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<document id="F538D208B9CB7409A4BAC22CC7889E59" ID-CLB-Dataset="31049" ID-DOI="10.11646/zootaxa.4370.1.8" ID-GBIF-Dataset="e3cfede2-b8fc-4c0a-9482-03411666548f" ID-ISSN="1175-5326" ID-Zenodo-Dep="1138554" ID-ZooBank="EC53F125-FAA9-4C3A-9F16-BCFDCB926EC4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1515488951785" checkinUser="plazi" docAuthor="Stauffer, Jay R." docDate="2018" docId="21673B771A16FFF5FF1DFC780139FAA4" docLanguage="en" docName="zootaxa.4370.1.7.pdf" docOrigin="Zootaxa 4370 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Metriaclima koningsi Stauffer 2018, new species" docType="treatment" docUuid="632F7DC1-2D74-47EF-9FD5-338D89BDC0FE" docUuidSource="ZooBank" docVersion="8" lastPageNumber="99" masterDocId="DD5E430F1A17FFF1FF8AFFA80577FC3C" masterDocTitle="Description of Metriaclima koningsi, a new species (Teleostei: Cichlidae) from Lake Malaŵi, Malaŵi, Africa" masterLastPageNumber="100" masterPageNumber="95" pageNumber="96" updateTime="1698513055472" updateUser="ExternalLinkService">
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<mods:title id="F90867E71FFF1CF7FB19EA4971B5567F">Description of Metriaclima koningsi, a new species (Teleostei: Cichlidae) from Lake Malaŵi, Malaŵi, Africa</mods:title>
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<mods:namePart id="0DEEA9D74A4D79FEC2AC64D4342F3F02">Stauffer, Jay R.</mods:namePart>
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<mods:date id="7A54351E5C64C9105B39A6E206B66BAF">2018</mods:date>
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<mods:number id="CE3F3801C6325868C21D56A83B1BD0A9">2018-01-09</mods:number>
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<mods:identifier id="8B57AF1C31201B3028B188567BB8DB39" type="DOI">10.11646/zootaxa.4370.1.8</mods:identifier>
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<treatment id="21673B771A16FFF5FF1DFC780139FAA4" ID-DOI="http://doi.org/10.5281/zenodo.5974884" ID-GBIF-Taxon="139690206" ID-Zenodo-Dep="5974884" LSID="urn:lsid:zoobank.org:act:632F7DC1-2D74-47EF-9FD5-338D89BDC0FE" httpUri="http://treatment.plazi.org/id/21673B771A16FFF5FF1DFC780139FAA4" lastPageId="4" lastPageNumber="99" pageId="1" pageNumber="96">
<subSubSection id="E1D4D9EA1A16FFF0FF1DFC780597F830" pageId="1" pageNumber="96" type="nomenclature">
<paragraph id="A9718A611A16FFF0FF1DFC78074EFFD7" blockId="1.[151,569,976,1037]" box="[151,569,976,1003]" pageId="1" pageNumber="96">
<heading id="F2393D0D1A16FFF0FF1DFC78074EFFD7" bold="true" box="[151,569,976,1003]" fontSize="11" level="1" pageId="1" pageNumber="96" reason="1">
<emphasis id="9BBA56731A16FFF0FF1DFC78074EFFD7" bold="true" box="[151,569,976,1003]" pageId="1" pageNumber="96">
<taxonomicName id="6ECEF1E21A16FFF0FF1DFC7804EDFFD6" authority="Stauffer, 2018" authorityName="Stauffer" authorityYear="2018" box="[151,410,976,1002]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="96" phylum="Chordata" rank="species" species="koningsi" status="sp. nov.">
<emphasis id="9BBA56731A16FFF0FF1DFC7804EDFFD6" bold="true" box="[151,410,976,1002]" italics="true" pageId="1" pageNumber="96">Metriaclima koningsi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="8089EB081A16FFF0FE23FC79074EFFD7" box="[425,569,977,1003]" pageId="1" pageNumber="96" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="A9718A611A16FFF0FF1DFC5C0597F830" blockId="1.[151,569,976,1037]" box="[151,224,1012,1037]" pageId="1" pageNumber="96">
<figureCitation id="31F596E41A16FFF0FF1DFC5C05AEF830" box="[151,217,1012,1037]" captionStart="FIGURE 1" captionStartId="3.[151,250,1095,1117]" captionTargetBox="[151,1437,81,1074]" captionTargetId="figure@3.[151,1437,81,1074]" captionTargetPageId="3" captionText="FIGURE 1. The holotype of Metriaclima koningsi (56.8 mm SL, male, PSU 12891)." httpUri="https://zenodo.org/record/1138556/files/figure.png" pageId="1" pageNumber="96">Fig. 1</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E1D4D9EA1A16FFF0FF1DFB94070DF84C" pageId="1" pageNumber="96" type="reference_group">
<paragraph id="A9718A611A16FFF0FF1DFB94070DF84C" blockId="1.[151,683,1084,1136]" pageId="1" pageNumber="96">
<taxonomicName id="6ECEF1E21A16FFF0FF1DFB94044EF86E" ID-CoL="73XT" authorityName="Regan" authorityYear="1922" box="[151,313,1084,1106]" class="Actinopterygii" family="Cichlidae" genus="Pseudotropheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="96" phylum="Chordata" rank="genus">
<emphasis id="9BBA56731A16FFF0FF1DFB94044EF86E" box="[151,313,1084,1106]" italics="true" pageId="1" pageNumber="96">Pseudotropheus</emphasis>
</taxonomicName>
‘membe deep’ (
<bibRefCitation id="CD5FF7901A16FFF0FE54FB9407D3F86E" author="Ribbink" box="[478,676,1084,1106]" pageId="1" pageNumber="96" refString="Ribbink, A. J., Marsh, B. A., Marsh, A. C., Ribbink, A. C. &amp; Sharp, B. J. (1983) A preliminary survey of the cichlid fishes of rocky habitats in Lake Malawi. South African Journal of Zoology, 18, 149 - 310. https: // doi. org / 10.1080 / 02541858.1983.11447831" type="journal article" year="1983">
Ribbink
<emphasis id="9BBA56731A16FFF0FDBDFB95071CF86E" box="[567,619,1084,1106]" italics="true" pageId="1" pageNumber="96">et al.</emphasis>
1983
</bibRefCitation>
)
<taxonomicName id="6ECEF1E21A16FFF0FF1DFBF20463F84C" authorityName="Stauffer, Bowers, Kellogg &amp; McKaye" authorityYear="1997" box="[151,276,1114,1136]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="96" phylum="Chordata" rank="genus">
<emphasis id="9BBA56731A16FFF0FF1DFBF20463F84C" box="[151,276,1114,1136]" italics="true" pageId="1" pageNumber="96">Metriaclima</emphasis>
</taxonomicName>
sp. “membe deep” (
<bibRefCitation id="CD5FF7901A16FFF0FE68FBF20704F84C" author="Konings" box="[482,627,1114,1136]" pageId="1" pageNumber="96" refString="Konings, A. F. (1990) Cichlids and all the other fishes of Lake Malawi. T. F. H. Publications, Inc., Neptune City, New Jersey, 495 pp." type="book" year="1990">Konings 1990</bibRefCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="E1D4D9EA1A16FFF0FF1DFB35044FF979" pageId="1" pageNumber="96" type="materials_examined">
<paragraph id="A9718A611A16FFF0FF1DFB350657F8E5" blockId="1.[151,1437,1181,2034]" pageId="1" pageNumber="96">
<emphasis id="9BBA56731A16FFF0FF1DFB35047CF889" bold="true" box="[151,267,1181,1205]" pageId="1" pageNumber="96">
<typeStatus id="767534C31A16FFF0FF1DFB350471F889" box="[151,262,1181,1205]" pageId="1" pageNumber="96" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="CFDF12A41A16FFF0FE98FB360432F88A" box="[274,325,1182,1206]" country="USA" httpUri="http://grbio.org/cool/5mg9-yyc1" name="Portland State University, Vertebrate Biology Museum" pageId="1" pageNumber="96">PSU</collectionCode>
12891, adult male
<quantity id="6E3627841A16FFF0FD95FB3507F3F889" box="[543,644,1181,1206]" metricMagnitude="-2" metricUnit="m" metricValue="5.68" pageId="1" pageNumber="96" unit="mm" value="56.8">56.8 mm</quantity>
SL,
<geoCoordinate id="CCFAECA61A16FFF0FD30FB350622F88A" box="[698,853,1181,1206]" direction="south" orientation="latitude" pageId="1" pageNumber="96" precision="1" value="-12.0587">12.058700° S</geoCoordinate>
,
<geoCoordinate id="CCFAECA61A16FFF0FCE8FB350688F889" box="[866,1023,1181,1206]" direction="east" orientation="longitude" pageId="1" pageNumber="96" precision="1" value="34.75242">34.752420° E</geoCoordinate>
, Membe Point,
<collectingRegion id="6B0A44831A16FFF0FB38FB35007EF88A" box="[1202,1289,1181,1206]" country="Malawi" name="Likoma" pageId="1" pageNumber="96">Likoma</collectingRegion>
Island, Lake
<collectingCountry id="D1D9CAF11A16FFF0FF1DFB690598F8E6" box="[151,239,1217,1242]" name="Malawi" pageId="1" pageNumber="96">Malaŵi</collectingCountry>
,
<collectingCountry id="D1D9CAF11A16FFF0FF70FB690425F8E6" box="[250,338,1217,1242]" name="Malawi" pageId="1" pageNumber="96">Malaŵi</collectingCountry>
, Africa,
<date id="DD70ACA11A16FFF0FE39FB69072CF8E6" box="[435,603,1217,1242]" pageId="1" pageNumber="96" value="2005-08-05">5 August 2005</date>
, J. R. Stauffer, Jr.
</paragraph>
<paragraph id="A9718A611A16FFF0FF4DFB4D044FF979" blockId="1.[151,1437,1181,2034]" pageId="1" pageNumber="96">
<emphasis id="9BBA56731A16FFF0FF4DFB4D043FF8C1" bold="true" box="[199,328,1253,1277]" pageId="1" pageNumber="96">
<typeStatus id="767534C31A16FFF0FF4DFB4D0434F8C1" box="[199,323,1253,1277]" pageId="1" pageNumber="96" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<collectionCode id="CFDF12A41A16FFF0FED8FB4E04F1F8C2" box="[338,390,1254,1278]" country="USA" httpUri="http://grbio.org/cool/5mg9-yyc1" name="Portland State University, Vertebrate Biology Museum" pageId="1" pageNumber="96">PSU</collectionCode>
12892, 16 (
<quantity id="6E3627841A16FFF0FD9DFB4D07C8F8C1" box="[535,703,1253,1278]" metricMagnitude="-2" metricUnit="m" metricValue="4.895" metricValueMax="5.529999999999999" metricValueMin="4.26" pageId="1" pageNumber="96" unit="mm" value="48.95" valueMax="55.3" valueMin="42.6">42.6–55.3 mm</quantity>
SL), data as for
<typeStatus id="767534C31A16FFF0FC07FB4D0684F8C2" box="[909,1011,1253,1278]" pageId="1" pageNumber="96" type="holotype">holotype</typeStatus>
;
<collectionCode id="CFDF12A41A16FFF0FB8BFB4E0143F8C2" box="[1025,1076,1254,1278]" country="USA" httpUri="http://grbio.org/cool/5mg9-yyc1" name="Portland State University, Vertebrate Biology Museum" pageId="1" pageNumber="96">PSU</collectionCode>
12893, 3 (40.3–49.0 mm SL),
<geoCoordinate id="CCFAECA61A16FFF0FF1DFAA10446F91E" box="[151,305,1289,1314]" direction="south" orientation="latitude" pageId="1" pageNumber="96" precision="1" value="-12.046202">12.046202° S</geoCoordinate>
,
<geoCoordinate id="CCFAECA61A16FFF0FEB7FAA104AFF91D" box="[317,472,1289,1314]" direction="east" orientation="longitude" pageId="1" pageNumber="96" precision="1" value="34.7594">34.759400° E</geoCoordinate>
, Maingano Island, Lake
<collectingCountry id="D1D9CAF11A16FFF0FD67FAA10632F91E" box="[749,837,1289,1314]" name="Malawi" pageId="1" pageNumber="96">Malaŵi</collectingCountry>
,
<collectingCountry id="D1D9CAF11A16FFF0FCDAFAA106DFF91E" box="[848,936,1289,1314]" name="Malawi" pageId="1" pageNumber="96">Malaŵi</collectingCountry>
, Africa,
<date id="DD70ACA11A16FFF0FB82FAA201E6F91E" box="[1032,1169,1289,1314]" pageId="1" pageNumber="96" value="1918-02">18 February</date>
, 2002, A. F. Konings, J. R. Stauffer, Jr.
</paragraph>
</subSubSection>
<subSubSection id="E1D4D9EA1A16FFF0FF4DFAF906B7FAEE" pageId="1" pageNumber="96" type="diagnosis">
<paragraph id="A9718A611A16FFF0FF4DFAF906B7FAEE" blockId="1.[151,1437,1181,2034]" pageId="1" pageNumber="96">
<emphasis id="9BBA56731A16FFF0FF4DFAF90436F956" bold="true" box="[199,321,1361,1386]" pageId="1" pageNumber="96">Diagnosis.</emphasis>
The moderately-sloped vomer (43.1º in
<typeStatus id="767534C31A16FFF0FC80FAF90619F956" box="[778,878,1361,1386]" pageId="1" pageNumber="96" type="holotype">holotype</typeStatus>
) with a swollen rostral tip (
<figureCitation id="31F596E41A16FFF0FB2FFAF9019EF956" box="[1189,1257,1361,1386]" captionStart="FIGURE 2" captionStartId="3.[151,250,1911,1933]" captionTargetBox="[151,1436,1167,1890]" captionTargetId="figure@3.[151,1436,1167,1890]" captionTargetPageId="3" captionText="FIGURE 2. Three-dimensional surface rendering of the cranium of the holotype of Metriaclima koningsi (PSU 12891), showing the position of the parashpenoid and the angle of the ethmo-vomerine bloc." httpUri="https://zenodo.org/record/1138558/files/figure.png" pageId="1" pageNumber="96">Fig. 2</figureCitation>
), bicuspid teeth in the anterior portion of the outer row of both upper and lower jaws, the lower jaw at a 45º angle to a line from the tip of the snout to the hypural plate, a lower jaw that is slightly longer than the upper, and its feeding at a 90º angle (
<bibRefCitation id="CD5FF7901A16FFF0FF14FA150435F9EA" author="Konings" box="[158,322,1469,1494]" pageId="1" pageNumber="96" refString="Konings, A. F. (1990) Cichlids and all the other fishes of Lake Malawi. T. F. H. Publications, Inc., Neptune City, New Jersey, 495 pp." type="book" year="1990">Konings 1990</bibRefCitation>
) to the substrate place this species in
<taxonomicName id="6ECEF1E21A16FFF0FD75FA1506FDF9E9" authorityName="Stauffer, Bowers, Kellogg &amp; McKaye" authorityYear="1997" box="[767,906,1469,1493]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="96" phylum="Chordata" rank="genus">
<emphasis id="9BBA56731A16FFF0FD75FA1506FDF9E9" box="[767,906,1469,1493]" italics="true" pageId="1" pageNumber="96">Metriaclima</emphasis>
</taxonomicName>
. The lack of conspicuous vertical bars on the flank and the dark submarginal band in the dorsal fin of male
<taxonomicName id="6ECEF1E21A16FFF0FCE6FA4A0683F9C5" authorityName="Stauffer" authorityYear="2018" box="[876,1012,1505,1529]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="96" phylum="Chordata" rank="species" species="koningsi">
<emphasis id="9BBA56731A16FFF0FCE6FA4A0683F9C5" box="[876,1012,1505,1529]" italics="true" pageId="1" pageNumber="96">M. koningsi</emphasis>
</taxonomicName>
(
<figureCitation id="31F596E41A16FFF0FB8CFA49013CF9C6" box="[1030,1099,1505,1530]" captionStart="FIGURE 3" captionStartId="4.[151,250,774,796]" captionTargetBox="[151,1437,193,752]" captionTargetId="figure@4.[151,1437,193,753]" captionTargetPageId="4" captionText="FIGURE 3. Metriaclima koningsi. A, Male from Mainango Island; B, Female from Membe Point, Likoma Island. Neither of the above were collected." httpUri="https://zenodo.org/record/1138560/files/figure.png" pageId="1" pageNumber="96">Fig. 3</figureCitation>
) distinguish it from all other species of
<emphasis id="9BBA56731A16FFF0FE99F9AD04D2FA21" box="[275,421,1541,1565]" italics="true" pageId="1" pageNumber="96">
<taxonomicName id="6ECEF1E21A16FFF0FE99F9AD04D7FA21" authorityName="Stauffer, Bowers, Kellogg &amp; McKaye" authorityYear="1997" box="[275,416,1541,1565]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="96" phylum="Chordata" rank="genus">Metriaclima</taxonomicName>
,
</emphasis>
except
<taxonomicName id="6ECEF1E21A16FFF0FD88F9AD0608FA22" authority="Li, Konings" authorityName="Li, Konings &amp; Stauffer" authorityYear="2016" box="[514,895,1541,1566]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="96" phylum="Chordata" rank="species" species="usisyae">
<emphasis id="9BBA56731A16FFF0FD88F9AD079EFA21" box="[514,745,1541,1565]" italics="true" pageId="1" pageNumber="96">Metriaclima usisyae</emphasis>
Li, Konings
</taxonomicName>
, &amp; Stauffer. Males of
<taxonomicName id="6ECEF1E21A16FFF0FB0CF9AE0075FA21" authorityName="Li, Konings &amp; Stauffer" authorityYear="2016" box="[1158,1282,1542,1565]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="96" phylum="Chordata" rank="species" species="usisyae">
<emphasis id="9BBA56731A16FFF0FB0CF9AE0075FA21" box="[1158,1282,1542,1565]" italics="true" pageId="1" pageNumber="96">M. usisyae</emphasis>
</taxonomicName>
have a goldcoloured flank while this is blue in male
<taxonomicName id="6ECEF1E21A16FFF0FDD6F98207A8FA7D" authorityName="Stauffer" authorityYear="2018" box="[604,735,1577,1601]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="96" phylum="Chordata" rank="species" species="koningsi">
<emphasis id="9BBA56731A16FFF0FDD6F98207A8FA7D" box="[604,735,1577,1601]" italics="true" pageId="1" pageNumber="96">M. koningsi</emphasis>
</taxonomicName>
. Females of
<taxonomicName id="6ECEF1E21A16FFF0FCE4F9820685FA7D" authorityName="Stauffer" authorityYear="2018" box="[878,1010,1577,1601]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="96" phylum="Chordata" rank="species" species="koningsi">
<emphasis id="9BBA56731A16FFF0FCE4F9820685FA7D" box="[878,1010,1577,1601]" italics="true" pageId="1" pageNumber="96">M. koningsi</emphasis>
</taxonomicName>
are entirely brownish yellow and thus distinguished from those of most other species of
<taxonomicName id="6ECEF1E21A16FFF0FD49F9E50639FA59" authorityName="Stauffer, Bowers, Kellogg &amp; McKaye" authorityYear="1997" box="[707,846,1613,1637]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="96" phylum="Chordata" rank="genus">
<emphasis id="9BBA56731A16FFF0FD49F9E50639FA59" box="[707,846,1613,1637]" italics="true" pageId="1" pageNumber="96">Metriaclima</emphasis>
</taxonomicName>
except for females of
<taxonomicName id="6ECEF1E21A16FFF0FBCDF9E6019DFA59" authorityName="Konings &amp; Stauffer" authorityYear="2006" box="[1095,1258,1613,1637]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="96" phylum="Chordata" rank="species" species="flavifemina">
<emphasis id="9BBA56731A16FFF0FBCDF9E6019DFA59" box="[1095,1258,1613,1637]" italics="true" pageId="1" pageNumber="96">M. flavifemina</emphasis>
</taxonomicName>
and
<taxonomicName id="6ECEF1E21A16FFF0FAAAF9E600E3FA59" authorityName="Stauffer, Bowers, Kellogg &amp; McKaye" authorityYear="1997" box="[1312,1428,1613,1637]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="96" phylum="Chordata" rank="species" species="phaeos">
<emphasis id="9BBA56731A16FFF0FAAAF9E600E3FA59" box="[1312,1428,1613,1637]" italics="true" pageId="1" pageNumber="96">M. phaeos</emphasis>
</taxonomicName>
. The snout of the latter species is relatively longer than that of
<taxonomicName id="6ECEF1E21A16FFF0FCEAF9DA0691FAB5" authorityName="Stauffer" authorityYear="2018" box="[864,998,1649,1673]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="96" phylum="Chordata" rank="species" species="koningsi">
<emphasis id="9BBA56731A16FFF0FCEAF9DA0691FAB5" box="[864,998,1649,1673]" italics="true" pageId="1" pageNumber="96">M. koningsi</emphasis>
</taxonomicName>
(37.9–44.7% vs. 26.6–33.1% HL) but females of
<taxonomicName id="6ECEF1E21A16FFF0FE9FF93E04CDFA91" authorityName="Konings &amp; Stauffer" authorityYear="2006" box="[277,442,1685,1709]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="96" phylum="Chordata" rank="species" species="flavifemina">
<emphasis id="9BBA56731A16FFF0FE9FF93E04CDFA91" box="[277,442,1685,1709]" italics="true" pageId="1" pageNumber="96">M. flavifemina</emphasis>
</taxonomicName>
cannot reliably be distinguished from those of
<taxonomicName id="6ECEF1E21A16FFF0FC58F93E0121FA91" authorityName="Stauffer" authorityYear="2018" box="[978,1110,1685,1709]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="96" phylum="Chordata" rank="species" species="koningsi">
<emphasis id="9BBA56731A16FFF0FC58F93E0121FA91" box="[978,1110,1685,1709]" italics="true" pageId="1" pageNumber="96">M. koningsi</emphasis>
</taxonomicName>
apart from the fact that they have a larger size (maximum
<quantity id="6E3627841A16FFF0FE68F9120746FAED" box="[482,561,1722,1746]" metricMagnitude="-2" metricUnit="m" metricValue="7.9" pageId="1" pageNumber="96" unit="mm" value="79.0">79 mm</quantity>
SL vs.
<quantity id="6E3627841A16FFF0FD0DF91207A2FAED" box="[647,725,1722,1746]" metricMagnitude="-2" metricUnit="m" metricValue="4.9" pageId="1" pageNumber="96" unit="mm" value="49.0">49 mm</quantity>
SL for
<taxonomicName id="6ECEF1E21A16FFF0FCA1F91206C7FAED" authorityName="Stauffer" authorityYear="2018" box="[811,944,1721,1745]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="96" phylum="Chordata" rank="species" species="koningsi">
<emphasis id="9BBA56731A16FFF0FCA1F91206C7FAED" box="[811,944,1721,1745]" italics="true" pageId="1" pageNumber="96">M. koningsi</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="E1D4D9EA1A16FFF5FF4DF974075CF864" lastPageId="4" lastPageNumber="99" pageId="1" pageNumber="96" type="description">
<paragraph id="A9718A611A16FFF0FF4DF974067EFB02" blockId="1.[151,1437,1181,2034]" pageId="1" pageNumber="96">
<emphasis id="9BBA56731A16FFF0FF4DF974042EFAC9" bold="true" box="[199,345,1756,1781]" pageId="1" pageNumber="96">Description.</emphasis>
Morphometric and meristic data in
<tableCitation id="E44CBFDA1A16FFF0FD65F9750633FACA" box="[751,836,1757,1782]" captionStart="TABLE 1" captionStartId="2.[151,239,152,175]" captionText="TABLE 1. Morphometric and meristic values of Metriaclima koningsi (n=20). The mean, standard deviation and range include holotype and paratypes." pageId="1" pageNumber="96">Table 1</tableCitation>
. Dorsal fin with 17–18 spines and 7–9 rays. Anal fin with three spines (one specimen with two) and 7–8 rays. Lateral line with 30–32 ctenoid scales with 1–2 pored scales posterior to hypural plate. Caudal fin emarginate.
</paragraph>
<paragraph id="A9718A611A16FFF0FF4DF8E10771FBCE" blockId="1.[151,1437,1181,2034]" pageId="1" pageNumber="96">Head of males grey with two bright interorbital bars; cheek with gray markings; gular dark grey/black. Laterally light blue dorsally and dark blue/black ventrally. Dorsal fin light blue with dark submarginal band and light blue/white lappets. Caudal-fin membranes blue, rays pale orange. Anal fin black with 1–2 yellow ocelli; tips of rays light blue. Pelvic fins with blue leading edge; anterior membranes black fading to clear. Pectoral fins with black rays and clear membranes.</paragraph>
<caption id="FDB1DAE91A15FFF3FF1DFF30049CFCF3" pageId="2" pageNumber="97">
<paragraph id="A9718A611A15FFF3FF1DFF30049CFCF3" blockId="2.[151,1435,151,207]" pageId="2" pageNumber="97">
<emphasis id="9BBA56731A15FFF3FF1DFF30047CFC93" bold="true" box="[151,267,152,175]" pageId="2" pageNumber="97">TABLE 1.</emphasis>
Morphometric and meristic values of
<taxonomicName id="6ECEF1E21A15FFF3FD23FF3F06F9FC93" authorityName="Stauffer" authorityYear="2018" box="[681,910,151,175]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="97" phylum="Chordata" rank="species" species="koningsi">
<emphasis id="9BBA56731A15FFF3FD23FF3F06F9FC93" box="[681,910,151,175]" italics="true" pageId="2" pageNumber="97">Metriaclima koningsi</emphasis>
</taxonomicName>
(n=20). The mean, standard deviation and range include holotype and paratypes.
</paragraph>
</caption>
<paragraph id="A9718A611A15FFF3FF15FF42016FFBD5" pageId="2" pageNumber="97">
<table id="DBCE78C11A15000EFF15FF420033FBD5" box="[159,1348,234,2025]" gridcols="5" gridrows="43" pageId="2" pageNumber="97">
<tr id="17FE88231A15000EFF15FF420033FD3C" box="[159,1348,234,256]" gridrow="0" pageId="2" pageNumber="97">
<th id="542FE15F1A15000EFF15FF420773FD3C" box="[159,516,234,256]" gridcol="0" gridrow="0" pageId="2" pageNumber="97">Variable</th>
<th id="542FE15F1A15000EFD2AFF420789FD3C" box="[672,766,234,256]" gridcol="1" gridrow="0" pageId="2" pageNumber="97">Holotype</th>
<th id="542FE15F1A15000EFCC2FF4206F4FD3C" box="[840,899,234,256]" gridcol="2" gridrow="0" pageId="2" pageNumber="97">Mean</th>
<th id="542FE15F1A15000EFC78FF42010EFD3C" box="[1010,1145,234,256]" gridcol="3" gridrow="0" pageId="2" pageNumber="97">SD</th>
<th id="542FE15F1A15000EFB58FF420033FD3C" box="[1234,1348,234,256]" gridcol="4" gridrow="0" pageId="2" pageNumber="97">Range</th>
</tr>
<tr id="17FE88231A15000EFF15FEBE0033FD10" box="[159,1348,278,300]" gridrow="1" pageId="2" pageNumber="97">
<th id="542FE15F1A15000EFF15FEBE0773FD10" box="[159,516,278,300]" gridcol="0" gridrow="1" pageId="2" pageNumber="97">Standard length, mm</th>
<td id="542FE15F1A15000EFD2AFEBE0789FD10" box="[672,766,278,300]" gridcol="1" gridrow="1" pageId="2" pageNumber="97">56.8</td>
<td id="542FE15F1A15000EFCC2FEBE06F4FD10" box="[840,899,278,300]" gridcol="2" gridrow="1" pageId="2" pageNumber="97">50.7</td>
<td id="542FE15F1A15000EFC78FEBE010EFD10" box="[1010,1145,278,300]" gridcol="3" gridrow="1" pageId="2" pageNumber="97">4.9</td>
<td id="542FE15F1A15000EFB58FEBE0033FD10" box="[1234,1348,278,300]" gridcol="4" gridrow="1" pageId="2" pageNumber="97">40.3–56.9</td>
</tr>
<tr id="17FE88231A15000EFF15FEE80033FD6A" box="[159,1348,320,342]" gridrow="2" pageId="2" pageNumber="97">
<th id="542FE15F1A15000EFF15FEE80773FD6A" box="[159,516,320,342]" gridcol="0" gridrow="2" pageId="2" pageNumber="97">Head length, mm</th>
<td id="542FE15F1A15000EFD2AFEE80789FD6A" box="[672,766,320,342]" gridcol="1" gridrow="2" pageId="2" pageNumber="97">18.7</td>
<td id="542FE15F1A15000EFCC2FEE806F4FD6A" box="[840,899,320,342]" gridcol="2" gridrow="2" pageId="2" pageNumber="97">16.2</td>
<td id="542FE15F1A15000EFC78FEE8010EFD6A" box="[1010,1145,320,342]" gridcol="3" gridrow="2" pageId="2" pageNumber="97">1.6</td>
<td id="542FE15F1A15000EFB58FEE80033FD6A" box="[1234,1348,320,342]" gridcol="4" gridrow="2" pageId="2" pageNumber="97">13.2–18.7</td>
</tr>
<tr id="17FE88231A15000EFF15FEC10033FD43" box="[159,1348,361,383]" gridrow="3" pageId="2" pageNumber="97" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="542FE15F1A15000EFF15FEC10773FD43" box="[159,516,361,383]" gridcol="0" gridrow="3" pageId="2" pageNumber="97">Percent of standard length</th>
</tr>
<tr id="17FE88231A15000EFF15FE3A0033FD94" box="[159,1348,402,424]" gridrow="4" pageId="2" pageNumber="97">
<th id="542FE15F1A15000EFF15FE3A0773FD94" box="[159,516,402,424]" gridcol="0" gridrow="4" pageId="2" pageNumber="97">Head length</th>
<td id="542FE15F1A15000EFD2AFE3A0789FD94" box="[672,766,402,424]" gridcol="1" gridrow="4" pageId="2" pageNumber="97">32.9</td>
<td id="542FE15F1A15000EFCC2FE3A06F4FD94" box="[840,899,402,424]" gridcol="2" gridrow="4" pageId="2" pageNumber="97">32.4</td>
<td id="542FE15F1A15000EFC78FE3A010EFD94" box="[1010,1145,402,424]" gridcol="3" gridrow="4" pageId="2" pageNumber="97">0.9</td>
<td id="542FE15F1A15000EFB58FE3A0033FD94" box="[1234,1348,402,424]" gridcol="4" gridrow="4" pageId="2" pageNumber="97">30.2–34.1</td>
</tr>
<tr id="17FE88231A15000EFF15FE140033FDEE" box="[159,1348,444,466]" gridrow="5" pageId="2" pageNumber="97">
<th id="542FE15F1A15000EFF15FE140773FDEE" box="[159,516,444,466]" gridcol="0" gridrow="5" pageId="2" pageNumber="97">Body depth</th>
<td id="542FE15F1A15000EFD2AFE140789FDEE" box="[672,766,444,466]" gridcol="1" gridrow="5" pageId="2" pageNumber="97">34.9</td>
<td id="542FE15F1A15000EFCC2FE1406F4FDEE" box="[840,899,444,466]" gridcol="2" gridrow="5" pageId="2" pageNumber="97">33.5</td>
<td id="542FE15F1A15000EFC78FE14010EFDEE" box="[1010,1145,444,466]" gridcol="3" gridrow="5" pageId="2" pageNumber="97">1.5</td>
<td id="542FE15F1A15000EFB58FE140033FDEE" box="[1234,1348,444,466]" gridcol="4" gridrow="5" pageId="2" pageNumber="97">31.3–37.1</td>
</tr>
<tr id="17FE88231A15000EFF15FE400033FDC2" box="[159,1348,488,510]" gridrow="6" pageId="2" pageNumber="97">
<th id="542FE15F1A15000EFF15FE400773FDC2" box="[159,516,488,510]" gridcol="0" gridrow="6" pageId="2" pageNumber="97">Snout to dorsal-fin origin</th>
<td id="542FE15F1A15000EFD2AFE400789FDC2" box="[672,766,488,510]" gridcol="1" gridrow="6" pageId="2" pageNumber="97">34.9</td>
<td id="542FE15F1A15000EFCC2FE4006F4FDC2" box="[840,899,488,510]" gridcol="2" gridrow="6" pageId="2" pageNumber="97">35.6</td>
<td id="542FE15F1A15000EFC78FE40010EFDC2" box="[1010,1145,488,510]" gridcol="3" gridrow="6" pageId="2" pageNumber="97">1.1</td>
<td id="542FE15F1A15000EFB58FE400033FDC2" box="[1234,1348,488,510]" gridcol="4" gridrow="6" pageId="2" pageNumber="97">33.3–37.8</td>
</tr>
<tr id="17FE88231A15000EFF15FDB90033FE1B" box="[159,1348,529,551]" gridrow="7" pageId="2" pageNumber="97">
<th id="542FE15F1A15000EFF15FDB90773FE1B" box="[159,516,529,551]" gridcol="0" gridrow="7" pageId="2" pageNumber="97">Snout to pelvic-fin origin</th>
<td id="542FE15F1A15000EFD2AFDB90789FE1B" box="[672,766,529,551]" gridcol="1" gridrow="7" pageId="2" pageNumber="97">36.5</td>
<td id="542FE15F1A15000EFCC2FDB906F4FE1B" box="[840,899,529,551]" gridcol="2" gridrow="7" pageId="2" pageNumber="97">37.2</td>
<td id="542FE15F1A15000EFC78FDB9010EFE1B" box="[1010,1145,529,551]" gridcol="3" gridrow="7" pageId="2" pageNumber="97">1.5</td>
<td id="542FE15F1A15000EFB58FDB90033FE1B" box="[1234,1348,529,551]" gridcol="4" gridrow="7" pageId="2" pageNumber="97">33.9–39.7</td>
</tr>
<tr id="17FE88231A15000EFF15FD930033FE6D" box="[159,1348,571,593]" gridrow="8" pageId="2" pageNumber="97">
<th id="542FE15F1A15000EFF15FD930773FE6D" box="[159,516,571,593]" gridcol="0" gridrow="8" pageId="2" pageNumber="97">Dorsal fin base length</th>
<td id="542FE15F1A15000EFD2AFD930789FE6D" box="[672,766,571,593]" gridcol="1" gridrow="8" pageId="2" pageNumber="97">61.4</td>
<td id="542FE15F1A15000EFCC2FD9306F4FE6D" box="[840,899,571,593]" gridcol="2" gridrow="8" pageId="2" pageNumber="97">61.4</td>
<td id="542FE15F1A15000EFC78FD93010EFE6D" box="[1010,1145,571,593]" gridcol="3" gridrow="8" pageId="2" pageNumber="97">1.2</td>
<td id="542FE15F1A15000EFB58FD930033FE6D" box="[1234,1348,571,593]" gridcol="4" gridrow="8" pageId="2" pageNumber="97">58.4–63.2</td>
</tr>
<tr id="17FE88231A15000EFF15FDCC0033FE46" box="[159,1348,612,634]" gridrow="9" pageId="2" pageNumber="97">
<th id="542FE15F1A15000EFF15FDCC0773FE46" box="[159,516,612,634]" gridcol="0" gridrow="9" pageId="2" pageNumber="97">Anterior dorsal to anterior anal</th>
<td id="542FE15F1A15000EFD2AFDCC0789FE46" box="[672,766,612,634]" gridcol="1" gridrow="9" pageId="2" pageNumber="97">51.5</td>
<td id="542FE15F1A15000EFCC2FDCC06F4FE46" box="[840,899,612,634]" gridcol="2" gridrow="9" pageId="2" pageNumber="97">51.4</td>
<td id="542FE15F1A15000EFC78FDCC010EFE46" box="[1010,1145,612,634]" gridcol="3" gridrow="9" pageId="2" pageNumber="97">1.5</td>
<td id="542FE15F1A15000EFB58FDCC0033FE46" box="[1234,1348,612,634]" gridcol="4" gridrow="9" pageId="2" pageNumber="97">48.2–54.1</td>
</tr>
<tr id="17FE88231A15000EFF15FD250033FE9F" box="[159,1348,653,675]" gridrow="10" pageId="2" pageNumber="97">
<th id="542FE15F1A15000EFF15FD250773FE9F" box="[159,516,653,675]" gridcol="0" gridrow="10" pageId="2" pageNumber="97">Anterior dorsal to posterior anal</th>
<td id="542FE15F1A15000EFD2AFD250789FE9F" box="[672,766,653,675]" gridcol="1" gridrow="10" pageId="2" pageNumber="97">62.7</td>
<td id="542FE15F1A15000EFCC2FD2506F4FE9F" box="[840,899,653,675]" gridcol="2" gridrow="10" pageId="2" pageNumber="97">63.7</td>
<td id="542FE15F1A15000EFC78FD25010EFE9F" box="[1010,1145,653,675]" gridcol="3" gridrow="10" pageId="2" pageNumber="97">1.3</td>
<td id="542FE15F1A15000EFB58FD250033FE9F" box="[1234,1348,653,675]" gridcol="4" gridrow="10" pageId="2" pageNumber="97">61.3–66.8</td>
</tr>
<tr id="17FE88231A15000EFF15FD1F0033FEF1" box="[159,1348,695,717]" gridrow="11" pageId="2" pageNumber="97">
<th id="542FE15F1A15000EFF15FD1F0773FEF1" box="[159,516,695,717]" gridcol="0" gridrow="11" pageId="2" pageNumber="97">Posterior dorsal to anterior anal</th>
<td id="542FE15F1A15000EFD2AFD1F0789FEF1" box="[672,766,695,717]" gridcol="1" gridrow="11" pageId="2" pageNumber="97">30.2</td>
<td id="542FE15F1A15000EFCC2FD1F06F4FEF1" box="[840,899,695,717]" gridcol="2" gridrow="11" pageId="2" pageNumber="97">30.6</td>
<td id="542FE15F1A15000EFC78FD1F010EFEF1" box="[1010,1145,695,717]" gridcol="3" gridrow="11" pageId="2" pageNumber="97">1.3</td>
<td id="542FE15F1A15000EFB58FD1F0033FEF1" box="[1234,1348,695,717]" gridcol="4" gridrow="11" pageId="2" pageNumber="97">27.0–32.2</td>
</tr>
<tr id="17FE88231A15000EFF15FD4F0033FEC1" box="[159,1348,743,765]" gridrow="12" pageId="2" pageNumber="97">
<th id="542FE15F1A15000EFF15FD4F0773FEC1" box="[159,516,743,765]" gridcol="0" gridrow="12" pageId="2" pageNumber="97">Posterior dorsal to posterior anal</th>
<td id="542FE15F1A15000EFD2AFD4F0789FEC1" box="[672,766,743,765]" gridcol="1" gridrow="12" pageId="2" pageNumber="97">15.2</td>
<td id="542FE15F1A15000EFCC2FD4F06F4FEC1" box="[840,899,743,765]" gridcol="2" gridrow="12" pageId="2" pageNumber="97">15.8</td>
<td id="542FE15F1A15000EFC78FD4F010EFEC1" box="[1010,1145,743,765]" gridcol="3" gridrow="12" pageId="2" pageNumber="97">1.0</td>
<td id="542FE15F1A15000EFB58FD4F0033FEC1" box="[1234,1348,743,765]" gridcol="4" gridrow="12" pageId="2" pageNumber="97">13.4–17.5</td>
</tr>
<tr id="17FE88231A15000EFF15FCB90033FF1B" box="[159,1348,785,807]" gridrow="13" pageId="2" pageNumber="97">
<th id="542FE15F1A15000EFF15FCB90773FF1B" box="[159,516,785,807]" gridcol="0" gridrow="13" pageId="2" pageNumber="97">Posterior dorsal ventral caudal</th>
<td id="542FE15F1A15000EFD2AFCB90789FF1B" box="[672,766,785,807]" gridcol="1" gridrow="13" pageId="2" pageNumber="97">18.3</td>
<td id="542FE15F1A15000EFCC2FCB906F4FF1B" box="[840,899,785,807]" gridcol="2" gridrow="13" pageId="2" pageNumber="97">17.5</td>
<td id="542FE15F1A15000EFC78FCB9010EFF1B" box="[1010,1145,785,807]" gridcol="3" gridrow="13" pageId="2" pageNumber="97">0.9</td>
<td id="542FE15F1A15000EFB58FCB90033FF1B" box="[1234,1348,785,807]" gridcol="4" gridrow="13" pageId="2" pageNumber="97">16.0–19.8</td>
</tr>
<tr id="17FE88231A15000EFF15FCE90033FF6B" box="[159,1348,833,855]" gridrow="14" pageId="2" pageNumber="97">
<th id="542FE15F1A15000EFF15FCE90773FF6B" box="[159,516,833,855]" gridcol="0" gridrow="14" pageId="2" pageNumber="97">Posterior anal to dorsal caudal</th>
<td id="542FE15F1A15000EFD2AFCE90789FF6B" box="[672,766,833,855]" gridcol="1" gridrow="14" pageId="2" pageNumber="97">20.5</td>
<td id="542FE15F1A15000EFCC2FCE906F4FF6B" box="[840,899,833,855]" gridcol="2" gridrow="14" pageId="2" pageNumber="97">20.1</td>
<td id="542FE15F1A15000EFC78FCE9010EFF6B" box="[1010,1145,833,855]" gridcol="3" gridrow="14" pageId="2" pageNumber="97">0.7</td>
<td id="542FE15F1A15000EFB58FCE90033FF6B" box="[1234,1348,833,855]" gridcol="4" gridrow="14" pageId="2" pageNumber="97">18.8–21.3</td>
</tr>
<tr id="17FE88231A15000EFF15FCC20033FFBC" box="[159,1348,874,896]" gridrow="15" pageId="2" pageNumber="97">
<th id="542FE15F1A15000EFF15FCC20773FFBC" box="[159,516,874,896]" gridcol="0" gridrow="15" pageId="2" pageNumber="97">Anterior dorsal to pelvic-fin origin</th>
<td id="542FE15F1A15000EFD2AFCC20789FFBC" box="[672,766,874,896]" gridcol="1" gridrow="15" pageId="2" pageNumber="97">36.0</td>
<td id="542FE15F1A15000EFCC2FCC206F4FFBC" box="[840,899,874,896]" gridcol="2" gridrow="15" pageId="2" pageNumber="97">34.7</td>
<td id="542FE15F1A15000EFC78FCC2010EFFBC" box="[1010,1145,874,896]" gridcol="3" gridrow="15" pageId="2" pageNumber="97">1.7</td>
<td id="542FE15F1A15000EFB58FCC20033FFBC" box="[1234,1348,874,896]" gridcol="4" gridrow="15" pageId="2" pageNumber="97">31.4–37.9</td>
</tr>
<tr id="17FE88231A15000EFF15FC3B0033FF95" box="[159,1348,915,937]" gridrow="16" pageId="2" pageNumber="97">
<th id="542FE15F1A15000EFF15FC3B0773FF95" box="[159,516,915,937]" gridcol="0" gridrow="16" pageId="2" pageNumber="97">Posterior dorsal to pelvic-fin origin</th>
<td id="542FE15F1A15000EFD2AFC3B0789FF95" box="[672,766,915,937]" gridcol="1" gridrow="16" pageId="2" pageNumber="97">57.3</td>
<td id="542FE15F1A15000EFCC2FC3B06F4FF95" box="[840,899,915,937]" gridcol="2" gridrow="16" pageId="2" pageNumber="97">58.4</td>
<td id="542FE15F1A15000EFC78FC3B010EFF95" box="[1010,1145,915,937]" gridcol="3" gridrow="16" pageId="2" pageNumber="97">1.5</td>
<td id="542FE15F1A15000EFB58FC3B0033FF95" box="[1234,1348,915,937]" gridcol="4" gridrow="16" pageId="2" pageNumber="97">54.6–61.3</td>
</tr>
<tr id="17FE88231A15000EFF15FC6B0033FFE5" box="[159,1348,963,985]" gridrow="17" pageId="2" pageNumber="97">
<th id="542FE15F1A15000EFF15FC6B0773FFE5" box="[159,516,963,985]" gridcol="0" gridrow="17" pageId="2" pageNumber="97">Caudal peduncle length</th>
<td id="542FE15F1A15000EFD2AFC6B0789FFE5" box="[672,766,963,985]" gridcol="1" gridrow="17" pageId="2" pageNumber="97">15.0</td>
<td id="542FE15F1A15000EFCC2FC6B06F4FFE5" box="[840,899,963,985]" gridcol="2" gridrow="17" pageId="2" pageNumber="97">16.0</td>
<td id="542FE15F1A15000EFC78FC6B010EFFE5" box="[1010,1145,963,985]" gridcol="3" gridrow="17" pageId="2" pageNumber="97">0.8</td>
<td id="542FE15F1A15000EFB58FC6B0033FFE5" box="[1234,1348,963,985]" gridcol="4" gridrow="17" pageId="2" pageNumber="97">15.0–17.0</td>
</tr>
<tr id="17FE88231A15000EFF15FC450033F83F" box="[159,1348,1005,1027]" gridrow="18" pageId="2" pageNumber="97">
<th id="542FE15F1A15000EFF15FC450773F83F" box="[159,516,1005,1027]" gridcol="0" gridrow="18" pageId="2" pageNumber="97">Least caudal peduncle depth</th>
<td id="542FE15F1A15000EFD2AFC450789F83F" box="[672,766,1005,1027]" gridcol="1" gridrow="18" pageId="2" pageNumber="97">11.7</td>
<td id="542FE15F1A15000EFCC2FC4506F4F83F" box="[840,899,1005,1027]" gridcol="2" gridrow="18" pageId="2" pageNumber="97">11.8</td>
<td id="542FE15F1A15000EFC78FC45010EF83F" box="[1010,1145,1005,1027]" gridcol="3" gridrow="18" pageId="2" pageNumber="97">0.7</td>
<td id="542FE15F1A15000EFB58FC450033F83F" box="[1234,1348,1005,1027]" gridcol="4" gridrow="18" pageId="2" pageNumber="97">10.4–13.3</td>
</tr>
<tr id="17FE88231A15000EFF15FBBF0033F811" box="[159,1348,1047,1069]" gridrow="19" pageId="2" pageNumber="97" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="542FE15F1A15000EFF15FBBF0773F811" box="[159,516,1047,1069]" gridcol="0" gridrow="19" pageId="2" pageNumber="97">Percent of head length</th>
</tr>
<tr id="17FE88231A15000EFF15FBE80033F86A" box="[159,1348,1088,1110]" gridrow="20" pageId="2" pageNumber="97">
<th id="542FE15F1A15000EFF15FBE80773F86A" box="[159,516,1088,1110]" gridcol="0" gridrow="20" pageId="2" pageNumber="97">Snout length</th>
<td id="542FE15F1A15000EFD2AFBE80789F86A" box="[672,766,1088,1110]" gridcol="1" gridrow="20" pageId="2" pageNumber="97">26.6</td>
<td id="542FE15F1A15000EFCC2FBE806F4F86A" box="[840,899,1088,1110]" gridcol="2" gridrow="20" pageId="2" pageNumber="97">29.8</td>
<td id="542FE15F1A15000EFC78FBE8010EF86A" box="[1010,1145,1088,1110]" gridcol="3" gridrow="20" pageId="2" pageNumber="97">1.8</td>
<td id="542FE15F1A15000EFB58FBE80033F86A" box="[1234,1348,1088,1110]" gridcol="4" gridrow="20" pageId="2" pageNumber="97">26.6–33.1</td>
</tr>
<tr id="17FE88231A15000EFF15FBC10033F843" box="[159,1348,1129,1151]" gridrow="21" pageId="2" pageNumber="97">
<th id="542FE15F1A15000EFF15FBC10773F843" box="[159,516,1129,1151]" gridcol="0" gridrow="21" pageId="2" pageNumber="97">Postorbital head length</th>
<td id="542FE15F1A15000EFD2AFBC10789F843" box="[672,766,1129,1151]" gridcol="1" gridrow="21" pageId="2" pageNumber="97">42.4</td>
<td id="542FE15F1A15000EFCC2FBC106F4F843" box="[840,899,1129,1151]" gridcol="2" gridrow="21" pageId="2" pageNumber="97">40.1</td>
<td id="542FE15F1A15000EFC78FBC1010EF843" box="[1010,1145,1129,1151]" gridcol="3" gridrow="21" pageId="2" pageNumber="97">1.9</td>
<td id="542FE15F1A15000EFB58FBC10033F843" box="[1234,1348,1129,1151]" gridcol="4" gridrow="21" pageId="2" pageNumber="97">37.6–44.7</td>
</tr>
<tr id="17FE88231A15000EFF15FB3B0033F895" box="[159,1348,1171,1193]" gridrow="22" pageId="2" pageNumber="97">
<th id="542FE15F1A15000EFF15FB3B0773F895" box="[159,516,1171,1193]" gridcol="0" gridrow="22" pageId="2" pageNumber="97">Horizontal eye diameter</th>
<td id="542FE15F1A15000EFD2AFB3B0789F895" box="[672,766,1171,1193]" gridcol="1" gridrow="22" pageId="2" pageNumber="97">32.7</td>
<td id="542FE15F1A15000EFCC2FB3B06F4F895" box="[840,899,1171,1193]" gridcol="2" gridrow="22" pageId="2" pageNumber="97">33.1</td>
<td id="542FE15F1A15000EFC78FB3B010EF895" box="[1010,1145,1171,1193]" gridcol="3" gridrow="22" pageId="2" pageNumber="97">1.7</td>
<td id="542FE15F1A15000EFB58FB3B0033F895" box="[1234,1348,1171,1193]" gridcol="4" gridrow="22" pageId="2" pageNumber="97">28.6–35.6</td>
</tr>
<tr id="17FE88231A15000EFF15FB680033F8EA" box="[159,1348,1216,1238]" gridrow="23" pageId="2" pageNumber="97">
<th id="542FE15F1A15000EFF15FB680773F8EA" box="[159,516,1216,1238]" gridcol="0" gridrow="23" pageId="2" pageNumber="97">Vertical eye diameter</th>
<td id="542FE15F1A15000EFD2AFB680789F8EA" box="[672,766,1216,1238]" gridcol="1" gridrow="23" pageId="2" pageNumber="97">29.0</td>
<td id="542FE15F1A15000EFCC2FB6806F4F8EA" box="[840,899,1216,1238]" gridcol="2" gridrow="23" pageId="2" pageNumber="97">31.4</td>
<td id="542FE15F1A15000EFC78FB68010EF8EA" box="[1010,1145,1216,1238]" gridcol="3" gridrow="23" pageId="2" pageNumber="97">2.2</td>
<td id="542FE15F1A15000EFB58FB680033F8EA" box="[1234,1348,1216,1238]" gridcol="4" gridrow="23" pageId="2" pageNumber="97">26.7–34.3</td>
</tr>
<tr id="17FE88231A15000EFF15FB420033F93C" box="[159,1348,1258,1280]" gridrow="24" pageId="2" pageNumber="97">
<th id="542FE15F1A15000EFF15FB420773F93C" box="[159,516,1258,1280]" gridcol="0" gridrow="24" pageId="2" pageNumber="97">Pre-orbital depth</th>
<td id="542FE15F1A15000EFD2AFB420789F93C" box="[672,766,1258,1280]" gridcol="1" gridrow="24" pageId="2" pageNumber="97">17.2</td>
<td id="542FE15F1A15000EFCC2FB4206F4F93C" box="[840,899,1258,1280]" gridcol="2" gridrow="24" pageId="2" pageNumber="97">18.5</td>
<td id="542FE15F1A15000EFC78FB42010EF93C" box="[1010,1145,1258,1280]" gridcol="3" gridrow="24" pageId="2" pageNumber="97">1.8</td>
<td id="542FE15F1A15000EFB58FB420033F93C" box="[1234,1348,1258,1280]" gridcol="4" gridrow="24" pageId="2" pageNumber="97">15.8–23.4</td>
</tr>
<tr id="17FE88231A15000EFF15FABB0033F915" box="[159,1348,1299,1321]" gridrow="25" pageId="2" pageNumber="97">
<th id="542FE15F1A15000EFF15FABB0773F915" box="[159,516,1299,1321]" gridcol="0" gridrow="25" pageId="2" pageNumber="97">Cheek depth</th>
<td id="542FE15F1A15000EFD2AFABB0789F915" box="[672,766,1299,1321]" gridcol="1" gridrow="25" pageId="2" pageNumber="97">26.8</td>
<td id="542FE15F1A15000EFCC2FABB06F4F915" box="[840,899,1299,1321]" gridcol="2" gridrow="25" pageId="2" pageNumber="97">25.1</td>
<td id="542FE15F1A15000EFC78FABB010EF915" box="[1010,1145,1299,1321]" gridcol="3" gridrow="25" pageId="2" pageNumber="97">2.3</td>
<td id="542FE15F1A15000EFB58FABB0033F915" box="[1234,1348,1299,1321]" gridcol="4" gridrow="25" pageId="2" pageNumber="97">19.7–28.4</td>
</tr>
<tr id="17FE88231A15000EFF15FA940033F96E" box="[159,1348,1340,1362]" gridrow="26" pageId="2" pageNumber="97">
<th id="542FE15F1A15000EFF15FA940773F96E" box="[159,516,1340,1362]" gridcol="0" gridrow="26" pageId="2" pageNumber="97">Lower jaw length</th>
<td id="542FE15F1A15000EFD2AFA940789F96E" box="[672,766,1340,1362]" gridcol="1" gridrow="26" pageId="2" pageNumber="97">38.0</td>
<td id="542FE15F1A15000EFCC2FA9406F4F96E" box="[840,899,1340,1362]" gridcol="2" gridrow="26" pageId="2" pageNumber="97">39.7</td>
<td id="542FE15F1A15000EFC78FA94010EF96E" box="[1010,1145,1340,1362]" gridcol="3" gridrow="26" pageId="2" pageNumber="97">2.6</td>
<td id="542FE15F1A15000EFB58FA940033F96E" box="[1234,1348,1340,1362]" gridcol="4" gridrow="26" pageId="2" pageNumber="97">34.4–43.3</td>
</tr>
<tr id="17FE88231A15000EFF15FACE0033F940" box="[159,1348,1382,1404]" gridrow="27" pageId="2" pageNumber="97">
<th id="542FE15F1A15000EFF15FACE0773F940" box="[159,516,1382,1404]" gridcol="0" gridrow="27" pageId="2" pageNumber="97">Head depth</th>
<td id="542FE15F1A15000EFD2AFACE0789F940" box="[672,766,1382,1404]" gridcol="1" gridrow="27" pageId="2" pageNumber="97">88.8</td>
<td id="542FE15F1A15000EFCC2FACE06F4F940" box="[840,899,1382,1404]" gridcol="2" gridrow="27" pageId="2" pageNumber="97">90.3</td>
<td id="542FE15F1A15000EFC78FACE010EF940" box="[1010,1145,1382,1404]" gridcol="3" gridrow="27" pageId="2" pageNumber="97">5.0</td>
<td id="542FE15F1A15000EFB58FACE0033F940" box="[1234,1348,1382,1404]" gridcol="4" gridrow="27" pageId="2" pageNumber="97">82.2–103.3</td>
</tr>
<tr id="17FE88231A15000EFF15FA380033F99A" box="[159,1348,1424,1446]" gridrow="28" pageId="2" pageNumber="97" rowspan-1="1">
<th id="542FE15F1A15000EFF15FA380773F99A" box="[159,516,1424,1446]" gridcol="0" gridrow="28" pageId="2" pageNumber="97">Counts</th>
<td id="542FE15F1A15000EFCC2FA3806F4F99A" box="[840,899,1424,1446]" gridcol="2" gridrow="28" pageId="2" pageNumber="97">Mode</td>
<td id="542FE15F1A15000EFC78FA38010EF99A" box="[1010,1145,1424,1446]" gridcol="3" gridrow="28" pageId="2" pageNumber="97">Frequency %</td>
<td id="542FE15F1A15000EFB58FA380033F99A" box="[1234,1348,1424,1446]" gridcol="4" gridrow="28" pageId="2" pageNumber="97">Range</td>
</tr>
<tr id="17FE88231A15000EFF15FA110033F9F3" box="[159,1348,1465,1487]" gridrow="29" pageId="2" pageNumber="97">
<th id="542FE15F1A15000EFF15FA110773F9F3" box="[159,516,1465,1487]" gridcol="0" gridrow="29" pageId="2" pageNumber="97">Dorsal-fin spines</th>
<td id="542FE15F1A15000EFD2AFA110789F9F3" box="[672,766,1465,1487]" gridcol="1" gridrow="29" pageId="2" pageNumber="97">18</td>
<td id="542FE15F1A15000EFCC2FA1106F4F9F3" box="[840,899,1465,1487]" gridcol="2" gridrow="29" pageId="2" pageNumber="97">17</td>
<td id="542FE15F1A15000EFC78FA11010EF9F3" box="[1010,1145,1465,1487]" gridcol="3" gridrow="29" pageId="2" pageNumber="97">60</td>
<td id="542FE15F1A15000EFB58FA110033F9F3" box="[1234,1348,1465,1487]" gridcol="4" gridrow="29" pageId="2" pageNumber="97">17–18</td>
</tr>
<tr id="17FE88231A15000EFF15FA4A0033F9C4" box="[159,1348,1506,1528]" gridrow="30" pageId="2" pageNumber="97">
<th id="542FE15F1A15000EFF15FA4A0773F9C4" box="[159,516,1506,1528]" gridcol="0" gridrow="30" pageId="2" pageNumber="97">Dorsal-fin rays</th>
<td id="542FE15F1A15000EFD2AFA4A0789F9C4" box="[672,766,1506,1528]" gridcol="1" gridrow="30" pageId="2" pageNumber="97">7</td>
<td id="542FE15F1A15000EFCC2FA4A06F4F9C4" box="[840,899,1506,1528]" gridcol="2" gridrow="30" pageId="2" pageNumber="97">8</td>
<td id="542FE15F1A15000EFC78FA4A010EF9C4" box="[1010,1145,1506,1528]" gridcol="3" gridrow="30" pageId="2" pageNumber="97">60</td>
<td id="542FE15F1A15000EFB58FA4A0033F9C4" box="[1234,1348,1506,1528]" gridcol="4" gridrow="30" pageId="2" pageNumber="97">7–9</td>
</tr>
<tr id="17FE88231A15000EFF15F9A40033FA1E" box="[159,1348,1548,1570]" gridrow="31" pageId="2" pageNumber="97">
<th id="542FE15F1A15000EFF15F9A40773FA1E" box="[159,516,1548,1570]" gridcol="0" gridrow="31" pageId="2" pageNumber="97">Anal-fin spines</th>
<td id="542FE15F1A15000EFD2AF9A40789FA1E" box="[672,766,1548,1570]" gridcol="1" gridrow="31" pageId="2" pageNumber="97">3</td>
<td id="542FE15F1A15000EFCC2F9A406F4FA1E" box="[840,899,1548,1570]" gridcol="2" gridrow="31" pageId="2" pageNumber="97">3</td>
<td id="542FE15F1A15000EFC78F9A4010EFA1E" box="[1010,1145,1548,1570]" gridcol="3" gridrow="31" pageId="2" pageNumber="97">95</td>
<td id="542FE15F1A15000EFB58F9A40033FA1E" box="[1234,1348,1548,1570]" gridcol="4" gridrow="31" pageId="2" pageNumber="97">2–3</td>
</tr>
<tr id="17FE88231A15000EFF15F99E0033FA70" box="[159,1348,1590,1612]" gridrow="32" pageId="2" pageNumber="97">
<th id="542FE15F1A15000EFF15F99E0773FA70" box="[159,516,1590,1612]" gridcol="0" gridrow="32" pageId="2" pageNumber="97">Anal-fin rays</th>
<td id="542FE15F1A15000EFD2AF99E0789FA70" box="[672,766,1590,1612]" gridcol="1" gridrow="32" pageId="2" pageNumber="97">7</td>
<td id="542FE15F1A15000EFCC2F99E06F4FA70" box="[840,899,1590,1612]" gridcol="2" gridrow="32" pageId="2" pageNumber="97">8</td>
<td id="542FE15F1A15000EFC78F99E010EFA70" box="[1010,1145,1590,1612]" gridcol="3" gridrow="32" pageId="2" pageNumber="97">75</td>
<td id="542FE15F1A15000EFB58F99E0033FA70" box="[1234,1348,1590,1612]" gridcol="4" gridrow="32" pageId="2" pageNumber="97">7–8</td>
</tr>
<tr id="17FE88231A15000EFF15F9F70033FA49" box="[159,1348,1631,1653]" gridrow="33" pageId="2" pageNumber="97">
<th id="542FE15F1A15000EFF15F9F70773FA49" box="[159,516,1631,1653]" gridcol="0" gridrow="33" pageId="2" pageNumber="97">Pelvic-fin rays</th>
<td id="542FE15F1A15000EFD2AF9F70789FA49" box="[672,766,1631,1653]" gridcol="1" gridrow="33" pageId="2" pageNumber="97">13</td>
<td id="542FE15F1A15000EFCC2F9F706F4FA49" box="[840,899,1631,1653]" gridcol="2" gridrow="33" pageId="2" pageNumber="97">13</td>
<td id="542FE15F1A15000EFC78F9F7010EFA49" box="[1010,1145,1631,1653]" gridcol="3" gridrow="33" pageId="2" pageNumber="97">95</td>
<td id="542FE15F1A15000EFB58F9F70033FA49" box="[1234,1348,1631,1653]" gridcol="4" gridrow="33" pageId="2" pageNumber="97">13–14</td>
</tr>
<tr id="17FE88231A15000EFF15F9200033FAA2" box="[159,1348,1672,1694]" gridrow="34" pageId="2" pageNumber="97" rowspan-4="1">
<th id="542FE15F1A15000EFF15F9200773FAA2" box="[159,516,1672,1694]" gridcol="0" gridrow="34" pageId="2" pageNumber="97">Pectoral-fin rays</th>
<td id="542FE15F1A15000EFD2AF9200789FAA2" box="[672,766,1672,1694]" gridcol="1" gridrow="34" pageId="2" pageNumber="97">5</td>
<td id="542FE15F1A15000EFCC2F92006F4FAA2" box="[840,899,1672,1694]" gridcol="2" gridrow="34" pageId="2" pageNumber="97">5</td>
<td id="542FE15F1A15000EFC78F920010EFAA2" box="[1010,1145,1672,1694]" gridcol="3" gridrow="34" pageId="2" pageNumber="97">100</td>
</tr>
<tr id="17FE88231A15000EFF15F91A0033FAF4" box="[159,1348,1714,1736]" gridrow="35" pageId="2" pageNumber="97">
<th id="542FE15F1A15000EFF15F91A0773FAF4" box="[159,516,1714,1736]" gridcol="0" gridrow="35" pageId="2" pageNumber="97">Lateral line scales</th>
<td id="542FE15F1A15000EFD2AF91A0789FAF4" box="[672,766,1714,1736]" gridcol="1" gridrow="35" pageId="2" pageNumber="97">30</td>
<td id="542FE15F1A15000EFCC2F91A06F4FAF4" box="[840,899,1714,1736]" gridcol="2" gridrow="35" pageId="2" pageNumber="97">31</td>
<td id="542FE15F1A15000EFC78F91A010EFAF4" box="[1010,1145,1714,1736]" gridcol="3" gridrow="35" pageId="2" pageNumber="97">50</td>
<td id="542FE15F1A15000EFB58F91A0033FAF4" box="[1234,1348,1714,1736]" gridcol="4" gridrow="35" pageId="2" pageNumber="97">30–32</td>
</tr>
<tr id="17FE88231A15000EFF15F9730033FACD" box="[159,1348,1755,1777]" gridrow="36" pageId="2" pageNumber="97">
<th id="542FE15F1A15000EFF15F9730773FACD" box="[159,516,1755,1777]" gridcol="0" gridrow="36" pageId="2" pageNumber="97">Pored scales posterior to LL</th>
<td id="542FE15F1A15000EFD2AF9730789FACD" box="[672,766,1755,1777]" gridcol="1" gridrow="36" pageId="2" pageNumber="97">2</td>
<td id="542FE15F1A15000EFCC2F97306F4FACD" box="[840,899,1755,1777]" gridcol="2" gridrow="36" pageId="2" pageNumber="97">1/2</td>
<td id="542FE15F1A15000EFC78F973010EFACD" box="[1010,1145,1755,1777]" gridcol="3" gridrow="36" pageId="2" pageNumber="97">50</td>
<td id="542FE15F1A15000EFB58F9730033FACD" box="[1234,1348,1755,1777]" gridcol="4" gridrow="36" pageId="2" pageNumber="97">1–2</td>
</tr>
<tr id="17FE88231A15000EFF15F8AC0033FB26" box="[159,1348,1796,1818]" gridrow="37" pageId="2" pageNumber="97">
<th id="542FE15F1A15000EFF15F8AC0773FB26" box="[159,516,1796,1818]" gridcol="0" gridrow="37" pageId="2" pageNumber="97">Cheek scale rows</th>
<td id="542FE15F1A15000EFD2AF8AC0789FB26" box="[672,766,1796,1818]" gridcol="1" gridrow="37" pageId="2" pageNumber="97">4</td>
<td id="542FE15F1A15000EFCC2F8AC06F4FB26" box="[840,899,1796,1818]" gridcol="2" gridrow="37" pageId="2" pageNumber="97">4</td>
<td id="542FE15F1A15000EFC78F8AC010EFB26" box="[1010,1145,1796,1818]" gridcol="3" gridrow="37" pageId="2" pageNumber="97">90</td>
<td id="542FE15F1A15000EFB58F8AC0033FB26" box="[1234,1348,1796,1818]" gridcol="4" gridrow="37" pageId="2" pageNumber="97">3–4</td>
</tr>
<tr id="17FE88231A15000EFF15F8860033FB78" box="[159,1348,1838,1860]" gridrow="38" pageId="2" pageNumber="97">
<th id="542FE15F1A15000EFF15F8860773FB78" box="[159,516,1838,1860]" gridcol="0" gridrow="38" pageId="2" pageNumber="97">Gill rakers on first epibranchial</th>
<td id="542FE15F1A15000EFD2AF8860789FB78" box="[672,766,1838,1860]" gridcol="1" gridrow="38" pageId="2" pageNumber="97">2</td>
<td id="542FE15F1A15000EFCC2F88606F4FB78" box="[840,899,1838,1860]" gridcol="2" gridrow="38" pageId="2" pageNumber="97">2/3</td>
<td id="542FE15F1A15000EFC78F886010EFB78" box="[1010,1145,1838,1860]" gridcol="3" gridrow="38" pageId="2" pageNumber="97">50</td>
<td id="542FE15F1A15000EFB58F8860033FB78" box="[1234,1348,1838,1860]" gridcol="4" gridrow="38" pageId="2" pageNumber="97">2–3</td>
</tr>
<tr id="17FE88231A15000EFF15F8FF0033FB51" box="[159,1348,1879,1901]" gridrow="39" pageId="2" pageNumber="97">
<th id="542FE15F1A15000EFF15F8FF0773FB51" box="[159,516,1879,1901]" gridcol="0" gridrow="39" pageId="2" pageNumber="97">Gill rakers on first ceratobranchial</th>
<td id="542FE15F1A15000EFD2AF8FF0789FB51" box="[672,766,1879,1901]" gridcol="1" gridrow="39" pageId="2" pageNumber="97">10</td>
<td id="542FE15F1A15000EFCC2F8FF06F4FB51" box="[840,899,1879,1901]" gridcol="2" gridrow="39" pageId="2" pageNumber="97">11</td>
<td id="542FE15F1A15000EFC78F8FF010EFB51" box="[1010,1145,1879,1901]" gridcol="3" gridrow="39" pageId="2" pageNumber="97">55</td>
<td id="542FE15F1A15000EFB58F8FF0033FB51" box="[1234,1348,1879,1901]" gridcol="4" gridrow="39" pageId="2" pageNumber="97">10–13</td>
</tr>
<tr id="17FE88231A15000EFF15F8280033FBAA" box="[159,1348,1920,1942]" gridrow="40" pageId="2" pageNumber="97">
<th id="542FE15F1A15000EFF15F8280773FBAA" box="[159,516,1920,1942]" gridcol="0" gridrow="40" pageId="2" pageNumber="97">Teeth outer row of left lower jaw</th>
<td id="542FE15F1A15000EFD2AF8280789FBAA" box="[672,766,1920,1942]" gridcol="1" gridrow="40" pageId="2" pageNumber="97">11</td>
<td id="542FE15F1A15000EFCC2F82806F4FBAA" box="[840,899,1920,1942]" gridcol="2" gridrow="40" pageId="2" pageNumber="97">11</td>
<td id="542FE15F1A15000EFC78F828010EFBAA" box="[1010,1145,1920,1942]" gridcol="3" gridrow="40" pageId="2" pageNumber="97">45</td>
<td id="542FE15F1A15000EFB58F8280033FBAA" box="[1234,1348,1920,1942]" gridcol="4" gridrow="40" pageId="2" pageNumber="97">9–13</td>
</tr>
<tr id="17FE88231A15000EFF15F8020033FBFC" box="[159,1348,1962,1984]" gridrow="41" pageId="2" pageNumber="97" rowspan-4="1">
<th id="542FE15F1A15000EFF15F8020773FBFC" box="[159,516,1962,1984]" gridcol="0" gridrow="41" pageId="2" pageNumber="97">Tooth rows on upper jaw</th>
<td id="542FE15F1A15000EFD2AF8020789FBFC" box="[672,766,1962,1984]" gridcol="1" gridrow="41" pageId="2" pageNumber="97">3</td>
<td id="542FE15F1A15000EFCC2F80206F4FBFC" box="[840,899,1962,1984]" gridcol="2" gridrow="41" pageId="2" pageNumber="97">3</td>
<td id="542FE15F1A15000EFC78F802010EFBFC" box="[1010,1145,1962,1984]" gridcol="3" gridrow="41" pageId="2" pageNumber="97">100</td>
</tr>
<tr id="17FE88231A15000EFF15F87B0033FBD5" box="[159,1348,2003,2025]" gridrow="42" pageId="2" pageNumber="97" rowspan-4="1">
<th id="542FE15F1A15000EFF15F87B0773FBD5" box="[159,516,2003,2025]" gridcol="0" gridrow="42" pageId="2" pageNumber="97">Tooth rows on lower jaw</th>
<td id="542FE15F1A15000EFD2AF87B0789FBD5" box="[672,766,2003,2025]" gridcol="1" gridrow="42" pageId="2" pageNumber="97">3</td>
<td id="542FE15F1A15000EFCC2F87B06F4FBD5" box="[840,899,2003,2025]" gridcol="2" gridrow="42" pageId="2" pageNumber="97">3</td>
<td id="542FE15F1A15000EFC78F87B010EFBD5" box="[1010,1145,2003,2025]" gridcol="3" gridrow="42" pageId="2" pageNumber="97">100</td>
</tr>
</table>
</paragraph>
<caption id="FDB1DAE91A14FFF2FF1DFBEF0683F861" box="[151,1012,1095,1117]" httpUri="https://zenodo.org/record/1138556/files/figure.png" pageId="3" pageNumber="98" targetBox="[151,1437,81,1074]" targetPageId="3">
<paragraph id="A9718A611A14FFF2FF1DFBEF0683F861" blockId="3.[151,1012,1095,1117]" box="[151,1012,1095,1117]" pageId="3" pageNumber="98">
<emphasis id="9BBA56731A14FFF2FF1DFBEF047AF860" bold="true" box="[151,269,1095,1117]" pageId="3" pageNumber="98">FIGURE 1</emphasis>
. The holotype of
<taxonomicName id="6ECEF1E21A14FFF2FE4BFBEF07EDF861" authorityName="Stauffer" authorityYear="2018" box="[449,666,1095,1117]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="98" phylum="Chordata" rank="species" species="koningsi">
<emphasis id="9BBA56731A14FFF2FE4BFBEF07EDF861" box="[449,666,1095,1117]" italics="true" pageId="3" pageNumber="98">Metriaclima koningsi</emphasis>
</taxonomicName>
(56.8 mm SL, male, PSU 12891).
</paragraph>
</caption>
<caption id="FDB1DAE91A14FFF2FF1DF8DF069BFB90" httpUri="https://zenodo.org/record/1138558/files/figure.png" pageId="3" pageNumber="98" targetBox="[151,1436,1167,1890]" targetPageId="3">
<paragraph id="A9718A611A14FFF2FF1DF8DF069BFB90" blockId="3.[151,1436,1911,1964]" pageId="3" pageNumber="98">
<emphasis id="9BBA56731A14FFF2FF1DF8DF0466FBB0" bold="true" box="[151,273,1911,1933]" pageId="3" pageNumber="98">FIGURE 2</emphasis>
. Three-dimensional surface rendering of the cranium of the holotype of
<taxonomicName id="6ECEF1E21A14FFF2FBA9F8DF0076FBB1" authorityName="Stauffer" authorityYear="2018" box="[1059,1281,1911,1933]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="98" phylum="Chordata" rank="species" species="koningsi">
<emphasis id="9BBA56731A14FFF2FBA9F8DF0076FBB1" box="[1059,1281,1911,1933]" italics="true" pageId="3" pageNumber="98">Metriaclima koningsi</emphasis>
</taxonomicName>
(PSU 12891), showing the position of the parashpenoid and the angle of the ethmo-vomerine bloc.
</paragraph>
</caption>
<caption id="FDB1DAE91A13FFF5FF1DFCAE04EEFF06" httpUri="https://zenodo.org/record/1138560/files/figure.png" pageId="4" pageNumber="99" targetBox="[151,1437,193,752]" targetPageId="4">
<paragraph id="A9718A611A13FFF5FF1DFCAE04EEFF06" blockId="4.[151,1436,774,826]" pageId="4" pageNumber="99">
<emphasis id="9BBA56731A13FFF5FF1DFCAE0479FF27" bold="true" box="[151,270,774,796]" pageId="4" pageNumber="99">FIGURE 3</emphasis>
.
<taxonomicName id="6ECEF1E21A13FFF5FE91FCAE0482FF20" authorityName="Stauffer" authorityYear="2018" box="[283,501,774,796]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="99" phylum="Chordata" rank="species" species="koningsi">
<emphasis id="9BBA56731A13FFF5FE91FCAE0482FF20" box="[283,501,774,796]" italics="true" pageId="4" pageNumber="99">Metriaclima koningsi</emphasis>
</taxonomicName>
.
<emphasis id="9BBA56731A13FFF5FD89FCAE0762FF27" bold="true" box="[515,533,774,795]" pageId="4" pageNumber="99">A</emphasis>
, Male from Mainango Island;
<emphasis id="9BBA56731A13FFF5FCDBFCAF061FFF20" bold="true" box="[849,872,775,796]" pageId="4" pageNumber="99">B,</emphasis>
Female from Membe Point, Likoma Island. Neither of the above were collected.
</paragraph>
</caption>
<paragraph id="A9718A611A13FFF5FF4DFCCF075CF864" blockId="4.[151,1437,871,1688]" pageId="4" pageNumber="99">Head of females yellow/brown with 1 light brown/green interorbital bar. Cheek yellowish brown with yellow/ brown operculum with green highlights; dark grey opercle spot; gular yellow. Laterally yellow/brown ground colour with yellow scale margins; centre of scales pale blue/white; belly and breast white. Dorsal fin yellow/brown with yellow lappets; distal one-third of rayed membranes bright yellow/orange. Caudal-fin membranes yellowish brown with yellow rays. Proximal one-quarter of anal fin clear; distal three-quarters yellow orange. Pectoral fins with brown rays and clear membranes. Anterior three membranes and rays of pelvic fin bright yellow/orange; posterior membranes and rays clear.</paragraph>
</subSubSection>
<subSubSection id="E1D4D9EA1A13FFF5FF4DFBCB0594F8F8" pageId="4" pageNumber="99" type="distribution">
<paragraph id="A9718A611A13FFF5FF4DFBCB0594F8F8" blockId="4.[151,1437,871,1688]" pageId="4" pageNumber="99">
<emphasis id="9BBA56731A13FFF5FF4DFBCB0417F840" bold="true" box="[199,352,1123,1148]" pageId="4" pageNumber="99">Distribution.</emphasis>
<taxonomicName id="6ECEF1E21A13FFF5FEE0FBCC0729F840" authorityName="Stauffer" authorityYear="2018" box="[362,606,1124,1148]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="99" phylum="Chordata" rank="species" species="koningsi">
<emphasis id="9BBA56731A13FFF5FEE0FBCC0729F840" box="[362,606,1124,1148]" italics="true" pageId="4" pageNumber="99">Metriaclima koningsi</emphasis>
</taxonomicName>
is a deep-water form and is usually found at depths greater than
<quantity id="6E3627841A13FFF5FAEAFBCB00EBF847" box="[1376,1436,1123,1148]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="99" unit="m" value="15.0">15 m</quantity>
(Konings 2016). It is only known from Membe Point and Maingano Island along the northeastern shore of
<collectingRegion id="6B0A44831A13FFF5FACFFB2F00EBF89C" box="[1349,1436,1159,1184]" country="Malawi" name="Likoma" pageId="4" pageNumber="99">Likoma</collectingRegion>
Island.
</paragraph>
</subSubSection>
<subSubSection id="E1D4D9EA1A13FFF5FF4DFB7807ECF930" pageId="4" pageNumber="99" type="etymology">
<paragraph id="A9718A611A13FFF5FF4DFB7807ECF930" blockId="4.[151,1437,871,1688]" pageId="4" pageNumber="99">
<emphasis id="9BBA56731A13FFF5FF4DFB780439F8D4" bold="true" box="[199,334,1232,1256]" pageId="4" pageNumber="99">Etymology.</emphasis>
Named after Adrianus Konings for his comprehensive knowledge and for teaching me most of what I know about the fishes of Lake
<collectingCountry id="D1D9CAF11A13FFF5FDB4FB5B07E0F930" box="[574,663,1267,1292]" name="Malawi" pageId="4" pageNumber="99">Malaŵi</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="E1D4D9EA1A13FFF5FF4DFABF069BF9A0" pageId="4" pageNumber="99" type="discussion">
<paragraph id="A9718A611A13FFF5FF4DFABF069BF9A0" blockId="4.[151,1437,871,1688]" pageId="4" pageNumber="99">
<emphasis id="9BBA56731A13FFF5FF4DFABF043BF90C" bold="true" box="[199,332,1303,1328]" pageId="4" pageNumber="99">Discussion.</emphasis>
<taxonomicName id="6ECEF1E21A13FFF5FEDFFABF073FF913" authorityName="Stauffer" authorityYear="2018" box="[341,584,1303,1327]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="99" phylum="Chordata" rank="species" species="koningsi">
<emphasis id="9BBA56731A13FFF5FEDFFABF073FF913" box="[341,584,1303,1327]" italics="true" pageId="4" pageNumber="99">Metriaclima koningsi</emphasis>
</taxonomicName>
usually occurs over the sediment-rich habitat between
<quantity id="6E3627841A13FFF5FB4FFABF005CF913" box="[1221,1323,1303,1328]" metricMagnitude="1" metricUnit="m" metricValue="2.25" metricValueMax="3.0" metricValueMin="1.5" pageId="4" pageNumber="99" unit="m" value="22.5" valueMax="30.0" valueMin="15.0">15–30 m</quantity>
(Konings 2016). Males have been observed feeding on the aufwuchs at a 90° angle, where they abduct their jaws to a near 180°; thus they can align their teeth of both the upper and lower jaws in the same plane. Both males and females will feed on zooplankton from the water column (Konings 2016, pers. obs.).
</paragraph>
</subSubSection>
<subSubSection id="E1D4D9EA1A13FFF5FF4DFA0F0139FAA4" pageId="4" pageNumber="99" type="reference_group">
<paragraph id="A9718A611A13FFF5FF4DFA0F0139FAA4" blockId="4.[151,1437,871,1688]" pageId="4" pageNumber="99">
<taxonomicName id="6ECEF1E21A13FFF5FF4DFA0F0425F983" authorityName="Stauffer, Bowers, Kellogg &amp; McKaye" authorityYear="1997" box="[199,338,1447,1471]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="99" phylum="Chordata" rank="genus">
<emphasis id="9BBA56731A13FFF5FF4DFA0F0425F983" box="[199,338,1447,1471]" italics="true" pageId="4" pageNumber="99">Metriaclima</emphasis>
</taxonomicName>
is a large genus of the endemic rock-dwelling cichlids in Lake
<collectingCountry id="D1D9CAF11A13FFF5FBCBFA0F01EDF9FC" box="[1089,1178,1447,1472]" name="Malawi" pageId="4" pageNumber="99">Malaŵi</collectingCountry>
. Stauffer
<emphasis id="9BBA56731A13FFF5FA81FA000030F983" box="[1291,1351,1447,1471]" italics="true" pageId="4" pageNumber="99">et al.</emphasis>
(2013) recognised 31 described species and subsequently
<bibRefCitation id="CD5FF7901A13FFF5FD5AFA63060DF9D8" box="[720,890,1483,1508]" pageId="4" pageNumber="99" refString="Li, S., Konings, A. F. &amp; Stauffer Jr., J. R. (2016) A revision of the Pseudotropheus elongatus species group (Teleostei: Cichlidae) with a description of a new genus and seven new species. Zootaxa, 4168 (2), 353 - 381. https: // doi. org / 10.11646 / zootaxa. 4168.2.9" type="journal article">
Li
<emphasis id="9BBA56731A13FFF5FD7AFA64065EF9D8" box="[752,809,1484,1508]" italics="true" pageId="4" pageNumber="99">et al.</emphasis>
(2016)
</bibRefCitation>
described two additional elongate species in the genus. Many authors (
<bibRefCitation id="CD5FF7901A13FFF5FE04FA5807EEFA34" author="Turner" box="[398,665,1519,1544]" pageId="4" pageNumber="99" refString="Turner, G. F. &amp; Burrows, M. T. (1995) A model of sympatric speciation by sexual selection. Proceedings of the Royal Society, London, Series B, 260, 287 - 292. https: // doi. org / 10.1098 / rspb. 1995.0093" type="journal article" year="1995">Turner &amp; Burrows1995</bibRefCitation>
,
<bibRefCitation id="CD5FF7901A13FFF5FD2EFA5806E9FA34" author="Van" box="[676,926,1519,1544]" pageId="4" pageNumber="99" refString="Van Doorn, G. S., Noest, A. J. &amp; Hogeweg, P. (1998) Sympatric speciation and extinction driven by environment dependent sexual selection. Proceedings of the Royal Society of London, Series B, 265, 1915 - 1919. https: // doi. org / 10.1098 / rspb. 1998.0520" type="journal article" year="1998">
Van Doorn
<emphasis id="9BBA56731A13FFF5FCAEFA58062BFA3B" box="[804,860,1519,1543]" italics="true" pageId="4" pageNumber="99">et al.</emphasis>
1998
</bibRefCitation>
,
<bibRefCitation id="CD5FF7901A13FFF5FC22FA470071FA34" author="Seehausen" box="[936,1286,1519,1544]" pageId="4" pageNumber="99" refString="Seehausen, O. &amp; Van Alphen, J. M. (1999) Can sympatric speciation by disruptive sexual selection explain rapid evolution of cichlid diversity in Lake Victoria? Ecology Letters, 2, 262 - 271. https: // doi. org / 10.1046 / j. 1461 - 0248.1999.00082. x" type="journal article" year="1999">Seehausen &amp; Van Alphen 1999</bibRefCitation>
) have argued that sexual selection on male breeding coloration and behavior has driven speciation of these fishes. Thus, in many cases male coloration is the primary trait used to delimit species (
<bibRefCitation id="CD5FF7901A13FFF5FCF4F99F0129FA6C" author="Ribbink" box="[894,1118,1591,1616]" pageId="4" pageNumber="99" refString="Ribbink, A. J., Marsh, B. A., Marsh, A. C., Ribbink, A. C. &amp; Sharp, B. J. (1983) A preliminary survey of the cichlid fishes of rocky habitats in Lake Malawi. South African Journal of Zoology, 18, 149 - 310. https: // doi. org / 10.1080 / 02541858.1983.11447831" type="journal article" year="1983">
Ribbink
<emphasis id="9BBA56731A13FFF5FC68F990016BFA73" box="[994,1052,1591,1615]" italics="true" pageId="4" pageNumber="99">et al.</emphasis>
1983
</bibRefCitation>
, Stauffer
<emphasis id="9BBA56731A13FFF5FB41F9900073FA73" box="[1227,1284,1591,1615]" italics="true" pageId="4" pageNumber="99">et al.</emphasis>
1995, 1997). There are now 35 described
<taxonomicName id="6ECEF1E21A13FFF5FE5EF9F40729FA48" authorityName="Stauffer, Bowers, Kellogg &amp; McKaye" authorityYear="1997" box="[468,606,1628,1652]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="99" phylum="Chordata" rank="genus">
<emphasis id="9BBA56731A13FFF5FE5EF9F40729FA48" box="[468,606,1628,1652]" italics="true" pageId="4" pageNumber="99">Metriaclima</emphasis>
</taxonomicName>
spp., but
<bibRefCitation id="CD5FF7901A13FFF5FD46F9F3060AFA48" author="Konings" box="[716,893,1627,1652]" pageId="4" pageNumber="99" refString="Konings, A. F. (2007) Malawi Cichlids in their natural habitat. 4 th Edition. Cichlid Press, El Paso, Texas, 424 pp." type="book" year="2007">Konings (2007)</bibRefCitation>
recognised at least 75 different forms within the genus; thus
<taxonomicName id="6ECEF1E21A13FFF5FE96F9D704D0FAAB" authorityName="Stauffer, Bowers, Kellogg &amp; McKaye" authorityYear="1997" box="[284,423,1663,1687]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="99" phylum="Chordata" rank="genus">
<emphasis id="9BBA56731A13FFF5FE96F9D704D0FAAB" box="[284,423,1663,1687]" italics="true" pageId="4" pageNumber="99">Metriaclima</emphasis>
</taxonomicName>
may be the most species-rich of the mbuna in Lake
<collectingCountry id="D1D9CAF11A13FFF5FC7AF9D7013EFAA4" box="[1008,1097,1663,1688]" name="Malawi" pageId="4" pageNumber="99">Malaŵi</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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