313 lines
40 KiB
XML
313 lines
40 KiB
XML
<document id="A7AE210724F02D657527C9B17E98E475" ID-DOI="10.5281/zenodo.186351" ID-GBIF-Dataset="bb17e039-5346-4e01-a9ba-59b87e9e6a4c" ID-ISSN="1175-5326" ID-Zenodo-Dep="186351" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461050352768" checkinUser="plazi" docAuthor="Maus, Mariana Chani-Posse De" docDate="2009" docId="03D787C9FFA8FF9AFF1BC909CB75E3C6" docLanguage="en" docName="zt02034p042.pdf" docOrigin="Zootaxa 2034" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Philonthus quadraticeps Boheman 1858" docType="treatment" docVersion="8" lastPageNumber="37" masterDocId="FFEEFFB1FFACFF9CFF8CCD70CA68E133" masterDocTitle="Revision of some types of Philonthus Stephens and Gabrius Stephens from southern South America (Coleoptera: Staphylinidae), with nomenclatural notes" masterLastPageNumber="42" masterPageNumber="31" pageNumber="35" updateTime="1698578662429" updateUser="plazi">
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<mods:title id="8D49A974DE12A5102911F3472D3E9017">Revision of some types of Philonthus Stephens and Gabrius Stephens from southern South America (Coleoptera: Staphylinidae), with nomenclatural notes</mods:title>
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<mods:namePart id="1BFF029AEC8EFDE4048D05CCA234A8C0">Maus, Mariana Chani-Posse De</mods:namePart>
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<mods:title id="86BEE3178988B67B4F375535F0B6CC3A">Zootaxa</mods:title>
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<mods:date id="CF4C668CF79B9A70E1239FC78A29A09E">2009</mods:date>
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<mods:detail id="2E61741D3168FFD1085F5A7F310B8DF2" type="volume">
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<mods:number id="CA646193ED89C9A863C847D44189785B">2034</mods:number>
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<subSubSection id="C3646554FFA8FF98FF1BC909C92BE585" pageId="4" pageNumber="35" type="nomenclature">
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<paragraph id="8BC136DFFFA8FF98FF1BC909C8EDE5A1" blockId="4.[151,835,1144,1206]" box="[151,645,1144,1171]" pageId="4" pageNumber="35">
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<taxonomicName id="4C7E4D5CFFA8FF98FF1BC909C8EDE5A1" ID-CoL="4GBN3" authority="Boheman, 1858" authorityName="Boheman" authorityYear="1858" box="[151,645,1144,1171]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="35" phylum="Arthropoda" rank="species" species="quadraticeps">
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<emphasis id="B90AEACDFFA8FF98FF1BC909CBD7E5A0" bold="true" box="[151,447,1145,1171]" italics="true" pageId="4" pageNumber="35">Philonthus quadraticeps</emphasis>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BC136DFFFA8FF98FF1BC9ECC92BE585" blockId="4.[151,835,1144,1206]" box="[151,835,1180,1206]" pageId="4" pageNumber="35">
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(
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<figureCitation id="13452A5AFFA8FF98FF13C9ECCB66E585" box="[159,270,1180,1206]" captionStart="FIGURES 1 – 12" captionStartId="6.[151,275,1634,1658]" captionText="FIGURES 1 – 12. Type specimens: habitus and labels; labels with lectotype designation are not included. (1 a – b) Philonthus argentinus Bernhauer (now in Gabrius), (lectotype with labels *). (2 a – b) Philonthus jujuyensis Bernhauer (now in Gabrius), (lectotype with labels *). (3 a – b) Philonthus tucumanensis Bernhauer (now in Gabrius), (lectotype with labels *). (4) Philonthus nidicola Bernhauer (now in Gabrius), (only labels are shown *). (5 a – b) Philonthus quadraticeps (lectotype with labels *). (6 a – b) P. tenebrosus (= P. quadraticeps), (lectotype with labels *). (7 a – b) Philonthus bonariensis (lectotype with labels *). (8 a – b) Philonthus weiseri (= P. bonariensis), (lectotype with labels *). (9 a – b) Philonthus catamarcanus (= P. bonariensis), (lectotype with labels *). (10 a – b) Philonthus cribriventris (lectotype with labels *). (11) Philonthus catamarcanus var. densior (= P. cribriventris), (lectotype, labels are not shown). (12) G. chiliensis (= G. nigritulus), (paratype, labels are not shown). * Labels with lectotype designation are not included (pictures were taken prior to any designation)." pageId="4" pageNumber="35">
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Figs.
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<quantity id="4C869B3AFFA8FF98FF6EC9ECCB66E585" box="[226,270,1180,1206]" metricMagnitude="-1" metricUnit="m" metricValue="1.4223999999999999" pageId="4" pageNumber="35" unit="in" value="5.6">5, 6</quantity>
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</figureCitation>
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in this paper; Figs.
|
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<quantity id="4C869B3AFFA8FF98FE7AC9ECC836E585" box="[502,606,1180,1206]" metricMagnitude="-1" metricUnit="m" metricValue="4.699" metricValueMax="5.334" metricValueMin="4.064" pageId="4" pageNumber="35" unit="in" value="18.5" valueMax="21.0" valueMin="16.0">16–21 in</quantity>
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Chani-Posse 2004)
|
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</paragraph>
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</subSubSection>
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<subSubSection id="C3646554FFA8FF98FF1BC999CE44E492" pageId="4" pageNumber="35" type="reference_group">
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<paragraph id="8BC136DFFFA8FF98FF1BC999C909E473" blockId="4.[151,1437,1257,1441]" pageId="4" pageNumber="35">
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<taxonomicName id="4C7E4D5CFFA8FF98FF1BC999C81DE433" ID-CoL="4GBN3" authority="Boheman,1858: 28" authorityName="Boheman" authorityPageNumber="28" authorityYear="1858" box="[151,629,1257,1280]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="35" phylum="Arthropoda" rank="species" species="quadraticeps">
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<emphasis id="B90AEACDFFA8FF98FF1BC999CBF6E433" box="[151,414,1257,1280]" italics="true" pageId="4" pageNumber="35">Philonthus quadraticeps</emphasis>
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<treatmentCitation id="0ADF10CEFFA8FF98FE2AC999C81DE433" author="Boheman" box="[422,629,1257,1280]" page="28" pageId="4" pageNumber="35" year="1858">
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<bibRefCitation id="EFEF4B2EFFA8FF98FE2AC999C824E433" author="Boheman" box="[422,588,1257,1280]" pageId="4" pageNumber="41" refString="Boheman, C. H. (1858) Coleoptera. Species novas descripsit. In: Virgin, C. (Ed.), Kongliga Svenska Fregatten Eugenies resa omkring Jorden, Zoologi I, Insecta. P. A. Norstedt & Soner, Stockholm, pp. 1 - 112." type="book chapter" year="1858">Boheman,1858</bibRefCitation>
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: 28
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||
</treatmentCitation>
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</taxonomicName>
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;
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||
<treatmentCitation id="0ADF10CEFFA8FF98FD0EC999C9AFE433" author="Lynch-Arribalzaga" box="[642,967,1257,1280]" page="150" pageId="4" pageNumber="35" year="1884">
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<bibRefCitation id="EFEF4B2EFFA8FF98FD0EC999C9FCE433" author="Lynch-Arribalzaga" box="[642,916,1257,1280]" pageId="4" pageNumber="42" refString="Lynch-Arribalzaga, F. (1884) Estafilinos de Buenos Aires. Boletin de la Academia Nacional de Ciencias, Cordoba, 7, 5 - 256." type="journal article" year="1884">Lynch-Arribálzaga, 1884</bibRefCitation>
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||
: 150
|
||
</treatmentCitation>
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||
(characters);
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||
<treatmentCitation id="0ADF10CEFFA8FF98FBD1C999CA87E413" author="Bernhauer" page="352" pageId="4" pageNumber="35" year="1914">
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<bibRefCitation id="EFEF4B2EFFA8FF98FBD1C999CFF0E433" author="Bernhauer" box="[1117,1432,1257,1280]" pageId="4" pageNumber="41" refString="Bernhauer, M. & Schubert, K. (1914) Staphylinidae IV. In: Schenkling, S. (Ed.), Coleopterorum Catalogus, 5 (57). W. Junk, Berlin, pp. 289 - 408." type="journal article" year="1914">Bernhauer & Schubert, 1914</bibRefCitation>
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: 352
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||
</treatmentCitation>
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(catalog);
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||
<treatmentCitation id="0ADF10CEFFA8FF98FEEEC879C873E413" author="Bruch" box="[354,539,1289,1312]" page="445" pageId="4" pageNumber="35" year="1915">
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<bibRefCitation id="EFEF4B2EFFA8FF98FEEEC879CB81E413" author="Bruch" box="[354,489,1289,1312]" pageId="4" pageNumber="41" refString="Bruch, C. (1915) Catalogo Sistematico de los coleopteros de la Republica Argentina. Pars VI. Revista del Museo de La Plata, 19 (Segunda Serie) (6), 235 - 302." type="journal article" year="1915">Bruch, 1915</bibRefCitation>
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: 445
|
||
</treatmentCitation>
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(catalog);
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||
<treatmentCitation id="0ADF10CEFFA8FF98FD02C879C9E3E413" author="Blackwelder" box="[654,907,1289,1312]" page="134" pageId="4" pageNumber="35" year="1944">
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<bibRefCitation id="EFEF4B2EFFA8FF98FD02C879C931E413" author="Blackwelder" box="[654,857,1289,1312]" pageId="4" pageNumber="41" refString="Blackwelder, R. E. (1944) Checklist of the Coleopterous insects of Mexico, Central America, the West Indies, and South America. Part I. United States National Museum Bulletin, 185, 1 - 188." type="journal article" year="1944">Blackwelder, 1944</bibRefCitation>
|
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: 134
|
||
</treatmentCitation>
|
||
(list);
|
||
<bibRefCitation id="EFEF4B2EFFA8FF98FC5EC879CE3DE413" author="Frank" box="[978,1109,1289,1312]" pageId="4" pageNumber="42" refString="Frank, J. H. (1982) The parasites of the Staphylinidae (Coleoptera). A contribution towards an encyclopedia of the Staphylinidae. University of Florida Agr. Exp. Stations, Technical Bulletin, 824, i - vii + 1 - 118 pp." type="journal article" year="1982">Frank, 1982</bibRefCitation>
|
||
; 43 (list);
|
||
<treatmentCitation id="0ADF10CEFFA8FF98FB4DC879CFF4E413" author="Herman" box="[1217,1436,1289,1312]" page="2928" pageId="4" pageNumber="35" year="2001">
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<bibRefCitation id="EFEF4B2EFFA8FF98FB4DC879CF35E413" author="Herman" box="[1217,1373,1289,1312]" pageId="4" pageNumber="42" refString="Herman, L. H. (2001) Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. Part V. Bulletin of the American Museum of Natural History, 265, 2441 - 3020." type="journal article" year="2001">Herman, 2001</bibRefCitation>
|
||
: 2928
|
||
</treatmentCitation>
|
||
(catalog); Chani-Posse, 2004: 224–227 (characters), 231 (list).
|
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BC136DFFFA8FF98FF1BC839CE44E492" blockId="4.[151,1437,1257,1441]" pageId="4" pageNumber="35">
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<treatmentCitationGroup id="AB6E11F1FFA8FF98FF1BC839CE44E492" pageId="4" pageNumber="35">
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<taxonomicName id="4C7E4D5CFFA8FF98FF1BC839C8E8E453" ID-CoL="4GBYC" authority="Boheman, 1858: 28" authorityName="Boheman" authorityPageNumber="28" authorityYear="1858" box="[151,640,1353,1376]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="35" phylum="Arthropoda" rank="species" species="tenebrosus">
|
||
<emphasis id="B90AEACDFFA8FF98FF1BC839CBE4E453" box="[151,396,1353,1376]" italics="true" pageId="4" pageNumber="35">Philonthus tenebrosus</emphasis>
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<treatmentCitation id="0ADF10CEFFA8FF98FE17C839C8E8E453" author="Boheman" box="[411,640,1353,1376]" page="28" pageId="4" pageNumber="35" year="1858">
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<bibRefCitation id="EFEF4B2EFFA8FF98FE17C839C838E453" author="Boheman" box="[411,592,1353,1376]" pageId="4" pageNumber="41" refString="Boheman, C. H. (1858) Coleoptera. Species novas descripsit. In: Virgin, C. (Ed.), Kongliga Svenska Fregatten Eugenies resa omkring Jorden, Zoologi I, Insecta. P. A. Norstedt & Soner, Stockholm, pp. 1 - 112." type="book chapter" year="1858">Boheman, 1858</bibRefCitation>
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||
: 28
|
||
</treatmentCitation>
|
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</taxonomicName>
|
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;
|
||
<treatmentCitation id="0ADF10CEFFA8FF98FD1FC839C98EE453" author="Lynch-Arribalzaga" box="[659,998,1353,1376]" page="149" pageId="4" pageNumber="35" year="1884">
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<bibRefCitation id="EFEF4B2EFFA8FF98FD1FC839C9C3E453" author="Lynch-Arribalzaga" box="[659,939,1353,1376]" pageId="4" pageNumber="42" refString="Lynch-Arribalzaga, F. (1884) Estafilinos de Buenos Aires. Boletin de la Academia Nacional de Ciencias, Cordoba, 7, 5 - 256." type="journal article" year="1884">Lynch-Arribálzaga, 1884</bibRefCitation>
|
||
: 149
|
||
</treatmentCitation>
|
||
(characters,
|
||
<collectingCountry id="F369764FFFA8FF98FB0CC839CE87E453" box="[1152,1263,1353,1376]" name="Argentina" pageId="4" pageNumber="35">Argentina</collectingCountry>
|
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);
|
||
<treatmentCitation id="0ADF10CEFFA8FF98FA85C839CBCAE4B3" author="Bernhauer" page="357" pageId="4" pageNumber="35" year="1914">
|
||
<bibRefCitation id="EFEF4B2EFFA8FF98FA85C839CB06E4B3" author="Bernhauer" pageId="4" pageNumber="41" refString="Bernhauer, M. & Schubert, K. (1914) Staphylinidae IV. In: Schenkling, S. (Ed.), Coleopterorum Catalogus, 5 (57). W. Junk, Berlin, pp. 289 - 408." type="journal article" year="1914">Bernhauer & Schubert, 1914</bibRefCitation>
|
||
: 357
|
||
</treatmentCitation>
|
||
(catalog);
|
||
<treatmentCitation id="0ADF10CEFFA8FF98FD95C819C8BEE4B3" author="Bruch" box="[537,726,1385,1408]" page="445" pageId="4" pageNumber="35" year="1915">
|
||
<bibRefCitation id="EFEF4B2EFFA8FF98FD95C819C8CAE4B3" author="Bruch" box="[537,674,1385,1408]" pageId="4" pageNumber="41" refString="Bruch, C. (1915) Catalogo Sistematico de los coleopteros de la Republica Argentina. Pars VI. Revista del Museo de La Plata, 19 (Segunda Serie) (6), 235 - 302." type="journal article" year="1915">Bruch, 1915</bibRefCitation>
|
||
: 445
|
||
</treatmentCitation>
|
||
(catalog);
|
||
<treatmentCitation id="0ADF10CEFFA8FF98FCC1C819CE3BE4B3" author="Scheerpeltz" box="[845,1107,1385,1408]" page="1363" pageId="4" pageNumber="35" year="1933">
|
||
<bibRefCitation id="EFEF4B2EFFA8FF98FCC1C819CE7AE4B3" author="Scheerpeltz" box="[845,1042,1385,1408]" pageId="4" pageNumber="42" refString="Scheerpeltz, O. (1933) Staphylinidae VII. In: Schenkling, S. (Ed.), Coleopterorum Catalogus. Pars 129. W. Junk, Berlin, pp. 989 - 1500." type="journal article" year="1933">Scheerpeltz, 1933</bibRefCitation>
|
||
: 1363
|
||
</treatmentCitation>
|
||
(catalog);
|
||
<treatmentCitation id="0ADF10CEFFA8FF98FB46C819CA87E493" author="Blackwelder" page="135" pageId="4" pageNumber="35" year="1944">
|
||
<bibRefCitation id="EFEF4B2EFFA8FF98FB46C819CFF0E4B3" author="Blackwelder" box="[1226,1432,1385,1408]" pageId="4" pageNumber="41" refString="Blackwelder, R. E. (1944) Checklist of the Coleopterous insects of Mexico, Central America, the West Indies, and South America. Part I. United States National Museum Bulletin, 185, 1 - 188." type="journal article" year="1944">Blackwelder, 1944</bibRefCitation>
|
||
: 135
|
||
</treatmentCitation>
|
||
(list);
|
||
<treatmentCitation id="0ADF10CEFFA8FF98FEBAC8F9C879E493" author="Herman" box="[310,529,1417,1440]" page="2977" pageId="4" pageNumber="35" year="2001">
|
||
<bibRefCitation id="EFEF4B2EFFA8FF98FEBAC8F9CBBAE493" author="Herman" box="[310,466,1417,1440]" pageId="4" pageNumber="42" refString="Herman, L. H. (2001) Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. Part V. Bulletin of the American Museum of Natural History, 265, 2441 - 3020." type="journal article" year="2001">Herman, 2001</bibRefCitation>
|
||
: 2977
|
||
</treatmentCitation>
|
||
(catalog); Chani-Posse, 2004: 231 (list),
|
||
<emphasis id="B90AEACDFFA8FF98FC45C8F9CE44E492" bold="true" box="[969,1068,1417,1441]" pageId="4" pageNumber="35">syn. nov.</emphasis>
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</treatmentCitationGroup>
|
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</paragraph>
|
||
</subSubSection>
|
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<subSubSection id="C3646554FFA8FF98FF1BC8BFCBFFE62C" pageId="4" pageNumber="35" type="materials_examined">
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<paragraph id="8BC136DFFFA8FF98FF1BC8BFCF21E704" blockId="4.[151,1437,1487,2016]" pageId="4" pageNumber="35">
|
||
<emphasis id="B90AEACDFFA8FF98FF1BC8BFCBA7E4DA" bold="true" box="[151,463,1487,1513]" pageId="4" pageNumber="35">
|
||
<typeStatus id="54C5887DFFA8FF98FF1BC8BFCABCE4DA" box="[151,212,1487,1513]" pageId="4" pageNumber="35">Type</typeStatus>
|
||
material.
|
||
<typeStatus id="54C5887DFFA8FF98FEDFC8BFCBA7E4DA" box="[339,463,1487,1513]" pageId="4" pageNumber="35" type="lectotype">Lectotype</typeStatus>
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</emphasis>
|
||
of
|
||
<taxonomicName id="4C7E4D5CFFA8FF98FE79C8A0C97DE4DA" box="[501,789,1488,1513]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="35" phylum="Arthropoda" rank="species" species="quadraticeps">
|
||
<emphasis id="B90AEACDFFA8FF98FE79C8A0C97DE4DA" box="[501,789,1488,1513]" italics="true" pageId="4" pageNumber="35">Philonthus quadraticeps</emphasis>
|
||
</taxonomicName>
|
||
(here designated;
|
||
<figureCitation id="13452A5AFFA8FF98FC67C8A0CE52E4D9" box="[1003,1082,1488,1514]" captionStart="FIGURES 1 – 12" captionStartId="6.[151,275,1634,1658]" captionText="FIGURES 1 – 12. Type specimens: habitus and labels; labels with lectotype designation are not included. (1 a – b) Philonthus argentinus Bernhauer (now in Gabrius), (lectotype with labels *). (2 a – b) Philonthus jujuyensis Bernhauer (now in Gabrius), (lectotype with labels *). (3 a – b) Philonthus tucumanensis Bernhauer (now in Gabrius), (lectotype with labels *). (4) Philonthus nidicola Bernhauer (now in Gabrius), (only labels are shown *). (5 a – b) Philonthus quadraticeps (lectotype with labels *). (6 a – b) P. tenebrosus (= P. quadraticeps), (lectotype with labels *). (7 a – b) Philonthus bonariensis (lectotype with labels *). (8 a – b) Philonthus weiseri (= P. bonariensis), (lectotype with labels *). (9 a – b) Philonthus catamarcanus (= P. bonariensis), (lectotype with labels *). (10 a – b) Philonthus cribriventris (lectotype with labels *). (11) Philonthus catamarcanus var. densior (= P. cribriventris), (lectotype, labels are not shown). (12) G. chiliensis (= G. nigritulus), (paratype, labels are not shown). * Labels with lectotype designation are not included (pictures were taken prior to any designation)." pageId="4" pageNumber="35">Figs. 5</figureCitation>
|
||
a–b), 3, “
|
||
<typeStatus id="54C5887DFFA8FF98FB3FC8A0CE9DE4D9" box="[1203,1269,1488,1514]" pageId="4" pageNumber="35">Type</typeStatus>
|
||
”, “Buen Ayr”, “Kinb”, “
|
||
<typeStatus id="54C5887DFFA8FF98FE89C886CB3CE723" box="[261,340,1526,1552]" pageId="4" pageNumber="35">Typus</typeStatus>
|
||
”, “
|
||
<taxonomicName id="4C7E4D5CFFA8FF98FEF4C887C83DE723" authority="Bhn." authorityName="Bhn." box="[376,597,1526,1552]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="35" phylum="Arthropoda" rank="species" species="quadraticeps">
|
||
<emphasis id="B90AEACDFFA8FF98FEF4C887C879E723" box="[376,529,1527,1552]" italics="true" pageId="4" pageNumber="35">quadraticeps</emphasis>
|
||
Bhn.
|
||
</taxonomicName>
|
||
”, “5374 E91”, “Naturhistoriska Riksmuseet Stockholm / Loan no 834 / 06”, “
|
||
<typeStatus id="54C5887DFFA8FF98FF56CB6DCB3AE704" box="[218,338,1565,1591]" pageId="4" pageNumber="35" type="lectotype">Lectotype</typeStatus>
|
||
<taxonomicName id="4C7E4D5CFFA8FF98FED5CB6DC950E704" authority="Boheman, 1858" authorityName="Boheman" authorityYear="1858" box="[345,824,1565,1591]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="35" phylum="Arthropoda" rank="species" species="quadraticeps">
|
||
<emphasis id="B90AEACDFFA8FF98FED5CB6DC811E705" box="[345,633,1565,1590]" italics="true" pageId="4" pageNumber="35">Philonthus quadraticeps</emphasis>
|
||
<bibRefCitation id="EFEF4B2EFFA8FF98FD0CCB6DC950E704" author="Boheman" box="[640,824,1565,1591]" pageId="4" pageNumber="41" refString="Boheman, C. H. (1858) Coleoptera. Species novas descripsit. In: Virgin, C. (Ed.), Kongliga Svenska Fregatten Eugenies resa omkring Jorden, Zoologi I, Insecta. P. A. Norstedt & Soner, Stockholm, pp. 1 - 112." type="book chapter" year="1858">Boheman, 1858</bibRefCitation>
|
||
</taxonomicName>
|
||
/ Des. Chani-Posse de Maus 2008” (
|
||
<collectionCode id="ED6FAE1AFFA8FF98FB64CB6DCF55E704" box="[1256,1341,1565,1591]" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="4" pageNumber="35">NHRS</collectionCode>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC136DFFFA8FF98FF4ACB33CBFFE62C" blockId="4.[151,1437,1487,2016]" pageId="4" pageNumber="35">
|
||
<emphasis id="B90AEACDFFA8FF98FF4ACB33CB2AE76E" bold="true" box="[198,322,1603,1629]" pageId="4" pageNumber="35">
|
||
<typeStatus id="54C5887DFFA8FF98FF4ACB33CB2AE76E" box="[198,322,1603,1629]" pageId="4" pageNumber="35" type="lectotype">Lectotype</typeStatus>
|
||
</emphasis>
|
||
of
|
||
<taxonomicName id="4C7E4D5CFFA8FF98FEFECB34C813E76E" box="[370,635,1604,1629]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="35" phylum="Arthropoda" rank="species" species="tenebrosus">
|
||
<emphasis id="B90AEACDFFA8FF98FEFECB34C813E76E" box="[370,635,1604,1629]" italics="true" pageId="4" pageNumber="35">Philonthus tenebrosus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<collectionCode id="ED6FAE1AFFA8FF98FD03CB34C889E76D" box="[655,737,1604,1630]" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="4" pageNumber="35">NHRS</collectionCode>
|
||
) (here designated;
|
||
<figureCitation id="13452A5AFFA8FF98FC42CB34CE4BE76D" box="[974,1059,1604,1630]" captionStart="FIGURES 1 – 12" captionStartId="6.[151,275,1634,1658]" captionText="FIGURES 1 – 12. Type specimens: habitus and labels; labels with lectotype designation are not included. (1 a – b) Philonthus argentinus Bernhauer (now in Gabrius), (lectotype with labels *). (2 a – b) Philonthus jujuyensis Bernhauer (now in Gabrius), (lectotype with labels *). (3 a – b) Philonthus tucumanensis Bernhauer (now in Gabrius), (lectotype with labels *). (4) Philonthus nidicola Bernhauer (now in Gabrius), (only labels are shown *). (5 a – b) Philonthus quadraticeps (lectotype with labels *). (6 a – b) P. tenebrosus (= P. quadraticeps), (lectotype with labels *). (7 a – b) Philonthus bonariensis (lectotype with labels *). (8 a – b) Philonthus weiseri (= P. bonariensis), (lectotype with labels *). (9 a – b) Philonthus catamarcanus (= P. bonariensis), (lectotype with labels *). (10 a – b) Philonthus cribriventris (lectotype with labels *). (11) Philonthus catamarcanus var. densior (= P. cribriventris), (lectotype, labels are not shown). (12) G. chiliensis (= G. nigritulus), (paratype, labels are not shown). * Labels with lectotype designation are not included (pictures were taken prior to any designation)." pageId="4" pageNumber="35">Figs. 6</figureCitation>
|
||
a–b), 3, “
|
||
<typeStatus id="54C5887DFFA8FF98FB25CB34CE83E76D" box="[1193,1259,1604,1630]" pageId="4" pageNumber="35">Type</typeStatus>
|
||
”, “Buen Ayr”, “Kinb”, “
|
||
<typeStatus id="54C5887DFFA8FF98FE84CB1ACB31E7B7" box="[264,345,1642,1668]" pageId="4" pageNumber="35">Typus</typeStatus>
|
||
”, “
|
||
<taxonomicName id="4C7E4D5CFFA8FF98FEF2CB1BC82DE7B7" authority="Bhn." authorityName="Bhn." box="[382,581,1642,1668]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="35" phylum="Arthropoda" rank="species" species="tenebrosus">
|
||
<emphasis id="B90AEACDFFA8FF98FEF2CB1BCB97E7B7" box="[382,511,1643,1668]" italics="true" pageId="4" pageNumber="35">tenebrosus</emphasis>
|
||
Bhn.
|
||
</taxonomicName>
|
||
”, “5373 E91”, “Naturhistoriska Riksmuseet Stockholm / Loan no 832/ 06”, “
|
||
<typeStatus id="54C5887DFFA8FF98FF6DCBE1CB0BE798" box="[225,355,1681,1707]" pageId="4" pageNumber="35" type="lectotype">Lectotype</typeStatus>
|
||
<taxonomicName id="4C7E4D5CFFA8FF98FEE0CBE1C932E798" authority="Boheman, 1858" authorityName="Boheman" authorityYear="1858" box="[364,858,1681,1707]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="35" phylum="Arthropoda" rank="species" species="tenebrosus">
|
||
<emphasis id="B90AEACDFFA8FF98FEE0CBE1C8E1E799" box="[364,649,1681,1706]" italics="true" pageId="4" pageNumber="35">Philonthus tenebrosus</emphasis>
|
||
<bibRefCitation id="EFEF4B2EFFA8FF98FD1ECBE1C932E798" author="Boheman" box="[658,858,1681,1707]" pageId="4" pageNumber="41" refString="Boheman, C. H. (1858) Coleoptera. Species novas descripsit. In: Virgin, C. (Ed.), Kongliga Svenska Fregatten Eugenies resa omkring Jorden, Zoologi I, Insecta. P. A. Norstedt & Soner, Stockholm, pp. 1 - 112." type="book chapter" year="1858">Boheman, 1858</bibRefCitation>
|
||
</taxonomicName>
|
||
/ Des. Chani-Posse de Maus 2008” (
|
||
<collectionCode id="ED6FAE1AFFA8FF98FAB9CBE1CFF8E798" box="[1333,1424,1681,1707]" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="4" pageNumber="35">NHRS</collectionCode>
|
||
).
|
||
<emphasis id="B90AEACDFFA8FF98FF1BCBC7CB30E7E2" bold="true" box="[151,344,1719,1745]" pageId="4" pageNumber="35">
|
||
<typeStatus id="54C5887DFFA8FF98FF1BCBC7CB3BE7E2" box="[151,339,1719,1745]" pageId="4" pageNumber="35" type="paralectotype">Paralectotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
3, “Buen Ayr”, “Kinb”, “
|
||
<taxonomicName id="4C7E4D5CFFA8FF98FD2ACBC8C91DE7E1" authority="Bhn." authorityName="Bhn." box="[678,885,1720,1746]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="35" phylum="Arthropoda" rank="species" species="tenebrosus">
|
||
<emphasis id="B90AEACDFFA8FF98FD2ACBC8C944E7E2" box="[678,812,1720,1745]" italics="true" pageId="4" pageNumber="35">tenebrosus</emphasis>
|
||
Bhn.
|
||
</taxonomicName>
|
||
”, “5375 E91”, “Naturhistoriska Riksmuseet Stockholm / Loan no 833/06”, “
|
||
<typeStatus id="54C5887DFFA8FF98FD98CBAEC8D1E7CB" box="[532,697,1758,1784]" pageId="4" pageNumber="35" type="paralectotype">Paralectotype</typeStatus>
|
||
<taxonomicName id="4C7E4D5CFFA8FF98FD4CCBAFCEE5E7CB" authority="Boheman, 1858" authorityName="Boheman" authorityYear="1858" box="[704,1165,1758,1784]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="35" phylum="Arthropoda" rank="species" species="tenebrosus">
|
||
<emphasis id="B90AEACDFFA8FF98FD4CCBAFC9A0E7CB" box="[704,968,1759,1784]" italics="true" pageId="4" pageNumber="35">Philonthus tenebrosus</emphasis>
|
||
<bibRefCitation id="EFEF4B2EFFA8FF98FC5DCBAECEE5E7CB" author="Boheman" box="[977,1165,1758,1784]" pageId="4" pageNumber="41" refString="Boheman, C. H. (1858) Coleoptera. Species novas descripsit. In: Virgin, C. (Ed.), Kongliga Svenska Fregatten Eugenies resa omkring Jorden, Zoologi I, Insecta. P. A. Norstedt & Soner, Stockholm, pp. 1 - 112." type="book chapter" year="1858">Boheman, 1858</bibRefCitation>
|
||
</taxonomicName>
|
||
/ Des. Chani-Posse de Maus 2008” (
|
||
<collectionCode id="ED6FAE1AFFA8FF98FEBACA75CBE3E62C" box="[310,395,1797,1823]" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="4" pageNumber="35">NHRS</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3646554FFA8FF9AFF4ACA5BC808E0F3" lastPageId="6" lastPageNumber="37" pageId="4" pageNumber="35" type="diagnosis">
|
||
<paragraph id="8BC136DFFFA8FF98FF4ACA5BCB5FE6D3" blockId="4.[151,1437,1487,2016]" pageId="4" pageNumber="35">
|
||
<emphasis id="B90AEACDFFA8FF98FF4ACA5BCB2EE676" bold="true" box="[198,326,1835,1861]" pageId="4" pageNumber="35">Diagnosis.</emphasis>
|
||
<taxonomicName id="4C7E4D5CFFA8FF98FEDCCA5CC81AE676" box="[336,626,1836,1861]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="35" phylum="Arthropoda" rank="species" species="quadraticeps">
|
||
<emphasis id="B90AEACDFFA8FF98FEDCCA5CC81AE676" box="[336,626,1836,1861]" italics="true" pageId="4" pageNumber="35">Philonthus quadraticeps</emphasis>
|
||
</taxonomicName>
|
||
may be identified by the following characters: head and pronotum distinctly quadrate, antennal segments 4 and 5 quadrate, eyes shorter than temples seen from above, postmandibular ridge absent, dorsal rows on pronotum each with four punctures, and transverse distance between punctures of elytra larger than diameters of punctures. For further details and illustrations see Chani-
|
||
<bibRefCitation id="EFEF4B2EFFA8FF98FF1BCAB6CB5BE6D3" author="Posse" box="[151,307,1990,2016]" pageId="4" pageNumber="41" refString="Chani Posse, M. R. (2004) Eight Argentinean species of dung-inhabiting Philonthus Stephens (Coleoptera: Staphylinidae). Studies on Neotropical Fauna and Environment, 39, 217 - 232." type="journal article" year="2004">Posse (2004)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC136DFFFAAFF9AFF4ACDE7CEB0E015" blockId="6.[151,1437,151,757]" pageId="6" pageNumber="37">
|
||
<emphasis id="B90AEACDFFAAFF9AFF4ACDE7CB7CE182" bold="true" box="[198,276,151,177]" pageId="6" pageNumber="37">Notes.</emphasis>
|
||
I designate the
|
||
<typeStatus id="54C5887DFFAAFF9AFE41CDE8C850E181" box="[461,568,152,178]" pageId="6" pageNumber="37" type="lectotype">lectotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C7E4D5CFFAAFF9AFDECCDE8C97EE182" box="[608,790,152,177]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="37" phylum="Arthropoda" rank="species" species="quadraticeps">
|
||
<emphasis id="B90AEACDFFAAFF9AFDECCDE8C97EE182" box="[608,790,152,177]" italics="true" pageId="6" pageNumber="37">P. quadraticeps</emphasis>
|
||
</taxonomicName>
|
||
rather than assume that the examined
|
||
<typeStatus id="54C5887DFFAAFF9AFB54CDE8CF62E181" box="[1240,1290,152,178]" pageId="6" pageNumber="37">type</typeStatus>
|
||
specimen is the
|
||
<typeStatus id="54C5887DFFAAFF9AFF49CDCECB58E1EB" box="[197,304,190,216]" pageId="6" pageNumber="37" type="holotype">holotype</typeStatus>
|
||
since in the original description
|
||
<bibRefCitation id="EFEF4B2EFFAAFF9AFD45CDCEC9FFE1EB" author="Boheman" box="[713,919,190,216]" pageId="6" pageNumber="41" refString="Boheman, C. H. (1858) Coleoptera. Species novas descripsit. In: Virgin, C. (Ed.), Kongliga Svenska Fregatten Eugenies resa omkring Jorden, Zoologi I, Insecta. P. A. Norstedt & Soner, Stockholm, pp. 1 - 112." type="book chapter" year="1858">Boheman (1858)</bibRefCitation>
|
||
gave no information about the number of specimens examined. Therefore, it is possible that this name was based on more than one specimen (
|
||
<bibRefCitation id="EFEF4B2EFFAAFF9AFAD9CD95CB80E015" author="ICZN" pageId="6" pageNumber="42" refString="ICZN (1999) International Code of Zoological Nomenclatur e. Fourth Edition. The International Trust for Zoological Nomenclature, London, xxix + 306 pp." type="book" year="1999">ICZN 1999, Recommendation 73F</bibRefCitation>
|
||
). The
|
||
<typeStatus id="54C5887DFFAAFF9AFDBDCC7CC8F4E015" box="[561,668,268,294]" pageId="6" pageNumber="37" type="lectotype">lectotype</typeStatus>
|
||
agrees with the description by
|
||
<bibRefCitation id="EFEF4B2EFFAAFF9AFB86CC7CCEBCE015" author="Boheman" box="[1034,1236,268,294]" pageId="6" pageNumber="41" refString="Boheman, C. H. (1858) Coleoptera. Species novas descripsit. In: Virgin, C. (Ed.), Kongliga Svenska Fregatten Eugenies resa omkring Jorden, Zoologi I, Insecta. P. A. Norstedt & Soner, Stockholm, pp. 1 - 112." type="book chapter" year="1858">Boheman (1858)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC136DFFFAAFF9AFF4ACC42C808E0F3" blockId="6.[151,1437,151,757]" pageId="6" pageNumber="37">
|
||
In the original description of
|
||
<taxonomicName id="4C7E4D5CFFAAFF9AFDBBCC42C9C6E07F" authority="Boheman 1858" authorityName="Boheman" authorityYear="1858" box="[567,942,306,332]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="37" phylum="Arthropoda" rank="species" species="tenebrosus">
|
||
<emphasis id="B90AEACDFFAAFF9AFDBBCC42C8BFE078" box="[567,727,306,331]" italics="true" pageId="6" pageNumber="37">P. tenebrosus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFEF4B2EFFAAFF9AFD67CC42C9CEE07F" author="Boheman" box="[747,934,306,332]" pageId="6" pageNumber="41" refString="Boheman, C. H. (1858) Coleoptera. Species novas descripsit. In: Virgin, C. (Ed.), Kongliga Svenska Fregatten Eugenies resa omkring Jorden, Zoologi I, Insecta. P. A. Norstedt & Soner, Stockholm, pp. 1 - 112." type="book chapter" year="1858">Boheman 1858</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
there is no indication about how many specimens were included in the
|
||
<typeStatus id="54C5887DFFAAFF9AFDA3CC29C80DE040" box="[559,613,345,371]" pageId="6" pageNumber="37">type</typeStatus>
|
||
series. Both specimens listed in the
|
||
<typeStatus id="54C5887DFFAAFF9AFBB5CC29CE07E040" box="[1081,1135,345,371]" pageId="6" pageNumber="37">type</typeStatus>
|
||
material agree with the description by
|
||
<bibRefCitation id="EFEF4B2EFFAAFF9AFEDECCF0C841E0A9" author="Boheman" box="[338,553,384,410]" pageId="6" pageNumber="41" refString="Boheman, C. H. (1858) Coleoptera. Species novas descripsit. In: Virgin, C. (Ed.), Kongliga Svenska Fregatten Eugenies resa omkring Jorden, Zoologi I, Insecta. P. A. Norstedt & Soner, Stockholm, pp. 1 - 112." type="book chapter" year="1858">Boheman (1858)</bibRefCitation>
|
||
. The specimen in a better condition is designated as the
|
||
<typeStatus id="54C5887DFFAAFF9AFB7ECCF0CF00E0A9" box="[1266,1384,384,410]" pageId="6" pageNumber="37" type="lectotype">lectotype</typeStatus>
|
||
, the remaining specimen is a
|
||
<typeStatus id="54C5887DFFAAFF9AFE36CCD6C834E0F3" box="[442,604,422,448]" pageId="6" pageNumber="37" type="paralectotype">paralectotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3646554FFAAFF9AFF4ACCBDCB75E3C6" pageId="6" pageNumber="37" type="discussion">
|
||
<paragraph id="8BC136DFFFAAFF9AFF4ACCBDCB75E3C6" blockId="6.[151,1437,151,757]" pageId="6" pageNumber="37">
|
||
<emphasis id="B90AEACDFFAAFF9AFF4ACCBDCB3CE0D4" bold="true" box="[198,340,461,487]" pageId="6" pageNumber="37">Comments.</emphasis>
|
||
My examination of the
|
||
<typeStatus id="54C5887DFFAAFF9AFD05CCBDC8AFE0D4" box="[649,711,461,487]" pageId="6" pageNumber="37">types</typeStatus>
|
||
of
|
||
<taxonomicName id="4C7E4D5CFFAAFF9AFD74CCBDC9DBE0D5" box="[760,947,461,486]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="37" phylum="Arthropoda" rank="species" species="quadraticeps">
|
||
<emphasis id="B90AEACDFFAAFF9AFD74CCBDC9DBE0D5" box="[760,947,461,486]" italics="true" pageId="6" pageNumber="37">P. quadraticeps</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7E4D5CFFAAFF9AFC7BCCBDCEFFE0D5" box="[1015,1175,461,486]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="37" phylum="Arthropoda" rank="species" species="tenebrosus">
|
||
<emphasis id="B90AEACDFFAAFF9AFC7BCCBDCEFFE0D5" box="[1015,1175,461,486]" italics="true" pageId="6" pageNumber="37">P. tenebrosus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13452A5AFFAAFF9AFB20CCBDCF6AE0D4" box="[1196,1282,461,487]" captionStart="FIGURES 1 – 12" captionStartId="6.[151,275,1634,1658]" captionText="FIGURES 1 – 12. Type specimens: habitus and labels; labels with lectotype designation are not included. (1 a – b) Philonthus argentinus Bernhauer (now in Gabrius), (lectotype with labels *). (2 a – b) Philonthus jujuyensis Bernhauer (now in Gabrius), (lectotype with labels *). (3 a – b) Philonthus tucumanensis Bernhauer (now in Gabrius), (lectotype with labels *). (4) Philonthus nidicola Bernhauer (now in Gabrius), (only labels are shown *). (5 a – b) Philonthus quadraticeps (lectotype with labels *). (6 a – b) P. tenebrosus (= P. quadraticeps), (lectotype with labels *). (7 a – b) Philonthus bonariensis (lectotype with labels *). (8 a – b) Philonthus weiseri (= P. bonariensis), (lectotype with labels *). (9 a – b) Philonthus catamarcanus (= P. bonariensis), (lectotype with labels *). (10 a – b) Philonthus cribriventris (lectotype with labels *). (11) Philonthus catamarcanus var. densior (= P. cribriventris), (lectotype, labels are not shown). (12) G. chiliensis (= G. nigritulus), (paratype, labels are not shown). * Labels with lectotype designation are not included (pictures were taken prior to any designation)." pageId="6" pageNumber="37">Figs. 5</figureCitation>
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a–b, 6 a–b) revealed that both are identical in all external characters and identical in characters of the male genitalia. Difference between the two nominal species as described by
|
||
<bibRefCitation id="EFEF4B2EFFAAFF9AFCE1CF6ACE0BE307" author="Boheman" box="[877,1123,538,564]" pageId="6" pageNumber="41" refString="Boheman, C. H. (1858) Coleoptera. Species novas descripsit. In: Virgin, C. (Ed.), Kongliga Svenska Fregatten Eugenies resa omkring Jorden, Zoologi I, Insecta. P. A. Norstedt & Soner, Stockholm, pp. 1 - 112." type="book chapter" year="1858">Boheman (1858: 28)</bibRefCitation>
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results mostly from slight variation in coloration of the antennae, mouthparts and abdomen. Since the
|
||
<typeStatus id="54C5887DFFAAFF9AFB85CF31CEE8E368" box="[1033,1152,577,603]" pageId="6" pageNumber="37" type="lectotype">lectotypes</typeStatus>
|
||
of both nominal species are conspecific and these two names were simultaneously established by
|
||
<bibRefCitation id="EFEF4B2EFFAAFF9AFBC0CF18CF4FE3B1" author="Boheman" box="[1100,1319,616,642]" pageId="6" pageNumber="41" refString="Boheman, C. H. (1858) Coleoptera. Species novas descripsit. In: Virgin, C. (Ed.), Kongliga Svenska Fregatten Eugenies resa omkring Jorden, Zoologi I, Insecta. P. A. Norstedt & Soner, Stockholm, pp. 1 - 112." type="book chapter" year="1858">Boheman (1858)</bibRefCitation>
|
||
, they are considered to be subjective synonyms. Under application of the article 24.2.2 of the Code (
|
||
<bibRefCitation id="EFEF4B2EFFAAFF9AFB35CFFECF2DE39B" author="ICZN" box="[1209,1349,654,680]" pageId="6" pageNumber="42" refString="ICZN (1999) International Code of Zoological Nomenclatur e. Fourth Edition. The International Trust for Zoological Nomenclature, London, xxix + 306 pp." type="book" year="1999">ICZN 1999</bibRefCitation>
|
||
), I give precedence to the name
|
||
<taxonomicName id="4C7E4D5CFFAAFF9AFE30CFC5C810E3FD" box="[444,632,693,718]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="37" phylum="Arthropoda" rank="species" species="quadraticeps">
|
||
<emphasis id="B90AEACDFFAAFF9AFE30CFC5C810E3FD" box="[444,632,693,718]" italics="true" pageId="6" pageNumber="37">P. quadraticeps</emphasis>
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||
</taxonomicName>
|
||
as valid name for the species and senior subjective synonym of
|
||
<emphasis id="B90AEACDFFAAFF9AFA0ACFC5CB75E3C6" italics="true" pageId="6" pageNumber="37">
|
||
<taxonomicName id="4C7E4D5CFFAAFF9AFA0ACFC5CB71E3C6" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="37" phylum="Arthropoda" rank="species" species="tenebrosus">P. tenebrosus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |