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<document id="33E92D47B8C43C253E0B12F50263637C" ID-ISSN="1447-2554" ID-ZooBank="urn:lsid:zoobank.org:pub:131EFBFB-3A77-4C35-9FC0-4451DDAC4A0A" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1718994616601" checkinUser="felipe" docAuthor="Davey, Nicola" docDate="2013" docId="039887A4FE24FD58FF704FDBEC66FBBD" docLanguage="en" docName="MemMusVictoria.70.61-67.pdf" docOrigin="Memoirs of Museum Victoria 70" docStyle="DocumentStyle:5B3B5C052A93BFEEE334AD6097A94E45.2:MemMusVictoria.2003-.journal_article.type1" docStyleId="5B3B5C052A93BFEEE334AD6097A94E45" docStyleName="MemMusVictoria.2003-.journal_article.type1" docStyleVersion="2" docTitle="Psolidium ramum Davey, 2013, sp. nov." docType="treatment" docUuid="A9BB906B-094B-4F5E-83C5-5FFD1C68CC9B" docUuidSource="ZooBank" docVersion="1" lastPageNumber="66" masterDocId="FFA1FFDCFE2AFD57FFFC4A74EE47FF9F" masterDocTitle="The Psolidae of New Zealand and some additions to the Macquarie Ridge fauna (Echinodermata: Holothuroidea: Psolidae)" masterLastPageNumber="67" masterPageNumber="51" pageNumber="65" updateTime="1719008751508" updateUser="GgImagineBatch" zenodo-license-document="CC-BY-4.0">
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<mods:title id="A47A03BE87BF4F80CDD08B112BFCF34D">The Psolidae of New Zealand and some additions to the Macquarie Ridge fauna (Echinodermata: Holothuroidea: Psolidae)</mods:title>
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<mods:namePart id="C31653678ACDD57AC95D0DEBAEABDCFB">Davey, Nicola</mods:namePart>
<mods:affiliation id="E9532BD3F56E5A1D4BB397D0A85B22A8">National Institute of Water and Atmosphere Research Ltd (NIWA), PO Box 893, Nelson 7040, New Zealand (niki. davey @ niwa. co. nz)</mods:affiliation>
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<mods:title id="15B79BC34EDC1C0BABFA5DA831970579">Memoirs of Museum Victoria</mods:title>
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<mods:part id="062AA655D1447F257104FB099CE79564">
<mods:date id="0704461CDBB83A93F088642FBA3CE35C">2013</mods:date>
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<mods:number id="D2636F28E0354B8F2B64B0C120155F4C">2013-12-18</mods:number>
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<mods:number id="9C172903E4F247D25AEA4A116C140B50">70</mods:number>
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<mods:identifier id="1DBEE25BAA53898DE69BB9727B1DB7C6" type="ISSN">1447-2554</mods:identifier>
<mods:identifier id="CDB58A47A7E26C1881AFC88BEA51F8CB" type="ZooBank">urn:lsid:zoobank.org:pub:131EFBFB-3A77-4C35-9FC0-4451DDAC4A0A</mods:identifier>
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<treatment id="039887A4FE24FD58FF704FDBEC66FBBD" LSID="urn:lsid:zoobank.org:act:A9BB906B-094B-4F5E-83C5-5FFD1C68CC9B" httpUri="http://treatment.plazi.org/id/039887A4FE24FD58FF704FDBEC66FBBD" lastPageId="15" lastPageNumber="66" pageId="14" pageNumber="65">
<subSubSection id="C32B6539FE24FD59FF704FDBED45FA71" pageId="14" pageNumber="65" type="nomenclature">
<paragraph id="8B8E36B2FE24FD59FF704FDBEFDAFA5A" blockId="14.[140,413,1455,1477]" box="[140,413,1455,1477]" pageId="14" pageNumber="65">
<heading id="D0C681DEFE24FD59FF704FDBEFDAFA5A" box="[140,413,1455,1477]" fontSize="9" level="2" pageId="14" pageNumber="65" reason="5">
<taxonomicName id="4C314D31FE24FD59FF704FDBEF0DFA5A" ID-CoL="4PL6Y" authorityName="Davey" authorityYear="2013" box="[140,330,1455,1477]" class="Holothuroidea" family="Psolidae" genus="Psolidium" kingdom="Animalia" order="Dendrochirotida" pageId="14" pageNumber="65" phylum="Echinodermata" rank="species" species="ramum" status="sp. nov.">
<emphasis id="B945EAA0FE24FD59FF704FDBEF0DFA5A" bold="true" box="[140,330,1455,1477]" italics="true" pageId="14" pageNumber="65">Psolidium ramum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27657DBFE24FD59FEAC4FC4EFDAFA5A" box="[336,413,1456,1477]" pageId="14" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8B8E36B2FE24FD59FF4E4FAFED45FA71" blockId="14.[140,770,1499,1544]" box="[178,770,1499,1518]" pageId="14" pageNumber="65">http://zoobank.org/urn:lsid:zoobank.org:act:A9BB906B-094B-</paragraph>
</subSubSection>
<subSubSection id="C32B6539FE24FD58FF704F81EC66FBBD" lastPageId="15" lastPageNumber="66" pageId="14" pageNumber="65" type="description">
<paragraph id="8B8E36B2FE24FD59FF704F81EFDFF997" blockId="14.[140,770,1499,1544]" box="[140,408,1525,1544]" pageId="14" pageNumber="65">4F5E-83C5-5FFD1C68CC9B</paragraph>
<paragraph id="8B8E36B2FE24FD59FF704C54EFA3F9AA" blockId="14.[140,484,1568,1590]" box="[140,484,1568,1590]" pageId="14" pageNumber="65">
<tableCitation id="C6B30309FE24FD59FF704C54EE92F9AA" box="[140,213,1568,1590]" captionStart="Table 1" captionStartId="2.[140,189,1107,1126]" captionText="Table 1. Distribution of New Zealand and Macquarie Ridge species of Psolidae Burmeister, 1837." pageId="14" pageNumber="65">Table 1</tableCitation>
,
<figureCitation id="130A2A37FE24FD59FF1D4C54EF05F9AA" box="[225,322,1568,1590]" captionStart="Figure 2" captionStartId="3.[816,876,955,974]" captionText="Figure 2. Map of area showing station locations of Psolidium species in the New Zealand and Macquarie Ridge regions." pageId="14" pageNumber="65">Figures 2</figureCitation>
,
<figureCitation id="130A2A37FE24FD59FEB14C54EFCDF9A9" box="[333,394,1568,1590]" captionStart="Figure 6" captionStartId="7.[147,207,772,791]" captionTargetBox="[147,1447,227,755]" captionTargetId="figure-8@7.[147,1447,227,755]" captionTargetPageId="7" captionText="Figure 6. Psolidium ramum sp. nov. holotype (A–B, NIWA 73660): A, dorsal view; B, ventral view. Scale bar 1 cm." pageId="14" pageNumber="65">6A–B</figureCitation>
,
<figureCitation id="130A2A37FE24FD59FE684C54EF98F9AA" box="[404,479,1568,1589]" captionStart="Figure 11" captionStartId="14.[141,201,1312,1331]" captionTargetBox="[140,1440,227,1295]" captionTargetId="figure-273@14.[140,1440,224,1298]" captionTargetPageId="14" captionText="Figure 11.Psolidium ramum sp. nov. holotype (A–H, NIWA 73660): A–B, dorsal scales with tube feet perforations visible; C, thorn ossicle from dorsal body wall (also present in ventral body wall); D, curved tube foot support plate from dorsal body wall; E–H, knobbed plates with predominantly 4 even perforations." pageId="14" pageNumber="65">11A–H</figureCitation>
.
</paragraph>
<paragraph id="8B8E36B2FE24FD59FF704C3FEF7FF90C" blockId="14.[140,771,1611,1683]" pageId="14" pageNumber="65">
<emphasis id="B945EAA0FE24FD59FF704C3FEF05F9C1" box="[140,322,1611,1630]" italics="true" pageId="14" pageNumber="65">Material examined.</emphasis>
<materialsCitation id="3B593CEFFE24FD59FEBB4C3FEF7FF90C" collectingDate="2011-04-01" country="New Zealand" elevation="199" latitude="-36.18" location="North Island" longLatPrecision="713" longitude="173.68" municipality="Stn" pageId="14" pageNumber="65" specimenCount="1" typeStatus="holotype">
<typeStatus id="548A8810FE24FD59FEBB4C3FEFD9F9C1" box="[327,414,1611,1630]" pageId="14" pageNumber="65" type="holotype">Holotype</typeStatus>
.
<collectingCountry id="F3267622FE24FD59FE5B4C38EC64F9C1" box="[423,547,1611,1631]" name="New Zealand" pageId="14" pageNumber="65">New Zealand</collectingCountry>
,
<location id="8EEE6069FE24FD59FDD04C3FECD8F9C1" LSID="urn:lsid:plazi:treatment:039887A4FE24FD58FF704FDBEC66FBBD:8EEE6069FE24FD59FDD04C3FECD8F9C1" box="[556,671,1611,1630]" country="New Zealand" latitude="-36.18" longLatPrecision="713" longitude="173.68" municipality="Stn" name="North Island" pageId="14" pageNumber="65">North Island</location>
west coast canyons: NIWA 73660 (1)
<collectingMunicipality id="6BEAACC8FE24FD59FE784C12EFE5F9E6" box="[388,418,1638,1657]" pageId="14" pageNumber="65">Stn</collectingMunicipality>
TAN1105/88,
<geoCoordinate id="EE055075FE24FD59FDD64C12EC37F9E6" box="[554,624,1638,1657]" degrees="36.18" direction="south" orientation="latitude" pageId="14" pageNumber="65" precision="555" value="-36.18">36.18°S</geoCoordinate>
<geoCoordinate id="EE055075FE24FD59FD8B4C12EC8CF9E6" box="[631,715,1638,1657]" degrees="173.68" direction="east" orientation="longitude" pageId="14" pageNumber="65" precision="555" value="173.68">173.68°E</geoCoordinate>
,
<quantity id="4CC99B57FE24FD59FD2A4C12EE82F90C" metricMagnitude="2" metricUnit="m" metricValue="1.9899999999999998" metricValueMax="2.1" metricValueMin="1.88" pageId="14" pageNumber="65" unit="m" value="199.0" valueMax="210.0" valueMin="188.0">
<elevation id="001CD181FE24FD59FD2A4C12EE82F90C" metricMagnitude="2" metricUnit="m" metricValue="1.9899999999999998" metricValueMax="2.1" metricValueMin="1.88" pageId="14" pageNumber="65" unit="m" value="199.0" valueMax="210.0" valueMin="188.0">188– 210 m</elevation>
</quantity>
,
<date id="FF8F1072FE24FD59FF324CF4EF73F90C" box="[206,308,1664,1683]" pageId="14" pageNumber="65" value="2011-04-01">
<collectingDate id="EFCBE99AFE24FD59FF324CF4EF73F90C" box="[206,308,1664,1683]" pageId="14" pageNumber="65" value="2011-04-01">01/04/2011</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="8B8E36B2FE24FD59FF704CDFEA17F9C1" blockId="14.[140,771,1706,1903]" lastBlockId="14.[807,1441,1456,1903]" pageId="14" pageNumber="65">
<emphasis id="B945EAA0FE24FD59FF704CDFEF4BF95F" box="[140,268,1707,1728]" italics="true" pageId="14" pageNumber="65">Description.</emphasis>
<taxonomicName id="4C314D31FE24FD59FEED4CDEEF8FF95F" box="[273,456,1706,1728]" pageId="14" pageNumber="65">
<emphasis id="B945EAA0FE24FD59FEED4CDEEF3FF920" box="[273,376,1706,1727]" italics="true" pageId="14" pageNumber="65">Psolidium</emphasis>
species
</taxonomicName>
up to
<quantity id="4CC99B57FE24FD59FDFB4CDEEC0AF95F" box="[519,589,1706,1728]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="14" pageNumber="65" unit="mm" value="16.0">16 mm</quantity>
long,
<quantity id="4CC99B57FE24FD59FD774CDFEC81F95F" box="[651,710,1707,1728]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="14" pageNumber="65" unit="mm" value="8.0">8 mm</quantity>
wide,
<quantity id="4CC99B57FE24FD59FF704CBCEE8CF942" box="[140,203,1736,1757]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="65" unit="mm" value="2.0">2 mm</quantity>
high. Profile low, body form oval. Oral cone slightly higher than anal cone; no distinct oral or anal valves or plates, approximately 230 µm wide, macroscopically smooth, microscopically glassy and beady texture, lacking significant bumps or pillars. Body wall dorsal and lateral scales macroscopically evident and continuous over body wall, up to
<quantity id="4CC99B57FE24FD59FCD64FC4ED30FA5A" box="[810,887,1456,1477]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="14" pageNumber="65" unit="mm" value="2.3">2.3 mm</quantity>
at widest point. Tube feet numerous, up to 10 per scale, evident throughout dorsal and lateral scales. Tentacles 10; 8 + 2 (ventral smaller). Sole largely destroyed, with peripheral single row of larger tube feet and smaller outer non-continuous ring of tube feet; midventral row of tube feet present, only
<quantity id="4CC99B57FE24FD59FADB4C5EEB18F9DF" box="[1319,1375,1578,1600]" metricMagnitude="-1" metricUnit="m" metricValue="6.096" pageId="14" pageNumber="65" unit="ft" value="2.0">2 feet</quantity>
found as a result of a damaged sole.
</paragraph>
<paragraph id="8B8E36B2FE24FD59FCB34C13EBE6F969" blockId="14.[807,1441,1456,1903]" pageId="14" pageNumber="65">Dorsal and lateral ossicles include large multilayered thick scales with small perforations and conspicuous round tube feet holes up to 60 µm in diameter; numerous tube feet support plates up to 85µm wide; rare single-layered plates with 4–9 perforations up to 70 µm wide; broken thorn (branching rod) ossicles present.</paragraph>
<paragraph id="8B8E36B2FE24FD58FCB34C8BEC3AFE88" blockId="14.[807,1441,1456,1903]" lastBlockId="15.[147,778,229,336]" lastPageId="15" lastPageNumber="66" pageId="14" pageNumber="65">Ventral ossicles knobbed perforated plates up to 100 µm wide with blunt marginal projections, predominantly 4 projections, sometimes more projections peripherally, and thick elongate plates up to 160 µm long, without knobs, with small perforations; small crosses up to 60 µm wide; thorn ossicles present, mainly broken, largest 135 µm.</paragraph>
<paragraph id="8B8E36B2FE25FD58FF454B6AEF60FECF" blockId="15.[147,778,229,336]" pageId="15" pageNumber="66">Colour. Preserved: white, with dorsal and lateral scales, grey centrally.</paragraph>
<paragraph id="8B8E36B2FE25FD58FF6F4B11ED4CFEE4" blockId="15.[147,779,357,379]" box="[147,779,357,379]" pageId="15" pageNumber="66">
<emphasis id="B945EAA0FE25FD58FF6F4B11EF51FEE5" box="[147,278,357,378]" italics="true" pageId="15" pageNumber="66">Distribution.</emphasis>
<collectingCountry id="F3267622FE25FD58FEE64B12EFE7FEE4" box="[282,416,357,379]" name="New Zealand" pageId="15" pageNumber="66">New Zealand</collectingCountry>
, North Island west coast,
<quantity id="4CC99B57FE25FD58FD644B12ED41FEE4" box="[664,774,358,379]" metricMagnitude="2" metricUnit="m" metricValue="1.9899999999999998" metricValueMax="2.1" metricValueMin="1.88" pageId="15" pageNumber="66" unit="m" value="199.0" valueMax="210.0" valueMin="188.0">
188–
<specimenCount id="9D37FD3BFE25FD58FD3B4B12ED41FEE4" box="[711,774,358,379]" count="210" pageId="15" pageNumber="66" type="generic">210 m</specimenCount>
</quantity>
.
</paragraph>
<paragraph id="8B8E36B2FE25FD58FF6F4BE5EF1BFE40" blockId="15.[147,778,401,479]" pageId="15" pageNumber="66">
<emphasis id="B945EAA0FE25FD58FF6F4BE5EF40FE39" box="[147,263,401,422]" italics="true" pageId="15" pageNumber="66">Etymology.</emphasis>
The Latin word
<emphasis id="B945EAA0FE25FD58FE4C4BE5EFFFFE39" box="[432,440,401,422]" italics="true" pageId="15" pageNumber="66">‘</emphasis>
ramum’ = branching, in reference to the branch-like thorn ossicles in the dorsal and ventral body wall of this species.
</paragraph>
<paragraph id="8B8E36B2FE25FD58FF6F4B80ED43FCE6" blockId="15.[146,780,500,1058]" pageId="15" pageNumber="66">
<emphasis id="B945EAA0FE25FD58FF6F4B80EEB3FD96" box="[147,244,500,521]" italics="true" pageId="15" pageNumber="66">Remarks.</emphasis>
This description is based on
<specimenCount id="9D37FD3BFE25FD58FDE74B81ECE0FD95" box="[539,679,500,522]" count="1" pageId="15" pageNumber="66" type="generic">one specimen</specimenCount>
, of which the ventral sole was partially destroyed. The distinctive thorn ossicles, while rare, were present in both the dorsal and ventral body wall and have not been reported for any other
<collectingCountry id="F3267622FE25FD58FD27483DEEA2FDE4" name="New Zealand" pageId="15" pageNumber="66">New Zealand</collectingCountry>
<taxonomicName id="4C314D31FE25FD58FF124811EFE9FDE4" box="[238,430,613,635]" class="Holothuroidea" family="Psolidae" genus="Psolidium" kingdom="Animalia" order="Dendrochirotida" pageId="15" pageNumber="66" phylum="Echinodermata" rank="species" species="undetermined">
<emphasis id="B945EAA0FE25FD58FF124811EF12FDE5" box="[238,341,613,634]" italics="true" pageId="15" pageNumber="66">Psolidium</emphasis>
species.
</taxonomicName>
In
<taxonomicName id="4C314D31FE25FD58FE284811ECCCFDE4" authorityName="Davey" authorityYear="2013" box="[468,651,613,635]" class="Holothuroidea" family="Psolidae" genus="Psolidium" kingdom="Animalia" order="Dendrochirotida" pageId="15" pageNumber="66" phylum="Echinodermata" rank="species" species="ramum" status="sp. nov.">
<emphasis id="B945EAA0FE25FD58FE284811ECCCFDE4" box="[468,651,613,635]" italics="true" pageId="15" pageNumber="66">Psolidium ramum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27657DBFE25FD58FD684812ECA4FDE4" box="[660,739,614,635]" pageId="15" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
the ventral ossicles have predominantly 4 perforations; in
<taxonomicName id="4C314D31FE25FD58FD0A48F6EEB7FD2C" authorityName="Davey" authorityYear="2013" class="Holothuroidea" family="Psolidae" genus="Psolidium" kingdom="Animalia" order="Dendrochirotida" pageId="15" pageNumber="66" phylum="Echinodermata" rank="species" species="marriotti" status="sp. nov.">
<emphasis id="B945EAA0FE25FD58FD0A48F6EEB7FD2C" italics="true" pageId="15" pageNumber="66">P. marriotti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27657DBFE25FD58FF0648EAEF0DFD2C" box="[250,330,670,691]" pageId="15" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
they are numerous.
<taxonomicName id="4C314D31FE25FD58FDD848EAECC1FD2C" authorityName="Davey" authorityYear="2013" box="[548,646,670,691]" class="Holothuroidea" family="Psolidae" genus="Psolidium" kingdom="Animalia" order="Dendrochirotida" pageId="15" pageNumber="66" phylum="Echinodermata" rank="species" species="aequm" status="sp. nov.">
<emphasis id="B945EAA0FE25FD58FDD848EAECC1FD2C" box="[548,646,670,691]" italics="true" pageId="15" pageNumber="66">P. aequm</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27657DBFE25FD58FD6C48EAECA7FD2C" box="[656,736,670,691]" pageId="15" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
has larger perforations, thicker ossicles and many more angular knobs on ossicle margins. The thickened elongated plates are unique to
<taxonomicName id="4C314D31FE25FD58FF064887EF1AFC97" authorityName="Davey" authorityYear="2013" box="[250,349,755,776]" class="Holothuroidea" family="Psolidae" genus="Psolidium" kingdom="Animalia" order="Dendrochirotida" pageId="15" pageNumber="66" phylum="Echinodermata" rank="species" species="ramum" status="sp. nov.">
<emphasis id="B945EAA0FE25FD58FF064887EF1AFC97" box="[250,349,755,776]" italics="true" pageId="15" pageNumber="66">P. ramum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27657DBFE25FD58FE984886EFF5FC98" box="[356,434,754,775]" pageId="15" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
The northwest Australian species
<taxonomicName id="4C314D31FE25FD58FF6F497BEFE3FCBC" authority="(Sluiter, 1901)" baseAuthorityName="Sluiter" baseAuthorityYear="1901" box="[147,420,782,804]" class="Holothuroidea" family="Psolidae" genus="Psolidium" kingdom="Animalia" order="Dendrochirotida" pageId="15" pageNumber="66" phylum="Echinodermata" rank="species" species="parmatus">
<emphasis id="B945EAA0FE25FD58FF6F497BEF56FCBB" box="[147,273,783,804]" italics="true" pageId="15" pageNumber="66">P. parmatus</emphasis>
(
<bibRefCitation id="EFA04B43FE25FD58FEE2497AEFDAFCBC" author="Sluiter, C. P." box="[286,413,782,804]" pageId="15" pageNumber="66" pagination="1 - 142" refId="ref12542" refString="Sluiter, C. P. 1901. Die Holothurien der Siboga - Expedition. Siboga- Expeditie 44: 1 - 142, 10 pls. Buchhandlung und Druckerei vormals E. J. Brill: Leiden." type="journal article" year="1901">Sluiter, 1901</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C314D31FE25FD58FE2B497BEC9CFCBC" authority="Clark, 1938" authorityName="Clark" authorityYear="1938" box="[471,731,782,804]" class="Holothuroidea" family="Psolidae" genus="Psolidium" kingdom="Animalia" order="Dendrochirotida" pageId="15" pageNumber="66" phylum="Echinodermata" rank="species" species="nigrescens">
<emphasis id="B945EAA0FE25FD58FE2B497BEC1AFCBB" box="[471,605,783,804]" italics="true" pageId="15" pageNumber="66">P. nigrescens</emphasis>
<bibRefCitation id="EFA04B43FE25FD58FD98497AEC9CFCBC" author="Clark, H. L." box="[612,731,782,804]" pageId="15" pageNumber="66" pagination="1 - 596" refId="ref11356" refString="Clark, H. L. 1938. Echinoderms from Australia. An account of collections made in 1929 and 1932. Memoirs of the Museum of Comparative Zoology at Harvard College 55: 1 - 596, 28 pls, 63 figs." type="journal article" year="1938">Clark, 1938</bibRefCitation>
</taxonomicName>
also contain thorn ossicles similar to those of
<taxonomicName id="4C314D31FE25FD58FDD0495FECC9FCDF" authorityName="Davey" authorityYear="2013" box="[556,654,811,832]" class="Holothuroidea" family="Psolidae" genus="Psolidium" kingdom="Animalia" order="Dendrochirotida" pageId="15" pageNumber="66" phylum="Echinodermata" rank="species" species="ramum" status="sp. nov.">
<emphasis id="B945EAA0FE25FD58FDD0495FECC9FCDF" box="[556,654,811,832]" italics="true" pageId="15" pageNumber="66">P. ramum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27657DBFE25FD58FD68495FECA5FCDF" box="[660,738,811,832]" pageId="15" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
, but
<taxonomicName id="4C314D31FE25FD58FF6F4933EF49FCC3" baseAuthorityName="Sluiter" baseAuthorityYear="1901" box="[147,270,839,860]" class="Holothuroidea" family="Psolidae" genus="Psolidium" kingdom="Animalia" order="Dendrochirotida" pageId="15" pageNumber="66" phylum="Echinodermata" rank="species" species="parmatus">
<emphasis id="B945EAA0FE25FD58FF6F4933EF49FCC3" box="[147,270,839,860]" italics="true" pageId="15" pageNumber="66">P. parmatus</emphasis>
</taxonomicName>
has bulbous pillars on the dorsal and lateral scales, and
<taxonomicName id="4C314D31FE25FD58FF424917EF03FCE7" authorityName="Clark" authorityYear="1938" box="[190,324,867,888]" class="Holothuroidea" family="Psolidae" genus="Psolidium" kingdom="Animalia" order="Dendrochirotida" pageId="15" pageNumber="66" phylum="Echinodermata" rank="species" species="nigrescens">
<emphasis id="B945EAA0FE25FD58FF424917EF03FCE7" box="[190,324,867,888]" italics="true" pageId="15" pageNumber="66">P. nigrescens</emphasis>
</taxonomicName>
is black, and has cups and crosses ventrally.
</paragraph>
<paragraph id="8B8E36B2FE25FD58FF45490BEC66FBBD" blockId="15.[146,780,500,1058]" pageId="15" pageNumber="66">Further specimens would contribute to this description. The structure and distribution of ventral tube feet are difficult to determine, and there was little material available for ossicle extraction and SEM study. The tentacle ossicles could not be described due to the damaged state of the specimen, and these will need to be examined in the future.</paragraph>
</subSubSection>
</treatment>
</document>
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