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<document id="D6119858AD2514D84604967344093452" ID-DOI="10.11646/zootaxa.4786.4.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="3876475" ID-ZooBank="677E748D-FA3D-4FE7-A0B3-F9FCFA13D8FF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1591279313921" checkinUser="plazi" docAuthor="Belousov, Igor A. &amp; Kabak, Ilya I." docDate="2020" docId="03C08A2BFFF7FFAD5FC6A149FD55916C" docLanguage="en" docName="zootaxa.4786.4.1.pdf" docOrigin="Zootaxa 4786 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trechus shangensis Belousov &amp; Kabak 2020, sp. n." docType="treatment" docUuid="1AE7B28D-4380-457B-B9A8-5B44659BFC1B" docUuidSource="ZooBank" docVersion="3" lastPageNumber="467" masterDocId="FFF9F253FFF9FFBD5F51A66AFFA59377" masterDocTitle="New Trechus species of the kozlovi group from Sichuan (Coleoptera: Carabidae)" masterLastPageNumber="484" masterPageNumber="451" pageNumber="465" updateTime="1700276390150" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="644626B992BE44A282D7B9B2827C78A8" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C393F26E2D82ECA677D1DF474843D14E">
<mods:title id="CEA4FEBD41B2F9BD04ED68379483AC6A">New Trechus species of the kozlovi group from Sichuan (Coleoptera: Carabidae)</mods:title>
</mods:titleInfo>
<mods:name id="57A14C901075B4738E2FCB2649C1457B" type="personal">
<mods:role id="68B873E926D5008F1E8C87AF4EFFC082">
<mods:roleTerm id="64C84C6838A84036DDE16C8BC88CE9B2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="00E80F05F64D697F5579F8C12B070395">Belousov, Igor A.</mods:namePart>
</mods:name>
<mods:name id="9EEDD92E0D40258D41B8C236004F99C9" type="personal">
<mods:role id="C81D7FCAAD01E07D1AE5F63BB35422E1">
<mods:roleTerm id="66B22EF3FAC189BA122F3B0B9D035808">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E3E56956D7CF6489FB657342D9D58D59">Kabak, Ilya I.</mods:namePart>
<mods:affiliation id="7F95988B3FCFB0E57EFE26CD24A4022D">All-Russian Institute of Plant Protection, Podbelskogo 3, St. Petersburg-Pushkin, 196608, Russia &amp; ilkabak @ yandex. ru; https: // orcid / 0000 - 0002 - 5549 - 6539</mods:affiliation>
</mods:name>
<mods:typeOfResource id="0DCD85D64D1224F5FB619EDD10D414C0">text</mods:typeOfResource>
<mods:relatedItem id="C5BFC126A2BC27ADAF9E14C5830180E9" type="host">
<mods:titleInfo id="F4E8324395797D7724EFADF134AB55D2">
<mods:title id="3888CE7CECF09FEB4FC46145649E1AE8">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="568FB34B813D49EE8C294C96930FB069">
<mods:date id="EEB1B4B2491ED9C86CCB73154DA7D457">2020</mods:date>
<mods:detail id="0350C461A69C0AFE4EF157D49CF5D8A8" type="pubDate">
<mods:number id="C94F74D0E1543211C91A69C29A0003F8">2020-06-04</mods:number>
</mods:detail>
<mods:detail id="451A41E8C6B794605531C0C60FFF4A2C" type="volume">
<mods:number id="FA6B36E659DBAECF0A763DD2D35B86AD">4786</mods:number>
</mods:detail>
<mods:detail id="5B93FF8B55B83DFED688BA1DA48A263E" type="issue">
<mods:number id="B45770D205E9A323684A93D90A703A91">4</mods:number>
</mods:detail>
<mods:extent id="03A81BB3A351D0D5EA6C910D8D46460E" unit="page">
<mods:start id="96B88AACF32F4366BEE9C376203FAA90">451</mods:start>
<mods:end id="3F4E61236092A891A7C3D293556689EA">484</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="AE3FB1CF2930C2912CBDD3312C6D1388">journal article</mods:classification>
<mods:identifier id="D27A87BA800DC2FB98EE9E2EEA68AE81" type="DOI">10.11646/zootaxa.4786.4.1</mods:identifier>
<mods:identifier id="1C19CB91B978D1E29E696A3E6DAF2BE4" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="2A4A6554AD52B61E894946EDDE643123" type="Zenodo-Dep">3876475</mods:identifier>
<mods:identifier id="BA19B8957D9B9BC8EEE9F925714D56CA" type="ZooBank">677E748D-FA3D-4FE7-A0B3-F9FCFA13D8FF</mods:identifier>
</mods:mods>
<treatment id="03C08A2BFFF7FFAD5FC6A149FD55916C" LSID="urn:lsid:zoobank.org:act:1AE7B28D-4380-457B-B9A8-5B44659BFC1B" httpUri="http://treatment.plazi.org/id/03C08A2BFFF7FFAD5FC6A149FD55916C" lastPageId="16" lastPageNumber="467" pageId="14" pageNumber="465">
<subSubSection id="C37368B6FFF7FFB35FC6A149FE759449" box="[151,464,1827,1854]" pageId="14" pageNumber="465" type="nomenclature">
<paragraph id="8BD63B3DFFF7FFB35FC6A149FE759449" blockId="14.[151,934,1827,1918]" box="[151,464,1827,1854]" pageId="14" pageNumber="465">
<heading id="D09E8C51FFF7FFB35FC6A149FE759449" bold="true" box="[151,464,1827,1854]" fontSize="11" level="1" pageId="14" pageNumber="465" reason="1">
<emphasis id="B91DE72FFFF7FFB35FC6A149FE759449" bold="true" box="[151,464,1827,1854]" pageId="14" pageNumber="465">
<taxonomicName id="4C6940BEFFF7FFB35FC6A149FE229449" ID-CoL="8MNCD" authority="Belousov &amp; Kabak, 2020" authorityName="Belousov &amp; Kabak" authorityYear="2020" box="[151,391,1827,1854]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="shangensis" status="sp. nov.">
<emphasis id="B91DE72FFFF7FFB35FC6A149FE229449" bold="true" box="[151,391,1827,1854]" italics="true" pageId="14" pageNumber="465">Trechus shangensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E5A54FFF7FFB35EDFA14EFE759449" box="[398,464,1828,1854]" pageId="14" pageNumber="465" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37368B6FFF7FFB35FC6A122FC039409" pageId="14" pageNumber="465" type="description">
<paragraph id="8BD63B3DFFF7FFB35FC6A122FE869415" blockId="14.[151,934,1827,1918]" box="[151,291,1863,1890]" pageId="14" pageNumber="465">
(
<figureCitation id="135227B8FFF7FFB35FF1A122FF579415" box="[160,242,1863,1890]" captionStart="FIGURE 15" captionStartId="18.[151,250,2008,2033]" captionText="FIGURE 15. Trechus shangensis sp. n., habitus, male, paratype." pageId="14" pageNumber="465">Fig. 15</figureCitation>
,
<figureCitation id="135227B8FFF7FFB35FAFA122FEBE9415" box="[254,283,1864,1890]" captionStart="FIGURES 18–20" captionStartId="29.[151,264,1948,1973]" captionTargetBox="[218,1371,204,1901]" captionTargetId="figure@29.[203,1383,181,1923]" captionTargetPageId="29" captionText="FIGURES 18–20. Male genitalia of Trechus spp. n., paratypes: a—lateral view, b—dorsal view. 18—T. shangensis sp. n.; 19—T. mengensis sp. n.; 20—T. cuspis sp. n." figureDoi="http://doi.org/10.5281/zenodo.3876485" httpUri="https://zenodo.org/record/3876485/files/figure.png" pageId="14" pageNumber="465">18</figureCitation>
)
</paragraph>
<paragraph id="8BD63B3DFFF7FFB35FC6A10CFC039409" blockId="14.[151,934,1827,1918]" box="[151,934,1894,1918]" pageId="14" pageNumber="465">
<uri id="FFF8373FFFF7FFB35FC6A10CFC039409" box="[151,934,1894,1918]" pageId="14" pageNumber="465">
urn:lsid:zoobank.org:act:
<uuid id="FFCF01E8FFF7FFB35EF1A10CFC039409" box="[416,934,1894,1918]" pageId="14" pageNumber="465">1AE7B28D-4380-457B-B9A8-5B44659BFC1B</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C37368B6FFF7FFB25FC6A1C7FCD39236" lastPageId="15" lastPageNumber="466" pageId="14" pageNumber="465" type="materials_examined">
<paragraph id="8BD63B3DFFF7FFB35FC6A1C7FA38949C" blockId="14.[151,1437,1964,2028]" pageId="14" pageNumber="465">
<emphasis id="B91DE72FFFF7FFB35FC6A1C7FEEF94B0" bold="true" box="[151,330,1964,1991]" pageId="14" pageNumber="465">Type material:</emphasis>
<materialsCitation id="3B013160FFF7FFB35E02A1C7FB04949C" collectingDate="2002-07-25" collectionCode="ZISP" collectorName="Kabak I." country="China" elevation="3925" latitude="31.66111" location="Lixian" longLatPrecision="20" longitude="103.030556" pageId="14" pageNumber="465" specimenCount="1" specimenCount-male="1" stateProvince="Sichuan" typeStatus="holotype">
<emphasis id="B91DE72FFFF7FFB35E02A1C7FE6594B0" bold="true" box="[339,448,1965,1991]" pageId="14" pageNumber="465">
<typeStatus id="54D2859FFFF7FFB35E02A1C7FE6594B0" box="[339,448,1965,1991]" pageId="14" pageNumber="465" type="holotype">Holotype</typeStatus>
</emphasis>
: 1(1)
<specimenCount id="9D6FF0B4FFF7FFB35EACA1C7FDB394B0" box="[509,534,1965,1991]" pageId="14" pageNumber="465" type="male">♂</specimenCount>
, “
<collectingCountry id="F37E7BADFFF7FFB35D61A1C7FDDD94B0" box="[560,632,1965,1991]" name="China" pageId="14" pageNumber="465">China</collectingCountry>
, NW
<collectingRegion id="49ADF5DFFFF7FFB35DECA1C7FCBF94B0" box="[701,794,1965,1991]" country="China" name="Sichuan" pageId="14" pageNumber="465">Sichuan</collectingRegion>
, NW of
<location id="8EB66DE6FFF7FFB35CD0A1C7FC6894B0" LSID="urn:lsid:plazi:treatment:03C08A2BFFF7FFAD5FC6A149FD55916C:8EB66DE6FFF7FFB35CD0A1C7FC6894B0" box="[897,973,1965,1991]" country="China" latitude="31.66111" longLatPrecision="20" longitude="103.030556" name="Lixian" pageId="14" pageNumber="465" stateProvince="Sichuan">Lixian</location>
,
<locationDeviation id="9A64221FFFF7FFB35C8BA1C7FA8794B0" box="[986,1314,1965,1991]" location="Shangmeng" pageId="14" pageNumber="465">
<quantity id="4C9196D8FFF7FFB35C8BA1C7FB8194B0" box="[986,1060,1965,1991]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="465" unit="km" value="10.0">10 km</quantity>
WSW of Shangmeng
</locationDeviation>
, alp. scree
<quantity id="4C9196D8FFF7FFB35FC6A1BAFE8B949C" box="[151,302,2000,2027]" metricMagnitude="3" metricUnit="m" metricValue="3.925" metricValueMax="4.0" metricValueMin="3.85" pageId="14" pageNumber="465" unit="m" value="3925.0" valueMax="4000.0" valueMin="3850.0">
<elevation id="0044DC0EFFF7FFB35FC6A1BAFE8B949C" box="[151,302,2000,2027]" metricMagnitude="3" metricUnit="m" metricValue="3.925" metricValueMax="4.0" metricValueMin="3.85" pageId="14" pageNumber="465" unit="m" value="3925.0" valueMax="4000.0" valueMin="3850.0">3850–4000m</elevation>
</quantity>
,
<date id="FFD71DFDFFF7FFB35E66A1BAFE1D949C" box="[311,440,2000,2027]" pageId="14" pageNumber="465" value="2002-07-25">
<collectingDate id="EF93E415FFF7FFB35E66A1BAFE1D949C" box="[311,440,2000,2027]" pageId="14" pageNumber="465" value="2002-07-25">25.07.2002</collectingDate>
</date>
, Belousov I. &amp;
<collectorName id="269C5EEBFFF7FFB35D31A1BBFD1B949B" box="[608,702,2001,2028]" pageId="14" pageNumber="465">Kabak I.</collectorName>
leg.” [
<geoCoordinate id="EE5D5DFAFFF7FFB35C54A1BBFC30949C" box="[773,917,2001,2027]" degrees="31" direction="north" minutes="39" orientation="latitude" pageId="14" pageNumber="465" precision="15" seconds="40" value="31.66111">31º39´40´´N</geoCoordinate>
/
<geoCoordinate id="EE5D5DFAFFF7FFB35CF6A1BAFBE0949C" box="[935,1093,2000,2027]" degrees="103" direction="east" minutes="01" orientation="longitude" pageId="14" pageNumber="465" precision="15" seconds="50" value="103.030556">103º01´50´´ E</geoCoordinate>
] (
<collectionCode id="ED78A3F8FFF7FFB35B0BA1BBFB3C949C" box="[1114,1177,2001,2027]" country="Russia" name="Zoological Institute, Russian Academy of Sciences" pageId="14" pageNumber="465">ZISP</collectionCode>
)
</materialsCitation>
.
<emphasis id="B91DE72FFFF7FFB35BFBA1BBFA87949C" bold="true" box="[1194,1314,2001,2027]" pageId="14" pageNumber="465">
<typeStatus id="54D2859FFFF7FFB35BFBA1BBFA87949C" box="[1194,1314,2001,2027]" pageId="14" pageNumber="465" type="paratype">Paratypes</typeStatus>
</emphasis>
: 43(8)
<specimenCount id="9D6FF0B4FFF7FFB35A3DA1BBFA3D949C" box="[1388,1432,2001,2027]" pageId="14" pageNumber="465" type="male">♂♂</specimenCount>
,
</paragraph>
<paragraph id="8BD63B3DFFF6FFB25FC6A6F2FCD39236" blockId="15.[151,1437,152,2025]" pageId="15" pageNumber="466">
<materialsCitation id="3B013160FFF6FFB25FC6A6F2FAB693C4" box="[151,1299,152,179]" collectingDate="2002-07-25" collectionCode="IZAS, BMNH, SMNS, ZISP, ZSM, CAG, CBK, CDW, CJS, CVZ" collectorName="Kabak I." country="China" elevation="3925" latitude="31.66111" location="Lixian" longLatPrecision="20" longitude="103.030556" pageId="15" pageNumber="466" specimenCount="25" specimenCount-female="25" stateProvince="Sichuan">
<specimenCount id="9D6FF0B4FFF6FFB25FC6A6F2FF7A93C4" box="[151,223,152,179]" pageId="15" pageNumber="466" type="female">25♀♀</specimenCount>
, collected with holotype (
<collectionCode id="ED78A3F8FFF6FFB25D5BA6F3FDE993C4" box="[522,588,153,179]" country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="15" pageNumber="466">IZAS</collectionCode>
,
<collectionCode id="ED78A3F8FFF6FFB25D0BA6F3FD1493C4" box="[602,689,153,179]" pageId="15" pageNumber="466">BMNH</collectionCode>
,
<collectionCode id="ED78A3F8FFF6FFB25DEEA6F3FCAB93C4" box="[703,782,153,179]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="15" pageNumber="466">SMNS</collectionCode>
,
<collectionCode id="ED78A3F8FFF6FFB25C4AA6F3FCF193C4" box="[795,852,153,179]" country="Russia" name="Zoological Institute, Russian Academy of Sciences" pageId="15" pageNumber="466">ZISP</collectionCode>
,
<collectionCode id="ED78A3F8FFF6FFB25C33A6F3FC3B93C4" box="[866,926,153,179]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="15" pageNumber="466" type="Museum">ZSM</collectionCode>
,
<collectionCode id="ED78A3F8FFF6FFB25CFDA6F3FC4F93C4" box="[940,1002,153,179]" pageId="15" pageNumber="466">CAG</collectionCode>
,
<collectionCode id="ED78A3F8FFF6FFB25CA9A6F3FB9193C4" box="[1016,1076,153,179]" pageId="15" pageNumber="466">CBK</collectionCode>
,
<collectionCode id="ED78A3F8FFF6FFB25B10A6F3FB2693C4" box="[1089,1155,153,179]" pageId="15" pageNumber="466">CDW</collectionCode>
,
<collectionCode id="ED78A3F8FFF6FFB25BC0A6F3FB6493C4" box="[1169,1217,153,179]" pageId="15" pageNumber="466">CJS</collectionCode>
,
<collectionCode id="ED78A3F8FFF6FFB25B9FA6F3FAAE93C4" box="[1230,1291,153,179]" pageId="15" pageNumber="466">CVZ</collectionCode>
)
</materialsCitation>
; 34(12)
<materialsCitation id="3B013160FFF6FFB25A3CA6F3FBE5938D" collectingDate="2002-07-26" collectionCode="MPU, CBK, CAK" collectorName="Belousov I. &amp; Kabak I." country="China" elevation="4250" latitude="31.65" location="Lixian" longLatPrecision="20" longitude="103.01945" pageId="15" pageNumber="466" specimenCount="28" specimenCount-female="27" specimenCount-male="1" stateProvince="Sichuan">
<specimenCount id="9D6FF0B4FFF6FFB25A3CA6F3FA3D93C4" box="[1389,1432,153,179]" pageId="15" pageNumber="466" type="male">♂♂</specimenCount>
,
<specimenCount id="9D6FF0B4FFF6FFB25FC6A6D6FF7A93A1" box="[151,223,188,214]" pageId="15" pageNumber="466" type="female">27♀♀</specimenCount>
, “
<collectingCountry id="F37E7BADFFF6FFB25FA6A6D6FE9A93A1" box="[247,319,188,214]" name="China" pageId="15" pageNumber="466">China</collectingCountry>
, NW
<collectingRegion id="49ADF5DFFFF6FFB25ED2A6D6FE4593A1" box="[387,480,188,214]" country="China" name="Sichuan" pageId="15" pageNumber="466">Sichuan</collectingRegion>
, NW of
<location id="8EB66DE6FFF6FFB25D15A6D6FD3593A1" LSID="urn:lsid:plazi:treatment:03C08A2BFFF7FFAD5FC6A149FD55916C:8EB66DE6FFF6FFB25D15A6D6FD3593A1" box="[580,656,188,214]" country="China" latitude="31.65" longLatPrecision="20" longitude="103.01945" name="Lixian" pageId="15" pageNumber="466" stateProvince="Sichuan">Lixian</location>
,
<locationDeviation id="9A64221FFFF6FFB25DCCA6D6FC4793A1" box="[669,994,188,214]" location="Shangmeng" pageId="15" pageNumber="466">
<quantity id="4C9196D8FFF6FFB25DCCA6D6FD4393A1" box="[669,742,188,214]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="15" pageNumber="466" unit="km" value="12.0">12 km</quantity>
WSW of Shangmeng
</locationDeviation>
, alp., scree
<quantity id="4C9196D8FFF6FFB25B3AA6D6FAAF93A0" box="[1131,1290,188,215]" metricMagnitude="3" metricUnit="m" metricValue="4.25" metricValueMax="4.5" metricValueMin="4.0" pageId="15" pageNumber="466" unit="m" value="4250.0" valueMax="4500.0" valueMin="4000.0">
<elevation id="0044DC0EFFF6FFB25B3AA6D6FAAF93A0" box="[1131,1290,188,215]" metricMagnitude="3" metricUnit="m" metricValue="4.25" metricValueMax="4.5" metricValueMin="4.0" pageId="15" pageNumber="466" unit="m" value="4250.0" valueMax="4500.0" valueMin="4000.0">4000–4500 m</elevation>
</quantity>
,
<date id="FFD71DFDFFF6FFB25A46A6D6FA3D93A1" box="[1303,1432,188,214]" pageId="15" pageNumber="466" value="2002-07-26">
<collectingDate id="EF93E415FFF6FFB25A46A6D6FA3D93A1" box="[1303,1432,188,214]" pageId="15" pageNumber="466" value="2002-07-26">26.07.2002</collectingDate>
</date>
,
<collectorName id="269C5EEBFFF6FFB25FC6A68AFEBE938D" box="[151,283,224,250]" pageId="15" pageNumber="466">Belousov I.</collectorName>
&amp;
<collectorName id="269C5EEBFFF6FFB25E15A68AFE02938D" box="[324,423,224,250]" pageId="15" pageNumber="466">Kabak I.</collectorName>
leg.” [
<geoCoordinate id="EE5D5DFAFFF6FFB25EA6A68AFD22938D" box="[503,647,224,250]" degrees="31" direction="north" minutes="39" orientation="latitude" pageId="15" pageNumber="466" precision="15" seconds="00" value="31.65">31º39´00´´N</geoCoordinate>
/
<geoCoordinate id="EE5D5DFAFFF6FFB25DF3A68AFCE3938D" box="[674,838,223,250]" degrees="103" direction="east" minutes="01" orientation="longitude" pageId="15" pageNumber="466" precision="15" seconds="10" value="103.01945">103º01´10´´ E</geoCoordinate>
] (
<collectionCode id="ED78A3F8FFF6FFB25C0FA68AFC3A938D" box="[862,927,224,250]" pageId="15" pageNumber="466">MPU</collectionCode>
,
<collectionCode id="ED78A3F8FFF6FFB25CFCA68AFC4C938D" box="[941,1001,224,250]" pageId="15" pageNumber="466">CBK</collectionCode>
,
<collectionCode id="ED78A3F8FFF6FFB25CA9A68AFB9D938D" box="[1016,1080,224,250]" pageId="15" pageNumber="466">CAK</collectionCode>
)
</materialsCitation>
; 8(7)
<materialsCitation id="3B013160FFF6FFB25B2DA68AFCD79236" collectingDate="2002-07-23" collectionCode="ZISP, ZSM, CBK" collectorName="Belousov I. &amp; Kabak I." country="China" elevation="3750" latitude="31.675" location="Lixian" longLatPrecision="20" longitude="103.040276" pageId="15" pageNumber="466" specimenCount="16" specimenCount-female="15" specimenCount-male="1" stateProvince="Sichuan">
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,
<specimenCount id="9D6FF0B4FFF6FFB25BE6A6B5FB5A938D" box="[1207,1279,223,250]" pageId="15" pageNumber="466" type="female">15♀♀</specimenCount>
, “
<collectingCountry id="F37E7BADFFF6FFB25A49A68AFAC5938D" box="[1304,1376,224,250]" name="China" pageId="15" pageNumber="466">China</collectingCountry>
, NW
<collectingRegion id="49ADF5DFFFF6FFB25FC6A769FF51926A" box="[151,244,259,285]" country="China" name="Sichuan" pageId="15" pageNumber="466">Sichuan</collectingRegion>
, NW of
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,
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W of Shangmeng
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, scree above timber-line,
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,
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<collectingDate id="EF93E415FFF6FFB25B22A769FB51926A" box="[1139,1268,259,285]" pageId="15" pageNumber="466" value="2002-07-23">23.07.2002</collectingDate>
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,
<collectorName id="269C5EEBFFF6FFB25BAEA769FADA9269" box="[1279,1407,259,286]" pageId="15" pageNumber="466">Belousov I.</collectorName>
&amp;
<collectorName id="269C5EEBFFF6FFB25FC6A74DFF529236" box="[151,247,295,321]" pageId="15" pageNumber="466">Kabak I.</collectorName>
leg.” [
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/
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] (
<collectionCode id="ED78A3F8FFF6FFB25DCFA74DFD7C9236" box="[670,729,295,321]" country="Russia" name="Zoological Institute, Russian Academy of Sciences" pageId="15" pageNumber="466">ZISP</collectionCode>
,
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,
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)
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.
</paragraph>
</subSubSection>
<subSubSection id="C37368B6FFF6FFAD5F96A723FED89234" lastPageId="16" lastPageNumber="467" pageId="15" pageNumber="466" type="description">
<paragraph id="8BD63B3DFFF6FFB25F96A723FDF69285" blockId="15.[151,1437,152,2025]" pageId="15" pageNumber="466">
<emphasis id="B91DE72FFFF6FFB25F96A723FEFC9213" bold="true" box="[199,345,329,356]" pageId="15" pageNumber="466">Description.</emphasis>
Body elongate-oval, moderately constricted at pronotal base, convex (
<figureCitation id="135227B8FFF6FFB25B3AA720FB1B9213" box="[1131,1214,329,356]" captionStart="FIGURE 15" captionStartId="18.[151,250,2008,2033]" captionText="FIGURE 15. Trechus shangensis sp. n., habitus, male, paratype." pageId="15" pageNumber="466">Fig. 15</figureCitation>
). Appendages moderately long and rather thick. Upperside pitchy black to reddish brown, often disc of pronotum, suture and margins of elytra paler, tinged with reddish. Anterior portion of head before clypeal suture amber brownish. Legs uniformly pale reddish.Antennae either uniformly pale reddish brown or vaguely obscured in middle and distal portions beginning from apical parts of segments 3–5.
</paragraph>
<paragraph id="8BD63B3DFFF6FFB25F96A791FAF391F7" blockId="15.[151,1437,152,2025]" pageId="15" pageNumber="466">Head of medium size, eyes average and subconvex, tempora rather long and slightly convex, with a few very short hairs. Frontal furrows evenly impressed, subangulate or arcuate, markedly approaching in middle, deepest just behind their mid-length. Supraorbital setae located in lines subparallel to body axis, posterior seta far behind posterior margin of eye. Pores of both anterior and posterior setae foveolate, though posterior one shallower.</paragraph>
<paragraph id="8BD63B3DFFF6FFB25F96A4E0FA3790EB" blockId="15.[151,1437,152,2025]" pageId="15" pageNumber="466">Pronotum convex, subcordate, moderately constricted at base, with maximum width in apical third. Its sides broadly arcuate in anterior part, moderately sinuate before hind angles, the latter large, subrectangular or acute, occasionally produced outwards. Posterior margin straight, oblique at sides; anterior margin straight. Anterior angles marked though rounded. Lateral margins moderately reflexed, lateral groove narrow to average, slightly widened posteriorly. Prebasal transverse impression parallel to basal margin, markedly curved near basal foveae, not sharply engraved. Basal foveae large and deep. Apical transverse impression distinct laterally, shallow medially. Anterior marginal seta located in anterior third of pronotum, posterior seta at hind angle. Base of pronotum slightly to markedly longitudinally rugose. Median line well impressed, deepest near its base, not reaching anterior margin of pronotum.</paragraph>
<paragraph id="8BD63B3DFFF6FFB25F96A5CFFDCE9632" blockId="15.[151,1437,152,2025]" pageId="15" pageNumber="466">Elytra oblong-oval, with sides evenly and broadly arcuate, widest slightly behind their mid-length. Humeri marked, but broadly rounded. Elytra conjointly rounded apically, each with a distinct subapical sinuation. Striation of elytra rather complete: striae 1–4 well impressed and entire, others shallow and shortened, becoming less distinguishable toward sides. All elytral striae slightly but distinctly punctured. Stria 2 surpassing level of preapical pore posteriad and usually reaching apical border of elytron, normally without distinct connection with stria 3; the latter joining stria 4 approximately at level of anterior end of apical striole; striae 5 and 6 joining each other far before elytral apex, near level of median group of umbilicate series; stria 7 joining apical striole and forming here a distinct sinuation. Parascutellar striole long and well engraved, parascutellar pore present. Apical recurrent striole deep, straight or slightly curved, rather long, though not reaching level of umbilicate pore 7, markedly divergent from suture anteriad, externally bordered by a well-developed carina. Apical triangle (consisting of three pores on apical slope) very long, angulo-apical pore located closer to exterior pore than to elytral suture. Lateral margins moderately reflexed, lateral groove of medium width.</paragraph>
<paragraph id="8BD63B3DFFF6FFB25F96A325FB1196C7" blockId="15.[151,1437,152,2025]" pageId="15" pageNumber="466">Microsculpture well developed over entire body surface, consisting of isodiametric meshes on head, irregular and slightly transverse meshes on disc of pronotum, becoming coarser and less transverse toward base, and distinctly transverse meshes, partially transverse lines on elytra, latter with faint iridescent luster.</paragraph>
<paragraph id="8BD63B3DFFF6FFB25F96A3D0FEDA956D" blockId="15.[151,1437,152,2025]" pageId="15" pageNumber="466">Fore tibiae flattened or shallowly grooved on exterior surface in middle part, occasionally with a few short and sparse hairs on anterior surface near inner apex of tibia. In male, two proximal segments of protarsi markedly dilated and inwards dentate.</paragraph>
<paragraph id="8BD63B3DFFF6FFB25F96A049FDE194D7" blockId="15.[151,1437,152,2025]" pageId="15" pageNumber="466">
Median lobe of aedeagus large, subcylindrical, rather thick, angularly bent at basal portion (
<figureCitation id="135227B8FFF6FFB25B9DA049FA8C9549" box="[1228,1321,1571,1598]" captionStart="FIGURES 18–20" captionStartId="29.[151,264,1948,1973]" captionTargetBox="[218,1371,204,1901]" captionTargetId="figure@29.[203,1383,181,1923]" captionTargetPageId="29" captionText="FIGURES 18–20. Male genitalia of Trechus spp. n., paratypes: a—lateral view, b—dorsal view. 18—T. shangensis sp. n.; 19—T. mengensis sp. n.; 20—T. cuspis sp. n." figureDoi="http://doi.org/10.5281/zenodo.3876485" httpUri="https://zenodo.org/record/3876485/files/figure.png" pageId="15" pageNumber="466">Fig. 18a</figureCitation>
) in lateral view. Both ventral and dorsal margins distinctly emarginate before apex, the latter in shape of oblique disc. In dorsal view (
<figureCitation id="135227B8FFF6FFB25F8EA000FEE795F2" box="[223,322,1642,1669]" captionStart="FIGURES 18–20" captionStartId="29.[151,264,1948,1973]" captionTargetBox="[218,1371,204,1901]" captionTargetId="figure@29.[203,1383,181,1923]" captionTargetPageId="29" captionText="FIGURES 18–20. Male genitalia of Trechus spp. n., paratypes: a—lateral view, b—dorsal view. 18—T. shangensis sp. n.; 19—T. mengensis sp. n.; 20—T. cuspis sp. n." figureDoi="http://doi.org/10.5281/zenodo.3876485" httpUri="https://zenodo.org/record/3876485/files/figure.png" pageId="15" pageNumber="466">Fig. 18b</figureCitation>
), median lobe rather wide, slightly dilated in apical third, gradually narrowing apically. Preputial field variable, always surpassing mid-length of median lobe. Ventral surface with a slight median carina. Apical lamella distinct, rather short, with convergent sides and broadly rounded apex in dorsal view. Sagittal lobe well developed, thickened but not grooved along its exterior surface. Parameres long and slender, straight in distal half, left paramere clearly longer, with a well-developed ventral apophysis; each paramere bearing 4 apical setae. Endophallus armature consisting of two slightly sclerotized pieces: proximal one semi-ring shaped, located near mid-length of median lobe close to its left wall, its dorsal part always with emargination of different shape and falcate lobe (see from above,
<figureCitation id="135227B8FFF6FFB25E76A109FE2C9409" box="[295,393,1891,1918]" captionStart="FIGURES 18–20" captionStartId="29.[151,264,1948,1973]" captionTargetBox="[218,1371,204,1901]" captionTargetId="figure@29.[203,1383,181,1923]" captionTargetPageId="29" captionText="FIGURES 18–20. Male genitalia of Trechus spp. n., paratypes: a—lateral view, b—dorsal view. 18—T. shangensis sp. n.; 19—T. mengensis sp. n.; 20—T. cuspis sp. n." figureDoi="http://doi.org/10.5281/zenodo.3876485" httpUri="https://zenodo.org/record/3876485/files/figure.png" pageId="15" pageNumber="466">Fig. 18b</figureCitation>
); distal piece smaller, located largely in horizontal plane, overlapped by small scaly patch if viewed from left side and from above.
</paragraph>
<paragraph id="8BD63B3DFFF6FFB25F96A1C3FCA6949F" blockId="15.[151,1437,152,2025]" pageId="15" pageNumber="466">
<emphasis id="B91DE72FFFF6FFB25F96A1C3FE1C94B3" bold="true" box="[199,441,1961,1988]" pageId="15" pageNumber="466">Sexual dimorphism.</emphasis>
Males differ in smaller head (EW/HW, on average, 1.97 vs.
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females, p≤0.001; PW/HW, on average, 1.28 vs.
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females, p≤0.05).
</paragraph>
<paragraph id="8BD63B3DFFE9FFAD5F96A6F2FED89234" blockId="16.[151,1437,152,539]" pageId="16" pageNumber="467">
<emphasis id="B91DE72FFFE9FFAD5F96A6F2FE1793C4" bold="true" box="[199,434,152,179]" pageId="16" pageNumber="467">Comparative notes.</emphasis>
Doubtless,
<taxonomicName id="4C6940BEFFE9FFAD5D68A6F0FD7593C4" authority="Belousov &amp; Kabak, 2020" authorityName="Belousov &amp; Kabak" authorityYear="2020" box="[569,720,153,179]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="467" phylum="Arthropoda" rank="species" species="shangensis" status="sp. nov.">
<emphasis id="B91DE72FFFE9FFAD5D68A6F0FD7593C4" box="[569,720,153,179]" italics="true" pageId="16" pageNumber="467">T. shangensis</emphasis>
</taxonomicName>
<emphasis id="B91DE72FFFE9FFAD5D89A6F3FCBC93C4" bold="true" box="[728,793,153,179]" pageId="16" pageNumber="467">
<taxonomicNameLabel id="A22E5A54FFE9FFAD5D89A6F3FCBC93C4" box="[728,793,153,179]" pageId="16" pageNumber="467" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
belongs to the
<taxonomicName id="4C6940BEFFE9FFAD5C99A6F0FB9193C4" authorityName="Jeannel" authorityYear="1935" box="[968,1076,153,179]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="467" phylum="Arthropoda" rank="species" species="kozlovi">
<emphasis id="B91DE72FFFE9FFAD5C99A6F0FB9193C4" box="[968,1076,153,179]" italics="true" pageId="16" pageNumber="467">T. kozlovi</emphasis>
</taxonomicName>
species group, as confirmed by both the external morphology and the structure of the male genitalia. Within this speciose group, the new species is easily recognized by the shape of the median lobe, which is large and rather thick, sharply curved at basal third and with a well-developed apical disc (
<figureCitation id="135227B8FFE9FFAD5D4DA76FFDCA9268" box="[540,623,261,287]" captionStart="FIGURES 18–20" captionStartId="29.[151,264,1948,1973]" captionTargetBox="[218,1371,204,1901]" captionTargetId="figure@29.[203,1383,181,1923]" captionTargetPageId="29" captionText="FIGURES 18–20. Male genitalia of Trechus spp. n., paratypes: a—lateral view, b—dorsal view. 18—T. shangensis sp. n.; 19—T. mengensis sp. n.; 20—T. cuspis sp. n." figureDoi="http://doi.org/10.5281/zenodo.3876485" httpUri="https://zenodo.org/record/3876485/files/figure.png" pageId="16" pageNumber="467">Fig. 18</figureCitation>
). It is most similar to the sympatric
<taxonomicName id="4C6940BEFFE9FFAD5B50A76CFB349268" authority="Belousov &amp; Kabak, 2020" authorityName="Belousov &amp; Kabak" authorityYear="2020" box="[1025,1169,262,287]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="467" phylum="Arthropoda" rank="species" species="mengensis" status="sp. nov.">
<emphasis id="B91DE72FFFE9FFAD5B50A76CFB349268" box="[1025,1169,262,287]" italics="true" pageId="16" pageNumber="467">T. mengensis</emphasis>
</taxonomicName>
<emphasis id="B91DE72FFFE9FFAD5BC9A76FFB729268" bold="true" box="[1176,1239,261,287]" pageId="16" pageNumber="467">
<taxonomicNameLabel id="A22E5A54FFE9FFAD5BC9A76FFB729268" box="[1176,1239,261,287]" pageId="16" pageNumber="467" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
Their differences are discussed below.
</paragraph>
</subSubSection>
<subSubSection id="C37368B6FFE9FFAD5F96A726FAC592A4" pageId="16" pageNumber="467" type="distribution">
<paragraph id="8BD63B3DFFE9FFAD5F96A726FC2E92FC" blockId="16.[151,1437,152,539]" pageId="16" pageNumber="467">
<emphasis id="B91DE72FFFE9FFAD5F96A726FEC49210" bold="true" box="[199,353,332,359]" pageId="16" pageNumber="467">Distribution.</emphasis>
<taxonomicName id="4C6940BEFFE9FFAD5E38A724FDA49210" authority="Belousov &amp; Kabak, 2020" authorityName="Belousov &amp; Kabak" authorityYear="2020" box="[361,513,333,359]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="467" phylum="Arthropoda" rank="species" species="shangensis" status="sp. nov.">
<emphasis id="B91DE72FFFE9FFAD5E38A724FDA49210" box="[361,513,333,359]" italics="true" pageId="16" pageNumber="467">T. shangensis</emphasis>
</taxonomicName>
<emphasis id="B91DE72FFFE9FFAD5D5BA727FDEE9210" bold="true" box="[522,587,333,359]" pageId="16" pageNumber="467">
<taxonomicNameLabel id="A22E5A54FFE9FFAD5D5BA727FDEE9210" box="[522,587,333,359]" pageId="16" pageNumber="467" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is only known from the upper valley of a small river located WSW of the village of Shangmeng, NW of Lixian City, Lixian County,
<collectingRegion id="49ADF5DFFFE9FFAD5C7BA71BFC2D92FC" box="[810,904,369,395]" country="China" name="Sichuan" pageId="16" pageNumber="467">Sichuan</collectingRegion>
.
</paragraph>
<paragraph id="8BD63B3DFFE9FFAD5F96A7FEFAC592A4" blockId="16.[151,1437,152,539]" pageId="16" pageNumber="467">
<emphasis id="B91DE72FFFE9FFAD5F96A7FEFEEE92D8" bold="true" box="[199,331,404,431]" pageId="16" pageNumber="467">Bionomics.</emphasis>
The species occurs in a wide range of altitudes from
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, in the alpine and upper forest zones. It was collected from under stones in humicolous habitats and by sifting from the
<taxonomicName id="4C6940BEFFE9FFAD5B28A7D3FABB92A4" box="[1145,1310,441,467]" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="16" pageNumber="467" phylum="Tracheophyta" rank="genus">
<emphasis id="B91DE72FFFE9FFAD5B28A7D3FABB92A4" box="[1145,1310,441,467]" italics="true" pageId="16" pageNumber="467">Rhododendron</emphasis>
</taxonomicName>
litter.
</paragraph>
</subSubSection>
<subSubSection id="C37368B6FFE9FFAD5F96A7B6FD55916C" pageId="16" pageNumber="467" type="etymology">
<paragraph id="8BD63B3DFFE9FFAD5F96A7B6FD55916C" blockId="16.[151,1437,152,539]" pageId="16" pageNumber="467">
<emphasis id="B91DE72FFFE9FFAD5F96A7B6FE059280" bold="true" box="[199,416,476,503]" pageId="16" pageNumber="467">Derivatio nominis</emphasis>
. The species epithet refers to the first part of the toponym “Shangmengxiang”, a village located near mountains where this species was found.
</paragraph>
</subSubSection>
</treatment>
</document>
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