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<document id="CB9C6ED982B67C6F639E44C3453F0984" ID-DOI="10.5281/zenodo.10831085" ID-ISSN="1942-1354" ID-Zenodo-Dep="10831085" ID-ZooBank="C1D5B819-A964-4679-B090-84CDBBC59D6A" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1688389961601" checkinUser="felipe" docAuthor="Otto, Robert L., Muona, Jyrki &amp; Córdoba-Alfaro, Jim" docDate="2023" docId="954287CDB712FFA969F0F94447A35E97" docLanguage="en" docName="InsectaMundi.2023.0991.1-36.pdf" docOrigin="Insecta Mundi 2023 (991)" docStyle="DocumentStyle:F9976E9B7F8A094D890004515A7A0592.4:InsectaMundi.2020-.journal_article" docStyleId="F9976E9B7F8A094D890004515A7A0592" docStyleName="InsectaMundi.2020-.journal_article" docStyleVersion="4" docTitle="Dromaeolus holdridgei Otto, Muona and Cordoba-Alfaro 2023, new species" docType="treatment" docVersion="4" lastPageNumber="29" masterDocId="697BFFB5B70FFFB76930FFD643695A4D" masterDocTitle="A new genus and sixteen new species of false click beetles (Coleoptera: Eucnemidae) described from the Heredia Province of Costa Rica with several additional records from the Osa Peninsula and Panama" masterLastPageNumber="36" masterPageNumber="1" pageNumber="28" updateTime="1710774778166" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-3.0">
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<mods:titleInfo id="C767B889A904959A0DF21EEA73A663AC">
<mods:title id="E1370D62B2232DED9C32FA57DC184C0C">A new genus and sixteen new species of false click beetles (Coleoptera: Eucnemidae) described from the Heredia Province of Costa Rica with several additional records from the Osa Peninsula and Panama</mods:title>
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<mods:name id="132ADB29158A8FE53FF801B3F755ACD7" type="personal">
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<mods:namePart id="22983E8513D519635D7C2B550A51460F">Otto, Robert L.</mods:namePart>
<mods:affiliation id="57AE8E2284F6E059E2E7A88AEC5842FC">W 4806 Chrissie Circle, Shawano, WI 54166, U. S. A.</mods:affiliation>
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<mods:namePart id="ADAAFABFC675AEF4498977F04D83577B">Muona, Jyrki</mods:namePart>
<mods:affiliation id="2D6F78DC90FDB9111144571F110DE1AB">Finnish Museum of Natural History, Zoology Unit PO Box 17, University of Helsinki, Finland FIN- 00014</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="1A9D36A365D473068D3E554E09A8AA16">Córdoba-Alfaro, Jim</mods:namePart>
<mods:affiliation id="6ECF6734015023E674BF9F7A1AFCDCC3">Insectopia Insect Museum, BioSur Foundation Puerto Jiménez, Península de Osa, Costa Rica jim. cordoba @ biosurcorcovado. org</mods:affiliation>
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<mods:title id="DDB4BC0D8ED3D73451B5B3C72B6F8D4E">Insecta Mundi</mods:title>
</mods:titleInfo>
<mods:part id="BF333A83E15978DE01A75114E455E9D4">
<mods:date id="BBBE1120279A254D4A6F1395AFBDE945">2023</mods:date>
<mods:detail id="6185306EBE47E0BD195B97E242C6D374" type="pubDate">
<mods:number id="FEB1E1B68E071F0475A810F84DCE8028">2023-05-26</mods:number>
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<mods:number id="A28E4287E06F2C73D7F4C3663912595B">2023</mods:number>
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<mods:number id="9F25BAAE357F03DE31064A28B01416B4">991</mods:number>
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<mods:classification id="49841CF88E91075468497725314CB9E6">journal article</mods:classification>
<mods:identifier id="C664E9E8305E4C7D683F57DB2011C195" type="DOI">10.5281/zenodo.10831085</mods:identifier>
<mods:identifier id="19912B43E7B0D1695C812E455591793B" type="ISSN">1942-1354</mods:identifier>
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<treatment id="954287CDB712FFA969F0F94447A35E97" ID-DOI="http://doi.org/10.5281/zenodo.8109109" ID-Zenodo-Dep="8109109" LSID="urn:lsid:plazi:treatment:954287CDB712FFA969F0F94447A35E97" httpUri="http://treatment.plazi.org/id/954287CDB712FFA969F0F94447A35E97" lastPageId="30" lastPageNumber="29" pageId="29" pageNumber="28">
<subSubSection id="55F16550B712FFAA69F0F94447555CE2" box="[192,1084,1681,1711]" pageId="29" pageNumber="28" type="nomenclature">
<paragraph id="1D5436DBB712FFAA69F0F94447555CE2" blockId="29.[192,1084,1681,1745]" box="[192,1084,1681,1711]" pageId="29" pageNumber="28">
<heading id="461C81B7B712FFAA69F0F94447555CE2" box="[192,1084,1681,1711]" fontSize="12" level="2" pageId="29" pageNumber="28" reason="2">
<emphasis id="2F9FEAC9B712FFAA69F0F94447555CE2" bold="true" box="[192,1084,1681,1711]" pageId="29" pageNumber="28">
<taxonomicName id="DAEB4D58B712FFAA69F0F94440F05CE3" authority="Otto, Muona and Cordoba-Alfaro" authorityName="Otto, Muona and Cordoba-Alfaro" authorityYear="2023" box="[192,921,1681,1711]" class="Insecta" family="Eucnemidae" genus="Dromaeolus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="28" phylum="Arthropoda" rank="species" species="holdridgei" status="sp. nov.">
<emphasis id="2F9FEAC9B712FFAA69F0F94442885CE2" bold="true" box="[192,481,1682,1711]" italics="true" pageId="29" pageNumber="28">Dromaeolus holdridgei</emphasis>
Otto, Muona and Córdoba-Alfaro
</taxonomicName>
,
<taxonomicNameLabel id="34AC57B2B712FFAA6A97F94447555CE2" box="[935,1084,1682,1711]" pageId="29" pageNumber="28" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="55F16550B712FFAA69F0F96142345C9C" box="[192,349,1719,1745]" pageId="29" pageNumber="28" type="description">
<paragraph id="1D5436DBB712FFAA69F0F96142345C9C" blockId="29.[192,1084,1681,1745]" box="[192,349,1719,1745]" pageId="29" pageNumber="28">
<figureCitation id="85D02A5EB712FFAA69F0F96142345C9C" box="[192,349,1719,1745]" captionStart="Figures 5862" captionStartId="28.[192,279,1209,1235]" captionTargetBox="[455,1176,213,1179]" captionTargetId="figure-277@28.[455,1176,213,1179]" captionTargetPageId="28" captionText="Figures 5862. Costa Rican Dromaeolus species. 58) Dromaeolus brunneus sp. nov., female allotype, dorsal habitus. 59) Dromaeolus herediensis sp. nov., dorsal habitus. 60) Dromaeolus herediensis sp. nov., ventral habitus. 61) Dromaeolus holdridgei sp. nov., dorsal habitus. 62) Dromaeolus holdridgei sp. nov., ventral habitus. (Scale: 5962 = 1.0 mm; 58 = 5.0 mm)" figureDoi="http://doi.org/10.5281/zenodo.10831111" httpUri="https://zenodo.org/record/10831111/files/figure.png" pageId="29" pageNumber="28">Figures 6162</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="55F16550B712FFAA69F0F93741435DCA" pageId="29" pageNumber="28" type="diagnosis">
<paragraph id="1D5436DBB712FFAA69F0F93741435DCA" blockId="29.[192,1441,1761,1927]" pageId="29" pageNumber="28">
<emphasis id="2F9FEAC9B712FFAA69F0F93742555CB6" bold="true" box="[192,316,1761,1787]" pageId="29" pageNumber="28">Diagnosis.</emphasis>
The new eucnemid species can be readily distinguished from any ornate members of
<taxonomicName id="DAEB4D58B712FFAA6C2BF93746C95CB6" authorityName="Kiesenwetter" authorityYear="1858" box="[1307,1440,1761,1787]" class="Insecta" family="Eucnemidae" genus="Dromaeolus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="2F9FEAC9B712FFAA6C2BF93746C95CB6" box="[1307,1440,1761,1787]" italics="true" pageId="29" pageNumber="28">Dromaeolus</emphasis>
</taxonomicName>
present in
<collectingCountry id="65FC764BB712FFAA680AF8D242DC5D53" box="[314,437,1796,1822]" name="Costa Rica" pageId="29" pageNumber="28">Costa Rica</collectingCountry>
by the presence of interantennal carina, very short, median carina on the frontoclypeal region and short, median carina present on the frons except
<taxonomicName id="DAEB4D58B712FFAA6A52F8F140935D0C" authority="Otto &amp; Muona &amp; Córdoba-Alfaro, 2023" authorityName="Otto &amp; Muona &amp; Córdoba-Alfaro" authorityYear="2023" box="[866,1018,1831,1857]" class="Insecta" family="Eucnemidae" genus="Dromaeolus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="28" phylum="Arthropoda" rank="species" species="herediensis" status="sp. nov.">
<emphasis id="2F9FEAC9B712FFAA6A52F8F140935D0C" box="[866,1018,1831,1857]" italics="true" pageId="29" pageNumber="28">D. herediensis</emphasis>
</taxonomicName>
<emphasis id="2F9FEAC9B712FFAA6D30F8F147E15D0D" bold="true" box="[1024,1160,1830,1857]" pageId="29" pageNumber="28">
<taxonomicNameLabel id="34AC57B2B712FFAA6D30F8F147E15D0D" box="[1024,1160,1830,1857]" pageId="29" pageNumber="28" rank="species">new species</taxonomicNameLabel>
</emphasis>
. Narrower silver-colored band on the elytra along with the absence of silver-colored setae at elytral apices will distinguish the new species from
<taxonomicName id="DAEB4D58B712FFAA69CEF8BB42FF5DCB" authority="Otto &amp; Muona &amp; Córdoba-Alfaro, 2023" authorityName="Otto &amp; Muona &amp; Córdoba-Alfaro" authorityYear="2023" box="[254,406,1900,1926]" class="Insecta" family="Eucnemidae" genus="Dromaeolus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="28" phylum="Arthropoda" rank="species" species="herediensis" status="sp. nov.">
<emphasis id="2F9FEAC9B712FFAA69CEF8BB42FF5DCB" box="[254,406,1900,1926]" italics="true" pageId="29" pageNumber="28">D. herediensis</emphasis>
</taxonomicName>
<emphasis id="2F9FEAC9B712FFAA68ACF8BA414D5DCB" bold="true" box="[412,548,1900,1926]" pageId="29" pageNumber="28">
<taxonomicNameLabel id="34AC57B2B712FFAA68ACF8BA414D5DCB" box="[412,548,1900,1926]" pageId="29" pageNumber="28" rank="species">new species</taxonomicNameLabel>
</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="55F16550B711FFA969F0FF0042335B15" pageId="30" pageNumber="29" type="materials_examined">
<paragraph id="1D5436DBB711FFA969F0FF0042335B15" blockId="30.[192,1441,213,344]" pageId="30" pageNumber="29">
<emphasis id="2F9FEAC9B711FFA969F0FF0042A45ABD" bold="true" box="[192,461,213,240]" pageId="30" pageNumber="29">Type material. Female</emphasis>
<materialsCitation id="AD833C86B711FFA968E6FF0042335B15" collectingDate="1993-07-01" collectionCode="ALAS, R, INBC" collectorName="L. Otto" country="Costa Rica" elevation="150" latitude="10.433333" location="Est. Bio. La Selva" longLatPrecision="1298" longitude="-84.01667" pageId="30" pageNumber="29" specimenCount="1" specimenCount-female="1" stateProvince="Heredia" typeStatus="holotype">
<emphasis id="2F9FEAC9B711FFA968E6FF0041565ABD" bold="true" box="[470,575,214,240]" pageId="30" pageNumber="29">
<typeStatus id="C2508879B711FFA968E6FF0041565ABD" box="[470,575,214,240]" pageId="30" pageNumber="29" type="holotype">holotype</typeStatus>
</emphasis>
: “
<collectingCountry id="65FC764BB711FFA96B68FF0040695ABD" box="[600,768,214,240]" name="Costa Rica" pageId="30" pageNumber="29">COSTA RICA</collectingCountry>
:
<collectingRegion id="DF2FF839B711FFA96A3CFF0040035ABD" box="[780,874,214,240]" country="Costa Rica" name="Heredia" pageId="30" pageNumber="29">Heredia</collectingRegion>
;
<location id="18346000B711FFA96A47FF0047535ABD" LSID="urn:lsid:plazi:treatment:954287CDB712FFA969F0F94447A35E97:18346000B711FFA96A47FF0047535ABD" box="[887,1082,214,240]" country="Costa Rica" latitude="10.433333" longLatPrecision="1298" longitude="-84.01667" name="Est. Bio. La Selva" pageId="30" pageNumber="29" stateProvince="Heredia">Est.Bio. La Selva</location>
, 50;
<quantity id="DA139B3EB711FFA96DB1FF0047A35ABD" box="[1153,1226,214,241]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="30" pageNumber="29" unit="m" value="150.0">
<elevation id="96C6D1E8B711FFA96DB1FF0047A35ABD" box="[1153,1226,214,241]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="30" pageNumber="29" unit="m" value="150.0">150 m</elevation>
</quantity>
<geoCoordinate id="78DF501CB711FFA96DE4FF0046595ABD" box="[1236,1328,214,240]" degrees="10" direction="north" minutes="26" orientation="latitude" pageId="30" pageNumber="29" precision="925" value="10.433333">10°26N</geoCoordinate>
<geoCoordinate id="78DF501CB711FFA96C09FF0046F45ABD" box="[1337,1437,214,240]" degrees="84" direction="west" minutes="01" orientation="longitude" pageId="30" pageNumber="29" precision="925" value="-84.01667">84°01W</geoCoordinate>
; Proy.
<collectionCode id="7BFAAE1EB711FFA969C9FF2F42285B5E" box="[249,321,249,275]" pageId="30" pageNumber="29">ALAS</collectionCode>
, INBio-OET” / “M/01/136;
<date id="6955101BB711FFA96BBDFF2F40455B5E" box="[653,812,248,275]" pageId="30" pageNumber="29" value="1993-07-01">
<collectingDate id="7911E9F3B711FFA96BBDFF2F40455B5E" box="[653,812,248,275]" pageId="30" pageNumber="29" value="1993-07-01">01 Julio 1993</collectingDate>
</date>
; Parcelas sucesionales” / “
<emphasis id="2F9FEAC9B711FFA96D52FF2E46685B5F" bold="true" box="[1122,1281,248,274]" pageId="30" pageNumber="29">
<typeStatus id="C2508879B711FFA96D52FF2E47955B5F" box="[1122,1276,248,274]" pageId="30" pageNumber="29" type="holotype">HOLOTYPE</typeStatus>
:
</emphasis>
;
<taxonomicName id="DAEB4D58B711FFA96C22FF2E46F05B5F" authorityName="Kiesenwetter" authorityYear="1858" box="[1298,1433,248,274]" class="Insecta" family="Eucnemidae" genus="Dromaeolus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="2F9FEAC9B711FFA96C22FF2E46F05B5F" box="[1298,1433,248,274]" italics="true" pageId="30" pageNumber="29">Dromaeolus</emphasis>
</taxonomicName>
;
<taxonomicName id="DAEB4D58B711FFA969F0FECD42585B78" authorityName="Otto, Muona and Cordoba-Alfaro" authorityYear="2023" box="[192,305,283,309]" class="Insecta" family="Eucnemidae" genus="Dromaeolus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="29" phylum="Arthropoda" rank="species" species="holdridgei">
<emphasis id="2F9FEAC9B711FFA969F0FECD42585B78" box="[192,305,283,309]" italics="true" pageId="30" pageNumber="29">holdridgei</emphasis>
</taxonomicName>
<specimenCount id="0BEDFD52B711FFA96807FEC942255B78" box="[311,332,287,309]" pageId="30" pageNumber="29" type="female"></specimenCount>
; Otto, Muona &amp; Córdoba-Alfaro; Det.
<collectionCode id="7BFAAE1EB711FFA96A38FECA40745B7B" box="[776,797,284,310]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="30" pageNumber="29">R</collectionCode>
.
<collectorName id="B01E530DB711FFA96A10FECA401A5B78" box="[800,883,283,310]" pageId="30" pageNumber="29">L. Otto</collectorName>
; 2019” (red printed label).
<typeStatus id="C2508879B711FFA96D94FECD46675B78" box="[1188,1294,283,309]" pageId="30" pageNumber="29" type="holotype">Holotype</typeStatus>
to be deposited in
<collectionCode id="7BFAAE1EB711FFA96820FEE8423F5B15" box="[272,342,318,344]" country="Costa Rica" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="30" pageNumber="29">INBC</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="55F16550B711FFA969F0FEBE46555E58" pageId="30" pageNumber="29" type="description">
<paragraph id="1D5436DBB711FFA969F0FEBE46555E58" blockId="30.[191,1442,360,1046]" pageId="30" pageNumber="29">
<emphasis id="2F9FEAC9B711FFA969F0FEBE41765BCE" bold="true" box="[192,543,360,387]" pageId="30" pageNumber="29">
Description. Female
<typeStatus id="C2508879B711FFA96887FEBF41765BCE" box="[439,543,361,387]" pageId="30" pageNumber="29" type="holotype">holotype</typeStatus>
</emphasis>
: Length, 6.0 mm. Width,
<quantity id="DA139B3EB711FFA96A76FEBE40C95BCE" box="[838,928,360,387]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="30" pageNumber="29" unit="mm" value="1.5">1.5 mm</quantity>
. Body elongate; black; antennae black; femur and tibiae black, tarsi dark brownish-black; head and pronotum clothed with short, recumbent silver-colored setae more apparent at base and along lateral sides of pronotum, elytra with silver-colored setae more apparent along lateral sides at basal half, narrow band across the middle of each elytron and narrow band along elytral suture terminating at the middle band (
<figureCitation id="85D02A5EB711FFA96BA4FE25406E5840" box="[660,775,499,525]" captionStart="Figures 5862" captionStartId="28.[192,279,1209,1235]" captionTargetBox="[455,1176,213,1179]" captionTargetId="figure-277@28.[455,1176,213,1179]" captionTargetPageId="28" captionText="Figures 5862. Costa Rican Dromaeolus species. 58) Dromaeolus brunneus sp. nov., female allotype, dorsal habitus. 59) Dromaeolus herediensis sp. nov., dorsal habitus. 60) Dromaeolus herediensis sp. nov., ventral habitus. 61) Dromaeolus holdridgei sp. nov., dorsal habitus. 62) Dromaeolus holdridgei sp. nov., ventral habitus. (Scale: 5962 = 1.0 mm; 58 = 5.0 mm)" figureDoi="http://doi.org/10.5281/zenodo.10831111" httpUri="https://zenodo.org/record/10831111/files/figure.png" pageId="30" pageNumber="29">Figure 61</figureCitation>
).
<emphasis id="2F9FEAC9B711FFA96A2EFE2540345840" bold="true" box="[798,861,499,525]" pageId="30" pageNumber="29">Head</emphasis>
: Surface dullish, subspherical with closely spaced punctures; frons convex with short median carina; frontoclypeal region with short, delicate median carina; interantennal carina present, sinuous; apical margin of frontoclypeal region weakly trilobed, more than 2 times wider than base; mandibles stout, bidentate, densely punctate.
<emphasis id="2F9FEAC9B711FFA96A93FD8D47605838" bold="true" box="[931,1033,603,629]" pageId="30" pageNumber="29">Antenna</emphasis>
: Filiform from flagellomeres IIX, attaining nearly 1/3the length of the body, never passed beyond pronotal hind angles; flagellomere I longer than wide, longer than II; flagellomere II quadrate, shorter than III; flagellomere III slightly longer than wide, shorter than IV; IVVIII each sub-equal, longer than wide; flagellomere IX slightly longer than VIII.
<emphasis id="2F9FEAC9B711FFA96DF6FD1546565890" bold="true" box="[1222,1343,707,733]" pageId="30" pageNumber="29">Pronotum</emphasis>
: Surface shiny, with evenly dispersed punctures; slightly longer than wide, with well-developed, sharp hind angles; sides subparallel-sided, slightly arcuate near craniad; disc convex without shallow median groove or circular fovea; base sinuous.
<emphasis id="2F9FEAC9B711FFA96851FCFD417E5908" bold="true" box="[353,535,811,837]" pageId="30" pageNumber="29">Scutellar shield</emphasis>
: Elongate, sub-triangular, shallowly punctate, setose and distally rounded.
<emphasis id="2F9FEAC9B711FFA96C42FCFD438B5925" bold="true" pageId="30" pageNumber="29">Elytra</emphasis>
: Very shallow striae indicated; interstices slightly elevated; surfaces shiny with transversely rugose to deep punctures at humeri, shallower punctures elsewhere.
<emphasis id="2F9FEAC9B711FFA96A25FCA6402359C7" bold="true" box="[789,842,880,906]" pageId="30" pageNumber="29">Legs</emphasis>
: First tarsomere longer than the combined lengths of the remaining four on meso- and metatarsi; tibiae rounded in cross section; metatarsomeres IIII simple; metatarsomere IV excavated; metatarsomere V short with simple claws.
<emphasis id="2F9FEAC9B711FFA96AF1FC604766599D" bold="true" box="[961,1039,950,976]" pageId="30" pageNumber="29">Venter</emphasis>
(
<figureCitation id="85D02A5EB711FFA96D2DFC6047E7599D" box="[1053,1166,950,976]" captionStart="Figures 5862" captionStartId="28.[192,279,1209,1235]" captionTargetBox="[455,1176,213,1179]" captionTargetId="figure-277@28.[455,1176,213,1179]" captionTargetPageId="28" captionText="Figures 5862. Costa Rican Dromaeolus species. 58) Dromaeolus brunneus sp. nov., female allotype, dorsal habitus. 59) Dromaeolus herediensis sp. nov., dorsal habitus. 60) Dromaeolus herediensis sp. nov., ventral habitus. 61) Dromaeolus holdridgei sp. nov., dorsal habitus. 62) Dromaeolus holdridgei sp. nov., ventral habitus. (Scale: 5962 = 1.0 mm; 58 = 5.0 mm)" figureDoi="http://doi.org/10.5281/zenodo.10831111" httpUri="https://zenodo.org/record/10831111/files/figure.png" pageId="30" pageNumber="29">Figure 62</figureCitation>
): Closely punctate, with short, recumbent silver-colored setae; hypomeron with basally opened, lateral antennal grooves; metepisterna parallel-sided; elytral epipleura punctate; metacoxal plates medially 3.06.0 times wider than laterally.
</paragraph>
</subSubSection>
<subSubSection id="55F16550B711FFA969F0FBF342575E2F" pageId="30" pageNumber="29" type="distribution">
<paragraph id="1D5436DBB711FFA969F0FBF342575E2F" blockId="30.[192,1440,1061,1122]" pageId="30" pageNumber="29">
<emphasis id="2F9FEAC9B711FFA969F0FBF342325E72" bold="true" box="[192,347,1061,1087]" pageId="30" pageNumber="29">Distribution.</emphasis>
This eucnemid species is known from a single specimen taken from La Selva within the province of
<collectingRegion id="DF2FF839B711FFA969EDFB9E42505E2F" box="[221,313,1096,1122]" country="Costa Rica" name="Heredia" pageId="30" pageNumber="29">Heredia</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="55F16550B711FFA969F0FBA547DD5EC0" box="[192,1204,1139,1166]" pageId="30" pageNumber="29" type="biology_ecology">
<paragraph id="1D5436DBB711FFA969F0FBA547DD5EC0" blockId="30.[192,1204,1139,1166]" box="[192,1204,1139,1166]" pageId="30" pageNumber="29">
<emphasis id="2F9FEAC9B711FFA969F0FBA542495EC0" bold="true" box="[192,288,1139,1165]" pageId="30" pageNumber="29">Biology.</emphasis>
The
<typeStatus id="C2508879B711FFA96866FBA542D15EC0" box="[342,440,1139,1165]" pageId="30" pageNumber="29" type="holotype">holotype</typeStatus>
was taken from a successional plot. Larvae and pupae are unknown.
</paragraph>
</subSubSection>
<subSubSection id="55F16550B711FFA969F0FB4847A35E97" pageId="30" pageNumber="29" type="etymology">
<paragraph id="1D5436DBB711FFA969F0FB4847A35E97" blockId="30.[192,1440,1182,1243]" pageId="30" pageNumber="29">
<emphasis id="2F9FEAC9B711FFA969F0FB48422D5EF5" bold="true" box="[192,324,1182,1208]" pageId="30" pageNumber="29">Etymology.</emphasis>
The specific epithet is dedicated in honor of Dr. Leslie Holdridge, an American botanist and climatologist who was the founding member of the La Selva Biological Station established in 1968.
</paragraph>
</subSubSection>
</treatment>
</document>