treatments-xml/data/03/F0/96/03F09661FFFA8063FEC3E2CE7A3F50AF.xml

<document ID-DOI="10.1080/00222933.2022.2115950" ID-ISSN="1464-5262" ID-Zenodo-Dep="7156474" approvalRequired="9" approvalRequired_for_textStreams="9" checkinTime="1665136984522" checkinUser="plazi" docAuthor="Medeiros, Rone A. F., Seidel, Matthias &amp; Grossi, Paschoal C." docDate="2022" docId="03F09661FFFA8063FEC3E2CE7A3F50AF" docLanguage="en" docName="JNATHIST.56.29-32.1315-1364.pdf" docOrigin="Journal of Natural History (J. Nat. Hist.) 56 (29 - 32)" docSource="http://dx.doi.org/10.1080/00222933.2022.2115950" docStyle="DocumentStyle:3C762F9783B768A90048B82ACD135A62.3:JNatHist.2017-.journal_article.0cover" docStyleId="3C762F9783B768A90048B82ACD135A62" docStyleName="JNatHist.2017-.journal_article.0cover" docStyleVersion="3" docTitle="Byrsopolis castanea Burmeister 1844" docType="treatment" docVersion="2" lastPageNumber="1323" masterDocId="FFC9EE19FFFF806BFFD0E116782B521F" masterDocTitle="Revision of the genus Byrsopolis Burmeister, 1844 (Coleoptera: Melolonthidae: Rutelinae: Rutelini), with the description of six new species endemic to Brazil and Paraguay" masterLastPageNumber="1364" masterPageNumber="1315" pageNumber="1320" updateTime="1665137547684" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of the genus Byrsopolis Burmeister, 1844 (Coleoptera: Melolonthidae: Rutelinae: Rutelini), with the description of six new species endemic to Brazil and Paraguay</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Medeiros, Rone A. F.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Seidel, Matthias</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Grossi, Paschoal C.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="series">
<mods:title>J. Nat. Hist.</mods:title>
</mods:detail>
<mods:detail type="pubDate">
<mods:number>2022-09-22</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>56</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>29 - 32</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1315</mods:start>
<mods:end>1364</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.1080/00222933.2022.2115950</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1080/00222933.2022.2115950</mods:identifier>
<mods:identifier type="ISSN">1464-5262</mods:identifier>
<mods:identifier type="Zenodo-Dep">7156474</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7156484" ID-Zenodo-Dep="7156484" LSID="urn:lsid:plazi:treatment:03F09661FFFA8063FEC3E2CE7A3F50AF" httpUri="http://treatment.plazi.org/id/03F09661FFFA8063FEC3E2CE7A3F50AF" lastPageId="8" lastPageNumber="1323" pageId="5" pageNumber="1320">
<subSubSection box="[275,1040,984,1010]" pageId="5" pageNumber="1320" type="nomenclature">
<paragraph blockId="5.[275,1040,984,1010]" box="[275,1040,984,1010]" pageId="5" pageNumber="1320">
<heading bold="true" box="[275,1040,984,1010]" centered="true" fontSize="11" level="2" pageId="5" pageNumber="1320" reason="3">
<emphasis bold="true" box="[275,1040,984,1010]" pageId="5" pageNumber="1320">
<taxonomicName authority="Burmeister, 1844" authorityName="Burmeister" authorityYear="1844" box="[275,742,984,1010]" class="Insecta" family="Rutelidae" genus="Byrsopolis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="1320" phylum="Arthropoda" rank="species" species="castanea">
<emphasis bold="true" box="[275,519,984,1010]" italics="true" pageId="5" pageNumber="1320">Byrsopolis castanea</emphasis>
<bibRefCitation author="Burmeister HCC" box="[527,742,984,1010]" pageId="5" pageNumber="1320" refId="ref25824" refString="Burmeister HCC. 1844. Besondere Entomologie, Fortsetzung. Erste Abtheilung. Coleoptera Lamellicornia Anthobia et Phyllophaga systellochela. Handuch der Entomologie. Vierter Band. Berlin: T. C. F. Enslin; p. xii. 588." type="book" year="1844">Burmeister, 1844</bibRefCitation>
</taxonomicName>
(
<figureCitation box="[759,870,984,1010]" captionStart="Figure 1" captionStartId="6.[160,225,908,931]" captionTargetBox="[163,1153,143,874]" captionTargetId="figure-234@6.[162,1154,142,876]" captionTargetPageId="6" captionText="Figure 1. Byrsopolis castanea Burmeister dorsal habitus (a, c, e) and lateral habitus (b, d, f). Byrsopolis castanea lectotype male (a–b) and paralectotype male and female (c–f), box series label (g). Images from Karla Schneider (MLUH)." figureDoi="http://doi.org/10.5281/zenodo.7156478" httpUri="https://zenodo.org/record/7156478/files/figure.png" pageId="5" pageNumber="1320">Figures 1</figureCitation>
(a–g), 5(a–h))
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="1320" type="reference_group">
<paragraph blockId="5.[160,1156,1037,1409]" pageId="5" pageNumber="1320">
<treatmentCitationGroup pageId="5" pageNumber="1320">
<treatmentCitation author="Burmeister HCC" box="[160,642,1037,1062]" page="425" pageId="5" pageNumber="1320" year="1844">
<taxonomicName authority="Burmeister, 1844: 425" authorityName="Burmeister" authorityPageNumber="425" authorityYear="1844" box="[160,642,1037,1062]" class="Insecta" family="Rutelidae" genus="Byrsopolis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="1320" phylum="Arthropoda" rank="species" species="castanea">
<emphasis box="[160,378,1037,1062]" italics="true" pageId="5" pageNumber="1320">Byrsopolis castanea</emphasis>
<bibRefCitation author="Burmeister HCC" box="[390,642,1037,1062]" pageId="5" pageNumber="1320" refId="ref25824" refString="Burmeister HCC. 1844. Besondere Entomologie, Fortsetzung. Erste Abtheilung. Coleoptera Lamellicornia Anthobia et Phyllophaga systellochela. Handuch der Entomologie. Vierter Band. Berlin: T. C. F. Enslin; p. xii. 588." type="book" year="1844">Burmeister, 1844: 425</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description); 
<treatmentCitation author="Burmeister HCC" box="[909,1155,1037,1062]" page="523" pageId="5" pageNumber="1320" year="1855">
<bibRefCitation author="Burmeister HCC" box="[909,1155,1037,1062]" pageId="5" pageNumber="1320" pagination="570" refId="ref25867" refString="Burmeister HCC. 1855. Besondere Entomologie, Fortsetzung. Zweite Abtheilung. Coleoptera Lamellicornia Phyllophaga chaenochela. Handuch der Entomologie. Vierter Band. Berlin: T. C. F. Enslin; p. 570." type="book chapter" year="1855">Burmeister 1855: 523</bibRefCitation>
</treatmentCitation>
(female diagnosis); 
<treatmentCitation author="Lacordaire JT" box="[382,619,1072,1097]" page="364" pageId="5" pageNumber="1320" year="1856">
<bibRefCitation author="Lacordaire JT" box="[382,619,1072,1097]" pageId="5" pageNumber="1320" pagination="594" refId="ref26568" refString="Lacordaire JT. 1856. Histoire naturelle des insectes. Genera des Coleopteres ou expose methodique et critique de tous les genres proposes jusqu'ici dans cet ordre d'insectes. Tome troisieme contenant les familles des pectinicornes et lamellicornes. Paris: Librairie Encyclopedique de Roret; p. 594." type="book chapter" year="1856">Lacordaire 1856: 364</bibRefCitation>
</treatmentCitation>
(citation); 
<treatmentCitation author="Castelnau FL &amp; Lucas MH" box="[743,1098,1072,1097]" page="130" pageId="5" pageNumber="1320" year="1857">
<bibRefCitation author="Castelnau FL &amp; Lucas MH" box="[743,1098,1072,1097]" pageId="5" pageNumber="1320" pagination="473" refId="ref26028" refString="Castelnau FL, Lucas MH. 1857. Animaux nouveaux ou rares recueilles pendant l'Expedition dans les parties centrales de l'Amerique du Sud, de Rio de Janeiro a Lima et de Lima au Para sous la direction du Comte Francis de Castelnau. Chez P. Bertrand. Paris: Librareie-Editeur; p. 473." type="book chapter" year="1857">Castelnau and Lucas 1857: 130</bibRefCitation>
</treatmentCitation>
(distribution and illustration); 
<treatmentCitation author="Gemminger M &amp; Harold B" box="[471,886,1106,1131]" page="1226" pageId="5" pageNumber="1320" year="1869">
<bibRefCitation author="Gemminger M &amp; Harold B" box="[471,886,1106,1131]" pageId="5" pageNumber="1320" pagination="1240" refId="ref26271" refString="Gemminger M, Harold B. 1869. Catalogus Coleopterum: Scarabaeidae, Tom. IV. 4: 1240." type="journal article" year="1869">Gemminger and Harold 1869: 1226</bibRefCitation>
</treatmentCitation>
(checklist); 
<treatmentCitation author="Heyne A &amp; Taschenberg O" page="88" pageId="5" pageNumber="1320" year="1908">
<bibRefCitation author="Heyne A &amp; Taschenberg O" pageId="5" pageNumber="1320" pagination="39" refId="ref26360" refString="Heyne A, Taschenberg O. 1908. Die Exotischen Kafer in Wort und Bild. Leipzig. vii + 262: 39." type="journal article" year="1908">Heyne and Taschenberg 1908: 88</bibRefCitation>
</treatmentCitation>
(diagnosis), 356 (illustration); 
<treatmentCitation author="Ohaus F" box="[787,988,1141,1166]" page="251" pageId="5" pageNumber="1320" year="1908">
<bibRefCitation author="Ohaus F" box="[787,988,1141,1166]" pageId="5" pageNumber="1320" pagination="239 - 262" refId="ref26967" refString="Ohaus F. 1908. Die Ruteliden meiner Sammelreisen in Sudamerika. (Col.). Deutsche entomologische Zeitschrift. 1908: 239 - 262." type="journal article" year="1908">Ohaus 1908: 251</bibRefCitation>
</treatmentCitation>
(distribution); 1912: 312 (citation); 
<treatmentCitation author="Hardy A" box="[388,570,1176,1201]" page="236" pageId="5" pageNumber="1320" year="1971">
<bibRefCitation author="Hardy A" box="[388,570,1176,1201]" pageId="5" pageNumber="1320" pagination="235 - 242" refId="ref26328" refString="Hardy A. 1971. The North American Areodina with a description of a new genus from California. Pan- Pac Entomol. 47 (3): 235 - 242." type="journal article" year="1971">Hardy 1971: 236</bibRefCitation>
</treatmentCitation>
(citation); 
<treatmentCitation author="Machatschke JW" box="[690,924,1176,1201]" page="3" pageId="5" pageNumber="1320" year="1972">
<bibRefCitation author="Machatschke JW" box="[690,924,1176,1201]" pageId="5" pageNumber="1320" pagination="361" refId="ref26699" refString="Machatschke JW. 1972. Scarabaeoidea: Melolonthinae, Rutelinae. Coleopterorum Catalogus. Supplementa. 66: 361." type="journal article" year="1972">Machatschke 1972: 3</bibRefCitation>
</treatmentCitation>
(catalogue, distribution); 
<treatmentCitation author="Krajcik M" box="[223,395,1210,1235]" page="58" pageId="5" pageNumber="1320" year="2008">
<bibRefCitation author="Krajcik M" box="[223,395,1210,1235]" pageId="5" pageNumber="1320" pagination="142" refId="ref26488" refString="Krajcik M. 2008. Animma. X. Supplement 4. Checklist of the Scarabaeoidea of the World. 2. Rutelinae (Coleoptera: Scarabaeidae: Rutelinae). 2 nd ed. Plzen, Czech Republic: Milan Krajcik; p. 142." type="book chapter" year="2008">Krajcik 2008: 58</bibRefCitation>
</treatmentCitation>
(checklist); 
<treatmentCitation author="Grossi PC &amp; Vaz-de-Mello FZ" box="[523,903,1210,1235]" page="223" pageId="5" pageNumber="1320" year="2015">
<bibRefCitation author="Grossi PC &amp; Vaz-de-Mello FZ" box="[523,903,1210,1235]" pageId="5" pageNumber="1320" pagination="221 - 226" refId="ref26292" refString="Grossi PC, Vaz-de-Mello FZ. 2015. Moronius miguelangeli new genus and new species of Areodina from western Brazil (Melolonthidae, Rutelinae, Rutelini). Dugesiana. 22: 221 - 226." type="journal article" year="2015">Grossi and Vaz-de-Mello 2015: 223</bibRefCitation>
</treatmentCitation>
(biology, distribution); 
<treatmentCitation author="Ferreira ADS" box="[160,357,1245,1270]" page="112" pageId="5" pageNumber="1320" year="2016">
<bibRefCitation author="Ferreira ADS" box="[160,357,1245,1270]" pageId="5" pageNumber="1320" refId="ref26223" refString="Ferreira ADS. 2016. Besouros rutelineos: beleza escondida no Semiarido, 111 - 119. In: Bravo F, Calor AR, editors. Conhecendo os artropodes do Semiario. 1 st ed. Sao Paulo: Metis Producao Editorial; p. 192." type="journal volume" year="2016">Ferreira 2016: 112</bibRefCitation>
</treatmentCitation>
(illustration); 
<treatmentCitation author="Carvalho TG &amp; Grossi PC" box="[507,839,1245,1270]" page="372" pageId="5" pageNumber="1320" year="2018">
<bibRefCitation author="Carvalho TG &amp; Grossi PC" box="[507,839,1245,1270]" pageId="5" pageNumber="1320" pagination="369 - 372" refId="ref25906" refString="Carvalho TG, Grossi PC. 2018. Description of a second species of the genus Moronius Grossi &amp; Vaz-deMello, 2015 (Coleoptera: Scarabaeidae: Rutelinae: Rutelini). Zootaxa. 4434 (2): 369 - 372. doi: 10. 11646 / zootaxa. 4434.2.7." type="journal article" year="2018">Carvalho and Grossi 2018: 372</bibRefCitation>
</treatmentCitation>
(checklist, distribution); Vazde-Mello and Grossi 2022 (online catalogue).
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="5.[160,1156,1037,1409]" pageId="5" pageNumber="1320">
<taxonomicName authority="(Burmeister, 1844)" baseAuthorityName="Burmeister" baseAuthorityYear="1844" box="[192,732,1314,1339]" class="Insecta" family="Rutelidae" genus="Cotalpa" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="1320" phylum="Arthropoda" rank="species" species="castanea" subGenus="Byrsopolis">
<emphasis box="[192,277,1314,1339]" italics="true" pageId="5" pageNumber="1320">Cotalpa</emphasis>
(
<emphasis box="[294,402,1314,1339]" italics="true" pageId="5" pageNumber="1320">Byrsopolis</emphasis>
) 
<emphasis box="[418,516,1314,1339]" italics="true" pageId="5" pageNumber="1320">castanea</emphasis>
(
<bibRefCitation author="Burmeister HCC" box="[531,723,1314,1339]" pageId="5" pageNumber="1320" refId="ref25824" refString="Burmeister HCC. 1844. Besondere Entomologie, Fortsetzung. Erste Abtheilung. Coleoptera Lamellicornia Anthobia et Phyllophaga systellochela. Handuch der Entomologie. Vierter Band. Berlin: T. C. F. Enslin; p. xii. 588." type="book" year="1844">Burmeister, 1844</bibRefCitation>
)
</taxonomicName>
: 
<bibRefCitation author="Ohaus F" box="[745,936,1314,1339]" pageId="5" pageNumber="1320" pagination="256 - 260" refId="ref27018" refString="Ohaus F. 1915. XVI Beitrag zur Kenntnis der Ruteliden. (Col. lamell.). Deutsche entomologische Zeitschrift. Berlin Beiheft; p. 256 - 260." type="book chapter" year="1915">Ohaus 1915: 257</bibRefCitation>
(citation); 1918: 10 (catalogue, distribution); 1934: 37 (citation, checklist), 173 (illustration); 
<bibRefCitation author="Blackwelder RE" pageId="5" pageNumber="1320" pagination="189 - 306" refId="ref25704" refString="Blackwelder RE. 1944. Checklist of the Coleopterous insects of Mexico, Central America, The West Indies, and South America. Washington: Smithsonian Institution, United States National Museum. Bulletin 185, Part 2; p. 189 - 306." type="journal article" year="1944">Blackwelder 1944: 235</bibRefCitation>
(checklist).
</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="1321" pageId="5" pageNumber="1320" type="materials_examined">
<paragraph blockId="5.[160,745,1466,1492]" box="[160,745,1466,1492]" pageId="5" pageNumber="1320">
<emphasis bold="true" box="[160,317,1466,1492]" italics="true" pageId="5" pageNumber="1320">Type locality.</emphasis>
Nova Friburgo, 
<collectingRegion box="[507,668,1467,1492]" country="Brazil" name="Rio de Janeiro" pageId="5" pageNumber="1320">Rio de Janeiro</collectingRegion>
, 
<collectingCountry box="[679,742,1467,1492]" name="Brazil" pageId="5" pageNumber="1320">Brazil</collectingCountry>
.
</paragraph>
<paragraph blockId="5.[160,1156,1550,1610]" pageId="5" pageNumber="1320">
<emphasis bold="true" box="[160,328,1550,1576]" italics="true" pageId="5" pageNumber="1320">Type material.</emphasis>
<materialsCitation collectionCode="ZMHB" pageId="5" pageNumber="1320" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
<emphasis box="[344,689,1551,1576]" italics="true" pageId="5" pageNumber="1320">
<typeStatus box="[344,448,1551,1576]" pageId="5" pageNumber="1320">Lectotype</typeStatus>
male, here designated
</emphasis>
. (a) ‘ 
<taxonomicName authorityName="Burmeister" authorityYear="1844" box="[735,852,1551,1576]" class="Insecta" family="Rutelidae" genus="Byrsopolis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="1320" phylum="Arthropoda" rank="genus">Byrsopolis</taxonomicName>
*’, (b) ‘castanea *./Bras. Bsk. [handwritten by Burmeister]’ (
<specimenCount box="[500,531,1585,1610]" pageId="5" pageNumber="1320" type="male">1♂</specimenCount>
<collectionCode box="[539,609,1585,1610]" pageId="5" pageNumber="1320">ZMHB</collectionCode>
) (
<figureCitation box="[629,716,1585,1610]" captionStart="Figure 1" captionStartId="6.[160,225,908,931]" captionTargetBox="[163,1153,143,874]" captionTargetId="figure-234@6.[162,1154,142,876]" captionTargetPageId="6" captionText="Figure 1. Byrsopolis castanea Burmeister dorsal habitus (a, c, e) and lateral habitus (b, d, f). Byrsopolis castanea lectotype male (a–b) and paralectotype male and female (c–f), box series label (g). Images from Karla Schneider (MLUH)." figureDoi="http://doi.org/10.5281/zenodo.7156478" httpUri="https://zenodo.org/record/7156478/files/figure.png" pageId="5" pageNumber="1320">Figure 1</figureCitation>
(a,b)).
</materialsCitation>
</paragraph>
<paragraph blockId="5.[160,1157,1668,1728]" pageId="5" pageNumber="1320">
<emphasis bold="true" box="[160,341,1668,1694]" italics="true" pageId="5" pageNumber="1320">
<typeStatus box="[160,336,1668,1694]" pageId="5" pageNumber="1320">Paralectotypes</typeStatus>
.
</emphasis>
(a) ‘ 
<taxonomicName authorityName="Burmeister" authorityYear="1844" box="[395,512,1669,1694]" class="Insecta" family="Rutelidae" genus="Byrsopolis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="1320" phylum="Arthropoda" rank="genus">Byrsopolis</taxonomicName>
*’, (b) ‘castanea *./Bras. Bsk. 
<materialsCitation collectionCode="ZMHB" pageId="5" pageNumber="1320" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paralectotype">
[handwritten by Burmeister]’ (
<specimenCount box="[170,201,1703,1728]" pageId="5" pageNumber="1320" type="female">1♀</specimenCount>
, 
<specimenCount box="[207,240,1703,1728]" pageId="5" pageNumber="1320" type="male">1♂</specimenCount>
<collectionCode box="[248,318,1703,1728]" pageId="5" pageNumber="1320">ZMHB</collectionCode>
) (
<figureCitation box="[339,426,1703,1728]" captionStart="Figure 1" captionStartId="6.[160,225,908,931]" captionTargetBox="[163,1153,143,874]" captionTargetId="figure-234@6.[162,1154,142,876]" captionTargetPageId="6" captionText="Figure 1. Byrsopolis castanea Burmeister dorsal habitus (a, c, e) and lateral habitus (b, d, f). Byrsopolis castanea lectotype male (a–b) and paralectotype male and female (c–f), box series label (g). Images from Karla Schneider (MLUH)." figureDoi="http://doi.org/10.5281/zenodo.7156478" httpUri="https://zenodo.org/record/7156478/files/figure.png" pageId="5" pageNumber="1320">Figure 1</figureCitation>
(c–1))
</materialsCitation>
.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7156478" ID-Zenodo-Dep="7156478" httpUri="https://zenodo.org/record/7156478/files/figure.png" pageId="6" pageNumber="1321" startId="6.[160,225,908,931]" targetBox="[163,1153,143,874]" targetPageId="6">
<paragraph blockId="6.[160,1155,908,990]" pageId="6" pageNumber="1321">
<emphasis bold="true" box="[160,250,908,931]" pageId="6" pageNumber="1321">Figure 1.</emphasis>
<taxonomicName authority="Burmeister" authorityName="Burmeister" authorityYear="1844" box="[257,557,908,931]" class="Insecta" family="Rutelidae" genus="Byrsopolis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1321" phylum="Arthropoda" rank="species" species="castanea">
<emphasis box="[257,444,908,931]" italics="true" pageId="6" pageNumber="1321">Byrsopolis castanea</emphasis>
Burmeister
</taxonomicName>
dorsal habitus (a, c, e) and lateral habitus (b, d, f). 
<taxonomicName authorityName="Burmeister" authorityYear="1844" class="Insecta" family="Rutelidae" genus="Byrsopolis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1321" phylum="Arthropoda" rank="species" species="castanea">
<emphasis italics="true" pageId="6" pageNumber="1321">Byrsopolis castanea</emphasis>
</taxonomicName>
lectotype male (a–b) and paralectotype male and female (c–f), box series label (g). Images from Karla Schneider (MLUH).
</paragraph>
</caption>
<paragraph blockId="6.[160,1156,1075,1552]" pageId="6" pageNumber="1321">
<emphasis bold="true" box="[160,398,1075,1101]" italics="true" pageId="6" pageNumber="1321">Additional material.</emphasis>
[
<specimenCount box="[420,579,1076,1101]" pageId="6" pageNumber="1321" type="generic">22 specimens</specimenCount>
] 
<emphasis box="[592,820,1076,1101]" italics="true" pageId="6" pageNumber="1321">
<collectingCountry box="[592,653,1076,1101]" name="Brazil" pageId="6" pageNumber="1321">Brazil</collectingCountry>
, 
<collectingRegion box="[665,820,1076,1101]" country="Brazil" name="Rio de Janeiro" pageId="6" pageNumber="1321">Rio de Janeiro</collectingRegion>
</emphasis>
. ‘ 
<collectingCountry box="[837,902,1076,1101]" name="Brazil" pageId="6" pageNumber="1321">Brasil</collectingCountry>
, 
<collectingRegion box="[913,1077,1076,1101]" country="Brazil" name="Rio de Janeiro" pageId="6" pageNumber="1321">Rio de Janeiro</collectingRegion>
, Nova/ Friburgo, Macaé de Cima,/ 
<date box="[464,530,1111,1136]" pageId="6" pageNumber="1321" value="2004-01">i.2004</date>
, 
<quantity box="[542,630,1111,1136]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="1321" unit="m" value="1500.0">1500 m</quantity>
, luz, D. &amp; I./Miller legs’ (
<specimenCount box="[910,941,1111,1136]" pageId="6" pageNumber="1321" type="female">1♀</specimenCount>
, 
<specimenCount box="[955,989,1111,1136]" pageId="6" pageNumber="1321" type="male">7♂</specimenCount>
<collectionCode box="[998,1074,1111,1136]" pageId="6" pageNumber="1321">CERPE</collectionCode>
); same, except ‘Conquista,/ 
<quantity box="[373,445,1145,1170]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="6" pageNumber="1321" unit="m" value="850.0">850 m</quantity>
– 
<date box="[473,564,1145,1170]" pageId="6" pageNumber="1321" value="2004-12">XII.2004</date>
/E. &amp; P. Grossi Leg’. // ‘COLEÇÃO E./ &amp; P. GROSSI’ (
<specimenCount box="[1118,1150,1145,1170]" pageId="6" pageNumber="1321" type="female">1♀</specimenCount>
, 
<specimenCount box="[160,194,1180,1205]" pageId="6" pageNumber="1321" type="male">2♂</specimenCount>
<collectionCode box="[199,264,1180,1205]" pageId="6" pageNumber="1321">EPGC</collectionCode>
); ‘ 
<collectingCountry box="[280,363,1180,1205]" name="Brazil" pageId="6" pageNumber="1321">BRASIL</collectingCountry>
, RJ, Teresópolis, 
<quantity box="[546,614,1180,1205]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="6" pageNumber="1321" unit="m" value="600.0">600 m</quantity>
/ 
<date box="[624,794,1180,1205]" pageId="6" pageNumber="1321" value="2006-12-01" valueMax="2006-12-31" valueMin="2006-12-01">01 a 31.xii.2006</date>
/E. &amp; P. Grossi Legs’. // ‘COLEÇÃO E. &amp; P. GROSSI’ (
<specimenCount box="[350,381,1215,1240]" pageId="6" pageNumber="1321" type="male">1♂</specimenCount>
<collectionCode box="[390,455,1215,1240]" pageId="6" pageNumber="1321">EPGC</collectionCode>
); ‘Brasília/Cantagallo’ // ‘castanea Brm’ // ‘ 
<taxonomicName authorityName="Burmeister" authorityYear="1844" box="[931,1046,1215,1240]" class="Insecta" family="Rutelidae" genus="Byrsopolis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1321" phylum="Arthropoda" rank="genus">Byrsopolis</taxonomicName>
/ 
<taxonomicName authority="Burmeister" authorityName="Burmeister" authorityYear="1844" class="Insecta" family="Rutelidae" genus="Byrsopolis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1321" phylum="Arthropoda" rank="species" species="castanea">castanea Burmeister</taxonomicName>
/Museum f. Naturkunde/Berlin’ (
<specimenCount box="[676,711,1249,1274]" pageId="6" pageNumber="1321" type="male">2♂</specimenCount>
<collectionCode box="[726,800,1249,1274]" pageId="6" pageNumber="1321">ZMHB</collectionCode>
). 
<emphasis box="[823,973,1249,1274]" italics="true" pageId="6" pageNumber="1321">
<collectingRegion box="[823,973,1249,1274]" country="Brazil" name="Minas Gerais" pageId="6" pageNumber="1321">Minas Gerais</collectingRegion>
</emphasis>
. ‘BR, MG, Alto Caparaó/PARNA Caparaó/Coleta noturna/ 
<date box="[630,785,1284,1309]" pageId="6" pageNumber="1321" value="2016-12-05" valueMax="2016-12-06" valueMin="2016-12-05">05-06.xii.2016</date>
/A. Orsetti &amp; S. Alóquio legs’ (
<specimenCount box="[1124,1155,1284,1309]" pageId="6" pageNumber="1321" type="male">1♂</specimenCount>
<collectionCode box="[160,236,1319,1344]" pageId="6" pageNumber="1321">CERPE</collectionCode>
). 
<emphasis box="[249,397,1319,1344]" italics="true" pageId="6" pageNumber="1321">
<collectingRegion box="[249,397,1319,1344]" country="Brazil" name="Espirito Santo" pageId="6" pageNumber="1321">Espírito Santo</collectingRegion>
</emphasis>
. ‘ 
<collectingCountry box="[410,493,1319,1344]" name="Brazil" pageId="6" pageNumber="1321">BRASIL</collectingCountry>
: 
<collectingRegion box="[502,658,1319,1344]" country="Brazil" name="Espirito Santo" pageId="6" pageNumber="1321">Espírito Santo</collectingRegion>
. Conceição do Castelo./Ribeirão do Meio. 20° 18’/31’S: 
<geoCoordinate box="[265,395,1353,1378]" degrees="41" direction="west" minutes="14" orientation="longitude" pageId="6" pageNumber="1321" precision="15" seconds="17" value="-41.238056">
41°14 
<emphasis bold="true" box="[328,334,1353,1377]" pageId="6" pageNumber="1321">ʹ</emphasis>
17”W
</geoCoordinate>
. 
<quantity box="[410,483,1353,1378]" metricMagnitude="2" metricUnit="m" metricValue="8.8" pageId="6" pageNumber="1321" unit="m" value="880.0">880 m</quantity>
/Light. 
<date box="[567,642,1353,1378]" pageId="6" pageNumber="1321" value="2011-02">ii.2011</date>
. R. Falqueto’ (
<specimenCount box="[813,844,1353,1378]" pageId="6" pageNumber="1321" type="male">1♂</specimenCount>
<collectionCode box="[856,924,1353,1378]" pageId="6" pageNumber="1321">CEMT</collectionCode>
). 
<emphasis box="[943,1149,1353,1378]" italics="true" pageId="6" pageNumber="1321">No collection data</emphasis>
. ‘ 
<taxonomicName authorityName="Burmeister" authorityYear="1844" box="[162,280,1388,1413]" class="Insecta" family="Rutelidae" genus="Byrsopolis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1321" phylum="Arthropoda" rank="genus">Byrsopolis</taxonomicName>
/castanea Burm’ (
<specimenCount box="[497,529,1388,1413]" pageId="6" pageNumber="1321" type="female">1♀</specimenCount>
<collectionCode box="[542,616,1388,1413]" pageId="6" pageNumber="1321">ZMHB</collectionCode>
); ‘ 
<specimenCount box="[643,663,1388,1411]" pageId="6" pageNumber="1321" type="male">♂</specimenCount>
’ // ‘11,445’ // ‘Hist. – Coll. (
<taxonomicName authorityName="Linnaeus" authorityYear="1758" box="[1009,1141,1388,1413]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1321" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
)/ Nr.11445/ 
<taxonomicName authority="Burm." authorityName="Burmeister" authorityYear="1844" box="[266,565,1423,1448]" class="Insecta" family="Rutelidae" genus="Byrsopolis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1321" phylum="Arthropoda" rank="species" species="castanea">Byrsopolis castanea Burm.</taxonomicName>
/Brasília, Riehl’ (
<specimenCount box="[748,782,1423,1448]" pageId="6" pageNumber="1321" type="male">2♂</specimenCount>
<collectionCode box="[793,866,1423,1448]" pageId="6" pageNumber="1321">ZMHB</collectionCode>
); ‘43,114’ // ‘ 
<specimenCount box="[1013,1033,1423,1446]" pageId="6" pageNumber="1321" type="female">♀</specimenCount>
’ // ‘Hist. – Coll. (
<taxonomicName authorityName="Linnaeus" authorityYear="1758" box="[226,356,1457,1482]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1321" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
)/Nr. 43,114/ 
<taxonomicName authority="Burm." authorityName="Burmeister" authorityYear="1844" box="[494,791,1457,1482]" class="Insecta" family="Rutelidae" genus="Byrsopolis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1321" phylum="Arthropoda" rank="species" species="castanea">Byrsopolis castanea Burm.</taxonomicName>
/Brasília, Coll. Schaum’ (
<specimenCount box="[1073,1105,1457,1482]" pageId="6" pageNumber="1321" type="female">1♀</specimenCount>
, 
<specimenCount box="[1121,1155,1457,1482]" pageId="6" pageNumber="1321" type="male">1♂</specimenCount>
<collectionCode box="[160,233,1492,1517]" pageId="6" pageNumber="1321">ZMHB</collectionCode>
); ‘ 
<taxonomicName authority="Burm." authorityName="Burmeister" authorityYear="1844" box="[255,437,1492,1517]" class="Insecta" family="Rutelidae" genus="Byrsopolis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1321" phylum="Arthropoda" rank="species" species="castanea">Castanea Burm.</taxonomicName>
Brésil’ // ‘Coll. Höhne’ // ‘ 
<taxonomicName authorityName="Burmeister" authorityYear="1844" box="[741,856,1492,1517]" class="Insecta" family="Rutelidae" genus="Byrsopolis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1321" phylum="Arthropoda" rank="genus">Byrsopolis</taxonomicName>
/castanea Burmeister’ (
<specimenCount box="[1124,1155,1492,1517]" pageId="6" pageNumber="1321" type="male">1♂</specimenCount>
<collectionCode box="[160,233,1527,1552]" pageId="6" pageNumber="1321">ZMHB</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="1322" pageId="6" pageNumber="1321" type="diagnosis">
<paragraph blockId="6.[160,1156,1598,1728]" lastBlockId="7.[160,1156,142,237]" lastPageId="7" lastPageNumber="1322" pageId="6" pageNumber="1321">
<emphasis bold="true" box="[160,282,1598,1624]" italics="true" pageId="6" pageNumber="1321">Diagnosis.</emphasis>
Body brown, reddish-brown to dark brown with metallic green reflections; clypeus subrectangular, anterior margin weakly emarginate; antennal club about 1.5x (female) to 2x (male) longer than antennomeres II–VII combined; scutellar plate with moderately rounded base; elytra with very distinct striae, densely punctate on the humeri; elytral apex truncated; protibial spur about 8x longer than wide; aedeagus with asymmetrical parameres, weakly to moderately divergent, weakly angulate laterally, apex of parameres weakly rounded.
</paragraph>
</subSubSection>
<subSubSection lastPageId="8" lastPageNumber="1323" pageId="7" pageNumber="1322" type="description">
<paragraph blockId="7.[160,1156,280,1346]" pageId="7" pageNumber="1322">
<emphasis bold="true" box="[160,503,280,306]" italics="true" pageId="7" pageNumber="1322">
Description of the 
<typeStatus box="[387,497,280,306]" pageId="7" pageNumber="1322">lectotype</typeStatus>
.
</emphasis>
Male. 
<emphasis box="[588,642,281,306]" italics="true" pageId="7" pageNumber="1322">Body</emphasis>
. Shape elongated, oval, with rounded sides; brown, reddish-brown to black colouration (
<figureCitation box="[671,760,316,341]" captionStart="Figure 1" captionStartId="6.[160,225,908,931]" captionTargetBox="[163,1153,143,874]" captionTargetId="figure-234@6.[162,1154,142,876]" captionTargetPageId="6" captionText="Figure 1. Byrsopolis castanea Burmeister dorsal habitus (a, c, e) and lateral habitus (b, d, f). Byrsopolis castanea lectotype male (a–b) and paralectotype male and female (c–f), box series label (g). Images from Karla Schneider (MLUH)." figureDoi="http://doi.org/10.5281/zenodo.7156478" httpUri="https://zenodo.org/record/7156478/files/figure.png" pageId="7" pageNumber="1322">Figure 1</figureCitation>
(a–b)); frontoclypeal suture darker than clypeus and frons; body with marginal areas from dark brown to black; metallic green and reddish reflections. Length 
<quantity box="[590,778,385,410]" metricMagnitude="-2" metricUnit="m" metricValue="2.748" metricValueMax="2.907" metricValueMin="2.589" pageId="7" pageNumber="1322" unit="mm" value="27.48" valueMax="29.07" valueMin="25.89">25.89–29.07 mm</quantity>
. Humeral width 
<quantity box="[964,1152,385,410]" metricMagnitude="-2" metricUnit="m" metricValue="1.392" metricValueMax="1.502" metricValueMin="1.282" pageId="7" pageNumber="1322" unit="mm" value="13.92" valueMax="15.02" valueMin="12.82">12.82–15.02 mm</quantity>
. Elytral width 
<quantity box="[310,499,420,445]" metricMagnitude="-2" metricUnit="m" metricValue="1.5785" metricValueMax="1.654" metricValueMin="1.503" pageId="7" pageNumber="1322" unit="mm" value="15.785" valueMax="16.54" valueMin="15.03">15.03–16.54 mm</quantity>
. 
<emphasis box="[510,567,420,445]" italics="true" pageId="7" pageNumber="1322">Head</emphasis>
. Clypeus about 2x wider than long, subrectangular, with angles weakly rounded, anterior margin weakly emarginate, surface weakly concave near the apex; frontoclypeal suture weakly bisinuate; surface strongly rugose; punctures small to moderately large, glabrous; frons as long as clypeus, punctures small to moderately large; moderately to densely punctate; glabrous; mandibles moderately sinuous at the border, scissorial region C-shaped (
<figureCitation box="[629,728,593,618]" captionStart="Figure 5" captionStartId="16.[160,225,913,936]" captionTargetBox="[163,1150,145,880]" captionTargetId="figure-176@16.[162,1154,142,882]" captionTargetPageId="16" captionText="Figure 5. Byrsopolis castanea Burmeister, Rio de Janeiro,Nova Friburgo. Male dorsal habitus (a); caudal view of aedeagus (b); female dorsal and lateral habitus (c–d); dorsal view of male left mandible (e); ventral view of left maxilla (f); ventral view of mentum (g); and dorsal view of labrum (h). Scale bars: a, c, d = 5 mm; b, e–h = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7156490" httpUri="https://zenodo.org/record/7156490/files/figure.png" pageId="7" pageNumber="1322">Figure 5</figureCitation>
(e)), rounded apex, small punctures, sparsely to densely punctate in the transverse median region; setae small to moderate in size near scissorial region, sparse to moderately distributed; lacinia with three teeth, equidistant; apical tooth longer and wider than basal one, basal tooth longer and wider than medium one (
<figureCitation box="[389,486,732,757]" captionStart="Figure 5" captionStartId="16.[160,225,913,936]" captionTargetBox="[163,1150,145,880]" captionTargetId="figure-176@16.[162,1154,142,882]" captionTargetPageId="16" captionText="Figure 5. Byrsopolis castanea Burmeister, Rio de Janeiro,Nova Friburgo. Male dorsal habitus (a); caudal view of aedeagus (b); female dorsal and lateral habitus (c–d); dorsal view of male left mandible (e); ventral view of left maxilla (f); ventral view of mentum (g); and dorsal view of labrum (h). Scale bars: a, c, d = 5 mm; b, e–h = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7156490" httpUri="https://zenodo.org/record/7156490/files/figure.png" pageId="7" pageNumber="1322">Figure 5</figureCitation>
(f)); basal tooth with long setae; last maxillary palpomere subrectangular sensorial area; premental apex moderately emarginated (
<figureCitation box="[975,1068,767,792]" captionStart="Figure 5" captionStartId="16.[160,225,913,936]" captionTargetBox="[163,1150,145,880]" captionTargetId="figure-176@16.[162,1154,142,882]" captionTargetPageId="16" captionText="Figure 5. Byrsopolis castanea Burmeister, Rio de Janeiro,Nova Friburgo. Male dorsal habitus (a); caudal view of aedeagus (b); female dorsal and lateral habitus (c–d); dorsal view of male left mandible (e); ventral view of left maxilla (f); ventral view of mentum (g); and dorsal view of labrum (h). Scale bars: a, c, d = 5 mm; b, e–h = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7156490" httpUri="https://zenodo.org/record/7156490/files/figure.png" pageId="7" pageNumber="1322">Figure 5</figureCitation>
(g)); last labial palpomere about 1.5x longer than palpomere II (
<figureCitation box="[792,887,801,826]" captionStart="Figure 5" captionStartId="16.[160,225,913,936]" captionTargetBox="[163,1150,145,880]" captionTargetId="figure-176@16.[162,1154,142,882]" captionTargetPageId="16" captionText="Figure 5. Byrsopolis castanea Burmeister, Rio de Janeiro,Nova Friburgo. Male dorsal habitus (a); caudal view of aedeagus (b); female dorsal and lateral habitus (c–d); dorsal view of male left mandible (e); ventral view of left maxilla (f); ventral view of mentum (g); and dorsal view of labrum (h). Scale bars: a, c, d = 5 mm; b, e–h = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7156490" httpUri="https://zenodo.org/record/7156490/files/figure.png" pageId="7" pageNumber="1322">Figure 5</figureCitation>
(g)); labrum moderately emarginate, moderately setose (
<figureCitation box="[536,635,836,861]" captionStart="Figure 5" captionStartId="16.[160,225,913,936]" captionTargetBox="[163,1150,145,880]" captionTargetId="figure-176@16.[162,1154,142,882]" captionTargetPageId="16" captionText="Figure 5. Byrsopolis castanea Burmeister, Rio de Janeiro,Nova Friburgo. Male dorsal habitus (a); caudal view of aedeagus (b); female dorsal and lateral habitus (c–d); dorsal view of male left mandible (e); ventral view of left maxilla (f); ventral view of mentum (g); and dorsal view of labrum (h). Scale bars: a, c, d = 5 mm; b, e–h = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7156490" httpUri="https://zenodo.org/record/7156490/files/figure.png" pageId="7" pageNumber="1322">Figure 5</figureCitation>
(h)); antennomere III subcylindrical, antennomeres IV and V subquadrate, antennomeres VI and VII subconical; antennal club about 2x longer than antennomeres II–VII combined. 
<emphasis box="[696,771,905,930]" italics="true" pageId="7" pageNumber="1322">Thorax</emphasis>
. Pronotum transverse, about 1.5x wider than long (
<figureCitation box="[351,443,940,965]" captionStart="Figure 5" captionStartId="16.[160,225,913,936]" captionTargetBox="[163,1150,145,880]" captionTargetId="figure-176@16.[162,1154,142,882]" captionTargetPageId="16" captionText="Figure 5. Byrsopolis castanea Burmeister, Rio de Janeiro,Nova Friburgo. Male dorsal habitus (a); caudal view of aedeagus (b); female dorsal and lateral habitus (c–d); dorsal view of male left mandible (e); ventral view of left maxilla (f); ventral view of mentum (g); and dorsal view of labrum (h). Scale bars: a, c, d = 5 mm; b, e–h = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7156490" httpUri="https://zenodo.org/record/7156490/files/figure.png" pageId="7" pageNumber="1322">Figure 5</figureCitation>
(a)); punctures small to moderately large, moderately to densely punctate, sparsely punctate at marginal areas, glabrous, two elongate oblique concavities at base, one on each side; scutellar plate subtriangular, sides straight, base moderately rounded; apex with punctures and setae small to moderate, disc moderately to densely punctate, marginal areas sparsely to moderately punctate. 
<emphasis box="[812,873,1079,1104]" italics="true" pageId="7" pageNumber="1322">Elytra</emphasis>
. About 1.25x longer than wide; surface glabrous, strongly striate and punctate, punctures small to large, moderately to densely punctate (
<figureCitation box="[415,512,1148,1173]" captionStart="Figure 5" captionStartId="16.[160,225,913,936]" captionTargetBox="[163,1150,145,880]" captionTargetId="figure-176@16.[162,1154,142,882]" captionTargetPageId="16" captionText="Figure 5. Byrsopolis castanea Burmeister, Rio de Janeiro,Nova Friburgo. Male dorsal habitus (a); caudal view of aedeagus (b); female dorsal and lateral habitus (c–d); dorsal view of male left mandible (e); ventral view of left maxilla (f); ventral view of mentum (g); and dorsal view of labrum (h). Scale bars: a, c, d = 5 mm; b, e–h = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7156490" httpUri="https://zenodo.org/record/7156490/files/figure.png" pageId="7" pageNumber="1322">Figure 5</figureCitation>
(a)); interstriae with strong coalescent punctures; apex truncate; glabrous. 
<emphasis box="[380,428,1183,1208]" italics="true" pageId="7" pageNumber="1322">Legs</emphasis>
. Protibial spur about 8x longer than wide; mesotibiae with one carina, metatibiae with two carinae. 
<emphasis box="[575,680,1217,1242]" italics="true" pageId="7" pageNumber="1322">Aedeagus</emphasis>
. Parameres with proximal margin moderately rounded; lateral margins excavated, forming a carina from near base to apex; apexes rounded and weakly sinuate internally, weakly divergent, dorsally symmetrical, laterally and ventrally asymmetrical; left paramere longer than righr paramere (
<figureCitation box="[956,1050,1321,1346]" captionStart="Figure 5" captionStartId="16.[160,225,913,936]" captionTargetBox="[163,1150,145,880]" captionTargetId="figure-176@16.[162,1154,142,882]" captionTargetPageId="16" captionText="Figure 5. Byrsopolis castanea Burmeister, Rio de Janeiro,Nova Friburgo. Male dorsal habitus (a); caudal view of aedeagus (b); female dorsal and lateral habitus (c–d); dorsal view of male left mandible (e); ventral view of left maxilla (f); ventral view of mentum (g); and dorsal view of labrum (h). Scale bars: a, c, d = 5 mm; b, e–h = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7156490" httpUri="https://zenodo.org/record/7156490/files/figure.png" pageId="7" pageNumber="1322">Figure 5</figureCitation>
(b)).
</paragraph>
<paragraph blockId="7.[160,897,1390,1416]" box="[160,897,1390,1416]" pageId="7" pageNumber="1322">
<emphasis bold="true" box="[160,253,1390,1416]" italics="true" pageId="7" pageNumber="1322">Female.</emphasis>
Similar to male, varying in the aspects described below.
</paragraph>
<paragraph blockId="7.[160,1156,1460,1728]" pageId="7" pageNumber="1322">
<emphasis bold="true" box="[160,225,1460,1486]" italics="true" pageId="7" pageNumber="1322">Body.</emphasis>
Length 
<quantity box="[332,524,1461,1486]" metricMagnitude="-2" metricUnit="m" metricValue="3.3755" metricValueMax="3.781" metricValueMin="2.9699999999999998" pageId="7" pageNumber="1322" unit="mm" value="33.755" valueMax="37.81" valueMin="29.7">29.70–37.81 mm</quantity>
. Humeral width 
<quantity box="[720,912,1461,1486]" metricMagnitude="-2" metricUnit="m" metricValue="1.9759999999999998" metricValueMax="2.073" metricValueMin="1.879" pageId="7" pageNumber="1322" unit="mm" value="19.759999999999998" valueMax="20.73" valueMin="18.79">18.79–20.73 mm</quantity>
. Elytral width 
<quantity metricMagnitude="-2" metricUnit="m" metricValue="2.1435" metricValueMax="2.237" metricValueMin="2.05" pageId="7" pageNumber="1322" unit="mm" value="21.435" valueMax="22.37" valueMin="20.5">20.50– 22.37 mm</quantity>
(
<figureCitation box="[288,387,1495,1520]" captionStart="Figure 1" captionStartId="6.[160,225,908,931]" captionTargetBox="[163,1153,143,874]" captionTargetId="figure-234@6.[162,1154,142,876]" captionTargetPageId="6" captionText="Figure 1. Byrsopolis castanea Burmeister dorsal habitus (a, c, e) and lateral habitus (b, d, f). Byrsopolis castanea lectotype male (a–b) and paralectotype male and female (c–f), box series label (g). Images from Karla Schneider (MLUH)." figureDoi="http://doi.org/10.5281/zenodo.7156478" httpUri="https://zenodo.org/record/7156478/files/figure.png" pageId="7" pageNumber="1322">Figures 1</figureCitation>
(e–f), 5(c-d)). 
<emphasis box="[540,597,1495,1520]" italics="true" pageId="7" pageNumber="1322">Head</emphasis>
. Clypeus about 2x wider than long, apical angles rounded with slight concavity near the apex; punctures small to large, densely punctate; frons with small to moderate punctures, moderately to densely punctate; in last maxillary palpomere with excavated area larger; antennal club about 1.5x as long as antennomeres II–VII combined. 
<emphasis box="[346,421,1634,1659]" italics="true" pageId="7" pageNumber="1322">Thorax</emphasis>
. Pronotum transverse about 2x wider than long; small punctures. 
<emphasis box="[160,221,1669,1694]" italics="true" pageId="7" pageNumber="1322">Elytra</emphasis>
. About 1.40x longer than wide; convex; punctures small to large, moderately to densely punctate.
</paragraph>
<paragraph blockId="8.[160,1156,142,237]" pageId="8" pageNumber="1323">
<emphasis bold="true" box="[160,277,142,168]" italics="true" pageId="8" pageNumber="1323">Variation.</emphasis>
Some specimens exhibit variation in colour, from reddish-brown to black, especially on marginal areas; apex of clypeus weakly to moderately emarginate; apex of parameres weakly to moderately divergent.
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="1323" type="etymology">
<paragraph blockId="8.[160,1155,281,341]" pageId="8" pageNumber="1323">
<emphasis bold="true" box="[160,291,281,307]" italics="true" pageId="8" pageNumber="1323">Etymology.</emphasis>
The species name 
<taxonomicName authorityName="Burmeister" authorityYear="1844" box="[516,614,282,307]" class="Insecta" family="Rutelidae" genus="Byrsopolis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1323" phylum="Arthropoda" rank="species" species="castanea">
<emphasis box="[516,614,282,307]" italics="true" pageId="8" pageNumber="1323">castanea</emphasis>
</taxonomicName>
is derived from Latin, in reference to the brown colouration of the specimens.
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="1323" type="distribution">
<paragraph blockId="8.[160,1155,385,445]" pageId="8" pageNumber="1323">
<emphasis bold="true" box="[160,307,385,411]" italics="true" pageId="8" pageNumber="1323">Distribution.</emphasis>
<collectingRegion box="[323,466,386,411]" country="Brazil" name="Minas Gerais" pageId="8" pageNumber="1323">Minas Gerais</collectingRegion>
(Alto Caparaó), 
<collectingRegion box="[648,804,386,411]" country="Brazil" name="Espirito Santo" pageId="8" pageNumber="1323">Espírito Santo</collectingRegion>
(Conceição do Castelo), 
<collectingRegion country="Brazil" name="Rio de Janeiro" pageId="8" pageNumber="1323">Rio de Janeiro</collectingRegion>
(Nova Friburgo [Conquista, Macaé de Cima] and Teresópolis) (
<figureCitation box="[945,1051,420,445]" captionStart="Figure 19" captionStartId="41.[160,225,961,984]" captionTargetBox="[162,1154,142,929]" captionTargetId="figure-144@41.[162,1154,142,929]" captionTargetPageId="41" captionText="Figure 19. Distribution map of Byrsopolis species." figureDoi="http://doi.org/10.5281/zenodo.7156532" httpUri="https://zenodo.org/record/7156532/files/figure.png" pageId="8" pageNumber="1323">Figure 19</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="1323" type="discussion">
<paragraph blockId="8.[160,1155,489,688]" pageId="8" pageNumber="1323">
<emphasis bold="true" box="[160,268,489,515]" italics="true" pageId="8" pageNumber="1323">Remarks.</emphasis>
This species is restricted to the Atlantic Forest of Serra do Mar and Serra dos Órgãos mountains in 
<collectingRegion box="[429,676,524,549]" country="Brazil" name="Rio de Janeiro" pageId="8" pageNumber="1323">Rio de Janeiro state</collectingRegion>
(near the cities of Nova Friburgo and Teresópolis), Serra de Conceição do Castelo in 
<collectingRegion box="[686,903,559,584]" country="Brazil" name="Espirito Santo" pageId="8" pageNumber="1323">Espírito Santo state</collectingRegion>
and Caparaó in 
<collectingRegion country="Brazil" name="Minas Gerais" pageId="8" pageNumber="1323">Minas Gerais state</collectingRegion>
. The locations of Alto Caparaó in 
<collectingRegion box="[679,822,594,619]" country="Brazil" name="Minas Gerais" pageId="8" pageNumber="1323">Minas Gerais</collectingRegion>
and Conceição do Castelo in 
<collectingRegion box="[160,316,628,653]" country="Brazil" name="Espirito Santo" pageId="8" pageNumber="1323">Espírito Santo</collectingRegion>
are new state records. Some specimens were collected in Nova Friburgo (
<collectingRegion box="[166,328,663,688]" country="Brazil" name="Rio de Janeiro" pageId="8" pageNumber="1323">Rio de Janeiro</collectingRegion>
) using light traps.
</paragraph>
</subSubSection>
</treatment>
</document>