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<document id="85595364170B610BB1AD8A160574214A" ID-CLB-Dataset="53278" ID-DOI="10.5852/ejt.2023.865.2097" ID-GBIF-Dataset="3a63a183-743e-4185-8fdc-1080072ada0a" ID-ISSN="2118-9773" ID-Zenodo-Dep="7867448" ID-ZooBank="845BAE1A-0E6E-484B-BE6D-17F866777B51" IM.bibliography_approvedBy="diego" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="diego" checkinTime="1682512407979" checkinUser="felipe" docAuthor="Rivera-Quiroz, F. Andrés &amp; Álvarez-Padilla, Fernando" docDate="2023" docId="03C8E927FFA6FF82FD41FA81FEA26689" docLanguage="en" docName="ejt-865_1-94.pdf" docOrigin="European Journal of Taxonomy 865" docSource="http://dx.doi.org/10.5852/ejt.2023.865.2097" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Anyphaena" docType="treatment" docVersion="8" lastPageNumber="19" masterDocId="FFF1915FFFB4FF90FF81FFF8FF856160" masterDocTitle="Integration or minimalism: twenty-one new species of ghost spiders (Anyphaenidae: Anyphaena) from Mexico" masterLastPageNumber="94" masterPageNumber="1" pageNumber="19" updateTime="1699514891957" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="2E6435C1EDC06A6B25D8B8B230329E2F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="4708EF0E2FD77951CEC65D1DEBDD1560">Integration or minimalism: twenty-one new species of ghost spiders (Anyphaenidae: Anyphaena) from Mexico</mods:title>
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<mods:name id="A504A163AA020309C2C0E759F94C6A11" type="personal">
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<mods:namePart id="CC9E863E397742C091C3B6EABD5A8C75">Rivera-Quiroz, F. Andrés</mods:namePart>
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<mods:affiliation id="51E589433CCD6A4B9361F454DB7E2BAE">Understanding Evolution Research Group, Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, the Netherlands. Laboratorio de Aracnología, Departamento de Biología Comparada, Facultad de Ciencias, Universidad Nacional Autónoma de México, Distrito Federal, México Ciudad Universitaria, México D. F. Del. Coyoacán, C. P. 04510, Mexico.</mods:affiliation>
<mods:nameIdentifier id="4297CA57CD47C1F9366F7B751F9630E0" type="email">andres.riveraquiroz@naturalis.nl</mods:nameIdentifier>
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<mods:namePart id="41B1232640F4F7AF4D900E5DE9E3D658">Álvarez-Padilla, Fernando</mods:namePart>
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<mods:nameIdentifier id="EC458FB8C4D9BE507C384435C83A31C1" type="email">fap@ciencias.unam.mx</mods:nameIdentifier>
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<mods:title id="1748E038756E98E9BAFE67C7C28AB46B">European Journal of Taxonomy</mods:title>
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<mods:date id="313A2BEF86408F36C8CC8626CA15D3FA">2023</mods:date>
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<mods:number id="C777FDBC50870BED1785231886471B94">865</mods:number>
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<treatment id="03C8E927FFA6FF82FD41FA81FEA26689" ID-DOI="http://doi.org/10.5281/zenodo.7867477" ID-GBIF-Taxon="209579072" ID-Zenodo-Dep="7867477" LSID="urn:lsid:plazi:treatment:03C8E927FFA6FF82FD41FA81FEA26689" httpUri="http://treatment.plazi.org/id/03C8E927FFA6FF82FD41FA81FEA26689" lastPageNumber="19" pageId="18" pageNumber="19">
<subSubSection id="C37B0BBAFFA6FF82FD41FA81FCF664F4" box="[704,883,1401,1428]" pageId="18" pageNumber="19" type="nomenclature">
<paragraph id="8BDE5831FFA6FF82FD41FA81FCF664F4" blockId="18.[704,883,1401,1428]" box="[704,883,1401,1428]" pageId="18" pageNumber="19">
<heading id="D096EF5DFFA6FF82FD41FA81FCF664F4" box="[704,883,1401,1428]" centered="true" fontSize="11" level="2" pageId="18" pageNumber="19" reason="2">
<emphasis id="B9158423FFA6FF82FD41FA81FCF664F4" bold="true" box="[704,883,1401,1428]" pageId="18" pageNumber="19">
<taxonomicName id="4C6123B2FFA6FF82FD41FA81FCA764F3" ID-CoL="F5JX" ID-ENA="1525866" baseAuthorityName="Banks" baseAuthorityYear="1896" box="[704,802,1401,1427]" class="Arachnida" family="Anyphaenidae" genus="Anyphaena" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B9158423FFA6FF82FD41FA81FCA764F3" bold="true" box="[704,802,1401,1427]" italics="true" pageId="18" pageNumber="19">Pacifica</emphasis>
</taxonomicName>
group
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37B0BBAFFA6FF82FF3CFA46FEA26689" pageId="18" pageNumber="19" type="diagnosis">
<paragraph id="8BDE5831FFA6FF82FF3CFA46FEB364B8" blockId="18.[189,310,1470,1496]" box="[189,310,1470,1496]" pageId="18" pageNumber="19">
<heading id="D096EF5DFFA6FF82FF3CFA46FEB364B8" bold="true" box="[189,310,1470,1496]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
<emphasis id="B9158423FFA6FF82FF3CFA46FEB364B8" bold="true" box="[189,310,1470,1496]" pageId="18" pageNumber="19">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BDE5831FFA6FF82FF3CFA17FAF26769" blockId="18.[189,1399,1519,1546]" box="[189,1399,1519,1546]" pageId="18" pageNumber="19">
The
<taxonomicName id="4C6123B2FFA6FF82FF71FA17FEC86769" baseAuthorityName="Banks" baseAuthorityYear="1896" box="[240,333,1519,1545]" class="Arachnida" family="Anyphaenidae" genus="Anyphaena" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="pacifica">
<emphasis id="B9158423FFA6FF82FF71FA17FEC86769" box="[240,333,1519,1545]" italics="true" pageId="18" pageNumber="19">pacifica</emphasis>
</taxonomicName>
group can be separated from other species groups of
<taxonomicName id="4C6123B2FFA6FF82FC2BFA17FBAB6769" authorityName="Sundevall" authorityYear="1833" box="[938,1070,1519,1545]" class="Arachnida" family="Anyphaenidae" genus="Anyphaena" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B9158423FFA6FF82FC2BFA17FBAB6769" box="[938,1070,1519,1545]" italics="true" pageId="18" pageNumber="19">Anyphaena</emphasis>
</taxonomicName>
by the following characters.
</paragraph>
<paragraph id="8BDE5831FFA6FF82FF6BF9CCFEB0672E" blockId="18.[189,1399,1588,1804]" box="[234,309,1588,1614]" pageId="18" pageNumber="19">
<emphasis id="B9158423FFA6FF82FF6BF9CCFEB0672E" bold="true" box="[234,309,1588,1614]" pageId="18" pageNumber="19">Males</emphasis>
</paragraph>
<paragraph id="8BDE5831FFA6FF82FF3CF9A1FB2F666C" blockId="18.[189,1399,1588,1804]" pageId="18" pageNumber="19">
Both of the species described here show modifications in the venter of coxae II, III and IV (
<figureCitation id="135A44B4FFA6FF82FB7DF9A1FF4B67FA" captionStart="Fig" captionStartId="22.[189,232,1861,1887]" captionTargetBox="[210,1392,267,1807]" captionTargetId="figure-15@22.[189,1398,265,1817]" captionTargetPageId="22" captionText="Fig. 12. Anyphaena triangularis sp. nov. A–F. Paratype, ♂ (CNAN-T01577). G–H. Paratype, ♀ (CNAN-T01579). A. Pedipalp, ventral view. B. Pedipalp, prolateral view. C. Pedipalp, dorsal view. D. Pedipalp, retrolateral view. E. Coxae, ventral view. F. Coxae, lateral view. G. Epigynum, ventral view. H. Epigynum, dorsal view. Scale bars: A–D, G–H =0.2 mm; E–F =1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7867493" httpUri="https://zenodo.org/record/7867493/files/figure.png" pageId="18" pageNumber="19">Figs 12E– F</figureCitation>
,
<figureCitation id="135A44B4FFA6FF82FF5CF987FEB567FA" box="[221,304,1663,1690]" captionStart="Fig" captionStartId="26.[189,232,1891,1917]" captionTargetBox="[232,1361,278,1841]" captionTargetId="figure-82@26.[225,1370,265,1860]" captionTargetPageId="26" captionText="Fig. 15. Anyphaena urieli sp. nov. A–F. Paratype, ♂ (CNAN-T01582). G–H. Paratype, ♀ (CNAN-T01581). A. Pedipalp, ventral view. B. Pedipalp, prolateral view. C. Pedipalp, dorsal view. D. Pedipalp, retrolateral view. E. Coxae, ventral view. F. Coxae, lateral view. G. Epigynum, ventral view. H. Epigynum, dorsal view. Scale bars: A–D, G–H =0.2 mm; E–F =1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7867503" httpUri="https://zenodo.org/record/7867503/files/figure.png" pageId="18" pageNumber="19">15E–F</figureCitation>
). Coxae II show a small and relatively inconspicuous bump. Coxae III and IV have fully developed ventral spurs. The palp of the
<taxonomicName id="4C6123B2FFA6FF82FD39F95EFC9067A0" baseAuthorityName="Banks" baseAuthorityYear="1896" box="[696,789,1702,1728]" class="Arachnida" family="Anyphaenidae" genus="Anyphaena" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="pacifica">
<emphasis id="B9158423FFA6FF82FD39F95EFC9067A0" box="[696,789,1702,1728]" italics="true" pageId="18" pageNumber="19">pacifica</emphasis>
</taxonomicName>
group shows a morphology similar to that of the
<taxonomicName id="4C6123B2FFA6FF82FF3CF935FEB56786" authorityName="L. Koch" authorityYear="1866" box="[189,304,1741,1766]" class="Arachnida" family="Anyphaenidae" genus="Anyphaena" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="pectorosa">
<emphasis id="B9158423FFA6FF82FF3CF935FEB56786" box="[189,304,1741,1766]" italics="true" pageId="18" pageNumber="19">pectorosa</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6123B2FFA6FF82FEEAF935FE2E6786" authorityName="Rivera-Quiroz &amp; Álvarez-Padilla" authorityYear="2023" box="[363,427,1741,1766]" class="Arachnida" family="Anyphaenidae" genus="Anyphaena" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="porta">
<emphasis id="B9158423FFA6FF82FEEAF935FE2E6786" box="[363,427,1741,1766]" italics="true" pageId="18" pageNumber="19">porta</emphasis>
</taxonomicName>
groups. It can be separated from both by the large dorsal cymbium bulge, and the larger RTA posterior branch that projects dorso-laterally (
<figureCitation id="135A44B4FFA6FF82FCDFF90AFB91666C" box="[862,1044,1778,1804]" captionStart="Fig" captionStartId="21.[189,232,1928,1954]" captionTargetBox="[189,1397,813,1883]" captionTargetId="figure-179@21.[189,1398,801,1886]" captionTargetPageId="21" captionText="Fig. 11. Anyphaena triangularis sp. nov. A–D. Paratype, ♂ (CNAN-T01577). E–F. Paratype, ♀ (CNAN-T01579). A. Pedipalp, ventral view. B. Pedipalp, prolateral view. C. Pedipalp, dorsal view. D. Pedipalp, retrolateral view.E. Epigynum, ventral view.F. Epigynum, dorsal view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7867491" httpUri="https://zenodo.org/record/7867491/files/figure.png" pageId="18" pageNumber="19">Figs 11A, C–D</figureCitation>
,
<figureCitation id="135A44B4FFA6FF82FBA1F90AFB18666C" box="[1056,1181,1778,1804]" captionStart="Fig" captionStartId="25.[191,234,1925,1951]" captionTargetBox="[190,1394,764,1883]" captionTargetId="figure-195@25.[189,1398,750,1883]" captionTargetPageId="25" captionText="Fig. 14. Anyphaena urieli sp. nov. A–D. Paratype, ♂ (CNAN-T01582). E–F. Paratype, ♀ (CNAN- T01581). A. Pedipalp, ventral view. B. Pedipalp, prolateral view. C. Pedipalp, dorsal view. D. Pedipalp, retrolateral view. E. Epigynum, ventral view. F. Epigynum, dorsal view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7867501" httpUri="https://zenodo.org/record/7867501/files/figure.png" pageId="18" pageNumber="19">14A, C–D</figureCitation>
).
</paragraph>
<paragraph id="8BDE5831FFA6FF82FF6BF8CEFED46630" blockId="18.[189,1400,1846,2025]" box="[234,337,1846,1872]" pageId="18" pageNumber="19">
<emphasis id="B9158423FFA6FF82FF6BF8CEFED46630" bold="true" box="[234,337,1846,1872]" pageId="18" pageNumber="19">Females</emphasis>
</paragraph>
<paragraph id="8BDE5831FFA6FF82FF3CF8A4FEA26689" blockId="18.[189,1400,1846,2025]" pageId="18" pageNumber="19">
Both new species differ from other
<taxonomicName id="4C6123B2FFA6FF82FDDEF8A4FD666616" authorityName="Sundevall" authorityYear="1833" box="[607,739,1884,1910]" class="Arachnida" family="Anyphaenidae" genus="Anyphaena" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B9158423FFA6FF82FDDEF8A4FD666616" box="[607,739,1884,1910]" italics="true" pageId="18" pageNumber="19">Anyphaena</emphasis>
</taxonomicName>
by having an enlarged epigynal plate bulging anteriorly (
<figureCitation id="135A44B4FFA6FF82FF44F87AFEB766FC" box="[197,306,1922,1948]" captionStart="Fig" captionStartId="20.[189,232,1891,1917]" captionTargetBox="[189,1396,732,1850]" captionTargetId="figure-246@20.[189,1399,731,1851]" captionTargetPageId="20" captionText="Fig. 10. Anyphaena triangularis sp. nov. A–D, F. Paratype, ♀ (CNAN-T01579). E. Paratype, ♂ (CNAN-T01577). A. Dorsal habitus. B. Lateral habitus. C. Ventral habitus. D. Prosoma, anterior view. E. Prosoma, anterior view. F. Epigynum, lateral view. Scale bars: A–C=1.0 mm; D–E =0.5 mm; F = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7867489" httpUri="https://zenodo.org/record/7867489/files/figure.png" pageId="18" pageNumber="19">Figs 10F</figureCitation>
,
<figureCitation id="135A44B4FFA6FF82FEC1F87AFEF466FC" box="[320,369,1922,1948]" captionStart="Fig" captionStartId="24.[189,232,1891,1917]" captionTargetBox="[190,1398,739,1847]" captionTargetId="figure-282@24.[189,1398,737,1847]" captionTargetPageId="24" captionText="Fig. 13. Anyphaena urieli sp. nov.A–D. Holotype, ♀ (CNAN-T01542). E. Paratype, ♂ (CNAN-T01582). F. Paratype, ♀ (CNAN-T01581). A. Dorsal habitus. B. Lateral habitus. C. Ventral habitus. D. Prosoma, anterior view. E. Prosoma, anterior view. F. Epigynum, lateral view. Scale bars: A–C= 1.0 mm; D–E= 0.5 mm; F =0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7867497" httpUri="https://zenodo.org/record/7867497/files/figure.png" pageId="18" pageNumber="19">13F</figureCitation>
). This enlarged plate shows two sclerotized spots that might serve as an anchor point for male palp structures; the spermathecae are usually oval to spherical (
<figureCitation id="135A44B4FFA6FF82FBA7F850FB3566A2" box="[1062,1200,1960,1986]" captionStart="Fig" captionStartId="21.[189,232,1928,1954]" captionTargetBox="[189,1397,813,1883]" captionTargetId="figure-179@21.[189,1398,801,1886]" captionTargetPageId="21" captionText="Fig. 11. Anyphaena triangularis sp. nov. A–D. Paratype, ♂ (CNAN-T01577). E–F. Paratype, ♀ (CNAN-T01579). A. Pedipalp, ventral view. B. Pedipalp, prolateral view. C. Pedipalp, dorsal view. D. Pedipalp, retrolateral view.E. Epigynum, ventral view.F. Epigynum, dorsal view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7867491" httpUri="https://zenodo.org/record/7867491/files/figure.png" pageId="18" pageNumber="19">Figs 11E–F</figureCitation>
,
<figureCitation id="135A44B4FFA6FF82FB3EF850FA9266A2" box="[1215,1303,1960,1986]" captionStart="Fig" captionStartId="22.[189,232,1861,1887]" captionTargetBox="[210,1392,267,1807]" captionTargetId="figure-15@22.[189,1398,265,1817]" captionTargetPageId="22" captionText="Fig. 12. Anyphaena triangularis sp. nov. A–F. Paratype, ♂ (CNAN-T01577). G–H. Paratype, ♀ (CNAN-T01579). A. Pedipalp, ventral view. B. Pedipalp, prolateral view. C. Pedipalp, dorsal view. D. Pedipalp, retrolateral view. E. Coxae, ventral view. F. Coxae, lateral view. G. Epigynum, ventral view. H. Epigynum, dorsal view. Scale bars: A–D, G–H =0.2 mm; E–F =1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7867493" httpUri="https://zenodo.org/record/7867493/files/figure.png" pageId="18" pageNumber="19">12G–H</figureCitation>
,
<figureCitation id="135A44B4FFA6FF82FAA7F850FAF166A2" box="[1318,1396,1960,1986]" captionStart="Fig" captionStartId="25.[191,234,1925,1951]" captionTargetBox="[190,1394,764,1883]" captionTargetId="figure-195@25.[189,1398,750,1883]" captionTargetPageId="25" captionText="Fig. 14. Anyphaena urieli sp. nov. A–D. Paratype, ♂ (CNAN-T01582). E–F. Paratype, ♀ (CNAN- T01581). A. Pedipalp, ventral view. B. Pedipalp, prolateral view. C. Pedipalp, dorsal view. D. Pedipalp, retrolateral view. E. Epigynum, ventral view. F. Epigynum, dorsal view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7867501" httpUri="https://zenodo.org/record/7867501/files/figure.png" pageId="18" pageNumber="19">14E–F</figureCitation>
,
<figureCitation id="135A44B4FFA6FF82FF3EF836FE9F6689" box="[191,282,1998,2025]" captionStart="Fig" captionStartId="26.[189,232,1891,1917]" captionTargetBox="[232,1361,278,1841]" captionTargetId="figure-82@26.[225,1370,265,1860]" captionTargetPageId="26" captionText="Fig. 15. Anyphaena urieli sp. nov. A–F. Paratype, ♂ (CNAN-T01582). G–H. Paratype, ♀ (CNAN-T01581). A. Pedipalp, ventral view. B. Pedipalp, prolateral view. C. Pedipalp, dorsal view. D. Pedipalp, retrolateral view. E. Coxae, ventral view. F. Coxae, lateral view. G. Epigynum, ventral view. H. Epigynum, dorsal view. Scale bars: A–D, G–H =0.2 mm; E–F =1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7867503" httpUri="https://zenodo.org/record/7867503/files/figure.png" pageId="18" pageNumber="19">15G–H</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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