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<document id="3B910AEC329D59B6A2234427CF654817" ID-CLB-Dataset="22298" ID-DOI="10.5852/ejt.2018.454" ID-GBIF-Dataset="654c9653-fbef-4d0d-9e07-619084ac1162" ID-Zenodo-Dep="3817358" ID-ZooBank="07F6B28D-ECA4-4F14-8132-0A58DE156057" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588950109658" checkinUser="valdenar" docAuthor="Mbanyana, Nokuthula, Garcia, Francisco Hita, Robertson, Hamish Gibson &amp; Roux, Johannes Jacobus Le" docDate="2018" docId="039887DCFF86A4622B0EA500FE11A18E" docLanguage="en" docName="ejt-454_mbanyana_hitagarcia_robertson.pdf.imf" docOrigin="European Journal of Taxonomy 454" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Tetramorium galoasanum Santschi 1910" docType="treatment" docVersion="12" lastPageNumber="23" masterDocId="FFA1FFA4FF90A474292DA61DFFFCA41A" masterDocTitle="A taxonomic revision of seed harvester ants of the Tetramorium solidum group (Hymenoptera: Formicidae) in southern Africa" masterLastPageNumber="59" masterPageNumber="1" pageNumber="23" updateTime="1698839042967" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="2E603D6F361625A8BBAE49462A823CAD">A taxonomic revision of seed harvester ants of the Tetramorium solidum group (Hymenoptera: Formicidae) in southern Africa</mods:title>
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<mods:namePart id="A0B2A447E51806C5B06E27647B1023F6">Mbanyana, Nokuthula</mods:namePart>
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<mods:affiliation id="574220B9E4DFC78EC0AF8F687613C815">Iziko Museums of South Africa, P. O. Box 61, Cape Town, 8000 South Africa. Okinawa Institute of Science and Technology Graduate University, 1919 - 1 Tancha, Onna-son, Okinawa, Japan. Department of Botany and Zoology, Stellenbosch University, Private Bag X 1, Matieland, 7602, South Africa.</mods:affiliation>
<mods:nameIdentifier id="A4D3A007630F19310F8DF86B6D3F1A41" type="email">nmbanyana@iziko.org.za</mods:nameIdentifier>
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<mods:namePart id="D30A744344020E3B000C1901774F464D">Garcia, Francisco Hita</mods:namePart>
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<mods:nameIdentifier id="AD01201891C618B71042F304C2960447" type="email">fhitagarcia@gmail.com</mods:nameIdentifier>
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<mods:namePart id="0113C4A902154D7E1FA9E87502AB6FDB">Robertson, Hamish Gibson</mods:namePart>
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<mods:affiliation id="1CE247B18A0024795095BC885340B040">Iziko Museums of South Africa, P. O. Box 61, Cape Town, 8000 South Africa. Okinawa Institute of Science and Technology Graduate University, 1919 - 1 Tancha, Onna-son, Okinawa, Japan. Department of Botany and Zoology, Stellenbosch University, Private Bag X 1, Matieland, 7602, South Africa.</mods:affiliation>
<mods:nameIdentifier id="7E0DE81659BC6DF7D038F547B60DEED2" type="email">hrobertson@iziko.org.za</mods:nameIdentifier>
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<mods:namePart id="C97D0424FAD2598920B94BD10EEA2D83">Roux, Johannes Jacobus Le</mods:namePart>
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<mods:nameIdentifier id="A5AC788310508FF55158D7FF4DC5466C" type="email">jleroux@sun.ac.za</mods:nameIdentifier>
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<mods:title id="8D87FB59259CCD59D095E6DE0C6A5A85">European Journal of Taxonomy</mods:title>
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<mods:date id="F732C2784B12CF805D8CC9805878B8C1">2018</mods:date>
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<mods:number id="B1526D267C6458E741E0C1CD4FA00673">2018-08-07</mods:number>
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<mods:number id="0623B42FFDEBB4073AB9768E686DC46A">454</mods:number>
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<mods:identifier id="669A50664798B7F924D71698E0416A34" type="DOI">10.5852/ejt.2018.454</mods:identifier>
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<treatment id="039887DCFF86A4622B0EA500FE11A18E" ID-DOI="http://doi.org/10.5281/zenodo.3818708" ID-GBIF-Taxon="163993638" ID-Zenodo-Dep="3818708" LSID="urn:lsid:plazi:treatment:039887DCFF86A4622B0EA500FE11A18E" httpUri="http://treatment.plazi.org/id/039887DCFF86A4622B0EA500FE11A18E" lastPageNumber="23" pageId="22" pageNumber="23">
<subSubSection id="C32B6541FF86A4622B0EA500FBEDA72D" box="[547,1041,796,823]" pageId="22" pageNumber="23" type="nomenclature">
<paragraph id="8B8E36CAFF86A4622B0EA500FBEDA72D" blockId="22.[547,1041,796,858]" box="[547,1041,796,823]" pageId="22" pageNumber="23">
<heading id="D0C681A6FF86A4622B0EA500FBEDA72D" box="[547,1041,796,823]" centered="true" fontSize="11" level="2" pageId="22" pageNumber="23" reason="2">
<taxonomicName id="4C314D49FF86A4622B0EA500FBEDA72D" ID-CoL="55PMG" authority="Santschi, 1910" authorityName="Santschi" authorityYear="1910" box="[547,1041,796,823]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="galoasanum">
<emphasis id="B945EAD8FF86A4622B0EA500FCA0A72D" bold="true" box="[547,860,796,823]" italics="true" pageId="22" pageNumber="23">Tetramorium galoasanum</emphasis>
<bibRefCitation id="EFA04B3BFF86A4622A49A500FBEDA72D" author="Santschi F." box="[868,1041,797,823]" pageId="22" pageNumber="23" pagination="349 - 400" refId="ref31139" refString="Santschi F. 1910. Formicides nouveaux ou peu connus du Congo francais. Annales de la Societe entomologique de France 78: 349 - 400." type="journal article" year="1910">Santschi, 1910</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32B6541FF86A4622BFAA55DFCA0A740" box="[727,860,832,858]" pageId="22" pageNumber="23" type="description">
<paragraph id="8B8E36CAFF86A4622BFAA55DFCA0A740" blockId="22.[547,1041,796,858]" box="[727,860,832,858]" pageId="22" pageNumber="23">
<figureCitation id="130A2A4FFF86A4622BFAA55DFCC8A740" box="[727,820,832,858]" captionStart="Fig" captionStartId="6.[189,232,1824,1850]" captionTargetBox="[189,1399,954,1804]" captionTargetId="figure@6.[189,1399,954,1804]" captionTargetPageId="6" captionText="Fig. 2. Body in profile showing pilosity and pubescence. A. T. dichroum Santschi, 1932 (CASENT0746320). B. T. peringueyi Arnold, 1926 (CASENT0250873). C. T. rothschildi (Forel, 1907) (CASENT0764434). D. T. clunum Forel, 1913 (CASENT0764600). E. T. galoanasum Santschi, 1910 (CASENT0915015 – Antweb, William Ericson). F. T. setuliferum Emery, 1895 (MCZ_ENT00512567). G. T. glabratum Stitz, 1923 (SAM-HYM-C024395). H. T. rufescens Stitz, 1923 (CASENT0250852). I. T. solidum Emery, 1886 (CASENT0250960)." figureDoi="http://doi.org/10.5281/zenodo.3817362" httpUri="https://zenodo.org/record/3817362/files/figure.png" pageId="22" pageNumber="23">Figs 2E</figureCitation>
,
<figureCitation id="130A2A4FFF86A4622A6DA55DFCA0A740" box="[832,860,832,858]" captionStart="Fig" captionStartId="23.[189,232,1965,1991]" captionTargetBox="[206,1382,265,1932]" captionTargetId="figure@23.[206,1382,265,1932]" captionTargetPageId="23" captionText="Fig. 11. Tetramorium galoasanum Santschi, 1910 (RMCAENT000017787 – Antweb, Will Ericson). A. Body in profile. B. Body in dorsal view. C. Head in full-face view." figureDoi="http://doi.org/10.5281/zenodo.3817380" httpUri="https://zenodo.org/record/3817380/files/figure.png" pageId="22" pageNumber="23">11</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32B6541FF86A4622990A59FFC70A781" box="[189,908,897,924]" pageId="22" pageNumber="23" type="reference_group">
<paragraph id="8B8E36CAFF86A4622990A59FFC70A781" blockId="22.[189,908,897,924]" box="[189,908,897,924]" pageId="22" pageNumber="23">
<treatmentCitationGroup id="AB2111E4FF86A4622990A59FFC70A781" box="[189,908,897,924]" pageId="22" pageNumber="23">
<treatmentCitation id="0A9010DBFF86A4622990A59FFC7BA781" author="Santschi F." box="[189,903,897,924]" page="381" pageId="22" pageNumber="23" year="1910">
<taxonomicName id="4C314D49FF86A4622990A59FFC7BA781" authority="Santschi, 1910: 381" authorityName="Santschi" authorityPageNumber="381" authorityYear="1910" box="[189,903,897,924]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="variety" species="setuliferum" variety="galaosana">
<emphasis id="B945EAD8FF86A4622990A59FFE1CA781" box="[189,480,897,923]" italics="true" pageId="22" pageNumber="23">Tetramorium setuliferum</emphasis>
var.
<emphasis id="B945EAD8FF86A4622B37A59CFD68A781" box="[538,660,897,923]" italics="true" pageId="22" pageNumber="23">galaosana</emphasis>
<bibRefCitation id="EFA04B3BFF86A4622BB1A59CFC7BA781" author="Santschi F." box="[668,903,897,923]" pageId="22" pageNumber="23" pagination="349 - 400" refId="ref31139" refString="Santschi F. 1910. Formicides nouveaux ou peu connus du Congo francais. Annales de la Societe entomologique de France 78: 349 - 400." type="journal article" year="1910">Santschi, 1910: 381</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="C32B6541FF86A4622990A5A6FD3BA050" pageId="22" pageNumber="23" type="diagnosis">
<paragraph id="8B8E36CAFF86A4622990A5A6FECAA7CF" blockId="22.[189,310,955,981]" box="[189,310,955,981]" pageId="22" pageNumber="23">
<heading id="D0C681A6FF86A4622990A5A6FECAA7CF" bold="true" box="[189,310,955,981]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="B945EAD8FF86A4622990A5A6FECAA7CF" bold="true" box="[189,310,955,981]" pageId="22" pageNumber="23">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8B8E36CAFF86A4622990A5F6FD3BA050" blockId="22.[189,1399,1002,1099]" pageId="22" pageNumber="23">
<taxonomicName id="4C314D49FF86A4622990A5F6FE90A01E" authorityName="Santschi" authorityYear="1910" box="[189,364,1002,1028]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="galoasanum">
<emphasis id="B945EAD8FF86A4622990A5F6FE90A01E" box="[189,364,1002,1028]" italics="true" pageId="22" pageNumber="23">T. galoasanum</emphasis>
</taxonomicName>
,
<taxonomicName id="4C314D49FF86A4622853A5F6FE0FA01E" authorityName="Forel" authorityYear="1913" box="[382,499,1002,1028]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="clunum">
<emphasis id="B945EAD8FF86A4622853A5F6FE0FA01E" box="[382,499,1002,1028]" italics="true" pageId="22" pageNumber="23">T. clunum</emphasis>
</taxonomicName>
and
<taxonomicName id="4C314D49FF86A4622B1FA5F6FD2AA01E" authorityName="Emery" authorityYear="1895" box="[562,726,1002,1028]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="setuliferum">
<emphasis id="B945EAD8FF86A4622B1FA5F6FD2AA01E" box="[562,726,1002,1028]" italics="true" pageId="22" pageNumber="23">T. setuliferum</emphasis>
</taxonomicName>
are the only species in the
<taxonomicName id="4C314D49FF86A4622D04A5F6FAD4A01E" authorityName="Emery" authorityYear="1886" box="[1065,1320,1002,1028]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="solidum">
<emphasis id="B945EAD8FF86A4622D04A5F6FAD4A01E" box="[1065,1320,1002,1028]" italics="true" pageId="22" pageNumber="23">Tetramorium solidum</emphasis>
</taxonomicName>
group with appressed silvery hairs covering the entire body (
<figureCitation id="130A2A4FFF86A4622A1CA210FC70A03D" box="[817,908,1037,1063]" captionStart="Fig" captionStartId="6.[189,232,1824,1850]" captionTargetBox="[189,1399,954,1804]" captionTargetId="figure@6.[189,1399,954,1804]" captionTargetPageId="6" captionText="Fig. 2. Body in profile showing pilosity and pubescence. A. T. dichroum Santschi, 1932 (CASENT0746320). B. T. peringueyi Arnold, 1926 (CASENT0250873). C. T. rothschildi (Forel, 1907) (CASENT0764434). D. T. clunum Forel, 1913 (CASENT0764600). E. T. galoanasum Santschi, 1910 (CASENT0915015 – Antweb, William Ericson). F. T. setuliferum Emery, 1895 (MCZ_ENT00512567). G. T. glabratum Stitz, 1923 (SAM-HYM-C024395). H. T. rufescens Stitz, 1923 (CASENT0250852). I. T. solidum Emery, 1886 (CASENT0250960)." figureDoi="http://doi.org/10.5281/zenodo.3817362" httpUri="https://zenodo.org/record/3817362/files/figure.png" pageId="22" pageNumber="23">Fig. 2E</figureCitation>
). The appressed silvery hairs are denser in
<taxonomicName id="4C314D49FF86A4622990A22CFE95A050" authorityName="Santschi" authorityYear="1910" box="[189,361,1072,1098]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="galoasanum">
<emphasis id="B945EAD8FF86A4622990A22CFE95A050" box="[189,361,1072,1098]" italics="true" pageId="22" pageNumber="23">T. galoasanum</emphasis>
</taxonomicName>
than in the other two species.
</paragraph>
</subSubSection>
<subSubSection id="C32B6541FF86A4622990A26CFE5AA13E" pageId="22" pageNumber="23" type="materials_examined">
<paragraph id="8B8E36CAFF86A4622990A26CFE56A091" blockId="22.[189,426,1137,1163]" box="[189,426,1137,1163]" pageId="22" pageNumber="23">
<heading id="D0C681A6FF86A4622990A26CFE56A091" bold="true" box="[189,426,1137,1163]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="B945EAD8FF86A4622990A26CFE56A091" bold="true" box="[189,426,1137,1163]" pageId="22" pageNumber="23">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8B8E36CAFF86A46229C7A2BDFEA6A0A0" blockId="22.[189,1399,1184,1316]" box="[234,346,1184,1210]" pageId="22" pageNumber="23">
<materialsCitation id="3B593C97FF86A46229C7A2BDFEA6A0A0" ID-GBIF-Occurrence="2611461323" box="[234,346,1184,1210]" collectingDate="1908-08" collectionCode="NHMB, RMCA, MCZC" collectorName="Weiss" country="Republic of the Congo" location="M'Bounion" municipality="Brazzaville" pageId="22" pageNumber="23" specimenCode="CASENT0915015, RMCAENT000017787" specimenCount="1" typeStatus="syntype">
<emphasis id="B945EAD8FF86A46229C7A2BDFEA6A0A0" bold="true" box="[234,346,1184,1210]" pageId="22" pageNumber="23">
<typeStatus id="548A8868FF86A46229C7A2BDFEA6A0A0" box="[234,346,1184,1210]" pageId="22" pageNumber="23" type="syntype">Syntypes</typeStatus>
</emphasis>
</materialsCitation>
</paragraph>
<paragraph id="8B8E36CAFF86A4622990A2DEFE5AA13E" blockId="22.[189,1399,1184,1316]" pageId="22" pageNumber="23">
<collectingCountry id="F326765AFF86A4622990A2DEFE0AA0C7" box="[189,502,1219,1245]" name="Republic of the Congo" pageId="22" pageNumber="23">REPUBLIC OF CONGO</collectingCountry>
:
<collectingMunicipality id="6BEAACB0FF86A4622B28A2DEFD72A0C7" box="[517,654,1219,1245]" pageId="22" pageNumber="23">Brazzaville</collectingMunicipality>
,
<location id="8EEE6011FF86A4622BB3A2DEFCD6A0C7" LSID="urn:lsid:plazi:treatment:039887DCFF86A4622B0EA500FE11A18E:8EEE6011FF86A4622BB3A2DEFCD6A0C7" box="[670,810,1219,1245]" country="Republic of the Congo" municipality="Brazzaville" name="M'Bounion" pageId="22" pageNumber="23">MʼBounion</location>
,
<location id="8EEE6011FF86A4622A16A2DEFC4BA0C7" LSID="urn:lsid:plazi:treatment:039887DCFF86A4622B0EA500FE11A18E:8EEE6011FF86A4622A16A2DEFC4BA0C7" box="[827,951,1219,1245]" country="Republic of the Congo" municipality="Brazzaville" name="Mindouga" pageId="22" pageNumber="23">Mindouga</location>
,
<location id="8EEE6011FF86A4622AEAA2DEFBA4A0C7" LSID="urn:lsid:plazi:treatment:039887DCFF86A4622B0EA500FE11A18E:8EEE6011FF86A4622AEAA2DEFBA4A0C7" box="[967,1112,1219,1245]" country="Republic of the Congo" municipality="Brazzaville" name="Comba-Ibre" pageId="22" pageNumber="23">Comba-Ibre</location>
,
<date id="FF8F100AFF86A4622D4BA2D9FB15A0C7" box="[1126,1257,1219,1246]" pageId="22" pageNumber="23" value="1908-08">
<collectingDate id="EFCBE9E2FF86A4622D4BA2D9FB15A0C7" box="[1126,1257,1219,1246]" pageId="22" pageNumber="23" value="1908-08">Aug. 1908</collectingDate>
</date>
,
<collectorName id="26C4531CFF86A4622DD7A2DEFAC3A0C7" box="[1274,1343,1219,1245]" pageId="22" pageNumber="23">Weiss</collectorName>
leg. (
<collectionCode id="ED20AE0FFF86A46229E8A2FAFEDFA11B" box="[197,291,1255,1281]" collectionName="Switzerland, Basel, Naturhistorisches Museum" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="22" pageNumber="23">NHMB</collectionCode>
:
<specimenCode id="DB979EB1FF86A4622816A2FBFDE4A11B" box="[315,536,1254,1281]" pageId="22" pageNumber="23">CASENT0915015</specimenCode>
;
<collectionCode id="ED20AE0FFF86A4622B1DA2FAFD77A11B" box="[560,651,1255,1281]" collectionName="Belgium, Tervuren, Musee Royal de l'Afrique Centrale" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="22" pageNumber="23" type="Museum">RMCA</collectionCode>
:
<specimenCode id="DB979EB1FF86A4622B8EA2FAFC46A11B" box="[675,954,1255,1281]" pageId="22" pageNumber="23">RMCAENT000017787</specimenCode>
;
<collectionCode id="ED20AE0FFF86A4622AFEA2FAFBDAA11B" box="[979,1062,1255,1281]" collectionName="USA, Massachusetts, Cambridge, Harvard University, Museum of Comparative Zoology" pageId="22" pageNumber="23">MCZC</collectionCode>
(not examined);
<materialsCitation id="3B593C97FF86A4622C3BA2FAFE5AA13E" ID-GBIF-Occurrence="3899602563" collectingDate="1908-08" collectionCode="NHMB, RMCA, MCZC, BMNH" country="Republic of the Congo" location="M'Bounion" municipality="Brazzaville" pageId="22" pageNumber="23" specimenCode="CASENT0915015, RMCAENT000017787, CASENT0901178" specimenCount="1" typeStatus="syntype">
<collectionCode id="ED20AE0FFF86A4622C3BA2FAFA8FA11B" box="[1302,1395,1255,1281]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="22" pageNumber="23">BMNH</collectionCode>
:
<specimenCode id="DB979EB1FF86A4622990A317FE67A13E" box="[189,411,1290,1316]" pageId="22" pageNumber="23">CASENT0901178</specimenCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32B6541FF86A4622990A356FE11A18E" box="[189,493,1355,1428]" pageId="22" pageNumber="23" type="distribution">
<paragraph id="8B8E36CAFF86A4622990A356FEAAA17F" blockId="22.[189,342,1355,1381]" box="[189,342,1355,1381]" pageId="22" pageNumber="23">
<heading id="D0C681A6FF86A4622990A356FEAAA17F" bold="true" box="[189,342,1355,1381]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="B945EAD8FF86A4622990A356FEAAA17F" bold="true" box="[189,342,1355,1381]" pageId="22" pageNumber="23">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8B8E36CAFF86A4622990A367FE11A18E" blockId="22.[189,493,1402,1428]" box="[189,493,1402,1428]" pageId="22" pageNumber="23">
Only known from
<collectingCountry id="F326765AFF86A46228BAA367FE15A18E" box="[407,489,1402,1428]" name="Democratic Republic of the Congo" pageId="22" pageNumber="23">Congo</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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