treatments-xml/data/03/F5/FA/03F5FA39BC63FFC1B27AFCCC307BFD10.xml

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<mods:title>A revision of Canarium L. (Burseraceae) in Madagascar</mods:title>
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<taxonomicName authority="Daly, Raharim. &amp; Federman" authorityName="Daly, Raharim. &amp; Federman" authorityYear="2015" box="[813,1365,852,879]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="59" pageNumber="336" phylum="Tracheophyta" rank="species" species="scholasticum" status="sp. nov.">
<emphasis bold="true" box="[813,1057,852,879]" italics="true" pageId="59" pageNumber="336">Canarium scholasticum</emphasis>
Daly, Raharim. &amp; Federman
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, 
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(
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; 
<figureCitation box="[1166,1195,885,911]" captionStart="FIG" captionStartId="60.[132,143,1894,1912]" captionTargetBox="[205,1386,214,1868]" captionTargetId="figure-19@60.[199,1390,210,1873]" captionTargetPageId="60" captionText="FIG. 28. — Canarium scholasticum Daly, Raharim. &amp; Federman, sp. nov.: A, flowering branchlet with detail of leaflet (A’); B, detail of shoot apex with stipules; C, apical view of stipules; D, staminate bud with bracteoles (right) and with part of calyx and two petals removed (left); E, longisection of ovariodisk and half of stamens; F, portion of pistillate inflorescence;G, pistillate flower with bracts and bracteoles; H, pistillate flower with perianth removed, plus apical view of stigma and abaxial view of pubescent staminode anther (right); I, portion of infructescence; A, F-H, Daly et al. 13075 (NY and field photos); B, Daly et al. 13015 (field photo by D. Daly); C, Daly et al. 12968 (field photo by D. Daly); D, E, from Service Forestier 2088 (NY); I, Malcomber 1046 (NY).Scale bars: A, I, 2 cm; B, C, 5 mm; D, E, G, H, 3 mm; F, 1 cm; A’, not to scale." figureDoi="http://doi.org/10.5281/zenodo.5208957" httpUri="https://zenodo.org/record/5208957/files/figure.png" pageId="59" pageNumber="336">28</figureCitation>
)
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Small to very large trees, leaves 3-4(6)-jugate, terminal leaflet usually obovate, leaflet apex usually abruptly and broadly short-acuminate with the acumen 
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long; distinguished from
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<taxonomicName authority="Daly, Raharim. &amp; Federman" authorityName="Daly, Raharim. &amp; Federman" authorityYear="2015" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="59" pageNumber="336" phylum="Tracheophyta" rank="species" species="lamianum" status="sp. nov.">
C. lamianum 
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, 
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by the bark finely fissured or smooth (vs deeply fissured in
</emphasis>
<taxonomicName authority="Daly &amp; Raharimampionona &amp; Federman, 2015" authorityName="Daly &amp; Raharimampionona &amp; Federman" authorityYear="2015" box="[1088,1218,1052,1076]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="59" pageNumber="336" phylum="Tracheophyta" rank="species" species="lamianum" status="sp. nov.">C. lamianum</taxonomicName>
, 
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<taxonomicNameLabel box="[1232,1308,1053,1076]" pageId="59" pageNumber="336" rank="species">sp. nov.</taxonomicNameLabel>
), lateral leaflets (oblong-)ovate (vs variously shaped but not ovate), lateral leaflet base cordate or less often truncate, very rarely acute (vs obtuse, truncate, or acute), secondary vein angle decreasing toward the base (vs increasing or uniform), on staminate inflorescences the secondary branch endings not congested (vs often congested).
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TYPUS. — 
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, on mild slope 
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N of reserve base, 
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elev., 
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13075
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(holo-, 
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!; iso-, G!, 
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!, 
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<collectorName box="[1132,1273,1421,1445]" pageId="59" pageNumber="336">M. Callmander</collectorName>
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429
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(G, 
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, P, 
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, 
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S.T. 
<collectorName box="[856,959,1527,1551]" pageId="59" pageNumber="336">Malcomber</collectorName>
1046
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(
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, 
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)
</materialsCitation>
. 
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P. 
<collectorName box="[834,963,1580,1604]" pageId="59" pageNumber="336">Antilahimena</collectorName>
2051
</emphasis>
(
<collectionCode box="[1037,1083,1580,1604]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="59" pageNumber="336" type="Herbarium">MO</collectionCode>
)
</materialsCitation>
; 
<materialsCitation collectingDate="2007-05-21" collectionCode="MO, TAN, TEF, US" collectorName="P. Antilahimena &amp; F. Edmond &amp; Tojoniaina" country="Madagascar" county="Alaotra-Mangoro" elevation="1221" latitude="-18.868334" location="Andasibe" longLatPrecision="21" longitude="48.303886" municipality="Moramanga" pageId="59" pageNumber="336" specimenCount="1" typeStatus="paratype">
<collectingCounty box="[1103,1275,1580,1604]" pageId="59" pageNumber="336">Alaotra-Mangoro</collectingCounty>
, 
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, 
<location LSID="urn:lsid:plazi:treatment:03F5FA39BC63FFC1B27AFCCC307BFD10:8E831DF4BC63FFC5B4DDF9B53267F9FE" country="Madagascar" county="Alaotra-Mangoro" latitude="-18.868334" longLatPrecision="21" longitude="48.303886" municipality="Moramanga" name="Andasibe" pageId="59" pageNumber="336">Andasibe</location>
, 
<location LSID="urn:lsid:plazi:treatment:03F5FA39BC63FFC1B27AFCCC307BFD10:8E831DF4BC63FFC5B22FF9DE32E5F9FE" box="[888,1007,1607,1631]" country="Madagascar" county="Alaotra-Mangoro" latitude="-18.868334" longLatPrecision="21" longitude="48.303886" municipality="Moramanga" name="Menalamba" pageId="59" pageNumber="336">Menalamba</location>
, 
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, 
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<elevation box="[1370,1451,1607,1631]" metricMagnitude="3" metricUnit="m" metricValue="1.2209999999999999" pageId="59" pageNumber="336" unit="m" value="1221.0">1221 m</elevation>
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, 
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, 
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, 
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<collectingDate box="[1061,1165,1633,1657]" pageId="59" pageNumber="336" value="2007-05-21">21.V.2007</collectingDate>
</date>
, 
<emphasis italics="true" pageId="59" pageNumber="336">
<collectorName box="[1170,1322,1633,1657]" pageId="59" pageNumber="336">P. Antilahimena</collectorName>
, 
<collectorName box="[1330,1429,1633,1657]" pageId="59" pageNumber="336">F. Edmond</collectorName>
&amp; 
<collectorName box="[811,909,1660,1683]" pageId="59" pageNumber="336">Tojoniaina</collectorName>
5540
</emphasis>
(
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, P, 
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, 
<collectionCode box="[1104,1151,1660,1684]" country="Madagascar" lsid="urn:lsid:biocol.org:col:12774" name="Centre National de la Recherche Appliquée au Developement Rural" pageId="59" pageNumber="336" type="Herbarium">TEF</collectionCode>
); 
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, 
<location LSID="urn:lsid:plazi:treatment:03F5FA39BC63FFC1B27AFCCC307BFD10:8E831DF4BC63FFC5B460F9E534A6F935" box="[1335,1452,1660,1684]" country="Madagascar" county="Alaotra-Mangoro" latitude="-18.868334" longLatPrecision="21" longitude="48.303886" municipality="Moramanga" name="Moramanga" pageId="59" pageNumber="336">Moramanga</location>
, 
<location LSID="urn:lsid:plazi:treatment:03F5FA39BC63FFC1B27AFCCC307BFD10:8E831DF4BC63FFC5B27CF90F32ACF90F" box="[811,934,1686,1710]" country="Madagascar" county="Alaotra-Mangoro" latitude="-18.868334" longLatPrecision="21" longitude="48.303886" municipality="Moramanga" name="Ambohibary" pageId="59" pageNumber="336">Ambohibary</location>
, 
<location LSID="urn:lsid:plazi:treatment:03F5FA39BC63FFC1B27AFCCC307BFD10:8E831DF4BC63FFC5B2F9F90F352EF90F" box="[942,1060,1686,1710]" country="Madagascar" county="Alaotra-Mangoro" latitude="-18.868334" longLatPrecision="21" longitude="48.303886" municipality="Moramanga" name="Ampitambe" pageId="59" pageNumber="336">Ampitambe</location>
, 
<location LSID="urn:lsid:plazi:treatment:03F5FA39BC63FFC1B27AFCCC307BFD10:8E831DF4BC63FFC5B57BF90F359BF90F" box="[1068,1169,1686,1710]" country="Madagascar" county="Alaotra-Mangoro" latitude="-18.868334" longLatPrecision="21" longitude="48.303886" municipality="Moramanga" name="Ambatovy" pageId="59" pageNumber="336">Ambatovy</location>
, 
<location LSID="urn:lsid:plazi:treatment:03F5FA39BC63FFC1B27AFCCC307BFD10:8E831DF4BC63FFC5B5CEF90F3459F90F" box="[1177,1363,1686,1710]" country="Madagascar" county="Alaotra-Mangoro" latitude="-18.868334" longLatPrecision="21" longitude="48.303886" municipality="Moramanga" name="Andranovery forest" pageId="59" pageNumber="336">Andranovery forest</location>
, 
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<elevation box="[1372,1449,1686,1710]" metricMagnitude="3" metricUnit="m" metricValue="1.028" pageId="59" pageNumber="336" unit="m" value="1028.0">1028 m</elevation>
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, 
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, 
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, 
<emphasis box="[1055,1304,1712,1737]" italics="true" pageId="59" pageNumber="336">
<collectorName box="[1055,1201,1712,1736]" pageId="59" pageNumber="336">P.Antilahimena</collectorName>
et al. 7098
</emphasis>
(
<collectionCode box="[1315,1349,1713,1737]" country="South Africa" name="University of Stellenbosch" pageId="59" pageNumber="336">US</collectionCode>
)
</materialsCitation>
; 
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<collectingCounty pageId="59" pageNumber="336">Analanjirofo</collectingCounty>
, 
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, 
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<location LSID="urn:lsid:plazi:treatment:03F5FA39BC63FFC1B27AFCCC307BFD10:8E831DF4BC63FFC5B47BF952326AF95C" country="Madagascar" county="Analanjirofo" latitude="-17.923056" longLatPrecision="21" longitude="49.19917" municipality="Befandriana Nord" name="Ampandisana River" pageId="59" pageNumber="336">Ampandisana River</location>
, 
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<elevation box="[874,940,1765,1789]" metricMagnitude="2" metricUnit="m" metricValue="1.49" pageId="59" pageNumber="336" unit="m" value="149.0">149 m</elevation>
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, 
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, 
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, 
<date box="[1201,1316,1765,1789]" pageId="59" pageNumber="336" value="2007-04-30">
<collectingDate box="[1201,1316,1765,1789]" pageId="59" pageNumber="336" value="2007-04-30">30.IV.2007</collectingDate>
</date>
, 
<emphasis italics="true" pageId="59" pageNumber="336">
<collectorName box="[1326,1429,1765,1789]" pageId="59" pageNumber="336">R. Bernard</collectorName>
&amp; 
<collectorName box="[812,952,1792,1816]" pageId="59" pageNumber="336">C. Birkinshaw</collectorName>
502
</emphasis>
(
<collectionCode box="[1016,1063,1792,1816]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="59" pageNumber="336" type="Herbarium">MO</collectionCode>
, P, 
<collectionCode box="[1097,1152,1792,1816]" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13842" lsid="urn:lsid:biocol.org:col:13842" name="Parc de Tsimbazaza" pageId="59" pageNumber="336" type="Herbarium">TAN</collectionCode>
); 
<locationDeviation box="[1173,1391,1792,1816]" location="Mandraka" pageId="59" pageNumber="336">
<quantity box="[1173,1226,1792,1816]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="59" pageNumber="336" unit="km" value="3.0">3 km</quantity>
E of Mandraka
</locationDeviation>
, along Antananarivo-Toamasina highway, 
<quantity box="[1176,1260,1818,1842]" metricMagnitude="3" metricUnit="m" metricValue="1.09" pageId="59" pageNumber="336" unit="m" value="1090.0">
<elevation box="[1176,1260,1818,1842]" metricMagnitude="3" metricUnit="m" metricValue="1.09" pageId="59" pageNumber="336" unit="m" value="1090.0">1090 m</elevation>
</quantity>
, 
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, 
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, 
<date box="[814,913,1845,1869]" pageId="59" pageNumber="336" value="1993-01-27">
<collectingDate box="[814,913,1845,1869]" pageId="59" pageNumber="336" value="1993-01-27">27.I.1993</collectingDate>
</date>
, 
<emphasis box="[923,1455,1845,1869]" italics="true" pageId="59" pageNumber="336">
<collectorName box="[923,1045,1845,1869]" pageId="59" pageNumber="336">G. E. Schatz</collectorName>
, 
<collectorName box="[1053,1183,1845,1868]" pageId="59" pageNumber="336">W. D. Stevens</collectorName>
&amp; 
<collectorName box="[1216,1398,1845,1869]" pageId="59" pageNumber="336">J. P. Rakotomazaza</collectorName>
3425
</emphasis>
(
<collectionCode box="[820,866,1871,1895]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="59" pageNumber="336" type="Herbarium">MO</collectionCode>
, 
<collectionCode box="[875,910,1872,1895]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="59" pageNumber="336" type="Herbarium">NY</collectionCode>
, P, 
<collectionCode box="[936,990,1871,1895]" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13842" lsid="urn:lsid:biocol.org:col:13842" name="Parc de Tsimbazaza" pageId="59" pageNumber="336" type="Herbarium">TAN</collectionCode>
); Atsinanana, Réserve Naturelle Intégrale Betampona, 
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<elevation box="[874,986,1898,1922]" metricMagnitude="2" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="59" pageNumber="336" unit="m" value="500.0" valueMax="600.0" valueMin="400.0">400-600 m</elevation>
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, 
<date box="[1248,1355,1898,1922]" pageId="59" pageNumber="336" value="2008-02-11">
<collectingDate box="[1248,1355,1898,1922]" pageId="59" pageNumber="336" value="2008-02-11">11.II.2008</collectingDate>
</date>
, 
<emphasis italics="true" pageId="59" pageNumber="336">
<collectorName box="[1366,1452,1898,1921]" pageId="59" pageNumber="336">T. Treuer</collectorName>
, 
<collectorName box="[811,929,1924,1948]" pageId="59" pageNumber="336">S. Federman</collectorName>
&amp; 
<collectorName box="[961,1052,1924,1948]" pageId="59" pageNumber="336">J. Sylvain</collectorName>
4
</emphasis>
(
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)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="61" lastPageNumber="338" pageId="59" pageNumber="336" type="distribution">
<paragraph blockId="59.[813,1457,1976,2027]" pageId="59" pageNumber="336">
DISTRIBUTION AND ECOLOGY. — 
<taxonomicName authority="Daly &amp; Raharimampionona &amp; Federman, 2015" authorityName="Daly &amp; Raharimampionona &amp; Federman" authorityYear="2015" box="[1147,1367,1976,2000]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="59" pageNumber="336" phylum="Tracheophyta" rank="species" species="scholasticum" status="sp. nov.">
<emphasis box="[1147,1367,1976,2000]" italics="true" pageId="59" pageNumber="336">Canarium scholasticum</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel box="[1380,1456,1977,2000]" pageId="59" pageNumber="336" rank="species">sp. nov.</taxonomicNameLabel>
has one of the broadest distributions of the genus in 
<collectingCountry box="[1337,1454,2003,2027]" name="Madagascar" pageId="59" pageNumber="336">Madagascar</collectingCountry>
.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.5208957" ID-Zenodo-Dep="5208957" httpUri="https://zenodo.org/record/5208957/files/figure.png" pageId="60" pageNumber="337" startId="60.[132,143,1894,1912]" targetBox="[205,1386,214,1868]" targetPageId="60">
<paragraph blockId="60.[132,1457,1894,2025]" pageId="60" pageNumber="337">
FIG. 28. — 
<taxonomicName authority="Daly, Raharim. &amp; Federman" authorityName="Daly, Raharim. &amp; Federman" authorityYear="2015" box="[226,660,1894,1912]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="60" pageNumber="337" phylum="Tracheophyta" rank="species" species="scholasticum" status="sp. nov.">
<emphasis box="[226,421,1894,1912]" italics="true" pageId="60" pageNumber="337">Canarium scholasticum</emphasis>
Daly, Raharim. &amp; Federman
</taxonomicName>
, 
<taxonomicNameLabel box="[669,739,1894,1912]" pageId="60" pageNumber="337" rank="species">sp. nov.</taxonomicNameLabel>
: 
<emphasis bold="true" box="[746,759,1894,1912]" pageId="60" pageNumber="337">A</emphasis>
, flowering branchlet with detail of leaflet (
<emphasis bold="true" box="[1113,1130,1894,1912]" pageId="60" pageNumber="337">A’</emphasis>
); 
<emphasis bold="true" box="[1146,1159,1894,1912]" pageId="60" pageNumber="337">B</emphasis>
, detail of shoot apex with stipules; 
<emphasis bold="true" box="[132,146,1916,1934]" pageId="60" pageNumber="337">C</emphasis>
, apical view of stipules; 
<emphasis bold="true" box="[351,365,1917,1935]" pageId="60" pageNumber="337">D</emphasis>
, staminate bud with bracteoles (right) and with part of calyx and two petals removed (left); 
<emphasis bold="true" box="[1125,1137,1917,1935]" pageId="60" pageNumber="337">E</emphasis>
, longisection of ovariodisk and half of stamens; 
<emphasis bold="true" box="[213,224,1939,1957]" pageId="60" pageNumber="337">F</emphasis>
, portion of pistillate inflorescence; 
<emphasis bold="true" box="[512,526,1939,1957]" pageId="60" pageNumber="337">G</emphasis>
, pistillate flower with bracts and bracteoles; 
<emphasis bold="true" box="[895,909,1939,1957]" pageId="60" pageNumber="337">H</emphasis>
, pistillate flower with perianth removed, plus apical view of stigma and abaxial view of pubescent staminode anther (right); 
<emphasis bold="true" box="[605,611,1962,1980]" pageId="60" pageNumber="337">I</emphasis>
, portion of infructescence; 
<emphasis bold="true" box="[842,855,1962,1980]" pageId="60" pageNumber="337">A</emphasis>
, 
<emphasis bold="true" box="[866,899,1962,1980]" pageId="60" pageNumber="337">F -H</emphasis>
, 
<emphasis box="[909,1050,1962,1980]" italics="true" pageId="60" pageNumber="337">Daly et al. 13075</emphasis>
(NY and field photos); 
<emphasis bold="true" box="[1245,1258,1962,1980]" pageId="60" pageNumber="337">B</emphasis>
, 
<emphasis box="[1269,1410,1962,1980]" italics="true" pageId="60" pageNumber="337">Daly et al. 13015</emphasis>
(field photo by D. Daly); 
<emphasis bold="true" box="[285,299,1984,2002]" pageId="60" pageNumber="337">C</emphasis>
, 
<emphasis box="[308,445,1985,2003]" italics="true" pageId="60" pageNumber="337">Daly et al. 12968</emphasis>
(field photo by D. Daly); 
<emphasis bold="true" box="[646,660,1985,2003]" pageId="60" pageNumber="337">D</emphasis>
, 
<emphasis bold="true" box="[669,681,1985,2003]" pageId="60" pageNumber="337">E</emphasis>
, from 
<emphasis box="[734,919,1984,2003]" italics="true" pageId="60" pageNumber="337">Service Forestier 2088</emphasis>
(NY); 
<emphasis bold="true" box="[968,974,1985,2003]" pageId="60" pageNumber="337">I</emphasis>
, 
<emphasis box="[983,1124,1985,2003]" italics="true" pageId="60" pageNumber="337">Malcomber 1046</emphasis>
(NY).Scale bars: A, I, 2 cm; B, C, 5 mm; D, E, G, H, 3 mm; F, 1 cm; A’, not to scale.
</paragraph>
</caption>
<paragraph blockId="61.[131,777,215,546]" pageId="61" pageNumber="338">
It ranges from Ranomafana National Park in the south, to Tsiroanomandidy in the west and Analamazaotra National Park in the center, to Maroantsetra, Vohémar and Sambava in the North; it is also broadly distributed in the Sambirano-Ambanja region. It is usually found in montane forests between (325) 
<quantity box="[651,777,321,345]" metricMagnitude="3" metricUnit="m" metricValue="1.2" metricValueMax="1.6" metricValueMin="0.8" pageId="61" pageNumber="338" unit="m" value="1200.0" valueMax="1600.0" valueMin="800.0">800-1600 m</quantity>
elevation. Flowering Jan.-Mar., fruiting Nov.-Apr.
</paragraph>
<paragraph blockId="61.[131,777,215,546]" pageId="61" pageNumber="338">COMMON NAMES. — Ramy blanc, ramy mena, ramy, tsiramiramy, aramy.</paragraph>
</subSubSection>
<subSubSection pageId="61" pageNumber="338" type="etymology">
<paragraph blockId="61.[131,777,215,546]" pageId="61" pageNumber="338">ETYMOLOGY. — The specific epithet refers to the occurrence of the species near places of study, the Cabine de Recherche and the Vato camp used by primatologists in Ranomafana National Park.</paragraph>
</subSubSection>
<subSubSection lastPageId="63" lastPageNumber="340" pageId="61" pageNumber="338" type="description">
<paragraph blockId="61.[130,777,598,2028]" box="[132,277,598,624]" pageId="61" pageNumber="338">DESCRIPTION</paragraph>
<paragraph blockId="61.[130,777,598,2028]" lastBlockId="61.[811,1457,215,1645]" pageId="61" pageNumber="338">
Trees, reproductive size 
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, to 
<quantity box="[518,601,630,656]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="61" pageNumber="338" unit="cm" value="300.0">300 cm</quantity>
diam, with low plank buttresses. Outer bark gray, thick, deeply fissured, lenticellate, shed in thick irregular plates, inner bark light orange. Resin white, flammable. Leaves 
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long, 3-4(6)-jugate; petiole (1.4) 
<quantity box="[429,522,757,784]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="61" pageNumber="338" unit="cm" value="7.5" valueMax="10.0" valueMin="5.0">5-10 cm</quantity>
, petiole and rachis with sparse to scattered erect ferrugineous hairs to 
<quantity box="[614,713,789,816]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="61" pageNumber="338" unit="mm" value="0.05">0.05 mm</quantity>
long, also sparse snail-shaped glands, in fruit often woody and lenticellate; stipules (1) 
<quantity box="[346,445,853,880]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="13.0" metricValueMin="5.0" pageId="61" pageNumber="338" unit="mm" value="9.0" valueMax="13.0" valueMin="5.0">5-13 mm</quantity>
from petiole insertion, 
<quantity box="[689,775,853,880]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="61" pageNumber="338" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">5-6 mm</quantity>
long, obovate and usually constricted at base, surfaces with dense ferrugineous ascending to appressed hairs to 
<quantity box="[677,776,917,944]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="61" pageNumber="338" unit="mm" value="0.25">0.25 mm</quantity>
long, the scar 
<quantity box="[289,379,948,975]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="61" pageNumber="338" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">3-5 mm</quantity>
long; basal petiolules (2) 4-14 (20) mm, other laterals (3) 
<quantity box="[363,462,980,1007]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="2.0" metricValueMin="0.6" pageId="61" pageNumber="338" unit="mm" value="13.0" valueMax="20.0" valueMin="6.0">6-20 mm</quantity>
long, terminal one (10) 14-40 (56) mm long, not canaliculate, lateral pulvinuli sometimes conspicuous; basal leaflets (1.8) 3-10.5 
<emphasis box="[563,579,1045,1070]" italics="true" pageId="61" pageNumber="338">×</emphasis>
(1.1) 
<quantity box="[649,772,1044,1071]" metricMagnitude="-2" metricUnit="m" metricValue="3.6999999999999997" metricValueMax="5.6" metricValueMin="1.8" pageId="61" pageNumber="338" unit="cm" value="3.6999999999999997" valueMax="5.6" valueMin="1.8">1.8-5.6 cm</quantity>
, (broadly) ovate to suborbicular, other laterals (3.5) 5.8-13 
<emphasis box="[759,775,1077,1102]" italics="true" pageId="61" pageNumber="338">×</emphasis>
(1.7) 
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, (oblong-)ovate, less often (oblong)obovate or rarely broadly elliptic, terminal one 4.7-14 
<emphasis box="[626,642,1141,1166]" italics="true" pageId="61" pageNumber="338">×</emphasis>
<quantity box="[649,770,1140,1167]" metricMagnitude="-2" metricUnit="m" metricValue="5.3" metricValueMax="8.2" metricValueMin="2.4" pageId="61" pageNumber="338" unit="cm" value="5.3" valueMax="8.2" valueMin="2.4">2.4-8.2 cm</quantity>
, obovate, less often broadly elliptic, rarely ovate; leaflet apex rounded to abruptly and broadly short-acuminate, the acumen 
<quantity box="[130,216,1236,1262]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="6.0" metricValueMin="2.0" pageId="61" pageNumber="338" unit="mm" value="4.0" valueMax="6.0" valueMin="2.0">2-6 mm</quantity>
long, leaflet base usually symmetric, sometimes basal insertion slightly asymmetric, cordate or less often truncate, very rarely acute; margin flat; leaflets coriaceous, sometimes splitting along midvein when pressed and dried, drying (greenish) brown, dull or sometimes glossy on adaxial side and rarely on abaxial side; secondary vein framework brochidodromous but usually looping near the margin, secondaries in 8-13 pairs, slightly arcuate to spreading, spacing decreasing toward apex and markedly toward base, angle increasing toward base, intercostal tertiaries alternate-percurrent or sometime irregular-reticulate, quaternaries regular-polygonal; on abaxial side the midvein narrowly prominent, secondaries prominent but slightly sunk in a groove, tertiaries usually narrowly prominulous and then often sunk, sometimes flat or broadly prominulous, the midvein with sparse to scattered erect ferrugineous hairs to 
<quantity box="[420,521,1714,1741]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="61" pageNumber="338" unit="mm" value="0.05">0.05 mm</quantity>
long, also flexuous ferrugineous hairs to 
<quantity box="[336,426,1746,1772]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="61" pageNumber="338" unit="mm" value="0.2">0.2 mm</quantity>
, also sparse snail-shaped glands, rest of surface with scattered short thick appressed hairs; on adaxial side the midvein narrowly prominulous but sunk in a groove, secondaries narrowly prominulous but usually sunk, rest of veins broad and almost flat, surface subglabrous or with fine erect or ascending white hairs, these sparse to scattered on midvein, relatively dense on petiolules, scattered on rest of surface plus a few along the margin. Inflorescences 
<quantity box="[684,775,1969,1996]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" metricValueMax="2.5" metricValueMin="0.5" pageId="61" pageNumber="338" unit="cm" value="15.0" valueMax="25.0" valueMin="5.0">5-25 cm</quantity>
long, axes with dense flexuous ferrugineous hairs to 
<quantity box="[689,775,2001,2028]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="61" pageNumber="338" unit="mm" value="0.3">0.3 mm</quantity>
long, also some elongate lenticels; staminate inflorescences with secondary axes to 
<quantity box="[1064,1118,247,274]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" pageId="61" pageNumber="338" unit="cm" value="9.0">9 cm</quantity>
long; bracts on axes 
<emphasis box="[1345,1362,248,273]" italics="true" pageId="61" pageNumber="338">c.</emphasis>
<quantity box="[1369,1455,247,274]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="61" pageNumber="338" unit="mm" value="7.8">7.8 mm</quantity>
long, with dense flexuous golden hairs to 
<quantity box="[1258,1343,279,306]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="61" pageNumber="338" unit="mm" value="0.2">0.2 mm</quantity>
long, narrowly elliptic, some ovate (especially on primary axes); bracteoles 
<emphasis box="[880,897,344,369]" italics="true" pageId="61" pageNumber="338">c.</emphasis>
<quantity box="[904,990,343,369]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="61" pageNumber="338" unit="mm" value="3.7">3.7 mm</quantity>
long, obovate and acuminate, often longer than buds; pedicel 3.4-4.6 
<emphasis box="[1101,1117,375,400]" italics="true" pageId="61" pageNumber="338">×</emphasis>
0.9-1.3. Staminate flowers 
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long overall; calyx 2.1-2.6 
<emphasis box="[1196,1212,407,432]" italics="true" pageId="61" pageNumber="338">×</emphasis>
<quantity box="[1220,1333,406,433]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" metricValueMax="4.0" metricValueMin="3.6" pageId="61" pageNumber="338" unit="mm" value="3.8" valueMax="4.0" valueMin="3.6">3.6-4 mm</quantity>
, taller than ovariodisk, deeply cupular, lobes 
<quantity box="[1151,1276,438,465]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" metricValueMax="1.7" metricValueMin="0.6" pageId="61" pageNumber="338" unit="mm" value="1.15" valueMax="1.7" valueMin="0.6">0.6-1.7 mm</quantity>
long, (depressed) rounded-deltate, abaxial surface with dense appressed flexuous ferrugineous hairs to 
<quantity box="[1081,1180,502,529]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="61" pageNumber="338" unit="mm" value="0.15">0.15 mm</quantity>
long, adaxial surface with dense ascending to appressed golden hairs to 
<quantity box="[1304,1391,534,561]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="61" pageNumber="338" unit="mm" value="0.3">0.3 mm</quantity>
long; petals 5-5.3 
<emphasis box="[945,961,567,592]" italics="true" pageId="61" pageNumber="338">×</emphasis>
<quantity box="[967,1056,566,593]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="61" pageNumber="338" unit="mm" value="3.4">3.4 mm</quantity>
, exposed part slightly shorter than or equal to calyx, ovate, abaxial surface with appressed golden hairs to 
<quantity box="[899,984,630,656]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="61" pageNumber="338" unit="mm" value="0.3">0.3 mm</quantity>
long, these retrorse at apex, oriented toward the apex up the center of the petals, and exmedial up the sides of the petals; stamens 
<quantity box="[1047,1177,693,720]" metricMagnitude="-3" metricUnit="m" metricValue="3.45" metricValueMax="3.5" metricValueMin="3.4" pageId="61" pageNumber="338" unit="mm" value="3.45" valueMax="3.5" valueMin="3.4">3.4-3.5 mm</quantity>
long (sometimes stamens slightly dimorphic in length), anthers 
<quantity box="[1216,1345,726,752]" metricMagnitude="-3" metricUnit="m" metricValue="1.2999999999999998" metricValueMax="1.4" metricValueMin="1.2" pageId="61" pageNumber="338" unit="mm" value="1.2999999999999998" valueMax="1.4" valueMin="1.2">1.2-1.4 mm</quantity>
long, narrowly oblong-ovate in dorsiventral view, lanceolate to narrowly ovate in lateral view; ovariodisk 0.8-1.5 
<emphasis box="[1231,1247,790,815]" italics="true" pageId="61" pageNumber="338">×</emphasis>
<quantity box="[1250,1380,789,816]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" metricValueMax="1.6" metricValueMin="1.2" pageId="61" pageNumber="338" unit="mm" value="1.4" valueMax="1.6" valueMin="1.2">1.2-1.6 mm</quantity>
, shortcolumnar to perdepressed-obovoid, apex truncate and craggy. Pistillate inflorescences with secondary axes to 
<quantity box="[1317,1393,853,880]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" pageId="61" pageNumber="338" unit="cm" value="5.5">5.5 cm</quantity>
long; bracts on axes 
<quantity box="[963,1091,885,911]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" metricValueMax="8.1" metricValueMin="6.1" pageId="61" pageNumber="338" unit="mm" value="7.1" valueMax="8.1" valueMin="6.1">6.1-8.1 mm</quantity>
long, ovate to obovate and foliose, with ascending to appressed flexuous golden hairs to 
<quantity box="[1369,1453,917,943]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="61" pageNumber="338" unit="mm" value="0.3">0.3 mm</quantity>
long; bracteoles 
<quantity box="[994,1089,948,975]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="61" pageNumber="338" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">5-6 mm</quantity>
, shorter than buds, obovate and acuminate; pedicel 2 
<emphasis box="[1044,1060,981,1006]" italics="true" pageId="61" pageNumber="338">×</emphasis>
<quantity box="[1067,1180,980,1007]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="61" pageNumber="338" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">1-1.5 mm</quantity>
, slightly clavate. Pistillate flowers 
<quantity box="[893,1022,1012,1039]" metricMagnitude="-3" metricUnit="m" metricValue="9.45" metricValueMax="9.5" metricValueMin="9.4" pageId="61" pageNumber="338" unit="mm" value="9.45" valueMax="9.5" valueMin="9.4">9.4-9.5 mm</quantity>
long overall; calyx 5.5-6.5 
<emphasis box="[1306,1322,1013,1038]" italics="true" pageId="61" pageNumber="338">×</emphasis>
<quantity box="[1325,1453,1012,1039]" metricMagnitude="-3" metricUnit="m" metricValue="6.199999999999999" metricValueMax="6.6" metricValueMin="5.8" pageId="61" pageNumber="338" unit="mm" value="6.199999999999999" valueMax="6.6" valueMin="5.8">5.8-6.6 mm</quantity>
overall, the lobes 
<quantity box="[1003,1135,1044,1071]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="61" pageNumber="338" unit="mm" value="1.0" valueMax="1.2" valueMin="0.8">0.8-1.2 mm</quantity>
long, rounded perdepresseddeltate; petals 6 
<emphasis box="[988,1004,1077,1102]" italics="true" pageId="61" pageNumber="338">×</emphasis>
<quantity box="[1011,1100,1076,1103]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="61" pageNumber="338" unit="mm" value="3.8">3.8 mm</quantity>
, exposed part much shorter than calyx, ovate, fleshy; staminodes 
<quantity box="[1164,1296,1108,1135]" metricMagnitude="-3" metricUnit="m" metricValue="2.5999999999999996" metricValueMax="2.8" metricValueMin="2.4" pageId="61" pageNumber="338" unit="mm" value="2.5999999999999996" valueMax="2.8" valueMin="2.4">2.4-2.8 mm</quantity>
long, inserted around rim of a hypanthium 
<quantity box="[1143,1275,1140,1167]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="61" pageNumber="338" unit="mm" value="1.4" valueMax="1.5" valueMin="1.3">1.3-1.5 mm</quantity>
deep, level with the base of the stigmas, the anthers 
<quantity box="[1196,1302,1172,1198]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="61" pageNumber="338" unit="mm" value="1.05" valueMax="1.1" valueMin="1.0">1-1.1 mm</quantity>
long, ovate in dorsiventral view and oblong in lateral view, pubescent; pistil 5-5.4 
<emphasis box="[877,893,1236,1261]" italics="true" pageId="61" pageNumber="338">×</emphasis>
<quantity box="[898,1028,1236,1262]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" metricValueMax="3.2" metricValueMin="2.6" pageId="61" pageNumber="338" unit="mm" value="2.9" valueMax="3.2" valueMin="2.6">2.6-3.2 mm</quantity>
overall, ovary ovoid and with appressed golden hairs to 
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long, the style 1.1 
<emphasis box="[1265,1281,1268,1293]" italics="true" pageId="61" pageNumber="338">×</emphasis>
<quantity box="[1285,1393,1267,1294]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" metricValueMax="1.6" metricValueMin="1.0" pageId="61" pageNumber="338" unit="mm" value="1.3" valueMax="1.6" valueMin="1.0">1-1.6 mm</quantity>
long, the stigmata area 1.1-1.4 tall, 3-lobed. Fruiting pedicel 3-10 
<emphasis box="[1440,1456,1300,1325]" italics="true" pageId="61" pageNumber="338">×</emphasis>
<quantity box="[811,928,1331,1358]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" metricValueMax="4.2" metricValueMin="3.0" pageId="61" pageNumber="338" unit="mm" value="3.6" valueMax="4.2" valueMin="3.0">3-4.2 mm</quantity>
, slightly clavate, with scattered erect ferrugineous to golden hairs 
<quantity box="[1025,1126,1363,1390]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="61" pageNumber="338" unit="mm" value="0.25">0.25 mm</quantity>
long, fruiting calyx 
<quantity box="[1346,1451,1363,1390]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="10.0" metricValueMin="4.0" pageId="61" pageNumber="338" unit="mm" value="7.0" valueMax="10.0" valueMin="4.0">4-10 mm</quantity>
, shallowly cupular, lobes sometimes distinct, 
<quantity box="[1301,1391,1395,1422]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="61" pageNumber="338" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">2-4 mm</quantity>
long, patent, with scattered lenticels. Fruits 3.5-5.1 
<emphasis box="[1307,1323,1428,1453]" italics="true" pageId="61" pageNumber="338">×</emphasis>
<quantity box="[1329,1451,1427,1454]" metricMagnitude="-2" metricUnit="m" metricValue="2.9" metricValueMax="3.5" metricValueMin="2.3" pageId="61" pageNumber="338" unit="cm" value="2.9" valueMax="3.5" valueMin="2.3">2.3-3.5 cm</quantity>
, brown or gray, ovoid to broadly oblong-ellipsoid to slightly obovoid, sometime slightly angular, rarely broadly ovoid, apex (slightly) tapering to rounded, base truncate to obtuse, surface with lenticels usually dense, relatively large, very raised, slightly ferrugineous, among the lenticels glabrous or some fine white hairs to 
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long.
</paragraph>
<paragraph blockId="61.[812,1457,1683,2028]" box="[813,887,1683,1709]" pageId="61" pageNumber="338">NOTES</paragraph>
<paragraph blockId="61.[812,1457,1683,2028]" pageId="61" pageNumber="338">
<taxonomicName authority="Daly &amp; Raharimampionona &amp; Federman, 2015" authorityName="Daly &amp; Raharimampionona &amp; Federman" authorityYear="2015" box="[813,1054,1714,1740]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="61" pageNumber="338" phylum="Tracheophyta" rank="species" species="scholasticum" status="sp. nov.">
<emphasis box="[813,1054,1714,1740]" italics="true" pageId="61" pageNumber="338">Canarium scholasticum</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel box="[1066,1147,1715,1741]" pageId="61" pageNumber="338" rank="species">sp. nov.</taxonomicNameLabel>
belongs to a group of 10 species that have relatively large, broad leaflets with truncate to slightly cordate base; within that group, it is contrasted with 
<taxonomicName authority="Daly &amp; Raharimampionona &amp; Federman, 2015" authorityName="Daly &amp; Raharimampionona &amp; Federman" authorityYear="2015" box="[812,958,1810,1836]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="61" pageNumber="338" phylum="Tracheophyta" rank="species" species="pilicarpum" status="sp. nov.">
<emphasis box="[812,958,1810,1836]" italics="true" pageId="61" pageNumber="338">C. pilicarpum</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel box="[974,1057,1810,1836]" pageId="61" pageNumber="338" rank="species">sp. nov.</taxonomicNameLabel>
in the discussion under that species, and outside that group it is contrasted with 
<taxonomicName authority="Daly &amp; Raharimampionona &amp; Federman, 2015" authorityName="Daly &amp; Raharimampionona &amp; Federman" authorityYear="2015" box="[1289,1448,1842,1868]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="61" pageNumber="338" phylum="Tracheophyta" rank="species" species="ferrugineum" status="sp. nov.">
<emphasis box="[1289,1448,1842,1868]" italics="true" pageId="61" pageNumber="338">C. ferrugineum</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel box="[813,894,1874,1900]" pageId="61" pageNumber="338" rank="species">sp. nov.</taxonomicNameLabel>
in the discussion under that species.
</paragraph>
<paragraph blockId="61.[812,1457,1683,2028]" lastBlockId="63.[131,775,215,689]" lastPageId="63" lastPageNumber="340" pageId="61" pageNumber="338">
Both 
<taxonomicName authority="Daly &amp; Raharimampionona &amp; Federman, 2015" authorityName="Daly &amp; Raharimampionona &amp; Federman" authorityYear="2015" box="[897,1041,1905,1932]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="61" pageNumber="338" phylum="Tracheophyta" rank="species" species="lamianum" status="sp. nov.">
<emphasis box="[897,1041,1905,1932]" italics="true" pageId="61" pageNumber="338">C. lamianum</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel box="[1055,1139,1906,1932]" pageId="61" pageNumber="338" rank="species">sp. nov.</taxonomicNameLabel>
and 
<taxonomicName authority="Daly &amp; Raharimampionona &amp; Federman, 2015" authorityName="Daly &amp; Raharimampionona &amp; Federman" authorityYear="2015" box="[1191,1356,1905,1932]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="61" pageNumber="338" phylum="Tracheophyta" rank="species" species="scholasticum" status="sp. nov.">
<emphasis box="[1191,1356,1905,1932]" italics="true" pageId="61" pageNumber="338">C. scholasticum</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel box="[1372,1455,1906,1932]" pageId="61" pageNumber="338" rank="species">sp. nov.</taxonomicNameLabel>
for the most part have usually 3-4-jugate leaves, stipules 
<quantity box="[815,921,1969,1996]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="13.0" metricValueMin="4.0" pageId="61" pageNumber="338" unit="mm" value="8.5" valueMax="13.0" valueMin="4.0">4-13 mm</quantity>
from petiole insertion and 
<quantity box="[1237,1328,1969,1996]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="7.0" metricValueMin="3.0" pageId="61" pageNumber="338" unit="mm" value="5.0" valueMax="7.0" valueMin="3.0">3-7 mm</quantity>
long, basal petiolules 
<quantity box="[930,1035,2001,2028]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="14.0" metricValueMin="4.0" pageId="61" pageNumber="338" unit="mm" value="9.0" valueMax="14.0" valueMin="4.0">4-14 mm</quantity>
long, terminal leaflet obovate, leaflet apex abruptly and broadly short-acuminate with the acumen 2-7 (11) mm long, and calyx deeply cupular in pistillate flowers. The latter differs by the bark finely fissured if at all (vs deeply fissured in 
<taxonomicName authority="Daly &amp; Raharimampionona &amp; Federman, 2015" authorityName="Daly &amp; Raharimampionona &amp; Federman" authorityYear="2015" box="[467,612,311,337]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="63" pageNumber="340" phylum="Tracheophyta" rank="species" species="lamianum" status="sp. nov.">
<emphasis box="[467,612,311,337]" italics="true" pageId="63" pageNumber="340">C. lamianum</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel box="[628,718,311,337]" pageId="63" pageNumber="340" rank="species">sp. nov.</taxonomicNameLabel>
), the lateral leaflets (oblong-)ovate (vs variously shaped but not ovate) and often folding lengthwise when dry (vs flat), the lateral leaflet base cordate or less often truncate, very rarely acute (vs obtuse to truncate to acute, sometimes shortattenuate), the secondary vein angle decreasing toward the base (vs increasing or uniform), on abaxial surface the secondary veins more prominent and the lamina often corrugated between secondary veins, on adaxial surface the secondary veins discolorous, and in staminate flowers the stamens inserted around the base of the ovariodisk (vs inserted on the side).
</paragraph>
</subSubSection>
</treatment>
</document>