219 lines
15 KiB
XML
219 lines
15 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.473.8659" ID-GBIF-Dataset="f0fbc7da-005d-4414-b66d-ca97ccf32a02" ID-PMC="PMC4304041" ID-Pensoft-Pub="1313-2970-473-1" ID-PubMed="25632253" ID-ZBK="02943D336D534CB6A6BD47526EC80C67" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-473-1" ModsDocOrigin="ZooKeys 473" ModsDocTitle="Systematics of the family Plectopylidae in Vietnam with additional information on Chinese taxa (Gastropoda, Pulmonata, Stylommatophora)" checkinTime="1451244823217" checkinUser="pensoft" docAuthor="Pall-Gergely, Barna, Hunyadi, Andras, Ablett, Jonathan, Lương, Hao Văn, Fred Naggs, & Asami, Takahiro" docDate="2015" docId="618DB50B05918F7A415EBB97C2DF3A38" docLanguage="en" docName="ZooKeys 473: 1-118" docOrigin="ZooKeys 473" docSource="http://dx.doi.org/10.3897/zookeys.473.8659" docTitle="Gudeodiscus (Gudeodiscus) hemmeni Pall-Gergely & Hunyadi, sp. n." docType="treatment" docUuid="5A98BC18-CF82-4C2F-BCE4-DCCA8DBBED3B" docUuidSource="ZooBank" docVersion="5" lastPageNumber="22" masterDocId="145AFF9BD02C0945FFA62A78FF974356" masterDocTitle="Systematics of the family Plectopylidae in Vietnam with additional information on Chinese taxa (Gastropoda, Pulmonata, Stylommatophora)" masterLastPageNumber="118" masterPageNumber="1" pageNumber="22" updateTime="1668159818540" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Systematics of the family Plectopylidae in Vietnam with additional information on Chinese taxa (Gastropoda, Pulmonata, Stylommatophora)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Pall-Gergely, Barna</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hunyadi, Andras</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ablett, Jonathan</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Lương, Hao Văn</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fred Naggs,</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Asami, Takahiro</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2015</mods:date>
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<mods:detail type="volume">
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<mods:number>473</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>118</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.473.8659</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.473.8659</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-473-1</mods:identifier>
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<mods:identifier type="ZBK">02943D336D534CB6A6BD47526EC80C67</mods:identifier>
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<mods:identifier type="ZooBank">02943D336D534CB6A6BD47526EC80C67</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152057804" LSID="urn:lsid:zoobank.org:act:5A98BC18-CF82-4C2F-BCE4-DCCA8DBBED3B" httpUri="http://treatment.plazi.org/id/618DB50B05918F7A415EBB97C2DF3A38" lastPageNumber="22" pageId="21" pageNumber="22">
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<subSubSection pageId="21" pageNumber="22" type="multiple">
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<paragraph pageId="21" pageNumber="22">Taxon classification Animalia Stylommatophora Plectopylidae</paragraph>
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</subSubSection>
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<subSubSection pageId="21" pageNumber="22" type="nomenclature">
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<paragraph pageId="21" pageNumber="22">
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<taxonomicName LSID="http://zoobank.org/5A98BC18-CF82-4C2F-BCE4-DCCA8DBBED3B" authority="Pall-Gergely & Hunyadi" class="Gastropoda" family="Plectopylidae" genus="Gudeodiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gudeodiscus (Gudeodiscus) hemmeni" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="hemmeni" subGenus="Gudeodiscus">
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Gudeodiscus (Gudeodiscus?) hemmeni
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<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
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& Hunyadi
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</taxonomicName>
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<taxonomicNameLabel pageId="21" pageNumber="22">sp. n.</taxonomicNameLabel>
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Figures 2
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<normalizedToken originalValue="C–D">C-D</normalizedToken>
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, 9F, 11
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<normalizedToken originalValue="G–J">G-J</normalizedToken>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="21" pageNumber="22" type="type material">
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<paragraph pageId="21" pageNumber="22">Type material.</paragraph>
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<paragraph pageId="21" pageNumber="22">
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2012/61
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<normalizedToken originalValue="Sơn">Sơn</normalizedToken>
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La Province,
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<normalizedToken originalValue="Hà">Ha</normalizedToken>
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<normalizedToken originalValue="Nội">Nội</normalizedToken>
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156 km towards
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<normalizedToken originalValue="Mộc">Mộc</normalizedToken>
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<normalizedToken originalValue="Châu">Chau</normalizedToken>
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, left side of the road nr. 6, rocky wall, 1110 m,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="20.76655">20°45.993'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="104.8978">104°53.868'E</geoCoordinate>
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, leg. Hunyadi, A., 06.06.2012., holotype HNHM 97458 (Figure 2C), HA/11 paratypes+4jb (not paratype), PGB/3 paratypes; 2012/62
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<normalizedToken originalValue="Sơn">Sơn</normalizedToken>
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La Province,
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<normalizedToken originalValue="Hà">Ha</normalizedToken>
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<normalizedToken originalValue="Nội">Nội</normalizedToken>
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156 km towards
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<normalizedToken originalValue="Mộc">Mộc</normalizedToken>
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<normalizedToken originalValue="Châu">Chau</normalizedToken>
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, right side of the road nr. 6, rocky wall, 1110 m,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="20.768084">20°46.085'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="104.89813">104°53.888'E</geoCoordinate>
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, leg. Hunyadi, A., 06.06.2012., HA/13 paratypes+2jb (not paratypes), PGB/1; Vn12-104
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<normalizedToken originalValue="Sơn">Sơn</normalizedToken>
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La Province, right side off road
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<normalizedToken originalValue="Mộc">Mộc</normalizedToken>
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<normalizedToken originalValue="Châu">Chau</normalizedToken>
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to
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<normalizedToken originalValue="Sơn">Sơn</normalizedToken>
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La,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="20.876116">20°52.567'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="104.5885">104°35.310'E</geoCoordinate>
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, leg. Hemmen, Ch., 02.10.2012., HE/1; Vn10-103A
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<normalizedToken originalValue="Hòa">Hoa</normalizedToken>
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<normalizedToken originalValue="Bình">Binh</normalizedToken>
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Province, ca. km 156 old road
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<normalizedToken originalValue="Hà">Ha</normalizedToken>
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<normalizedToken originalValue="Nội">Nội</normalizedToken>
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to
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<normalizedToken originalValue="Sơn">Sơn</normalizedToken>
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La (right side off road),
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="20.766666">20°46.000'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="104.89809">104°53.885'E</geoCoordinate>
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, leg. Hemmen, Ch. & J., 15.10.2010., HE/1 (Figure 2D); Vn10-76A
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<normalizedToken originalValue="Sơn">Sơn</normalizedToken>
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La Province, ca. 32 km from
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<normalizedToken originalValue="Mộc">Mộc</normalizedToken>
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<normalizedToken originalValue="Châu">Chau</normalizedToken>
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to
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<normalizedToken originalValue="Hà">Ha</normalizedToken>
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<normalizedToken originalValue="Nội">Nội</normalizedToken>
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(old road),
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="20.789183">20°47.351'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="104.83438">104°50.063'E</geoCoordinate>
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, leg. Hemmen, Ch. & J., 02.04.2010., HE/1.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="21" pageNumber="22" type="diagnosis">
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<paragraph pageId="21" pageNumber="22">Diagnosis.</paragraph>
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<paragraph pageId="21" pageNumber="22">
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Shell small, with slightly elevated spire, characteristically shaped aperture having wide upper sinulus and small apertural fold (Figure 9F); parietal wall with two lamellae and horizontal plicae above and below; palatal plicae depressed Z-shaped; free from each other, or connected to each other with a ridge (Figures 11
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<normalizedToken originalValue="G–J">G-J</normalizedToken>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="21" pageNumber="22" type="description">
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<paragraph pageId="21" pageNumber="22">Description.</paragraph>
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<paragraph pageId="21" pageNumber="22">Shell very small to small, light brown to chocolate brown, with slightly elevated spire, consists of 5.25-5.5 whorls; suture relatively shallow, especially at the first 3-4 whorls; protoconch (2.25-2.5 whorls) glossy, very finely, regularly ribbed, but the ribs are sometimes hardly visible, they are more prominent at the upper part of the whorls, close to the suture; teleoconch without notable spiral lines, very finely regularly ribbed; sculpture strength equal on ventral and dorsal side; umbilicus narrow and deep; aperture with widened upper part (sinulus), apertural lip whitish, thin, slightly expanded but not reflexed; apertural denticle (fold) always present, very small, free from the callus or connected to it.</paragraph>
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<paragraph pageId="21" pageNumber="22">Two specimens were opened. Parietal side with a stronger anterior lamella with anteriorly widened lower part, and a slimmer posterior lamella; shorter upper and longer lower horizontal plicae free from the anterior lamella, the lower one a bit extends beyond the anterior lamella in the anterior direction. Palatal side with six plicae; first and last are straight, the others are depressed Z-shaped and are connected with a ridge.</paragraph>
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</subSubSection>
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<subSubSection pageId="21" pageNumber="22" type="measurements">
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<paragraph pageId="21" pageNumber="22">Measurements</paragraph>
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<paragraph pageId="21" pageNumber="22">(in mm): D = 9.5-10.1, H = 4.3-5.2 (n=5, belonging to different populations).</paragraph>
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</subSubSection>
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<subSubSection pageId="21" pageNumber="22" type="differential diagnosis">
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<paragraph pageId="21" pageNumber="22">Differential diagnosis.</paragraph>
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<paragraph pageId="21" pageNumber="22">
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Gudeodiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gudeodiscus hemmeni" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="hemmeni">Gudeodiscus hemmeni</taxonomicName>
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sp. n. differs from most
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Gudeodiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gudeodiscus phlyarius" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="phlyarius">Gudeodiscus phlyarius</taxonomicName>
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populations by the smaller shell, shorter denticle (fold) in the aperture, thinner apertural lip, the wider and reflexed apertural rim, the wide upper sinus of the aperture, lack of spiral lines in the sculpture and narrower umbilicus.
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Gudeodiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gudeodiscus anceyi" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="anceyi">Gudeodiscus anceyi</taxonomicName>
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is usually smaller, has a longer apertural fold, prominent spiral sculpture, a weaker callus and differently shaped aperture.
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</paragraph>
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<paragraph pageId="21" pageNumber="22">
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In all localities,
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Gudeodiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gudeodiscus hemmeni" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="hemmeni">Gudeodiscus hemmeni</taxonomicName>
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sp. n. lives sympatrically with
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Gudeodiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gudeodiscus messageri" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="subSpecies" species="messageri" subSpecies="raheemi">Gudeodiscus messageri raheemi</taxonomicName>
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ssp. n., which is much larger, lacks the apertural fold, and usually has an anterior lamella which is dissolved into small denticles.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="21" pageNumber="22" type="intraspecific diversity">
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<paragraph pageId="21" pageNumber="22">Intraspecific diversity.</paragraph>
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<paragraph pageId="21" pageNumber="22">Low; shell characters are stable, although only a few shells are known.</paragraph>
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</subSubSection>
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<subSubSection pageId="21" pageNumber="22" type="etymology">
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<paragraph pageId="21" pageNumber="22">Etymology.</paragraph>
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<paragraph pageId="21" pageNumber="22">The new species is dedicated to Jens Hemmen (1944-2012), malacologist and much-valued friend, who contributed to our revision by providing shell and ethanol-preserved material.</paragraph>
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</subSubSection>
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<subSubSection pageId="21" pageNumber="22" type="type locality">
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<paragraph pageId="21" pageNumber="22">Type locality.</paragraph>
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<paragraph pageId="21" pageNumber="22">
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<normalizedToken originalValue="Sơn">Sơn</normalizedToken>
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La Province,
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<normalizedToken originalValue="Hà">Ha</normalizedToken>
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<normalizedToken originalValue="Nội">Nội</normalizedToken>
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156 km towards
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<normalizedToken originalValue="Mộc">Mộc</normalizedToken>
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<normalizedToken originalValue="Châu">Chau</normalizedToken>
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, left side of the road nr. 6, rocky wall, 1110 m,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="20.76655">20°45.993'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="104.8978">104°53.868'E</geoCoordinate>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="21" pageNumber="22" type="distribution">
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<paragraph pageId="21" pageNumber="22">Distribution</paragraph>
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<paragraph pageId="21" pageNumber="22">
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(see Figure 43). The new species is known from few locations in south-eastern
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<normalizedToken originalValue="Sơn">Sơn</normalizedToken>
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La province.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |