279 lines
34 KiB
XML
279 lines
34 KiB
XML
<document id="E47F0519C7395D886E5B0AE95AFDCD63" ID-DOI="10.5281/zenodo.185684" ID-GBIF-Dataset="c2695917-74c7-4122-97b9-1ed7984e8874" ID-ISSN="1175-5326" ID-Zenodo-Dep="185684" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460006098723" checkinUser="plazi" docAuthor="Hershler, Robert & Liu, Hsiu-Ping" docDate="2009" docId="03A1AA57DC58FFD36AFBFEA7FCE5F833" docLanguage="en" docName="zt02006p022.pdf" docOrigin="Zootaxa 2006" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Pyrgulopsis intermedia Tryon 1865" docType="treatment" docVersion="8" lastPageNumber="20" masterDocId="FF98D22FDC4AFFC06A6CFFD8FFEBFF8C" masterDocTitle="New species and records of Pyrgulopsis (Gastropoda: Hydrobiidae) from the Snake River basin, southeastern Oregon: further delineation of a highly imperiled fauna" masterLastPageNumber="22" masterPageNumber="1" pageNumber="19" updateTime="1698233312231" updateUser="plazi">
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<mods:title id="7079940438BE436E6CFF0F429962145E">New species and records of Pyrgulopsis (Gastropoda: Hydrobiidae) from the Snake River basin, southeastern Oregon: further delineation of a highly imperiled fauna</mods:title>
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<mods:namePart id="A74EA8E1194EA264ABCF048B248108FA">Hershler, Robert</mods:namePart>
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<mods:namePart id="71447911E9DE0964BB4885BFADE6130A">Liu, Hsiu-Ping</mods:namePart>
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<mods:title id="A3FD26F67CC3FC075D0C86BC508821BC">Zootaxa</mods:title>
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<mods:date id="5CA88810C0C6CB67E04019106958E15D">2009</mods:date>
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<mods:detail id="232CA221442778B3ED969E524DA884A5" type="volume">
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<mods:number id="316B5E98D788551FFC6E7D38FA505366">2006</mods:number>
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<mods:identifier id="317548509F394D8C06C5093A5DE3712C" type="DOI">10.5281/zenodo.185684</mods:identifier>
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<mods:identifier id="5D5404C775F41F5E7B018EE9E891D2C9" type="Zenodo-Dep">185684</mods:identifier>
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<treatment id="03A1AA57DC58FFD36AFBFEA7FCE5F833" ID-DOI="http://doi.org/10.5281/zenodo.6219810" ID-GBIF-Taxon="119381003" ID-Zenodo-Dep="6219810" LSID="urn:lsid:plazi:treatment:03A1AA57DC58FFD36AFBFEA7FCE5F833" httpUri="http://treatment.plazi.org/id/03A1AA57DC58FFD36AFBFEA7FCE5F833" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
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<subSubSection id="C31248CADC58FFD26AFBFEA7FEC6FE4C" pageId="18" pageNumber="19" type="nomenclature">
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<paragraph id="8BB71B41DC58FFD26AFBFEA7FDB6FE15" blockId="18.[151,605,383,448]" box="[151,605,383,409]" pageId="18" pageNumber="19">
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<heading id="D0FFAC2DDC58FFD26AFBFEA7FDB6FE15" bold="true" box="[151,605,383,409]" fontSize="11" level="1" pageId="18" pageNumber="19" reason="1">
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<taxonomicName id="4C0860C2DC58FFD26AFBFEA7FDB6FE15" ID-CoL="7SYTL" authority="Tryon, 1865" authorityName="Tryon" authorityYear="1865" box="[151,605,383,409]" class="Gastropoda" family="Hydrobiidae" genus="Pyrgulopsis" kingdom="Animalia" order="Littorinimorpha" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="intermedia">
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<emphasis id="B97CC753DC58FFD26AFBFEA7FDB6FE15" bold="true" box="[151,605,383,409]" pageId="18" pageNumber="19">
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<emphasis id="B97CC753DC58FFD26AFBFEA7FE45FE15" bold="true" box="[151,430,383,409]" italics="true" pageId="18" pageNumber="19">Pyrgulopsis intermedia</emphasis>
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(
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<bibRefCitation id="EF9966B0DC58FFD26BD1FEA7FDBEFE15" author="Tryon" box="[445,597,383,409]" pageId="18" pageNumber="22" refString="Tryon, G. W. (1865) Descriptions of new species of Amnicola, Pomatiopsis, Somatogyrus, Gabbia, Hydrobia and Rissoa. American Journal of Conchology, 1, 219 - 220." type="journal article" year="1865">Tryon, 1865</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BB71B41DC58FFD26AFBFE7EFEC6FE4C" blockId="18.[151,605,383,448]" box="[151,301,422,448]" pageId="18" pageNumber="19">
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(
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<figureCitation id="133307C4DC58FFD26AF3FE7EFECFFE4C" box="[159,292,422,448]" captionStart-0="FIGURE 10. P" captionStart-1="FIGURE 11. P" captionStartId-0="19.[151,257,933,957]" captionStartId-1="19.[151,256,1698,1722]" captionTargetBox-0="[157,1430,310,909]" captionTargetBox-1="[397,1197,1074,1657]" captionTargetId-0="figure@19.[157,1431,310,910]" captionTargetId-1="figure@19.[355,1233,1046,1674]" captionTargetPageId-0="19" captionTargetPageId-1="19" captionText-0="FIGURE 10. P. intermedia (Tryon). A – D. Shells. Scale = 1.0 mm. A. JFBM 20507. B. USNM 1107071. C. USNM 1107076. D. USNM 1075362." captionText-1="FIGURE 11. P. intermedia (Tryon). A – B. Penis, USNM 1075362. Scale bar = 0.5 mm. A. Dorsal surface. B. Ventral surface. Pd = penial duct, Pf = penial filament, Pg = penial gland, Pl = penial lobe, Tg = terminal gland, Vg = ventral gland." httpUri-0="https://zenodo.org/record/185694/files/figure.png" httpUri-1="https://zenodo.org/record/185695/files/figure.png" pageId="18" pageNumber="19">Figs 10–11</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C31248CADC58FFD26AFBFE2CFA77FDC7" pageId="18" pageNumber="19" type="reference_group">
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<paragraph id="8BB71B41DC58FFD26AFBFE2CFE1CFDA7" blockId="18.[151,1436,500,587]" pageId="18" pageNumber="19">
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<treatmentCitationGroup id="AB183C6FDC58FFD26AFBFE2CFE1CFDA7" pageId="18" pageNumber="19">
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<taxonomicName id="4C0860C2DC58FFD26AFBFE2CFDBCFD87" authority="Tryon, 1865: 220" authorityName="Tryon" authorityPageNumber="220" authorityYear="1865" box="[151,599,500,523]" class="Gastropoda" family="Pomatiopsidae" genus="Pomatiopsis" kingdom="Animalia" order="Littorinimorpha" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="intermedia">
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<emphasis id="B97CC753DC58FFD26AFBFE2CFE7DFD87" box="[151,406,500,523]" italics="true" pageId="18" pageNumber="19">Pomatiopsis intermedia</emphasis>
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<treatmentCitation id="0AA93D50DC58FFD26BF1FE2CFDBCFD87" author="Tryon" box="[413,599,500,523]" page="220" pageId="18" pageNumber="19" year="1865">
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<bibRefCitation id="EF9966B0DC58FFD26BF1FE2CFDCFFD87" author="Tryon" box="[413,548,500,523]" pageId="18" pageNumber="22" refString="Tryon, G. W. (1865) Descriptions of new species of Amnicola, Pomatiopsis, Somatogyrus, Gabbia, Hydrobia and Rissoa. American Journal of Conchology, 1, 219 - 220." type="journal article" year="1865">Tryon, 1865</bibRefCitation>
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: 220
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</treatmentCitation>
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</taxonomicName>
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[
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<typeStatus id="54B3A5E3DC58FFD26809FE2CFD7DFD87" box="[613,662,500,523]" pageId="18" pageNumber="19">type</typeStatus>
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locality, Owyhee R., S. E. Oregon; original label states Crooked Creek, Owyhee, southeast Oregon].
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BB71B41DC58FFD26AFBFDECFA77FDC7" blockId="18.[151,1436,500,587]" box="[151,1436,564,587]" pageId="18" pageNumber="19">
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<treatmentCitationGroup id="AB183C6FDC58FFD26AFBFDECFA77FDC7" box="[151,1436,564,587]" pageId="18" pageNumber="19">
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<taxonomicName id="4C0860C2DC58FFD26AFBFDECFDCCFDC7" ID-CoL="7SYTL" authority="Tryon, 1865" authorityName="Tryon" authorityYear="1865" box="[151,551,564,587]" class="Gastropoda" family="Hydrobiidae" genus="Pyrgulopsis" kingdom="Animalia" order="Littorinimorpha" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="intermedia">
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<emphasis id="B97CC753DC58FFD26AFBFDECFE64FDC7" box="[151,399,564,587]" italics="true" pageId="18" pageNumber="19">Pyrgulopsis intermedia</emphasis>
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(
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<bibRefCitation id="EF9966B0DC58FFD26BF1FDECFDCBFDC7" author="Tryon" box="[413,544,564,587]" pageId="18" pageNumber="22" refString="Tryon, G. W. (1865) Descriptions of new species of Amnicola, Pomatiopsis, Somatogyrus, Gabbia, Hydrobia and Rissoa. American Journal of Conchology, 1, 219 - 220." type="journal article" year="1865">Tryon, 1865</bibRefCitation>
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)
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</taxonomicName>
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.—
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<treatmentCitation id="0AA93D50DC58FFD2682BFDECFCE5FDC7" author="Hershler" box="[583,782,564,587]" page="42" pageId="18" pageNumber="19" year="1994">
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<bibRefCitation id="EF9966B0DC58FFD2682BFDECFD0CFDC7" author="Hershler" box="[583,743,564,587]" pageId="18" pageNumber="21" refString="Hershler, R. (1994) A review of the North American freshwater snail genus Pyrgulopsis. Smithsonian Contributions to Zoology, 554, 1 - 115." type="journal article" year="1994">Hershler 1994</bibRefCitation>
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: 42
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</treatmentCitation>
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, 44, figs 17a–d, 36c, 47b (synonymy, diagnosis, description).
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31248CADC58FFD26AFBFDA2FDF4FC2F" pageId="18" pageNumber="19" type="materials_examined">
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<paragraph id="8BB71B41DC58FFD26AFBFDA2FDF4FC2F" blockId="18.[151,1437,634,2014]" pageId="18" pageNumber="19">
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<emphasis id="B97CC753DC58FFD26AFBFDA2FE91FD18" bold="true" box="[151,378,634,660]" pageId="18" pageNumber="19">Referred material</emphasis>
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. OREGON. Malheur County.
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<collectionCode id="ED198384DC58FFD26888FDA2FCC5FD18" box="[740,814,634,660]" httpUri="http://grbio.org/cool/v57z-1dzw" name="James Ford Bell Museum of Natural History" pageId="18" pageNumber="19">JFBM</collectionCode>
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20497,
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<collectionCode id="ED198384DC58FFD269E3FDA2FC32FD18" box="[911,985,634,660]" httpUri="http://grbio.org/cool/v57z-1dzw" name="James Ford Bell Museum of Natural History" pageId="18" pageNumber="19">JFBM</collectionCode>
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20507, Crooked Creek Spring, ca.
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<quantity id="4CF0B6A4DC58FFD26F12FDA2FF56FD37" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="18" pageNumber="19" unit="km" value="16.0">16 km</quantity>
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southwest of Burns Junction, T. 33 S, R. 39 E, sec. 22,
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<date id="FFB63D81DC58FFD2690AFD79FC08FD37" box="[870,995,673,699]" pageId="18" pageNumber="19" value="1963-05-23">23/v/1963</date>
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,
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<date id="FFB63D81DC58FFD2699CFD79FB86FD37" box="[1008,1133,673,699]" pageId="18" pageNumber="19" value="1982-05-19">19/v/1982</date>
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.
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<collectionCode id="ED198384DC58FFD26E16FD79FB3BFD37" LSID="urn:lsid:biocol.org:col:34871" box="[1146,1232,673,699]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="18" pageNumber="19">USNM</collectionCode>
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1075362, spring tributary to Birch Creek, 459964 E, 4781865 N,
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<date id="FFB63D81DC58FFD268B3FD10FC8FFD6E" box="[735,868,712,738]" pageId="18" pageNumber="19" value="2003-04-25">25/iv/2003</date>
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.
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<collectionCode id="ED198384DC58FFD2691CFD10FC2DFD6E" LSID="urn:lsid:biocol.org:col:34871" box="[880,966,712,738]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="18" pageNumber="19">USNM</collectionCode>
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1075684,
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<emphasis id="B97CC753DC58FFD26E2AFD10FB9FFD6D" box="[1094,1140,712,737]" italics="true" pageId="18" pageNumber="19">ibid</emphasis>
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., 459871 E, 4782621 N,
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<date id="FFB63D81DC58FFD26AFBFD36FEF0FC84" box="[151,283,750,776]" pageId="18" pageNumber="19" value="2003-06-25">25/vi/2003</date>
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.
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<collectionCode id="ED198384DC58FFD26B4AFD36FE90FC84" LSID="urn:lsid:biocol.org:col:34871" box="[294,379,750,776]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="18" pageNumber="19">USNM</collectionCode>
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1092855,
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<emphasis id="B97CC753DC58FFD26B96FD37FDC3FC84" box="[506,552,751,776]" italics="true" pageId="18" pageNumber="19">ibid</emphasis>
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., 460058 E, 4781808 N,
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<date id="FFB63D81DC58FFD26939FD36FC0AFC84" box="[853,993,750,776]" pageId="18" pageNumber="19" value="2006-07-18">18/vii/2006</date>
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.
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<collectionCode id="ED198384DC58FFD26980FD36FBAAFC84" LSID="urn:lsid:biocol.org:col:34871" box="[1004,1089,750,776]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="18" pageNumber="19">USNM</collectionCode>
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1107071, spring on east side Owyhee River, above Long Sweetwater rapids, 442355 E, 4753770 N,
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<date id="FFB63D81DC58FFD269B1FCCDFBB9FCA3" box="[989,1106,789,815]" pageId="18" pageNumber="19" value="2007-09-06">6/ix/2007</date>
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.
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<collectionCode id="ED198384DC58FFD26E30FCCDFB5AFCA3" LSID="urn:lsid:biocol.org:col:34871" box="[1116,1201,789,815]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="18" pageNumber="19">USNM</collectionCode>
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1107076, spring on hillside, Jackson Hole, 447566 E, 4781838 N,
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<date id="FFB63D81DC58FFD268AFFCE4FCD2FCDA" box="[707,825,828,854]" pageId="18" pageNumber="19" value="2007-09-08">8/ix/2007</date>
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.
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<collectionCode id="ED198384DC58FFD26929FCE4FC71FCDA" LSID="urn:lsid:biocol.org:col:34871" box="[837,922,828,854]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="18" pageNumber="19">USNM</collectionCode>
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1107079, spring west of Two Mile Spring, 453320 E, 4782496 N,
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<date id="FFB63D81DC58FFD26BDFFCBAFDC1FCF0" box="[435,554,866,892]" pageId="18" pageNumber="19" value="2007-09-09">9/ix/2007</date>
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.
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<collectionCode id="ED198384DC58FFD2685BFCBAFD66FCF0" LSID="urn:lsid:biocol.org:col:34871" box="[567,653,866,892]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="18" pageNumber="19">USNM</collectionCode>
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1107077, mouth of Rinehart Creek, west side of Owyhee River, 449007 E, 4783611 N,
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<date id="FFB63D81DC58FFD26BCBFC51FDF7FC2F" box="[423,540,905,931]" pageId="18" pageNumber="19" value="2007-09-09">9/ix/2007</date>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31248CADC58FFD26AA9FC77FC6EFBB2" pageId="18" pageNumber="19" type="distribution">
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<paragraph id="8BB71B41DC58FFD26AA9FC77FC6EFBB2" blockId="18.[151,1437,634,2014]" pageId="18" pageNumber="19">
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<emphasis id="B97CC753DC58FFD26AA9FC77FE14FC45" bold="true" box="[197,511,943,969]" pageId="18" pageNumber="19">Distribution and habitat.</emphasis>
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Crooked Creek (Owyhee River basin), Barren Valley (Great Basin), plus new records from the Crooked Creek headspring and along the lower Owyhee River, southeastern Oregon (
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<figureCitation id="133307C4DC58FFD26F29FC0EFA61FC7C" box="[1349,1418,982,1008]" captionStart="FIGURE 1" captionStartId="2.[151,255,1934,1958]" captionTargetBox="[274,1314,197,1897]" captionTargetId="figure@2.[274,1315,197,1898]" captionTargetPageId="2" captionText="FIGURE 1. Map showing the distributions of Pyrgulopsis species in southeastern Oregon. Arrows indicate previously documented localities; all others are newly reported herein. Some of the symbols refer to multiple, closely proximal localities." httpUri="https://zenodo.org/record/185685/files/figure.png" pageId="18" pageNumber="19">Fig. 1</figureCitation>
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). This species lives in springs and small streams. It was collected in association with
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<taxonomicName id="4C0860C2DC58FFD26E88FC25FEC5FBB1" class="Gastropoda" family="Hydrobiidae" genus="Potamopyrgus" kingdom="Animalia" order="Littorinimorpha" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="antipodarum">
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<emphasis id="B97CC753DC58FFD26E88FC25FEC5FBB1" italics="true" pageId="18" pageNumber="19">Potamopyrgus antipodarum</emphasis>
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</taxonomicName>
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(USNM 1107069) at the mouth of Rinehart Creek.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31248CADC58FFD36AA9FB92FCE5F833" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19" type="discussion">
|
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<paragraph id="8BB71B41DC58FFD26AA9FB92FD4EF8CF" blockId="18.[151,1437,634,2014]" pageId="18" pageNumber="19">
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<emphasis id="B97CC753DC58FFD26AA9FB92FED1FBE8" bold="true" box="[197,314,1098,1124]" pageId="18" pageNumber="19">Remarks</emphasis>
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. In an unpublished checklist,
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<bibRefCitation id="EF9966B0DC58FFD268C0FB92FCBFFBE8" author="Taylor" box="[684,852,1098,1124]" pageId="18" pageNumber="22" refString="Taylor, D. W. (1977) Rocky Mountain and intermountain freshwater molluscs: an annotated list. Unpublished mss, 40 pp." type="book" year="1977">Taylor (1977)</bibRefCitation>
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contended that the
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<typeStatus id="54B3A5E3DC58FFD26E2AFB92FB91FBE8" box="[1094,1146,1098,1124]" pageId="18" pageNumber="19">type</typeStatus>
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locality of
|
||
<taxonomicName id="4C0860C2DC58FFD26F65FB93FEFDFB06" class="Gastropoda" family="Hydrobiidae" genus="Fontelicella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="intermedia">
|
||
<emphasis id="B97CC753DC58FFD26F65FB93FEFDFB06" italics="true" pageId="18" pageNumber="19">Fontelicella intermedia</emphasis>
|
||
</taxonomicName>
|
||
(=
|
||
<taxonomicName id="4C0860C2DC58FFD26B52FBA9FDA4FB06" box="[318,591,1137,1162]" class="Gastropoda" family="Hydrobiidae" genus="Pyrgulopsis" kingdom="Animalia" order="Littorinimorpha" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="intermedia">
|
||
<emphasis id="B97CC753DC58FFD26B52FBA9FDA4FB06" box="[318,591,1137,1162]" italics="true" pageId="18" pageNumber="19">Pyrgulopsis intermedia</emphasis>
|
||
</taxonomicName>
|
||
) may have been incorrectly attributed to Crooked Creek (Owyhee River drainage) instead of Crooked Creek (Deschutes River drainage) and consequently provided a new species name,
|
||
<taxonomicName id="4C0860C2DC58FFD26A8AFB67FE13FB54" box="[230,504,1215,1240]" class="Gastropoda" family="Hydrobiidae" genus="Fontelicella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="malhorica">
|
||
<emphasis id="B97CC753DC58FFD26A8AFB67FE13FB54" box="[230,504,1215,1240]" italics="true" pageId="18" pageNumber="19">Fontelicella malhorica</emphasis>
|
||
</taxonomicName>
|
||
, for the congener that lives in the headspring of the former stream. (Note that Taylor apparently changed his mind on this subject as he later stated that
|
||
<taxonomicName id="4C0860C2DC58FFD26E48FB3DFB23FB72" box="[1060,1224,1253,1278]" class="Gastropoda" family="Hydrobiidae" genus="Pyrgulopsis" kingdom="Animalia" order="Littorinimorpha" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="intermedia">
|
||
<emphasis id="B97CC753DC58FFD26E48FB3DFB23FB72" box="[1060,1224,1253,1278]" italics="true" pageId="18" pageNumber="19">P. intermedia</emphasis>
|
||
</taxonomicName>
|
||
lives in Crooked Creek, Owyhee River drainage [
|
||
<bibRefCitation id="EF9966B0DC58FFD2685FFAD4FCB8FAAA" author="Taylor" box="[563,851,1292,1318]" pageId="18" pageNumber="22" refString="Taylor, D. W. (1985) Evolution of freshwater drainages and molluscs in western North America. In: Smiley, C. J. & Leviton, A. J. (Eds), Late Cenozoic History of the Pacific Northwest, Interdisciplinary Studies on the Clarkia Fossil Beds of Northern Idaho. American Association for the Advancement of Science, San Francisco, pp. 265 - 321." type="book chapter" year="1985">Taylor 1985:309–310</bibRefCitation>
|
||
].) In an unpublished report,
|
||
<bibRefCitation id="EF9966B0DC58FFD26ED2FAD4FEB8FAC0" author="Frest" pageId="18" pageNumber="21" refString="Frest, T. J. & Johannes, E. J. (1995) Interior Columbia Basin mollusk species of special concern. Final Report to the Interior Columbia Basin Ecosystem Management Project, Walla Walla, Washington, Contract # 43 - OEOO- 4 - 9112, xi + 274 pp. Available from http: // www. icbemp. gov / science / frest _ 1. pdf (accessed 10 September 2008)." type="book" year="1995">Frest & Johannes (1995:198, 202)</bibRefCitation>
|
||
listed this
|
||
<emphasis id="B97CC753DC58FFD26BBEFAEBFD9CFAC0" box="[466,631,1331,1356]" italics="true" pageId="18" pageNumber="19">nomen nudum</emphasis>
|
||
(to which they applied a common name, “Malheur springsnail”) as a different species than
|
||
<taxonomicName id="4C0860C2DC58FFD26BCEFA81FDA9FAFE" box="[418,578,1369,1394]" class="Gastropoda" family="Hydrobiidae" genus="Pyrgulopsis" kingdom="Animalia" order="Littorinimorpha" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="intermedia">
|
||
<emphasis id="B97CC753DC58FFD26BCEFA81FDA9FAFE" box="[418,578,1369,1394]" italics="true" pageId="18" pageNumber="19">P. intermedia</emphasis>
|
||
</taxonomicName>
|
||
and indicated that both of these snails live in Crooked Creek, Owyhee River drainage.
|
||
<taxonomicName id="4C0860C2DC58FFD26B3FFA58FD8FFA15" box="[339,612,1408,1433]" class="Gastropoda" family="Hydrobiidae" genus="Pyrgulopsis" kingdom="Animalia" order="Littorinimorpha" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="intermedia">
|
||
<emphasis id="B97CC753DC58FFD26B3FFA58FD8FFA15" box="[339,612,1408,1433]" italics="true" pageId="18" pageNumber="19">Pyrgulopsis intermedia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B97CC753DC58FFD268F0FA58FD40FA15" box="[668,683,1408,1433]" italics="true" pageId="18" pageNumber="19">P</emphasis>
|
||
. sp. nov. (Malheur springsnail) have subsequently been listed in various conservation-related documents and publications (cited in the Introduction; also see
|
||
<bibRefCitation id="EF9966B0DC58FFD26E9FFA7EFF32FA6B" author="Wisdom" pageId="18" pageNumber="22" refString="Wisdom, M. J., Rowland, M. M., Suring, L. H., Schueck, L., Wolff Meinke, C. & Wales, B. C. (2003) Procedures for regional assessment of habitats for species of conservation concern in the sagebrush ecosystem. March 2003 report, Version 1. Pacific Northwest Research Station, La Grande, Oregon. 153 pp. Available from http: // www. fs. fed. us / pnw / lagrande / sagebrush / docs / Procedures % 20 for % 20 Regional % 20 Asessment /" type="book" year="2003">
|
||
Wisdom
|
||
<emphasis id="B97CC753DC58FFD26F33FA7FFA7FFA4C" box="[1375,1428,1447,1472]" italics="true" pageId="18" pageNumber="19">et al</emphasis>
|
||
. 2003
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9966B0DC58FFD26A8AFA15FE24FA6B" author="Brown" box="[230,463,1485,1511]" pageId="18" pageNumber="21" refString="Brown J., Wyers, A., Aldous A. & Bach, L. (2007) Groundwater and biodiversity conservation: a methods guide for integrating groundwater needs of ecosystems into conservation plans in the Pacific Northwest. The Nature Conservancy. 123 pp + appendices A-G. Available from http: // www. waconservation. org / data / collins / GroundwaterMethodsGuideTNC _ Jan 08. pdf (accessed 10 September 2008)." type="book" year="2007">
|
||
Brown
|
||
<emphasis id="B97CC753DC58FFD26B2FFA15FE91FA6A" box="[323,378,1485,1510]" italics="true" pageId="18" pageNumber="19">et al</emphasis>
|
||
. 2007
|
||
</bibRefCitation>
|
||
). Taylor’s (1977) manuscript name is not valid because his checklist does not constitute published work under ICZN Articles 8.1 and 9.7 and the only subsequent citation (
|
||
<bibRefCitation id="EF9966B0DC58FFD26EBCFA2CFF38F9B9" author="Frest" pageId="18" pageNumber="21" refString="Frest, T. J. & Johannes, E. J. (1995) Interior Columbia Basin mollusk species of special concern. Final Report to the Interior Columbia Basin Ecosystem Management Project, Walla Walla, Washington, Contract # 43 - OEOO- 4 - 9112, xi + 274 pp. Available from http: // www. icbemp. gov / science / frest _ 1. pdf (accessed 10 September 2008)." type="book" year="1995">Frest & Johannes 1995</bibRefCitation>
|
||
) was also in an unpublished venue. (Also note in this context that the name was not accompanied by a description or definition.) There is no reason to suspect that
|
||
<bibRefCitation id="EF9966B0DC58FFD26930F999FC12F9D7" author="Tryon" box="[860,1017,1601,1627]" pageId="18" pageNumber="22" refString="Tryon, G. W. (1865) Descriptions of new species of Amnicola, Pomatiopsis, Somatogyrus, Gabbia, Hydrobia and Rissoa. American Journal of Conchology, 1, 219 - 220." type="journal article" year="1865">Tryon (1865)</bibRefCitation>
|
||
made an error in reporting the
|
||
<typeStatus id="54B3A5E3DC58FFD26F05F999FA77F9D7" box="[1385,1436,1601,1627]" pageId="18" pageNumber="19">type</typeStatus>
|
||
locality of
|
||
<taxonomicName id="4C0860C2DC58FFD26B70F9B0FE54F90D" box="[284,447,1640,1665]" class="Gastropoda" family="Hydrobiidae" genus="Pyrgulopsis" kingdom="Animalia" order="Littorinimorpha" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="intermedia">
|
||
<emphasis id="B97CC753DC58FFD26B70F9B0FE54F90D" box="[284,447,1640,1665]" italics="true" pageId="18" pageNumber="19">P. intermedia</emphasis>
|
||
</taxonomicName>
|
||
as his
|
||
<typeStatus id="54B3A5E3DC58FFD26879F9B0FDBEF90E" box="[533,597,1640,1666]" pageId="18" pageNumber="19">types</typeStatus>
|
||
well conform to snails now living in Crooked Creek, Owyhee River drainage (
|
||
<bibRefCitation id="EF9966B0DC58FFD26B7CF956FE2DF924" author="Hershler" box="[272,454,1678,1704]" pageId="18" pageNumber="21" refString="Hershler, R. (1994) A review of the North American freshwater snail genus Pyrgulopsis. Smithsonian Contributions to Zoology, 554, 1 - 115." type="journal article" year="1994">Hershler 1994</bibRefCitation>
|
||
). Snails (collected by Taylor) from Crooked Creek Spring (JFBM 20497, JFBM 20507,
|
||
<figureCitation id="133307C4DC58FFD26A99F96DFEA6F943" box="[245,333,1717,1743]" captionStart="FIGURE 10. P" captionStartId="19.[151,257,933,957]" captionTargetBox="[157,1430,310,909]" captionTargetId="figure@19.[157,1431,310,910]" captionTargetPageId="19" captionText="FIGURE 10. P. intermedia (Tryon). A – D. Shells. Scale = 1.0 mm. A. JFBM 20507. B. USNM 1107071. C. USNM 1107076. D. USNM 1075362." httpUri="https://zenodo.org/record/185694/files/figure.png" pageId="18" pageNumber="19">Fig. 10</figureCitation>
|
||
A) conform morphologically to
|
||
<taxonomicName id="4C0860C2DC58FFD268B1F96DFC68F942" box="[733,899,1717,1742]" class="Gastropoda" family="Hydrobiidae" genus="Pyrgulopsis" kingdom="Animalia" order="Littorinimorpha" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="intermedia">
|
||
<emphasis id="B97CC753DC58FFD268B1F96DFC68F942" box="[733,899,1717,1742]" italics="true" pageId="18" pageNumber="19">P. intermedia</emphasis>
|
||
</taxonomicName>
|
||
in all respects and our genetic studies also suggest that only one congener lives in Crooked Creek. Thus, there is no basis for recognizing the Malheur springsnail as a validly described and/or distinct species and this name therefore should be removed from conservation watch lists and other databases.
|
||
</paragraph>
|
||
<paragraph id="8BB71B41DC58FFD36AA9F888FCC1FF73" blockId="18.[151,1437,634,2014]" lastBlockId="19.[151,1436,152,255]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
|
||
Snails from the newly reported localities are referable to
|
||
<taxonomicName id="4C0860C2DC58FFD26937F888FC1DF8E5" box="[859,1014,1872,1897]" class="Gastropoda" family="Hydrobiidae" genus="Pyrgulopsis" kingdom="Animalia" order="Littorinimorpha" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="intermedia">
|
||
<emphasis id="B97CC753DC58FFD26937F888FC1DF8E5" box="[859,1014,1872,1897]" italics="true" pageId="18" pageNumber="19">P. intermedia</emphasis>
|
||
</taxonomicName>
|
||
based on their relatively large (>
|
||
<quantity id="4CF0B6A4DC58FFD26F1AF888FF21F81C" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="18" pageNumber="19" unit="mm" value="3.3">3.3 mm</quantity>
|
||
), solid, ovate conic shell (
|
||
<figureCitation id="133307C4DC58FFD2686BF8AEFDB7F81C" box="[519,604,1910,1936]" captionStart="FIGURE 10. P" captionStartId="19.[151,257,933,957]" captionTargetBox="[157,1430,310,909]" captionTargetId="figure@19.[157,1431,310,910]" captionTargetPageId="19" captionText="FIGURE 10. P. intermedia (Tryon). A – D. Shells. Scale = 1.0 mm. A. JFBM 20507. B. USNM 1107071. C. USNM 1107076. D. USNM 1075362." httpUri="https://zenodo.org/record/185694/files/figure.png" pageId="18" pageNumber="19">Fig. 10</figureCitation>
|
||
A–D) with prominent spire; and details of penis shape and glandular ornament (
|
||
<figureCitation id="133307C4DC58FFD26B7AF845FE83F83B" box="[278,360,1949,1975]" captionStart="FIGURE 11. P" captionStartId="19.[151,256,1698,1722]" captionTargetBox="[397,1197,1074,1657]" captionTargetId="figure@19.[355,1233,1046,1674]" captionTargetPageId="19" captionText="FIGURE 11. P. intermedia (Tryon). A – B. Penis, USNM 1075362. Scale bar = 0.5 mm. A. Dorsal surface. B. Ventral surface. Pd = penial duct, Pf = penial filament, Pg = penial gland, Pl = penial lobe, Tg = terminal gland, Vg = ventral gland." httpUri="https://zenodo.org/record/185695/files/figure.png" pageId="18" pageNumber="19">Fig. 11</figureCitation>
|
||
A–B), particularly the short penial filament and medially positioned ventral gland. Samples from Crooked Creek (IP60) and the lower Owyhee River drainage (IP67, P217, P222, P223, P224) that were newly sequenced for this project were closely similar (0.2% divergence) to previously analyzed specimens from Crooked Creek (P1, P2) and Barren Valley (P4). The mean uncorrected sequence divergence within
|
||
<taxonomicName id="4C0860C2DC59FFD36FEAFF67FEFDFF72" class="Gastropoda" family="Hydrobiidae" genus="Pyrgulopsis" kingdom="Animalia" order="Littorinimorpha" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="intermedia">
|
||
<emphasis id="B97CC753DC59FFD36FEAFF67FEFDFF72" italics="true" pageId="19" pageNumber="20">P. intermedia</emphasis>
|
||
</taxonomicName>
|
||
was quite low (0.2%), ranging from 0–0.8%.
|
||
</paragraph>
|
||
<caption id="DF774BC9DC59FFD36AFBFC7DFE32FC6C" httpUri="https://zenodo.org/record/185694/files/figure.png" pageId="19" pageNumber="20" targetBox="[157,1430,310,909]" targetPageId="19">
|
||
<paragraph id="8BB71B41DC59FFD36AFBFC7DFE32FC6C" blockId="19.[151,1436,933,992]" pageId="19" pageNumber="20">
|
||
<emphasis id="B97CC753DC59FFD36AFBFC7DFECFFC31" bold="true" box="[151,292,933,957]" pageId="19" pageNumber="20">FIGURE 10</emphasis>
|
||
.
|
||
<taxonomicName id="4C0860C2DC59FFD36B5FFC7EFDCCFC31" authority="Tryon" authorityName="Tryon" box="[307,551,934,957]" class="Gastropoda" family="Hydrobiidae" genus="Pyrgulopsis" kingdom="Animalia" order="Littorinimorpha" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="intermedia">
|
||
<emphasis id="B97CC753DC59FFD36B5FFC7EFEA3FC31" box="[307,328,934,957]" italics="true" pageId="19" pageNumber="20">P.</emphasis>
|
||
<emphasis id="B97CC753DC59FFD36B23FC7EFE2CFC31" box="[335,455,934,957]" italics="true" pageId="19" pageNumber="20">intermedia</emphasis>
|
||
(Tryon)
|
||
</taxonomicName>
|
||
. A–D. Shells. Scale = 1.0 mm. A. JFBM 20507. B. USNM 1107071. C. USNM 1107076. D. USNM 1075362.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF774BC9DC59FFD36AFBF97AFF32F973" httpUri="https://zenodo.org/record/185695/files/figure.png" pageId="19" pageNumber="20" targetBox="[397,1197,1074,1657]" targetPageId="19">
|
||
<paragraph id="8BB71B41DC59FFD36AFBF97AFF32F973" blockId="19.[151,1436,1698,1791]" pageId="19" pageNumber="20">
|
||
<emphasis id="B97CC753DC59FFD36AFBF97AFECAF936" bold="true" box="[151,289,1698,1722]" pageId="19" pageNumber="20">FIGURE 11</emphasis>
|
||
.
|
||
<taxonomicName id="4C0860C2DC59FFD36B43F97BFDCBF936" authority="Tryon" authorityName="Tryon" box="[303,544,1699,1722]" class="Gastropoda" family="Hydrobiidae" genus="Pyrgulopsis" kingdom="Animalia" order="Littorinimorpha" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="intermedia">
|
||
<emphasis id="B97CC753DC59FFD36B43F97BFEAFF936" box="[303,324,1699,1722]" italics="true" pageId="19" pageNumber="20">P.</emphasis>
|
||
<emphasis id="B97CC753DC59FFD36B27F97BFE2AF936" box="[331,449,1699,1722]" italics="true" pageId="19" pageNumber="20">intermedia</emphasis>
|
||
(Tryon)
|
||
</taxonomicName>
|
||
. A–B. Penis, USNM 1075362. Scale bar = 0.5 mm. A. Dorsal surface. B. Ventral surface. Pd = penial duct, Pf = penial filament, Pg = penial gland, Pl = penial lobe, Tg = terminal gland, Vg = ventral gland.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB71B41DC59FFD36AA9F880FCE5F833" blockId="19.[151,1436,1880,1983]" pageId="19" pageNumber="20">
|
||
The new records detailed herein extend the range of
|
||
<taxonomicName id="4C0860C2DC59FFD36955F880FC3CF8FD" box="[825,983,1880,1905]" class="Gastropoda" family="Hydrobiidae" genus="Pyrgulopsis" kingdom="Animalia" order="Littorinimorpha" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="intermedia">
|
||
<emphasis id="B97CC753DC59FFD36955F880FC3CF8FD" box="[825,983,1880,1905]" italics="true" pageId="19" pageNumber="20">P. intermedia</emphasis>
|
||
</taxonomicName>
|
||
about
|
||
<quantity id="4CF0B6A4DC59FFD36E45F880FB9EF8FE" box="[1065,1141,1880,1906]" metricMagnitude="4" metricUnit="m" metricValue="6.5" pageId="19" pageNumber="20" unit="km" value="65.0">65 km</quantity>
|
||
north of its
|
||
<typeStatus id="54B3A5E3DC59FFD36F64F880FAD0F8FE" box="[1288,1339,1880,1906]" pageId="19" pageNumber="20">type</typeStatus>
|
||
locality area into the lower Owyhee River basin. It may be appropriate to reconsider the conservation status of this species in the light of this significant range extension.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |