288 lines
34 KiB
XML
288 lines
34 KiB
XML
<document id="56A057A1151125F75B8DEA29524A21EF" ID-CLB-Dataset="20789" ID-DOI="10.11646/zootaxa.5093.5.2" ID-GBIF-Dataset="f349242c-a281-4590-aff8-0e52538be142" ID-ISSN="1175-5326" ID-Zenodo-Dep="5964429" ID-ZooBank="6D56585F-68D1-4AD0-83B6-87207480735B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1643698181357" checkinUser="plazi" docAuthor="Kamiński, Marcin Jan, Lumen, Ryan, Kanda, Kojun & Smith, Aaron Dennis" docDate="2022" docId="038187A07D0F151C7986CB44FA90FF79" docLanguage="en" docName="zootaxa.5093.5.2.pdf" docOrigin="Zootaxa 5093 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Blapstinus punctulatus Kamiński & Lumen & Kanda & Smith 2022" docType="treatment" docVersion="9" lastPageNumber="522" masterDocId="FFB8FFD87D0D15187911CD3AFFACFF80" masterDocTitle="Phylogenetic status of some unique species representing Blapstinina Mulsant & Rey (Tenebrionidae: Blaptinae: Opatrini), and implications for continued study of the subtribe" masterLastPageNumber="532" masterPageNumber="519" pageNumber="521" updateTime="1699304908880" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="9F9A9019719EA3408FA2620D2E77245F">Phylogenetic status of some unique species representing Blapstinina Mulsant & Rey (Tenebrionidae: Blaptinae: Opatrini), and implications for continued study of the subtribe</mods:title>
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<mods:namePart id="2A934D154FC258BC9DCB4E2A1F03140D">Kamiński, Marcin Jan</mods:namePart>
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<mods:affiliation id="508B5E152C0BC19467FEEA9ABE0FFDBA">Zoological Museum, Museum and Institute of Zoology, Polish Academy of Sciences, Wilcza 64, 00 - 679 Warszawa, Poland & Purdue University, Department of Entomology, 901 W. State Street, West Lafayette, IN 47907, USA https: // orcid. org / 0000 - 0002 - 1286 - 950 X</mods:affiliation>
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<mods:namePart id="4BFADD6155031995CB6AD40F948D4562">Lumen, Ryan</mods:namePart>
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<mods:nameIdentifier id="63194E3E08569ED4D6EE72C078F603EF" type="ORCID">0000-0002-3958-7596</mods:nameIdentifier>
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<mods:affiliation id="691C0B189B08C7B5CB329524D4969787">Zoological Museum, Museum and Institute of Zoology, Polish Academy of Sciences, Wilcza 64, 00 - 679 Warszawa, Poland & https: // orcid. org / 0000 - 0002 - 3958 - 7596</mods:affiliation>
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<mods:namePart id="7F4ADD1A4EA4BD17A6EEBF0CD7CE1B01">Kanda, Kojun</mods:namePart>
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<mods:nameIdentifier id="8B9505225F0FF602B9E2634A4E4B2F7C" type="ORCID">0000-0001-5561-8471</mods:nameIdentifier>
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<mods:affiliation id="40888E64AE583094415259437D0CEE5E">USDA Systematic Entomology Laboratory, c / o Smithsonian Institution, National Museum of Natural History, Washington, DC, U. S. A. https: // orcid. org / 0000 - 0001 - 5561 - 8471</mods:affiliation>
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<mods:namePart id="C37E4B88ECF19FDC135A097123940D9D">Smith, Aaron Dennis</mods:namePart>
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<mods:nameIdentifier id="88C1F7B84A5DA9F2485A7AA135E0F253" type="ORCID">0000-0002-1286-950X</mods:nameIdentifier>
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<mods:affiliation id="B35AEC2D13BB37CE9C6DB11E576B8AE7">Purdue University, Department of Entomology, 901 W. State Street, West Lafayette, IN 47907, USA https: // orcid. org / 0000 - 0002 - 1286 - 950 X</mods:affiliation>
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<mods:title id="59B47CD56F439312BDC3E71EC2CE6B85">Zootaxa</mods:title>
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<mods:part id="832B1A96924A2D5536DC1A928A9C7E8D">
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<mods:date id="804B525BB30A1E943DABCCB1F6424A1B">2022</mods:date>
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<mods:number id="CD653B3CD73DA6926B49CCC57EBEC1A5">5093</mods:number>
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<treatment id="038187A07D0F151C7986CB44FA90FF79" ID-DOI="http://doi.org/10.5281/zenodo.6204791" ID-GBIF-Taxon="193277090" ID-Zenodo-Dep="6204791" LSID="urn:lsid:plazi:treatment:038187A07D0F151C7986CB44FA90FF79" httpUri="http://treatment.plazi.org/id/038187A07D0F151C7986CB44FA90FF79" lastPageId="4" lastPageNumber="522" pageId="2" pageNumber="521">
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<subSubSection id="C332653D7D0F151A7986CB44FDBEF919" box="[151,530,1662,1689]" pageId="2" pageNumber="521" type="nomenclature">
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<paragraph id="8B9736B67D0F151A7986CB44FDBEF919" blockId="2.[151,1437,1662,2013]" box="[151,530,1662,1689]" pageId="2" pageNumber="521">
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<heading id="D0DF81DA7D0F151A7986CB44FDBEF919" bold="true" box="[151,530,1662,1689]" fontSize="11" level="1" pageId="2" pageNumber="521" reason="1">
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<emphasis id="B95CEAA47D0F151A7986CB44FDBEF919" bold="true" box="[151,530,1662,1689]" pageId="2" pageNumber="521">
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Status of
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<taxonomicName id="4C284D357D0F151A7816CB44FDBEF919" authorityName="Kamiński & Lumen & Kanda & Smith" authorityYear="2022" baseAuthorityName="Kamiński & Lumen & Kanda & Smith" baseAuthorityYear="1851" box="[263,530,1662,1689]" class="Insecta" family="Tenebrionidae" genus="Blapstinus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="521" phylum="Arthropoda" rank="species" species="punctulatus">
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<emphasis id="B95CEAA47D0F151A7816CB44FDBEF919" bold="true" box="[263,530,1662,1689]" italics="true" pageId="2" pageNumber="521">Blapstinus punctulatus</emphasis>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C332653D7D0F151C79D6CB99FA90FF79" lastPageId="4" lastPageNumber="523" pageId="2" pageNumber="521" type="description">
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<paragraph id="8B9736B67D0F151B79D6CB99FE25FE75" blockId="2.[151,1437,1662,2013]" lastBlockId="3.[151,1437,151,501]" lastPageId="3" lastPageNumber="522" pageId="2" pageNumber="521">
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Although molecular support for the placement of
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<taxonomicName id="4C284D357D0F151A7BFFCB99FC3CF93D" authorityName="Kamiński & Lumen & Kanda & Smith" authorityYear="2022" baseAuthorityName="Kamiński & Lumen & Kanda & Smith" baseAuthorityYear="1851" box="[750,912,1699,1725]" class="Insecta" family="Tenebrionidae" genus="Blapstinus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="521" phylum="Arthropoda" rank="species" species="punctulatus">
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<emphasis id="B95CEAA47D0F151A7BFFCB99FC3CF93D" box="[750,912,1699,1725]" italics="true" pageId="2" pageNumber="521">B. punctulatus</emphasis>
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</taxonomicName>
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is low, a review of available literature revealed that this species fell within the morphological concept of
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<taxonomicName id="4C284D357D0F151A7A30CBFDFC6AF961" authorityName="Marcuzzi" authorityYear="1954" box="[801,966,1735,1761]" class="Insecta" family="Tenebrionidae" genus="Austrocaribius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="521" phylum="Arthropoda" rank="genus">
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<emphasis id="B95CEAA47D0F151A7A30CBFDFC6AF961" box="[801,966,1735,1761]" italics="true" pageId="2" pageNumber="521">Austrocaribius</emphasis>
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</taxonomicName>
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(Marcuzzi 1982, 1986).
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<bibRefCitation id="EFB94B477D0F151A7DF1CBFDFA30F961" author="Marcuzzi, G." box="[1248,1436,1734,1761]" pageId="2" pageNumber="521" pagination="1 - 36" refId="ref9624" refString="Marcuzzi, G. (1954) Studies on the fauna of Curacao and other Caribbean Islands: no. 22. Tenebrionid beetles of Curacao, Aruba, Bonaire, and the Venezuelan Islands. Natuurwetenschappelijke Studiekring voor Suriname, 10, 1 - 36." type="journal article" year="1954">Marcuzzi (1954)</bibRefCitation>
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distinguished
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<taxonomicName id="4C284D357D0F151A7822CBD1FE74F885" authorityName="Marcuzzi" authorityYear="1954" box="[307,472,1771,1797]" class="Insecta" family="Tenebrionidae" genus="Austrocaribius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="521" phylum="Arthropoda" rank="genus">
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<emphasis id="B95CEAA47D0F151A7822CBD1FE74F885" box="[307,472,1771,1797]" italics="true" pageId="2" pageNumber="521">Austrocaribius</emphasis>
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</taxonomicName>
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from
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<taxonomicName id="4C284D357D0F151A7B0DCBD1FD3EF885" authorityName="Sturm" authorityYear="1826" box="[540,658,1771,1797]" class="Insecta" family="Tenebrionidae" genus="Blapstinus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="521" phylum="Arthropoda" rank="genus">
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<emphasis id="B95CEAA47D0F151A7B0DCBD1FD3EF885" box="[540,658,1771,1797]" italics="true" pageId="2" pageNumber="521">Blapstinus</emphasis>
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</taxonomicName>
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using the following characters: antennal segments eight-eleven transverse, segments six and seven more or less square, the remaining segments longer than wide; intercoxal process of prosternum developed posteriorly in an oval-lanceolate point, seen in profile prominent and abruptly truncate; penis widely projecting between the parameres, apical part of tegmen truncate, and contiguous. Upon examination of
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<typeStatus id="549388147D0F151A79F8CA41FEB5F815" box="[233,281,1915,1941]" pageId="2" pageNumber="521">type</typeStatus>
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material of
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<taxonomicName id="4C284D357D0F151A78B2CA41FDE4F815" authorityName="Marcuzzi" authorityYear="1954" box="[419,584,1915,1941]" class="Insecta" family="Tenebrionidae" genus="Austrocaribius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="521" phylum="Arthropoda" rank="genus">
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<emphasis id="B95CEAA47D0F151A78B2CA41FDE4F815" box="[419,584,1915,1941]" italics="true" pageId="2" pageNumber="521">Austrocaribius</emphasis>
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</taxonomicName>
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and comparison with representatives of
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<taxonomicName id="4C284D357D0F151A7D03CA41FB24F815" authorityName="Sturm" authorityYear="1826" box="[1042,1160,1915,1941]" class="Insecta" family="Tenebrionidae" genus="Blapstinus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="521" phylum="Arthropoda" rank="genus">
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<emphasis id="B95CEAA47D0F151A7D03CA41FB24F815" box="[1042,1160,1915,1941]" italics="true" pageId="2" pageNumber="521">Blapstinus</emphasis>
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</taxonomicName>
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, it is difficult to reliably separate
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<taxonomicName id="4C284D357D0F151A79EBCAA5FE33F839" authorityName="Marcuzzi" authorityYear="1954" box="[250,415,1951,1977]" class="Insecta" family="Tenebrionidae" genus="Austrocaribius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="521" phylum="Arthropoda" rank="genus">
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<emphasis id="B95CEAA47D0F151A79EBCAA5FE33F839" box="[250,415,1951,1977]" italics="true" pageId="2" pageNumber="521">Austrocaribius</emphasis>
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</taxonomicName>
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from
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<taxonomicName id="4C284D357D0F151A78F7CAA5FDF0F839" authorityName="Sturm" authorityYear="1826" box="[486,604,1951,1977]" class="Insecta" family="Tenebrionidae" genus="Blapstinus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="521" phylum="Arthropoda" rank="genus">
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<emphasis id="B95CEAA47D0F151A78F7CAA5FDF0F839" box="[486,604,1951,1977]" italics="true" pageId="2" pageNumber="521">Blapstinus</emphasis>
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</taxonomicName>
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using most of the characters Marcuzzi outlined (e.g., both
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<taxonomicName id="4C284D357D0F151A7DE7CAA5FAC0F839" authorityName="Sturm" authorityYear="1826" box="[1270,1388,1951,1977]" class="Insecta" family="Tenebrionidae" genus="Blapstinus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="521" phylum="Arthropoda" rank="genus">
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<emphasis id="B95CEAA47D0F151A7DE7CAA5FAC0F839" box="[1270,1388,1951,1977]" italics="true" pageId="2" pageNumber="521">Blapstinus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C284D357D0F151A7986CAF9FE90F85D" authorityName="Marcuzzi" authorityYear="1954" box="[151,316,1987,2013]" class="Insecta" family="Tenebrionidae" genus="Austrocaribius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="521" phylum="Arthropoda" rank="genus">
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<emphasis id="B95CEAA47D0F151A7986CAF9FE90F85D" box="[151,316,1987,2013]" italics="true" pageId="2" pageNumber="521">Austrocaribius</emphasis>
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</taxonomicName>
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have clavate antennae, resulting in the last 4 antennomeres becoming more-or-less transverse). The aedeagal characters remain useful and, as of this study, can be used to consistently delineate the two genera (
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<figureCitation id="13132A337D0E151B7C7ECDADFF08FF55" captionStart="FIGURE 1" captionStartId="3.[151,250,1675,1699]" captionTargetBox="[151,1419,525,1640]" captionTargetId="figure-169@3.[151,1436,525,1651]" captionTargetPageId="3" captionText="FIGURE 1. Phylogeny of subtribe Blapstinina. Figured topology was recovered in maximum likelihood analysis using IQ- Tree. Ultrafast bootstrap values (1,000 replicates) (integers) and posterior probabilities (20 million generations) acquired in Bayesian analysis (decimals) are displayed in red. The three investigated Blapstinina species are displayed in blue. Male genitalia (1–4) of selected species (dorsal views) are illustrated on the left. Topologies of collapsed clades (‘remaining Opatrini’ & ‘Dendarini + Pedininini + Playtonotini’) are fully consistent with those presented in Lumen et al. (2020)." figureDoi="http://doi.org/10.5281/zenodo.5964431" httpUri="https://zenodo.org/record/5964431/files/figure.png" pageId="3" pageNumber="522">Fig. 1</figureCitation>
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). However, the relationship uncovered by morphological comparison between
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<taxonomicName id="4C284D357D0E151B7D30CD81FA8EFF55" authorityName="Kamiński & Lumen & Kanda & Smith" authorityYear="2022" baseAuthorityName="Kamiński & Lumen & Kanda & Smith" baseAuthorityYear="1851" box="[1057,1314,187,213]" class="Insecta" family="Tenebrionidae" genus="Blapstinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="522" phylum="Arthropoda" rank="species" species="punctulatus">
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<emphasis id="B95CEAA47D0E151B7D30CD81FA8EFF55" box="[1057,1314,187,213]" italics="true" pageId="3" pageNumber="522">Blapstinus punctulatus</emphasis>
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</taxonomicName>
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and previously known species of
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<taxonomicName id="4C284D357D0E151B78B9CDE5FDE1FF79" authorityName="Marcuzzi" authorityYear="1954" box="[424,589,223,249]" class="Insecta" family="Tenebrionidae" genus="Austrocaribius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="522" phylum="Arthropoda" rank="genus">
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<emphasis id="B95CEAA47D0E151B78B9CDE5FDE1FF79" box="[424,589,223,249]" italics="true" pageId="3" pageNumber="522">Austrocaribius</emphasis>
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</taxonomicName>
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necessitates the following nomenclatural acts: Namely,
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<taxonomicName id="4C284D357D0E151B7DD8CDE5FF61FE9D" authorityName="Kamiński & Lumen & Kanda & Smith" authorityYear="2022" baseAuthorityName="Kamiński & Lumen & Kanda & Smith" baseAuthorityYear="1851" class="Insecta" family="Tenebrionidae" genus="Blapstinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="522" phylum="Arthropoda" rank="species" species="punctulatus">
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<emphasis id="B95CEAA47D0E151B7DD8CDE5FF61FE9D" italics="true" pageId="3" pageNumber="522">Blapstinus punctulatus</emphasis>
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</taxonomicName>
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is a synonym of
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<taxonomicName id="4C284D357D0E151B7881CC39FCEFFE9D" authority="Mulsant and Rey" authorityName="Mulsant and Rey" box="[400,835,259,285]" class="Insecta" family="Tenebrionidae" genus="Lodinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="522" phylum="Arthropoda" rank="species" species="nigroaeneus">
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<emphasis id="B95CEAA47D0E151B7881CC39FDD7FE9C" box="[400,635,259,285]" italics="true" pageId="3" pageNumber="522">Lodinus nigroaeneus</emphasis>
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Mulsant and Rey
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</taxonomicName>
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(see
|
||
<bibRefCitation id="EFB94B477D0E151B7A6ECC39FBDCFE9D" author="Bouchard, P. & Bousquet, Y. & Aalbu, R. L. & Alonso-Zarazaga, M. A. & Merkl, O. & Davies, A. E." box="[895,1136,259,285]" pageId="3" pageNumber="522" pagination="1 - 633" refId="ref7977" refString="Bouchard, P., Bousquet, Y., Aalbu, R. L., Alonso-Zarazaga, M. A., Merkl, O. & Davies, A. E. (2021) Review of genus group names in the family Tenebrionidae (Insecta, Coleoptera). ZooKeys, 1050, 1 - 633. https: // doi. org / 10.3897 / zookeys. 1050.64217" type="journal article" year="2021">
|
||
Bouchard
|
||
<emphasis id="B95CEAA47D0E151B7AE5CC39FB8AFE9D" box="[1012,1062,259,285]" italics="true" pageId="3" pageNumber="522">et al</emphasis>
|
||
. 2021
|
||
</bibRefCitation>
|
||
), which is the
|
||
<typeStatus id="549388147D0E151B7C05CC39FAE8FE9D" box="[1300,1348,259,285]" pageId="3" pageNumber="522">type</typeStatus>
|
||
species of the monotypic genus
|
||
<taxonomicName id="4C284D357D0E151B78B8CC1DFDAFFEC1" box="[425,515,295,321]" class="Insecta" family="Tenebrionidae" genus="Lodinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="522" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95CEAA47D0E151B78B8CC1DFDAFFEC1" box="[425,515,295,321]" italics="true" pageId="3" pageNumber="522">Lodinus</emphasis>
|
||
</taxonomicName>
|
||
that was synonymised with
|
||
<taxonomicName id="4C284D357D0E151B7A55CC1DFC16FEC1" authorityName="Sturm" authorityYear="1826" box="[836,954,295,321]" class="Insecta" family="Tenebrionidae" genus="Blapstinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="522" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95CEAA47D0E151B7A55CC1DFC16FEC1" box="[836,954,295,321]" italics="true" pageId="3" pageNumber="522">Blapstinus</emphasis>
|
||
</taxonomicName>
|
||
by Gemminger in
|
||
<bibRefCitation id="EFB94B477D0E151B7D80CC1DFF49FEE5" author="Gemminger, M. & Harold, E. von" pageId="3" pageNumber="522" pagination="1801 - 2179" refId="ref8577" refString="Gemminger, M. & Harold, E. von (1870) a. n. In: Catalogus coleopterorum hucusque descriptorum synonymicus et systematicus. Tom. VII. Tenebrionidae, Nilionidae, Pythidae, Melandryidae, Lagriidae, Pedilidae, Anthicidae, Pyrochroidae, Mordellidae, Rhipidophoridae, Cantharidae, Oedemeridae. E. H. Gummi, Monachium [Munchen], pp. 1801 - 2179." type="book chapter" year="1870">Gemminger and Harold (1870)</bibRefCitation>
|
||
. Since the name
|
||
<taxonomicName id="4C284D357D0E151B78B1CC71FE56FEE5" box="[416,506,331,357]" class="Insecta" family="Tenebrionidae" genus="Lodinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="522" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95CEAA47D0E151B78B1CC71FE56FEE5" box="[416,506,331,357]" italics="true" pageId="3" pageNumber="522">Lodinus</emphasis>
|
||
</taxonomicName>
|
||
predates
|
||
<taxonomicName id="4C284D357D0E151B7B75CC71FCA5FEE5" authorityName="Marcuzzi" authorityYear="1954" box="[612,777,331,357]" class="Insecta" family="Tenebrionidae" genus="Austrocaribius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="522" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95CEAA47D0E151B7B75CC71FCA5FEE5" box="[612,777,331,357]" italics="true" pageId="3" pageNumber="522">Austrocaribius</emphasis>
|
||
</taxonomicName>
|
||
by 95 years, the change in status of
|
||
<taxonomicName id="4C284D357D0E151B7D8CCC71FA31FEE5" authorityName="Kamiński & Lumen & Kanda & Smith" authorityYear="2022" baseAuthorityName="Kamiński & Lumen & Kanda & Smith" baseAuthorityYear="1851" box="[1181,1437,331,357]" class="Insecta" family="Tenebrionidae" genus="Blapstinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="522" phylum="Arthropoda" rank="species" species="punctulatus">
|
||
<emphasis id="B95CEAA47D0E151B7D8CCC71FA31FEE5" box="[1181,1437,331,357]" italics="true" pageId="3" pageNumber="522">Blapstinus punctulatus</emphasis>
|
||
</taxonomicName>
|
||
therefore requires introduction of the following synonymy and combinations:
|
||
<taxonomicName id="4C284D357D0E151B7AEBCC55FAB4FE09" authority="Mulsant and Rey" authorityName="Mulsant and Rey" box="[1018,1304,367,393]" class="Insecta" family="Tenebrionidae" genus="Lodinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="522" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95CEAA47D0E151B7AEBCC55FBF8FE09" box="[1018,1108,367,393]" italics="true" pageId="3" pageNumber="522">Lodinus</emphasis>
|
||
Mulsant and Rey
|
||
</taxonomicName>
|
||
<emphasis id="B95CEAA47D0E151B7C0ECC55FA30FE08" bold="true" box="[1311,1436,366,393]" pageId="3" pageNumber="522">stat. restit.</emphasis>
|
||
(=
|
||
<taxonomicName id="4C284D357D0E151B79A1CCA9FE6BFE2D" authority="Marcuzzi" authorityName="Marcuzzi" authorityYear="1954" box="[176,455,403,429]" class="Insecta" family="Tenebrionidae" genus="Austrocaribius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="522" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95CEAA47D0E151B79A1CCA9FEF9FE2D" box="[176,341,403,429]" italics="true" pageId="3" pageNumber="522">Austrocaribius</emphasis>
|
||
Marcuzzi
|
||
</taxonomicName>
|
||
<emphasis id="B95CEAA47D0E151B78DFCCA9FD9AFE2D" bold="true" box="[462,566,403,429]" pageId="3" pageNumber="522">
|
||
<taxonomicNameLabel id="A26F57DF7D0E151B78DFCCA9FD9AFE2D" box="[462,566,403,429]" pageId="3" pageNumber="522">syn. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
),
|
||
<taxonomicName id="4C284D357D0E151B7B5CCCA9FC3CFE2E" authority="(Marcuzzi)" baseAuthorityName="Marcuzzi" box="[589,912,403,430]" class="Insecta" family="Tenebrionidae" genus="Lodinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="522" phylum="Arthropoda" rank="species" species="araguae" status="comb. et stat. nov.">
|
||
<emphasis id="B95CEAA47D0E151B7B5CCCA9FCA0FE2C" box="[589,780,403,429]" italics="true" pageId="3" pageNumber="522">Lodinus araguae</emphasis>
|
||
(Marcuzzi)
|
||
</taxonomicName>
|
||
<emphasis id="B95CEAA47D0E151B7A86CCA9FBC2FE2D" bold="true" box="[919,1134,403,429]" pageId="3" pageNumber="522">
|
||
<taxonomicNameLabel id="A26F57DF7D0E151B7A86CCA9FBC2FE2D" box="[919,1134,403,429]" pageId="3" pageNumber="522">comb. et stat. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
[transferred from
|
||
<taxonomicName id="4C284D357D0E151B7C2CCCA9FF4AFE51" authorityName="Marcuzzi" authorityYear="1954" class="Insecta" family="Tenebrionidae" genus="Austrocaribius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="522" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95CEAA47D0E151B7C2CCCA9FF4AFE51" italics="true" pageId="3" pageNumber="522">Austrocaribius</emphasis>
|
||
</taxonomicName>
|
||
],
|
||
<taxonomicName id="4C284D357D0E151B79EECC8DFE0CFE51" authorityName="Kamiński & Lumen & Kanda & Smith" authorityYear="2022" baseAuthorityName="Kamiński & Lumen & Kanda & Smith" baseAuthorityYear="1851" box="[255,416,439,465]" class="Insecta" family="Tenebrionidae" genus="Blapstinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="522" phylum="Arthropoda" rank="species" species="punctulatus" status="comb. nov.">
|
||
<emphasis id="B95CEAA47D0E151B79EECC8DFE0CFE51" box="[255,416,439,465]" italics="true" pageId="3" pageNumber="522">L. punctulatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95CEAA47D0E151B78B9CC8DFD85FE51" bold="true" box="[424,553,439,465]" pageId="3" pageNumber="522">
|
||
<taxonomicNameLabel id="A26F57DF7D0E151B78B9CC8DFD85FE51" box="[424,553,439,465]" pageId="3" pageNumber="522" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(=
|
||
<taxonomicName id="4C284D357D0E151B7B5BCC8DFC99FE50" box="[586,821,439,465]" class="Insecta" family="Tenebrionidae" genus="Lodinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="522" phylum="Arthropoda" rank="species" species="nigroaeneus">
|
||
<emphasis id="B95CEAA47D0E151B7B5BCC8DFC99FE50" box="[586,821,439,465]" italics="true" pageId="3" pageNumber="522">Lodinus nigroaeneus</emphasis>
|
||
</taxonomicName>
|
||
),
|
||
<taxonomicName id="4C284D357D0E151B7A5FCC8DFB25FE52" authority="(Marcuzzi)" baseAuthorityName="Marcuzzi" box="[846,1161,439,466]" class="Insecta" family="Tenebrionidae" genus="Lodinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="522" phylum="Arthropoda" rank="species" species="venezuelensis" status="comb. nov.">
|
||
<emphasis id="B95CEAA47D0E151B7A5FCC8DFBA8FE51" box="[846,1028,439,465]" italics="true" pageId="3" pageNumber="522">L. venezuelensis</emphasis>
|
||
(Marcuzzi)
|
||
</taxonomicName>
|
||
<emphasis id="B95CEAA47D0E151B7D80CC8DFABEFE51" bold="true" box="[1169,1298,439,465]" pageId="3" pageNumber="522">
|
||
<taxonomicNameLabel id="A26F57DF7D0E151B7D80CC8DFABEFE51" box="[1169,1298,439,465]" pageId="3" pageNumber="522" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
[transferred from
|
||
<taxonomicName id="4C284D357D0E151B79C2CCE0FED3FE75" authorityName="Marcuzzi" authorityYear="1954" box="[211,383,474,501]" class="Insecta" family="Tenebrionidae" genus="Austrocaribius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="524" phylum="Arthropoda" rank="genus">Austrocaribius</taxonomicName>
|
||
].
|
||
</paragraph>
|
||
<caption id="DF57663E7D0E151B7986CBB1FB6CF8B3" ID-DOI="http://doi.org/10.5281/zenodo.5964431" ID-Zenodo-Dep="5964431" httpUri="https://zenodo.org/record/5964431/files/figure.png" pageId="3" pageNumber="522" startId="3.[151,250,1675,1699]" targetBox="[151,1419,525,1640]" targetPageId="3">
|
||
<paragraph id="8B9736B67D0E151B7986CBB1FB6CF8B3" blockId="3.[151,1437,1675,1843]" pageId="3" pageNumber="522">
|
||
<emphasis id="B95CEAA47D0E151B7986CBB1FEB9F923" bold="true" box="[151,277,1675,1699]" pageId="3" pageNumber="522">FIGURE 1.</emphasis>
|
||
Phylogeny of subtribe
|
||
<taxonomicName id="4C284D357D0E151B7B1ACBB1FD2EF923" baseAuthorityName="Mulsant & Rey" baseAuthorityYear="1853" box="[523,642,1675,1699]" pageId="3" pageNumber="522" rank="subTribe" subTribe="Blapstinina">Blapstinina</taxonomicName>
|
||
. Figured topology was recovered in maximum likelihood analysis using IQ-Tree. Ultrafast bootstrap values (1,000 replicates) (integers) and posterior probabilities (20 million generations) acquired in Bayesian analysis (decimals) are displayed in red. The three investigated
|
||
<taxonomicName id="4C284D357D0E151B7A92CBE9FC5AF96B" baseAuthorityName="Mulsant & Rey" baseAuthorityYear="1853" box="[899,1014,1747,1771]" pageId="3" pageNumber="522" rank="subTribe" subTribe="Blapstinina">Blapstinina</taxonomicName>
|
||
species are displayed in blue. Male genitalia (1–4) of selected species (dorsal views) are illustrated on the left. Topologies of collapsed clades (‘remaining Opatrini’ & ‘Dendarini + Pedininini + Playtonotini’) are fully consistent with those presented in
|
||
<bibRefCitation id="EFB94B477D0E151B7AFFCA21FB17F8B3" author="Lumen, R. & Kanda, K. & Iwan, D. & Smith, A. D. & Kaminski, M." box="[1006,1211,1819,1843]" pageId="3" pageNumber="522" pagination="337 - 348" refId="ref9448" refString="Lumen, R., Kanda, K., Iwan, D., Smith, A. D. & Kaminski, M. (2020) Molecular insights into the phylogeny of Blapstinina (Coleoptera: Tenebrionidae: Opatrini). Systematic Entomology, 45, 337 - 348. https: // doi. org / 10.1111 / syen. 12398" type="journal article" year="2020">
|
||
Lumen
|
||
<emphasis id="B95CEAA47D0E151B7D2DCA26FBC4F8B3" box="[1084,1128,1819,1843]" italics="true" pageId="3" pageNumber="522">et al</emphasis>
|
||
. (2020)
|
||
</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9736B67D0E151C79D6CA5BFA90FF79" blockId="3.[151,1437,1889,2024]" lastBlockId="4.[151,1436,151,249]" lastPageId="4" lastPageNumber="523" pageId="3" pageNumber="522">
|
||
The unexpected phylogenetic position of
|
||
<taxonomicName id="4C284D357D0E151B7B84CA58FC99F8FB" authorityName="Kamiński & Lumen & Kanda & Smith" authorityYear="2022" baseAuthorityName="Kamiński & Lumen & Kanda & Smith" baseAuthorityYear="1851" box="[661,821,1889,1915]" class="Insecta" family="Tenebrionidae" genus="Blapstinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="522" phylum="Arthropoda" rank="species" species="punctulatus">
|
||
<emphasis id="B95CEAA47D0E151B7B84CA58FC99F8FB" box="[661,821,1889,1915]" italics="true" pageId="3" pageNumber="522">L. punctulatus</emphasis>
|
||
</taxonomicName>
|
||
highlights an urgent need for additional morphological investigation of South American
|
||
<taxonomicName id="4C284D357D0E151B7B1BCABFFD20F81F" baseAuthorityName="Mulsant & Rey" baseAuthorityYear="1853" box="[522,652,1925,1951]" pageId="3" pageNumber="522" rank="subTribe" subTribe="Blapstinina">Blapstinina</taxonomicName>
|
||
, especially those superficially resembling
|
||
<emphasis id="B95CEAA47D0E151B7D79CABFFB4AF81F" box="[1128,1254,1925,1951]" italics="true" pageId="3" pageNumber="522">
|
||
<taxonomicName id="4C284D357D0E151B7D79CABFFB4EF81F" authorityName="Sturm" authorityYear="1826" box="[1128,1250,1925,1951]" class="Insecta" family="Tenebrionidae" genus="Blapstinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="522" phylum="Arthropoda" rank="genus">Blapstinus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Taxonomic acts in this paper greatly extend the range and change the status of a previously geographically restricted genus
|
||
<taxonomicName id="4C284D357D0E151B7C2ACA93FA39F843" box="[1339,1429,1961,1987]" class="Insecta" family="Tenebrionidae" genus="Lodinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="522" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95CEAA47D0E151B7C2ACA93FA39F843" box="[1339,1429,1961,1987]" italics="true" pageId="3" pageNumber="522">Lodinus</emphasis>
|
||
</taxonomicName>
|
||
. Furthermore, there are 75 species and subspecies of
|
||
<emphasis id="B95CEAA47D0E151B7BC8CAF7FCEDF867" box="[729,833,1997,2023]" italics="true" pageId="3" pageNumber="522">Blastinus</emphasis>
|
||
recorded North of Mexico (
|
||
<bibRefCitation id="EFB94B477D0E151B7D66CAF7FACEF867" author="Bousquet, Y. & Thomas, D. B. & Bouchard, P. & Smith, A. D. & Aalbu, R. L. & Johnston, M. A. & Steiner, W." box="[1143,1378,1997,2024]" pageId="3" pageNumber="522" pagination="1 - 455" refId="ref8050" refString="Bousquet, Y., Thomas, D. B., Bouchard, P., Smith, A. D., Aalbu, R. L., Johnston, M. A. & Steiner, W. (2018) Catalogue of Tene- brionidae (Coleoptera) of North America. Zookeys, 728, 1 - 455. https: // doi. org / 10.3897 / zookeys. 728.20602" type="journal article" year="2018">
|
||
Bousquet
|
||
<emphasis id="B95CEAA47D0E151B7DF8CAF4FAB5F867" box="[1257,1305,1997,2023]" italics="true" pageId="3" pageNumber="522">et al</emphasis>
|
||
. 2018
|
||
</bibRefCitation>
|
||
), and no official records available for
|
||
<collectingCountry id="F33F76267D09151C78E5CDADFDCDFF31" box="[500,609,151,177]" name="Argentina" pageId="4" pageNumber="523">Argentina</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFB94B477D09151C7B7FCDADFCB6FF31" author="Fairmaire, L." box="[622,794,151,177]" pageId="4" pageNumber="523" pagination="507 - 516" refId="ref8516" refString="Fairmaire, L. (1883) Descriptions de Coleopteres de la Republique-Argentine. Annales de la Societe entomologique de France, 6, 507 - 516." type="journal article" year="1883">Fairmaire 1883</bibRefCitation>
|
||
, present paper, Gustavo Flores pers. comm.). Additionally, as a result of investigating taxon records for this study, it was revealed to the authors that the
|
||
<typeStatus id="549388147D09151C7DB3CD81FAAEFF55" box="[1186,1282,187,213]" pageId="4" pageNumber="523" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C284D357D09151C7C39CD81FE39FF79" authority="Fairmaire" authorityName="Fairmaire" class="Insecta" family="Tenebrionidae" genus="Blapstinus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="523" phylum="Arthropoda" rank="species" species="metallescens">
|
||
<emphasis id="B95CEAA47D09151C7C39CD81FE8FFF79" italics="true" pageId="4" pageNumber="523">Blapstinus metallescens</emphasis>
|
||
Fairmaire
|
||
</taxonomicName>
|
||
does not represent
|
||
<taxonomicName id="4C284D357D09151C7B7BCDE5FCABFF79" authorityName="Latreille" authorityYear="1802" box="[618,775,223,249]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="523" phylum="Arthropoda" rank="family">Tenebrionidae</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFB94B477D09151C7A05CDE5FC40FF79" author="Cabrera, N. & Fernandez, L. A. & Flores, G. E. & Lattari, M." box="[788,1004,223,249]" pageId="4" pageNumber="523" pagination="107 - 123" refId="ref8126" refString="Cabrera, N., Fernandez, L. A., Flores, G. E. & Lattari, M. (2010) Catalog of the types of Tenebrionidae and Perirnylopidae (Insecta, Coleoptera) deposited at Museo de La Plata (Argentina). Transactions of the American Entomological Society, 136, 107 - 123." type="journal article" year="2010">
|
||
Cabrera
|
||
<emphasis id="B95CEAA47D09151C7A65CDE5FC08FF79" box="[884,932,223,249]" italics="true" pageId="4" pageNumber="523">et al</emphasis>
|
||
. 2010
|
||
</bibRefCitation>
|
||
, Gustavo Flores pers. comm.).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |