treatments-xml/data/95/42/87/954287CDB715FFAB69F0FA3246415C9F.xml
2024-06-21 12:44:36 +02:00

797 lines
86 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="68714F12324B9E047A3CE5A549B0244E" ID-DOI="10.5281/zenodo.10831085" ID-ISSN="1942-1354" ID-Zenodo-Dep="10831085" ID-ZooBank="C1D5B819-A964-4679-B090-84CDBBC59D6A" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1688389961601" checkinUser="felipe" docAuthor="Otto, Robert L., Muona, Jyrki &amp; Córdoba-Alfaro, Jim" docDate="2023" docId="954287CDB715FFAB69F0FA3246415C9F" docLanguage="en" docName="InsectaMundi.2023.0991.1-36.pdf" docOrigin="Insecta Mundi 2023 (991)" docStyle="DocumentStyle:F9976E9B7F8A094D890004515A7A0592.4:InsectaMundi.2020-.journal_article" docStyleId="F9976E9B7F8A094D890004515A7A0592" docStyleName="InsectaMundi.2020-.journal_article" docStyleVersion="4" docTitle="Dromaeolus brunneus Otto, Muona and Cordoba-Alfaro 2023, new species" docType="treatment" docVersion="4" lastPageNumber="27" masterDocId="697BFFB5B70FFFB76930FFD643695A4D" masterDocTitle="A new genus and sixteen new species of false click beetles (Coleoptera: Eucnemidae) described from the Heredia Province of Costa Rica with several additional records from the Osa Peninsula and Panama" masterLastPageNumber="36" masterPageNumber="1" pageNumber="25" updateTime="1710774778166" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-3.0">
<mods:mods id="AA5B0714F982EDB9799F35083D521915" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3D7F94D4A41208C75D792E677A8105C9">
<mods:title id="A15E612203AEB8354FF23E2998195AFC">A new genus and sixteen new species of false click beetles (Coleoptera: Eucnemidae) described from the Heredia Province of Costa Rica with several additional records from the Osa Peninsula and Panama</mods:title>
</mods:titleInfo>
<mods:name id="1275A990BA6A642B8CAB3D9DC6A23F8B" type="personal">
<mods:role id="368E33BBACAA67F4A4BA89AB2319B180">
<mods:roleTerm id="334325B0498295531C75855CCCAA3BCB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D5F91FAE3106657C0B49DF1CFAEA0AC4">Otto, Robert L.</mods:namePart>
<mods:affiliation id="68B032DA6E56074E5E5522EBA489AEB1">W 4806 Chrissie Circle, Shawano, WI 54166, U. S. A.</mods:affiliation>
</mods:name>
<mods:name id="0B6677A4B15613AA1E757EE3A091C8D8" type="personal">
<mods:role id="EE00D5F2B26731FFA2ED22B78E9134B8">
<mods:roleTerm id="92DE085530986061E9EB4E1C3C62FC89">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="569D02D0C911E3D36892F545B25E8D21">Muona, Jyrki</mods:namePart>
<mods:affiliation id="D5B1AD013CC41E89843CDDD989332ABF">Finnish Museum of Natural History, Zoology Unit PO Box 17, University of Helsinki, Finland FIN- 00014</mods:affiliation>
</mods:name>
<mods:name id="6CC14EB19790AB4F8F243A10AEA89D81" type="personal">
<mods:role id="FD082514127A266692B612E04906696E">
<mods:roleTerm id="52225650789AFD6F6A244561FAF5AB5C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="02F3DFFF4CD77947B00272DA6407E122">Córdoba-Alfaro, Jim</mods:namePart>
<mods:affiliation id="6BAC2CC5ECABF5E0CA1306175C3178D8">Insectopia Insect Museum, BioSur Foundation Puerto Jiménez, Península de Osa, Costa Rica jim. cordoba @ biosurcorcovado. org</mods:affiliation>
</mods:name>
<mods:typeOfResource id="6B03B448726CE582EE7C5BF712FA001A">text</mods:typeOfResource>
<mods:relatedItem id="E135680227ECCE01560F3063ED263184" type="host">
<mods:titleInfo id="8B5EB6CBF05BE7C6EF5BAC05B607EBF4">
<mods:title id="BA39A0172EC467C7CEBAD27E7BDF542D">Insecta Mundi</mods:title>
</mods:titleInfo>
<mods:part id="8F61A7355496C6C347929BF838E8FEF6">
<mods:date id="E09A7620720589681365B8463CE906D2">2023</mods:date>
<mods:detail id="9CC0A6A5D84326262B83D4F025805DE3" type="pubDate">
<mods:number id="926277E81BC9A6E6027542E2157981A1">2023-05-26</mods:number>
</mods:detail>
<mods:detail id="C42A2D948B468DE69A6403A5C5D8F367" type="volume">
<mods:number id="0B8D75437DE556628E1D9945FD7B7E10">2023</mods:number>
</mods:detail>
<mods:detail id="A67575E1B1ECA2E582796F55F6755E18" type="issue">
<mods:number id="70D31C1A530BCC4C9E4A56B73A7DD705">991</mods:number>
</mods:detail>
<mods:extent id="F74CCE1A25BB0DEC970659198A1D9E08" unit="page">
<mods:start id="4478F02BC38EFC6298E21C467482C715">1</mods:start>
<mods:end id="465D8AF6CA644DF2D10D28BB5FFE8EEE">36</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="EE1FF37D5EC23C88A93DAA30E5BD71D4">journal article</mods:classification>
<mods:identifier id="B8128C0367DEF2BBD3C8ABF28D317BE2" type="DOI">10.5281/zenodo.10831085</mods:identifier>
<mods:identifier id="1EA394897A0C866C1FE766432629A53D" type="ISSN">1942-1354</mods:identifier>
<mods:identifier id="96C988897AA2FCCEADF1ED213AAE59AB" type="Zenodo-Dep">10831085</mods:identifier>
<mods:identifier id="18F0B717A68BB50883A4742A5409002B" type="ZooBank">C1D5B819-A964-4679-B090-84CDBBC59D6A</mods:identifier>
</mods:mods>
<treatment id="954287CDB715FFAB69F0FA3246415C9F" ID-DOI="http://doi.org/10.5281/zenodo.8109103" ID-Zenodo-Dep="8109103" LSID="urn:lsid:plazi:treatment:954287CDB715FFAB69F0FA3246415C9F" httpUri="http://treatment.plazi.org/id/954287CDB715FFAB69F0FA3246415C9F" lastPageId="28" lastPageNumber="27" pageId="26" pageNumber="25">
<subSubSection id="55F16550B715FFAD69F0FA3247585C4F" box="[192,1073,1508,1538]" pageId="26" pageNumber="25" type="nomenclature">
<paragraph id="1D5436DBB715FFAD69F0FA3247585C4F" blockId="26.[192,1073,1508,1572]" box="[192,1073,1508,1538]" pageId="26" pageNumber="25">
<heading id="461C81B7B715FFAD69F0FA3247585C4F" box="[192,1073,1508,1538]" fontSize="12" level="2" pageId="26" pageNumber="25" reason="2">
<emphasis id="2F9FEAC9B715FFAD69F0FA3247585C4F" bold="true" box="[192,1073,1508,1538]" pageId="26" pageNumber="25">
<taxonomicName id="DAEB4D58B715FFAD69F0FA3240E75C4C" authority="Otto, Muona and Cordoba-Alfaro" authorityName="Otto, Muona and Cordoba-Alfaro" authorityYear="2023" box="[192,910,1508,1538]" class="Insecta" family="Eucnemidae" genus="Dromaeolus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="25" phylum="Arthropoda" rank="species" species="brunneus" status="sp. nov.">
<emphasis id="2F9FEAC9B715FFAD69F0FA3242BE5C4C" bold="true" box="[192,471,1508,1537]" italics="true" pageId="26" pageNumber="25">Dromaeolus brunneus</emphasis>
Otto, Muona and Córdoba-Alfaro
</taxonomicName>
,
<taxonomicNameLabel id="34AC57B2B715FFAD6AACFA3347585C4F" box="[924,1073,1509,1538]" pageId="26" pageNumber="25" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="55F16550B715FFAD69F0F9DF42375C69" box="[192,350,1545,1572]" pageId="26" pageNumber="25" type="description">
<paragraph id="1D5436DBB715FFAD69F0F9DF42375C69" blockId="26.[192,1073,1508,1572]" box="[192,350,1545,1572]" pageId="26" pageNumber="25">
<figureCitation id="85D02A5EB715FFAD69F0F9DF42375C69" box="[192,350,1545,1572]" captionStart-0="Figures 5457" captionStart-1="Figures 5862" captionStartId-0="26.[192,279,709,735]" captionStartId-1="28.[192,279,1209,1235]" captionTargetBox-0="[219,1413,213,679]" captionTargetBox-1="[455,1176,213,1179]" captionTargetId-0="figure-487@26.[219,1414,213,679]" captionTargetId-1="figure-277@28.[455,1176,213,1179]" captionTargetPageId-0="26" captionTargetPageId-1="28" captionText-0="Figures 5457. Costa Rican Dromaeolus species. 54) Dromaeolus americanus sp. nov., dorsal habitus. 55) Dromaeolus americanus sp. nov., ventral habitus. 56) Dromaeolus brunneus sp. nov., male holotype, dorsal habitus. 57) Dromaeolus brunneus sp. nov., male holotype, ventral habitus. (Scale: 5457 = 1.0 mm)" captionText-1="Figures 5862. Costa Rican Dromaeolus species. 58) Dromaeolus brunneus sp. nov., female allotype, dorsal habitus. 59) Dromaeolus herediensis sp. nov., dorsal habitus. 60) Dromaeolus herediensis sp. nov., ventral habitus. 61) Dromaeolus holdridgei sp. nov., dorsal habitus. 62) Dromaeolus holdridgei sp. nov., ventral habitus. (Scale: 5962 = 1.0 mm; 58 = 5.0 mm)" figureDoi-0="http://doi.org/10.5281/zenodo.10831109" figureDoi-1="http://doi.org/10.5281/zenodo.10831111" httpUri-0="https://zenodo.org/record/10831109/files/figure.png" httpUri-1="https://zenodo.org/record/10831111/files/figure.png" pageId="26" pageNumber="25">Figures 5658</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="55F16550B715FFAD69F0F9E447FD5CFF" pageId="26" pageNumber="25" type="diagnosis">
<paragraph id="1D5436DBB715FFAD69F0F9E447FD5CFF" blockId="26.[192,1440,1586,1715]" pageId="26" pageNumber="25">
<emphasis id="2F9FEAC9B715FFAD69F0F9E442555C01" bold="true" box="[192,316,1586,1612]" pageId="26" pageNumber="25">Diagnosis.</emphasis>
The new eucnemid species can be readily distinguished by its elongate form, brown colored dorsum and presence of interantennal carina from any
<taxonomicName id="DAEB4D58B715FFAD6BF6F98340C95C22" box="[710,928,1621,1647]" pageId="26" pageNumber="25">
<emphasis id="2F9FEAC9B715FFAD6BF6F98340225C22" box="[710,843,1621,1647]" italics="true" pageId="26" pageNumber="25">Dromaeolus</emphasis>
species
</taxonomicName>
present in
<collectingCountry id="65FC764BB715FFAD6D2BF98347FC5C22" box="[1051,1173,1621,1647]" name="Costa Rica" pageId="26" pageNumber="25">Costa Rica</collectingCountry>
. The new species is very similar to the Nearctic
<taxonomicName id="DAEB4D58B715FFAD688EF9A141AD5CDC" authority="(Melsheimer)" baseAuthorityName="Melsheimer" baseAuthorityYear="1845" box="[446,708,1655,1681]" class="Insecta" family="Eucnemidae" genus="Dromaeolus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="25" phylum="Arthropoda" rank="species" species="badius">
<emphasis id="2F9FEAC9B715FFAD688EF9A1414E5CDC" box="[446,551,1655,1681]" italics="true" pageId="26" pageNumber="25">D. badius</emphasis>
(Melsheimer)
</taxonomicName>
and
<taxonomicName id="DAEB4D58B715FFAD6BCBF9A1408D5CDC" authority="Bonvouloir." authorityName="Bonvouloir." box="[763,996,1655,1681]" class="Insecta" family="Eucnemidae" genus="Dromaeolus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="25" phylum="Arthropoda" rank="species" species="salsus">
<emphasis id="2F9FEAC9B715FFAD6BCBF9A140335CDC" box="[763,858,1655,1681]" italics="true" pageId="26" pageNumber="25">D. salsus</emphasis>
Bonvouloir.
</taxonomicName>
The presence of an interantennal carina and shorter flagellomere II will distinguish the new species from either Nearctic species.
</paragraph>
</subSubSection>
<subSubSection id="55F16550B715FFAC69F0F91541265EC5" lastPageId="27" lastPageNumber="26" pageId="26" pageNumber="25" type="materials_examined">
<paragraph id="1D5436DBB715FFAD69F0F91547E85DCB" blockId="26.[192,1441,1730,1927]" pageId="26" pageNumber="25">
<emphasis id="2F9FEAC9B715FFAD69F0F91542C55C90" bold="true" box="[192,428,1730,1757]" pageId="26" pageNumber="25">Type material. Male</emphasis>
<materialsCitation id="AD833C86B715FFAD6883F91547145DCB" collectingDate="1993-09-16" collectingDateMax="1994-03-15" collectingDateMin="1993-09-16" collectionCode="ALAS, R, INBC" collectorName="L. Otto" country="Costa Rica" county="Est. Bio. La Selva" elevation="150" latitude="10.433333" location="Est. Bio. La Selva" longLatPrecision="1298" longitude="-84.01667" municipality="Parcelas" pageId="26" pageNumber="25" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Heredia" typeStatus="holotype">
<emphasis id="2F9FEAC9B715FFAD6883F91541735C90" bold="true" box="[435,538,1731,1757]" pageId="26" pageNumber="25">
<typeStatus id="C2508879B715FFAD6883F91541735C90" box="[435,538,1731,1757]" pageId="26" pageNumber="25" type="holotype">holotype</typeStatus>
</emphasis>
: “
<collectingCountry id="65FC764BB715FFAD6B02F91541BF5C90" box="[562,726,1731,1757]" name="Costa Rica" pageId="26" pageNumber="25">COSTA RICA</collectingCountry>
:
<collectingRegion id="DF2FF839B715FFAD6BD1F91440545C91" box="[737,829,1730,1756]" country="Costa Rica" name="Heredia" pageId="26" pageNumber="25">Heredia</collectingRegion>
;
<collectingCounty id="F4354E57B715FFAD6A79F914476C5C91" box="[841,1029,1730,1757]" pageId="26" pageNumber="25">Est.Bio. La Selva</collectingCounty>
, 50;
<quantity id="DA139B3EB715FFAD6D79F91447F95C90" box="[1097,1168,1730,1757]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="26" pageNumber="25" unit="m" value="150.0">
<elevation id="96C6D1E8B715FFAD6D79F91447F95C90" box="[1097,1168,1730,1757]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="26" pageNumber="25" unit="m" value="150.0">150 m</elevation>
</quantity>
<geoCoordinate id="78DF501CB715FFAD6DA8F914479A5C90" box="[1176,1267,1730,1757]" degrees="10" direction="north" minutes="26" orientation="latitude" pageId="26" pageNumber="25" precision="925" value="10.433333">10°26N</geoCoordinate>
<geoCoordinate id="78DF501CB715FFAD6DCAF91546345C90" box="[1274,1373,1731,1757]" degrees="84" direction="west" minutes="01" orientation="longitude" pageId="26" pageNumber="25" precision="925" value="-84.01667">84°01W</geoCoordinate>
; Proy.
<collectionCode id="7BFAAE1EB715FFAD69F0F932426F5CB3" box="[192,262,1764,1790]" pageId="26" pageNumber="25">ALAS</collectionCode>
, INBio-OET” / “M/01/208;
<date id="6955101BB715FFAD6B08F93240645CB3" box="[568,781,1764,1790]" pageId="26" pageNumber="25" value="1993-09-16">
<collectingDate id="7911E9F3B715FFAD6B08F93240645CB3" box="[568,781,1764,1790]" pageId="26" pageNumber="25" value="1993-09-16">16 Septembre 1993</collectingDate>
</date>
;
<collectingMunicipality id="FD30ACA1B715FFAD6A26F93240195CB3" box="[790,880,1764,1790]" pageId="26" pageNumber="25">Parcelas</collectingMunicipality>
sucesionales” / “
<emphasis id="2F9FEAC9B715FFAD6D19F93247AC5CB3" bold="true" box="[1065,1221,1764,1790]" pageId="26" pageNumber="25">
<typeStatus id="C2508879B715FFAD6D19F93247A95CB3" box="[1065,1216,1764,1790]" pageId="26" pageNumber="25" type="holotype">HOLOTYPE</typeStatus>
:
</emphasis>
;
<taxonomicName id="DAEB4D58B715FFAD6DE1F932463F5CB3" authorityName="Kiesenwetter" authorityYear="1858" box="[1233,1366,1764,1790]" class="Insecta" family="Eucnemidae" genus="Dromaeolus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="2F9FEAC9B715FFAD6DE1F932463F5CB3" box="[1233,1366,1764,1790]" italics="true" pageId="26" pageNumber="25">Dromaeolus</emphasis>
</taxonomicName>
;
<taxonomicName id="DAEB4D58B715FFAD6C51F932439B5D6D" authorityName="Otto, Muona and Cordoba-Alfaro" authorityYear="2023" class="Insecta" family="Eucnemidae" genus="Dromaeolus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="25" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="2F9FEAC9B715FFAD6C51F932439B5D6D" italics="true" pageId="26" pageNumber="25">brunneus</emphasis>
</taxonomicName>
<specimenCount id="0BEDFD52B715FFAD69CBF8DC42795D6D" box="[251,272,1802,1824]" pageId="26" pageNumber="25" type="male"></specimenCount>
; Otto, Muona &amp; Córdoba-Alfaro; Det.
<collectionCode id="7BFAAE1EB715FFAD6BE2F8D1418F5D6C" box="[722,742,1799,1825]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="26" pageNumber="25">R</collectionCode>
.
<collectorName id="B01E530DB715FFAD6BDAF8D140565D6D" box="[746,831,1798,1825]" pageId="26" pageNumber="25">L. Otto</collectorName>
; 2019” (red printed label).
<emphasis id="2F9FEAC9B715FFAD6D45F8D046465D6D" bold="true" box="[1141,1327,1798,1824]" pageId="26" pageNumber="25">
Female
<typeStatus id="C2508879B715FFAD6DE1F8D046465D6D" box="[1233,1327,1798,1824]" pageId="26" pageNumber="25" type="allotype">allotype</typeStatus>
</emphasis>
: “
<collectingCountry id="65FC764BB715FFAD6C78F8D0426D5D0F" name="Costa Rica" pageId="26" pageNumber="25">COSTA RICA</collectingCountry>
:
<collectingRegion id="DF2FF839B715FFAD683DF8FE42005D0F" box="[269,361,1832,1858]" country="Costa Rica" name="Heredia" pageId="26" pageNumber="25">Heredia</collectingRegion>
;
<location id="18346000B715FFAD6843F8FE41455D0F" LSID="urn:lsid:plazi:treatment:954287CDB715FFAB69F0FA3246415C9F:18346000B715FFAD6843F8FE41455D0F" box="[371,556,1832,1859]" country="Costa Rica" county="Est. Bio. La Selva" latitude="10.433333" longLatPrecision="1298" longitude="-84.01667" municipality="Parcelas" name="Est. Bio. La Selva" pageId="26" pageNumber="25" stateProvince="Heredia">Est.Bio. La Selva</location>
, 50;
<quantity id="DA139B3EB715FFAD6B5CF8FE41D85D0E" box="[620,689,1832,1859]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="26" pageNumber="25" unit="m" value="150.0">
<elevation id="96C6D1E8B715FFAD6B5CF8FE41D85D0E" box="[620,689,1832,1859]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="26" pageNumber="25" unit="m" value="150.0">150 m</elevation>
</quantity>
<geoCoordinate id="78DF501CB715FFAD6B87F8FE407B5D0E" box="[695,786,1832,1859]" degrees="10" direction="north" minutes="26" orientation="latitude" pageId="26" pageNumber="25" precision="925" value="10.433333">10°26N</geoCoordinate>
<geoCoordinate id="78DF501CB715FFAD6A28F8FE40115D0E" box="[792,888,1832,1859]" degrees="84" direction="west" minutes="01" orientation="longitude" pageId="26" pageNumber="25" precision="925" value="-84.01667">84°01W</geoCoordinate>
; Proy.
<collectionCode id="7BFAAE1EB715FFAD6A8AF8FE47685D0F" box="[954,1025,1832,1858]" pageId="26" pageNumber="25">ALAS</collectionCode>
, INBio-OET” / “M/10/380;
<date id="6955101BB715FFAD6C06F8FE43915D29" pageId="26" pageNumber="25" value="1994-03-15">
<collectingDate id="7911E9F3B715FFAD6C06F8FE43915D29" pageId="26" pageNumber="25" value="1994-03-15">15 Marzo 1994</collectingDate>
</date>
; Bosque primario” / “
<emphasis id="2F9FEAC9B715FFAD68C3F89C41EE5D29" bold="true" box="[499,647,1866,1892]" pageId="26" pageNumber="25">
<typeStatus id="C2508879B715FFAD68C3F89C41EA5D29" box="[499,643,1866,1892]" pageId="26" pageNumber="25" type="allotype">ALLOTYPE</typeStatus>
:
</emphasis>
;
<taxonomicName id="DAEB4D58B715FFAD6BA7F89C40755D29" authorityName="Kiesenwetter" authorityYear="1858" box="[663,796,1866,1892]" class="Insecta" family="Eucnemidae" genus="Dromaeolus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="2F9FEAC9B715FFAD6BA7F89C40755D29" box="[663,796,1866,1892]" italics="true" pageId="26" pageNumber="25">Dromaeolus</emphasis>
</taxonomicName>
;
<taxonomicName id="DAEB4D58B715FFAD6A1BF89C40FB5D29" authorityName="Otto, Muona and Cordoba-Alfaro" authorityYear="2023" box="[811,914,1866,1892]" class="Insecta" family="Eucnemidae" genus="Dromaeolus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="25" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="2F9FEAC9B715FFAD6A1BF89C40FB5D29" box="[811,914,1866,1892]" italics="true" pageId="26" pageNumber="25">brunneus</emphasis>
</taxonomicName>
<specimenCount id="0BEDFD52B715FFAD6AABF89840D95D29" box="[923,944,1870,1892]" pageId="26" pageNumber="25" type="female"></specimenCount>
; Otto, Muona &amp; Córdoba-Alfaro; Det.
<collectionCode id="7BFAAE1EB715FFAD6C42F89D46EF5D28" box="[1394,1414,1867,1893]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="26" pageNumber="25">R</collectionCode>
.
<collectorName id="B01E530DB715FFAD6CBAF89D439F5DCB" pageId="26" pageNumber="25">L. Otto</collectorName>
; 2019” (yellow printed label).
<typeStatus id="C2508879B715FFAD6B74F8BA41C55DCB" box="[580,684,1900,1926]" pageId="26" pageNumber="25" type="holotype">Holotype</typeStatus>
and
<typeStatus id="C2508879B715FFAD6BD2F8BA40525DCB" box="[738,827,1900,1926]" pageId="26" pageNumber="25" type="allotype">allotype</typeStatus>
are to be deposited in
<collectionCode id="7BFAAE1EB715FFAD6D08F8BA47145DCB" box="[1080,1149,1900,1926]" country="Costa Rica" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="26" pageNumber="25">INBC</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="1D5436DBB714FFAC6830FF0041265EC5" blockId="27.[192,1441,213,1161]" pageId="27" pageNumber="26">
<materialsCitation id="AD833C86B714FFAC6830FF00460D5B79" collectingDate="1993-03-03" collectionCode="ALAS, R, INBC" collectorName="L. Otto" country="Costa Rica" elevation="150" latitude="10.433333" location="Est. Bio. La Selva" longLatPrecision="1298" longitude="-84.01667" pageId="27" pageNumber="26" specimenCount="13" specimenCount-female="10" specimenCount-male="3" stateProvince="Heredia" typeStatus="paratype">
<emphasis id="2F9FEAC9B714FFAC6830FF0042135ABD" bold="true" box="[256,378,214,240]" pageId="27" pageNumber="26">
<typeStatus id="C2508879B714FFAC6830FF00421C5ABD" box="[256,373,214,240]" pageId="27" pageNumber="26" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="0BEDFD52B714FFAC684FFF0042D25AA2" box="[383,443,214,240]" pageId="27" pageNumber="26" type="male">3 ♂♂</specimenCount>
<specimenCount id="0BEDFD52B714FFAC68F1FF0042945AA2" box="[449,509,214,240]" pageId="27" pageNumber="26" type="female">8 ♀♀</specimenCount>
:
<emphasis id="2F9FEAC9B714FFAC6B3AFF0341DD5AA2" bold="true" box="[522,692,213,239]" pageId="27" pageNumber="26">
<collectingCountry id="65FC764BB714FFAC6B3AFF0341C65AA2" box="[522,687,213,239]" name="Costa Rica" pageId="27" pageNumber="26">COSTA RICA</collectingCountry>
:
</emphasis>
<specimenCount id="0BEDFD52B714FFAC6B89FF0041895AA2" box="[697,736,214,240]" pageId="27" pageNumber="26" type="female">1 ♀</specimenCount>
, “COSTA RICA:
<collectingRegion id="DF2FF839B714FFAC6A90FF0040955ABD" box="[928,1020,214,240]" country="Costa Rica" name="Heredia" pageId="27" pageNumber="26">Heredia</collectingRegion>
;
<location id="18346000B714FFAC6D35FF0047D45ABD" LSID="urn:lsid:plazi:treatment:954287CDB715FFAB69F0FA3246415C9F:18346000B714FFAC6D35FF0047D45ABD" box="[1029,1213,214,240]" country="Costa Rica" latitude="10.433333" longLatPrecision="1298" longitude="-84.01667" name="Est. Bio. La Selva" pageId="27" pageNumber="26" stateProvince="Heredia">Est.Bio. La Selva</location>
, 50;
<quantity id="DA139B3EB714FFAC6DCCFF0046295ABD" box="[1276,1344,214,241]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="27" pageNumber="26" unit="m" value="150.0">
<elevation id="96C6D1E8B714FFAC6DCCFF0046295ABD" box="[1276,1344,214,241]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="27" pageNumber="26" unit="m" value="150.0">150 m</elevation>
</quantity>
<geoCoordinate id="78DF501CB714FFAC6C76FF0046C95ABD" box="[1350,1440,214,240]" degrees="10" direction="north" minutes="26" orientation="latitude" pageId="27" pageNumber="26" precision="925" value="10.433333">10°26N</geoCoordinate>
<geoCoordinate id="78DF501CB714FFAC69F0FF2E42495B5F" box="[192,288,248,274]" degrees="84" direction="west" minutes="01" orientation="longitude" pageId="27" pageNumber="26" precision="925" value="-84.01667">84°01W</geoCoordinate>
; Proy.
<collectionCode id="7BFAAE1EB714FFAC6856FF2E42C45B5F" box="[358,429,248,274]" pageId="27" pageNumber="26">ALAS</collectionCode>
, INBio-OET” / “M/13/028;
<date id="6955101BB714FFAC6BDFFF2E40F75B5F" box="[751,926,248,274]" pageId="27" pageNumber="26" value="1993-03-03">
<collectingDate id="7911E9F3B714FFAC6BDFFF2E40F75B5F" box="[751,926,248,274]" pageId="27" pageNumber="26" value="1993-03-03">03 Marzo 1993</collectingDate>
</date>
; Bosque secundario” / “
<emphasis id="2F9FEAC9B714FFAC6D84FF2E462F5B5F" bold="true" box="[1204,1350,248,274]" pageId="27" pageNumber="26">
<typeStatus id="C2508879B714FFAC6D84FF2E462B5B5F" box="[1204,1346,248,274]" pageId="27" pageNumber="26" type="paratype">PARATYPE</typeStatus>
:
</emphasis>
;
<taxonomicName id="DAEB4D58B714FFAC6C66FF2F426C5B79" authorityName="Kiesenwetter" authorityYear="1858" class="Insecta" family="Eucnemidae" genus="Dromaeolus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="2F9FEAC9B714FFAC6C66FF2F426C5B79" italics="true" pageId="27" pageNumber="26">Dromaeolus</emphasis>
</taxonomicName>
;
<taxonomicName id="DAEB4D58B714FFAC6823FECC42135B79" authorityName="Otto, Muona and Cordoba-Alfaro" authorityYear="2023" box="[275,378,282,308]" class="Insecta" family="Eucnemidae" genus="Dromaeolus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="26" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="2F9FEAC9B714FFAC6823FECC42135B79" box="[275,378,282,308]" italics="true" pageId="27" pageNumber="26">brunneus</emphasis>
</taxonomicName>
<specimenCount id="0BEDFD52B714FFAC68B2FECB42FE5B7E" box="[386,407,285,307]" pageId="27" pageNumber="26" type="female"></specimenCount>
; Otto, Muona &amp; Córdoba-Alfaro; Det.
<collectionCode id="7BFAAE1EB714FFAC6A63FECC400E5B79" box="[851,871,282,308]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="27" pageNumber="26">R</collectionCode>
.
<collectorName id="B01E530DB714FFAC6A5AFECC40D75B79" box="[874,958,282,308]" pageId="27" pageNumber="26">L. Otto</collectorName>
; 2019” (yellow printed label) (
<collectionCode id="7BFAAE1EB714FFAC6C24FECC46355B79" box="[1300,1372,282,308]" country="Costa Rica" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="27" pageNumber="26">INBC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="AD833C86B714FFAC6C5FFECC40625BD7" collectingDate="1993-03-03" collectionCode="ALAS, R, INBC" collectorName="L. Otto" country="Costa Rica" elevation="150" latitude="10.433333" location="Proy." longLatPrecision="1298" longitude="-84.01667" municipality="Est. Bio. La Selva" pageId="27" pageNumber="26" specimenCount="2" specimenCount-female="2" stateProvince="Heredia" typeStatus="paratype">
<specimenCount id="0BEDFD52B714FFAC6C5FFECC46F35B7E" box="[1391,1434,282,308]" pageId="27" pageNumber="26" type="female">1 ♀</specimenCount>
, “COSTA RICA:
<collectingRegion id="DF2FF839B714FFAC6846FEEA42BB5B1B" box="[374,466,316,342]" country="Costa Rica" name="Heredia" pageId="27" pageNumber="26">Heredia</collectingRegion>
;
<collectingMunicipality id="FD30ACA1B714FFAC68ECFEEA41FF5B1B" box="[476,662,316,343]" pageId="27" pageNumber="26">Est.Bio. La Selva</collectingMunicipality>
, 50;
<quantity id="DA139B3EB714FFAC6BE7FEEA40745B1A" box="[727,797,316,343]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="27" pageNumber="26" unit="m" value="150.0">
<elevation id="96C6D1E8B714FFAC6BE7FEEA40745B1A" box="[727,797,316,343]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="27" pageNumber="26" unit="m" value="150.0">150 m</elevation>
</quantity>
<geoCoordinate id="78DF501CB714FFAC6A14FEEA40165B1A" box="[804,895,316,343]" degrees="10" direction="north" minutes="26" orientation="latitude" pageId="27" pageNumber="26" precision="925" value="10.433333">10°26N</geoCoordinate>
<geoCoordinate id="78DF501CB714FFAC6AB5FEEA408F5B1A" box="[901,998,316,343]" degrees="84" direction="west" minutes="01" orientation="longitude" pageId="27" pageNumber="26" precision="925" value="-84.01667">84°01W</geoCoordinate>
;
<location id="18346000B714FFAC6AC1FEEA47405B1B" LSID="urn:lsid:plazi:treatment:954287CDB715FFAB69F0FA3246415C9F:18346000B714FFAC6AC1FEEA47405B1B" box="[1009,1065,316,342]" country="Costa Rica" latitude="10.433333" longLatPrecision="1298" longitude="-84.01667" municipality="Est. Bio. La Selva" name="Proy." pageId="27" pageNumber="26" stateProvince="Heredia">Proy.</location>
<collectionCode id="7BFAAE1EB714FFAC6D18FEEA47065B1B" box="[1064,1135,316,342]" pageId="27" pageNumber="26">ALAS</collectionCode>
, INBio-OET” / “M/10/025;
<date id="6955101BB714FFAC69F0FE8842045B35" box="[192,365,350,377]" pageId="27" pageNumber="26" value="1993-03-03">
<collectingDate id="7911E9F3B714FFAC69F0FE8842045B35" box="[192,365,350,377]" pageId="27" pageNumber="26" value="1993-03-03">03 Marzo 1993</collectingDate>
</date>
; Bosque primario” / “
<emphasis id="2F9FEAC9B714FFAC6B56FE8841905B35" bold="true" box="[614,761,350,376]" pageId="27" pageNumber="26">
<typeStatus id="C2508879B714FFAC6B56FE88419D5B35" box="[614,756,350,376]" pageId="27" pageNumber="26" type="paratype">PARATYPE</typeStatus>
:
</emphasis>
;
<taxonomicName id="DAEB4D58B714FFAC6A38FE8840E45B35" authorityName="Kiesenwetter" authorityYear="1858" box="[776,909,350,376]" class="Insecta" family="Eucnemidae" genus="Dromaeolus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="2F9FEAC9B714FFAC6A38FE8840E45B35" box="[776,909,350,376]" italics="true" pageId="27" pageNumber="26">Dromaeolus</emphasis>
</taxonomicName>
;
<taxonomicName id="DAEB4D58B714FFAC6AABFE88476B5B35" authorityName="Otto, Muona and Cordoba-Alfaro" authorityYear="2023" box="[923,1026,350,376]" class="Insecta" family="Eucnemidae" genus="Dromaeolus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="26" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="2F9FEAC9B714FFAC6AABFE88476B5B35" box="[923,1026,350,376]" italics="true" pageId="27" pageNumber="26">brunneus</emphasis>
</taxonomicName>
<specimenCount id="0BEDFD52B714FFAC6D3BFEB447495B35" box="[1035,1056,354,376]" pageId="27" pageNumber="26" type="female"></specimenCount>
; Otto, Muona &amp; Córdoba-Alfaro; Det.
<collectionCode id="7BFAAE1EB714FFAC69C9FE5742645BD6" box="[249,269,385,411]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="27" pageNumber="26">R</collectionCode>
.
<collectorName id="B01E530DB714FFAC6821FE57420C5BD7" box="[273,357,384,411]" pageId="27" pageNumber="26">L. Otto</collectorName>
; 2019” (yellow printed label) (
<collectionCode id="7BFAAE1EB714FFAC6B8CFE56406D5BD7" box="[700,772,384,410]" country="Costa Rica" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="27" pageNumber="26">INBC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="AD833C86B714FFAC6A27FE56464C5B93" collectingDate="1993-04-01" collectionCode="R, INBC" collectorName="L. Otto" country="Costa Rica" latitude="10.433333" location="La Selva Bio. Sta." longLatPrecision="1298" longitude="-84.01667" municipality="Pr" pageId="27" pageNumber="26" specimenCount="2" specimenCount-female="2" stateProvince="Heredia" typeStatus="paratype">
<specimenCount id="0BEDFD52B714FFAC6A27FE5640565BD7" box="[791,831,384,410]" pageId="27" pageNumber="26" type="female">1 ♀</specimenCount>
, “COSTA RICA:
<collectingRegion id="DF2FF839B714FFAC6D36FE56470B5BD7" box="[1030,1122,384,410]" country="Costa Rica" name="Heredia" pageId="27" pageNumber="26">Heredia</collectingRegion>
;
<collectingMunicipality id="FD30ACA1B714FFAC6D5EFE5747E25BD6" box="[1134,1163,385,411]" pageId="27" pageNumber="26">Pr</collectingMunicipality>
:
<location id="18346000B714FFAC6DA7FE5746335BD7" LSID="urn:lsid:plazi:treatment:954287CDB715FFAB69F0FA3246415C9F:18346000B714FFAC6DA7FE5746335BD7" box="[1175,1370,384,411]" country="Costa Rica" latitude="10.433333" longLatPrecision="1298" longitude="-84.01667" municipality="Pr" name="La Selva Bio. Sta." pageId="27" pageNumber="26" stateProvince="Heredia">La Selva Bio. Sta.</location>
;
<locationDeviation id="0CE62FF9B714FFAC6C56FE5642225BF1" location="Pto. Viejo" pageId="27" pageNumber="26">
<quantity id="DA139B3EB714FFAC6C56FE5646C95BD7" box="[1382,1440,384,410]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="27" pageNumber="26" unit="km" value="3.0">3 km</quantity>
S Pto. Viejo
</locationDeviation>
;
<geoCoordinate id="78DF501CB714FFAC6868FE7442D85BF0" box="[344,433,418,445]" degrees="10" direction="north" minutes="26" orientation="latitude" pageId="27" pageNumber="26" precision="925" value="10.433333">10°26N</geoCoordinate>
<geoCoordinate id="78DF501CB714FFAC688AFE7441725BF1" box="[442,539,418,444]" degrees="84" direction="west" minutes="01" orientation="longitude" pageId="27" pageNumber="26" precision="925" value="-84.01667">84°01W</geoCoordinate>
” / “
<date id="6955101BB714FFAC6B7FFE7441B75BF1" box="[591,734,418,444]" pageId="27" pageNumber="26" value="1993-04-01">
<collectingDate id="7911E9F3B714FFAC6B7FFE7441B75BF1" box="[591,734,418,444]" pageId="27" pageNumber="26" value="1993-04-01">1 Abril 1993</collectingDate>
</date>
; Bosque secundario; M/02/049” / “
<emphasis id="2F9FEAC9B714FFAC6D42FE74466C5BF1" bold="true" box="[1138,1285,418,444]" pageId="27" pageNumber="26">
<typeStatus id="C2508879B714FFAC6D42FE7446695BF1" box="[1138,1280,418,444]" pageId="27" pageNumber="26" type="paratype">PARATYPE</typeStatus>
:
</emphasis>
;
<taxonomicName id="DAEB4D58B714FFAC6C25FE7446F35BF1" authorityName="Kiesenwetter" authorityYear="1858" box="[1301,1434,418,444]" class="Insecta" family="Eucnemidae" genus="Dromaeolus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="2F9FEAC9B714FFAC6C25FE7446F35BF1" box="[1301,1434,418,444]" italics="true" pageId="27" pageNumber="26">Dromaeolus</emphasis>
</taxonomicName>
;
<taxonomicName id="DAEB4D58B714FFAC69F0FE12424E5B93" authorityName="Otto, Muona and Cordoba-Alfaro" authorityYear="2023" box="[192,295,452,478]" class="Insecta" family="Eucnemidae" genus="Dromaeolus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="26" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="2F9FEAC9B714FFAC69F0FE12424E5B93" box="[192,295,452,478]" italics="true" pageId="27" pageNumber="26">brunneus</emphasis>
</taxonomicName>
<specimenCount id="0BEDFD52B714FFAC6801FE1E422F5B93" box="[305,326,456,478]" pageId="27" pageNumber="26" type="female"></specimenCount>
; Otto, Muona &amp; Córdoba-Alfaro; Det.
<collectionCode id="7BFAAE1EB714FFAC6A3BFE1340765B92" box="[779,799,453,479]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="27" pageNumber="26">R</collectionCode>
.
<collectorName id="B01E530DB714FFAC6A13FE1340115B93" box="[803,888,452,479]" pageId="27" pageNumber="26">L. Otto</collectorName>
; 2019” (yellow printed label) (
<collectionCode id="7BFAAE1EB714FFAC6DE6FE1246775B93" box="[1238,1310,452,478]" country="Costa Rica" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="27" pageNumber="26">INBC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="AD833C86B714FFAC6C03FE12460C5808" collectingDate="1993-04-02" collectionCode="ALAS, R, INBC" collectorName="L. Otto" country="Costa Rica" elevation="150" latitude="10.433333" location="Proy." longLatPrecision="1298" longitude="-84.01667" municipality="Est. Bio. La Selva" pageId="27" pageNumber="26" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Heredia" typeStatus="paratype">
<specimenCount id="0BEDFD52B714FFAC6C03FE1246375B93" box="[1331,1374,452,478]" pageId="27" pageNumber="26" type="male">1 ♂</specimenCount>
,
<specimenCount id="0BEDFD52B714FFAC6C5EFE1246F35B93" box="[1390,1434,452,478]" pageId="27" pageNumber="26" type="female">1 ♀</specimenCount>
, “COSTA RICA:
<collectingRegion id="DF2FF839B714FFAC6846FE3042BB584D" box="[374,466,486,512]" country="Costa Rica" name="Heredia" pageId="27" pageNumber="26">Heredia</collectingRegion>
;
<collectingMunicipality id="FD30ACA1B714FFAC68ECFE3041FF584D" box="[476,662,486,513]" pageId="27" pageNumber="26">Est.Bio. La Selva</collectingMunicipality>
, 50;
<quantity id="DA139B3EB714FFAC6BE7FE304074584C" box="[727,797,486,513]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="27" pageNumber="26" unit="m" value="150.0">
<elevation id="96C6D1E8B714FFAC6BE7FE304074584C" box="[727,797,486,513]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="27" pageNumber="26" unit="m" value="150.0">150 m</elevation>
</quantity>
<geoCoordinate id="78DF501CB714FFAC6A14FE304016584C" box="[804,895,486,513]" degrees="10" direction="north" minutes="26" orientation="latitude" pageId="27" pageNumber="26" precision="925" value="10.433333">10°26N</geoCoordinate>
<geoCoordinate id="78DF501CB714FFAC6AB5FE31408F584C" box="[901,998,487,513]" degrees="84" direction="west" minutes="01" orientation="longitude" pageId="27" pageNumber="26" precision="925" value="-84.01667">84°01W</geoCoordinate>
;
<location id="18346000B714FFAC6AC1FE314740584C" LSID="urn:lsid:plazi:treatment:954287CDB715FFAB69F0FA3246415C9F:18346000B714FFAC6AC1FE314740584C" box="[1009,1065,487,513]" country="Costa Rica" latitude="10.433333" longLatPrecision="1298" longitude="-84.01667" municipality="Est. Bio. La Selva" name="Proy." pageId="27" pageNumber="26" stateProvince="Heredia">Proy.</location>
<collectionCode id="7BFAAE1EB714FFAC6D18FE314706584C" box="[1064,1135,487,513]" pageId="27" pageNumber="26">ALAS</collectionCode>
, INBio-OET” / “M/16/063;
<date id="6955101BB714FFAC69F0FDDF4208586E" box="[192,353,520,547]" pageId="27" pageNumber="26" value="1993-04-02">
<collectingDate id="7911E9F3B714FFAC69F0FDDF4208586E" box="[192,353,520,547]" pageId="27" pageNumber="26" value="1993-04-02">02 Abril 1993</collectingDate>
</date>
; Bosque secundario” / “
<emphasis id="2F9FEAC9B714FFAC6B49FDDE4065586F" bold="true" box="[633,780,520,546]" pageId="27" pageNumber="26">
<typeStatus id="C2508879B714FFAC6B49FDDE406E586F" box="[633,775,520,546]" pageId="27" pageNumber="26" type="paratype">PARATYPE</typeStatus>
:
</emphasis>
;
<taxonomicName id="DAEB4D58B714FFAC6A2DFDDE40CB586F" authorityName="Kiesenwetter" authorityYear="1858" box="[797,930,520,546]" class="Insecta" family="Eucnemidae" genus="Dromaeolus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="2F9FEAC9B714FFAC6A2DFDDE40CB586F" box="[797,930,520,546]" italics="true" pageId="27" pageNumber="26">Dromaeolus</emphasis>
</taxonomicName>
;
<taxonomicName id="DAEB4D58B714FFAC6A82FDDE4770586F" authorityName="Otto, Muona and Cordoba-Alfaro" authorityYear="2023" box="[946,1049,520,546]" class="Insecta" family="Eucnemidae" genus="Dromaeolus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="26" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="2F9FEAC9B714FFAC6A82FDDE4770586F" box="[946,1049,520,546]" italics="true" pageId="27" pageNumber="26">brunneus</emphasis>
</taxonomicName>
; Otto, Muona &amp; Córdoba-Alfaro; Det.
<collectionCode id="7BFAAE1EB714FFAC69C6FDFD42635808" box="[246,266,555,581]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="27" pageNumber="26">R</collectionCode>
.
<collectorName id="B01E530DB714FFAC683EFDFD42085808" box="[270,353,555,581]" pageId="27" pageNumber="26">L. Otto</collectorName>
; 2019” (either
<specimenCount id="0BEDFD52B714FFAC6B36FDF841725809" box="[518,539,558,580]" pageId="27" pageNumber="26" type="male"></specimenCount>
or
<specimenCount id="0BEDFD52B714FFAC6B71FDF8413F5809" box="[577,598,558,580]" pageId="27" pageNumber="26" type="female"></specimenCount>
typed behind species name on label) [yellow printed label] (2,
<collectionCode id="7BFAAE1EB714FFAC6C27FDFD46345808" box="[1303,1373,555,581]" country="Costa Rica" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="27" pageNumber="26">INBC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="AD833C86B714FFAC6C40FDFD406858E6" collectingDate="1993-04-15" collectionCode="ALAS, R, INBC" collectorName="L. Otto" country="Costa Rica" elevation="150" latitude="10.433333" location="Proy." longLatPrecision="1298" longitude="-84.01667" municipality="Est. Bio. La Selva" pageId="27" pageNumber="26" specimenCount="2" specimenCount-male="2" stateProvince="Heredia" typeStatus="paratype">
<specimenCount id="0BEDFD52B714FFAC6C40FDFD46F05809" box="[1392,1433,555,581]" pageId="27" pageNumber="26" type="male">1 ♂</specimenCount>
, “COSTA RICA:
<collectingRegion id="DF2FF839B714FFAC6846FD9B42BB582A" box="[374,466,589,615]" country="Costa Rica" name="Heredia" pageId="27" pageNumber="26">Heredia</collectingRegion>
;
<collectingMunicipality id="FD30ACA1B714FFAC68ECFD9B41FF582A" box="[476,662,589,615]" pageId="27" pageNumber="26">Est.Bio. La Selva</collectingMunicipality>
, 50;
<quantity id="DA139B3EB714FFAC6BE7FD9A4074582A" box="[727,797,588,615]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="27" pageNumber="26" unit="m" value="150.0">
<elevation id="96C6D1E8B714FFAC6BE7FD9A4074582A" box="[727,797,588,615]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="27" pageNumber="26" unit="m" value="150.0">150 m</elevation>
</quantity>
<geoCoordinate id="78DF501CB714FFAC6A14FD9B4016582A" box="[804,895,589,615]" degrees="10" direction="north" minutes="26" orientation="latitude" pageId="27" pageNumber="26" precision="925" value="10.433333">10°26N</geoCoordinate>
<geoCoordinate id="78DF501CB714FFAC6AB5FD9B408F582A" box="[901,998,589,615]" degrees="84" direction="west" minutes="01" orientation="longitude" pageId="27" pageNumber="26" precision="925" value="-84.01667">84°01W</geoCoordinate>
;
<location id="18346000B714FFAC6AC1FD9B4740582A" LSID="urn:lsid:plazi:treatment:954287CDB715FFAB69F0FA3246415C9F:18346000B714FFAC6AC1FD9B4740582A" box="[1009,1065,589,615]" country="Costa Rica" latitude="10.433333" longLatPrecision="1298" longitude="-84.01667" municipality="Est. Bio. La Selva" name="Proy." pageId="27" pageNumber="26" stateProvince="Heredia">Proy.</location>
<collectionCode id="7BFAAE1EB714FFAC6D18FD9B4706582A" box="[1064,1135,589,615]" pageId="27" pageNumber="26">ALAS</collectionCode>
, INBio-OET” / “M/04/067;
<date id="6955101BB714FFAC69F0FDB9420958C4" box="[192,352,623,650]" pageId="27" pageNumber="26" value="1993-04-15">
<collectingDate id="7911E9F3B714FFAC69F0FDB9420958C4" box="[192,352,623,650]" pageId="27" pageNumber="26" value="1993-04-15">15 Abril 1993</collectingDate>
</date>
; Bosque primario” / “
<emphasis id="2F9FEAC9B714FFAC6B6EFDB8419958C5" bold="true" box="[606,752,622,648]" pageId="27" pageNumber="26">
<typeStatus id="C2508879B714FFAC6B6EFDB8418558C5" box="[606,748,622,648]" pageId="27" pageNumber="26" type="paratype">PARATYPE</typeStatus>
:
</emphasis>
;
<taxonomicName id="DAEB4D58B714FFAC6A31FDB940EF58C4" authorityName="Kiesenwetter" authorityYear="1858" box="[769,902,623,649]" class="Insecta" family="Eucnemidae" genus="Dromaeolus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="2F9FEAC9B714FFAC6A31FDB940EF58C4" box="[769,902,623,649]" italics="true" pageId="27" pageNumber="26">Dromaeolus</emphasis>
</taxonomicName>
;
<taxonomicName id="DAEB4D58B714FFAC6AA6FDB9409458C4" authorityName="Otto, Muona and Cordoba-Alfaro" authorityYear="2023" box="[918,1021,623,649]" class="Insecta" family="Eucnemidae" genus="Dromaeolus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="26" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="2F9FEAC9B714FFAC6AA6FDB9409458C4" box="[918,1021,623,649]" italics="true" pageId="27" pageNumber="26">brunneus</emphasis>
</taxonomicName>
<specimenCount id="0BEDFD52B714FFAC6D36FDA4477258C5" box="[1030,1051,626,648]" pageId="27" pageNumber="26" type="male"></specimenCount>
; Otto, Muona &amp; Córdoba-Alfaro; Det.
<collectionCode id="7BFAAE1EB714FFAC69C6FD47426358E6" box="[246,266,657,683]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="27" pageNumber="26">R</collectionCode>
.
<collectorName id="B01E530DB714FFAC683EFD47420858E6" box="[270,353,657,683]" pageId="27" pageNumber="26">L. Otto</collectorName>
; 2019” (yellow printed label) (
<collectionCode id="7BFAAE1EB714FFAC6B82FD47419358E6" box="[690,762,657,683]" country="Costa Rica" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="27" pageNumber="26">INBC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="AD833C86B714FFAC6A3CFD47471C58A2" collectingDate="1993-05-18" collectionCode="ALAS" collectorName="Robert L. Otto" country="Costa Rica" elevation="150" latitude="10.433333" location="Est. Bio. La Selva" longLatPrecision="1298" longitude="-84.01667" pageId="27" pageNumber="26" specimenCount="1" specimenCount-female="1" stateProvince="Heredia" typeStatus="paratype">
<specimenCount id="0BEDFD52B714FFAC6A3CFD47405F58E7" box="[780,822,657,683]" pageId="27" pageNumber="26" type="female">1 ♀</specimenCount>
, “COSTA RICA:
<collectingRegion id="DF2FF839B714FFAC6AC8FD47473D58E6" box="[1016,1108,657,683]" country="Costa Rica" name="Heredia" pageId="27" pageNumber="26">Heredia</collectingRegion>
;
<location id="18346000B714FFAC6D6FFD47467058E6" LSID="urn:lsid:plazi:treatment:954287CDB715FFAB69F0FA3246415C9F:18346000B714FFAC6D6FFD47467058E6" box="[1119,1305,657,683]" country="Costa Rica" latitude="10.433333" longLatPrecision="1298" longitude="-84.01667" name="Est. Bio. La Selva" pageId="27" pageNumber="26" stateProvince="Heredia">Est.Bio. La Selva</location>
, 50;
<quantity id="DA139B3EB714FFAC6C6AFD4746C958E6" box="[1370,1440,657,684]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="27" pageNumber="26" unit="m" value="150.0">
<elevation id="96C6D1E8B714FFAC6C6AFD4746C958E6" box="[1370,1440,657,684]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="27" pageNumber="26" unit="m" value="150.0">150 m</elevation>
</quantity>
<geoCoordinate id="78DF501CB714FFAC69F0FD6542735883" box="[192,282,691,718]" degrees="10" direction="north" minutes="26" orientation="latitude" pageId="27" pageNumber="26" precision="925" value="10.433333">10°26N</geoCoordinate>
<geoCoordinate id="78DF501CB714FFAC6812FD6542EA5883" box="[290,387,691,718]" degrees="84" direction="west" minutes="01" orientation="longitude" pageId="27" pageNumber="26" precision="925" value="-84.01667">84°01W</geoCoordinate>
; Proy.
<collectionCode id="7BFAAE1EB714FFAC68F7FD6541675880" box="[455,526,691,717]" pageId="27" pageNumber="26">ALAS</collectionCode>
, INBio-OET” / “M/10/104;
<date id="6955101BB714FFAC6A79FD6540825880" box="[841,1003,691,718]" pageId="27" pageNumber="26" value="1993-05-18">
<collectingDate id="7911E9F3B714FFAC6A79FD6540825880" box="[841,1003,691,718]" pageId="27" pageNumber="26" value="1993-05-18">18 Mayo 1993</collectingDate>
</date>
; Bosque primario” / “Collection of the Global; Eucnemid Research Project; (
<collectorName id="B01E530DB714FFAC6B54FD03406758A2" box="[612,782,725,752]" pageId="27" pageNumber="26">Robert L. Otto</collectorName>
)” (green framed white label) / “
</materialsCitation>
<emphasis id="2F9FEAC9B714FFAC6D44FD03466E58A2" bold="true" box="[1140,1287,725,751]" pageId="27" pageNumber="26">
<typeStatus id="C2508879B714FFAC6D44FD03466B58A2" box="[1140,1282,725,751]" pageId="27" pageNumber="26" type="paratype">PARATYPE</typeStatus>
:
</emphasis>
;
<taxonomicName id="DAEB4D58B714FFAC6C25FD0346F358A2" authorityName="Kiesenwetter" authorityYear="1858" box="[1301,1434,725,751]" class="Insecta" family="Eucnemidae" genus="Dromaeolus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="2F9FEAC9B714FFAC6C25FD0346F358A2" box="[1301,1434,725,751]" italics="true" pageId="27" pageNumber="26">Dromaeolus</emphasis>
</taxonomicName>
;
<taxonomicName id="DAEB4D58B714FFAC69F0FD21424E595C" authorityName="Otto, Muona and Cordoba-Alfaro" authorityYear="2023" box="[192,295,759,785]" class="Insecta" family="Eucnemidae" genus="Dromaeolus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="26" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="2F9FEAC9B714FFAC69F0FD21424E595C" box="[192,295,759,785]" italics="true" pageId="27" pageNumber="26">brunneus</emphasis>
</taxonomicName>
<specimenCount id="0BEDFD52B714FFAC681DFD2D422B595C" box="[301,322,763,785]" pageId="27" pageNumber="26" type="female"></specimenCount>
;
<materialsCitation id="AD833C86B714FFAC687FFD2141DC595C" box="[335,693,759,786]" collectorName="Muona &amp; Cordoba-Alfaro" country="Costa Rica" location="Otto" pageId="27" pageNumber="26" specimenCount="1" stateProvince="Heredia" typeStatus="paratype">
<location id="18346000B714FFAC687FFD2142ED595C" LSID="urn:lsid:plazi:treatment:954287CDB715FFAB69F0FA3246415C9F:18346000B714FFAC687FFD2142ED595C" box="[335,388,759,785]" country="Costa Rica" name="Otto" pageId="27" pageNumber="26" stateProvince="Heredia">Otto</location>
,
<collectorName id="B01E530DB714FFAC68BEFD2E42B7595F" box="[398,478,760,786]" pageId="27" pageNumber="26">Muona</collectorName>
&amp;
<collectorName id="B01E530DB714FFAC68CFFD2141DC595C" box="[511,693,759,785]" pageId="27" pageNumber="26">Córdoba-Alfaro</collectorName>
</materialsCitation>
;
<materialsCitation id="AD833C86B714FFAC6B8FFD2E4036595C" box="[703,863,759,786]" collectionCode="R" collectorName="Det. &amp; L. Otto" country="Costa Rica" pageId="27" pageNumber="26" specimenCount="1" stateProvince="Heredia" typeStatus="paratype">
<collectorName id="B01E530DB714FFAC6B8FFD2E4186595F" box="[703,751,760,786]" pageId="27" pageNumber="26">Det.</collectorName>
<collectionCode id="7BFAAE1EB714FFAC6BC5FD2E4060595F" box="[757,777,760,786]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="27" pageNumber="26">R</collectionCode>
.
<collectorName id="B01E530DB714FFAC6A3DFD2E4036595C" box="[781,863,759,786]" pageId="27" pageNumber="26">L. Otto</collectorName>
</materialsCitation>
; 2019” (yellow printed label) (
<collectionCode id="7BFAAE1EB714FFAC6D9CFD21479E595C" box="[1196,1271,759,785]" pageId="27" pageNumber="26">GERP</collectionCode>
);
<materialsCitation id="AD833C86B714FFAC6C39FD2141F05935" collectingDate="1993-06-01" collectionCode="ALAS, R, INBC" collectorName="L. Otto" country="Costa Rica" elevation="150" latitude="10.433333" location="Proy." longLatPrecision="1298" longitude="-84.01667" municipality="Est. Bio. La Selva" pageId="27" pageNumber="26" specimenCount="2" specimenCount-female="2" stateProvince="Heredia" typeStatus="paratype">
<specimenCount id="0BEDFD52B714FFAC6C39FD21465B595C" box="[1289,1330,759,785]" pageId="27" pageNumber="26" type="female">1 ♀</specimenCount>
, “COSTA RICA:
<collectingRegion id="DF2FF839B714FFAC683EFCCF4203597E" box="[270,362,793,819]" country="Costa Rica" name="Heredia" pageId="27" pageNumber="26">Heredia</collectingRegion>
;
<collectingMunicipality id="FD30ACA1B714FFAC6844FCCF4146597E" box="[372,559,793,820]" pageId="27" pageNumber="26">Est.Bio. La Selva</collectingMunicipality>
, 50;
<quantity id="DA139B3EB714FFAC6B41FCCF41DE5979" box="[625,695,793,820]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="27" pageNumber="26" unit="m" value="150.0">
<elevation id="96C6D1E8B714FFAC6B41FCCF41DE5979" box="[625,695,793,820]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="27" pageNumber="26" unit="m" value="150.0">150 m</elevation>
</quantity>
<geoCoordinate id="78DF501CB714FFAC6B8EFCCF40715979" box="[702,792,793,820]" degrees="10" direction="north" minutes="26" orientation="latitude" pageId="27" pageNumber="26" precision="925" value="10.433333">10°26N</geoCoordinate>
<geoCoordinate id="78DF501CB714FFAC6A2FFCCF40E95979" box="[799,896,793,820]" degrees="84" direction="west" minutes="01" orientation="longitude" pageId="27" pageNumber="26" precision="925" value="-84.01667">84°01W</geoCoordinate>
;
<location id="18346000B714FFAC6ABBFCCF40AD597E" LSID="urn:lsid:plazi:treatment:954287CDB715FFAB69F0FA3246415C9F:18346000B714FFAC6ABBFCCF40AD597E" box="[907,964,793,819]" country="Costa Rica" latitude="10.433333" longLatPrecision="1298" longitude="-84.01667" municipality="Est. Bio. La Selva" name="Proy." pageId="27" pageNumber="26" stateProvince="Heredia">Proy.</location>
<collectionCode id="7BFAAE1EB714FFAC6AF3FCCF4763597E" box="[963,1034,793,819]" pageId="27" pageNumber="26">ALAS</collectionCode>
, INBio-OET” / “M/04/111;
<date id="6955101BB714FFAC6C72FCCF43905918" pageId="27" pageNumber="26" value="1993-06-01">
<collectingDate id="7911E9F3B714FFAC6C72FCCF43905918" pageId="27" pageNumber="26" value="1993-06-01">01 Junio 1993</collectingDate>
</date>
; Bosque primario” / “
<emphasis id="2F9FEAC9B714FFAC68C4FCED41EF5918" bold="true" box="[500,646,827,853]" pageId="27" pageNumber="26">
<typeStatus id="C2508879B714FFAC68C4FCED41EB5918" box="[500,642,827,853]" pageId="27" pageNumber="26" type="paratype">PARATYPE</typeStatus>
:
</emphasis>
;
<taxonomicName id="DAEB4D58B714FFAC6BA6FCED40725918" authorityName="Kiesenwetter" authorityYear="1858" box="[662,795,827,853]" class="Insecta" family="Eucnemidae" genus="Dromaeolus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="2F9FEAC9B714FFAC6BA6FCED40725918" box="[662,795,827,853]" italics="true" pageId="27" pageNumber="26">Dromaeolus</emphasis>
</taxonomicName>
;
<taxonomicName id="DAEB4D58B714FFAC6A1AFCED40F85918" authorityName="Otto, Muona and Cordoba-Alfaro" authorityYear="2023" box="[810,913,827,853]" class="Insecta" family="Eucnemidae" genus="Dromaeolus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="26" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="2F9FEAC9B714FFAC6A1AFCED40F85918" box="[810,913,827,853]" italics="true" pageId="27" pageNumber="26">brunneus</emphasis>
</taxonomicName>
<specimenCount id="0BEDFD52B714FFAC6AAAFCE940C65918" box="[922,943,831,853]" pageId="27" pageNumber="26" type="female"></specimenCount>
; Otto, Muona &amp; Córdoba-Alfaro; Det.
<collectionCode id="7BFAAE1EB714FFAC6C42FCEA46EF591B" box="[1394,1414,828,854]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="27" pageNumber="26">R</collectionCode>
.
<collectorName id="B01E530DB714FFAC6CBAFCEA439F5935" pageId="27" pageNumber="26">L. Otto</collectorName>
; 2019” (yellow printed label) (
<collectionCode id="7BFAAE1EB714FFAC6B7AFC8841FB5935" box="[586,658,862,888]" country="Costa Rica" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="27" pageNumber="26">INBC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="AD833C86B714FFAC6B95FC88479659F1" collectingDate="1994-04-04" collectionCode="ALAS, R, INBC" collectorName="L. Otto" country="Costa Rica" elevation="150" latitude="10.433333" location="Proy." longLatPrecision="1298" longitude="-84.01667" municipality="Est. Bio. La Selva" pageId="27" pageNumber="26" specimenCount="2" specimenCount-male="2" stateProvince="Heredia" typeStatus="paratype">
<specimenCount id="0BEDFD52B714FFAC6B95FC8841A6593A" box="[677,719,862,888]" pageId="27" pageNumber="26" type="male">1 ♂</specimenCount>
, “COSTA RICA:
<collectingRegion id="DF2FF839B714FFAC6AA3FC8B4086593A" box="[915,1007,861,887]" country="Costa Rica" name="Heredia" pageId="27" pageNumber="26">Heredia</collectingRegion>
;
<collectingMunicipality id="FD30ACA1B714FFAC6ACAFC8B47DF593A" box="[1018,1206,861,888]" pageId="27" pageNumber="26">Est.Bio. La Selva</collectingMunicipality>
, 50;
<quantity id="DA139B3EB714FFAC6DC8FC8B46565935" box="[1272,1343,861,888]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="27" pageNumber="26" unit="m" value="150.0">
<elevation id="96C6D1E8B714FFAC6DC8FC8B46565935" box="[1272,1343,861,888]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="27" pageNumber="26" unit="m" value="150.0">150 m</elevation>
</quantity>
<geoCoordinate id="78DF501CB714FFAC6C76FC8B46C95935" box="[1350,1440,861,888]" degrees="10" direction="north" minutes="26" orientation="latitude" pageId="27" pageNumber="26" precision="925" value="10.433333">10°26N</geoCoordinate>
<geoCoordinate id="78DF501CB714FFAC69F0FC56424959D7" box="[192,288,896,922]" degrees="84" direction="west" minutes="01" orientation="longitude" pageId="27" pageNumber="26" precision="925" value="-84.01667">84°01W</geoCoordinate>
;
<location id="18346000B714FFAC681AFC56420A59D7" LSID="urn:lsid:plazi:treatment:954287CDB715FFAB69F0FA3246415C9F:18346000B714FFAC681AFC56420A59D7" box="[298,355,896,922]" country="Costa Rica" latitude="10.433333" longLatPrecision="1298" longitude="-84.01667" municipality="Est. Bio. La Selva" name="Proy." pageId="27" pageNumber="26" stateProvince="Heredia">Proy.</location>
<collectionCode id="7BFAAE1EB714FFAC6852FC5642C059D7" box="[354,425,896,922]" pageId="27" pageNumber="26">ALAS</collectionCode>
, INBio-OET” / “M/11/389;
<date id="6955101BB714FFAC6BEAFC56401859D7" box="[730,881,895,922]" pageId="27" pageNumber="26" value="1994-04-04">
<collectingDate id="7911E9F3B714FFAC6BEAFC56401859D7" box="[730,881,895,922]" pageId="27" pageNumber="26" value="1994-04-04">04 Abril 1994</collectingDate>
</date>
; Bosque secundario” / “
<emphasis id="2F9FEAC9B714FFAC6D46FCA9466059D4" bold="true" box="[1142,1289,895,921]" pageId="27" pageNumber="26">
<typeStatus id="C2508879B714FFAC6D46FCA9466D59D4" box="[1142,1284,895,921]" pageId="27" pageNumber="26" type="paratype">PARATYPE</typeStatus>
:
</emphasis>
;
<taxonomicName id="DAEB4D58B714FFAC6C25FCA946F359D4" authorityName="Kiesenwetter" authorityYear="1858" box="[1301,1434,895,921]" class="Insecta" family="Eucnemidae" genus="Dromaeolus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="2F9FEAC9B714FFAC6C25FCA946F359D4" box="[1301,1434,895,921]" italics="true" pageId="27" pageNumber="26">Dromaeolus</emphasis>
</taxonomicName>
;
<taxonomicName id="DAEB4D58B714FFAC69F0FC77424E59F6" authorityName="Otto, Muona and Cordoba-Alfaro" authorityYear="2023" box="[192,295,929,955]" class="Insecta" family="Eucnemidae" genus="Dromaeolus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="26" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="2F9FEAC9B714FFAC69F0FC77424E59F6" box="[192,295,929,955]" italics="true" pageId="27" pageNumber="26">brunneus</emphasis>
</taxonomicName>
<specimenCount id="0BEDFD52B714FFAC681EFC73422A59F6" box="[302,323,933,955]" pageId="27" pageNumber="26" type="male"></specimenCount>
; Otto, Muona &amp; Córdoba-Alfaro; Det.
<collectionCode id="7BFAAE1EB714FFAC6BC6FC74406359F1" box="[758,778,930,956]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="27" pageNumber="26">R</collectionCode>
.
<collectorName id="B01E530DB714FFAC6A3EFC74400959F1" box="[782,864,930,956]" pageId="27" pageNumber="26">L. Otto</collectorName>
; 2019” (yellow printed label) (
<collectionCode id="7BFAAE1EB714FFAC6D9FFC74479E59F1" box="[1199,1271,930,956]" country="Costa Rica" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="27" pageNumber="26">INBC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="AD833C86B714FFAC6C39FC7441F05E6F" collectingDate="1996-05-02" collectionCode="ALAS, R, INBC" collectorName="L. Otto" country="Costa Rica" elevation="150" latitude="10.433333" location="Proy." longLatPrecision="1298" longitude="-84.01667" municipality="Est. Bio. La Selva" pageId="27" pageNumber="26" specimenCount="2" specimenCount-female="2" stateProvince="Heredia" typeStatus="paratype">
<specimenCount id="0BEDFD52B714FFAC6C39FC74465B59F6" box="[1289,1330,930,956]" pageId="27" pageNumber="26" type="female">1 ♀</specimenCount>
, “COSTA RICA:
<collectingRegion id="DF2FF839B714FFAC683EFC1242035993" box="[270,362,964,990]" country="Costa Rica" name="Heredia" pageId="27" pageNumber="26">Heredia</collectingRegion>
;
<collectingMunicipality id="FD30ACA1B714FFAC6844FC1241475993" box="[372,558,964,990]" pageId="27" pageNumber="26">Est.Bio. La Selva</collectingMunicipality>
, 50;
<quantity id="DA139B3EB714FFAC6B40FC1541DF5993" box="[624,694,963,990]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="27" pageNumber="26" unit="m" value="150.0">
<elevation id="96C6D1E8B714FFAC6B40FC1541DF5993" box="[624,694,963,990]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="27" pageNumber="26" unit="m" value="150.0">150 m</elevation>
</quantity>
<geoCoordinate id="78DF501CB714FFAC6B8CFC12407E5993" box="[700,791,964,990]" degrees="10" direction="north" minutes="26" orientation="latitude" pageId="27" pageNumber="26" precision="925" value="10.433333">10°26N</geoCoordinate>
<geoCoordinate id="78DF501CB714FFAC6A2DFC1240175993" box="[797,894,964,990]" degrees="84" direction="west" minutes="01" orientation="longitude" pageId="27" pageNumber="26" precision="925" value="-84.01667">84°01W</geoCoordinate>
;
<location id="18346000B714FFAC6AB9FC1240AB5993" LSID="urn:lsid:plazi:treatment:954287CDB715FFAB69F0FA3246415C9F:18346000B714FFAC6AB9FC1240AB5993" box="[905,962,964,990]" country="Costa Rica" latitude="10.433333" longLatPrecision="1298" longitude="-84.01667" municipality="Est. Bio. La Selva" name="Proy." pageId="27" pageNumber="26" stateProvince="Heredia">Proy.</location>
<collectionCode id="7BFAAE1EB714FFAC6AF1FC1247615993" box="[961,1032,964,990]" pageId="27" pageNumber="26">ALAS</collectionCode>
, INBio-OET” / “M/10/635;
<date id="6955101BB714FFAC6C0FFC1243915E4D" pageId="27" pageNumber="26" value="1996-05-02">
<collectingDate id="7911E9F3B714FFAC6C0FFC1243915E4D" pageId="27" pageNumber="26" value="1996-05-02">02 Mayo 1996</collectingDate>
</date>
; Bosque primario” / “
<emphasis id="2F9FEAC9B714FFAC68C4FC3341EF59B2" bold="true" box="[500,646,997,1023]" pageId="27" pageNumber="26">
<typeStatus id="C2508879B714FFAC68C4FC3341EB59B2" box="[500,642,997,1023]" pageId="27" pageNumber="26" type="paratype">PARATYPE</typeStatus>
:
</emphasis>
;
<taxonomicName id="DAEB4D58B714FFAC6BA6FC3040725E4D" authorityName="Kiesenwetter" authorityYear="1858" box="[662,795,998,1024]" class="Insecta" family="Eucnemidae" genus="Dromaeolus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="2F9FEAC9B714FFAC6BA6FC3040725E4D" box="[662,795,998,1024]" italics="true" pageId="27" pageNumber="26">Dromaeolus</emphasis>
</taxonomicName>
;
<taxonomicName id="DAEB4D58B714FFAC6A1AFC3040F85E4D" authorityName="Otto, Muona and Cordoba-Alfaro" authorityYear="2023" box="[810,913,998,1024]" class="Insecta" family="Eucnemidae" genus="Dromaeolus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="26" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="2F9FEAC9B714FFAC6A1AFC3040F85E4D" box="[810,913,998,1024]" italics="true" pageId="27" pageNumber="26">brunneus</emphasis>
</taxonomicName>
<specimenCount id="0BEDFD52B714FFAC6AAAFC3F40C659B2" box="[922,943,1001,1023]" pageId="27" pageNumber="26" type="female"></specimenCount>
; Otto, Muona &amp; Córdoba-Alfaro; Det.
<collectionCode id="7BFAAE1EB714FFAC6C42FC3046EF5E4D" box="[1394,1414,998,1024]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="27" pageNumber="26">R</collectionCode>
.
<collectorName id="B01E530DB714FFAC6CBAFC30439F5E6F" pageId="27" pageNumber="26">L. Otto</collectorName>
; 2019” (yellow printed label) (
<collectionCode id="7BFAAE1EB714FFAC6B7AFBDE41FB5E6F" box="[586,658,1032,1058]" country="Costa Rica" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="27" pageNumber="26">INBC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="AD833C86B714FFAC6B95FBDE466B5E2B" collectingDate="2000-04" collectingDateMax="2000-04-17" collectingDateMin="2000-04" collectionCode="R, INBC" collectorName="L. Otto" country="Costa Rica" elevation="150" latitude="10.433333" location="Est. Bio. La Selva" longLatPrecision="1298" longitude="-84.01667" pageId="27" pageNumber="26" specimenCount="2" specimenCount-female="2" stateProvince="Heredia" typeStatus="paratype">
<specimenCount id="0BEDFD52B714FFAC6B95FBDE41A65E6C" box="[677,719,1032,1058]" pageId="27" pageNumber="26" type="female">1 ♀</specimenCount>
, “COSTA RICA:
<collectingRegion id="DF2FF839B714FFAC6AA3FBDE40865E6F" box="[915,1007,1032,1058]" country="Costa Rica" name="Heredia" pageId="27" pageNumber="26">Heredia</collectingRegion>
;
<location id="18346000B714FFAC6ACAFBDE47DF5E6F" LSID="urn:lsid:plazi:treatment:954287CDB715FFAB69F0FA3246415C9F:18346000B714FFAC6ACAFBDE47DF5E6F" box="[1018,1206,1032,1058]" country="Costa Rica" latitude="10.433333" longLatPrecision="1298" longitude="-84.01667" name="Est. Bio. La Selva" pageId="27" pageNumber="26" stateProvince="Heredia">Est.Bio. La Selva</location>
, 50;
<quantity id="DA139B3EB714FFAC6DC8FBDE46565E6F" box="[1272,1343,1032,1059]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="27" pageNumber="26" unit="m" value="150.0">
<elevation id="96C6D1E8B714FFAC6DC8FBDE46565E6F" box="[1272,1343,1032,1059]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="27" pageNumber="26" unit="m" value="150.0">150 m</elevation>
</quantity>
<geoCoordinate id="78DF501CB714FFAC6C76FBDE46C95E6F" box="[1350,1440,1032,1058]" degrees="10" direction="north" minutes="26" orientation="latitude" pageId="27" pageNumber="26" precision="925" value="10.433333">10°26N</geoCoordinate>
<geoCoordinate id="78DF501CB714FFAC69F0FBFC42495E09" box="[192,288,1066,1092]" degrees="84" direction="west" minutes="01" orientation="longitude" pageId="27" pageNumber="26" precision="925" value="-84.01667">84°01W</geoCoordinate>
;
<date id="6955101BB714FFAC681BFBFC42F05E09" box="[299,409,1066,1092]" pageId="27" pageNumber="26" value="2000-04">
<collectingDate id="7911E9F3B714FFAC681BFBFC42F05E09" box="[299,409,1066,1092]" pageId="27" pageNumber="26" value="2000-04">Apr. 2000</collectingDate>
</date>
, INBio-OET” / “
<date id="6955101BB714FFAC6B65FBFC41875E09" box="[597,750,1066,1092]" pageId="27" pageNumber="26" value="2000-04-17">
<collectingDate id="7911E9F3B714FFAC6B65FBFC41875E09" box="[597,750,1066,1092]" pageId="27" pageNumber="26" value="2000-04-17">17 Abril 2000</collectingDate>
</date>
; Bosque secundario; M/19/746” / “
<emphasis id="2F9FEAC9B714FFAC6D45FBFC46615E09" bold="true" box="[1141,1288,1066,1092]" pageId="27" pageNumber="26">
<typeStatus id="C2508879B714FFAC6D45FBFC466A5E09" box="[1141,1283,1066,1092]" pageId="27" pageNumber="26" type="paratype">PARATYPE</typeStatus>
:
</emphasis>
;
<taxonomicName id="DAEB4D58B714FFAC6C25FBFC46F35E09" authorityName="Kiesenwetter" authorityYear="1858" box="[1301,1434,1066,1092]" class="Insecta" family="Eucnemidae" genus="Dromaeolus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="2F9FEAC9B714FFAC6C25FBFC46F35E09" box="[1301,1434,1066,1092]" italics="true" pageId="27" pageNumber="26">Dromaeolus</emphasis>
</taxonomicName>
;
<taxonomicName id="DAEB4D58B714FFAC69F0FB9A424E5E2B" authorityName="Otto, Muona and Cordoba-Alfaro" authorityYear="2023" box="[192,295,1100,1126]" class="Insecta" family="Eucnemidae" genus="Dromaeolus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="26" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="2F9FEAC9B714FFAC69F0FB9A424E5E2B" box="[192,295,1100,1126]" italics="true" pageId="27" pageNumber="26">brunneus</emphasis>
</taxonomicName>
<specimenCount id="0BEDFD52B714FFAC681EFB99422A5E28" box="[302,323,1103,1125]" pageId="27" pageNumber="26" type="female"></specimenCount>
; Otto, Muona &amp; Córdoba-Alfaro; Det.
<collectionCode id="7BFAAE1EB714FFAC6BC8FB9B40655E2A" box="[760,780,1101,1127]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="27" pageNumber="26">R</collectionCode>
.
<collectorName id="B01E530DB714FFAC6A3FFB9B400B5E2B" box="[783,866,1100,1127]" pageId="27" pageNumber="26">L. Otto</collectorName>
; 2019” (yellow printed label) (
<collectionCode id="7BFAAE1EB714FFAC6D82FB9A47935E2B" box="[1202,1274,1100,1126]" country="Costa Rica" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="27" pageNumber="26">INBC</collectionCode>
)
</materialsCitation>
.
<typeStatus id="C2508879B714FFAC6C3CFB9B46115E2A" box="[1292,1400,1101,1127]" pageId="27" pageNumber="26" type="paratype">Paratypes</typeStatus>
are to be deposited in
<collectionCode id="7BFAAE1EB714FFAC68BEFBB842B95EC5" box="[398,464,1134,1160]" country="Costa Rica" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="27" pageNumber="26">INBC</collectionCode>
and
<collectionCode id="7BFAAE1EB714FFAC6B37FBB841255EC5" box="[519,588,1134,1160]" pageId="27" pageNumber="26">GERP</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="55F16550B714FFAB69F0FB4E46CB5C5D" lastPageId="28" lastPageNumber="27" pageId="27" pageNumber="26" type="description">
<paragraph id="1D5436DBB714FFAC69F0FB4E47525CFC" blockId="27.[191,1443,1176,1850]" pageId="27" pageNumber="26">
<emphasis id="2F9FEAC9B714FFAC69F0FB4E416A5EFF" bold="true" box="[192,515,1176,1202]" pageId="27" pageNumber="26">
Description. Male
<typeStatus id="C2508879B714FFAC68ACFB4E416A5EFF" box="[412,515,1176,1202]" pageId="27" pageNumber="26" type="holotype">holotype</typeStatus>
</emphasis>
: Length, 6.0 mm. Width,
<quantity id="DA139B3EB714FFAC6A1DFB4E40EE5EFE" box="[813,903,1176,1203]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="27" pageNumber="26" unit="mm" value="1.5">1.5 mm</quantity>
. Body elongate; dark brown; antennae and legs dark reddish-brown; head, pronotum and elytra clothed with short, recumbent yellowish setae (
<figureCitation id="85D02A5EB714FFAC6DEEFB6C46255E98" box="[1246,1356,1210,1237]" captionStart="Figures 5457" captionStartId="26.[192,279,709,735]" captionTargetBox="[219,1413,213,679]" captionTargetId="figure-487@26.[219,1414,213,679]" captionTargetPageId="26" captionText="Figures 5457. Costa Rican Dromaeolus species. 54) Dromaeolus americanus sp. nov., dorsal habitus. 55) Dromaeolus americanus sp. nov., ventral habitus. 56) Dromaeolus brunneus sp. nov., male holotype, dorsal habitus. 57) Dromaeolus brunneus sp. nov., male holotype, ventral habitus. (Scale: 5457 = 1.0 mm)" figureDoi="http://doi.org/10.5281/zenodo.10831109" httpUri="https://zenodo.org/record/10831109/files/figure.png" pageId="27" pageNumber="26">Figure 56</figureCitation>
).
<emphasis id="2F9FEAC9B714FFAC6C6AFB6C46F15E99" bold="true" box="[1370,1432,1210,1236]" pageId="27" pageNumber="26">Head</emphasis>
: Surface dullish, subspherical, closely punctate; frons convex, simple without impression or carina; apical margin of frontoclypeal region weakly trilobed, about 2 times wider than base; interantennal carina complete; mandibles stout, bidentate, densely punctate.
<emphasis id="2F9FEAC9B714FFAC6B0EFAF641CA5F77" bold="true" box="[574,675,1312,1338]" pageId="27" pageNumber="26">Antenna</emphasis>
: Filiform from flagellomeres IIX, attaining up to 1/3the length of the body, just beyond pronotal hind angles; flagellomere I longer than wide, longer than II; flagellomeres IIIII each sub-equal, longer than wide, shorter than IV; IVVIII each longer than wide, sub-equal; flagellomere IX slightly longer than VIII.
<emphasis id="2F9FEAC9B714FFAC68BBFA51416B5FEC" bold="true" box="[395,514,1415,1441]" pageId="27" pageNumber="26">Pronotum</emphasis>
: Surface somewhat dullish, with closely spaced punctures; longer than wide, with well-developed, sharp hind angles; sides arcuate; disc convex without fovea or carina; base sinuous.
<emphasis id="2F9FEAC9B714FFAC6C09FA7F426F5FA8" bold="true" pageId="27" pageNumber="26">Scutellar shield</emphasis>
: Longer than wide, sub-triangular, shallowly punctate, setose and distally rounded.
<emphasis id="2F9FEAC9B714FFAC6D8AFA1D46695FA8" bold="true" box="[1210,1280,1483,1509]" pageId="27" pageNumber="26">Elytra</emphasis>
: Very shallow striae indicated at humeri region, indistinct elsewhere; interstices flattened; surfaces shiny, transversely rugose at basal 1/2, closely punctate at apical 1/2.
<emphasis id="2F9FEAC9B714FFAC6B6BF9D941F95C64" bold="true" box="[603,656,1551,1577]" pageId="27" pageNumber="26">Legs</emphasis>
: First tarsomere as long as the combined lengths of the remaining four on meso- and metatarsi; tibiae rounded in cross section; metatarsomeres IIII simple; metatarsomere IV excavated; metatarsomere V elongate with simple claws.
<emphasis id="2F9FEAC9B714FFAC6A34F985403B5C20" bold="true" box="[772,850,1619,1645]" pageId="27" pageNumber="26">Venter</emphasis>
(
<figureCitation id="85D02A5EB714FFAC6A6FF98540A65C23" box="[863,975,1619,1646]" captionStart="Figures 5457" captionStartId="26.[192,279,709,735]" captionTargetBox="[219,1413,213,679]" captionTargetId="figure-487@26.[219,1414,213,679]" captionTargetPageId="26" captionText="Figures 5457. Costa Rican Dromaeolus species. 54) Dromaeolus americanus sp. nov., dorsal habitus. 55) Dromaeolus americanus sp. nov., ventral habitus. 56) Dromaeolus brunneus sp. nov., male holotype, dorsal habitus. 57) Dromaeolus brunneus sp. nov., male holotype, ventral habitus. (Scale: 5457 = 1.0 mm)" figureDoi="http://doi.org/10.5281/zenodo.10831109" httpUri="https://zenodo.org/record/10831109/files/figure.png" pageId="27" pageNumber="26">Figure 57</figureCitation>
): Closely punctate, with short, recumbent yellowish setae; hypomeron with basally opened, lateral antennal grooves; metepisterna parallel-sided; elytral epipleura punctate; metacoxal plates medially 3.06.0 times wider than laterally.
</paragraph>
<paragraph id="1D5436DBB714FFAC6830F96C42DB5D77" blockId="27.[191,1443,1176,1850]" pageId="27" pageNumber="26">
<emphasis id="2F9FEAC9B714FFAC6830F96C42D05C99" bold="true" box="[256,441,1722,1748]" pageId="27" pageNumber="26">
Female
<typeStatus id="C2508879B714FFAC686BF96C42D05C99" box="[347,441,1722,1748]" pageId="27" pageNumber="26" type="allotype">allotype</typeStatus>
</emphasis>
(
<figureCitation id="85D02A5EB714FFAC68F8F96C41505C99" box="[456,569,1721,1748]" captionStart="Figures 5862" captionStartId="28.[192,279,1209,1235]" captionTargetBox="[455,1176,213,1179]" captionTargetId="figure-277@28.[455,1176,213,1179]" captionTargetPageId="28" captionText="Figures 5862. Costa Rican Dromaeolus species. 58) Dromaeolus brunneus sp. nov., female allotype, dorsal habitus. 59) Dromaeolus herediensis sp. nov., dorsal habitus. 60) Dromaeolus herediensis sp. nov., ventral habitus. 61) Dromaeolus holdridgei sp. nov., dorsal habitus. 62) Dromaeolus holdridgei sp. nov., ventral habitus. (Scale: 5962 = 1.0 mm; 58 = 5.0 mm)" figureDoi="http://doi.org/10.5281/zenodo.10831111" httpUri="https://zenodo.org/record/10831111/files/figure.png" pageId="27" pageNumber="26">Figure 58</figureCitation>
): Length, 9.0 mm. Width, 2.0 mm. Short carina extending from interantennal carina down the frontoclypeal region. Flagellomeres filiform, less than 1/3 the length of the body; flagellomere I longer than wide, longer than II; flagellomere II longer than wide, as long as III; flagellomeres IVVIII longer than wide, sub-equal.
</paragraph>
<paragraph id="1D5436DBB714FFAB69F0F89C46CB5C5D" blockId="27.[192,1440,1866,1927]" lastBlockId="28.[191,1442,1389,1552]" lastPageId="28" lastPageNumber="27" pageId="27" pageNumber="26">
<emphasis id="2F9FEAC9B714FFAC69F0F89C422B5D29" bold="true" box="[192,322,1866,1892]" pageId="27" pageNumber="26">Variations.</emphasis>
Three male and
<specimenCount id="0BEDFD52B714FFAC6B31F89C41E55D29" box="[513,652,1866,1892]" pageId="27" pageNumber="26" type="female">eight female</specimenCount>
<typeStatus id="C2508879B714FFAC6BA3F89C40695D29" box="[659,768,1866,1892]" pageId="27" pageNumber="26" type="paratype">paratypes</typeStatus>
were examined. Males measured 6.0 mm long and
<quantity id="DA139B3EB714FFAC6C79F89C46C95D29" box="[1353,1440,1866,1893]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="27" pageNumber="26" unit="mm" value="1.5">1.5 mm</quantity>
wide. Females measured 6.09.0 mm long and
<quantity id="DA139B3EB714FFAC6BE8F8BA40095DCA" box="[728,864,1900,1927]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="27" pageNumber="26" unit="mm" value="2.0" valueMax="2.5" valueMin="1.5">1.52.5 mm</quantity>
wide. All male
<typeStatus id="C2508879B714FFAC6D24F8BB47E85DCA" box="[1044,1153,1901,1927]" pageId="27" pageNumber="26" type="paratype">paratypes</typeStatus>
are just as long as and as wide as the
<typeStatus id="C2508879B713FFAB6875FABB42C25FCA" box="[325,427,1389,1415]" pageId="28" pageNumber="27" type="holotype">holotype</typeStatus>
. Two female
<typeStatus id="C2508879B713FFAB6B70FAB841C45FC5" box="[576,685,1390,1416]" pageId="28" pageNumber="27" type="paratype">paratypes</typeStatus>
are just as large and as wide as the
<typeStatus id="C2508879B713FFAB6D0DFABB47F35FCA" box="[1085,1178,1389,1415]" pageId="28" pageNumber="27" type="allotype">allotype</typeStatus>
. Four female
<typeStatus id="C2508879B713FFAB6C03FAB846C95FC5" box="[1331,1440,1390,1416]" pageId="28" pageNumber="27" type="paratype">paratypes</typeStatus>
are shorter than and just as wide as the
<typeStatus id="C2508879B713FFAB6B4DFA5941B35FE4" box="[637,730,1423,1449]" pageId="28" pageNumber="27" type="allotype">allotype</typeStatus>
. The remaining
<specimenCount id="0BEDFD52B713FFAB6AA0FA4647655FE4" box="[912,1036,1423,1450]" pageId="28" pageNumber="27" type="female">two female</specimenCount>
<typeStatus id="C2508879B713FFAB6D22FA4647165FE7" box="[1042,1151,1424,1450]" pageId="28" pageNumber="27" type="paratype">paratypes</typeStatus>
are shorter than and narrower than the
<typeStatus id="C2508879B713FFAB685AFA6742AE5F86" box="[362,455,1457,1483]" pageId="28" pageNumber="27" type="allotype">allotype</typeStatus>
. None of the
<typeStatus id="C2508879B713FFAB6B69FA6441D25F81" box="[601,699,1458,1484]" pageId="28" pageNumber="27" type="paratype">paratype</typeStatus>
have a short carina extending down the frontoclypeal region from the interantennal carina. Four
<typeStatus id="C2508879B713FFAB6B11FA0241E75FA3" box="[545,654,1492,1518]" pageId="28" pageNumber="27" type="paratype">paratypes</typeStatus>
out of 11 have the same elytral microsculpture as the
<typeStatus id="C2508879B713FFAB6C33FA0246005FA3" box="[1283,1385,1492,1518]" pageId="28" pageNumber="27" type="holotype">holotype</typeStatus>
. The remaining
<specimenCount id="0BEDFD52B713FFAB680CFA2042875C5D" box="[316,494,1526,1552]" pageId="28" pageNumber="27" type="generic" typeStatus="paratypes">seven paratypes</specimenCount>
exhibit transversely rugose surfaces at the basal 1/3of the elytra instead of the basal 1/2.
</paragraph>
</subSubSection>
<caption id="49946653B713FFAB69F0FB6F42B95F75" ID-DOI="http://doi.org/10.5281/zenodo.10831111" ID-Zenodo-Dep="10831111" httpUri="https://zenodo.org/record/10831111/files/figure.png" pageId="28" pageNumber="27" startId="28.[192,279,1209,1235]" targetBox="[455,1176,213,1179]" targetPageId="28" targetType="figure">
<paragraph id="1D5436DBB713FFAB69F0FB6F42B95F75" blockId="28.[192,1440,1209,1337]" pageId="28" pageNumber="27">
<emphasis id="2F9FEAC9B713FFAB69F0FB6F42035E9E" bold="true" box="[192,362,1209,1235]" pageId="28" pageNumber="27">Figures 5862.</emphasis>
Costa Rican
<taxonomicName id="DAEB4D58B713FFAB68CBFB6F41B55E9E" box="[507,732,1209,1235]" class="Insecta" family="Eucnemidae" genus="Dromaeolus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="2F9FEAC9B713FFAB68CBFB6F41E95E9E" box="[507,640,1209,1235]" italics="true" pageId="28" pageNumber="27">Dromaeolus</emphasis>
species.
</taxonomicName>
<emphasis id="2F9FEAC9B713FFAB6BD0FB6F406F5E9E" bold="true" box="[736,774,1209,1235]" pageId="28" pageNumber="27">58)</emphasis>
<taxonomicName id="DAEB4D58B713FFAB6A3CFB6F40975E9E" authority="Otto &amp; Muona &amp; Córdoba-Alfaro, 2023" authorityName="Otto &amp; Muona &amp; Córdoba-Alfaro" authorityYear="2023" box="[780,1022,1209,1235]" class="Insecta" family="Eucnemidae" genus="Dromaeolus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="brunneus" status="sp. nov.">
<emphasis id="2F9FEAC9B713FFAB6A3CFB6F40975E9E" box="[780,1022,1209,1235]" italics="true" pageId="28" pageNumber="27">Dromaeolus brunneus</emphasis>
</taxonomicName>
<emphasis id="2F9FEAC9B713FFAB6D34FB6C47335E99" bold="true" box="[1028,1114,1210,1236]" pageId="28" pageNumber="27">
<taxonomicNameLabel id="34AC57B2B713FFAB6D34FB6C47335E99" box="[1028,1114,1210,1236]" pageId="28" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female allotype, dorsal habitus.
<emphasis id="2F9FEAC9B713FFAB69DEFB0D427D5EB8" bold="true" box="[238,276,1243,1269]" pageId="28" pageNumber="27">59)</emphasis>
<taxonomicName id="DAEB4D58B713FFAB682AFB0D41745EB8" authority="Otto &amp; Muona &amp; Córdoba-Alfaro, 2023" authorityName="Otto &amp; Muona &amp; Córdoba-Alfaro" authorityYear="2023" box="[282,541,1243,1269]" class="Insecta" family="Eucnemidae" genus="Dromaeolus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="herediensis" status="sp. nov.">
<emphasis id="2F9FEAC9B713FFAB682AFB0D41745EB8" box="[282,541,1243,1269]" italics="true" pageId="28" pageNumber="27">Dromaeolus herediensis</emphasis>
</taxonomicName>
<emphasis id="2F9FEAC9B713FFAB6B13FB0D41135EB8" bold="true" box="[547,634,1243,1269]" pageId="28" pageNumber="27">
<taxonomicNameLabel id="34AC57B2B713FFAB6B13FB0D41135EB8" box="[547,634,1243,1269]" pageId="28" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, dorsal habitus.
<emphasis id="2F9FEAC9B713FFAB6A1FFB0D403C5EB8" bold="true" box="[815,853,1243,1269]" pageId="28" pageNumber="27">60)</emphasis>
<taxonomicName id="DAEB4D58B713FFAB6A6BFB0D47365EB8" authority="Otto &amp; Muona &amp; Córdoba-Alfaro, 2023" authorityName="Otto &amp; Muona &amp; Córdoba-Alfaro" authorityYear="2023" box="[859,1119,1243,1269]" class="Insecta" family="Eucnemidae" genus="Dromaeolus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="herediensis" status="sp. nov.">
<emphasis id="2F9FEAC9B713FFAB6A6BFB0D47365EB8" box="[859,1119,1243,1269]" italics="true" pageId="28" pageNumber="27">Dromaeolus herediensis</emphasis>
</taxonomicName>
<emphasis id="2F9FEAC9B713FFAB6D55FB0D47D55EB8" bold="true" box="[1125,1212,1243,1269]" pageId="28" pageNumber="27">
<taxonomicNameLabel id="34AC57B2B713FFAB6D55FB0D47D55EB8" box="[1125,1212,1243,1269]" pageId="28" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, ventral habitus.
<emphasis id="2F9FEAC9B713FFAB6C4AFB0D46F65EB8" bold="true" box="[1402,1439,1243,1269]" pageId="28" pageNumber="27">61)</emphasis>
<taxonomicName id="DAEB4D58B713FFAB69F0FB2B42D05F5A" authority="Otto &amp; Muona &amp; Córdoba-Alfaro, 2023" authorityName="Otto &amp; Muona &amp; Córdoba-Alfaro" authorityYear="2023" box="[192,441,1277,1303]" class="Insecta" family="Eucnemidae" genus="Dromaeolus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="holdridgei" status="sp. nov.">
<emphasis id="2F9FEAC9B713FFAB69F0FB2B42D05F5A" box="[192,441,1277,1303]" italics="true" pageId="28" pageNumber="27">Dromaeolus holdridgei</emphasis>
</taxonomicName>
<emphasis id="2F9FEAC9B713FFAB68F0FB2B417F5F5A" bold="true" box="[448,534,1277,1303]" pageId="28" pageNumber="27">
<taxonomicNameLabel id="34AC57B2B713FFAB68F0FB2B417F5F5A" box="[448,534,1277,1303]" pageId="28" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, dorsal habitus.
<emphasis id="2F9FEAC9B713FFAB6BFBFB2A41985F5B" bold="true" box="[715,753,1276,1302]" pageId="28" pageNumber="27">62)</emphasis>
<taxonomicName id="DAEB4D58B713FFAB6BC8FB2B40985F5A" authority="Otto &amp; Muona &amp; Córdoba-Alfaro, 2023" authorityName="Otto &amp; Muona &amp; Córdoba-Alfaro" authorityYear="2023" box="[760,1009,1277,1303]" class="Insecta" family="Eucnemidae" genus="Dromaeolus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="holdridgei" status="sp. nov.">
<emphasis id="2F9FEAC9B713FFAB6BC8FB2B40985F5A" box="[760,1009,1277,1303]" italics="true" pageId="28" pageNumber="27">Dromaeolus holdridgei</emphasis>
</taxonomicName>
<emphasis id="2F9FEAC9B713FFAB6AC7FB2B47275F5A" bold="true" box="[1015,1102,1277,1303]" pageId="28" pageNumber="27">
<taxonomicNameLabel id="34AC57B2B713FFAB6AC7FB2B47275F5A" box="[1015,1102,1277,1303]" pageId="28" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, ventral habitus. (Scale: 5962 = 1.0 mm; 58 = 5.0 mm)
</paragraph>
</caption>
<subSubSection id="55F16550B713FFAB69F0F9F6424B5C11" pageId="28" pageNumber="27" type="distribution">
<paragraph id="1D5436DBB713FFAB69F0F9F6424B5C11" blockId="28.[192,1440,1568,1628]" pageId="28" pageNumber="27">
<emphasis id="2F9FEAC9B713FFAB69F0F9F642325C77" bold="true" box="[192,347,1568,1594]" pageId="28" pageNumber="27">Distribution.</emphasis>
This eucnemid species is known from
<specimenCount id="0BEDFD52B713FFAB6A28F9F640D95C77" box="[792,944,1568,1594]" pageId="28" pageNumber="27" type="generic">13 specimens</specimenCount>
taken from La Selva within the province of
<collectingRegion id="DF2FF839B713FFAB69F2F99442775C11" box="[194,286,1602,1628]" country="Costa Rica" name="Heredia" pageId="28" pageNumber="27">Heredia</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="55F16550B713FFAB69F0F9BA471A5CE5" pageId="28" pageNumber="27" type="biology_ecology">
<paragraph id="1D5436DBB713FFAB69F0F9BA471A5CE5" blockId="28.[191,1441,1644,1705]" pageId="28" pageNumber="27">
<emphasis id="2F9FEAC9B713FFAB69F0F9BA42495CCB" bold="true" box="[192,288,1644,1670]" pageId="28" pageNumber="27">Biology.</emphasis>
Six specimens were taken from a primary forest. Six specimens were taken from a second growth forest. One specimen was taken from a successional parcel. Larvae and pupae are unknown.
</paragraph>
</subSubSection>
<subSubSection id="55F16550B713FFAB69F0F96E46415C9F" box="[192,1320,1720,1747]" pageId="28" pageNumber="27" type="etymology">
<paragraph id="1D5436DBB713FFAB69F0F96E46415C9F" blockId="28.[192,1320,1720,1747]" box="[192,1320,1720,1747]" pageId="28" pageNumber="27">
<emphasis id="2F9FEAC9B713FFAB69F0F96E422D5C9F" bold="true" box="[192,324,1720,1746]" pageId="28" pageNumber="27">Etymology.</emphasis>
The specific epithet is named from its brown colored dorsum present on the new species.
</paragraph>
</subSubSection>
</treatment>
</document>