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<document id="709DD5D6E5E3B9E7C3214B3BC8DBA6CB" ID-CLB-Dataset="53278" ID-DOI="10.5852/ejt.2023.865.2097" ID-GBIF-Dataset="3a63a183-743e-4185-8fdc-1080072ada0a" ID-ISSN="2118-9773" ID-Zenodo-Dep="7867448" ID-ZooBank="845BAE1A-0E6E-484B-BE6D-17F866777B51" IM.bibliography_approvedBy="diego" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="diego" checkinTime="1682512407979" checkinUser="felipe" docAuthor="Rivera-Quiroz, F. Andrés &amp; Álvarez-Padilla, Fernando" docDate="2023" docId="03C8E927FFB1FF98FDC9FB11FB8B600E" docLanguage="en" docName="ejt-865_1-94.pdf" docOrigin="European Journal of Taxonomy 865" docSource="http://dx.doi.org/10.5852/ejt.2023.865.2097" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Anyphaena Sundevall 1833" docType="treatment" docVersion="6" lastPageNumber="7" masterDocId="FFF1915FFFB4FF90FF81FFF8FF856160" masterDocTitle="Integration or minimalism: twenty-one new species of ghost spiders (Anyphaenidae: Anyphaena) from Mexico" masterLastPageNumber="94" masterPageNumber="1" pageNumber="6" updateTime="1699514891957" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="EB4C0FBCBF734193EBA28CAA5BA0145F">Integration or minimalism: twenty-one new species of ghost spiders (Anyphaenidae: Anyphaena) from Mexico</mods:title>
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<mods:namePart id="6880298E761520FDB6A4B3089425AFDE">Rivera-Quiroz, F. Andrés</mods:namePart>
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<mods:affiliation id="195B678BEEA6BEB759986D0373A832CC">Understanding Evolution Research Group, Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, the Netherlands. Laboratorio de Aracnología, Departamento de Biología Comparada, Facultad de Ciencias, Universidad Nacional Autónoma de México, Distrito Federal, México Ciudad Universitaria, México D. F. Del. Coyoacán, C. P. 04510, Mexico.</mods:affiliation>
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<treatment id="03C8E927FFB1FF98FDC9FB11FB8B600E" ID-DOI="http://doi.org/10.5281/zenodo.7867463" ID-GBIF-Taxon="209579070" ID-Zenodo-Dep="7867463" LSID="urn:lsid:plazi:treatment:03C8E927FFB1FF98FDC9FB11FB8B600E" httpUri="http://treatment.plazi.org/id/03C8E927FFB1FF98FDC9FB11FB8B600E" lastPageId="8" lastPageNumber="7" pageId="5" pageNumber="6">
<subSubSection id="C37B0BBAFFB1FF95FDC9FB11FC6E6463" box="[584,1003,1257,1284]" pageId="5" pageNumber="6" type="nomenclature">
<paragraph id="8BDE5831FFB1FF95FDC9FB11FC6E6463" blockId="5.[584,1003,1257,1284]" box="[584,1003,1257,1284]" pageId="5" pageNumber="6">
<heading id="D096EF5DFFB1FF95FDC9FB11FC6E6463" box="[584,1003,1257,1284]" centered="true" fontSize="11" level="2" pageId="5" pageNumber="6" reason="2">
Genus
<taxonomicName id="4C6123B2FFB1FF95FD1AFB11FC6E6463" ID-CoL="62DGM" ID-ENA="336631" authority="Sundevall, 1833" authorityName="Sundevall" authorityYear="1833" box="[667,1003,1257,1284]" class="Arachnida" family="Anyphaenidae" genus="Anyphaena" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B9158423FFB1FF95FD1AFB11FCA36464" bold="true" box="[667,806,1257,1284]" italics="true" pageId="5" pageNumber="6">Anyphaena</emphasis>
Sundevall, 1833
</taxonomicName>
</heading>
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<subSubSection id="C37B0BBAFFB1FF95FF3CFAD3FCCE6413" pageId="5" pageNumber="6" type="type_taxon">
<paragraph id="8BDE5831FFB1FF95FF3CFAD3FEDD6424" blockId="5.[189,344,1322,1349]" box="[189,344,1322,1349]" pageId="5" pageNumber="6">
<heading id="D096EF5DFFB1FF95FF3CFAD3FEDD6424" bold="true" box="[189,344,1322,1349]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="B9158423FFB1FF95FF3CFAD3FEDD6424" bold="true" box="[189,344,1322,1349]" pageId="5" pageNumber="6">
<typeStatus id="54DAE693FFB1FF95FF3CFAD3FF7F6425" box="[189,250,1323,1349]" pageId="5" pageNumber="6">Type</typeStatus>
species
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</heading>
</paragraph>
<paragraph id="8BDE5831FFB1FF95FF3CFAA1FCCE6413" blockId="5.[189,843,1369,1395]" box="[189,843,1369,1395]" pageId="5" pageNumber="6">
<taxonomicName id="4C6123B2FFB1FF95FF3CFAA1FDF26413" authority="Walckenaer, 1802" authorityName="Walckenaer" authorityYear="1802" box="[189,631,1369,1395]" class="Gastropoda" family="Muricidae" genus="Aranea" kingdom="Animalia" order="Neogastropoda" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="accentuata">
<emphasis id="B9158423FFB1FF95FF3CFAA1FE196413" box="[189,412,1369,1395]" italics="true" pageId="5" pageNumber="6">Aranea accentuata</emphasis>
Walckenaer, 1802
</taxonomicName>
; gender feminine.
</paragraph>
</subSubSection>
<subSubSection id="C37B0BBAFFB1FF95FD51FA62FCE664D4" box="[720,867,1434,1461]" pageId="5" pageNumber="6" type="multiple">
<paragraph id="8BDE5831FFB1FF95FD51FA62FCE664D4" blockId="5.[720,867,1434,1461]" box="[720,867,1434,1461]" pageId="5" pageNumber="6">
<heading id="D096EF5DFFB1FF95FD51FA62FCE664D4" box="[720,867,1434,1461]" centered="true" fontSize="11" level="2" pageId="5" pageNumber="6" reason="2">
<emphasis id="B9158423FFB1FF95FD51FA62FCE664D4" bold="true" box="[720,867,1434,1461]" pageId="5" pageNumber="6">
<taxonomicName id="4C6123B2FFB1FF95FD51FA62FC9464D5" ID-CoL="F5HM" ID-ENA="1109461" baseAuthorityName="Hentz" baseAuthorityYear="1847" box="[720,785,1434,1461]" class="Arachnida" family="Anyphaenidae" genus="Anyphaena" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="celer">
<emphasis id="B9158423FFB1FF95FD51FA62FC9464D5" bold="true" box="[720,785,1434,1461]" italics="true" pageId="5" pageNumber="6">Celer</emphasis>
</taxonomicName>
group
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C37B0BBAFFB1FF98FF3CFA23FB8B600E" lastPageId="8" lastPageNumber="9" pageId="5" pageNumber="6" type="diagnosis">
<paragraph id="8BDE5831FFB1FF95FF3CFA23FEB36495" blockId="5.[189,310,1499,1525]" box="[189,310,1499,1525]" pageId="5" pageNumber="6">
<heading id="D096EF5DFFB1FF95FF3CFA23FEB36495" bold="true" box="[189,310,1499,1525]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="B9158423FFB1FF95FF3CFA23FEB36495" bold="true" box="[189,310,1499,1525]" pageId="5" pageNumber="6">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BDE5831FFB1FF95FF3CF9F2FAD66744" blockId="5.[189,1363,1546,1573]" box="[189,1363,1546,1573]" pageId="5" pageNumber="6">
The
<taxonomicName id="4C6123B2FFB1FF95FF73F9F2FEA86744" baseAuthorityName="Hentz" baseAuthorityYear="1847" box="[242,301,1546,1572]" class="Arachnida" family="Anyphaenidae" genus="Anyphaena" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="celer">
<emphasis id="B9158423FFB1FF95FF73F9F2FEA86744" box="[242,301,1546,1572]" italics="true" pageId="5" pageNumber="6">celer</emphasis>
</taxonomicName>
group can be separated from other
<taxonomicName id="4C6123B2FFB1FF95FD4CF9F2FC286744" box="[717,941,1546,1572]" pageId="5" pageNumber="6">
<emphasis id="B9158423FFB1FF95FD4CF9F2FCD46744" box="[717,849,1546,1572]" italics="true" pageId="5" pageNumber="6">Anyphaena</emphasis>
species
</taxonomicName>
groups by the following characters.
</paragraph>
<paragraph id="8BDE5831FFB1FF95FF6BF9B4FEB06706" blockId="5.[187,1401,1612,2025]" box="[234,309,1612,1638]" pageId="5" pageNumber="6">
<emphasis id="B9158423FFB1FF95FF6BF9B4FEB06706" bold="true" box="[234,309,1612,1638]" pageId="5" pageNumber="6">Males</emphasis>
</paragraph>
<paragraph id="8BDE5831FFB1FF96FF3CF997FD636026" blockId="5.[187,1401,1612,2025]" lastBlockId="6.[188,1398,265,326]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
Coxae lacking ventral spurs (spine-like modifications) but might present clusters of setae (
<figureCitation id="135A44B4FFB1FF95FA92F997FAE867E9" box="[1299,1389,1647,1673]" captionStart="Fig" captionStartId="9.[189,232,1927,1953]" captionTargetBox="[189,1399,775,1879]" captionTargetId="figure-209@9.[189,1399,771,1882]" captionTargetPageId="9" captionText="Fig. 3. Anyphaena natachae sp. nov., paratypes (CNAN-T01562). A. ♀, dorsal habitus. B. ♀, lateral habitus. C. ♀, ventral habitus. D. ♀, prosoma, anterior view. E. ♂, prosoma, anterior view. F. ♂, prosoma, oblique view. Scale bars: A–C= 1.0 mm; D–F =0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7867465" httpUri="https://zenodo.org/record/7867465/files/figure.png" pageId="5" pageNumber="6">Fig. 3F</figureCitation>
) or tubercles (cuticle blunt protuberances of various shapes) (
<figureCitation id="135A44B4FFB1FF95FC0CF96AFB9C67CC" box="[909,1049,1682,1708]" captionStart="Fig" captionStartId="66.[189,232,1892,1918]" captionTargetBox="[258,1334,266,1850]" captionTargetId="figure-82@66.[247,1341,251,1860]" captionTargetPageId="66" captionText="Fig. 41. Anyphaena porta sp. nov. A–F. Paratype, ♂ (CNAN-T01563). G–H. Paratype, ♀ (CNAN-T01565). A. Pedipalp, ventral view. B. Pedipalp, prolateral view. C. Pedipalp, dorsal view. D. Pedipalp, retrolateral view. E. Coxae, ventral view. F. Coxae, lateral view. G. Epigynum, ventral view. H. Epigynum, dorsal view. Scale bars: A–D, G–H =0.2 mm; E–F =1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7867578" httpUri="https://zenodo.org/record/7867578/files/figure.png" pageId="5" pageNumber="6">Fig. 41E–F</figureCitation>
). The palp of the
<taxonomicName id="4C6123B2FFB1FF95FB70F96AFAAE67CC" baseAuthorityName="Hentz" baseAuthorityYear="1847" box="[1265,1323,1682,1708]" class="Arachnida" family="Anyphaenidae" genus="Anyphaena" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="celer">
<emphasis id="B9158423FFB1FF95FB70F96AFAAE67CC" box="[1265,1323,1682,1708]" italics="true" pageId="5" pageNumber="6">celer</emphasis>
</taxonomicName>
group (
<figureCitation id="135A44B4FFB1FF95FF47F94DFEC067AF" box="[198,325,1717,1743]" captionStart="Fig" captionStartId="6.[189,232,1926,1952]" captionTargetBox="[197,1396,585,1886]" captionTargetId="figure-103@6.[189,1399,585,1887]" captionTargetPageId="6" captionText="Fig. 1. Anyphaena spp., expanded pedipalps. A–C. A. natachae sp. nov., ♂ (PXF302). A. Ventral view. B. Retrolateral view. C. Prolateral view. – D–F. A. urieli sp. nov., allotype, ♂ (CNAN-T01521). D. Ventral view. E. Retrolateral view. F. Prolateral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7867455" httpUri="https://zenodo.org/record/7867455/files/figure.png" pageId="5" pageNumber="6">Fig. 1A–C</figureCitation>
) (
<bibRefCitation id="EFF025C0FFB1FF95FEDCF94DFD8367AF" author="Platnick N. I." box="[349,518,1717,1743]" pageId="5" pageNumber="6" pagination="205 - 266" refId="ref43826" refString="Platnick N. I. 1974. The spider family Anyphaenidae in America north of Mexico. Bulletin of the Museum of Comparative Zoology 146: 205 - 266." type="journal article" year="1974">Platnick 1974</bibRefCitation>
;
<bibRefCitation id="EFF025C0FFB1FF95FD92F94DFC8867B0" author="Platnick N. I. &amp; A. Lau" box="[531,781,1717,1744]" pageId="5" pageNumber="6" pagination="1 - 36" refId="ref43889" refString="Platnick N. I. &amp; A. Lau. 1975. A revision of the celer group of the spider genus Anyphaena (Araneae, Anyphaenidae) in Mexico and Central America. American Museum Novitates 2575: 1 - 36." type="journal article" year="1975">Platnick &amp; Lau 1975</bibRefCitation>
) presents a tegulum divided in three lamella-shaped projections, described as prolateral tegular projection, retrolateral tegular projection and ventral tegular projection following the homologies in
<bibRefCitation id="EFF025C0FFB1FF95FD03F904FCC16676" author="Brescovit A. D." box="[642,836,1788,1814]" pageId="5" pageNumber="6" pagination="1 - 187" refId="ref41217" refString="Brescovit A. D. 1997. Revisao de Anyphaeninae Bertkau a nivel de generos na regiao Neotropical (Araneae, Anyphaenidae). Revista brasileira de Zoologia 13 (Suppl. 1): 1 - 187. https: // doi. org / 10.1590 / S 0101 - 81751996000500001" type="journal article" year="1997">Brescovit (1997)</bibRefCitation>
and
<bibRefCitation id="EFF025C0FFB1FF95FCFBF904FB446676" author="Oliveira L. F. M. de &amp; Brescovit A. D." box="[890,1217,1788,1814]" pageId="5" pageNumber="6" pagination="1 - 77" refId="ref43466" refString="Oliveira L. F. M. de &amp; Brescovit A. D. 2021. Taxonomic revision and cladistic analysis of ghost spiders of the genus Tafana Simon, 1903 (Araneae: Dionycha, Anyphaenidae), with the descriptions of twelve new species. European Journal of Taxonomy 742: 1 - 77. https: // doi. org / 10.5852 / ejt. 2021.742.1291" type="journal article" year="2021">Oliveira &amp; Brescovit (2021)</bibRefCitation>
. The embolus is a visible sclerite (in the un-expanded palp), tubular in shape, and curved dorsally. The median apophysis (or conductor in
<bibRefCitation id="EFF025C0FFB1FF95FEFFF8BAFDF6663D" author="Platnick N. I. &amp; A. Lau" box="[382,627,1858,1885]" pageId="5" pageNumber="6" pagination="1 - 36" refId="ref43889" refString="Platnick N. I. &amp; A. Lau. 1975. A revision of the celer group of the spider genus Anyphaena (Araneae, Anyphaenidae) in Mexico and Central America. American Museum Novitates 2575: 1 - 36." type="journal article" year="1975">Platnick &amp; Lau 1975</bibRefCitation>
) is a thin, transparent and curved sclerite, articulated at the base of the ventral tegular projection by flexible cuticle. In expanded pedipalps, the retrolateral tegular projection and prolateral tegular projection are lamella-shaped and partially articulated. In the unexpanded bulb, these projections lie dorsally like two folded tortillas around the other sclerites (
<figureCitation id="135A44B4FFB1FF95FBE7F853FB6366A5" box="[1126,1254,1963,1989]" captionStart="Fig" captionStartId="6.[189,232,1926,1952]" captionTargetBox="[197,1396,585,1886]" captionTargetId="figure-103@6.[189,1399,585,1887]" captionTargetPageId="6" captionText="Fig. 1. Anyphaena spp., expanded pedipalps. A–C. A. natachae sp. nov., ♂ (PXF302). A. Ventral view. B. Retrolateral view. C. Prolateral view. – D–F. A. urieli sp. nov., allotype, ♂ (CNAN-T01521). D. Ventral view. E. Retrolateral view. F. Prolateral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7867455" httpUri="https://zenodo.org/record/7867455/files/figure.png" pageId="5" pageNumber="6">Fig. 1A–C</figureCitation>
). Their RTA is dominated by the anterior and posterior branches, the former usually larger and shaped as a folded cuticular lamella, the latter is usually smaller and its shape varies considerably between species (
<bibRefCitation id="EFF025C0FFB2FF96FA94FEF1FF7C6026" author="Platnick N. I." pageId="6" pageNumber="7" pagination="205 - 266" refId="ref43826" refString="Platnick N. I. 1974. The spider family Anyphaenidae in America north of Mexico. Bulletin of the Museum of Comparative Zoology 146: 205 - 266." type="journal article" year="1974">Platnick 1974</bibRefCitation>
,
<bibRefCitation id="EFF025C0FFB2FF96FE84FED4FEC66026" author="Platnick N. I." box="[261,323,300,326]" pageId="6" pageNumber="7" pagination="207 - 210" refId="ref43856" refString="Platnick N. I. 1977. New species and records of the Anyphaena celer group in Mexico (Araneae, Anyphaenidae). Journal of Arachnology 4: 207 - 210." type="journal article" year="1977">1977</bibRefCitation>
;
<bibRefCitation id="EFF025C0FFB2FF96FECFFED4FDC66026" author="Platnick N. I. &amp; A. Lau" box="[334,579,299,326]" pageId="6" pageNumber="7" pagination="1 - 36" refId="ref43889" refString="Platnick N. I. &amp; A. Lau. 1975. A revision of the celer group of the spider genus Anyphaena (Araneae, Anyphaenidae) in Mexico and Central America. American Museum Novitates 2575: 1 - 36." type="journal article" year="1975">Platnick &amp; Lau 1975</bibRefCitation>
) (
<figureCitation id="135A44B4FFB2FF96FDDBFED4FD5C6026" box="[602,729,299,326]" captionStart="Fig" captionStartId="11.[191,234,1926,1952]" captionTargetBox="[206,1372,611,1877]" captionTargetId="figure-162@11.[189,1398,595,1887]" captionTargetPageId="11" captionText="Fig. 5. Anyphaena natachae sp. nov., paratypes (CNAN-T01562). A. ♂, pedipalp, ventral view. B. ♂, pedipalp, prolateral view. C. ♂, pedipalp, dorsal view. D. ♂, pedipalp, retrolateral view. E. ♀, epigynum, ventral view. F. ♀, epigynum, dorsal view. Scale bars: A–D=0.2 mm; E–F =0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7867473" httpUri="https://zenodo.org/record/7867473/files/figure.png" pageId="6" pageNumber="7">Fig. 5A, C</figureCitation>
).
</paragraph>
<paragraph id="8BDE5831FFB2FF96FF6BFE95FED460E7" blockId="6.[189,1399,365,531]" box="[234,337,365,391]" pageId="6" pageNumber="7">
<emphasis id="B9158423FFB2FF96FF6BFE95FED460E7" bold="true" box="[234,337,365,391]" pageId="6" pageNumber="7">Females</emphasis>
</paragraph>
<paragraph id="8BDE5831FFB2FF98FF3CFE68FB8B600E" blockId="6.[189,1399,365,531]" lastBlockId="8.[189,1400,265,367]" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7">
Externally the epigynum has a hood close to the anterior margin, a sclerotized median plate below the hood, and a transparent atrium delineated by two sclerotized lateral borders. The copulatory openings are difficult to observe; however, they are located near the lateral border's posterior edge (
<figureCitation id="135A44B4FFB2FF96FA8FFE2DFAF4608F" box="[1294,1393,469,495]" captionStart="Fig" captionStartId="7.[189,232,1783,1809]" captionTargetBox="[192,1396,266,1720]" captionTargetId="figure-80@7.[189,1399,265,1728]" captionTargetPageId="7" captionText="Fig. 2. Anyphaena spp., epigynum anatomies. A–B. A. adnani sp. nov., paratype, ♀ (CNAN-T01546). A. Epigynum, ventral view. B. Epigynum, dorsal view. – C–D. A. rebecae sp. nov., holotype, ♀ (CNAN-T01536). C. Epigynum, ventral view. D. Epigynum, dorsal view. – E–F. A. salgueiroi sp. nov., holotype, ♀ (CNAN-T01537). E. Epigynum, ventral view. F. Epigynum, dorsal view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7867461" httpUri="https://zenodo.org/record/7867461/files/figure.png" pageId="6" pageNumber="7">Figs 2A</figureCitation>
,
<figureCitation id="135A44B4FFB2FF96FF3CFE00FF646372" box="[189,225,504,530]" captionStart="Fig" captionStartId="10.[189,232,1925,1951]" captionTargetBox="[193,1397,799,1875]" captionTargetId="figure-231@10.[189,1399,782,1878]" captionTargetPageId="10" captionText="Fig. 4. Anyphaena natachae sp. nov., paratypes (CNAN-T01562). A. ♂, pedipalp, ventral view. B. ♂, pedipalp, prolateral view. C. ♂, pedipalp, dorsal view. D. ♂, pedipalp, retrolateral view. E. ♀, epigynum, ventral view. F. ♀, epigynum, dorsal view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7867471" httpUri="https://zenodo.org/record/7867471/files/figure.png" pageId="6" pageNumber="7">4E</figureCitation>
,
<figureCitation id="135A44B4FFB2FF96FF6FFE00FE946373" box="[238,273,504,531]" captionStart="Fig" captionStartId="11.[191,234,1926,1952]" captionTargetBox="[206,1372,611,1877]" captionTargetId="figure-162@11.[189,1398,595,1887]" captionTargetPageId="11" captionText="Fig. 5. Anyphaena natachae sp. nov., paratypes (CNAN-T01562). A. ♂, pedipalp, ventral view. B. ♂, pedipalp, prolateral view. C. ♂, pedipalp, dorsal view. D. ♂, pedipalp, retrolateral view. E. ♀, epigynum, ventral view. F. ♀, epigynum, dorsal view. Scale bars: A–D=0.2 mm; E–F =0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7867473" httpUri="https://zenodo.org/record/7867473/files/figure.png" pageId="6" pageNumber="7">5E</figureCitation>
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<figureCitation id="135A44B4FFB2FF96FE9FFE01FEC46373" box="[286,321,505,531]" captionStart="Fig" captionStartId="14.[189,232,1927,1953]" captionTargetBox="[189,1401,817,1885]" captionTargetId="figure-238@14.[189,1402,812,1886]" captionTargetPageId="14" captionText="Fig. 7. Anyphaena adnani sp. nov. A–D. Paratype, ♂ (CNAN-T01545).E–F. Paratype, ♀ (CNAN- T01546). A. Pedipalp, ventral view. B. Pedipalp, prolateral view. C. Pedipalp, dorsal view. D. Pedipalp, retrolateral view. E. Epigynum, ventral view. F. Epigynum, dorsal view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7867479" httpUri="https://zenodo.org/record/7867479/files/figure.png" pageId="6" pageNumber="7">7E</figureCitation>
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<figureCitation id="135A44B4FFB2FF96FECEFE00FEF76372" box="[335,370,504,530]" captionStart="Fig" captionStartId="15.[189,232,1890,1916]" captionTargetBox="[211,1384,493,1842]" captionTargetId="figure-131@15.[189,1397,480,1852]" captionTargetPageId="15" captionText="Fig. 8. Anyphaena adnani sp. nov. A–D. Paratype, ♂ (CNAN-T01545). E–F. Paratype, ♀ (CNAN-T01546). A. Pedipalp, ventral view. B. Pedipalp, prolateral view. C. Pedipalp, dorsal view. D. Pedipalp, retrolateral view. E. Epigynum, ventral view. F. Epigynum, dorsal view. Scale bars: A–D =0.2 mm; E–F =0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7867481" httpUri="https://zenodo.org/record/7867481/files/figure.png" pageId="6" pageNumber="7">8E</figureCitation>
,
<figureCitation id="135A44B4FFB2FF96FEFEFE00FE276372" box="[383,418,504,530]" captionStart="Fig" captionStartId="17.[190,233,1926,1952]" captionTargetBox="[190,1394,698,1879]" captionTargetId="figure-138@17.[189,1398,697,1885]" captionTargetPageId="17" captionText="Fig. 9. Anyphaena fernandae sp. nov., holotype, ♀ (CNAN-T01526). A. Dorsal habitus. B. Lateral habitus. C. Ventral habitus. D. Prosoma, anterior view. E. Epigynum, ventral view. F. Epigynum, dorsal view. Scale bars: A–C = 1.0 mm; D =0.5 mm; E–F= 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7867485" httpUri="https://zenodo.org/record/7867485/files/figure.png" pageId="6" pageNumber="7">9E</figureCitation>
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<figureCitation id="135A44B4FFB2FF96FE31FE00FE626372" box="[432,487,504,530]" captionStart="Fig" captionStartId="32.[189,232,1892,1918]" captionTargetBox="[196,1390,437,1825]" captionTargetId="figure-92@32.[189,1398,421,1833]" captionTargetPageId="32" captionText="Fig. 18. Anyphaena spp. A–B. A. fernandae sp. nov., holotype, ♀ (CNAN-T01526). A. Epigynum, ventral view. B. Epigynum, dorsal view. – C–D. A. jimenezi sp. nov., holotype, ♀ (CNAN-T01529). C. Epigynum, ventral view. D. Epigynum, dorsal view. – E–F. A. franciscoi sp. nov., holotype, ♀ (CNAN-T01527). E. Epigynum, ventral view. F. Epigynum, dorsal view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7867521" httpUri="https://zenodo.org/record/7867521/files/figure.png" pageId="6" pageNumber="7">18A</figureCitation>
). Internally, the copulatory and fertilization ducts are usually sclerotized and variable in length and shape (
<bibRefCitation id="EFF025C0FFBCFF98FD9AFEF1FD466043" author="Platnick N. I." box="[539,707,265,291]" pageId="8" pageNumber="9" pagination="205 - 266" refId="ref43826" refString="Platnick N. I. 1974. The spider family Anyphaenidae in America north of Mexico. Bulletin of the Museum of Comparative Zoology 146: 205 - 266." type="journal article" year="1974">Platnick 1974</bibRefCitation>
,
<bibRefCitation id="EFF025C0FFBCFF98FD51FEF1FC8B6043" author="Platnick N. I." box="[720,782,265,291]" pageId="8" pageNumber="9" pagination="207 - 210" refId="ref43856" refString="Platnick N. I. 1977. New species and records of the Anyphaena celer group in Mexico (Araneae, Anyphaenidae). Journal of Arachnology 4: 207 - 210." type="journal article" year="1977">1977</bibRefCitation>
;
<bibRefCitation id="EFF025C0FFBCFF98FC9DFEF1FB9C6044" author="Platnick N. I. &amp; A. Lau" box="[796,1049,265,292]" pageId="8" pageNumber="9" pagination="1 - 36" refId="ref43889" refString="Platnick N. I. &amp; A. Lau. 1975. A revision of the celer group of the spider genus Anyphaena (Araneae, Anyphaenidae) in Mexico and Central America. American Museum Novitates 2575: 1 - 36." type="journal article" year="1975">Platnick &amp; Lau 1975</bibRefCitation>
). The seminal receptacles are cylindrical in shape and also located near the posterior edge. The spermathecae are usually spherical to oval and close to the posterior margin (
<figureCitation id="135A44B4FFBCFF98FD06FEACFD6D600E" box="[647,744,340,366]" captionStart="Fig" captionStartId="7.[189,232,1783,1809]" captionTargetBox="[192,1396,266,1720]" captionTargetId="figure-80@7.[189,1399,265,1728]" captionTargetPageId="7" captionText="Fig. 2. Anyphaena spp., epigynum anatomies. A–B. A. adnani sp. nov., paratype, ♀ (CNAN-T01546). A. Epigynum, ventral view. B. Epigynum, dorsal view. – C–D. A. rebecae sp. nov., holotype, ♀ (CNAN-T01536). C. Epigynum, ventral view. D. Epigynum, dorsal view. – E–F. A. salgueiroi sp. nov., holotype, ♀ (CNAN-T01537). E. Epigynum, ventral view. F. Epigynum, dorsal view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7867461" httpUri="https://zenodo.org/record/7867461/files/figure.png" pageId="8" pageNumber="9">Figs 2B</figureCitation>
,
<figureCitation id="135A44B4FFBCFF98FD75FEACFC96600E" box="[756,787,340,366]" captionStart="Fig" captionStartId="10.[189,232,1925,1951]" captionTargetBox="[193,1397,799,1875]" captionTargetId="figure-231@10.[189,1399,782,1878]" captionTargetPageId="10" captionText="Fig. 4. Anyphaena natachae sp. nov., paratypes (CNAN-T01562). A. ♂, pedipalp, ventral view. B. ♂, pedipalp, prolateral view. C. ♂, pedipalp, dorsal view. D. ♂, pedipalp, retrolateral view. E. ♀, epigynum, ventral view. F. ♀, epigynum, dorsal view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7867471" httpUri="https://zenodo.org/record/7867471/files/figure.png" pageId="8" pageNumber="9">4F</figureCitation>
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<figureCitation id="135A44B4FFBCFF98FC9EFEACFCBB600F" box="[799,830,340,367]" captionStart="Fig" captionStartId="11.[191,234,1926,1952]" captionTargetBox="[206,1372,611,1877]" captionTargetId="figure-162@11.[189,1398,595,1887]" captionTargetPageId="11" captionText="Fig. 5. Anyphaena natachae sp. nov., paratypes (CNAN-T01562). A. ♂, pedipalp, ventral view. B. ♂, pedipalp, prolateral view. C. ♂, pedipalp, dorsal view. D. ♂, pedipalp, retrolateral view. E. ♀, epigynum, ventral view. F. ♀, epigynum, dorsal view. Scale bars: A–D=0.2 mm; E–F =0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7867473" httpUri="https://zenodo.org/record/7867473/files/figure.png" pageId="8" pageNumber="9">5F</figureCitation>
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<figureCitation id="135A44B4FFBCFF98FCCBFEADFCEC600F" box="[842,873,341,367]" captionStart="Fig" captionStartId="14.[189,232,1927,1953]" captionTargetBox="[189,1401,817,1885]" captionTargetId="figure-238@14.[189,1402,812,1886]" captionTargetPageId="14" captionText="Fig. 7. Anyphaena adnani sp. nov. A–D. Paratype, ♂ (CNAN-T01545).E–F. Paratype, ♀ (CNAN- T01546). A. Pedipalp, ventral view. B. Pedipalp, prolateral view. C. Pedipalp, dorsal view. D. Pedipalp, retrolateral view. E. Epigynum, ventral view. F. Epigynum, dorsal view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7867479" httpUri="https://zenodo.org/record/7867479/files/figure.png" pageId="8" pageNumber="9">7F</figureCitation>
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<figureCitation id="135A44B4FFBCFF98FCF7FEACFC10600E" box="[886,917,340,366]" captionStart="Fig" captionStartId="15.[189,232,1890,1916]" captionTargetBox="[211,1384,493,1842]" captionTargetId="figure-131@15.[189,1397,480,1852]" captionTargetPageId="15" captionText="Fig. 8. Anyphaena adnani sp. nov. A–D. Paratype, ♂ (CNAN-T01545). E–F. Paratype, ♀ (CNAN-T01546). A. Pedipalp, ventral view. B. Pedipalp, prolateral view. C. Pedipalp, dorsal view. D. Pedipalp, retrolateral view. E. Epigynum, ventral view. F. Epigynum, dorsal view. Scale bars: A–D =0.2 mm; E–F =0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7867481" httpUri="https://zenodo.org/record/7867481/files/figure.png" pageId="8" pageNumber="9">8F</figureCitation>
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<figureCitation id="135A44B4FFBCFF98FC20FEACFC45600E" box="[929,960,340,366]" captionStart="Fig" captionStartId="17.[190,233,1926,1952]" captionTargetBox="[190,1394,698,1879]" captionTargetId="figure-138@17.[189,1398,697,1885]" captionTargetPageId="17" captionText="Fig. 9. Anyphaena fernandae sp. nov., holotype, ♀ (CNAN-T01526). A. Dorsal habitus. B. Lateral habitus. C. Ventral habitus. D. Prosoma, anterior view. E. Epigynum, ventral view. F. Epigynum, dorsal view. Scale bars: A–C = 1.0 mm; D =0.5 mm; E–F= 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7867485" httpUri="https://zenodo.org/record/7867485/files/figure.png" pageId="8" pageNumber="9">9F</figureCitation>
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<figureCitation id="135A44B4FFBCFF98FC4DFEACFB85600E" box="[972,1024,340,366]" captionStart="Fig" captionStartId="32.[189,232,1892,1918]" captionTargetBox="[196,1390,437,1825]" captionTargetId="figure-92@32.[189,1398,421,1833]" captionTargetPageId="32" captionText="Fig. 18. Anyphaena spp. A–B. A. fernandae sp. nov., holotype, ♀ (CNAN-T01526). A. Epigynum, ventral view. B. Epigynum, dorsal view. – C–D. A. jimenezi sp. nov., holotype, ♀ (CNAN-T01529). C. Epigynum, ventral view. D. Epigynum, dorsal view. – E–F. A. franciscoi sp. nov., holotype, ♀ (CNAN-T01527). E. Epigynum, ventral view. F. Epigynum, dorsal view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7867521" httpUri="https://zenodo.org/record/7867521/files/figure.png" pageId="8" pageNumber="9">18B</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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