochila sp. (A, B), Engynoma spinicollis (Horváth 1925) (C), Epimixia sp. (D), and Eritingis trivirgata (Horváth 1925) (EG.). Abbreviations: fmr, femur; frn, frons; mdsp, medial spine; para, paranotum; pdsp, paired spines; pmg, peritreme of the metathoracic gland, tro, trochanter." figureDoi="http://doi.org/10.5281/zenodo.11240247" httpUri="https://zenodo.org/record/11240247/files/figure.png" pageId="15" pageNumber="375">Fig. 4F, G</figureCitation>
); costal area areolate................................ ...................................................................................................................................................................
<taxonomicName id="4C284D47FFE3364DFBB5777316C56B13" authority="Drake and Ruhoff 1962" authorityName="Drake and Ruhoff" authorityYear="1962" box="[1121,1447,1884,1908]" class="Insecta" family="Tingidae" genus="Eritingis" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="375" phylum="Arthropoda" rank="genus">
<emphasis id="B95CEAD6FFE3364DFBB5777317D16B13" box="[1121,1203,1885,1908]" italics="true" pageId="15" pageNumber="375">Eritingis</emphasis>
<bibRefCitation id="EFB94B35FFE3364DFB6C777216C56B13" author="Drake CJ &amp; Ruhoff FA" box="[1208,1447,1884,1908]" pageId="15" pageNumber="375" pagination="489 - 506" refId="ref49295" refString="Drake CJ, Ruhoff FA. Synomic notes and descriptions of new Tingidae (Hemiptera). Studia Entomologica 1962; 5: 489 - 506." type="journal article" year="1962">Drake and Ruhoff 1962</bibRefCitation>
</taxonomicName>
</keyStep>
</keyLead>
</paragraph>
<paragraph id="8B9736C4FFE33652FF4F775216FA6CB1" blockId="15.[155,1448,1224,1971]" lastBlockId="16.[140,1433,159,466]" lastPageId="16" lastPageNumber="376" pageId="15" pageNumber="375">
<keyLead id="30D995F1FFE3364DFF4F775216C56BD4" pageId="15" pageNumber="375">
<keyStep id="30DC2E61FFE3364DFF4F775216C56BD4" pageId="15" pageNumber="375">
16 Peritreme of metathoracic gland absent (
<figureCitation id="13132A41FFE3364DFDBE775211CD6BF3" box="[618,687,1916,1940]" captionStart="Figure 5" captionStartId="17.[129,194,1596,1620]" captionTargetBox="[166,1436,143,1568]" captionTargetId="figure-188@17.[801,1437,618,1095]" captionTargetPageId="17" captionText="Figure 5. Scanning electron micrographs of Proteatingis xouthos sp. nov. (A, B), Malandiola semota Drake 1942b (C, D), and Cercotingis tasmaniensis sp. nov. (E, F). Abbreviations:hd, hood; mdsp, medial spine; para, paranotum; pmg, peritreme of the metathoracic gland." figureDoi="http://doi.org/10.5281/zenodo.11240249" httpUri="https://zenodo.org/record/11240249/files/figure.png" pageId="15" pageNumber="375">Fig. 5d</figureCitation>
); medial spine absent or greatly reduced in size (
<figureCitation id="13132A41FFE3364DFB757752179E6BF3" box="[1185,1276,1916,1940]" captionStart="Figure 5" captionStartId="17.[129,194,1596,1620]" captionTargetBox="[166,1436,143,1568]" captionTargetId="figure-188@17.[801,1437,618,1095]" captionTargetPageId="17" captionText="Figure 5. Scanning electron micrographs of Proteatingis xouthos sp. nov. (A, B), Malandiola semota Drake 1942b (C, D), and Cercotingis tasmaniensis sp. nov. (E, F). Abbreviations:hd, hood; mdsp, medial spine; para, paranotum; pmg, peritreme of the metathoracic gland." figureDoi="http://doi.org/10.5281/zenodo.11240249" httpUri="https://zenodo.org/record/11240249/files/figure.png" pageId="15" pageNumber="375">Fig. 5c, d</figureCitation>
); if medial spine present, costal area reduced to carina-like structure................................................................................................................................9
</keyStep>
</keyLead>
<keyLead id="30D995F1FFFC3652FF5870B116FA6CB1" pageId="16" pageNumber="376">
<keyStep id="30DC2E61FFFC3652FF5870B116FA6CB1" pageId="16" pageNumber="376">
17 Costal area areolate for at least portion of length; mesosternal and metasternal carinae straight, forming narrow rostral channel; dorsal surface of pronotum and hemelytra glabrous ..........................................................................
<taxonomicName id="4C284D47FFFC3652FB45709016FA6CB1" authority="Horvath 1925" authorityName="Horvath" authorityYear="1925" box="[1169,1432,190,214]" class="Insecta" family="Tingidae" genus="Malandiola" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="376" phylum="Arthropoda" rank="genus">
<emphasis id="B95CEAD6FFFC3652FB45709016616CB1" box="[1169,1283,190,214]" italics="true" pageId="16" pageNumber="376">Malandiola</emphasis>
<bibRefCitation id="EFB94B35FFFC3652FADC709016FA6CB1" author="Horvath G" box="[1288,1432,190,214]" pageId="16" pageNumber="376" pagination="1 - 17" refId="ref49693" refString="Horvath G. Results of Dr. E. Mjoberg's Swedish scientific expeditions to Australia 1910 - 1913. 45. Tingitidae. Arkiv for Zoologi 1925; 17 A: 1 - 17." type="journal article" year="1925">Horváth 1925</bibRefCitation>
</taxonomicName>
</keyStep>
</keyLead>
</paragraph>
<paragraph id="8B9736C4FFFC3652FF5870F016FA6D72" blockId="16.[140,1433,159,466]" pageId="16" pageNumber="376">
<keyLead id="30D995F1FFFC3652FF5870F016FA6D72" pageId="16" pageNumber="376">
<keyStep id="30DC2E61FFFC3652FF5870F016FA6D72" pageId="16" pageNumber="376">
18 Costal area reduced to carina-like structure, lacking areolae; mesosternal and metasternal carinae curved, forming broad rostral channel; dorsal surface of pronotum and hemelytra usually with setae or hairs .........................
<taxonomicName id="4C284D47FFFC3652FB7270D316FA6D72" authority="Horvath 1925" authorityName="Horvath" authorityYear="1925" box="[1190,1432,253,277]" class="Insecta" family="Tingidae" genus="Nethersia" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="376" phylum="Arthropoda" rank="genus">
<emphasis id="B95CEAD6FFFC3652FB7270D316616D72" box="[1190,1283,253,277]" italics="true" pageId="16" pageNumber="376">Nethersia</emphasis>
<bibRefCitation id="EFB94B35FFFC3652FADC70D316FA6D72" author="Horvath G" box="[1288,1432,253,277]" pageId="16" pageNumber="376" pagination="1 - 17" refId="ref49693" refString="Horvath G. Results of Dr. E. Mjoberg's Swedish scientific expeditions to Australia 1910 - 1913. 45. Tingitidae. Arkiv for Zoologi 1925; 17 A: 1 - 17." type="journal article" year="1925">Horváth 1925</bibRefCitation>
</taxonomicName>
</keyStep>
</keyLead>
</paragraph>
<paragraph id="8B9736C4FFFC3652FF58713316FB6D34" blockId="16.[140,1433,159,466]" pageId="16" pageNumber="376">
<keyLead id="30D995F1FFFC3652FF58713316FB6D34" pageId="16" pageNumber="376">
<keyStep id="30DC2E61FFFC3652FF58713316FB6D34" pageId="16" pageNumber="376">
19 Collum raised anteriorly, forming hood (
<figureCitation id="13132A41FFFC3652FD83713211FF6D53" box="[599,669,284,308]" captionStart="Figure 5" captionStartId="17.[129,194,1596,1620]" captionTargetBox="[166,1436,143,1568]" captionTargetId="figure-188@17.[801,1437,618,1095]" captionTargetPageId="17" captionText="Figure 5. Scanning electron micrographs of Proteatingis xouthos sp. nov. (A, B), Malandiola semota Drake 1942b (C, D), and Cercotingis tasmaniensis sp. nov. (E, F). Abbreviations:hd, hood; mdsp, medial spine; para, paranotum; pmg, peritreme of the metathoracic gland." figureDoi="http://doi.org/10.5281/zenodo.11240249" httpUri="https://zenodo.org/record/11240249/files/figure.png" pageId="16" pageNumber="376">Fig. 5B</figureCitation>
,
<emphasis id="B95CEAD6FFFC3652FD73713211A26D53" box="[679,704,284,308]" italics="true" pageId="16" pageNumber="376">hd</emphasis>
); paranota never more than two areolae wide anteriorly, one areola wide at humeral angles (
<figureCitation id="13132A41FFFC3652FEA9711212A26D33" box="[381,448,316,340]" captionStart="Figure 5" captionStartId="17.[129,194,1596,1620]" captionTargetBox="[166,1436,143,1568]" captionTargetId="figure-188@17.[801,1437,618,1095]" captionTargetPageId="17" captionText="Figure 5. Scanning electron micrographs of Proteatingis xouthos sp. nov. (A, B), Malandiola semota Drake 1942b (C, D), and Cercotingis tasmaniensis sp. nov. (E, F). Abbreviations:hd, hood; mdsp, medial spine; para, paranotum; pmg, peritreme of the metathoracic gland." figureDoi="http://doi.org/10.5281/zenodo.11240249" httpUri="https://zenodo.org/record/11240249/files/figure.png" pageId="16" pageNumber="376">Fig. 5a</figureCitation>
); costal area one areolae wide over entire length..................................................
<taxonomicName id="4C284D47FFFC3652FB127112165B6D34" authority="Shofner &amp; Cassis, 2024" authorityName="Shofner &amp; Cassis" authorityYear="2024" box="[1222,1337,316,339]" class="Insecta" family="Tingidae" genus="Proteatingis" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="376" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B95CEAD6FFFC3652FB127112165B6D34" box="[1222,1337,316,339]" italics="true" pageId="16" pageNumber="376">Proteatingis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F57ADFFFC3652FAEB711216FB6D34" box="[1343,1433,316,340]" pageId="16" pageNumber="376" rank="genus">gen. nov.</taxonomicNameLabel>
</keyStep>
</keyLead>
</paragraph>
<paragraph id="8B9736C4FFFC3652FF58717516FA6DF5" blockId="16.[140,1433,159,466]" pageId="16" pageNumber="376">
<keyLead id="30D995F1FFFC3652FF58717516FA6DF5" pageId="16" pageNumber="376">
<keyStep id="30DC2E61FFFC3652FF58717516FA6DF5" pageId="16" pageNumber="376">20 Collum flattened, or if raised, only posteriorly, never forming hood; paranota two or more areolae wide, one to three areolae wide at humeral angles or carina-like; costal area one or two areolae wide, at least partially ...................................................... 11</keyStep>
</keyLead>
</paragraph>
<paragraph id="8B9736C4FFFC3652FF5871B416FA6DD5" blockId="16.[140,1433,159,466]" box="[140,1432,410,434]" pageId="16" pageNumber="376">
<keyLead id="30D995F1FFFC3652FF5871B416FA6DD5" box="[140,1432,410,434]" pageId="16" pageNumber="376">
<keyStep id="30DC2E61FFFC3652FF5871B416FA6DD5" box="[140,1432,410,434]" pageId="16" pageNumber="376">
21 Costal area one areolae wide over entire length; dorsal surface of pronotum glabrous ..........
<taxonomicName id="4C284D47FFFC3652FB9A71B416FA6DD5" authority="Drake and Poor 1937 a" authorityName="Drake and Poor" authorityYear="1937" box="[1102,1432,410,434]" class="Insecta" family="Tingidae" genus="Ulonemia" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="376" phylum="Arthropoda" rank="genus">
<emphasis id="B95CEAD6FFFC3652FB9A71B417CC6DD5" box="[1102,1198,410,434]" italics="true" pageId="16" pageNumber="376">Ulonemia</emphasis>
<bibRefCitation id="EFB94B35FFFC3652FB6771B416FA6DD5" author="Drake CJ &amp; Poor ME" box="[1203,1432,410,434]" pageId="16" pageNumber="376" pagination="1 - 18" refId="ref48996" refString="Drake CJ, Poor ME. Tingitidae from Malaysia and Madagascar (Hemiptera). The Philippine Journal of Science 1937 a; 62: 1 - 18." type="journal article" year="1937">Drake and Poor 1937a</bibRefCitation>
</taxonomicName>
</keyStep>
</keyLead>
</paragraph>
<paragraph id="8B9736C4FFFC3652FF58719716FB6DB6" blockId="16.[140,1433,159,466]" box="[140,1433,441,466]" pageId="16" pageNumber="376">
<keyLead id="30D995F1FFFC3652FF58719716FB6DB6" box="[140,1433,441,466]" pageId="16" pageNumber="376">
<keyStep id="30DC2E61FFFC3652FF58719716FB6DB6" box="[140,1433,441,466]" pageId="16" pageNumber="376">
22 Costal area two areolae wide anteriorly; dorsal surface of pronotum with scale-like setae...........................
<taxonomicName id="4C284D47FFFC3652FB1A7197165B6DB7" authority="Shofner &amp; Cassis, 2024" authorityName="Shofner &amp; Cassis" authorityYear="2024" box="[1230,1337,441,464]" class="Insecta" family="Tingidae" genus="Cercotingis" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="376" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B95CEAD6FFFC3652FB1A7197165B6DB7" box="[1230,1337,441,464]" italics="true" pageId="16" pageNumber="376">Cercotingis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F57ADFFFC3652FAEB719416FB6DB6" box="[1343,1433,442,466]" pageId="16" pageNumber="376" rank="genus">gen. nov.</taxonomicNameLabel>
</keyStep>
</keyLead>
</paragraph>
</key>
</subSubSection>
</treatment>
</document>