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<document id="09B63E95C4AB7DD151B75423D97E2067" ID-CLB-Dataset="2390" ID-DOI="10.11646/zootaxa.5093.3.8" ID-GBIF-Dataset="42d4c4ac-b6d2-472d-a0f3-77dc3fc8cb58" ID-ISSN="1175-5326" ID-Zenodo-Dep="5910029" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1643277067791" checkinUser="plazi" docAuthor="Takano, Tsuyoshi, Tsuzuki, Shouji &amp; Kano, Yasunori" docDate="2022" docId="038187C9FF8EFFF396E8A8F6FD76F8A9" docLanguage="en" docName="zootaxa.5093.3.8.pdf" docOrigin="Zootaxa 5093 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Concavibalcis haterumaensis Takano &amp; Tsuzuki &amp; Kano 2022, n. sp." docType="treatment" docVersion="5" lastPageNumber="398" masterDocId="FFB8FFB1FF8FFFF2967FA861FFBBFFE5" masterDocTitle="Description of a second species of the eulimid genus Concavibalcis (Gastropoda: Vanikoroidea)" masterLastPageNumber="400" masterPageNumber="397" pageNumber="398" updateTime="1699304549584" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="5EE5A847A0A39BB1F6E8FC0CE2ADA51E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0DED1063F2375F3DF9E33BBCDF70129A">
<mods:title id="9C2811A136739949DAEF6C44E0F6E569">Description of a second species of the eulimid genus Concavibalcis (Gastropoda: Vanikoroidea)</mods:title>
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<mods:name id="0C3B7568AEBDED9053E69513A4E8BF1B" type="personal">
<mods:role id="B268DAB9D1B3281A05A0462B16477926">
<mods:roleTerm id="55A9999A39901D81E8633B16032DFD01">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="149F5AD7F3ECC2832481A217166EAA7A">Takano, Tsuyoshi</mods:namePart>
<mods:affiliation id="A38E3D29FEE68A87BA711841503E6C87">Meguro Parasitological Museum, 4 - 1 - 1 Shimomeguro, Meguro, Tokyo 153 - 0064, Japan.</mods:affiliation>
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<mods:role id="DC432EB539867CFC2B5721076872783C">
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<mods:namePart id="A875F34E8FB00438953A0558A39E355C">Tsuzuki, Shouji</mods:namePart>
<mods:affiliation id="7C61391B094C77864CDF97CAA5CE8DB3">53 - 17 Oyumi, Chiba, Chiba 260 - 0813, Japan. urumashell 9860 @ gmail. com</mods:affiliation>
</mods:name>
<mods:name id="4A23B9530451099E1DA686319EC15868" type="personal">
<mods:role id="C62B3DA414839AF4ABE89A8411702B1F">
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<mods:namePart id="74C5F7E248EA77A2D6AB044810BEACF4">Kano, Yasunori</mods:namePart>
<mods:nameIdentifier id="B1D4366B21011FA3D7C3DCE1B43FEF67" type="ORCID">0000-0002-9527-5116</mods:nameIdentifier>
<mods:affiliation id="7329645F4D24C34625E2FA406DB7B269">Atmosphere and Ocean Research Institute, The University of Tokyo, 5 - 1 - 5 Kashiwanoha, Kashiwa, Chiba 277 - 8564, Japan. kano @ aori. u-tokyo. ac. jp; https: // orcid. org / 0000 - 0002 - 9527 - 5116</mods:affiliation>
<mods:nameIdentifier id="B4293DE868AE320C96E6C9670C8D80A1" type="email">kano@aori.u-tokyo.ac.jp</mods:nameIdentifier>
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<mods:typeOfResource id="8BE33E61BAAD128446A673812C5515C3">text</mods:typeOfResource>
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<mods:title id="F81B58A3213E9B29D5433E25D5493EAD">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="3015BA953E4A9CA343169E363E6AAA50">
<mods:date id="E89F82B0165823B31A94544243305365">2022</mods:date>
<mods:detail id="D1EDB793E755A1941CA33D9BB2FC1909" type="pubDate">
<mods:number id="91F0DE0042D123C981188F4A4A4CA16D">2022-01-27</mods:number>
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<mods:number id="AA997F27B3A5E763A1FCAD3404940649">5093</mods:number>
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<mods:start id="1529F75CC6C80F95C3DD4E2155EF14DD">397</mods:start>
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<mods:identifier id="63AFA7BF5D43A530E647AED8D3BB9E63" type="DOI">10.11646/zootaxa.5093.3.8</mods:identifier>
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<treatment id="038187C9FF8EFFF396E8A8F6FD76F8A9" ID-DOI="http://doi.org/10.5281/zenodo.5910037" ID-GBIF-Taxon="192651053" ID-Zenodo-Dep="5910037" LSID="urn:lsid:plazi:treatment:038187C9FF8EFFF396E8A8F6FD76F8A9" httpUri="http://treatment.plazi.org/id/038187C9FF8EFFF396E8A8F6FD76F8A9" lastPageNumber="398" pageId="1" pageNumber="398">
<subSubSection id="C3326554FF8EFFF396E8A8F6FDF8FF54" box="[151,579,151,178]" pageId="1" pageNumber="398" type="nomenclature">
<paragraph id="8B9736DFFF8EFFF396E8A8F6FDF8FF54" blockId="1.[151,1437,151,1869]" box="[151,579,151,178]" pageId="1" pageNumber="398">
<heading id="D0DF81B3FF8EFFF396E8A8F6FDF8FF54" bold="true" box="[151,579,151,178]" fontSize="11" level="1" pageId="1" pageNumber="398" reason="1">
<emphasis id="B95CEACDFF8EFFF396E8A8F6FDF8FF54" bold="true" box="[151,579,151,178]" pageId="1" pageNumber="398">
<taxonomicName id="4C284D5CFF8EFFF396E8A8F6FE41FF57" ID-CoL="8VQ32" authority="Takano &amp; Tsuzuki &amp; Kano, 2022" authorityName="Takano &amp; Tsuzuki &amp; Kano" authorityYear="2022" box="[151,506,151,178]" class="Gastropoda" family="Eulimidae" genus="Concavibalcis" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="398" phylum="Mollusca" rank="species" species="haterumaensis" status="sp. nov.">
<emphasis id="B95CEACDFF8EFFF396E8A8F6FE41FF57" bold="true" box="[151,506,151,178]" italics="true" pageId="1" pageNumber="398">Concavibalcis haterumaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F57B6FF8EFFF3947EA8F6FDF8FF54" box="[513,579,151,177]" pageId="1" pageNumber="398" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3326554FF8EFFF396E8A8DCFF53FF30" box="[151,232,189,213]" pageId="1" pageNumber="398" type="description">
<paragraph id="8B9736DFFF8EFFF396E8A8DCFF53FF30" blockId="1.[151,1437,151,1869]" box="[151,232,189,213]" pageId="1" pageNumber="398">
(
<figureCitation id="13132A5AFF8EFFF396E0A8DCFF66FF30" box="[159,221,189,213]" captionStart="FIGURE 1" captionStartId="2.[151,250,1431,1455]" captionTargetBox="[151,1436,181,1408]" captionTargetId="figure-27@2.[151,1436,181,1408]" captionTargetPageId="2" captionText="FIGURE 1. Concavibalcis haterumaensis n. sp. A. Holotype NSMT-Mo 79320, 3.0 mm in shell height. B–E. Paratype MPM Coll. No. 21794a, 2.0 mm, SEM. B. Lateral view showing incremental scars in penultimate and last whorls. C. Close-up view of blunt apex; arrowheads denote demarcation line between protoconch and teleoconch. D. Penultimate whorl with an incremental line and fine spiral ridges. E. Surface of first teleoconch whorl showing short oblique ridges and continuous spiral ridges. Scale bars = 1 mm (A), 500 µm (B) and 100 µm (C–E)." figureDoi="http://doi.org/10.5281/zenodo.5910031" httpUri="https://zenodo.org/record/5910031/files/figure.png" pageId="1" pageNumber="398">Fig. 1</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3326554FF8EFFF396E8A880FAEDFE48" pageId="1" pageNumber="398" type="materials_examined">
<paragraph id="8B9736DFFF8EFFF396E8A880FE9DFEF9" blockId="1.[151,1437,151,1869]" pageId="1" pageNumber="398">
<materialsCitation id="3B403C82FF8EFFF396E8A880FE9DFEF9" ID-GBIF-Occurrence="3458045302" country="Japan" county="Taketomi" elevation="4" latitude="24.068335" location="Nishi-hama" longLatPrecision="21" longitude="123.76278" municipality="Hateruma Island" pageId="1" pageNumber="398" specimenCount="4" stateProvince="Okinawa" typeStatus="holotype">
<emphasis id="B95CEACDFF8EFFF396E8A880FE94FF1C" bold="true" box="[151,303,225,249]" inLineHeading="true" pageId="1" pageNumber="398" reason="1">
<typeStatus id="5493887DFF8EFFF396E8A880FF74FF1C" box="[151,207,225,249]" pageId="1" pageNumber="398">Type</typeStatus>
locality.
</emphasis>
<location id="8EF76004FF8EFFF39748A880FE0DFF1C" LSID="urn:lsid:plazi:treatment:038187C9FF8EFFF396E8A8F6FD76F8A9:8EF76004FF8EFFF39748A880FE0DFF1C" box="[311,438,225,249]" country="Japan" county="Taketomi" latitude="24.068335" longLatPrecision="21" longitude="123.76278" municipality="Hateruma Island" name="Nishi-hama" pageId="1" pageNumber="398" stateProvince="Okinawa">Nishi-hama</location>
,
<collectingMunicipality id="6BF3ACA5FF8EFFF397BDA880FDCCFF1C" box="[450,631,225,249]" pageId="1" pageNumber="398">Hateruma Island</collectingMunicipality>
,
<collectingCounty id="62F64E53FF8EFFF394FDA880FD52FF1C" box="[642,745,225,249]" pageId="1" pageNumber="398">Taketomi</collectingCounty>
,
<collectingRegion id="49ECF83DFF8EFFF3948BA880FCECFF1C" box="[756,855,225,249]" country="Japan" name="Okinawa" pageId="1" pageNumber="398">Okinawa</collectingRegion>
,
<collectingCountry id="F33F764FFF8EFFF3951CA880FC1BFF1D" box="[867,928,225,248]" name="Japan" pageId="1" pageNumber="398">Japan</collectingCountry>
(
<geoCoordinate id="EE1C5018FF8EFFF395CFA880FB8BFF1C" box="[944,1072,225,249]" degrees="24" direction="north" minutes="04" orientation="latitude" pageId="1" pageNumber="398" precision="15" seconds="06" value="24.068335">24°04'06&quot;N</geoCoordinate>
,
<geoCoordinate id="EE1C5018FF8EFFF39242A880FB7CFF1C" box="[1085,1223,225,249]" degrees="123" direction="east" minutes="45" orientation="longitude" pageId="1" pageNumber="398" precision="15" seconds="46" value="123.76278">123°45'46&quot;E</geoCoordinate>
), sandy bottom at a depth of
<specimenCount id="9D2EFD56FF8EFFF39689A964FE9DFEF9" box="[246,294,261,285]" pageId="1" pageNumber="398" type="generic">
<quantity id="4CD09B3AFF8EFFF39689A964FE99FEF9" box="[246,290,261,285]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="398" unit="m" value="4.0">
<elevation id="0005D1ECFF8EFFF39689A964FE99FEF9" box="[246,290,261,285]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="398" unit="m" value="4.0">4 m</elevation>
</quantity>
.
</specimenCount>
</materialsCitation>
</paragraph>
<paragraph id="8B9736DFFF8EFFF396B8A948FEE2FE6C" blockId="1.[151,1437,151,1869]" pageId="1" pageNumber="398">
<emphasis id="B95CEACDFF8EFFF396B8A948FED6FEA4" bold="true" box="[199,365,297,321]" inLineHeading="true" pageId="1" pageNumber="398" reason="1">Type material.</emphasis>
<materialsCitation id="3B403C82FF8EFFF3970BA948FEE2FE6C" ID-GBIF-Occurrence="3458045303" collectingDate="2019-05-30" collectionCode="MPM" collectorName="Coll. No." pageId="1" pageNumber="398" specimenCount="13" typeStatus="holotype">
<typeStatus id="5493887DFF8EFFF3970BA948FE6CFEA4" box="[372,471,297,321]" pageId="1" pageNumber="398" type="holotype">Holotype</typeStatus>
NSMT-Mo 79320;
<specimenCount id="9D2EFD56FF8EFFF394D2A948FC9CFEA4" box="[685,807,297,321]" pageId="1" pageNumber="398" type="generic" typeStatus="paratypes">8 paratypes</specimenCount>
NSMT-Mo 79321;
<specimenCount id="9D2EFD56FF8EFFF39583A948FBCCFEA4" box="[1020,1143,297,321]" pageId="1" pageNumber="398" type="generic" typeStatus="paratypes">3 paratypes</specimenCount>
<collectionCode id="ED39AE1AFF8EFFF39201A948FB07FEA5" box="[1150,1212,297,320]" country="USA" httpUri="http://grbio.org/cool/7hds-psx6" name="Milwaukee Public Museum" pageId="1" pageNumber="398" type="Museum">MPM</collectionCode>
<collectorName id="26DD5309FF8EFFF392BBA948FA9CFEA5" box="[1220,1319,297,321]" pageId="1" pageNumber="398">Coll. No.</collectorName>
21794a–c, all collected from the type locality on
<date id="FF96101FFF8EFFF3944AA92CFD6BFE80" box="[565,720,333,357]" pageId="1" pageNumber="398" value="2019-05-30">
<collectingDate id="EFD2E9F7FF8EFFF3944AA92CFD6BFE80" box="[565,720,333,357]" pageId="1" pageNumber="398" value="2019-05-30">May 30, 2019</collectingDate>
</date>
.
<specimenCount id="9D2EFD56FF8EFFF394A5A92CFCEEFE80" box="[730,853,333,357]" pageId="1" pageNumber="398" type="generic" typeStatus="paratypes">2 paratypes</specimenCount>
(
<collectionCode id="ED39AE1AFF8EFFF3951BA92CFC1FFE80" box="[868,932,333,357]" country="USA" httpUri="http://grbio.org/cool/7hds-psx6" name="Milwaukee Public Museum" pageId="1" pageNumber="398" type="Museum">MPM</collectionCode>
<collectorName id="26DD5309FF8EFFF395D3A92CFBABFE81" box="[940,1040,333,357]" pageId="1" pageNumber="398">Coll. No.</collectorName>
21794a, b) coated with platinum for SEM observation.
</materialsCitation>
</paragraph>
<paragraph id="8B9736DFFF8EFFF396B8A9F4FAEDFE48" blockId="1.[151,1437,151,1869]" box="[199,1366,405,429]" pageId="1" pageNumber="398">
<materialsCitation id="3B403C82FF8EFFF396B8A9F4FAE9FE48" ID-GBIF-Occurrence="3458045301" box="[199,1362,405,429]" collectionCode="MPM" collectorName="Coll. No." pageId="1" pageNumber="398" specimenCount="12">
<emphasis id="B95CEACDFF8EFFF396B8A9F4FE56FE48" bold="true" box="[199,493,405,429]" inLineHeading="true" pageId="1" pageNumber="398" reason="1">Other material examined.</emphasis>
<specimenCount id="9D2EFD56FF8EFFF3978CA9F4FD3FFE48" box="[499,644,405,429]" pageId="1" pageNumber="398" type="generic">12 specimens</specimenCount>
<collectionCode id="ED39AE1AFF8EFFF394F5A9F4FD73FE49" box="[650,712,405,428]" country="USA" httpUri="http://grbio.org/cool/7hds-psx6" name="Milwaukee Public Museum" pageId="1" pageNumber="398" type="Museum">MPM</collectionCode>
<collectorName id="26DD5309FF8EFFF394B0A9F4FC88FE49" box="[719,819,405,429]" pageId="1" pageNumber="398">Coll. No.</collectorName>
21795, collected together with the
<typeStatus id="5493887DFF8EFFF392D4A9F4FB62FE48" box="[1195,1241,405,429]" pageId="1" pageNumber="398">type</typeStatus>
specimens
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3326554FF8EFFF396B8A9D8FC36FE34" box="[199,909,441,465]" pageId="1" pageNumber="398" type="etymology">
<paragraph id="8B9736DFFF8EFFF396B8A9D8FC36FE34" blockId="1.[151,1437,151,1869]" box="[199,909,441,465]" pageId="1" pageNumber="398">
<emphasis id="B95CEACDFF8EFFF396B8A9D8FEF3FE34" bold="true" box="[199,328,441,465]" inLineHeading="true" pageId="1" pageNumber="398" reason="1">Etymology.</emphasis>
The name refers to the
<typeStatus id="5493887DFF8EFFF3943BA9D8FDC9FE34" box="[580,626,441,465]" pageId="1" pageNumber="398">type</typeStatus>
locality, Hateruma Island.
</paragraph>
</subSubSection>
<subSubSection id="C3326554FF8EFFF396B8A9BCFD6CFE10" box="[199,727,477,501]" pageId="1" pageNumber="398" type="distribution">
<paragraph id="8B9736DFFF8EFFF396B8A9BCFD6CFE10" blockId="1.[151,1437,151,1869]" box="[199,727,477,501]" pageId="1" pageNumber="398">
<emphasis id="B95CEACDFF8EFFF396B8A9BCFEE1FE10" bold="true" box="[199,346,477,501]" inLineHeading="true" pageId="1" pageNumber="398" reason="1">Distribution.</emphasis>
Known only from the
<typeStatus id="5493887DFF8EFFF39432A9BCFDC0FE10" box="[589,635,477,501]" pageId="1" pageNumber="398">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C3326554FF8EFFF396B8AA60FC49FD84" pageId="1" pageNumber="398" type="diagnosis">
<paragraph id="8B9736DFFF8EFFF396B8AA60FC49FD84" blockId="1.[151,1437,151,1869]" pageId="1" pageNumber="398">
<emphasis id="B95CEACDFF8EFFF396B8AA60FE87FDFC" bold="true" box="[199,316,513,537]" inLineHeading="true" pageId="1" pageNumber="398" reason="1">Diagnosis.</emphasis>
Species of
<taxonomicName id="4C284D5CFF8EFFF397B4AA60FDDFFDFC" authorityName="Waren" authorityYear="1980" box="[459,612,513,537]" class="Gastropoda" family="Eulimidae" genus="Concavibalcis" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="398" phylum="Mollusca" rank="genus">
<emphasis id="B95CEACDFF8EFFF397B4AA60FDDFFDFC" box="[459,612,513,537]" italics="true" pageId="1" pageNumber="398">Concavibalcis</emphasis>
</taxonomicName>
with a small, thick, white translucent shell, lacking chalky appearance. Teleoconch surface decorated with very fine spiral ridges. Aperture ovate, callused; outer lip simple, curved, constricted below suture, with most protruding part at half of aperture height in lateral view.
</paragraph>
</subSubSection>
<subSubSection id="C3326554FF8EFFF396B8AA0CFA34FC08" pageId="1" pageNumber="398" type="description">
<paragraph id="8B9736DFFF8EFFF396B8AA0CFB82FC2C" blockId="1.[151,1437,151,1869]" pageId="1" pageNumber="398">
<emphasis id="B95CEACDFF8EFFF396B8AA0CFEE9FD60" bold="true" box="[199,338,621,645]" inLineHeading="true" pageId="1" pageNumber="398" reason="1">Description.</emphasis>
Shell conical with blunt apex, up to
<quantity id="4CD09B3AFF8EFFF39577AA0CFCE4FD61" box="[776,863,621,645]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="1" pageNumber="398" unit="mm" value="3.1">3.1 mm</quantity>
high, thick, solid, white translucent (
<figureCitation id="13132A5AFF8EFFF39370AA0CFA34FD60" box="[1295,1423,621,645]" captionStart="FIGURE 1" captionStartId="2.[151,250,1431,1455]" captionTargetBox="[151,1436,181,1408]" captionTargetId="figure-27@2.[151,1436,181,1408]" captionTargetPageId="2" captionText="FIGURE 1. Concavibalcis haterumaensis n. sp. A. Holotype NSMT-Mo 79320, 3.0 mm in shell height. B–E. Paratype MPM Coll. No. 21794a, 2.0 mm, SEM. B. Lateral view showing incremental scars in penultimate and last whorls. C. Close-up view of blunt apex; arrowheads denote demarcation line between protoconch and teleoconch. D. Penultimate whorl with an incremental line and fine spiral ridges. E. Surface of first teleoconch whorl showing short oblique ridges and continuous spiral ridges. Scale bars = 1 mm (A), 500 µm (B) and 100 µm (C–E)." figureDoi="http://doi.org/10.5281/zenodo.5910031" httpUri="https://zenodo.org/record/5910031/files/figure.png" pageId="1" pageNumber="398">Fig. 1A, B</figureCitation>
). Protoconch dome-shaped, smooth, glassy translucent white, with 1.3 whorls; demarcation line with teleoconch faint, straight, opisthocline; exposed part 220 μm wide and 250 μm high (
<figureCitation id="13132A5AFF8EFFF3950BAAD4FC73FD28" box="[884,968,693,717]" captionStart="FIGURE 1" captionStartId="2.[151,250,1431,1455]" captionTargetBox="[151,1436,181,1408]" captionTargetId="figure-27@2.[151,1436,181,1408]" captionTargetPageId="2" captionText="FIGURE 1. Concavibalcis haterumaensis n. sp. A. Holotype NSMT-Mo 79320, 3.0 mm in shell height. B–E. Paratype MPM Coll. No. 21794a, 2.0 mm, SEM. B. Lateral view showing incremental scars in penultimate and last whorls. C. Close-up view of blunt apex; arrowheads denote demarcation line between protoconch and teleoconch. D. Penultimate whorl with an incremental line and fine spiral ridges. E. Surface of first teleoconch whorl showing short oblique ridges and continuous spiral ridges. Scale bars = 1 mm (A), 500 µm (B) and 100 µm (C–E)." figureDoi="http://doi.org/10.5281/zenodo.5910031" httpUri="https://zenodo.org/record/5910031/files/figure.png" pageId="1" pageNumber="398">Fig. 1C</figureCitation>
). Up to 6.4 teleconch whorls, concave near shoulder and convex below; suture well defined and encircled by a rounded keel (
<figureCitation id="13132A5AFF8EFFF3958EAAB8FBF9FD14" box="[1009,1090,729,753]" captionStart="FIGURE 1" captionStartId="2.[151,250,1431,1455]" captionTargetBox="[151,1436,181,1408]" captionTargetId="figure-27@2.[151,1436,181,1408]" captionTargetPageId="2" captionText="FIGURE 1. Concavibalcis haterumaensis n. sp. A. Holotype NSMT-Mo 79320, 3.0 mm in shell height. B–E. Paratype MPM Coll. No. 21794a, 2.0 mm, SEM. B. Lateral view showing incremental scars in penultimate and last whorls. C. Close-up view of blunt apex; arrowheads denote demarcation line between protoconch and teleoconch. D. Penultimate whorl with an incremental line and fine spiral ridges. E. Surface of first teleoconch whorl showing short oblique ridges and continuous spiral ridges. Scale bars = 1 mm (A), 500 µm (B) and 100 µm (C–E)." figureDoi="http://doi.org/10.5281/zenodo.5910031" httpUri="https://zenodo.org/record/5910031/files/figure.png" pageId="1" pageNumber="398">Fig. 1B</figureCitation>
); incremental scars distinct (
<figureCitation id="13132A5AFF8EFFF3930EAAB8FF02FCF0" captionStart="FIGURE 1" captionStartId="2.[151,250,1431,1455]" captionTargetBox="[151,1436,181,1408]" captionTargetId="figure-27@2.[151,1436,181,1408]" captionTargetPageId="2" captionText="FIGURE 1. Concavibalcis haterumaensis n. sp. A. Holotype NSMT-Mo 79320, 3.0 mm in shell height. B–E. Paratype MPM Coll. No. 21794a, 2.0 mm, SEM. B. Lateral view showing incremental scars in penultimate and last whorls. C. Close-up view of blunt apex; arrowheads denote demarcation line between protoconch and teleoconch. D. Penultimate whorl with an incremental line and fine spiral ridges. E. Surface of first teleoconch whorl showing short oblique ridges and continuous spiral ridges. Scale bars = 1 mm (A), 500 µm (B) and 100 µm (C–E)." figureDoi="http://doi.org/10.5281/zenodo.5910031" httpUri="https://zenodo.org/record/5910031/files/figure.png" pageId="1" pageNumber="398">Fig. 1D</figureCitation>
), situated at 0.7, 1.6, 2.0, 2.7, 3.4, 4.1, 4.5, 5.0 and 5.5 whorls from aperture in
<typeStatus id="5493887DFF8EFFF3927DAA9CFBD8FCF0" box="[1026,1123,765,789]" pageId="1" pageNumber="398" type="holotype">holotype</typeStatus>
. First two to three teleoconch whorls bear short, oblique, slightly curved ridges on upper two-thirds and continuous spiral ridges on lower one-third of each whorl (
<figureCitation id="13132A5AFF8EFFF39762AB24FED4FCB8" box="[285,367,837,861]" captionStart="FIGURE 1" captionStartId="2.[151,250,1431,1455]" captionTargetBox="[151,1436,181,1408]" captionTargetId="figure-27@2.[151,1436,181,1408]" captionTargetPageId="2" captionText="FIGURE 1. Concavibalcis haterumaensis n. sp. A. Holotype NSMT-Mo 79320, 3.0 mm in shell height. B–E. Paratype MPM Coll. No. 21794a, 2.0 mm, SEM. B. Lateral view showing incremental scars in penultimate and last whorls. C. Close-up view of blunt apex; arrowheads denote demarcation line between protoconch and teleoconch. D. Penultimate whorl with an incremental line and fine spiral ridges. E. Surface of first teleoconch whorl showing short oblique ridges and continuous spiral ridges. Scale bars = 1 mm (A), 500 µm (B) and 100 µm (C–E)." figureDoi="http://doi.org/10.5281/zenodo.5910031" httpUri="https://zenodo.org/record/5910031/files/figure.png" pageId="1" pageNumber="398">Fig. 1E</figureCitation>
). Later whorls bear spirals only, which are stronger and more widely spaced towards suture of next whorl (
<figureCitation id="13132A5AFF8EFFF3969AAB08FE81FC64" box="[229,314,873,897]" captionStart="FIGURE 1" captionStartId="2.[151,250,1431,1455]" captionTargetBox="[151,1436,181,1408]" captionTargetId="figure-27@2.[151,1436,181,1408]" captionTargetPageId="2" captionText="FIGURE 1. Concavibalcis haterumaensis n. sp. A. Holotype NSMT-Mo 79320, 3.0 mm in shell height. B–E. Paratype MPM Coll. No. 21794a, 2.0 mm, SEM. B. Lateral view showing incremental scars in penultimate and last whorls. C. Close-up view of blunt apex; arrowheads denote demarcation line between protoconch and teleoconch. D. Penultimate whorl with an incremental line and fine spiral ridges. E. Surface of first teleoconch whorl showing short oblique ridges and continuous spiral ridges. Scale bars = 1 mm (A), 500 µm (B) and 100 µm (C–E)." figureDoi="http://doi.org/10.5281/zenodo.5910031" httpUri="https://zenodo.org/record/5910031/files/figure.png" pageId="1" pageNumber="398">Fig. 1D</figureCitation>
). Last adult whorl occupies 55% of total shell height. Aperture ovate; outer lip simple, curved, retracted near suture, sigmoid with most protruding part at half of aperture height in lateral view; parietal wall and columellar lip joining in a slight arc; columellar and parietal callus well developed; basal lip rounded.
</paragraph>
<paragraph id="8B9736DFFF8EFFF396B8ABB4FA34FC08" blockId="1.[151,1437,151,1869]" box="[199,1423,981,1005]" pageId="1" pageNumber="398">
<emphasis id="B95CEACDFF8EFFF396B8ABB4FEE9FC08" bold="true" box="[199,338,981,1005]" inLineHeading="true" pageId="1" pageNumber="398" reason="1">Dimensions.</emphasis>
<typeStatus id="5493887DFF8EFFF39727ABB4FE04FC08" box="[344,447,981,1005]" pageId="1" pageNumber="398" type="holotype">Holotype</typeStatus>
: 3.0 mm in SH and
<quantity id="4CD09B3AFF8EFFF394ECABB4FD5FFC09" box="[659,740,981,1005]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="398" unit="mm" value="1.2">1.2 mm</quantity>
in SD;
<specimenCount id="9D2EFD56FF8EFFF39548ABB4FC7AFC08" box="[823,961,981,1005]" pageId="1" pageNumber="398" type="generic" typeStatus="paratypes">11 paratypes</specimenCount>
: 2.0–
<quantity id="4CD09B3AFF8EFFF39583ABB4FBF0FC09" box="[1020,1099,981,1005]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="1" pageNumber="398" unit="mm" value="3.1">3.1 mm</quantity>
in SH and
<quantity id="4CD09B3AFF8EFFF392BBABB4FAF8FC09" box="[1220,1347,981,1005]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="1" pageNumber="398" unit="mm" value="1.0" valueMax="1.2" valueMin="0.8">0.8–1.2 mm</quantity>
in SD.
</paragraph>
</subSubSection>
<subSubSection id="C3326554FF8EFFF396B8AB98FD76F8A9" pageId="1" pageNumber="398" type="discussion">
<paragraph id="8B9736DFFF8EFFF396B8AB98FC60FAE8" blockId="1.[151,1437,151,1869]" pageId="1" pageNumber="398">
<emphasis id="B95CEACDFF8EFFF396B8AB98FE8DFBF4" bold="true" box="[199,310,1017,1041]" inLineHeading="true" pageId="1" pageNumber="398" reason="1">Remarks.</emphasis>
The assignment of the present new species to
<taxonomicName id="4C284D5CFF8EFFF3953BAB98FC66FBF4" authorityName="Waren" authorityYear="1980" box="[836,989,1017,1041]" class="Gastropoda" family="Eulimidae" genus="Concavibalcis" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="398" phylum="Mollusca" rank="genus">
<emphasis id="B95CEACDFF8EFFF3953BAB98FC66FBF4" box="[836,989,1017,1041]" italics="true" pageId="1" pageNumber="398">Concavibalcis</emphasis>
</taxonomicName>
is supported by its shouldered, concave teleoconch whorls, and its blunt apex with a dome-shaped paucispiral protoconch (
<figureCitation id="13132A5AFF8EFFF39259AC7CFBD2FBD0" box="[1062,1129,1053,1077]" captionStart="FIGURE 1" captionStartId="2.[151,250,1431,1455]" captionTargetBox="[151,1436,181,1408]" captionTargetId="figure-27@2.[151,1436,181,1408]" captionTargetPageId="2" captionText="FIGURE 1. Concavibalcis haterumaensis n. sp. A. Holotype NSMT-Mo 79320, 3.0 mm in shell height. B–E. Paratype MPM Coll. No. 21794a, 2.0 mm, SEM. B. Lateral view showing incremental scars in penultimate and last whorls. C. Close-up view of blunt apex; arrowheads denote demarcation line between protoconch and teleoconch. D. Penultimate whorl with an incremental line and fine spiral ridges. E. Surface of first teleoconch whorl showing short oblique ridges and continuous spiral ridges. Scale bars = 1 mm (A), 500 µm (B) and 100 µm (C–E)." figureDoi="http://doi.org/10.5281/zenodo.5910031" httpUri="https://zenodo.org/record/5910031/files/figure.png" pageId="1" pageNumber="398">Fig. 1</figureCitation>
;
<bibRefCitation id="EFB94B2EFF8EFFF39207AC7CFAB6FBD0" author="Waren, A." box="[1144,1293,1053,1077]" pageId="1" pageNumber="398" pagination="283 - 306" refId="ref2452" refString="Waren, A. (1980 b) Descriptions of new taxa of Eulimidae (Mollusca, Prosobranchia), with notes on some previously described genera. Zoologica Scripta, 9, 283 - 306. https: // doi. org / 10.1111 / j. 1463 - 6409.1980. tb 00668. x" type="journal article" year="1980">Warén 1980b</bibRefCitation>
). The former condition is rare among the
<taxonomicName id="4C284D5CFF8EFFF397B1AC20FD87FBBC" authorityName="Philippi" authorityYear="1853" box="[462,572,1089,1113]" class="Gastropoda" family="Eulimidae" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="398" phylum="Mollusca" rank="family">Eulimidae</taxonomicName>
and found in only a few other genera, including
<taxonomicName id="4C284D5CFF8EFFF39223AC20FAD7FBBC" authority="de Folin, 1870" authorityName="de Folin" authorityYear="1870" box="[1116,1388,1089,1113]" class="Gastropoda" family="Eulimidae" genus="Oceanida" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="398" phylum="Mollusca" rank="genus">
<emphasis id="B95CEACDFF8EFFF39223AC20FB7FFBBC" box="[1116,1220,1089,1113]" italics="true" pageId="1" pageNumber="398">Oceanida</emphasis>
de Folin, 1870
</taxonomicName>
and
<taxonomicName id="4C284D5CFF8EFFF396E8AC04FE06FB98" authority="Waren, 1980" authorityName="Waren" authorityYear="1980" box="[151,445,1125,1149]" class="Gastropoda" family="Eulimidae" genus="Scalaribalcis" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="398" phylum="Mollusca" rank="genus">
<emphasis id="B95CEACDFF8EFFF396E8AC04FE9FFB98" box="[151,292,1125,1149]" italics="true" pageId="1" pageNumber="398">Scalaribalcis</emphasis>
Warén, 1980
</taxonomicName>
. However,
<taxonomicName id="4C284D5CFF8EFFF39444AC04FD18FB98" authorityName="de Folin" authorityYear="1870" box="[571,675,1125,1149]" class="Gastropoda" family="Eulimidae" genus="Oceanida" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="398" phylum="Mollusca" rank="genus">
<emphasis id="B95CEACDFF8EFFF39444AC04FD18FB98" box="[571,675,1125,1149]" italics="true" pageId="1" pageNumber="398">Oceanida</emphasis>
</taxonomicName>
differs in having a taller, pupiform protoconch, and
<taxonomicName id="4C284D5CFF8EFFF3928EAC04FAC5FB98" authorityName="Waren" authorityYear="1980" box="[1265,1406,1125,1149]" class="Gastropoda" family="Eulimidae" genus="Scalaribalcis" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="398" phylum="Mollusca" rank="genus">
<emphasis id="B95CEACDFF8EFFF3928EAC04FAC5FB98" box="[1265,1406,1125,1149]" italics="true" pageId="1" pageNumber="398">Scalaribalcis</emphasis>
</taxonomicName>
in having an ovate outline of the teleoconch with a much broader aperture (
<bibRefCitation id="EFB94B2EFF8EFFF395D5ACE8FB84FB44" author="Waren, A." box="[938,1087,1161,1185]" pageId="1" pageNumber="398" pagination="1 - 96" refId="ref2568" refString="Waren, A. (1983 a) A generic revision of the family Eulimidae (Gastropoda, Prosobranchia). Journal of Molluscan Studies, Supplement 13, 1 - 96. https: // doi. org / 10.1093 / mollus / 49. Supplement _ 13.1" type="journal article" year="1983">Warén 1983a</bibRefCitation>
). The shell apex is blunt in such genera as
<taxonomicName id="4C284D5CFF8EFFF39774ACCCFD88FB20" authority="Laseron, 1955" authorityName="Laseron" authorityYear="1955" box="[267,563,1197,1221]" class="Gastropoda" family="Eulimidae" genus="Fusceulima" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="398" phylum="Mollusca" rank="genus">
<emphasis id="B95CEACDFF8EFFF39774ACCCFE3CFB20" box="[267,391,1197,1221]" italics="true" pageId="1" pageNumber="398">Fusceulima</emphasis>
Laseron, 1955
</taxonomicName>
,
<taxonomicName id="4C284D5CFF8EFFF3943CACCCFC5AFB20" authority="Bouchet &amp; Waren, 1986" authorityName="Bouchet &amp; Waren" authorityYear="1986" box="[579,993,1197,1221]" class="Gastropoda" family="Eulimidae" genus="Halielloides" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="398" phylum="Mollusca" rank="genus">
<emphasis id="B95CEACDFF8EFFF3943CACCCFD7EFB20" box="[579,709,1197,1221]" italics="true" pageId="1" pageNumber="398">Halielloides</emphasis>
Bouchet &amp; Warén, 1986
</taxonomicName>
and
<taxonomicName id="4C284D5CFF8EFFF39260ACCFFA95FB20" authority="Iredale, 1918" authorityName="Iredale" authorityYear="1918" box="[1055,1326,1197,1221]" class="Gastropoda" family="Eulimidae" genus="Teretianax" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="398" phylum="Mollusca" rank="genus">
<emphasis id="B95CEACDFF8EFFF39260ACCFFB34FB20" box="[1055,1167,1198,1221]" italics="true" pageId="1" pageNumber="398">Teretianax</emphasis>
Iredale, 1918
</taxonomicName>
, but their teleoconch features differ greatly from those of the present species and of
<taxonomicName id="4C284D5CFF8EFFF395D8ACB0FBA1FB0C" authorityName="Waren" authorityYear="1980" box="[935,1050,1233,1257]" class="Gastropoda" family="Eulimidae" genus="Concavibalcis" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="398" phylum="Mollusca" rank="species" species="scalaris">
<emphasis id="B95CEACDFF8EFFF395D8ACB0FBA1FB0C" box="[935,1050,1233,1257]" italics="true" pageId="1" pageNumber="398">C. scalaris</emphasis>
</taxonomicName>
, the
<typeStatus id="5493887DFF8EFFF39231ACB0FBC7FB0C" box="[1102,1148,1233,1257]" pageId="1" pageNumber="398">type</typeStatus>
and only known species of
<taxonomicName id="4C284D5CFF8EFFF396E8AC94FE8BFAE8" authorityName="Waren" authorityYear="1980" box="[151,304,1269,1293]" class="Gastropoda" family="Eulimidae" genus="Concavibalcis" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="398" phylum="Mollusca" rank="genus">
<emphasis id="B95CEACDFF8EFFF396E8AC94FE8BFAE8" box="[151,304,1269,1293]" italics="true" pageId="1" pageNumber="398">Concavibalcis</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFB94B2EFF8EFFF39742AC94FE6BFAE8" author="Waren, A." box="[317,464,1269,1293]" pageId="1" pageNumber="398" pagination="273 - 294" refId="ref2617" refString="Waren, A. (1983 b) An anatomical description of Eulima bilineata Alder with remarks on and a revision of Pyramidelloides Nevill (Mollusca, Prosobranchia, Eulimidae). Zoologica Scripta, 12, 273 - 294. https: // doi. org / 10.1111 / j. 1463 - 6409.1983. tb 00511. x" type="journal article" year="1983">Warén 1983b</bibRefCitation>
;
<bibRefCitation id="EFB94B2EFF8EFFF397A3AC94FD6DFAE8" author="Souza, L. S. &amp; Pimenta, A. D." box="[476,726,1269,1293]" pageId="1" pageNumber="398" pagination="621 - 633" refId="ref2221" refString="Souza, L. S. &amp; Pimenta, A. D. (2014) Fusceulima and Halielloides (Gastropoda: Eulimidae) in the southwestern Atlantic, with descriptions of two new species of Fusceulima. Zoologia, 31, 621 - 633. https: // doi. org / 10.1590 / S 1984 - 46702014000600008" type="journal article" year="2014">Souza &amp; Pimenta 2014</bibRefCitation>
;
<bibRefCitation id="EFB94B2EFF8EFFF3949FAC94FC6BFAE8" author="Hori, S. &amp; Matsuda, H." box="[736,976,1269,1293]" pageId="1" pageNumber="398" pagination="823 - 832" refId="ref2118" refString="Hori, S. &amp; Matsuda, H. (2017) Family Eulimidae. In: Okutani, T. (Ed.), Marine Mollusks in Japan, the Second Edition. Tokai University Press, Hiratsuka, pp. 142 - 151, 823 - 832." type="journal article" year="2017">Hori &amp; Matsuda 2017</bibRefCitation>
).
</paragraph>
<paragraph id="8B9736DFFF8EFFF396B8AD78FE61F958" blockId="1.[151,1437,151,1869]" pageId="1" pageNumber="398">
<taxonomicName id="4C284D5CFF8EFFF396B8AD78FDBFFAD4" authority="Takano &amp; Tsuzuki &amp; Kano, 2022" authorityName="Takano &amp; Tsuzuki &amp; Kano" authorityYear="2022" box="[199,516,1305,1329]" class="Gastropoda" family="Eulimidae" genus="Concavibalcis" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="398" phylum="Mollusca" rank="species" species="haterumaensis" status="sp. nov.">
<emphasis id="B95CEACDFF8EFFF396B8AD78FDBFFAD4" box="[199,516,1305,1329]" italics="true" pageId="1" pageNumber="398">Concavibalcis haterumaensis</emphasis>
</taxonomicName>
<emphasis id="B95CEACDFF8EFFF39473AD78FDF2FAD4" bold="true" box="[524,585,1305,1329]" pageId="1" pageNumber="398">
<taxonomicNameLabel id="A26F57B6FF8EFFF39473AD78FDF2FAD4" box="[524,585,1305,1329]" pageId="1" pageNumber="398" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is distinguished from
<taxonomicName id="4C284D5CFF8EFFF39541AD78FC08FAD4" authorityName="Waren" authorityYear="1980" box="[830,947,1305,1329]" class="Gastropoda" family="Eulimidae" genus="Concavibalcis" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="398" phylum="Mollusca" rank="species" species="scalaris">
<emphasis id="B95CEACDFF8EFFF39541AD78FC08FAD4" box="[830,947,1305,1329]" italics="true" pageId="1" pageNumber="398">C. scalaris</emphasis>
</taxonomicName>
in having (1) a smaller, thicker shell and (2) fine spiral ridges on teleoconch whorls. The two species have nearly the same number of teleoconch whorls (up to 6.4 and 6.5, respectively), meaning that
<taxonomicName id="4C284D5CFF8EFFF39792AD00FD1CFA9C" authority="Takano &amp; Tsuzuki &amp; Kano, 2022" authorityName="Takano &amp; Tsuzuki &amp; Kano" authorityYear="2022" box="[493,679,1377,1401]" class="Gastropoda" family="Eulimidae" genus="Concavibalcis" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="398" phylum="Mollusca" rank="species" species="haterumaensis" status="sp. nov.">
<emphasis id="B95CEACDFF8EFFF39792AD00FD1CFA9C" box="[493,679,1377,1401]" italics="true" pageId="1" pageNumber="398">C. haterumaensis</emphasis>
</taxonomicName>
<emphasis id="B95CEACDFF8EFFF394D2AD00FD52FA9C" bold="true" box="[685,745,1377,1401]" pageId="1" pageNumber="398">
<taxonomicNameLabel id="A26F57B6FF8EFFF394D2AD00FD52FA9C" box="[685,745,1377,1401]" pageId="1" pageNumber="398" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<quantity id="4CD09B3AFF8EFFF39488AD00FCF2FA9D" box="[759,841,1377,1401]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="1" pageNumber="398" unit="mm" value="3.1">3.1 mm</quantity>
high) is more tightly coiled with a lower expansion rate of the aperture than
<taxonomicName id="4C284D5CFF8EFFF39712ADE4FE5BFA78" authorityName="Waren" authorityYear="1980" box="[365,480,1413,1437]" class="Gastropoda" family="Eulimidae" genus="Concavibalcis" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="398" phylum="Mollusca" rank="species" species="scalaris">
<emphasis id="B95CEACDFF8EFFF39712ADE4FE5BFA78" box="[365,480,1413,1437]" italics="true" pageId="1" pageNumber="398">C. scalaris</emphasis>
</taxonomicName>
(
<quantity id="4CD09B3AFF8EFFF39791ADE4FDFBFA79" box="[494,576,1413,1437]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="1" pageNumber="398" unit="mm" value="5.1">5.1 mm</quantity>
high). The presence or absence of fine spirals is a species-specific character also in other eulimid genera (e.g.
<taxonomicName id="4C284D5CFF8EFFF397B5ADC8FD61FA24" authority="Habe, 1965" authorityName="Habe" authorityYear="1965" box="[458,730,1449,1473]" class="Gastropoda" family="Eulimidae" genus="Annulobalcis" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="398" phylum="Mollusca" rank="genus">
<emphasis id="B95CEACDFF8EFFF397B5ADC8FDECFA24" box="[458,599,1449,1473]" italics="true" pageId="1" pageNumber="398">Annulobalcis</emphasis>
Habe, 1965
</taxonomicName>
,
<taxonomicName id="4C284D5CFF8EFFF3949AADC8FC42FA24" authority="Bowdich, 1822" authorityName="Bowdich" authorityYear="1822" box="[741,1017,1449,1473]" class="Gastropoda" family="Eulimidae" genus="Melanella" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="398" phylum="Mollusca" rank="genus">
<emphasis id="B95CEACDFF8EFFF3949AADC8FCEAFA24" box="[741,849,1449,1473]" italics="true" pageId="1" pageNumber="398">Melanella</emphasis>
Bowdich, 1822
</taxonomicName>
and
<taxonomicName id="4C284D5CFF8EFFF39252ADC8FAF5FA24" authority="Waren, 1980" authorityName="Waren" authorityYear="1980" box="[1069,1358,1449,1473]" class="Gastropoda" family="Eulimidae" genus="Trochostilifer" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="398" phylum="Mollusca" rank="genus">
<emphasis id="B95CEACDFF8EFFF39252ADC8FB06FA24" box="[1069,1213,1449,1473]" italics="true" pageId="1" pageNumber="398">Trochostilifer</emphasis>
Warén, 1980
</taxonomicName>
;
<bibRefCitation id="EFB94B2EFF8EFFF39328ADC8FF66FA00" author="Waren, A." pageId="1" pageNumber="398" pagination="283 - 306" refId="ref2452" refString="Waren, A. (1980 b) Descriptions of new taxa of Eulimidae (Mollusca, Prosobranchia), with notes on some previously described genera. Zoologica Scripta, 9, 283 - 306. https: // doi. org / 10.1111 / j. 1463 - 6409.1980. tb 00668. x" type="journal article" year="1980">Warén 1980b</bibRefCitation>
;
<bibRefCitation id="EFB94B2EFF8EFFF39694ADACFE16FA00" author="Gofas, S. &amp; Moreno, D. &amp; Salas, C." box="[235,429,1485,1509]" pageId="1" pageNumber="398" refId="ref2038" refString="Gofas, S., Moreno, D. &amp; Salas, C. (2011) Moluscos marinos de Andalucia, vol. I. Servicio de Publicaciones e Intercambio Cientifico. Universidad de Malaga, Malaga, 342 pp." type="book" year="2011">
Gofas
<emphasis id="B95CEACDFF8EFFF3974BADAFFED7FA00" box="[308,364,1485,1509]" italics="true" pageId="1" pageNumber="398">et al.</emphasis>
2011
</bibRefCitation>
;
<bibRefCitation id="EFB94B2EFF8EFFF397C5ADACFD0CFA00" author="Dgebuadze, P. Y. &amp; Fedosov, A. E. &amp; Kantor, Y. I." box="[442,695,1485,1509]" pageId="1" pageNumber="398" pagination="139 - 155" refId="ref1959" refString="Dgebuadze, P. Y., Fedosov, A. E. &amp; Kantor, Y. I. (2012) Host specificity of parasitic gastropods of the genus Annulobalcis Habe, 1965 (Mollusca, Gastropoda, Eulimidae) from crinoids in Vietnam, with descriptions of four new species. Zoosystema, 34, 139 - 155." type="journal article" year="2012">
Dgebuadze
<emphasis id="B95CEACDFF8EFFF39444ADAFFDCFFA00" box="[571,628,1485,1509]" italics="true" pageId="1" pageNumber="398">et al.</emphasis>
2012
</bibRefCitation>
). Furthermore, (3) the polished translucent shell of the new species differs from the chalky shell of the
<typeStatus id="5493887DFF8EFFF39472AD90FD80F9EC" box="[525,571,1521,1545]" pageId="1" pageNumber="398">type</typeStatus>
species, although the latter chalky appearance may be attributable to corrosion in the specimens examined (see
<bibRefCitation id="EFB94B2EFF8EFFF397A8AE74FDD1F9C8" author="Waren, A." box="[471,618,1557,1581]" pageId="1" pageNumber="398" pagination="283 - 306" refId="ref2452" refString="Waren, A. (1980 b) Descriptions of new taxa of Eulimidae (Mollusca, Prosobranchia), with notes on some previously described genera. Zoologica Scripta, 9, 283 - 306. https: // doi. org / 10.1111 / j. 1463 - 6409.1980. tb 00668. x" type="journal article" year="1980">Warén 1980b</bibRefCitation>
). Their habitats also differ from each other: (4)
<taxonomicName id="4C284D5CFF8EFFF39213AE74FA93F9C8" authority="Takano &amp; Tsuzuki &amp; Kano, 2022" authorityName="Takano &amp; Tsuzuki &amp; Kano" authorityYear="2022" box="[1132,1320,1557,1581]" class="Gastropoda" family="Eulimidae" genus="Concavibalcis" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="398" phylum="Mollusca" rank="species" species="haterumaensis" status="sp. nov.">
<emphasis id="B95CEACDFF8EFFF39213AE74FA93F9C8" box="[1132,1320,1557,1581]" italics="true" pageId="1" pageNumber="398">C. haterumaensis</emphasis>
</taxonomicName>
<emphasis id="B95CEACDFF8EFFF39350AE74FAD0F9C8" bold="true" box="[1327,1387,1557,1581]" pageId="1" pageNumber="398">
<taxonomicNameLabel id="A26F57B6FF8EFFF39350AE74FAD0F9C8" box="[1327,1387,1557,1581]" pageId="1" pageNumber="398" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
was collected from a shallow subtidal site, whereas
<taxonomicName id="4C284D5CFF8EFFF394E3AE58FCABF9B4" authorityName="Waren" authorityYear="1980" box="[668,784,1593,1617]" class="Gastropoda" family="Eulimidae" genus="Concavibalcis" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="398" phylum="Mollusca" rank="species" species="scalaris">
<emphasis id="B95CEACDFF8EFFF394E3AE58FCABF9B4" box="[668,784,1593,1617]" italics="true" pageId="1" pageNumber="398">C. scalaris</emphasis>
</taxonomicName>
lives in deeper, bathyal waters (
<quantity id="4CD09B3AFF8EFFF3920DAE58FB4AF9B5" box="[1138,1265,1593,1617]" metricMagnitude="2" metricUnit="m" metricValue="3.825" metricValueMax="4.0" metricValueMin="3.65" pageId="1" pageNumber="398" unit="m" value="382.5" valueMax="400.0" valueMin="365.0">365–400 m</quantity>
;
<bibRefCitation id="EFB94B2EFF8EFFF39282AE58FA2AF9B4" author="Waren, A." box="[1277,1425,1593,1617]" pageId="1" pageNumber="398" pagination="283 - 306" refId="ref2452" refString="Waren, A. (1980 b) Descriptions of new taxa of Eulimidae (Mollusca, Prosobranchia), with notes on some previously described genera. Zoologica Scripta, 9, 283 - 306. https: // doi. org / 10.1111 / j. 1463 - 6409.1980. tb 00668. x" type="journal article" year="1980">Warén 1980b</bibRefCitation>
). Several eulimid genera exhibit a similar broad bathymetric range (e.g.
<taxonomicName id="4C284D5CFF8EFFF395E9AE3CFBB9F990" authorityName="Bowdich" authorityYear="1822" box="[918,1026,1629,1653]" class="Gastropoda" family="Eulimidae" genus="Melanella" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="398" phylum="Mollusca" rank="genus">
<emphasis id="B95CEACDFF8EFFF395E9AE3CFBB9F990" box="[918,1026,1629,1653]" italics="true" pageId="1" pageNumber="398">Melanella</emphasis>
</taxonomicName>
,
<taxonomicName id="4C284D5CFF8EFFF3926EAE3CFA23F990" authority="Koehler &amp; Vaney, 1908" authorityName="Koehler &amp; Vaney" authorityYear="1908" box="[1041,1432,1629,1653]" class="Gastropoda" family="Eulimidae" genus="Pelseneeria" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="398" phylum="Mollusca" rank="genus">
<emphasis id="B95CEACDFF8EFFF3926EAE3CFB34F990" box="[1041,1167,1629,1653]" italics="true" pageId="1" pageNumber="398">Pelseneeria</emphasis>
Koehler &amp; Vaney, 1908
</taxonomicName>
,
<taxonomicName id="4C284D5CFF8EFFF396E8AEE0FE3FF97C" authority="Iredale, 1925" authorityName="Iredale" authorityYear="1925" box="[151,388,1665,1689]" class="Gastropoda" family="Eulimidae" genus="Stilapex" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="398" phylum="Mollusca" rank="genus">
<emphasis id="B95CEACDFF8EFFF396E8AEE0FF56F97C" box="[151,237,1665,1689]" italics="true" pageId="1" pageNumber="398">Stilapex</emphasis>
Iredale, 1925
</taxonomicName>
and
<taxonomicName id="4C284D5CFF8EFFF397C4AEE0FCF2F97C" authority="H. Adams &amp; A. Adams, 1854" authorityName="H. Adams &amp; A. Adams" authorityYear="1854" box="[443,841,1665,1689]" class="Gastropoda" family="Eulimidae" genus="Thyca" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="398" phylum="Mollusca" rank="genus">
<emphasis id="B95CEACDFF8EFFF397C4AEE0FE47F97C" box="[443,508,1665,1689]" italics="true" pageId="1" pageNumber="398">Thyca</emphasis>
H. Adams &amp; A. Adams, 1854
</taxonomicName>
;
<bibRefCitation id="EFB94B2EFF8EFFF3952AAEE0FC5DF97C" author="Waren, A." box="[853,998,1665,1689]" pageId="1" pageNumber="398" pagination="187 - 210" refId="ref2395" refString="Waren, A. (1980 a) Revision of the genera Thyca, Stilifer, Scalenostoma, Mucronalia and Echineulima (Mollusca, Prosobranchia, Eulimidae). Zoologica Scripta, 9, 187 - 210. https: // doi. org / 10.1111 / j. 1463 - 6409.1980. tb 00663. x" type="journal article" year="1980">
Warén 1980
<bibRefCitation id="EFB94B2EFF8EFFF395A5AEE0FC5DF97C" author="Waren, A." box="[986,998,1665,1689]" pageId="1" pageNumber="398" pagination="133 - 154" refId="ref2508" refString="Waren, A. (1981) Revision of the genera Apicalia A. Adams and Stilapex Iredale and description of two new genera (Mollusca, Prosobranchia, Eulimidae). Zoologica Scripta, 10, 133 - 154. https: // doi. org / 10.1111 / j. 1463 - 6409.1981. tb 00491. x" type="journal article" year="1981">a</bibRefCitation>
</bibRefCitation>
, 1981,
<bibRefCitation id="EFB94B2EFF8EFFF39248AEE0FBC0F97C" author="Waren, A." box="[1079,1147,1665,1689]" pageId="1" pageNumber="398" pagination="1 - 96" refId="ref2568" refString="Waren, A. (1983 a) A generic revision of the family Eulimidae (Gastropoda, Prosobranchia). Journal of Molluscan Studies, Supplement 13, 1 - 96. https: // doi. org / 10.1093 / mollus / 49. Supplement _ 13.1" type="journal article" year="1983">1983a</bibRefCitation>
;
<bibRefCitation id="EFB94B2EFF8EFFF392F7AEE0FAC7F97C" author="Hori, S. &amp; Matsuda, H." box="[1160,1404,1665,1689]" pageId="1" pageNumber="398" pagination="823 - 832" refId="ref2118" refString="Hori, S. &amp; Matsuda, H. (2017) Family Eulimidae. In: Okutani, T. (Ed.), Marine Mollusks in Japan, the Second Edition. Tokai University Press, Hiratsuka, pp. 142 - 151, 823 - 832." type="journal article" year="2017">Hori &amp; Matsuda 2017</bibRefCitation>
; T. Takano personal observation).
</paragraph>
<paragraph id="8B9736DFFF8EFFF396B8AEA8FD76F8A9" blockId="1.[151,1437,151,1869]" pageId="1" pageNumber="398">
The genus
<taxonomicName id="4C284D5CFF8EFFF3973EAEA8FE61F904" authorityName="Waren" authorityYear="1980" box="[321,474,1737,1761]" class="Gastropoda" family="Eulimidae" genus="Concavibalcis" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="398" phylum="Mollusca" rank="genus">
<emphasis id="B95CEACDFF8EFFF3973EAEA8FE61F904" box="[321,474,1737,1761]" italics="true" pageId="1" pageNumber="398">Concavibalcis</emphasis>
</taxonomicName>
remains to be the least studied among the
<taxonomicName id="4C284D5CFF8EFFF395C7AEA8FB9DF904" authorityName="Philippi" authorityYear="1853" box="[952,1062,1737,1761]" class="Gastropoda" family="Eulimidae" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="398" phylum="Mollusca" rank="family">Eulimidae</taxonomicName>
from phylogenetic and ecological perspectives. Eulimid species in closely related genera exploit hosts of the same echinoderm class (
<bibRefCitation id="EFB94B2EFF8EFFF392CCAE8CFAFFF8E0" author="Waren, A." box="[1203,1348,1773,1797]" pageId="1" pageNumber="398" pagination="1 - 96" refId="ref2568" refString="Waren, A. (1983 a) A generic revision of the family Eulimidae (Gastropoda, Prosobranchia). Journal of Molluscan Studies, Supplement 13, 1 - 96. https: // doi. org / 10.1093 / mollus / 49. Supplement _ 13.1" type="journal article" year="1983">Warén 1983a</bibRefCitation>
;
<bibRefCitation id="EFB94B2EFF8EFFF39331AE8CFE92F8CC" author="Takano, T. &amp; Goto, R." pageId="1" pageNumber="398" pagination="5" refId="ref2334" refString="Takano, T. &amp; Goto, R. (2021) Molecular and morphological systematics of the crinoid-parasitic snail genus Goodingia (Mollusca: Caenogastropoda: Eulimidae) with new insights into intrafamilial phylogenetic relationships. Marine Biodiversity, 51, 5. https: // doi. org / 10.1007 / s 12526 - 020 - 01141 - 0" type="journal article" year="2021">Takano &amp; Goto 2021</bibRefCitation>
). Future phylogenetic reconstruction for
<taxonomicName id="4C284D5CFF8EFFF394A2AF70FCCDF8CC" authorityName="Waren" authorityYear="1980" box="[733,886,1809,1833]" class="Gastropoda" family="Eulimidae" genus="Concavibalcis" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="398" phylum="Mollusca" rank="genus">
<emphasis id="B95CEACDFF8EFFF394A2AF70FCCDF8CC" box="[733,886,1809,1833]" italics="true" pageId="1" pageNumber="398">Concavibalcis</emphasis>
</taxonomicName>
based on live-taken specimens may lead to a better understanding of its parasitic ecology and
<emphasis id="B95CEACDFF8EFFF39424AF57FD7CF8A8" box="[603,711,1846,1869]" italics="true" pageId="1" pageNumber="398">vice versa</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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