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<document id="95F2CFF27CF89E67CE95E0EC1E64C48F" ID-DOI="10.11646/zootaxa.5497.3.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="13618611" ID-ZooBank="C107D617-5F70-42C5-8FD4-E5C2DE0E48B1" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725026357643" checkinUser="plazi" docAuthor="Khaustov, Alexander A., Frolov, Andrey V. &amp; Akhmetova, Lilia A." docDate="2024" docId="03EB878FFFAF745E81B43CEEFB9DFE1A" docLanguage="en" docName="zootaxa.5497.3.2.pdf" docOrigin="Zootaxa 5497 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5497.3.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Spatulaphorus spinisetus Khaustov, Frolov &amp; Akhmetova, 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="350" masterDocId="FFD2FFF7FFA6745381233D58FFDEFFEF" masterDocTitle="Seven new species of Spatulaphorus Rack (Acari: Pygmephoridae) phoretic on scarab beetles (Coleoptera: Scarabaeidae)" masterLastPageNumber="368" masterPageNumber="337" pageNumber="346" updateTime="1725028125811" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="D7357C7C0F04AEEC3FA4515DFD826D8C">Seven new species of Spatulaphorus Rack (Acari: Pygmephoridae) phoretic on scarab beetles (Coleoptera: Scarabaeidae)</mods:title>
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<mods:namePart id="4D520273EDA3C106BBE3D5C6E6CEC678">Khaustov, Alexander A.</mods:namePart>
<mods:affiliation id="69554FAF2B6DF1774D360B59DEB281D8">Tyumen State University, Tyumen, 6 Volodarskogo Str., 625003 Russia.</mods:affiliation>
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<mods:namePart id="AC717FC99824E36AB7544F905A31BC89">Frolov, Andrey V.</mods:namePart>
<mods:affiliation id="C94692745CE2F6DBE1E31F46A7725B09">Zoological Institute of Russian Academy of Sciences, Universitetskaya Embankment 1, 199034, Saint Petersburg, Russia</mods:affiliation>
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<mods:namePart id="4ED167C04F1BBA2094F1C58BB3D27ED9">Akhmetova, Lilia A.</mods:namePart>
<mods:affiliation id="6ED7F3EBE8685E37723C6636B4DAE108">Zoological Institute of Russian Academy of Sciences, Universitetskaya Embankment 1, 199034, Saint Petersburg, Russia</mods:affiliation>
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<mods:title id="E86C73B5CB1106A61AD07FB300DA8651">Zootaxa</mods:title>
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<mods:date id="CB42F58C50E314100DAB6ACB50180447">2024</mods:date>
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<mods:number id="361B24C94F8E0C8BEE8A17E89043F120">2024-08-26</mods:number>
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<mods:number id="F5B65FF9814557616444D8572B8A6476">5497</mods:number>
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<mods:number id="C45B7EA9160CD7A1DC5502FDC09AF949">3</mods:number>
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<treatment id="03EB878FFFAF745E81B43CEEFB9DFE1A" LSID="urn:lsid:plazi:treatment:03EB878FFFAF745E81B43CEEFB9DFE1A" httpUri="http://treatment.plazi.org/id/03EB878FFFAF745E81B43CEEFB9DFE1A" lastPageId="13" lastPageNumber="350" pageId="9" pageNumber="346">
<subSubSection id="C3586512FFAF745A81B43CEEFDF2FE3E" box="[151,556,438,465]" pageId="9" pageNumber="346" type="nomenclature">
<paragraph id="8BFD3699FFAF745A81B43CEEFDF2FE3E" blockId="9.[151,556,438,501]" box="[151,556,438,465]" pageId="9" pageNumber="346">
<heading id="D0B581F5FFAF745A81B43CEEFDF2FE3E" bold="true" box="[151,556,438,465]" fontSize="11" level="1" pageId="9" pageNumber="346" reason="1">
<emphasis id="B936EA8BFFAF745A81B43CEEFDF2FE3E" bold="true" box="[151,556,438,465]" pageId="9" pageNumber="346">
<taxonomicName id="4C424D1AFFAF745A81B43CEEFE19FE3E" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[151,455,438,465]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="9" pageNumber="346" phylum="Arthropoda" rank="species" species="spinisetus" status="sp. nov.">
<emphasis id="B936EA8BFFAF745A81B43CEEFE19FE3E" bold="true" box="[151,455,438,465]" italics="true" pageId="9" pageNumber="346">Spatulaphorus spinisetus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20557F0FFAF745A80EC3CEFFDF2FE3E" box="[463,556,439,465]" pageId="9" pageNumber="346" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3586512FFAF745F81B43C83FE4FF9FC" lastPageId="12" lastPageNumber="349" pageId="9" pageNumber="346" type="description">
<paragraph id="8BFD3699FFAF745A81B43C83FEC7FE1A" blockId="9.[151,556,438,501]" box="[151,281,475,501]" pageId="9" pageNumber="346">
(
<figureCitation id="13792A1CFFAF745A81BC3C83FECFFE1A" box="[159,273,475,501]" captionStart-0="FIGURE 9" captionStart-1="FIGURE 10" captionStart-2="FIGURE 11" captionStart-3="FIGURE 12" captionStartId-0="9.[151,250,1795,1819]" captionStartId-1="10.[151,250,1311,1335]" captionStartId-2="11.[151,250,1860,1884]" captionStartId-3="12.[151,250,1422,1446]" captionTargetBox-0="[151,1435,610,1772]" captionTargetBox-1="[151,1435,241,1288]" captionTargetBox-2="[151,1418,190,1836]" captionTargetBox-3="[151,1436,633,1399]" captionTargetId-0="figure-146@9.[151,1435,597,1772]" captionTargetId-1="figure-50@10.[151,1435,237,1288]" captionTargetId-2="figure-17@11.[151,1436,181,1836]" captionTargetId-3="figure-287@12.[151,1436,633,1399]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="11" captionTargetPageId-3="12" captionText-0="FIGURE 9. Spatulaphorus spinisetus sp. nov., female: A—dorsum of body; B—venter of body. Legs omitted." captionText-1="FIGURE 10. Spatulaphorus spinisetus sp. nov., female: A—right leg I, dorsal aspect; B—right leg II, dorsal aspect." captionText-2="FIGURE 11. Spatulaphorus spinisetus sp. nov., female: A—right leg III, dorsal aspect; B—right leg IV, dorsal aspect." captionText-3="FIGURE 12. DIC micrographs of Spatulaphorus spinisetus sp. nov., female: A—general view dorsally; B—general view ventrally." figureDoi-0="http://doi.org/10.5281/zenodo.13618629" figureDoi-1="http://doi.org/10.5281/zenodo.13618631" figureDoi-2="http://doi.org/10.5281/zenodo.13618637" figureDoi-3="http://doi.org/10.5281/zenodo.13618639" httpUri-0="https://zenodo.org/record/13618629/files/figure.png" httpUri-1="https://zenodo.org/record/13618631/files/figure.png" httpUri-2="https://zenodo.org/record/13618637/files/figure.png" httpUri-3="https://zenodo.org/record/13618639/files/figure.png" pageId="9" pageNumber="346">Figs 912</figureCitation>
)
</paragraph>
<paragraph id="8BFD3699FFAF745A81B43F7AFAE1FDD2" blockId="9.[151,1343,546,573]" box="[151,1343,546,573]" pageId="9" pageNumber="346">
<emphasis id="B936EA8BFFAF745A81B43F7AFEFFFDD3" bold="true" box="[151,289,546,572]" pageId="9" pageNumber="346">Description</emphasis>
. FEMALE. Body well sclerotized. Length of idiosoma 245 (190265), width 155 (105155).
</paragraph>
<caption id="DF3D6611FFAF745A81B43A5BFB25F8F4" ID-DOI="http://doi.org/10.5281/zenodo.13618629" ID-Zenodo-Dep="13618629" box="[151,1275,1795,1820]" httpUri="https://zenodo.org/record/13618629/files/figure.png" pageId="9" pageNumber="346" startId="9.[151,250,1795,1819]" targetBox="[151,1435,610,1772]" targetPageId="9" targetType="figure">
<paragraph id="8BFD3699FFAF745A81B43A5BFB25F8F4" blockId="9.[151,1275,1795,1820]" box="[151,1275,1795,1820]" pageId="9" pageNumber="346">
<emphasis id="B936EA8BFFAF745A81B43A5BFED3F8F4" bold="true" box="[151,269,1795,1819]" pageId="9" pageNumber="346">FIGURE 9</emphasis>
.
<taxonomicName id="4C424D1AFFAF745A803A3A5BFDC6F8F4" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[281,536,1795,1819]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="9" pageNumber="346" phylum="Arthropoda" rank="species" species="spinisetus" status="sp. nov.">
<emphasis id="B936EA8BFFAF745A803A3A5BFDC6F8F4" box="[281,536,1795,1819]" italics="true" pageId="9" pageNumber="346">Spatulaphorus spinisetus</emphasis>
</taxonomicName>
<emphasis id="B936EA8BFFAF745A833D3A5BFDB0F8F4" bold="true" box="[542,622,1795,1819]" pageId="9" pageNumber="346">
<taxonomicNameLabel id="A20557F0FFAF745A833D3A5BFDB0F8F4" box="[542,622,1795,1819]" pageId="9" pageNumber="346" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: A—dorsum of body; B—venter of body. Legs omitted.
</paragraph>
</caption>
<paragraph id="8BFD3699FFAF745981E43A11FCC9FF3A" blockId="9.[151,1437,1865,2036]" lastBlockId="10.[151,1436,151,213]" lastPageId="10" lastPageNumber="347" pageId="9" pageNumber="346">
<emphasis id="B936EA8BFFAF745A81E43A11FE88F88C" box="[199,342,1865,1891]" italics="true" pageId="9" pageNumber="346">Gnathosoma</emphasis>
. Gnathosomal capsule, excluding palps, subrectangular, length 21 (1922), width 22 (1922). Dorsum with one pair of needle-like cheliceral setae (
<emphasis id="B936EA8BFFAF745A823D3A35FC98F868" box="[798,838,1901,1927]" italics="true" pageId="9" pageNumber="346">cha</emphasis>
) 6 (56). Dorsal median apodeme present, poorly developed. Postpalpal setae (
<emphasis id="B936EA8BFFAF745A80FC3ACAFE25F844" box="[479,507,1938,1963]" italics="true" pageId="9" pageNumber="346">pp</emphasis>
) 2 (2) needle-like, situated laterad setae
<emphasis id="B936EA8BFFAF745A82EA3AC9FC2FF844" box="[969,1009,1937,1963]" italics="true" pageId="9" pageNumber="346">cha</emphasis>
. Ventral gnathosoma with one pair of smooth, weakly blunt-tipped subcapitular setae
<emphasis id="B936EA8BFFAF745A838D3AEEFD1CF820" box="[686,706,1974,1999]" italics="true" pageId="9" pageNumber="346">m</emphasis>
6 (67) and round pits
<emphasis id="B936EA8BFFAF745A82E93AEEFC06F820" box="[970,984,1974,1999]" italics="true" pageId="9" pageNumber="346">n</emphasis>
situated posteriad
<emphasis id="B936EA8BFFAF745A85883AEEFB61F820" box="[1195,1215,1974,1999]" italics="true" pageId="9" pageNumber="346">m</emphasis>
. Palps with smooth and weakly blunt-tipped setae
<emphasis id="B936EA8BFFAF745A80C93A81FDC8F81C" box="[490,534,2009,2035]" italics="true" pageId="9" pageNumber="346">dFe</emphasis>
4 (34) and
<emphasis id="B936EA8BFFAF745A83813A81FD0FF81C" box="[674,721,2009,2035]" italics="true" pageId="9" pageNumber="346">dGe</emphasis>
8 (78) dorsolaterally. Palps ventrally each with mushroom-like accessory setigenous structure (
<emphasis id="B936EA8BFFAC745980DA3DCFFDC3FF5F" box="[505,541,151,176]" italics="true" pageId="10" pageNumber="347">ass</emphasis>
) and tiny solenidion
<emphasis id="B936EA8BFFAC745982293DCFFCC0FF5F" box="[778,798,151,176]" italics="true" pageId="10" pageNumber="347">ω</emphasis>
. Palps terminated with small blunt-tipped tibial claw and tiny eupathid-like seta. Pharyngeal pumps poorly visible.
</paragraph>
<caption id="DF3D6611FFAC745981B43847FAEAFAD8" ID-DOI="http://doi.org/10.5281/zenodo.13618631" ID-Zenodo-Dep="13618631" box="[151,1332,1311,1335]" httpUri="https://zenodo.org/record/13618631/files/figure.png" pageId="10" pageNumber="347" startId="10.[151,250,1311,1335]" targetBox="[151,1435,241,1288]" targetPageId="10" targetType="figure">
<paragraph id="8BFD3699FFAC745981B43847FAEAFAD8" blockId="10.[151,1332,1311,1335]" box="[151,1332,1311,1335]" pageId="10" pageNumber="347">
<emphasis id="B936EA8BFFAC745981B43847FEC7FAD8" bold="true" box="[151,281,1311,1335]" pageId="10" pageNumber="347">FIGURE 10</emphasis>
.
<taxonomicName id="4C424D1AFFAC745980053847FDFAFAD8" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[294,548,1311,1335]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="10" pageNumber="347" phylum="Arthropoda" rank="species" species="spinisetus" status="sp. nov.">
<emphasis id="B936EA8BFFAC745980053847FDFAFAD8" box="[294,548,1311,1335]" italics="true" pageId="10" pageNumber="347">Spatulaphorus spinisetus</emphasis>
</taxonomicName>
<emphasis id="B936EA8BFFAC745983093847FDA5FAD8" bold="true" box="[554,635,1311,1335]" pageId="10" pageNumber="347">
<taxonomicNameLabel id="A20557F0FFAC745983093847FDA5FAD8" box="[554,635,1311,1335]" pageId="10" pageNumber="347" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: A—right leg I, dorsal aspect; B—right leg II, dorsal aspect.
</paragraph>
</caption>
<paragraph id="8BFD3699FFAC745981E4383DFC95F994" blockId="10.[151,1437,1381,1984]" pageId="10" pageNumber="347">
<emphasis id="B936EA8BFFAC745981E4383DFE48FA90" box="[199,406,1381,1407]" italics="true" pageId="10" pageNumber="347">Idiosomal dorsum</emphasis>
(
<figureCitation id="13792A1CFFAC7459808A383DFDD4FA90" box="[425,522,1381,1407]" captionStart="FIGURE 9" captionStartId="9.[151,250,1795,1819]" captionTargetBox="[151,1435,610,1772]" captionTargetId="figure-146@9.[151,1435,597,1772]" captionTargetPageId="9" captionText="FIGURE 9. Spatulaphorus spinisetus sp. nov., female: A—dorsum of body; B—venter of body. Legs omitted." figureDoi="http://doi.org/10.5281/zenodo.13618629" httpUri="https://zenodo.org/record/13618629/files/figure.png" pageId="10" pageNumber="347">Figs 9A</figureCitation>
,
<figureCitation id="13792A1CFFAC7459833A383DFD93FA90" box="[537,589,1381,1407]" captionStart="FIGURE 12" captionStartId="12.[151,250,1422,1446]" captionTargetBox="[151,1436,633,1399]" captionTargetId="figure-287@12.[151,1436,633,1399]" captionTargetPageId="12" captionText="FIGURE 12. DIC micrographs of Spatulaphorus spinisetus sp. nov., female: A—general view dorsally; B—general view ventrally." figureDoi="http://doi.org/10.5281/zenodo.13618639" httpUri="https://zenodo.org/record/13618639/files/figure.png" pageId="10" pageNumber="347">12A</figureCitation>
). All dorsal shields with big puncta. Prodorsal shield with three pairs of setae (
<emphasis id="B936EA8BFFAC745981C338D1FF24FA4C" box="[224,250,1417,1443]" italics="true" pageId="10" pageNumber="347">v1</emphasis>
,
<emphasis id="B936EA8BFFAC7459802A38D1FEFDFA4C" box="[265,291,1417,1443]" italics="true" pageId="10" pageNumber="347">v2</emphasis>
,
<emphasis id="B936EA8BFFAC7459801238D1FE89FA4C" box="[305,343,1417,1443]" italics="true" pageId="10" pageNumber="347">sc2</emphasis>
), one pair of weakly barbed capitate trichobothria (
<emphasis id="B936EA8BFFAC745982B838D1FC1FFA4C" box="[923,961,1417,1443]" italics="true" pageId="10" pageNumber="347">sc1</emphasis>
) and one pair of round stigmata. Stigmata associated with well-developed tracheal trunks.All dorsal idiosomal setae blunt-tipped; seta
<emphasis id="B936EA8BFFAC745985BA38F5FB6DFA28" box="[1177,1203,1453,1479]" italics="true" pageId="10" pageNumber="347">v2</emphasis>
smooth, other dorsal setae distinctly barbed.Alveoli of setae
<emphasis id="B936EA8BFFAC7459836D388AFD84FA04" box="[590,602,1490,1515]" italics="true" pageId="10" pageNumber="347">e</emphasis>
and
<emphasis id="B936EA8BFFAC745983AC3889FD49FA04" box="[655,663,1489,1515]" italics="true" pageId="10" pageNumber="347">f</emphasis>
almost contiguous. Cupules
<emphasis id="B936EA8BFFAC745982F6388AFC35FA04" box="[981,1003,1490,1515]" italics="true" pageId="10" pageNumber="347">ia</emphasis>
on tergite D,
<emphasis id="B936EA8BFFAC745985A3388AFB42FA04" box="[1152,1180,1490,1515]" italics="true" pageId="10" pageNumber="347">im</emphasis>
on tergite EF and
<emphasis id="B936EA8BFFAC745984463889FAA5FA04" box="[1381,1403,1489,1515]" italics="true" pageId="10" pageNumber="347">ih</emphasis>
on tergite H large, round, subequal. Lengths of dorsal setae:
<emphasis id="B936EA8BFFAC7459823138ADFCF2F9E0" box="[786,812,1525,1551]" italics="true" pageId="10" pageNumber="347">v1</emphasis>
26 (2127),
<emphasis id="B936EA8BFFAC7459829838ADFC08F9E0" box="[955,982,1525,1551]" italics="true" pageId="10" pageNumber="347">v2</emphasis>
9 (810),
<emphasis id="B936EA8BFFAC7459856B38ADFBB0F9E0" box="[1096,1134,1525,1551]" italics="true" pageId="10" pageNumber="347">sc2</emphasis>
47 (3849),
<emphasis id="B936EA8BFFAC745985DF38ADFAC9F9E0" box="[1276,1303,1525,1551]" italics="true" pageId="10" pageNumber="347">c1</emphasis>
32 (2734),
<emphasis id="B936EA8BFFAC745981B43B41FF6CF9DC" box="[151,178,1561,1587]" italics="true" pageId="10" pageNumber="347">c2</emphasis>
45 (3648),
<emphasis id="B936EA8BFFAC745980663B41FE8DF9DC" box="[325,339,1561,1587]" italics="true" pageId="10" pageNumber="347">d</emphasis>
39 (3039),
<emphasis id="B936EA8BFFAC745980C43B42FE2DF9DC" box="[487,499,1562,1587]" italics="true" pageId="10" pageNumber="347">e</emphasis>
24 (2027),
<emphasis id="B936EA8BFFAC745983A43B41FD51F9DC" box="[647,655,1561,1587]" italics="true" pageId="10" pageNumber="347">f</emphasis>
41 (3242),
<emphasis id="B936EA8BFFAC745982003B41FCE0F9DC" box="[803,830,1561,1587]" italics="true" pageId="10" pageNumber="347">h1</emphasis>
43 (3444),
<emphasis id="B936EA8BFFAC745982F03B41FC31F9DC" box="[979,1007,1561,1587]" italics="true" pageId="10" pageNumber="347">h2</emphasis>
24 (1924). Distances between setae:
<emphasis id="B936EA8BFFAC745981B43B65FF04F9B8" box="[151,218,1597,1624]" italics="true" pageId="10" pageNumber="347">v1v1</emphasis>
27 (2429),
<emphasis id="B936EA8BFFAC745980533B65FE6CF9B8" box="[368,434,1597,1623]" italics="true" pageId="10" pageNumber="347">v2v2</emphasis>
45 (4049),
<emphasis id="B936EA8BFFAC7459836A3B65FD7FF9B8" box="[585,673,1597,1623]" italics="true" pageId="10" pageNumber="347">sc2sc2</emphasis>
56 (4661),
<emphasis id="B936EA8BFFAC745982143B65FCA4F9B8" box="[823,890,1597,1624]" italics="true" pageId="10" pageNumber="347">c1c1</emphasis>
44 (3547),
<emphasis id="B936EA8BFFAC7459852D3B65FB8FF9B8" box="[1038,1105,1597,1624]" italics="true" pageId="10" pageNumber="347">c1c2</emphasis>
32 (2432),
<emphasis id="B936EA8BFFAC745985C43B65FACFF9B8" box="[1255,1297,1597,1624]" italics="true" pageId="10" pageNumber="347">dd</emphasis>
76 (6580),
<emphasis id="B936EA8BFFAC745981B43B3AFF64F994" box="[151,186,1633,1660]" italics="true" pageId="10" pageNumber="347">ef</emphasis>
5 (45),
<emphasis id="B936EA8BFFAC745980033B39FEE0F994" box="[288,318,1633,1660]" italics="true" pageId="10" pageNumber="347">ff</emphasis>
73 (5875),
<emphasis id="B936EA8BFFAC745980ED3B39FDCAF994" box="[462,532,1633,1660]" italics="true" pageId="10" pageNumber="347">h1h1</emphasis>
62 (4762),
<emphasis id="B936EA8BFFAC745983863B39FD35F994" box="[677,747,1633,1660]" italics="true" pageId="10" pageNumber="347">h1h2</emphasis>
7 (78).
</paragraph>
<paragraph id="8BFD3699FFAC745981E43BDDFE3DF82F" blockId="10.[151,1437,1381,1984]" pageId="10" pageNumber="347">
<emphasis id="B936EA8BFFAC745981E43BDDFE57F970" box="[199,393,1669,1695]" italics="true" pageId="10" pageNumber="347">Idiosomal venter</emphasis>
(
<figureCitation id="13792A1CFFAC745980BF3BDDFE22F970" box="[412,508,1669,1695]" captionStart="FIGURE 9" captionStartId="9.[151,250,1795,1819]" captionTargetBox="[151,1435,610,1772]" captionTargetId="figure-146@9.[151,1435,597,1772]" captionTargetPageId="9" captionText="FIGURE 9. Spatulaphorus spinisetus sp. nov., female: A—dorsum of body; B—venter of body. Legs omitted." figureDoi="http://doi.org/10.5281/zenodo.13618629" httpUri="https://zenodo.org/record/13618629/files/figure.png" pageId="10" pageNumber="347">Figs 9B</figureCitation>
,
<figureCitation id="13792A1CFFAC7459832E3BDDFDE1F970" box="[525,575,1669,1695]" captionStart="FIGURE 12" captionStartId="12.[151,250,1422,1446]" captionTargetBox="[151,1436,633,1399]" captionTargetId="figure-287@12.[151,1436,633,1399]" captionTargetPageId="12" captionText="FIGURE 12. DIC micrographs of Spatulaphorus spinisetus sp. nov., female: A—general view dorsally; B—general view ventrally." figureDoi="http://doi.org/10.5281/zenodo.13618639" httpUri="https://zenodo.org/record/13618639/files/figure.png" pageId="10" pageNumber="347">12B</figureCitation>
). Coxisternal fields I-II, III-IV and aggenital plate laterally with sparsely distributed big puncta. Setae
<emphasis id="B936EA8BFFAC745980C53BF1FDDCF92C" box="[486,514,1705,1731]" italics="true" pageId="10" pageNumber="347">1b</emphasis>
pointed; other ventral setae blunt-tipped; setae
<emphasis id="B936EA8BFFAC745985063BF1FB9CF92C" box="[1061,1090,1705,1731]" italics="true" pageId="10" pageNumber="347">1b</emphasis>
and
<emphasis id="B936EA8BFFAC7459855F3BF1FB7DF92C" box="[1148,1187,1705,1731]" italics="true" pageId="10" pageNumber="347">ps3</emphasis>
weakly barbed, other ventral setae smooth. Bases of setae
<emphasis id="B936EA8BFFAC745983053B95FD93F908" box="[550,589,1741,1767]" italics="true" pageId="10" pageNumber="347">ps3</emphasis>
situated distinctly anteriad bases of setae
<emphasis id="B936EA8BFFAC745985353B95FB82F908" box="[1046,1116,1741,1767]" italics="true" pageId="10" pageNumber="347">ps1-2.</emphasis>
Ap1 and ap2 well-developed and joined with thick appr; apsej developed only laterally; ap3 weak, not reaching appo; ap4 well developed, exceeding beyond bases of setae
<emphasis id="B936EA8BFFAC745983323A4DFDF3F8C0" box="[529,557,1813,1839]" italics="true" pageId="10" pageNumber="347">3b</emphasis>
. Ap5 short, joined with appo and reaching bases of setae
<emphasis id="B936EA8BFFAC7459859C3A4DFB02F8C0" box="[1215,1244,1813,1839]" italics="true" pageId="10" pageNumber="347">4a</emphasis>
. Ags small, cup-like; pgs small, oval. Posterior margin of posterior sternal plate straight in middle part. Posterior margin of aggenital plate weakly concave. Lengths of ventral setae:
<emphasis id="B936EA8BFFAC745983983A05FD09F898" box="[699,727,1885,1911]" italics="true" pageId="10" pageNumber="347">1a</emphasis>
9 (810),
<emphasis id="B936EA8BFFAC7459826C3A05FCB2F898" box="[847,876,1885,1911]" italics="true" pageId="10" pageNumber="347">1b</emphasis>
20 (1722),
<emphasis id="B936EA8BFFAC745985233A05FBC4F898" box="[1024,1050,1885,1911]" italics="true" pageId="10" pageNumber="347">1с</emphasis>
11 (1012),
<emphasis id="B936EA8BFFAC7459858D3A05FB14F898" box="[1198,1226,1885,1911]" italics="true" pageId="10" pageNumber="347">2a</emphasis>
12 (1114),
<emphasis id="B936EA8BFFAC7459847E3A05FAA9F898" box="[1373,1399,1885,1911]" italics="true" pageId="10" pageNumber="347">2с</emphasis>
13 (1113),
<emphasis id="B936EA8BFFAC745981DE3AD9FEC7F874" box="[253,281,1921,1947]" italics="true" pageId="10" pageNumber="347">3a</emphasis>
14 (1114),
<emphasis id="B936EA8BFFAC7459808A3AD9FE1BF874" box="[425,453,1921,1947]" italics="true" pageId="10" pageNumber="347">3b</emphasis>
12 (1013),
<emphasis id="B936EA8BFFAC745983753AD9FDAEF874" box="[598,624,1921,1947]" italics="true" pageId="10" pageNumber="347">3c</emphasis>
16 (1216),
<emphasis id="B936EA8BFFAC745982213AD9FCC0F874" box="[770,798,1921,1947]" italics="true" pageId="10" pageNumber="347">4a</emphasis>
12 (1012),
<emphasis id="B936EA8BFFAC7459828C3AD9FC15F874" box="[943,971,1921,1947]" italics="true" pageId="10" pageNumber="347">4b</emphasis>
19 (1520),
<emphasis id="B936EA8BFFAC7459857F3AD9FBA8F874" box="[1116,1142,1921,1947]" italics="true" pageId="10" pageNumber="347">4c</emphasis>
14 (1216),
<emphasis id="B936EA8BFFAC745984243AD9FAF0F874" box="[1287,1326,1921,1947]" italics="true" pageId="10" pageNumber="347">ps1</emphasis>
9 (710),
<emphasis id="B936EA8BFFAC745981B43AFDFF60F850" box="[151,190,1957,1983]" italics="true" pageId="10" pageNumber="347">ps2</emphasis>
9 (810),
<emphasis id="B936EA8BFFAC745980103AFDFE84F850" box="[307,346,1957,1983]" italics="true" pageId="10" pageNumber="347">ps3</emphasis>
19 (1519).
</paragraph>
<caption id="DF3D6611FFAD745881B43A1CFA94F8B3" ID-DOI="http://doi.org/10.5281/zenodo.13618637" ID-Zenodo-Dep="13618637" box="[151,1354,1860,1884]" httpUri="https://zenodo.org/record/13618637/files/figure.png" pageId="11" pageNumber="348" startId="11.[151,250,1860,1884]" targetBox="[151,1418,190,1836]" targetPageId="11" targetType="figure">
<paragraph id="8BFD3699FFAD745881B43A1CFA94F8B3" blockId="11.[151,1354,1860,1884]" box="[151,1354,1860,1884]" pageId="11" pageNumber="348">
<emphasis id="B936EA8BFFAD745881B43A1CFEC6F8B3" bold="true" box="[151,280,1860,1884]" pageId="11" pageNumber="348">FIGURE 11</emphasis>
.
<taxonomicName id="4C424D1AFFAD745880063A1CFDFDF8B3" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[293,547,1860,1884]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="11" pageNumber="348" phylum="Arthropoda" rank="species" species="spinisetus" status="sp. nov.">
<emphasis id="B936EA8BFFAD745880063A1CFDFDF8B3" box="[293,547,1860,1884]" italics="true" pageId="11" pageNumber="348">Spatulaphorus spinisetus</emphasis>
</taxonomicName>
<emphasis id="B936EA8BFFAD7458830A3A1CFDA4F8B3" bold="true" box="[553,634,1860,1884]" pageId="11" pageNumber="348">
<taxonomicNameLabel id="A20557F0FFAD7458830A3A1CFDA4F8B3" box="[553,634,1860,1884]" pageId="11" pageNumber="348" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: A—right leg III, dorsal aspect; B—right leg IV, dorsal aspect.
</paragraph>
</caption>
<paragraph id="8BFD3699FFAD745F81E43AD3FF23FD8E" blockId="11.[151,1437,1930,2029]" lastBlockId="12.[151,1437,151,609]" lastPageId="12" lastPageNumber="349" pageId="11" pageNumber="348">
<emphasis id="B936EA8BFFAD745881E43AD3FF22F84B" box="[199,252,1931,1956]" italics="true" pageId="11" pageNumber="348">Legs</emphasis>
(
<figureCitation id="13792A1CFFAD7458802D3AD2FEB9F84B" box="[270,359,1930,1956]" captionStart="FIGURE 10" captionStartId="10.[151,250,1311,1335]" captionTargetBox="[151,1435,241,1288]" captionTargetId="figure-50@10.[151,1435,237,1288]" captionTargetPageId="10" captionText="FIGURE 10. Spatulaphorus spinisetus sp. nov., female: A—right leg I, dorsal aspect; B—right leg II, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.13618631" httpUri="https://zenodo.org/record/13618631/files/figure.png" pageId="11" pageNumber="348">Figs 10</figureCitation>
,
<figureCitation id="13792A1CFFAD745880553AD2FE4DF84B" box="[374,403,1930,1956]" captionStart="FIGURE 11" captionStartId="11.[151,250,1860,1884]" captionTargetBox="[151,1418,190,1836]" captionTargetId="figure-17@11.[151,1436,181,1836]" captionTargetPageId="11" captionText="FIGURE 11. Spatulaphorus spinisetus sp. nov., female: A—right leg III, dorsal aspect; B—right leg IV, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.13618637" httpUri="https://zenodo.org/record/13618637/files/figure.png" pageId="11" pageNumber="348">11</figureCitation>
). Setation of legs as in
<taxonomicName id="4C424D1AFFAD7458838B3AD3FC8EF84B" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[680,848,1930,1957]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="11" pageNumber="348" phylum="Arthropoda" rank="species" species="brevisetosus">
<emphasis id="B936EA8BFFAD7458838B3AD3FC8EF84B" box="[680,848,1930,1957]" italics="true" pageId="11" pageNumber="348">S. brevisetosus</emphasis>
</taxonomicName>
. Leg I (
<figureCitation id="13792A1CFFAD745882923AD2FBC2F84B" box="[945,1052,1930,1956]" captionStart="FIGURE 10" captionStartId="10.[151,250,1311,1335]" captionTargetBox="[151,1435,241,1288]" captionTargetId="figure-50@10.[151,1435,237,1288]" captionTargetPageId="10" captionText="FIGURE 10. Spatulaphorus spinisetus sp. nov., female: A—right leg I, dorsal aspect; B—right leg II, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.13618631" httpUri="https://zenodo.org/record/13618631/files/figure.png" pageId="11" pageNumber="348">Fig. 10A</figureCitation>
). Tibiotarsus enlarged, with large claw; tibiotarsus internally with oval strongly sclerotized structure near solenidion
<emphasis id="B936EA8BFFAD745885623AF6FB81F827" box="[1089,1119,1966,1992]" italics="true" pageId="11" pageNumber="348">φ2</emphasis>
. Setae (
<emphasis id="B936EA8BFFAD7458859F3AF7FB14F827" box="[1212,1226,1967,1992]" italics="true" pageId="11" pageNumber="348">u</emphasis>
) consolidated into a horn-like structure opposing claw. Tarsal claw very thick and blunt-tipped. Seta
<emphasis id="B936EA8BFFAD7458851B3A8AFB98F803" box="[1080,1094,2002,2028]" italics="true" pageId="11" pageNumber="348">d</emphasis>
of femur spatulate, with short subterminal projection. Setae
<emphasis id="B936EA8BFFAA745F80C43DCFFE26FF5E" box="[487,504,151,177]" italics="true" pageId="12" pageNumber="349">l</emphasis>
of femur and
<emphasis id="B936EA8BFFAA745F83BB3DCFFD7AFF5E" box="[664,676,151,177]" italics="true" pageId="12" pageNumber="349">k</emphasis>
of tibiotarsus smooth, other leg setae (except eupathidia
<emphasis id="B936EA8BFFAA745F84053DCFFAE3FF5E" box="[1318,1341,151,177]" italics="true" pageId="12" pageNumber="349">p</emphasis>
,
<emphasis id="B936EA8BFFAA745F846F3DCFFAB7FF5E" box="[1356,1385,151,177]" italics="true" pageId="12" pageNumber="349">p”</emphasis>
,
<emphasis id="B936EA8BFFAA745F845B3DCFFA48FF5E" box="[1400,1430,151,177]" italics="true" pageId="12" pageNumber="349">tc</emphasis>
,
<emphasis id="B936EA8BFFAA745F81B43DE3FF65FF3A" box="[151,187,187,213]" italics="true" pageId="12" pageNumber="349">tc”</emphasis>
,
<emphasis id="B936EA8BFFAA745F81EB3DE3FF3FFF3A" box="[200,225,187,213]" italics="true" pageId="12" pageNumber="349">ft</emphasis>
,
<emphasis id="B936EA8BFFAA745F81CE3DE3FED3FF3A" box="[237,269,187,213]" italics="true" pageId="12" pageNumber="349">ft”</emphasis>
) sparsely barbed. Setae
<emphasis id="B936EA8BFFAA745F833A3DE3FDF4FF3A" box="[537,554,187,213]" italics="true" pageId="12" pageNumber="349">l</emphasis>
,
<emphasis id="B936EA8BFFAA745F83143DE3FD90FF3A" box="[567,590,187,213]" italics="true" pageId="12" pageNumber="349">l”</emphasis>
of femur,
<emphasis id="B936EA8BFFAA745F83E33DE3FD08FF3A" box="[704,726,187,213]" italics="true" pageId="12" pageNumber="349">l”</emphasis>
of genu and
<emphasis id="B936EA8BFFAA745F82473DE3FCAEFF3A" box="[868,880,187,213]" italics="true" pageId="12" pageNumber="349">k</emphasis>
of tibiotarsus blunt-tipped; other leg setae (except eupathidia) pointed. Lengths of solenidia
<emphasis id="B936EA8BFFAA745F834E3D87FD50FF16" box="[621,654,223,249]" italics="true" pageId="12" pageNumber="349">ω1</emphasis>
6 (67),
<emphasis id="B936EA8BFFAA745F83D43D87FCC7FF16" box="[759,793,223,249]" italics="true" pageId="12" pageNumber="349">ω2</emphasis>
5 (5),
<emphasis id="B936EA8BFFAA745F82463D87FC5CFF16" box="[869,898,223,249]" italics="true" pageId="12" pageNumber="349">φ1</emphasis>
7 (67),
<emphasis id="B936EA8BFFAA745F82C83D87FBD7FF16" box="[1003,1033,223,249]" italics="true" pageId="12" pageNumber="349">φ2</emphasis>
5 (45); solenidia
<emphasis id="B936EA8BFFAA745F85FC3D87FADEFF16" box="[1247,1280,223,249]" italics="true" pageId="12" pageNumber="349">ω1</emphasis>
and
<emphasis id="B936EA8BFFAA745F841B3D87FA8BFF16" box="[1336,1365,223,249]" italics="true" pageId="12" pageNumber="349">φ1</emphasis>
thick, clavate; other solenidia weakly clavate. Leg II (
<figureCitation id="13792A1CFFAA745F83873C5BFCD7FEF2" box="[676,777,259,285]" captionStart="FIGURE 10" captionStartId="10.[151,250,1311,1335]" captionTargetBox="[151,1435,241,1288]" captionTargetId="figure-50@10.[151,1435,237,1288]" captionTargetPageId="10" captionText="FIGURE 10. Spatulaphorus spinisetus sp. nov., female: A—right leg I, dorsal aspect; B—right leg II, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.13618631" httpUri="https://zenodo.org/record/13618631/files/figure.png" pageId="12" pageNumber="349">Fig. 10B</figureCitation>
). Tarsus with thickened basally claws and pad-like empodium. Solenidion
<emphasis id="B936EA8BFFAA745F806E3C7FFEBFFEAF" box="[333,353,295,320]" italics="true" pageId="12" pageNumber="349">ω</emphasis>
6 (56) weakly clavate, solenidion
<emphasis id="B936EA8BFFAA745F83DD3C7FFCD3FEAF" box="[766,781,295,320]" italics="true" pageId="12" pageNumber="349">φ</emphasis>
absent. Setae
<emphasis id="B936EA8BFFAA745F82973C7FFC04FEAE" box="[948,986,295,321]" italics="true" pageId="12" pageNumber="349">pl”</emphasis>
,
<emphasis id="B936EA8BFFAA745F82CA3C7FFBD9FEAE" box="[1001,1031,295,321]" italics="true" pageId="12" pageNumber="349">tc</emphasis>
,
<emphasis id="B936EA8BFFAA745F85343C7FFBE4FEAE" box="[1047,1082,295,321]" italics="true" pageId="12" pageNumber="349">tc”</emphasis>
and
<emphasis id="B936EA8BFFAA745F85563C7FFB52FEAE" box="[1141,1164,295,321]" italics="true" pageId="12" pageNumber="349">u</emphasis>
of tarsus smooth, other setae barbed; setae
<emphasis id="B936EA8BFFAA745F804C3C13FE53FE8A" box="[367,397,331,357]" italics="true" pageId="12" pageNumber="349">tc</emphasis>
and
<emphasis id="B936EA8BFFAA745F80E03C13FE36FE8A" box="[451,488,331,357]" italics="true" pageId="12" pageNumber="349">pl”</emphasis>
of tarsus spiniform, blunt-tipped; setae
<emphasis id="B936EA8BFFAA745F82863C13FC6DFE8A" box="[933,947,331,357]" italics="true" pageId="12" pageNumber="349">d</emphasis>
,
<emphasis id="B936EA8BFFAA745F82E23C13FC0CFE8A" box="[961,978,331,357]" italics="true" pageId="12" pageNumber="349">l</emphasis>
, and
<emphasis id="B936EA8BFFAA745F852C3C13FBF4FE8A" box="[1039,1066,331,357]" italics="true" pageId="12" pageNumber="349">v”</emphasis>
of femur blunt-tipped; setae (
<emphasis id="B936EA8BFFAA745F845B3C13FA4CFE8B" box="[1400,1426,331,356]" italics="true" pageId="12" pageNumber="349">pv</emphasis>
) of tarsus weakly foliate distally; other leg setae pointed. Leg III (
<figureCitation id="13792A1CFFAA745F824B3C37FC11FE66" box="[872,975,367,393]" captionStart="FIGURE 11" captionStartId="11.[151,250,1860,1884]" captionTargetBox="[151,1418,190,1836]" captionTargetId="figure-17@11.[151,1436,181,1836]" captionTargetPageId="11" captionText="FIGURE 11. Spatulaphorus spinisetus sp. nov., female: A—right leg III, dorsal aspect; B—right leg IV, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.13618637" httpUri="https://zenodo.org/record/13618637/files/figure.png" pageId="12" pageNumber="349">Fig. 11A</figureCitation>
). Femur divided into basi- and telofemur. Claws and empodium of same shape as on tarsus II. Setae
<emphasis id="B936EA8BFFAA745F82133CCBFC9DFE42" box="[816,835,403,429]" italics="true" pageId="12" pageNumber="349">v</emphasis>
of trochanter,
<emphasis id="B936EA8BFFAA745F82CA3CCBFBD9FE42" box="[1001,1031,403,429]" italics="true" pageId="12" pageNumber="349">tc</emphasis>
,
<emphasis id="B936EA8BFFAA745F85363CCBFBE5FE42" box="[1045,1083,403,429]" italics="true" pageId="12" pageNumber="349">pl”</emphasis>
, and
<emphasis id="B936EA8BFFAA745F85593CCBFB4FFE42" box="[1146,1169,403,429]" italics="true" pageId="12" pageNumber="349">u</emphasis>
of tarsus smooth, other setae barbed; setae
<emphasis id="B936EA8BFFAA745F804C3CEFFE53FE3E" box="[367,397,439,465]" italics="true" pageId="12" pageNumber="349">tc</emphasis>
and
<emphasis id="B936EA8BFFAA745F80E13CEFFE36FE3E" box="[450,488,439,465]" italics="true" pageId="12" pageNumber="349">pl”</emphasis>
of tarsus spiniform; setae
<emphasis id="B936EA8BFFAA745F822E3CEFFCFDFE3E" box="[781,803,439,465]" italics="true" pageId="12" pageNumber="349">v</emphasis>
of trochanter,
<emphasis id="B936EA8BFFAA745F82E53CEFFC0AFE3E" box="[966,980,439,465]" italics="true" pageId="12" pageNumber="349">d</emphasis>
,
<emphasis id="B936EA8BFFAA745F82C23CEFFC29FE3E" box="[993,1015,439,465]" italics="true" pageId="12" pageNumber="349">v</emphasis>
of femur, and
<emphasis id="B936EA8BFFAA745F85B93CEFFB75FE3E" box="[1178,1195,439,465]" italics="true" pageId="12" pageNumber="349">l</emphasis>
of genu blunt-tipped, other leg setae pointed; solenidion
<emphasis id="B936EA8BFFAA745F830D3C83FDE3FE1B" box="[558,573,475,500]" italics="true" pageId="12" pageNumber="349">φ</emphasis>
absent. Leg IV (
<figureCitation id="13792A1CFFAA745F82253C83FCB6FE1A" box="[774,872,475,501]" captionStart="FIGURE 11" captionStartId="11.[151,250,1860,1884]" captionTargetBox="[151,1418,190,1836]" captionTargetId="figure-17@11.[151,1436,181,1836]" captionTargetPageId="11" captionText="FIGURE 11. Spatulaphorus spinisetus sp. nov., female: A—right leg III, dorsal aspect; B—right leg IV, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.13618637" httpUri="https://zenodo.org/record/13618637/files/figure.png" pageId="12" pageNumber="349">Fig 11B</figureCitation>
). Femur divided into basi- and telofemur. Claws simple, empodium as on tarsi II and III. Setae
<emphasis id="B936EA8BFFAA745F83B83CA7FD6FFDF6" box="[667,689,511,537]" italics="true" pageId="12" pageNumber="349">v</emphasis>
of trochanter and
<emphasis id="B936EA8BFFAA745F825F3CA7FC7CFDF6" box="[892,930,511,537]" italics="true" pageId="12" pageNumber="349">pl”</emphasis>
of tarsus smooth, other setae barbed; setae
<emphasis id="B936EA8BFFAA745F84A53CA7FA42FDF6" box="[1414,1436,511,537]" italics="true" pageId="12" pageNumber="349">v</emphasis>
of trochanter,
<emphasis id="B936EA8BFFAA745F80113F7BFE9EFDD2" box="[306,320,547,573]" italics="true" pageId="12" pageNumber="349">d</emphasis>
,
<emphasis id="B936EA8BFFAA745F806E3F7BFEBDFDD2" box="[333,355,547,573]" italics="true" pageId="12" pageNumber="349">v</emphasis>
of femur,
<emphasis id="B936EA8BFFAA745F80F63F7BFE2FFDD2" box="[469,497,547,573]" italics="true" pageId="12" pageNumber="349">v”</emphasis>
of tibia blunt-tipped; seta
<emphasis id="B936EA8BFFAA745F82363F7BFCE5FDD2" box="[789,827,547,573]" italics="true" pageId="12" pageNumber="349">pl”</emphasis>
of tarsus spiniform, other leg setae pointed; solenidion
<emphasis id="B936EA8BFFAA745F81B43F1FFF78FD8F" box="[151,166,583,608]" italics="true" pageId="12" pageNumber="349">φ</emphasis>
absent.
</paragraph>
<caption id="DF3D6611FFAA745F81B438D6FF29FA25" ID-DOI="http://doi.org/10.5281/zenodo.13618639" ID-Zenodo-Dep="13618639" httpUri="https://zenodo.org/record/13618639/files/figure.png" pageId="12" pageNumber="349" startId="12.[151,250,1422,1446]" targetBox="[151,1436,633,1399]" targetPageId="12" targetType="figure">
<paragraph id="8BFD3699FFAA745F81B438D6FF29FA25" blockId="12.[151,1436,1422,1482]" pageId="12" pageNumber="349">
<emphasis id="B936EA8BFFAA745F81B438D6FEC3FA49" bold="true" box="[151,285,1422,1446]" pageId="12" pageNumber="349">FIGURE 12</emphasis>
. DIC micrographs of
<taxonomicName id="4C424D1AFFAA745F832938D6FCD2FA49" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[522,780,1422,1446]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="12" pageNumber="349" phylum="Arthropoda" rank="species" species="spinisetus" status="sp. nov.">
<emphasis id="B936EA8BFFAA745F832938D6FCD2FA49" box="[522,780,1422,1446]" italics="true" pageId="12" pageNumber="349">Spatulaphorus spinisetus</emphasis>
</taxonomicName>
<emphasis id="B936EA8BFFAA745F823538D7FCB4FA48" bold="true" box="[790,874,1423,1447]" pageId="12" pageNumber="349">
<taxonomicNameLabel id="A20557F0FFAA745F823538D7FCB4FA48" box="[790,874,1423,1447]" pageId="12" pageNumber="349" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: A—general view dorsally; B—general view ventrally.
</paragraph>
</caption>
<paragraph id="8BFD3699FFAA745F81EB38A1FE4FF9FC" blockId="12.[151,1436,1529,1987]" box="[200,401,1529,1555]" pageId="12" pageNumber="349">MALE unknown.</paragraph>
</subSubSection>
<subSubSection id="C3586512FFAA745F81E43B45FD9BF904" pageId="12" pageNumber="349" type="materials_examined">
<paragraph id="8BFD3699FFAA745F81E43B45FE56F990" blockId="12.[151,1436,1529,1987]" pageId="12" pageNumber="349">
<emphasis id="B936EA8BFFAA745F81E43B45FEB5F9D9" bold="true" box="[199,363,1564,1591]" pageId="12" pageNumber="349">Type material</emphasis>
.
<materialsCitation id="3B2A3CC4FFAA745F80543B45FE5FF990" collectingDate="2004-01-19" collectingDateMax="2005-01" collectingDateMin="2004-01-19" collectionCode="ZISP, CEMT" country="Brazil" location="Campos do Jordao" municipality="Female" pageId="12" pageNumber="349" specimenCount="36" specimenCount-female="36" stateProvince="Sao Paulo" typeStatus="holotype">
<collectingMunicipality id="6B99ACE3FFAA745F80543B45FE17F9D8" box="[375,457,1565,1591]" pageId="12" pageNumber="349">Female</collectingMunicipality>
<typeStatus id="54F9883BFFAA745F80EE3B45FDECF9D8" box="[461,562,1565,1591]" pageId="12" pageNumber="349" type="holotype">holotype</typeStatus>
, slide
<collectionCode id="ED53AE5CFFAA745F83573B45FD73F9D8" box="[628,685,1565,1591]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="12" pageNumber="349">ZISP</collectionCode>
T-Pygm-012,
<collectingCountry id="F3557609FFAA745F826B3B45FC51F9D8" box="[840,911,1565,1591]" name="Brazil" pageId="12" pageNumber="349">Brazil</collectingCountry>
,
<collectingRegion id="4986F87BFFAA745F82BB3B45FBD7F9D8" box="[920,1033,1565,1591]" country="Brazil" name="Sao Paulo" pageId="12" pageNumber="349">São Paulo</collectingRegion>
,
<location id="8E9D6042FFAA745F85313B45FB3EF9D8" LSID="urn:lsid:plazi:treatment:03EB878FFFAF745E81B43CEEFB9DFE1A:8E9D6042FFAA745F85313B45FB3EF9D8" box="[1042,1248,1565,1591]" country="Brazil" municipality="Female" name="Campos do Jordao" pageId="12" pageNumber="349" stateProvince="Sao Paulo">Campos do Jordão</location>
,
<date id="FFFC1059FFAA745F85CB3B44FAEFF9D8" box="[1256,1329,1564,1591]" pageId="12" pageNumber="349" value="2005-01">
<collectingDate id="EFB8E9B1FFAA745F85CB3B44FAEFF9D8" box="[1256,1329,1564,1591]" pageId="12" pageNumber="349" value="2005-01">I.2005</collectingDate>
</date>
(
<collectionCode id="ED53AE5CFFAA745F841E3B45FA4EF9D8" box="[1341,1424,1565,1591]" pageId="12" pageNumber="349">CEMT</collectionCode>
),
<collectedFrom id="385E7564FFAA745F81B43B19FDD2F9B4" box="[151,524,1601,1627]" pageId="12" pageNumber="349">
on
<taxonomicName id="4C424D1AFFAA745F819E3B19FDD2F9B4" authority="(Lucas)" baseAuthorityName="Lucas" baseAuthorityYear="1859" box="[189,524,1601,1627]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="349" phylum="Arthropoda" rank="species" species="buqueti">
<emphasis id="B936EA8BFFAA745F819E3B19FE74F9B4" box="[189,426,1601,1627]" italics="true" pageId="12" pageNumber="349">Dichotomius buqueti</emphasis>
(Lucas)
</taxonomicName>
</collectedFrom>
;
<typeStatus id="54F9883BFFAA745F83393B19FD54F9B4" box="[538,650,1601,1627]" pageId="12" pageNumber="349" type="paratype">paratypes</typeStatus>
:
<specimenCount id="9D44FD10FFAA745F83B43B19FCC9F9B4" box="[663,791,1601,1627]" count="10" pageId="12" pageNumber="349" type="female">10 females</specimenCount>
, same data;
<specimenCount id="9D44FD10FFAA745F828B3B19FBF6F9B4" box="[936,1064,1601,1627]" count="26" pageId="12" pageNumber="349" type="female">26 females</specimenCount>
, same locality but
<date id="FFFC1059FFAA745F84273B19FAAFF9B4" box="[1284,1393,1601,1627]" pageId="12" pageNumber="349" value="2004-01-19">
<collectingDate id="EFB8E9B1FFAA745F84273B19FAAFF9B4" box="[1284,1393,1601,1627]" pageId="12" pageNumber="349" value="2004-01-19">19.I.2004</collectingDate>
</date>
,
<collectedFrom id="385E7564FFAA745F84A33B19FE5FF990" pageId="12" pageNumber="349">
on
<taxonomicName id="4C424D1AFFAA745F81B43B3DFE5FF990" baseAuthorityName="Lucas" baseAuthorityYear="1859" box="[151,385,1637,1663]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="349" phylum="Arthropoda" rank="species" species="buqueti">
<emphasis id="B936EA8BFFAA745F81B43B3DFE5FF990" box="[151,385,1637,1663]" italics="true" pageId="12" pageNumber="349">Dichotomius buqueti</emphasis>
</taxonomicName>
</collectedFrom>
</materialsCitation>
.
</paragraph>
<paragraph id="8BFD3699FFAA745F81E43BD1FD9BF904" blockId="12.[151,1436,1529,1987]" pageId="12" pageNumber="349">
<emphasis id="B936EA8BFFAA745F81E43BD1FE58F94D" bold="true" box="[199,390,1672,1699]" pageId="12" pageNumber="349">Type deposition</emphasis>
.
<materialsCitation id="3B2A3CC4FFAA745F80B43BD1FE25F928" country="Russia" location="Zoological Institute" pageId="12" pageNumber="349" specimenCount="4" stateProvince="Saint Petersburg" typeStatus="holotype">
The
<typeStatus id="54F9883BFFAA745F80EF3BD1FDF0F94C" box="[460,558,1673,1699]" pageId="12" pageNumber="349" type="holotype">holotype</typeStatus>
and
<specimenCount id="9D44FD10FFAA745F83493BD1FCD3F94C" box="[618,781,1673,1699]" count="4" pageId="12" pageNumber="349" type="generic" typeStatus="paratype">four paratypes</specimenCount>
are deposited in the collection of
<location id="8E9D6042FFAA745F85BF3BD1FAA5F94C" LSID="urn:lsid:plazi:treatment:03EB878FFFAF745E81B43CEEFB9DFE1A:8E9D6042FFAA745F85BF3BD1FAA5F94C" box="[1180,1403,1673,1699]" country="Russia" name="Zoological Institute" pageId="12" pageNumber="349" stateProvince="Saint Petersburg">Zoological Institute</location>
of RAS,
<collectingRegion id="4986F87BFFAA745F81FC3BF5FE40F928" box="[223,414,1709,1735]" country="Russia" name="Saint Petersburg" pageId="12" pageNumber="349">Saint Petersburg</collectingRegion>
,
<collectingCountry id="F3557609FFAA745F808E3BF5FE25F928" box="[429,507,1709,1735]" name="Russia" pageId="12" pageNumber="349">Russia</collectingCountry>
</materialsCitation>
;
<materialsCitation id="3B2A3CC4FFAA745F83293BF5FAC5F928" box="[522,1307,1709,1735]" country="Russia" location="Tyumen State" pageId="12" pageNumber="349" specimenCount="1" stateProvince="Tyumen'" typeStatus="paratype">
other
<typeStatus id="54F9883BFFAA745F836D3BF5FD67F928" box="[590,697,1709,1735]" pageId="12" pageNumber="349" type="paratype">paratypes</typeStatus>
are deposited in the collection of the
<collectingRegion id="4986F87BFFAA745F855E3BF5FAC5F928" box="[1149,1307,1709,1735]" country="Russia" name="Tyumen'" pageId="12" pageNumber="349">Tyumen State</collectingRegion>
</materialsCitation>
<materialsCitation id="3B2A3CC4FFAA745F84053BF5FD9EF904" country="Russia" location="University Museum of Zoology" pageId="12" pageNumber="349" specimenCount="1" stateProvince="Tyumen'" typeStatus="paratype">
<location id="8E9D6042FFAA745F84053BF5FEA1F904" LSID="urn:lsid:plazi:treatment:03EB878FFFAF745E81B43CEEFB9DFE1A:8E9D6042FFAA745F84053BF5FEA1F904" country="Russia" name="University Museum of Zoology" pageId="12" pageNumber="349" stateProvince="Tyumen'">University Museum of Zoology</location>
,
<collectingRegion id="4986F87BFFAA745F80AA3B89FE39F904" box="[393,487,1745,1771]" country="Russia" name="Tyumen'" pageId="12" pageNumber="349">Tyumen</collectingRegion>
,
<collectingCountry id="F3557609FFAA745F80D13B89FD9EF904" box="[498,576,1745,1771]" name="Russia" pageId="12" pageNumber="349">Russia</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3586512FFAA745F81E43BACFE26F82C" pageId="12" pageNumber="349" type="diagnosis">
<paragraph id="8BFD3699FFAA745F81E43BACFE26F82C" blockId="12.[151,1436,1529,1987]" pageId="12" pageNumber="349">
<emphasis id="B936EA8BFFAA745F81E43BACFE15F8E1" bold="true" box="[199,459,1780,1806]" pageId="12" pageNumber="349">Differential diagnosis</emphasis>
. The new species is very similar to
<taxonomicName id="4C424D1AFFAA745F82A73BADFADAF8E0" authority="(Mahunka, 1980)" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" baseAuthorityName="Mahunka" baseAuthorityYear="1980" box="[900,1284,1781,1807]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="12" pageNumber="349" phylum="Arthropoda" rank="species" species="imbricatipes" status="comb. nov.">
<emphasis id="B936EA8BFFAA745F82A73BADFC4CF8E0" box="[900,914,1781,1807]" italics="true" pageId="12" pageNumber="349">S</emphasis>
.
<emphasis id="B936EA8BFFAA745F82863BADFBEFF8E0" box="[933,1073,1781,1807]" italics="true" pageId="12" pageNumber="349">imbricatipes</emphasis>
(
<bibRefCitation id="EFD34B68FFAA745F85673BADFB22F8E0" author="Mahunka, S." box="[1092,1276,1781,1807]" pageId="12" pageNumber="349" pagination="377 - 399" refId="ref16707" refString="Mahunka, S. (1980) Data to the knowledge of mites preserved in the &quot; Berlese collection &quot; (Acari: Tarsonemida, Oribatida). I. Acta Zoologica Academiae Scientiarum Hungaricae, 26, 377 - 399." type="journal article" year="1980">Mahunka, 1980</bibRefCitation>
)
</taxonomicName>
<emphasis id="B936EA8BFFAA745F84333BADFA4BF8E0" bold="true" box="[1296,1429,1781,1807]" pageId="12" pageNumber="349">
<taxonomicNameLabel id="A20557F0FFAA745F84333BADFA4BF8E0" box="[1296,1429,1781,1807]" pageId="12" pageNumber="349" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
, described from
<collectingCountry id="F3557609FFAA745F80663A41FE6AF8DC" box="[325,436,1817,1843]" name="Argentina" pageId="12" pageNumber="349">Argentina</collectingCountry>
(
<bibRefCitation id="EFD34B68FFAA745F80E13A41FDB0F8DC" author="Mahunka, S." box="[450,622,1817,1843]" pageId="12" pageNumber="349" pagination="377 - 399" refId="ref16707" refString="Mahunka, S. (1980) Data to the knowledge of mites preserved in the &quot; Berlese collection &quot; (Acari: Tarsonemida, Oribatida). I. Acta Zoologica Academiae Scientiarum Hungaricae, 26, 377 - 399." type="journal article" year="1980">Mahunka 1980</bibRefCitation>
) in having spiniform setae
<emphasis id="B936EA8BFFAA745F82BD3A41FC31F8DC" box="[926,1007,1817,1843]" italics="true" pageId="12" pageNumber="349">pl”, tc</emphasis>
on tarsi II and III, enlarged tibiotarsus I and basally thickened claws on tarsi II and III. The new species differs from
<taxonomicName id="4C424D1AFFAA745F85323A65FB64F8B8" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" baseAuthorityName="Mahunka" baseAuthorityYear="1980" box="[1041,1210,1853,1879]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="12" pageNumber="349" phylum="Arthropoda" rank="species" species="imbricatipes">
<emphasis id="B936EA8BFFAA745F85323A65FBC1F8B8" box="[1041,1055,1853,1879]" italics="true" pageId="12" pageNumber="349">S</emphasis>
.
<emphasis id="B936EA8BFFAA745F850D3A65FB64F8B8" box="[1070,1210,1853,1879]" italics="true" pageId="12" pageNumber="349">imbricatipes</emphasis>
</taxonomicName>
in having distinctly longer dorsal idiosomal setae, especially
<emphasis id="B936EA8BFFAA745F834E3A39FD57F894" box="[621,649,1889,1915]" italics="true" pageId="12" pageNumber="349">h1</emphasis>
which are longer than the distance between their bases (vs. setae
<emphasis id="B936EA8BFFAA745F84A33A39FA42F894" box="[1408,1436,1889,1915]" italics="true" pageId="12" pageNumber="349">h1</emphasis>
shorter than distance between their bases in
<taxonomicName id="4C424D1AFFAA745F83A03ADDFCF4F870" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" baseAuthorityName="Mahunka" baseAuthorityYear="1980" box="[643,810,1925,1951]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="12" pageNumber="349" phylum="Arthropoda" rank="species" species="imbricatipes">
<emphasis id="B936EA8BFFAA745F83A03ADDFCF4F870" box="[643,810,1925,1951]" italics="true" pageId="12" pageNumber="349">S. imbricatipes</emphasis>
</taxonomicName>
) and setae
<emphasis id="B936EA8BFFAA745F828B3ADDFC1DF870" box="[936,963,1925,1951]" italics="true" pageId="12" pageNumber="349">c1</emphasis>
distinctly shorter than
<emphasis id="B936EA8BFFAA745F85E03ADDFB00F870" box="[1219,1246,1925,1951]" italics="true" pageId="12" pageNumber="349">c2</emphasis>
(vs. setae
<emphasis id="B936EA8BFFAA745F84703ADDFAB0F870" box="[1363,1390,1925,1951]" italics="true" pageId="12" pageNumber="349">c1</emphasis>
and
<emphasis id="B936EA8BFFAA745F81B43AF1FF6CF82C" box="[151,178,1961,1987]" italics="true" pageId="12" pageNumber="349">c2</emphasis>
subequal in
<taxonomicName id="4C424D1AFFAA745F80633AF1FE36F82C" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" baseAuthorityName="Mahunka" baseAuthorityYear="1980" box="[320,488,1961,1987]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="12" pageNumber="349" phylum="Arthropoda" rank="species" species="imbricatipes">
<emphasis id="B936EA8BFFAA745F80633AF1FE36F82C" box="[320,488,1961,1987]" italics="true" pageId="12" pageNumber="349">S. imbricatipes</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C3586512FFAB745E81E43DCFFC60FF3A" pageId="13" pageNumber="350" type="etymology">
<paragraph id="8BFD3699FFAB745E81E43DCFFC60FF3A" blockId="13.[151,1437,151,501]" pageId="13" pageNumber="350">
<emphasis id="B936EA8BFFAB745E81E43DCFFE96FF5E" bold="true" box="[199,328,151,177]" pageId="13" pageNumber="350">Etymology</emphasis>
. The name of the new species is a combination of two Latin words:
<emphasis id="B936EA8BFFAB745E85613DCFFB7CFF5F" box="[1090,1186,151,176]" italics="true" pageId="13" pageNumber="350">spinosus</emphasis>
meaning
<emphasis id="B936EA8BFFAB745E84333DCFFAB8FF5F" box="[1296,1382,151,176]" italics="true" pageId="13" pageNumber="350">spinous</emphasis>
, and
<emphasis id="B936EA8BFFAB745E81B43DE3FF1AFF3B" box="[151,196,187,212]" italics="true" pageId="13" pageNumber="350">seta</emphasis>
meaning
<emphasis id="B936EA8BFFAB745E80103DE3FEA5FF3A" box="[307,379,187,213]" italics="true" pageId="13" pageNumber="350">bristle</emphasis>
, and refers to the spiniform setae on tarsi II and III.
</paragraph>
</subSubSection>
<subSubSection id="C3586512FFAB745E81E43D87FB9DFE1A" pageId="13" pageNumber="350" type="discussion">
<paragraph id="8BFD3699FFAB745E81E43D87FB9DFE1A" blockId="13.[151,1437,151,501]" pageId="13" pageNumber="350">
<emphasis id="B936EA8BFFAB745E81E43D87FEEEFF16" bold="true" box="[199,304,223,249]" pageId="13" pageNumber="350">Remark.</emphasis>
<bibRefCitation id="EFD34B68FFAB745E80153D87FE2EFF16" author="Mahunka, S." box="[310,496,223,249]" pageId="13" pageNumber="350" pagination="377 - 399" refId="ref16707" refString="Mahunka, S. (1980) Data to the knowledge of mites preserved in the &quot; Berlese collection &quot; (Acari: Tarsonemida, Oribatida). I. Acta Zoologica Academiae Scientiarum Hungaricae, 26, 377 - 399." type="journal article" year="1980">Mahunka (1980)</bibRefCitation>
described
<emphasis id="B936EA8BFFAB745E83453D87FC73FF16" box="[614,941,223,249]" italics="true" pageId="13" pageNumber="350">
<taxonomicName id="4C424D1AFFAB745E83453D87FCC5FF16" box="[614,795,223,249]" genus="Pygmephorellus" pageId="13" pageNumber="367" rank="genus">Pygmephorellus</taxonomicName>
<taxonomicName id="4C424D1AFFAB745E82023D87FC73FF16" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" baseAuthorityName="Mahunka" baseAuthorityYear="1980" box="[801,941,223,249]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="13" pageNumber="350" phylum="Arthropoda" rank="species" species="imbricatipes">imbricatipes</taxonomicName>
</emphasis>
based on specimens deposited in the Berlese collection and collected on
<taxonomicName id="4C424D1AFFAB745E80FB3C5BFDAEFEF2" box="[472,624,259,285]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="350" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B936EA8BFFAB745E80FB3C5BFD98FEF2" box="[472,582,259,285]" italics="true" pageId="13" pageNumber="350">Phanaeus</emphasis>
sp.
</taxonomicName>
in
<collectingCountry id="F3557609FFAB745E83BA3C5BFCD2FEF2" box="[665,780,259,285]" name="Argentina" pageId="13" pageNumber="350">Argentina</collectingCountry>
. In the description he did not describe the chaetotaxy of legs. However, illustrations of legs IIV show character states typical for
<taxonomicName id="4C424D1AFFAB745E82C23C7FFB5AFEAE" authorityName="Rack" authorityYear="1993" box="[993,1156,295,321]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="13" pageNumber="350" phylum="Arthropoda" rank="genus">
<emphasis id="B936EA8BFFAB745E82C23C7FFB5AFEAE" box="[993,1156,295,321]" italics="true" pageId="13" pageNumber="350">Spatulaphorus</emphasis>
</taxonomicName>
, namely the presence of only two setae on genu I, only one seta on genu II and genu IV without setae. Also, setae
<emphasis id="B936EA8BFFAB745E85A73C13FB75FE8A" box="[1156,1195,331,357]" italics="true" pageId="13" pageNumber="350">ps3</emphasis>
are located distinctly anteriad setae
<emphasis id="B936EA8BFFAB745E80143C37FEA4FE66" box="[311,378,367,393]" italics="true" pageId="13" pageNumber="350">ps12</emphasis>
which is characteristic for most Neotropical
<taxonomicName id="4C424D1AFFAB745E82503C37FBABFE66" box="[883,1141,367,393]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" kingdom="Animalia" order="Trombidiformes" pageId="13" pageNumber="350" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B936EA8BFFAB745E82503C37FBC8FE66" box="[883,1046,367,393]" italics="true" pageId="13" pageNumber="350">Spatulaphorus</emphasis>
species.
</taxonomicName>
<bibRefCitation id="EFD34B68FFAB745E855F3C37FA42FE66" author="Khaustov, A. A. &amp; Trach, V. A." box="[1148,1436,367,393]" pageId="13" pageNumber="350" pagination="480 - 485" refId="ref16479" refString="Khaustov, A. A. &amp; Trach, V. A. (2012) A new species and new records of mites of the genus Spatulaphorus Rack (Acari: Heterostigmata: Pygmephoridae) from Ukraine. International Journal of Acarology, 38, 480 - 485. https: // doi. org / 10.1080 / 01647954.2012.677475" type="journal article" year="2012">Khaustov &amp; Trach (2012)</bibRefCitation>
and
<bibRefCitation id="EFD34B68FFAB745E81E53CCBFE35FE42" author="Khaustov, A. A. &amp; Frolov, A. V." box="[198,491,403,429]" pageId="13" pageNumber="350" pagination="501 - 521" refId="ref16128" refString="Khaustov, A. A. &amp; Frolov, A. V. (2017) New species of heterostigmatic mites (Acari: Heterostigmata: Athyreacaridae, Dolichocybidae, Pygmephoridae) associated with scarab beetles (Coleoptera: Geotrupidae, Scarabaeidae) from Brazil. Zootaxa, 4294 (5), 501 - 521. https: // doi. org / 10.11646 / zootaxa. 4294.5.1" type="journal article" year="2017">Khaustov &amp; Frolov (2017</bibRefCitation>
;
<bibRefCitation id="EFD34B68FFAB745E80D53CCBFDF1FE42" author="Khaustov, A. A. &amp; Frolov, A. V." box="[502,559,403,429]" pageId="13" pageNumber="350" pagination="333 - 345" refId="ref16337" refString="Khaustov, A. A. &amp; Frolov, A. V. (2021) Two new species and a new record of Spatulaphorus (Acari &quot; Pygmephoridae) phoretic on scarab beetles (Coleoptera: Scarabaeidae) from French Guiana. Acarina, 29 (2), 333 - 345. https: // doi. org / 10.21684 / 0132 - 8077 - 2021 - 29 - 2 - 233 - 245" type="journal article" year="2021">2021</bibRefCitation>
) suggested that
<taxonomicName id="4C424D1AFFAB745E83C73CCBFC55FE42" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" baseAuthorityName="Mahunka" baseAuthorityYear="1980" box="[740,907,403,429]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="13" pageNumber="350" phylum="Arthropoda" rank="species" species="imbricatipes">
<emphasis id="B936EA8BFFAB745E83C73CCBFC55FE42" box="[740,907,403,429]" italics="true" pageId="13" pageNumber="350">P. imbricatipes</emphasis>
</taxonomicName>
could be a species of the genus
<taxonomicName id="4C424D1AFFAB745E85D13CCBFA4BFE42" authorityName="Rack" authorityYear="1993" box="[1266,1429,403,429]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="13" pageNumber="350" phylum="Arthropoda" rank="genus">
<emphasis id="B936EA8BFFAB745E85D13CCBFA4BFE42" box="[1266,1429,403,429]" italics="true" pageId="13" pageNumber="350">Spatulaphorus</emphasis>
</taxonomicName>
. Discovery of the closely related species
<taxonomicName id="4C424D1AFFAB745E83483CEFFC58FE3F" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[619,902,439,465]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="13" pageNumber="350" phylum="Arthropoda" rank="species" species="spinisetus" status="sp. nov.">
<emphasis id="B936EA8BFFAB745E83483CEFFC58FE3F" box="[619,902,439,465]" italics="true" pageId="13" pageNumber="350">Spatulaphorus spinisetus</emphasis>
</taxonomicName>
<emphasis id="B936EA8BFFAB745E82B23CEFFC33FE3E" bold="true" box="[913,1005,439,465]" pageId="13" pageNumber="350">
<taxonomicNameLabel id="A20557F0FFAB745E82B23CEFFC33FE3E" box="[913,1005,439,465]" pageId="13" pageNumber="350" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
confirms this opinion. Therefore, we move
<emphasis id="B936EA8BFFAB745E81FF3C83FDF9FE1A" box="[220,551,475,501]" italics="true" pageId="13" pageNumber="350">
<taxonomicName id="4C424D1AFFAB745E81FF3C83FE4FFE1A" box="[220,401,475,501]" genus="Pygmephorellus" pageId="13" pageNumber="367" rank="genus">Pygmephorellus</taxonomicName>
imbricaripes
</emphasis>
to the genus
<taxonomicName id="4C424D1AFFAB745E839E3C83FCBEFE1A" authorityName="Rack" authorityYear="1993" box="[701,864,475,501]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="13" pageNumber="350" phylum="Arthropoda" rank="genus">
<emphasis id="B936EA8BFFAB745E839E3C83FCBEFE1A" box="[701,864,475,501]" italics="true" pageId="13" pageNumber="350">Spatulaphorus</emphasis>
</taxonomicName>
(new combination).
</paragraph>
</subSubSection>
</treatment>
</document>