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<mods:title id="C1988DF5717F30D14E1649757F9B6992">Croton (Euphorbiaceae) of the Brazilian state of Paraná: an annotated checklist, species distribution, and identification key</mods:title>
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<mods:namePart id="8F57CE84919C7FD6B1F300B7F8E9B7D0">Pereira, Amanda Da Paixão Noronha</mods:namePart>
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<mods:affiliation id="76E5BC2053477BCC2A1D1800CF7DAA46">Instituto de Pesquisas Ambientais, Núcleo de Pesquisa Curadoria do Herbário, São Paulo, SP 01061 - 970, Brazil &amp; Universidade Federal de São Paulo, Instituto de Ciências Ambientais, Químicas e Farmacêuticas, Departamento de Ecologia e Biologia Evolutiva, 09972 - 270, Diadema, SP, Brazil. &amp; amanda. pnoronhap @ gmail. com; https: // orcid. org / 0000 - 0003 - 4874 - 129 X</mods:affiliation>
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<mods:namePart id="9E042D4FF094D18A98B57C0FD97742CF">Riina, Ricarda</mods:namePart>
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<mods:affiliation id="F299E81DDFAE1D8ECB1910F64DEC5D1C">Real Jardín Botánico (RJB), CSIC, Plaza de Murillo 2, 28014, Madrid, Spain. &amp; rriina @ rjb. csic. es; https: // orcid. org / 0000 - 0002 - 7423 - 899 X</mods:affiliation>
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<treatment id="039887C690080473FF44FAEC28CB8061" ID-DOI="http://doi.org/10.5281/zenodo.7268530" ID-GBIF-Taxon="203712492" ID-Zenodo-Dep="7268530" LSID="urn:lsid:plazi:treatment:039887C690080473FF44FAEC28CB8061" httpUri="http://treatment.plazi.org/id/039887C690080473FF44FAEC28CB8061" lastPageId="4" lastPageNumber="235" pageId="2" pageNumber="233">
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<paragraph id="8B8E36D090080475FF44FAEC2FD087CB" blockId="2.[136,689,1381,1408]" box="[136,689,1381,1408]" pageId="2" pageNumber="233">
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Key to the species of
<taxonomicName id="4C314D5390080475FE4EFAEC2CB287CB" ID-CoL="8VWKB" authority="Linnaeus (1753: 1004)" authorityName="Linnaeus" authorityPageNumber="1004" authorityYear="1753" box="[386,467,1382,1408]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="233" phylum="Tracheophyta" rank="genus">
<emphasis id="B945EAC290080475FE4EFAEC2CB287CB" bold="true" box="[386,467,1382,1408]" italics="true" pageId="2" pageNumber="233">Croton</emphasis>
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from
<collectingRegion id="49F5F83290080475FDD7FAEF2FD087CB" box="[539,689,1381,1408]" country="Brazil" name="Parana" pageId="2" pageNumber="233">Paraná state</collectingRegion>
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<keyStep id="30C52E7590080475FF44FA2128CA845D" pageId="2" pageNumber="233">
<paragraph id="8B8E36D090080475FF44FA2128CA8796" blockId="2.[136,1452,1451,2077]" pageId="2" pageNumber="233">
<keyLead id="30C095E590080475FF44FA2128CA8796" pageId="2" pageNumber="233">
1. Prostrate subshrubs. Young branches and leaves with whitish to blackish stellate trichomes. Leaves suborbiculate to reniform ..... ................................................................................................................................................................................
<taxonomicName id="4C314D5390080475FB30FA4D28CA8796" ID-CoL="ZQ5J" authorityName="Muller Argoviensis" authorityYear="1865" box="[1276,1451,1479,1501]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="233" phylum="Tracheophyta" rank="species" species="glechomifolius">
<emphasis id="B945EAC290080475FB30FA4D28CA8796" bold="true" box="[1276,1451,1479,1501]" italics="true" pageId="2" pageNumber="233">C. glechomifolius</emphasis>
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<paragraph id="8B8E36D090080475FF44FA6F28CA845D" blockId="2.[136,1452,1451,2077]" pageId="2" pageNumber="233">
<keyLead id="30C095E590080475FF44FA6F28CA845D" pageId="2" pageNumber="233">- Trees, shrubs or erect subshrubs. Young branches without blackish stellate trichomes. Leaves never suborbiculate or reniform ..... ........................................................................................................................................................................................................... 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E7590080475FF44F99428CA841B" pageId="2" pageNumber="233">
<paragraph id="8B8E36D090080475FF44F99428CA8478" blockId="2.[136,1452,1451,2077]" box="[136,1451,1566,1588]" pageId="2" pageNumber="233">
<keyLead id="30C095E590080475FF44F99428CA8478" box="[136,1451,1566,1588]" pageId="2" pageNumber="233">
2. Leaves with acropetiolar or basilaminar nectary glands (rarely absent in some leaves of
<taxonomicName id="4C314D5390080475FBD2F99429D08478" ID-CoL="ZQ9X" authorityName="Smith &amp; Downs" authorityYear="1959" box="[1054,1201,1566,1587]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="233" phylum="Tracheophyta" rank="species" species="ichthygaster">
<emphasis id="B945EAC290080475FBD2F99429D08478" box="[1054,1201,1566,1587]" italics="true" pageId="2" pageNumber="233">C. ichthygaster</emphasis>
</taxonomicName>
) .................................... 3
</keyLead>
</paragraph>
<paragraph id="8B8E36D090080475FF44F9B128CA841B" blockId="2.[136,1452,1451,2077]" box="[136,1451,1595,1617]" pageId="2" pageNumber="233">
<keyLead id="30C095E590080475FF44F9B128CA841B" box="[136,1451,1595,1617]" pageId="2" pageNumber="233">- Leaves without acropetiolar or basilaminar nectary glands ........................................................................................................... 21</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E7590080475FF44F9D228CA84ED" pageId="2" pageNumber="233">
<paragraph id="8B8E36D090080475FF44F9D228CB8426" blockId="2.[136,1452,1451,2077]" box="[136,1450,1623,1645]" pageId="2" pageNumber="233">
<keyLead id="30C095E590080475FF44F9D228CB8426" box="[136,1450,1623,1645]" pageId="2" pageNumber="233">
3. Young branches angular, costate; with lepidote subentire trichomes. Pistillate flowers urceolate .................................
<taxonomicName id="4C314D5390080475FAF5F9DD28CB8426" authorityName="Casaretto" authorityYear="1845" box="[1337,1450,1623,1645]" class="Magnoliopsida" family="Euphorbiaceae" genus="Cleodora" higherTaxonomySource="GBIF" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="233" phylum="Tracheophyta" rank="species" species="salutaris">
<emphasis id="B945EAC290080475FAF5F9DD28CB8426" bold="true" box="[1337,1450,1623,1645]" italics="true" pageId="2" pageNumber="233">C. salutaris</emphasis>
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<paragraph id="8B8E36D090080475FF44F9FF28CA84ED" blockId="2.[136,1452,1451,2077]" pageId="2" pageNumber="233">
<keyLead id="30C095E590080475FF44F9FF28CA84ED" pageId="2" pageNumber="233">- Young branches cylindrical to slightly flattened, never costate; with stellate, stellate-porrect, stellate-lepidote, rosulate or dendritic trichomes. Pistillate flowers rotate, campanulate or subcampanulate .............................................................................................. 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E7590080475FF44F92428CA8551" pageId="2" pageNumber="233">
<paragraph id="8B8E36D090080475FF44F92428CA84AB" blockId="2.[136,1452,1451,2077]" pageId="2" pageNumber="233">
<keyLead id="30C095E590080475FF44F92428CA84AB" pageId="2" pageNumber="233">
4. Young branches with a dense, hirsute indumentum of stellate-porrect trichomes. Inflorescences densely congested. ...................... .........................................................................................................................................................................................
<taxonomicName id="4C314D5390080475FAFBF94028CA84AB" ID-CoL="ZPFM" authorityName="Muller Argoviensis" authorityYear="1873" box="[1335,1451,1738,1760]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="233" phylum="Tracheophyta" rank="species" species="aberrans">
<emphasis id="B945EAC290080475FAFBF94028CA84AB" bold="true" box="[1335,1451,1738,1760]" italics="true" pageId="2" pageNumber="233">C. aberrans</emphasis>
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<paragraph id="8B8E36D090080475FF44F96228CA8551" blockId="2.[136,1452,1451,2077]" pageId="2" pageNumber="233">
<keyLead id="30C095E590080475FF44F96228CA8551" pageId="2" pageNumber="233">- Young branches pilose, sparsely to densely pubescent with stellate, stellate-lepidote, rosulate or dendritic trichomes. Inflorescences lax or congested ................................................................................................................................................................................ 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E7590080475FF44F8AB28CA853A" pageId="2" pageNumber="233">
<paragraph id="8B8E36D090080475FF44F8AB28CA857C" blockId="2.[136,1452,1451,2077]" box="[136,1451,1825,1847]" pageId="2" pageNumber="233">
<keyLead id="30C095E590080475FF44F8AB28CA857C" box="[136,1451,1825,1847]" pageId="2" pageNumber="233">
5. Stipitate acropetiolar nectary glands, facing up (adaxial side). Leaves with stellate-lepidote trichomes ..............
<taxonomicName id="4C314D5390080475FACBF8AB28CA857C" ID-CoL="ZQH6" authorityName="Baillon" authorityYear="1864" box="[1287,1451,1825,1847]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="233" phylum="Tracheophyta" rank="species" species="macrobothrys">
<emphasis id="B945EAC290080475FACBF8AB28CA857C" bold="true" box="[1287,1451,1825,1847]" italics="true" pageId="2" pageNumber="233">C. macrobothrys</emphasis>
</taxonomicName>
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</paragraph>
<paragraph id="8B8E36D090080475FF44F8B428CA853A" blockId="2.[136,1452,1451,2077]" pageId="2" pageNumber="233">
<keyLead id="30C095E590080475FF44F8B428CA853A" pageId="2" pageNumber="233">- Stipitate acropetiolar, facing down (abaxial side), sessile acropetiolar or basilaminar nectary glands. Leaves with stellate, rosulate or dendritic trichomes ....................................................................................................................................................................... 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E7590080475FF44F8FD28CA85E1" pageId="2" pageNumber="233">
<paragraph id="8B8E36D090080475FF44F8FD28CA85C6" blockId="2.[136,1452,1451,2077]" box="[136,1451,1911,1933]" pageId="2" pageNumber="233">
<keyLead id="30C095E590080475FF44F8FD28CA85C6" box="[136,1451,1911,1933]" pageId="2" pageNumber="233">
6. Stamens 80130. Styles basally connate, forming a column at the top of the ovary .................................................
<taxonomicName id="4C314D5390080475FAD0F8FD28CA85C6" ID-CoL="6BK6Y" authorityName="Baillon" authorityYear="1864" box="[1308,1451,1911,1933]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="233" phylum="Tracheophyta" rank="species" species="vulnerarius">
<emphasis id="B945EAC290080475FAD0F8FD28CA85C6" bold="true" box="[1308,1451,1911,1933]" italics="true" pageId="2" pageNumber="233">C. vulnerarius</emphasis>
</taxonomicName>
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</paragraph>
<paragraph id="8B8E36D090080475FF44F81F28CA85E1" blockId="2.[136,1452,1451,2077]" box="[136,1451,1940,1962]" pageId="2" pageNumber="233">
<keyLead id="30C095E590080475FF44F81F28CA85E1" box="[136,1451,1940,1962]" pageId="2" pageNumber="233">- Stamens up to 60. Styles free, not forming a column at the top of the ovary .................................................................................. 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E7590080475FF44F83B28CA8A4B" pageId="2" pageNumber="233">
<paragraph id="8B8E36D090080475FF44F83B28CA858C" blockId="2.[136,1452,1451,2077]" box="[136,1451,1969,1991]" pageId="2" pageNumber="233">
<keyLead id="30C095E590080475FF44F83B28CA858C" box="[136,1451,1969,1991]" pageId="2" pageNumber="233">
7. Adaxial leaf surface glabrescent. Pistillate and staminate flowers reduplicate-valvate. Fruits strongly muricate ...........
<taxonomicName id="4C314D5390080475FA88F83B28CA858C" ID-CoL="ZPZT" authorityName="Muller Argoviensis" authorityYear="1873" box="[1348,1451,1969,1991]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="233" phylum="Tracheophyta" rank="species" species="eichleri">
<emphasis id="B945EAC290080475FA88F83B28CA858C" bold="true" box="[1348,1451,1969,1991]" italics="true" pageId="2" pageNumber="233">C. eichleri</emphasis>
</taxonomicName>
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</paragraph>
<paragraph id="8B8E36D090080475FF44F84428CA8A4B" blockId="2.[136,1452,1451,2077]" pageId="2" pageNumber="233">
<keyLead id="30C095E590080475FF44F84428CA8A4B" pageId="2" pageNumber="233">- Adaxial leaf surface glabrescent to pubescent. Pistillate and staminate flowers valvate. Fruits smooth or inconspicuously verrucose ........................................................................................................................................................................................... 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E7590080474FF44F78228CA82FF" lastPageId="3" lastPageNumber="234" pageId="2" pageNumber="233">
<paragraph id="8B8E36D090080475FF44F78228CA8A56" blockId="2.[136,1452,1451,2077]" box="[136,1451,2055,2077]" pageId="2" pageNumber="233">
<keyLead id="30C095E590080475FF44F78228CA8A56" box="[136,1451,2055,2077]" pageId="2" pageNumber="233">
8. Inflorescences with a sterile zone between pistillate and staminate flowers ...............................................................
<taxonomicName id="4C314D5390080475FAE7F78D28CA8A56" ID-CoL="ZQGT" authorityName="Muller Argoviensis" authorityYear="1866" baseAuthorityName="Didrichsen" baseAuthorityYear="1857" box="[1323,1451,2055,2077]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="233" phylum="Tracheophyta" rank="species" species="lundianus">
<emphasis id="B945EAC290080475FAE7F78D28CA8A56" bold="true" box="[1323,1451,2055,2077]" italics="true" pageId="2" pageNumber="233">C. lundianus</emphasis>
</taxonomicName>
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<paragraph id="8B8E36D090090474FF44FF2A28CA82FF" blockId="3.[136,1453,159,2082]" box="[136,1451,159,181]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44FF2A28CA82FF" box="[136,1451,159,181]" pageId="3" pageNumber="234">- Inflorescences without a sterile zone between pistillate and staminate flowers .............................................................................. 9</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E7590090474FF44FF3628CA82A5" pageId="3" pageNumber="234">
<paragraph id="8B8E36D090090474FF44FF3628CA829A" blockId="3.[136,1453,159,2082]" box="[136,1451,187,210]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44FF3628CA829A" box="[136,1451,187,210]" pageId="3" pageNumber="234">
9. Nectary glands stipitate to shortly stipitate, acropetiolar. Stamens ca. 60. Fruits inconspicuously verrucose ...........
<taxonomicName id="4C314D5390090474FAD7FF3628CA829A" ID-CoL="ZPS4" authorityName="Baillon" authorityYear="1864" box="[1307,1451,187,209]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="234" phylum="Tracheophyta" rank="species" species="celtidifolius">
<emphasis id="B945EAC290090474FAD7FF3628CA829A" bold="true" box="[1307,1451,187,209]" italics="true" pageId="3" pageNumber="234">C. celtidifolius</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8E36D090090474FF44FF5328CA82A5" blockId="3.[136,1453,159,2082]" box="[136,1451,217,239]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44FF5328CA82A5" box="[136,1451,217,239]" pageId="3" pageNumber="234">- Nectary glands, sessile, basilaminar, sometimes absent in a few leaves of the same plant. Stamens up to 20. Fruits smooth ...... 10</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E7590090474FF44FF7C28CA836C" pageId="3" pageNumber="234">
<paragraph id="8B8E36D090090474FF44FF7C28CB8340" blockId="3.[136,1453,159,2082]" box="[136,1450,245,267]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44FF7C28CB8340" box="[136,1450,245,267]" pageId="3" pageNumber="234">
10. Leaf apex slightly emarginate to emarginate, sometimes mucronate. Nectary glands absent in some leaves ..........
<taxonomicName id="4C314D5390090474FAD9FF7F28CB8340" ID-CoL="ZQ9X" authorityName="Smith &amp; Downs" authorityYear="1959" box="[1301,1450,245,267]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="234" phylum="Tracheophyta" rank="species" species="ichthygaster">
<emphasis id="B945EAC290090474FAD9FF7F28CB8340" bold="true" box="[1301,1450,245,267]" italics="true" pageId="3" pageNumber="234">C. ichthygaster</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8E36D090090474FF44FE9928CA836C" blockId="3.[136,1453,159,2082]" box="[136,1451,274,296]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44FE9928CA836C" box="[136,1451,274,296]" pageId="3" pageNumber="234">- Leaf apex neither emarginate nor mucronate. Nectary glands present in all leaves ...................................................................... 11</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E7590090474FF44FEA528CA832A" pageId="3" pageNumber="234">
<paragraph id="8B8E36D090090474FF44FEA528CB830E" blockId="3.[136,1453,159,2082]" box="[136,1450,303,325]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44FEA528CB830E" box="[136,1450,303,325]" pageId="3" pageNumber="234">
11. Adaxial leaf surface glabrous ..................................................................................................................................
<taxonomicName id="4C314D5390090474FAC1FEA528CB830E" ID-CoL="6BK6T" authorityName="Smith &amp; Downs" authorityYear="1959" box="[1293,1450,303,325]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="234" phylum="Tracheophyta" rank="species" species="polygonoides">
<emphasis id="B945EAC290090474FAC1FEA528CB830E" bold="true" box="[1293,1450,303,325]" italics="true" pageId="3" pageNumber="234">C. polygonoides</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8E36D090090474FF44FEC628CA832A" blockId="3.[136,1453,159,2082]" box="[136,1451,332,354]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44FEC628CA832A" box="[136,1451,332,354]" pageId="3" pageNumber="234">- Adaxial leaf surface with trichomes ............................................................................................................................................... 12</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E7590090474FF44FEE328CA83D0" pageId="3" pageNumber="234">
<paragraph id="8B8E36D090090474FF44FEE328CA8335" blockId="3.[136,1453,159,2082]" box="[136,1451,360,383]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44FEE328CA8335" box="[136,1451,360,383]" pageId="3" pageNumber="234">
12. Xylopodium present. Leaves with patelliform nectary glands along the margin .................................................
<taxonomicName id="4C314D5390090474FB34FEE328CA8335" ID-CoL="ZPJJ" box="[1272,1451,360,382]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="234" phylum="Tracheophyta" rank="species" species="antisyphiliticus">
<emphasis id="B945EAC290090474FB34FEE328CA8335" bold="true" box="[1272,1451,360,382]" italics="true" pageId="3" pageNumber="234">C. antisyphiliticus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8E36D090090474FF44FE0C28CA83D0" blockId="3.[136,1453,159,2082]" box="[136,1451,389,411]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44FE0C28CA83D0" box="[136,1451,389,411]" pageId="3" pageNumber="234">- Xylopodium absent. Leaves without patelliform nectary glands along the margin ....................................................................... 13</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E7590090474FF44FE2828CA839E" pageId="3" pageNumber="234">
<paragraph id="8B8E36D090090474FF44FE2828CA83FC" blockId="3.[136,1453,159,2082]" box="[136,1451,418,440]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44FE2828CA83FC" box="[136,1451,418,440]" pageId="3" pageNumber="234">13. Subshrubs. Leaf margin irregularly dentate ................................................................................................................................... 14</keyLead>
</paragraph>
<paragraph id="8B8E36D090090474FF44FE4A28CA839E" blockId="3.[136,1453,159,2082]" box="[136,1451,447,469]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44FE4A28CA839E" box="[136,1451,447,469]" pageId="3" pageNumber="234">- Shrubs to trees. Leaf margin entire to slightly serrate..................................................................................................................... 15</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E7590090474FF44FE5628CA8045" pageId="3" pageNumber="234">
<paragraph id="8B8E36D090090474FF44FE5628CA83BA" blockId="3.[136,1453,159,2082]" box="[136,1451,475,498]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44FE5628CA83BA" box="[136,1451,475,498]" pageId="3" pageNumber="234">
14. Bracts with glands. Pistillate flowers sessile; sepals unequal; styles pubescent .......................................................
<taxonomicName id="4C314D5390090474FADBFE5628CA83BA" ID-CoL="ZQ5G" authorityName="Linnaeus (1759:" authorityYear="1275" box="[1303,1451,475,497]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="234" phylum="Tracheophyta" rank="species" species="glandulosus">
<emphasis id="B945EAC290090474FADBFE5628CA83BA" bold="true" box="[1303,1451,475,497]" italics="true" pageId="3" pageNumber="234">C. glandulosus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8E36D090090474FF44FE7328CA8045" blockId="3.[136,1453,159,2082]" box="[136,1451,505,527]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44FE7328CA8045" box="[136,1451,505,527]" pageId="3" pageNumber="234">
- Bracts without glands. Pistillate flowers pedicellate, sometimes pedicels recurved; sepals equal; styles glabrous .......
<taxonomicName id="4C314D5390090474FAF7FE7328CA8045" ID-CoL="ZRBM" authorityName="Millspaugh" authorityYear="1900" box="[1339,1451,505,526]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="234" phylum="Tracheophyta" rank="species" species="trinitatis">
<emphasis id="B945EAC290090474FAF7FE7328CA8045" bold="true" box="[1339,1451,505,526]" italics="true" pageId="3" pageNumber="234">C. trinitatis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E7590090474FF44FD9F28CA80C9" pageId="3" pageNumber="234">
<paragraph id="8B8E36D090090474FF44FD9F28CA8003" blockId="3.[136,1453,159,2082]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44FD9F28CA8003" pageId="3" pageNumber="234">
15. Acropetiolar nectary glands only visible from below. Inflorescences with bisexual lower cymules. Pistillate flowers pedicellate. Columella apex inconspicuous and unlobed ...............................................................................................................
<taxonomicName id="4C314D5390090474FAE8FDB828CA8003" ID-CoL="ZRCL" authorityName="Baillon" authorityYear="1864" box="[1316,1451,562,584]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="234" phylum="Tracheophyta" rank="species" species="urucurana">
<emphasis id="B945EAC290090474FAE8FDB828CA8003" bold="true" box="[1316,1451,562,584]" italics="true" pageId="3" pageNumber="234">C. urucurana</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8E36D090090474FF44FDDA28CA80C9" blockId="3.[136,1453,159,2082]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44FDDA28CA80C9" pageId="3" pageNumber="234">- Acropetiolar nectary glands only visible from above. Inflorescences with unisexual lower cymules. Pistillate flowers sessile to subsessile. Columella apex with three inflated lobes ..................................................................................................................... 16</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E7590090474FF44FD0328CA80F0" pageId="3" pageNumber="234">
<paragraph id="8B8E36D090090474FF44FD0328CA80D5" blockId="3.[136,1453,159,2082]" box="[136,1451,649,671]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44FD0328CA80D5" box="[136,1451,649,671]" pageId="3" pageNumber="234">
16. Inflorescences congested ......................................................................................................................................................
<taxonomicName id="4C314D5390090474FA9BFD0328CA80D5" ID-CoL="ZQYG" authorityName="Smith &amp; Downs" authorityYear="1959" box="[1367,1451,649,670]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="234" phylum="Tracheophyta" rank="species" species="reitzii">
<emphasis id="B945EAC290090474FA9BFD0328CA80D5" bold="true" box="[1367,1451,649,670]" italics="true" pageId="3" pageNumber="234">C. reitzii</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8E36D090090474FF44FD2C28CA80F0" blockId="3.[136,1453,159,2082]" box="[136,1451,677,699]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44FD2C28CA80F0" box="[136,1451,677,699]" pageId="3" pageNumber="234">- Inflorescences lax ........................................................................................................................................................................... 17</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E7590090474FF44FD4828CA80BF" pageId="3" pageNumber="234">
<paragraph id="8B8E36D090090474FF44FD4828CA8093" blockId="3.[136,1453,159,2082]" box="[136,1451,706,728]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44FD4828CA8093" box="[136,1451,706,728]" pageId="3" pageNumber="234">
17. Styles tetrafid (12 terminal tips). Fruits with stellate-rosulate trichomes ..................................................................
<taxonomicName id="4C314D5390090474FAD3FD4828CA8093" ID-CoL="75FSP" authorityName="Muller Argoviensis" authorityYear="1873" box="[1311,1451,706,728]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="234" phylum="Tracheophyta" rank="species" species="thermarum">
<emphasis id="B945EAC290090474FAD3FD4828CA8093" bold="true" box="[1311,1451,706,728]" italics="true" pageId="3" pageNumber="234">C. thermarum</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8E36D090090474FF44FD6A28CA80BF" blockId="3.[136,1453,159,2082]" box="[136,1451,735,757]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44FD6A28CA80BF" box="[136,1451,735,757]" pageId="3" pageNumber="234">- Styles bifid (6 terminal tips). Fruits with stellate (no rosulate) trichomes ..................................................................................... 18</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E7590090474FF44FD7628CA8165" pageId="3" pageNumber="234">
<paragraph id="8B8E36D090090474FF44FD7628CB815A" blockId="3.[136,1453,159,2082]" box="[136,1450,763,786]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44FD7628CB815A" box="[136,1450,763,786]" pageId="3" pageNumber="234">
18. Young branches smooth. Styles densely pubescent (at least half of its length) ................................................................
<taxonomicName id="4C314D5390090474FA89FD7628CB815A" ID-CoL="ZQDP" authorityName="Lamarck" authorityYear="1786" box="[1349,1450,763,785]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="234" phylum="Tracheophyta" rank="species" species="lanatus">
<emphasis id="B945EAC290090474FA89FD7628CB815A" bold="true" box="[1349,1450,763,785]" italics="true" pageId="3" pageNumber="234">C. lanatus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8E36D090090474FF44FC9328CA8165" blockId="3.[136,1453,159,2082]" box="[136,1451,793,815]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44FC9328CA8165" box="[136,1451,793,815]" pageId="3" pageNumber="234">- Young branches striate. Styles sparsely pubescent (just at the base) or glabrous .......................................................................... 19</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E7590090474FF44FCBC28CA812C" pageId="3" pageNumber="234">
<paragraph id="8B8E36D090090474FF44FCBC28CA8101" blockId="3.[136,1453,159,2082]" box="[136,1451,821,843]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44FCBC28CA8101" box="[136,1451,821,843]" pageId="3" pageNumber="234">
19. Inflorescences axis cylindrical. Staminate flowers sessile to subsessile ..............................................................
<taxonomicName id="4C314D5390090474FACCFCBF28CA8101" ID-CoL="ZR2W" authorityName="Moore" authorityYear="1865" box="[1280,1451,821,842]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="234" phylum="Tracheophyta" rank="species" species="sanctae-crucis">
<emphasis id="B945EAC290090474FACCFCBF28CA8101" bold="true" box="[1280,1451,821,842]" italics="true" pageId="3" pageNumber="234">C. sanctae-crucis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8E36D090090474FF44FCD928CA812C" blockId="3.[136,1453,159,2082]" box="[136,1451,850,872]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44FCD928CA812C" box="[136,1451,850,872]" pageId="3" pageNumber="234">- Inflorescences axis flattened. Staminate flowers pedicellate ......................................................................................................... 20</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E7590090474FF44FCE528CA81B3" pageId="3" pageNumber="234">
<paragraph id="8B8E36D090090474FF44FCE528CA81F5" blockId="3.[136,1453,159,2082]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44FCE528CA81F5" pageId="3" pageNumber="234">
20. Young branches glabrescent. Leaves membranous, ovate-lanceolate to lanceolate, abaxial surface glabrescent. Stipules without a minute globose gland (colleter) at the apex. Pistillate flowers without nectary glands in place of petals. Fruits ellipsoid to rarely obovoid .................................................................................................................................................................
<taxonomicName id="4C314D5390090474FB37FC2328CA81F5" ID-CoL="ZPNW" authorityName="Baillon" authorityYear="1864" box="[1275,1451,936,958]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="234" phylum="Tracheophyta" rank="species" species="bonplandianus">
<emphasis id="B945EAC290090474FB37FC2328CA81F5" bold="true" box="[1275,1451,936,958]" italics="true" pageId="3" pageNumber="234">C. bonplandianus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8E36D090090474FF44FC4C28CA81B3" blockId="3.[136,1453,159,2082]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44FC4C28CA81B3" pageId="3" pageNumber="234">
- Young branches pubescent. Leaves chartaceus, ovate to cordate, abaxial surface pubescent. Stipules with a minute globose gland (colleter) at the apex. Pistillate flowers with nectary glands in place of petals. Fruits obovoid ...................................
<taxonomicName id="4C314D5390090474FAE3FC6828CA81B3" ID-CoL="ZQ66" authorityName="Baillon" authorityYear="1864" box="[1327,1451,994,1016]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="234" phylum="Tracheophyta" rank="species" species="gracilipes">
<emphasis id="B945EAC290090474FAE3FC6828CA81B3" bold="true" box="[1327,1451,994,1016]" italics="true" pageId="3" pageNumber="234">C. gracilipes</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E7590090474FF44FC7528CA867A" pageId="3" pageNumber="234">
<paragraph id="8B8E36D090090474FF44FC7528CA865E" blockId="3.[136,1453,159,2082]" box="[136,1451,1023,1045]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44FC7528CA865E" box="[136,1451,1023,1045]" pageId="3" pageNumber="234">
21. Trees. Adaxial leaf surface with hispid indumentum. Fruits conspicuously muricate ...............................................
<taxonomicName id="4C314D5390090474FAD7FC7528CA865E" ID-CoL="ZQ3Y" authorityName="Sprengel" authorityYear="1826" box="[1307,1451,1023,1045]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="234" phylum="Tracheophyta" rank="species" species="floribundus">
<emphasis id="B945EAC290090474FAD7FC7528CA865E" bold="true" box="[1307,1451,1023,1045]" italics="true" pageId="3" pageNumber="234">C. floribundus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8E36D090090474FF44FB9628CA867A" blockId="3.[136,1453,159,2082]" box="[136,1451,1052,1074]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44FB9628CA867A" box="[136,1451,1052,1074]" pageId="3" pageNumber="234">
- Shrubs to subshrubs. Adaxial leaf surface glabrous or other
<typeStatus id="548A887290090474FCDCFB962E58867A" box="[784,825,1052,1073]" pageId="3" pageNumber="234">type</typeStatus>
indumentum. Fruits smooth .................................................... 22
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E7590090474FF44FBB328CA8620" pageId="3" pageNumber="234">
<paragraph id="8B8E36D090090474FF44FBB328CB8605" blockId="3.[136,1453,159,2082]" box="[136,1450,1080,1103]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44FBB328CB8605" box="[136,1450,1080,1103]" pageId="3" pageNumber="234">
22. Staminate flowers reduplicate-valvate. Styles multifid .....................................................................................................
<taxonomicName id="4C314D5390090474FA89FBB328CB8605" ID-CoL="ZRB9" authorityName="Klotzsch ex Baillon" authorityYear="1864" box="[1349,1450,1080,1102]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="234" phylum="Tracheophyta" rank="species" species="tricolor">
<emphasis id="B945EAC290090474FA89FBB328CB8605" bold="true" box="[1349,1450,1080,1102]" italics="true" pageId="3" pageNumber="234">C. tricolor</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8E36D090090474FF44FBDC28CA8620" blockId="3.[136,1453,159,2082]" box="[136,1451,1109,1131]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44FBDC28CA8620" box="[136,1451,1109,1131]" pageId="3" pageNumber="234">- Staminate flowers valvate. Styles bifid, twice bifid or tetrafid ...................................................................................................... 23</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E7590090474FF44FBF828CA86EE" pageId="3" pageNumber="234">
<paragraph id="8B8E36D090090474FF44FBF828CA86CC" blockId="3.[136,1453,159,2082]" box="[136,1451,1138,1160]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44FBF828CA86CC" box="[136,1451,1138,1160]" pageId="3" pageNumber="234">23. Columella apex with three inflated lobes ....................................................................................................................................... 24</keyLead>
</paragraph>
<paragraph id="8B8E36D090090474FF44FB1A28CA86EE" blockId="3.[136,1453,159,2082]" box="[136,1451,1167,1189]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44FB1A28CA86EE" box="[136,1451,1167,1189]" pageId="3" pageNumber="234">- Columella apex inconspicuous and unlobed .................................................................................................................................. 26</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E7590090474FF44FB2628CA8695" pageId="3" pageNumber="234">
<paragraph id="8B8E36D090090474FF44FB2628CB868A" blockId="3.[136,1453,159,2082]" box="[136,1450,1195,1218]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44FB2628CB868A" box="[136,1450,1195,1218]" pageId="3" pageNumber="234">
24. Shrubs
<quantity id="4CC99B3590090474FED7FB262C35868A" box="[283,340,1196,1217]" metricMagnitude="0" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="3" pageNumber="234" unit="m" value="1.5" valueMax="2.0" valueMin="1.0">1-2 m</quantity>
tall, densely branched, stems striate ..............................................................................................................
<taxonomicName id="4C314D5390090474FA9DFB2628CB868A" ID-CoL="ZQ4M" box="[1361,1450,1195,1217]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="234" phylum="Tracheophyta" rank="species" species="fulvus">
<emphasis id="B945EAC290090474FA9DFB2628CB868A" bold="true" box="[1361,1450,1195,1217]" italics="true" pageId="3" pageNumber="234">C. fulvus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
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- Subshrubs to shrubs up to
<quantity id="4CC99B3590090474FE1DFB422F6D8695" box="[465,524,1224,1246]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="234" unit="cm" value="50.0">50 cm</quantity>
tall, little branched, stems smooth ............................................................................................. 25
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E7590090474FF44FB6F28CB8753" pageId="3" pageNumber="234">
<paragraph id="8B8E36D090090474FF44FB6F28CA86B0" blockId="3.[136,1453,159,2082]" box="[136,1451,1253,1275]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44FB6F28CA86B0" box="[136,1451,1253,1275]" pageId="3" pageNumber="234">
25. Stipules narrowly lanceolate with numerous glands (colleters). Staminate bracts narrowly lanceolate ....................
<taxonomicName id="4C314D5390090474FAD7FB6F28CA86B0" ID-CoL="5NCMK" authorityName="Baillon" authorityYear="1864" box="[1307,1451,1253,1275]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="234" phylum="Tracheophyta" rank="species" species="grandivelus">
<emphasis id="B945EAC290090474FAD7FB6F28CA86B0" bold="true" box="[1307,1451,1253,1275]" italics="true" pageId="3" pageNumber="234">C. grandivelus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8E36D090090474FF44FA8928CB8753" blockId="3.[136,1453,159,2082]" box="[136,1450,1282,1304]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44FA8928CB8753" box="[136,1450,1282,1304]" pageId="3" pageNumber="234">
- Stipules glanduliform, with 5 glands (colleters). Staminate bracts ovate or lanceolate .............................................
<taxonomicName id="4C314D5390090474FAEDFA8828CB8753" ID-CoL="ZR8G" authorityName="Muller Argoviensis" authorityYear="1873" box="[1313,1450,1282,1304]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="234" phylum="Tracheophyta" rank="species" species="subvillosus">
<emphasis id="B945EAC290090474FAEDFA8828CB8753" bold="true" box="[1313,1450,1282,1304]" italics="true" pageId="3" pageNumber="234">C. subvillosus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E7590090474FF44FA9528CA871A" pageId="3" pageNumber="234">
<paragraph id="8B8E36D090090474FF44FA9528CB877E" blockId="3.[136,1453,159,2082]" box="[136,1450,1311,1333]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44FA9528CB877E" box="[136,1450,1311,1333]" pageId="3" pageNumber="234">
26. Leaves adaxial surface with lepidote subentire trichomes .........................................................................................
<taxonomicName id="4C314D5390090474FAD1FA9528CB877E" ID-CoL="ZQNR" authorityName="Muller Argoviensis" authorityYear="1865" box="[1309,1450,1311,1333]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="234" phylum="Tracheophyta" rank="species" species="myrianthus">
<emphasis id="B945EAC290090474FAD1FA9528CB877E" bold="true" box="[1309,1450,1311,1333]" italics="true" pageId="3" pageNumber="234">C. myrianthus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8E36D090090474FF44FAB628CA871A" blockId="3.[136,1453,159,2082]" box="[136,1451,1340,1362]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44FAB628CA871A" box="[136,1451,1340,1362]" pageId="3" pageNumber="234">- Leaves adaxial surface glabrous or with stellate or appressed-stellate trichomes .......................................................................... 27</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E7590090474FF44FAD328CA87C0" pageId="3" pageNumber="234">
<paragraph id="8B8E36D090090474FF44FAD328CA8725" blockId="3.[136,1453,159,2082]" box="[136,1451,1368,1391]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44FAD328CA8725" box="[136,1451,1368,1391]" pageId="3" pageNumber="234">
27. Leaves venation hyphodromous (only midrib visible) .............................................................................................
<taxonomicName id="4C314D5390090474FAC3FAD328CA8725" ID-CoL="ZR4J" box="[1295,1451,1368,1390]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="234" phylum="Tracheophyta" rank="species" species="serpyllifolius">
<emphasis id="B945EAC290090474FAC3FAD328CA8725" bold="true" box="[1295,1451,1368,1390]" italics="true" pageId="3" pageNumber="234">C. serpyllifolius</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8E36D090090474FF44FAFC28CA87C0" blockId="3.[136,1453,159,2082]" box="[136,1451,1397,1419]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44FAFC28CA87C0" box="[136,1451,1397,1419]" pageId="3" pageNumber="234">
- Leaves with other venation
<typeStatus id="548A887290090474FE11FAFC2F6787C0" box="[477,518,1398,1419]" pageId="3" pageNumber="234">type</typeStatus>
..................................................................................................................................................... 28
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E7590090474FF44FA1828CA87B5" pageId="3" pageNumber="234">
<paragraph id="8B8E36D090090474FF44FA1828CA878E" blockId="3.[136,1453,159,2082]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44FA1828CA878E" pageId="3" pageNumber="234">
28. Indumentum yellowish to orange. Stipules reduced to minute conic glands (colleters). Staminate petals cream with a central dark line .............................................................................................................................................................................
<taxonomicName id="4C314D5390090474FAD4FA2528CA878E" box="[1304,1451,1455,1477]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="234" phylum="Tracheophyta" rank="species" species="chamaepitis">
<emphasis id="B945EAC290090474FAD4FA2528CA878E" bold="true" box="[1304,1451,1455,1477]" italics="true" pageId="3" pageNumber="234">C. chamaepitis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8E36D090090474FF44FA4628CA87B5" blockId="3.[136,1453,159,2082]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44FA4628CA87B5" pageId="3" pageNumber="234">- Indumentum whitish, yellowish, cinereous, silvery, cream or ferrugineous. Stipules inconspicuous, not reduced to glands (colleters). Staminate petals cream or white without a central dark line .......................................................................................................... 29</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E7590090474FF44F98C28CA847C" pageId="3" pageNumber="234">
<paragraph id="8B8E36D090090474FF44F98C28CA8450" blockId="3.[136,1453,159,2082]" box="[136,1451,1541,1563]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44F98C28CA8450" box="[136,1451,1541,1563]" pageId="3" pageNumber="234">
29. Leaves up to
<quantity id="4CC99B3590090474FE9FF98F2CF58450" box="[339,404,1541,1563]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="234" unit="cm" value="1.5">1.5 cm</quantity>
long. Styles twice bifid .........................................................................................................
<taxonomicName id="4C314D5390090474FB30F98F28CA8450" ID-CoL="ZQXC" authorityName="Baillon" authorityYear="1864" box="[1276,1451,1541,1563]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="234" phylum="Tracheophyta" rank="species" species="pycnocephalus">
<emphasis id="B945EAC290090474FB30F98F28CA8450" bold="true" box="[1276,1451,1541,1563]" italics="true" pageId="3" pageNumber="234">C. pycnocephalus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8E36D090090474FF44F9A928CA847C" blockId="3.[136,1453,159,2082]" box="[136,1451,1570,1592]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44F9A928CA847C" box="[136,1451,1570,1592]" pageId="3" pageNumber="234">
- Leaves
<quantity id="4CC99B3590090474FED1F9A82C2D8473" box="[285,332,1570,1592]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="234" unit="cm" value="3.0">3 cm</quantity>
long or longer. Styles bifid or tetrafid ....................................................................................................................... 30
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E7590090474FF44F9B528CA843A" pageId="3" pageNumber="234">
<paragraph id="8B8E36D090090474FF44F9B528CA841F" blockId="3.[136,1453,159,2082]" box="[136,1451,1599,1621]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44F9B528CA841F" box="[136,1451,1599,1621]" pageId="3" pageNumber="234">30. Leaves with abaxial surface covered by lepidote subentire or stellate-lepidote trichomes. Styles bifid ....................................... 31</keyLead>
</paragraph>
<paragraph id="8B8E36D090090474FF44F9D628CA843A" blockId="3.[136,1453,159,2082]" box="[136,1451,1628,1650]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44F9D628CA843A" box="[136,1451,1628,1650]" pageId="3" pageNumber="234">- Leaves with abaxial surface covered by stellate trichomes. Styles tetrafid ................................................................................... 33</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E7590090474FF44F9F328CA84E0" pageId="3" pageNumber="234">
<paragraph id="8B8E36D090090474FF44F9F328CA84C5" blockId="3.[136,1453,159,2082]" box="[136,1451,1657,1679]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44F9F328CA84C5" box="[136,1451,1657,1679]" pageId="3" pageNumber="234">31. Leaves adaxial surface glabrous. Pistillate flowers subcampanulate ............................................................................................. 32</keyLead>
</paragraph>
<paragraph id="8B8E36D090090474FF44F91C28CA84E0" blockId="3.[136,1453,159,2082]" box="[136,1451,1685,1707]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44F91C28CA84E0" box="[136,1451,1685,1707]" pageId="3" pageNumber="234">- Leaves adaxial surface pubescent, rarely glabrescent. Pistillate flowers rotate ............................................................................. 33</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E7590090474FF44F93828CA84AE" pageId="3" pageNumber="234">
<paragraph id="8B8E36D090090474FF44F93828CB848C" blockId="3.[136,1453,159,2082]" box="[136,1450,1714,1736]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44F93828CB848C" box="[136,1450,1714,1736]" pageId="3" pageNumber="234">
32. Leaf blade narrowly elliptic to narrowly lanceolate. Inflorescences unisexual ......................................................
<taxonomicName id="4C314D5390090474FAC5F93828CB848C" ID-CoL="ZRCM" authorityName="Baillon" authorityYear="1864" box="[1289,1450,1714,1735]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="234" phylum="Tracheophyta" rank="species" species="uruguayensis">
<emphasis id="B945EAC290090474FAC5F93828CB848C" bold="true" box="[1289,1450,1714,1735]" italics="true" pageId="3" pageNumber="234">C. uruguayensis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8E36D090090474FF44F95A28CA84AE" blockId="3.[136,1453,159,2082]" box="[136,1451,1743,1765]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44F95A28CA84AE" box="[136,1451,1743,1765]" pageId="3" pageNumber="234">
- Leaf blade broadly ovate, broadly elliptic or rarely obovate. Inflorescences bisexual ...........................................
<taxonomicName id="4C314D5390090474FAC0F94528CA84AE" ID-CoL="ZQN6" box="[1292,1451,1743,1765]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="234" phylum="Tracheophyta" rank="species" species="muellerianus">
<emphasis id="B945EAC290090474FAC0F94528CA84AE" bold="true" box="[1292,1451,1743,1765]" italics="true" pageId="3" pageNumber="234">C. muellerianus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E7590090474FF44F96628CA8555" pageId="3" pageNumber="234">
<paragraph id="8B8E36D090090474FF44F96628CB854A" blockId="3.[136,1453,159,2082]" box="[136,1450,1771,1794]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44F96628CB854A" box="[136,1450,1771,1794]" pageId="3" pageNumber="234">
33. Abaxial leaf surface covered by stellate-lepidote trichomes; secondary veins conspicuously raised ...................
<taxonomicName id="4C314D5390090474FACDF96628CB854A" ID-CoL="ZPRY" authorityName="Baillon" authorityYear="1864" box="[1281,1450,1771,1793]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="234" phylum="Tracheophyta" rank="species" species="ceanothifolius">
<emphasis id="B945EAC290090474FACDF96628CB854A" bold="true" box="[1281,1450,1771,1793]" italics="true" pageId="3" pageNumber="234">C. ceanothifolius</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8E36D090090474FF44F88328CA8555" blockId="3.[136,1453,159,2082]" box="[136,1451,1801,1823]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44F88328CA8555" box="[136,1451,1801,1823]" pageId="3" pageNumber="234">- Abaxial leaf surface covered by lepidote subentire trichomes; secondary veins inconspicuous, not raised .................................. 34</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E7590090474FF44F8AC28CB85DA" pageId="3" pageNumber="234">
<paragraph id="8B8E36D090090474FF44F8AC28CB8513" blockId="3.[136,1453,159,2082]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44F8AC28CB8513" pageId="3" pageNumber="234">
34. Leaf blade narrowly elliptic to broadly elliptic. Pistillate sepals obovate to widely elliptic. Ovaries and fruits globose. Seeds brown without spots ................................................................................................................................................................
<taxonomicName id="4C314D5390090474FAE9F8C828CB8513" ID-CoL="ZR6B" authorityName="Martius ex Colla" authorityYear="1836" box="[1317,1450,1858,1880]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="234" phylum="Tracheophyta" rank="species" species="splendidus">
<emphasis id="B945EAC290090474FAE9F8C828CB8513" bold="true" box="[1317,1450,1858,1880]" italics="true" pageId="3" pageNumber="234">C. splendidus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8E36D090090474FF44F8EA28CB85DA" blockId="3.[136,1453,159,2082]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44F8EA28CB85DA" pageId="3" pageNumber="234">
- Leaf blade narrowly lanceolate to lanceolate. Pistillate sepals narrowly spatulate. Ovaries and fruits ellipsoid. Seeds gray with brown spots ................................................................................................................................................................
<taxonomicName id="4C314D5390090474FAD5F8F628CB85DA" ID-CoL="8WT3R" authorityName="Pereira, Caruzo &amp; Riina" authorityYear="2017" box="[1305,1450,1915,1937]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="234" phylum="Tracheophyta" rank="species" species="longicarpus">
<emphasis id="B945EAC290090474FAD5F8F628CB85DA" bold="true" box="[1305,1450,1915,1937]" italics="true" pageId="3" pageNumber="234">C. longicarpus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E7590090474FF44F81228CA8A4F" pageId="3" pageNumber="234">
<paragraph id="8B8E36D090090474FF44F81228CA8580" blockId="3.[136,1453,159,2082]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44F81228CA8580" pageId="3" pageNumber="234">35. Bracts with glands (colleters), viscid. Staminate flowers without cucullate sepals. Pistillate sepals equal, entire or slightly laciniate ........................................................................................................................................................................................... 36</keyLead>
</paragraph>
<paragraph id="8B8E36D090090474FF44F85928CA8A4F" blockId="3.[136,1453,159,2082]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44F85928CA8A4F" pageId="3" pageNumber="234">- Bracts without glands (colleters), not viscid. Staminate flowers with 3 cucullate sepals. Pistillate sepals unequal (3 conspicuous), pinnatifid ......................................................................................................................................................................................... 40</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E7590090473FF44F78628CA82FF" lastPageId="4" lastPageNumber="235" pageId="3" pageNumber="234">
<paragraph id="8B8E36D090090474FF44F78628CB8A6A" blockId="3.[136,1453,159,2082]" box="[136,1450,2059,2082]" pageId="3" pageNumber="234">
<keyLead id="30C095E590090474FF44F78628CB8A6A" box="[136,1450,2059,2082]" pageId="3" pageNumber="234">
36. Leaves with lanate indumentum, margin entire. Inflorescences densely congested .................................................
<taxonomicName id="4C314D5390090474FAD9F78628CB8A6A" ID-CoL="ZQ8C" authorityName="Baillon" authorityYear="1864" box="[1301,1450,2059,2081]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="234" phylum="Tracheophyta" rank="species" species="heterodoxus">
<emphasis id="B945EAC290090474FAD9F78628CB8A6A" bold="true" box="[1301,1450,2059,2081]" italics="true" pageId="3" pageNumber="234">C. heterodoxus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8E36D0900E0473FF44FF2A28CA82FF" blockId="4.[136,1452,159,555]" box="[136,1451,159,181]" pageId="4" pageNumber="235">
<keyLead id="30C095E5900E0473FF44FF2A28CA82FF" box="[136,1451,159,181]" pageId="4" pageNumber="235">- Leaves pubescent, margin entire to serrate. Inflorescences lax. ..................................................................................................... 37</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E75900E0473FF44FF3628CA82A5" pageId="4" pageNumber="235">
<paragraph id="8B8E36D0900E0473FF44FF3628CA829A" blockId="4.[136,1452,159,555]" box="[136,1451,187,210]" pageId="4" pageNumber="235">
<keyLead id="30C095E5900E0473FF44FF3628CA829A" box="[136,1451,187,210]" pageId="4" pageNumber="235">
37. Stipules with glands (colleters) along the margin .....................................................................................................
<taxonomicName id="4C314D53900E0473FADBFF3628CA829A" ID-CoL="ZR4L" authorityName="Baillon" authorityYear="1864" box="[1303,1451,187,209]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="4" pageNumber="235" phylum="Tracheophyta" rank="species" species="serratifolius">
<emphasis id="B945EAC2900E0473FADBFF3628CA829A" bold="true" box="[1303,1451,187,209]" italics="true" pageId="4" pageNumber="235">C. serratifolius</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8E36D0900E0473FF44FF5328CA82A5" blockId="4.[136,1452,159,555]" box="[136,1451,217,239]" pageId="4" pageNumber="235">
<keyLead id="30C095E5900E0473FF44FF5328CA82A5" box="[136,1451,217,239]" pageId="4" pageNumber="235">- Stipules with glands (colleters) only at the base ............................................................................................................................ 38</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E75900E0473FF44FF7C28CA830F" pageId="4" pageNumber="235">
<paragraph id="8B8E36D0900E0473FF44FF7C28CA8340" blockId="4.[136,1452,159,555]" box="[136,1451,245,267]" pageId="4" pageNumber="235">
<keyLead id="30C095E5900E0473FF44FF7C28CA8340" box="[136,1451,245,267]" pageId="4" pageNumber="235">
38. Leaves pubescent. Bracts lanceolate-rhombic. Pistillate sepals entirely covered by glands (colleters) .................
<taxonomicName id="4C314D53900E0473FAC4FF7F28CA8340" ID-CoL="6BJ79" authorityName="Muller Argoviensis" authorityYear="1865" box="[1288,1451,245,267]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="4" pageNumber="235" phylum="Tracheophyta" rank="species" species="chaetophorus">
<emphasis id="B945EAC2900E0473FAC4FF7F28CA8340" bold="true" box="[1288,1451,245,267]" italics="true" pageId="4" pageNumber="235">C. chaetophorus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8E36D0900E0473FF44FE9928CA830F" blockId="4.[136,1452,159,555]" pageId="4" pageNumber="235">
<keyLead id="30C095E5900E0473FF44FE9928CA830F" pageId="4" pageNumber="235">- Leaves glabrescent to sparsely pubescent. Bracts lanceolate. Pistillate sepals with glands (colleters) along the margin or absent ... ......................................................................................................................................................................................................... 39</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E75900E0473FF44FEC628CA83F3" pageId="4" pageNumber="235">
<paragraph id="8B8E36D0900E0473FF44FEC628CA8335" blockId="4.[136,1452,159,555]" pageId="4" pageNumber="235">
<keyLead id="30C095E5900E0473FF44FEC628CA8335" pageId="4" pageNumber="235">
39. Leaves rhombic, rarely obovate, margin irregularly serrate. Inflorescences bisexual. Pistillate sepals entire, glands (colleters) along the margin ...........................................................................................................................................................................
<taxonomicName id="4C314D53900E0473FA83FEE328CA8335" ID-CoL="ZQ4S" authorityName="Muller Argoviensis" authorityYear="1865" baseAuthorityName="Didr." box="[1359,1451,360,382]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="4" pageNumber="235" phylum="Tracheophyta" rank="species" species="fuscus">
<emphasis id="B945EAC2900E0473FA83FEE328CA8335" bold="true" box="[1359,1451,360,382]" italics="true" pageId="4" pageNumber="235">C. fuscus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8E36D0900E0473FF44FE0C28CA83F3" blockId="4.[136,1452,159,555]" pageId="4" pageNumber="235">
<keyLead id="30C095E5900E0473FF44FE0C28CA83F3" pageId="4" pageNumber="235">
- Leaves elliptic, rarely oblanceolate, margin entire. Inflorescences unisexual. Pistillate sepals slightly laciniate, without glands (colleters) .........................................................................................................................................................
<taxonomicName id="4C314D53900E0473FB17FE2828CA83F3" ID-CoL="ZPQL" authorityName="Allem" authorityYear="1979" box="[1243,1451,418,440]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="4" pageNumber="235" phylum="Tracheophyta" rank="species" species="calycireduplicatus">
<emphasis id="B945EAC2900E0473FB17FE2828CA83F3" bold="true" box="[1243,1451,418,440]" italics="true" pageId="4" pageNumber="235">C. calycireduplicatus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E75900E0473FF44FE3528CB8061" pageId="4" pageNumber="235">
<paragraph id="8B8E36D0900E0473FF44FE3528CB83BA" blockId="4.[136,1452,159,555]" pageId="4" pageNumber="235">
<keyLead id="30C095E5900E0473FF44FE3528CB83BA" pageId="4" pageNumber="235">
40. Subshrubs with xylopodium. Leaves with actinodromous venation. Bracts linear, entire. Plants from open vegetation and cerrado ....................................................................................................................................................................................
<taxonomicName id="4C314D53900E0473FAD5FE5628CB83BA" ID-CoL="ZPXY" authorityName="Webster" authorityYear="1992" box="[1305,1450,475,497]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="4" pageNumber="235" phylum="Tracheophyta" rank="species" species="didrichsenii">
<emphasis id="B945EAC2900E0473FAD5FE5628CB83BA" bold="true" box="[1305,1450,475,497]" italics="true" pageId="4" pageNumber="235">C. didrichsenii</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8E36D0900E0473FF44FE7328CB8061" blockId="4.[136,1452,159,555]" pageId="4" pageNumber="235">
<keyLead id="30C095E5900E0473FF44FE7328CB8061" pageId="4" pageNumber="235">
- Shrubs without xylopodium. Leaves with camptodromous venation. Bracts ovate, pinnatifid. Plants from edges of forest and disturbed areas .................................................................................................................................................................
<taxonomicName id="4C314D53900E0473FAF5FD9F28CB8061" ID-CoL="ZRBN" authorityName="Lamarck" authorityYear="1786" box="[1337,1450,533,554]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="4" pageNumber="235" phylum="Tracheophyta" rank="species" species="triqueter">
<emphasis id="B945EAC2900E0473FAF5FD9F28CB8061" bold="true" box="[1337,1450,533,554]" italics="true" pageId="4" pageNumber="235">C. triqueter</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>